The first proandric species of Tritogenia Kinberg, 1867 from Botswana, Tritogenia talana sp. n. (Clitellata, Crassiclitellata, Tritogeniidae)
Author
Nxele, Thembeka Clara
Author
Plisko, Jadwiga Danuta
Author
Ramongalo, Boipelo K.
text
Zootaxa
2018
2018-10-05
4496
1
214
217
journal article
29267
10.11646/zootaxa.4496.1.17
a161220f-7db6-4ccc-bfaa-de528dab984c
1175-5326
1446761
68B7C32E-BDEB-4C7C-A4F8-BC242DE24351
Genus
Tritogenia
Kinberg, 1867
Brachydrilus
:
Benham 1888
: 72
;
Michaelsen 1900
: 462
;
Michaelsen 1918
: 332
;
Stephenson 1930
: 900
;
Reynolds & Cook 1976
: 77
.
Microchaetus
:
Michaelsen 1902
: 33
;
Michaelsen 1907
: 6
;
Michaelsen 1913
: 431
;
Reynolds & Cook 1976
: 77
, 192.
Type species.
Tritogenia sulcata
Kinberg, 1867
.
Diagnosis.
One oesophageal gizzard in 6–7. Four or five septa variably thickened, with thickened septa 6/7 attached externally to the gizzard. Dorsal blood vessel double in segments 4–10, 11, 12; double when crossing septa; single after segment 10, 11 or 12. Two or more small nephridia per segment (may be missing in some posterior segments). Testes and their funnels in the species earlier accredited to this genus in holandric arrangement, in presently described new species these are proandric. Description based on
Plisko (2013)
.
Description.
Annulation of pre-clitellar segments present; 1–3 usually simple, 4–9, 10 ringleted, with 2–4 ringlets annulated or not, with an external segmental subdivision; first ringlet may be longer than second. Postclitellar segments may be externally subdivided, or not. Setae: minute and irregularly placed; on preclitellar segments, may be recognizable under high magnification (120–250×) and only as a few pairs of
ab
or
cd
; sometimes
ab
seen only on a few segments. As from segment 9,
ab
may be associated with papillae. On postclitellar segments, setae easier to notice, although variably placed; in four pairs located mostly ventrally, with
ab
and
cd
paired very closely, or distantly, with
aa
2–3 times as
bc
;
dd
nearly 0.6–0.7 of U. Occasionally
cd
not noticeable, when
ab
easily noted. Papillae and genital glands variably shaped, with segments on/in which they occur varying, occasionally associated with genital setae. Male pores occur variably in the area of tubercula pubertatis, although the pores are difficult to detect externally. Spermathecae small, present or absent, often deeply embedded in body tissue; variably shaped, paired or multiple; in the majority of species they are close to septa
11/ 12 and 12/13
, although in some species may be located in more segments, sometimes up to six.