182 lines
19 KiB
XML
182 lines
19 KiB
XML
<document id="1B1DB1B233D2FACCA1170A520CD7473F" ID-CLB-Dataset="26449" ID-DOI="10.11646/zootaxa.4619.2.8" ID-GBIF-Dataset="aa446d35-b5ef-4885-8bb8-8ddcc1f2e2da" ID-ISSN="1175-5326" ID-Zenodo-Dep="3249661" ID-ZooBank="643CEA27-5DBD-4EE6-B02B-B42E1AC0168F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1560917014283" checkinUser="plazi" docAuthor="Calcetas, Orlando A." docDate="2019" docId="620587D9B434941EFF2AFA52FEA85E69" docLanguage="en" docName="zootaxa.4619.2.8.pdf" docOrigin="Zootaxa 4619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carlschoenherria adoradae Calcetas 2019, new species" docType="treatment" docUuid="526C8351-687D-4131-A4CF-30A218183145" docUuidSource="ZooBank" docVersion="16" lastPageNumber="333" masterDocId="9E3CFFA1B435941DFFBDFFB4FFC95D7E" masterDocTitle="Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines" masterLastPageNumber="346" masterPageNumber="330" pageNumber="331" updateTime="1698735450490" updateUser="ExternalLinkService">
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<mods:mods id="1D70296665D9C19F12FFA88C020B5599" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2E93E6840EA884BFCF38B3DD2D8871DF">
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<mods:title id="22C11230F9A62CC1BC012DE130FF83CF">Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name id="D4129E7796E089BC6D8CD5A7EE3A7A41" type="personal">
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<mods:roleTerm id="D24EDA768675AEEE71EC753F40E913EB">Author</mods:roleTerm>
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<mods:namePart id="84EA4113A061658588D4E0C5E472A303">Calcetas, Orlando A.</mods:namePart>
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<mods:title id="C7A511A9C2E7EC51DB51C5AFF517BC9D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B79A3718BCE0A6B903474060360F694C">
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<mods:date id="7E6BEB1551A402A6F82D5C9280C923BF">2019</mods:date>
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<mods:number id="5CAF97D354EC7539B80B891AE697377D">2019-06-19</mods:number>
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<mods:number id="C795E871CB7070FA6A272F0DF11E11A5">4619</mods:number>
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<mods:detail id="B79A028335D2A75EF8234CE4B678EFFA" type="issue">
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<mods:classification id="30E1F8D27247CD53E532CF5783B657AC">journal article</mods:classification>
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<mods:identifier id="BB9AAE9FBF9741AABF451BC659A2B5BA" type="CLB-Dataset">26449</mods:identifier>
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<mods:identifier id="4707B3AA773578222AEF862640D19972" type="DOI">10.11646/zootaxa.4619.2.8</mods:identifier>
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<treatment id="620587D9B434941EFF2AFA52FEA85E69" ID-GBIF-Taxon="158297467" LSID="urn:lsid:zoobank.org:act:526C8351-687D-4131-A4CF-30A218183145" httpUri="http://treatment.plazi.org/id/620587D9B434941EFF2AFA52FEA85E69" lastPageId="3" lastPageNumber="333" pageId="1" pageNumber="331">
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<subSubSection id="A2B66544B434941CFF2AFA52FEF55B58" pageId="1" pageNumber="331" type="nomenclature">
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<paragraph id="EA1336CFB434941CFF2AFA52FD3D5B7F" blockId="1.[151,756,1510,1574]" box="[151,756,1510,1537]" pageId="1" pageNumber="331">
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<heading id="B15B81A3B434941CFF2AFA52FD3D5B7F" bold="true" box="[151,756,1510,1537]" fontSize="11" level="1" pageId="1" pageNumber="331" reason="1">
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<emphasis id="D8D8EADDB434941CFF2AFA52FD3D5B7F" bold="true" box="[151,756,1510,1537]" pageId="1" pageNumber="331">
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<taxonomicName id="2DAC4D4CB434941CFF2AFA52FD915B7F" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[151,600,1510,1537]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="adoradae" status="sp. nov.">
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<emphasis id="D8D8EADDB434941CFF2AFA52FE2D5B7F" bold="true" box="[151,484,1510,1537]" italics="true" pageId="1" pageNumber="331">Carlschoenherria adoradae</emphasis>
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Calcetas
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</taxonomicName>
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,
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<taxonomicNameLabel id="C3EB57A6B434941CFDD9FA53FD3D5B7F" box="[612,756,1511,1537]" pageId="1" pageNumber="331" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EA1336CFB434941CFF2AF9BFFEF55B58" blockId="1.[151,756,1510,1574]" box="[151,316,1547,1574]" pageId="1" pageNumber="331">(Figs. 1–4, 36)</paragraph>
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</subSubSection>
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<subSubSection id="A2B66544B434941CFF2AF9E7FAB15BEF" pageId="1" pageNumber="331" type="materials_examined">
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<paragraph id="EA1336CFB434941CFF2AF9E7FC915B13" blockId="1.[151,1436,1618,2041]" box="[151,856,1618,1645]" pageId="1" pageNumber="331">
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<materialsCitation id="5AC43C92B434941CFF2AF9E7FC9D5B13" ID-GBIF-Occurrence="2268317642" box="[151,852,1618,1645]" country="Philippines" location="Luzon Island" municipality="The" pageId="1" pageNumber="331" specimenCount="1" stateProvince="Kalinga Apayao" typeStatus="holotype">
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<emphasis id="D8D8EADDB434941CFF2AF9E7FEFF5B13" bold="true" box="[151,310,1618,1645]" pageId="1" pageNumber="331">
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<typeStatus id="3517886DB434941CFF2AF9E7FF1B5B13" box="[151,210,1619,1645]" pageId="1" pageNumber="331">Type</typeStatus>
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locality.
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</emphasis>
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||
<collectingMunicipality id="0A77ACB5B434941CFE81F9E7FEA15B13" box="[316,360,1619,1645]" pageId="1" pageNumber="331">The</collectingMunicipality>
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<collectingCountry id="92BB765FB434941CFED3F9E7FE265B13" box="[366,495,1619,1645]" name="Philippines" pageId="1" pageNumber="331">Philippines</collectingCountry>
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||
,
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||
<location id="EF736014B434941CFE47F9E7FD595B13" LSID="urn:lsid:plazi:treatment:620587D9B434941EFF2AFA52FEA85E69:EF736014B434941CFE47F9E7FD595B13" box="[506,656,1619,1645]" country="Philippines" municipality="The" name="Luzon Island" pageId="1" pageNumber="331" stateProvince="Kalinga Apayao">Luzon Island</location>
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,
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<collectingRegion id="2868F82DB434941CFD26F9E7FC9D5B13" box="[667,852,1619,1645]" isEnumeration="true" pageId="1" pageNumber="331">
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<collectingRegion id="2868F82DB434941CFD26F9E7FD3D5B13" box="[667,756,1619,1645]" country="Philippines" name="Kalinga" pageId="1" pageNumber="331">Kalinga</collectingRegion>
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<collectingRegion id="2868F82DB434941CFD44F9E7FC9D5B13" box="[761,852,1619,1645]" country="Philippines" name="Apayao" pageId="1" pageNumber="331">Apayao</collectingRegion>
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</collectingRegion>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="EA1336CFB434941CFF7AF9C3FAB15BEF" blockId="1.[151,1436,1618,2041]" box="[199,1400,1654,1682]" pageId="1" pageNumber="331">
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<emphasis id="D8D8EADDB434941CFF7AF9C3FEBC5BEF" bold="true" box="[199,373,1654,1681]" pageId="1" pageNumber="331">Type material.</emphasis>
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<materialsCitation id="5AC43C92B434941CFEC1F9C3FAB15BEF" ID-GBIF-Occurrence="2268317644" box="[380,1400,1654,1682]" collectingDate="1952-05-15" collectionCode="MNH" collectorName="S. M. Cendana" elevation="91" location="Bayog" pageId="1" pageNumber="331" specimenCount="1" stateProvince="Apayao" typeStatus="holotype">
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<typeStatus id="3517886DB434941CFEC1F9C3FE2D5BEF" box="[380,484,1655,1681]" pageId="1" pageNumber="331" type="holotype">Holotype</typeStatus>
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(male), labeled:
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<collectingRegion id="2868F82DB434941CFD1DF9C3FD335BEF" box="[672,762,1655,1681]" country="Philippines" name="Apayao" pageId="1" pageNumber="331">Apayao</collectingRegion>
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,
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<location id="EF736014B434941CFCB8F9C3FC985BEF" LSID="urn:lsid:plazi:treatment:620587D9B434941EFF2AFA52FEA85E69:EF736014B434941CFCB8F9C3FC985BEF" box="[773,849,1655,1681]" name="Bayog" pageId="1" pageNumber="331" stateProvince="Apayao">Bayog</location>
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, Elev.
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<elevation id="6181D1FCB434941CFC21F9C3FC2E5BEF" box="[924,999,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="1" pageNumber="331" unit="ft" value="300.0">
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<quantity id="2D549B2AB434941CFC21F9C3FC2E5BEF" box="[924,999,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="1" pageNumber="331" unit="ft" value="300.0">300 ft.</quantity>
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</elevation>
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,
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<collectingDate id="8E56E9E7B434941CFC4EF9C2FBA35BEF" box="[1011,1130,1654,1681]" pageId="1" pageNumber="331" value="1952-05-15">
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<date id="9E12100FB434941CFC4EF9C2FBA35BEF" box="[1011,1130,1654,1681]" pageId="1" pageNumber="331" value="1952-05-15">15-v-1952</date>
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</collectingDate>
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,
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<collectorName id="47595319B434941CFBCBF9C3FADC5BEF" box="[1142,1301,1655,1681]" pageId="1" pageNumber="331">S.M. Cendana</collectorName>
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(
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<collectionCode id="8CBDAE0AB434941CFA99F9C3FAA55BEF" box="[1316,1388,1655,1681]" country="Hungary" httpUri="http://grbio.org/cool/34nf-ps25" name="Musei Nacionalis Hungarici" pageId="1" pageNumber="331">MNH</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2B66544B434941EFF7AF92EFD375CAD" lastPageId="3" lastPageNumber="333" pageId="1" pageNumber="331" type="description">
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||
<paragraph id="EA1336CFB434941CFF7AF92EFAAB5B83" blockId="1.[151,1436,1618,2041]" pageId="1" pageNumber="331">
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||
<emphasis id="D8D8EADDB434941CFF7AF92EFDE15BCB" bold="true" box="[199,552,1690,1717]" pageId="1" pageNumber="331">
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||
Description of
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<typeStatus id="3517886DB434941CFEC5F92FFE295BCB" box="[376,480,1691,1717]" pageId="1" pageNumber="331" type="holotype">holotype</typeStatus>
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, male
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</emphasis>
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. Length 20.5 mm, width 8.5 mm. Body dichromatic, head, pronotum and scu- tellum dark reddish brown; uniformly covered with short, brownish to whitish setae; elytron light reddish brown, covered with short brownish to yellowish setae, with few short whitish acicular-like scales; leg black (Fig. 1).
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</paragraph>
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<paragraph id="EA1336CFB434941CFF7AF8B3FCD05A87" blockId="1.[151,1436,1618,2041]" pageId="1" pageNumber="331">
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<emphasis id="D8D8EADDB434941CFF7AF8B3FEC25A5F" box="[199,267,1799,1825]" italics="true" pageId="1" pageNumber="331">Head.</emphasis>
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Mesal carina starting at junction of fronto-clypeal suture, terminating far behind vertex. Anterior margin of clypeus straight dorsally; surface subrugosely punctate anteriorly; with very narrow, smooth margin, impunctate, non-setose area adjacent to anterior margin; dorsal surface of clypeus sloping, deeply depressed adjacent to anterior margin. Antennomere III with short stalk, widely expanded apically. Lamellae length 2.3 mm. Terminal maxillary palpomere with shallow nearly flattened lanceolate shaped groove or depression, as long as antennomere III. Men- tum with distinct roof-like mesal carina, slanted on each side, with deep depression below it; surface subrugosely punctate, covered with few setae and acicular-like scales.
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</paragraph>
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<paragraph id="EA1336CFB436941EFF7AFF2DFA455D85" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
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<emphasis id="D8D8EADDB436941EFF7AFF2DFE055DCD" box="[199,460,153,179]" italics="true" pageId="3" pageNumber="333">Ventral side of thorax.</emphasis>
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Prosternal process rectangulate, elongate, slender or narrow longitudinally, truncate apically. Prosternum and mesoventrum densely covered with long yellowish hairs. Metaventral process little prolonged, not reaching procoxae, sub-apically attached to mesoventral process; metaventral process length 1.5 mm.
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</paragraph>
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||
<paragraph id="EA1336CFB436941EFF7AFEB1FB725C19" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
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||
<emphasis id="D8D8EADDB436941EFF7AFEB1FE655C61" box="[199,428,261,287]" italics="true" pageId="3" pageNumber="333">Abdominal ventrites</emphasis>
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not uniformly covered with setation, bare at least near each margin or rather narrowly restricted in distribution of setation; posterior part of abdominal ventrites II–IV narrow, smooth, bare; sides of II–V with large and small ovoid shaped patches of yellowish-white scales or abdominal maculation.
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</paragraph>
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<paragraph id="EA1336CFB436941EFF7AFEC5FC1D5CF5" blockId="3.[151,1437,153,791]" box="[199,980,369,395]" pageId="3" pageNumber="333">
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<emphasis id="D8D8EADDB436941EFF7AFEC5FEFB5CF5" box="[199,306,369,395]" italics="true" pageId="3" pageNumber="333">Pygidium</emphasis>
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with anterior margin nearly straight; posterior margin wide.
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</paragraph>
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<paragraph id="EA1336CFB436941EFF7AFE21FD375CAD" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
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<emphasis id="D8D8EADDB436941EFF7AFE21FEFD5CD1" box="[199,308,405,431]" italics="true" pageId="3" pageNumber="333">Aedeagus</emphasis>
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. Posterior margin of parameres ribbon-like, longitudinally undulated towards apex. Length of param- eres and phallobase combined 6.4 mm; as in Figs. 3, 4.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2B66544B436941EFF7AFE68FD495C89" box="[199,640,476,503]" pageId="3" pageNumber="333" type="distribution">
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||
<paragraph id="EA1336CFB436941EFF7AFE68FD495C89" blockId="3.[151,1437,153,791]" box="[199,640,476,503]" pageId="3" pageNumber="333">
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||
<emphasis id="D8D8EADDB436941EFF7AFE68FEA85C89" bold="true" box="[199,353,476,503]" pageId="3" pageNumber="333">Distribution.</emphasis>
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The
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<collectingCountry id="92BB765FB436941EFE24FE69FDDE5C89" box="[409,535,477,503]" name="Philippines" pageId="3" pageNumber="333">Philippines</collectingCountry>
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(Luzon).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2B66544B436941EFF7AFDB5FB005F41" pageId="3" pageNumber="333" type="etymology">
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||
<paragraph id="EA1336CFB436941EFF7AFDB5FB005F41" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
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<emphasis id="D8D8EADDB436941EFF7AFDB5FE875F65" bold="true" box="[199,334,513,539]" pageId="3" pageNumber="333">Etymology.</emphasis>
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This new species is named after Dr. Jessamyn R. Adorada, professor of taxonomy and systematics at the University of the
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<collectingCountry id="92BB765FB436941EFE1DFD91FDE85F41" box="[416,545,549,575]" name="Philippines" pageId="3" pageNumber="333">Philippines</collectingCountry>
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, Los Baños. She is also an expert on Philippine coccinellids.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A2B66544B436941EFF7AFDFCFEA85E69" pageId="3" pageNumber="333" type="diagnosis">
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||
<paragraph id="EA1336CFB436941EFF7AFDFCFEA85E69" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
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||
<emphasis id="D8D8EADDB436941EFF7AFDFCFE195F1D" bold="true" box="[199,464,584,611]" pageId="3" pageNumber="333">Differential diagnosis.</emphasis>
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||
Based on the species-group classification of
|
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<bibRefCitation id="8E3D4B3EB436941EFC61FDFDFB965F1D" author="Itoh, T." box="[988,1119,585,611]" pageId="3" pageNumber="333" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
|
||
, this species belongs to the
|
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<taxonomicName id="2DAC4D4CB436941EFF2AFDD9FEF05FF9" authorityName="Laporte" authorityYear="1840" box="[151,313,621,647]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="sulcipennis">
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<emphasis id="D8D8EADDB436941EFF2AFDD9FEF05FF9" box="[151,313,621,647]" italics="true" pageId="3" pageNumber="333">C. sulcipennis</emphasis>
|
||
</taxonomicName>
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group wherein the parameres are laterally compressed and bird-bill shaped at apex. However, in contrast to other members of
|
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<taxonomicName id="2DAC4D4CB436941EFE58FD25FD4C5FD5" authorityName="Laporte" authorityYear="1840" box="[485,645,657,683]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="sulcipennis">
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<emphasis id="D8D8EADDB436941EFE58FD25FD4C5FD5" box="[485,645,657,683]" italics="true" pageId="3" pageNumber="333">C. sulcipennis</emphasis>
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</taxonomicName>
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group, the anterior margin of the parameres of
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<taxonomicName id="2DAC4D4CB436941EFB1CFD25FA515FD5" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[1185,1432,657,683]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="adoradae" status="sp. nov.">
|
||
<emphasis id="D8D8EADDB436941EFB1CFD25FAE75FD5" box="[1185,1326,657,683]" italics="true" pageId="3" pageNumber="333">C. adoradae</emphasis>
|
||
Calcetas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C3EB57A6B436941EFF2AFD01FED45FB1" box="[151,285,693,719]" pageId="3" pageNumber="333" rank="species">new species</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="2DAC4D4CB436941EFEEEFD01FD815FB1" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[339,584,693,719]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="hadsallae" status="sp. nov.">
|
||
<emphasis id="D8D8EADDB436941EFEEEFD01FE295FB1" box="[339,480,693,719]" italics="true" pageId="3" pageNumber="333">C. hadsallae</emphasis>
|
||
Calcetas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C3EB57A6B436941EFDEEFD01FD105FB1" box="[595,729,693,719]" pageId="3" pageNumber="333" rank="species">new species</taxonomicNameLabel>
|
||
are wide and laterally expanded while it’s laterally compressed and bird-bill shaped at apex for all other species. Thus, they are classified under the newly proposed
|
||
<taxonomicName id="2DAC4D4CB436941EFAB2FD6DFA555F8D" authorityName="Calcetas" authorityYear="2019" box="[1295,1436,729,755]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="adoradae">
|
||
<emphasis id="D8D8EADDB436941EFAB2FD6DFA555F8D" box="[1295,1436,729,755]" italics="true" pageId="3" pageNumber="333">C. adoradae</emphasis>
|
||
</taxonomicName>
|
||
species-subgroup.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |