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<document id="E9306B41593FF5B21D06FFB7AD763F92" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D6430650FDA6FD87C186F831" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Xystopyge bentemarieae Enghoff 2022, sp. nov." docType="treatment" docUuid="0968ADE8-6469-44B3-AE8D-D427EED0A597" docUuidSource="ZooBank" docVersion="7" lastPageNumber="104" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="103" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A7D7409851B566051616733BA045869D">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<mods:namePart id="284FB70B647779A48DA3409043471B44">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="79B4572BFFACC237CBD9E28F683B1E43">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark.</mods:affiliation>
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<mods:title id="08F5C7CC3FE91861568D85809A2FF0C4">European Journal of Taxonomy</mods:title>
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<mods:date id="62E073F9FB305F385EA4CB55134D1BB6">2022</mods:date>
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<subSubSection id="C342BEA4D6430651FDA6FD87C6F3FDA6" box="[597,991,611,638]" pageId="102" pageNumber="103" type="nomenclature">
<paragraph id="8BE7ED2FD6430651FDA6FD87C6F3FDA6" blockId="102.[361,1226,611,709]" box="[597,991,611,638]" pageId="102" pageNumber="103">
<heading id="D0AF5A43D6430651FDA6FD87C6F3FDA6" box="[597,991,611,638]" centered="true" fontSize="11" level="2" pageId="102" pageNumber="103" reason="2">
<taxonomicName id="4C5896ACD6430651FDA6FD87C651FDA6" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[597,893,611,638]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="bentemarieae" status="sp. nov.">
<emphasis id="B92C313DD6430651FDA6FD87C651FDA6" bold="true" box="[597,893,611,638]" italics="true" pageId="102" pageNumber="103">Xystopyge bentemarieae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6430651FC77FD87C6F3FDA6" box="[900,991,612,638]" pageId="102" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BE7ED2FD6430651FD33FD48C658FD1D" blockId="102.[361,1226,611,709]" box="[704,884,683,709]" pageId="102" pageNumber="103">
<figureCitation id="1363F1AAD6430651FD33FD48C632FD1D" box="[704,798,683,709]" captionStart="Fig" captionStartId="9.[191,234,1714,1740]" captionTargetBox="[189,1398,504,1672]" captionTargetId="figure-242@9.[189,1398,504,1672]" captionTargetPageId="9" captionText="Fig. 4. Telsons of odontopygids. A. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), telson and last six body rings, lateral view. B. Xystopyge bentemarieae sp. nov., paratype, ♂ (NHMD 621767), telson, lateral view, showing two setae per anal valve as characteristic of the genus Xystopyge. CD. Aquattuor udzungwensis Enghoff, 2015, telson and last body ring (from Enghoff &amp; Frederiksen 2015), showing three setae per anal valve as characteristic of most odontopygids; setae in this species placed on ravelins on raised valve margins. C. Lateral view, also showing the rectangular limbus flaps characteristic of the genus Aquattuor. D. Posterior view. Stars indicate the dorsal anal valve spines characteristic of family Odontopygidae; white arrows indicate anal valve setae. Scale bars: A=1 mm; BD=0.1 mm. AB: Photo S.G. Selvantharan." figureDoi="http://doi.org/10.5281/zenodo.6359081" httpUri="https://zenodo.org/record/6359081/files/figure.png" pageId="102" pageNumber="103">Figs 4B</figureCitation>
,
<figureCitation id="1363F1AAD6430651FCD9FD48C658FD1D" box="[810,884,683,709]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="104.[189,232,1750,1776]" captionStartId-1="105.[189,232,1857,1883]" captionStartId-2="106.[189,232,1860,1886]" captionTargetBox-0="[189,1398,287,1717]" captionTargetBox-1="[263,1324,266,1829]" captionTargetBox-2="[236,1352,265,1827]" captionTargetId-0="figure-86@104.[189,1398,287,1717]" captionTargetId-1="figure-90@105.[263,1324,266,1829]" captionTargetId-2="figure-83@106.[236,1352,265,1827]" captionTargetPageId-0="104" captionTargetPageId-1="105" captionTargetPageId-2="106" captionText-0="Fig. 62. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766). A. Gonopod sternum (sternum 8). B. Sternum 9. C. Midbody dorsal limbus. DF. First pair of legs. D. Anterior view. E. Lateral view. F. Ventral view. Abbreviations: APS=mesapical prefemoral sensilla; CXS =coxosternal setae; LPS =lateral prefemoral sensilla. Scale bars: A= 0.05 mm; B, E=0.1 mm; C=0.01 mm; D, F =0.2 mm." captionText-1="Fig. 63. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766), left gonopod coxa. A. Anterior view. B. Posterior view. C. Mesal view. D. Apical view. Abbreviations: MP=metaplica; mpo =metaplical ridged process; mpp =metaplical posterior process; mpv =metaplical vertical lobe; mtp =metaplical triangular process; PP =proplica; PPL = proplical lobe; rx =metaplical subrectangular process; sx =slender metaplical process. Scale bars: AC=0.2 mm; D =0.1 mm." captionText-2="Fig. 64. Xystopyge bentemarieae sp. nov. AD. Holotype, ♂ (NHMD 621766), left gonopod telopodite. A. Anterior view. B. Posterior view. C. Dorsal view. D. Ventral view. EH. Paratype, ♂ (NHMD 621767), isolated solenomere. Abbreviations: sdl =solenomeral distal microstriate branch; sdp =solenomeral distal spinelike branch; SLM =solenomere; th =telomeral hook; TM =telomere. Scale bars: A E= 0.1 mm; FH =0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6359251" figureDoi-1="http://doi.org/10.5281/zenodo.6359254" figureDoi-2="http://doi.org/10.5281/zenodo.6359256" httpUri-0="https://zenodo.org/record/6359251/files/figure.png" httpUri-1="https://zenodo.org/record/6359254/files/figure.png" httpUri-2="https://zenodo.org/record/6359256/files/figure.png" pageId="102" pageNumber="103">6264</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6430651FF4EFD0EC1B7FBC8" pageId="102" pageNumber="103" type="diagnosis">
<paragraph id="8BE7ED2FD6430651FF4EFD0EC41AFCDF" blockId="102.[189,310,749,775]" box="[189,310,749,775]" pageId="102" pageNumber="103">
<heading id="D0AF5A43D6430651FF4EFD0EC41AFCDF" bold="true" box="[189,310,749,775]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="B92C313DD6430651FF4EFD0EC41AFCDF" bold="true" box="[189,310,749,775]" pageId="102" pageNumber="103">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EFCFEC1B7FBC8" blockId="102.[188,1399,797,1040]" pageId="102" pageNumber="103">
Differs from other species of
<taxonomicName id="4C5896ACD6430651FDE6FCFDC7A2FCEF" authorityName="Attems" authorityYear="1909" box="[533,654,798,823]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD6430651FDE6FCFDC7A2FCEF" box="[533,654,798,823]" italics="true" pageId="102" pageNumber="103">Xystopyge</emphasis>
</taxonomicName>
by the combination of a hammerlike profile of the distal part of the gonopod coxa in certain views (
<figureCitation id="1363F1AAD6430651FD93FCA2C7E5FC84" box="[608,713,833,860]" captionStart="Fig" captionStartId="105.[189,232,1857,1883]" captionTargetBox="[263,1324,266,1829]" captionTargetId="figure-90@105.[263,1324,266,1829]" captionTargetPageId="105" captionText="Fig. 63. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766), left gonopod coxa. A. Anterior view. B. Posterior view. C. Mesal view. D. Apical view. Abbreviations: MP=metaplica; mpo =metaplical ridged process; mpp =metaplical posterior process; mpv =metaplical vertical lobe; mtp =metaplical triangular process; PP =proplica; PPL = proplical lobe; rx =metaplical subrectangular process; sx =slender metaplical process. Scale bars: AC=0.2 mm; D =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6359254" httpUri="https://zenodo.org/record/6359254/files/figure.png" pageId="102" pageNumber="103">Fig. 63C</figureCitation>
) (shared only with
<taxonomicName id="4C5896ACD6430651FC5CFCA1C5D7FCA7" authority="VandenSpiegel &amp; Pierrard, 2004" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2004" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="martella">
<emphasis id="B92C313DD6430651FC5CFCA1C11FFC83" box="[943,1075,833,859]" italics="true" pageId="102" pageNumber="103">X. martella</emphasis>
<bibRefCitation id="EFC990DED6430651FBC8FCA2C5D7FCA7" author="VandenSpiegel D. &amp; Pierrard G." pageId="102" pageNumber="103" pagination="31 - 46" refId="ref49620" refString="VandenSpiegel D. &amp; Pierrard G. 2004. Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Journal of Afrotropical Zoology 1: 31 - 46." type="journal article" year="2004">VandenSpiegel &amp; Pierrard, 2004</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5896ACD6430651FEC5FC85C4E3FCA7" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[310,463,869,895]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6430651FEC5FC85C4E3FCA7" box="[310,463,869,895]" italics="true" pageId="102" pageNumber="103">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6430651FE25FC85C71FFC58" box="[470,563,870,896]" pageId="102" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
), an only moderately long, tongue-shaped gonopod sternum (very long, slender-triangular in
<taxonomicName id="4C5896ACD6430651FE46FC69C716FC7B" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2004" box="[437,570,905,931]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="martella">
<emphasis id="B92C313DD6430651FE46FC69C716FC7B" box="[437,570,905,931]" italics="true" pageId="102" pageNumber="103">X. martella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5896ACD6430651FD86FC69C623FC7B" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[629,783,905,931]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6430651FD86FC69C623FC7B" box="[629,783,905,931]" italics="true" pageId="102" pageNumber="103">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6430651FCE5FC69C659FC7C" box="[790,885,906,932]" pageId="102" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
), a very stout, moderately curved hook (
<emphasis id="B92C313DD6430651FAA5FC6AC041FC7B" box="[1366,1389,905,931]" italics="true" pageId="102" pageNumber="103">th</emphasis>
) at midlength of the gonopod telomere (very strongly curved in
<taxonomicName id="4C5896ACD6430651FC41FC4DC161FC1F" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[946,1101,941,967]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6430651FC41FC4DC161FC1F" box="[946,1101,941,967]" italics="true" pageId="102" pageNumber="103">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6430651FBA7FC4DC19FFC10" box="[1108,1203,942,968]" pageId="102" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
), lack of a basal solenomeral spine (
<emphasis id="B92C313DD6430651FE50FC31C4FEFC34" box="[419,466,978,1004]" italics="true" pageId="102" pageNumber="103">BSS</emphasis>
) (a small tubercle in
<taxonomicName id="4C5896ACD6430651FD38FC31C663FC33" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2004" box="[715,847,977,1003]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="martella">
<emphasis id="B92C313DD6430651FD38FC31C663FC33" box="[715,847,977,1003]" italics="true" pageId="102" pageNumber="103">X. martella</emphasis>
</taxonomicName>
, a long basal spine in in
<taxonomicName id="4C5896ACD6430651FB81FC31C020FC33" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1138,1292,977,1003]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6430651FB81FC31C020FC33" box="[1138,1292,977,1003]" italics="true" pageId="102" pageNumber="103">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6430651FAE0FC31C042FC34" box="[1299,1390,978,1004]" pageId="102" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
) and a distal division of the solenomere into two branches (undivided in
<taxonomicName id="4C5896ACD6430651FBF5FC15C1A5FBD7" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2004" box="[1030,1161,1013,1039]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="martella">
<emphasis id="B92C313DD6430651FBF5FC15C1A5FBD7" box="[1030,1161,1013,1039]" italics="true" pageId="102" pageNumber="103">X. martella</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6430651FF4EFBDBC64EFB59" box="[189,866,1080,1153]" pageId="102" pageNumber="103" type="etymology">
<paragraph id="8BE7ED2FD6430651FF4EFBDBC468FB8A" blockId="102.[189,324,1080,1106]" box="[189,324,1080,1106]" pageId="102" pageNumber="103">
<heading id="D0AF5A43D6430651FF4EFBDBC468FB8A" bold="true" box="[189,324,1080,1106]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="B92C313DD6430651FF4EFBDBC468FB8A" bold="true" box="[189,324,1080,1106]" pageId="102" pageNumber="103">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EFB84C64EFB59" blockId="102.[189,866,1127,1153]" box="[189,866,1127,1153]" pageId="102" pageNumber="103">After my granddaughter Bente Marie Enghoff Mogensen.</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6430651FF4EFB4AC11EFA79" pageId="102" pageNumber="103" type="materials_examined">
<paragraph id="8BE7ED2FD6430651FF4EFB4AC76FFB1C" blockId="102.[189,579,1193,1220]" box="[189,579,1193,1220]" pageId="102" pageNumber="103">
<emphasis id="B92C313DD6430651FF4EFB4AC486FB1B" bold="true" box="[189,426,1193,1219]" pageId="102" pageNumber="103">Material examined</emphasis>
(total
<specimenCount id="9D5E26A6D6430651FE04FB49C716FB1C" box="[503,570,1194,1220]" pageId="102" pageNumber="103" type="male">7 ♂♂</specimenCount>
)
</paragraph>
<paragraph id="8BE7ED2FD6430651FF19FB3AC7EDFAE2" blockId="102.[189,1399,1241,1339]" pageId="102" pageNumber="103">
<materialsCitation id="3B30E772D6430651FF19FB3AC7EDFAE2" ID-GBIF-Occurrence="3712026312" collectedFrom="lowland rain forest" collectingDate="1984-11-22" collectingDateMax="1984-11-25" collectingDateMin="1984-11-22" collectionCode="NHMD" collectorName="N. Scharff" country="Tanzania" elevation="450" location="Kanga Mts" pageId="102" pageNumber="103" specimenCode="NHMD 621766" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
<emphasis id="B92C313DD6430651FF19FB3AC470FB2B" bold="true" box="[234,348,1241,1267]" pageId="102" pageNumber="103">
<typeStatus id="54E3538DD6430651FF19FB3AC470FB2B" box="[234,348,1241,1267]" pageId="102" pageNumber="103" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34FADBFD6430651FF4EFB1EC47EFACF" box="[189,338,1277,1303]" name="Tanzania" pageId="102" pageNumber="103">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D6430651FE95FB1EC453FACF" box="[358,383,1277,1303]" pageId="102" pageNumber="103" type="male"></specimenCount>
;
<collectingRegion id="499C23CDD6430651FE7AFB1EC772FACE" box="[393,606,1276,1303]" country="Tanzania" name="Morogoro" pageId="102" pageNumber="103">Morogoro Region</collectingRegion>
,
<location id="8E87BBF4D6430651FD94FB1EC7C5FACF" LSID="urn:lsid:plazi:treatment:03F15C39D6430650FDA6FD87C186F831:8E87BBF4D6430651FD94FB1EC7C5FACF" box="[615,745,1277,1303]" country="Tanzania" name="Kanga Mts" pageId="102" pageNumber="103" stateProvince="Morogoro Region">Kanga Mts</location>
,
<location id="8E87BBF4D6430651FD07FB1EC641FACF" LSID="urn:lsid:plazi:treatment:03F15C39D6430650FDA6FD87C186F831:8E87BBF4D6430651FD07FB1EC641FACF" box="[756,877,1277,1303]" country="Tanzania" name="Kanga FR" pageId="102" pageNumber="103" stateProvince="Morogoro Region">Kanga FR</location>
;
<elevation id="00750A1CD6430651FC84FB1FC114FACE" box="[887,1080,1276,1303]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="102" pageNumber="103" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<quantity id="4CA040CAD6430651FC84FB1FC6D6FACF" box="[887,1018,1276,1303]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="102" pageNumber="103" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE6CBEFD6430651FBB7FB1FC022FACF" box="[1092,1294,1276,1303]" pageId="102" pageNumber="103" value="1984-11-22" valueMax="1984-11-25" valueMin="1984-11-22">
<collectingDate id="EFA23207D6430651FBB7FB1FC022FACF" box="[1092,1294,1276,1303]" pageId="102" pageNumber="103" value="1984-11-22" valueMax="1984-11-25" valueMin="1984-11-22">2225 Nov. 1984</collectingDate>
</date>
;
<collectedFrom id="3844AED2D6430651FAEBFB1FC415FAE3" pageId="102" pageNumber="103">lowland rain forest</collectedFrom>
;
<collectorName id="26AD88F9D6430651FEB7FAC2C492FAE3" box="[324,446,1313,1339]" pageId="102" pageNumber="103">N. Scharff</collectorName>
leg.;
<specimenCode id="DBFE4554D6430651FDF3FAC2C791FAE2" box="[512,701,1312,1339]" collectionCode="NHMD" pageId="102" pageNumber="103">NHMD 621766</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE7ED2FD6430651FF19FA80C11EFA79" blockId="102.[189,1074,1379,1441]" pageId="102" pageNumber="103">
<materialsCitation id="3B30E772D6430651FF19FA80C102FA79" ID-GBIF-Occurrence="3712026338" collectedFrom="lowland rain forest" collectingDate="1984-11-22" collectingDateMax="1984-11-25" collectingDateMin="1984-11-22" collectionCode="NHMD" collectorName="Kanga FR &amp; N. Scharff" country="Tanzania" elevation="450" location="TANZANIA" pageId="102" pageNumber="103" specimenCode="NHMD 621767" specimenCount="6" specimenCount-male="6" stateProvince="Morogoro" typeStatus="paratype">
<emphasis id="B92C313DD6430651FF19FA80C44BFAA5" bold="true" box="[234,359,1379,1405]" pageId="102" pageNumber="103">
<typeStatus id="54E3538DD6430651FF19FA80C44BFAA5" box="[234,359,1379,1405]" pageId="102" pageNumber="103" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F34FADBFD6430651FF4EFA64C47EFA79" box="[189,338,1415,1441]" name="Tanzania" pageId="102" pageNumber="103">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D6430651FE9AFA64C482FA79" box="[361,430,1415,1441]" pageId="102" pageNumber="103" type="male">6 ♂♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBFE4554D6430651FC82FA64C102FA79" box="[881,1070,1415,1441]" collectionCode="NHMD" pageId="102" pageNumber="103">NHMD 621767</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6430650FF4EFA2AC6F1FB9E" lastPageId="103" lastPageNumber="104" pageId="102" pageNumber="103" type="description">
<paragraph id="8BE7ED2FD6430651FF4EFA2AC463FA3B" blockId="102.[189,335,1481,1507]" box="[189,335,1481,1507]" pageId="102" pageNumber="103">
<heading id="D0AF5A43D6430651FF4EFA2AC463FA3B" bold="true" box="[189,335,1481,1507]" fontSize="11" level="3" pageId="102" pageNumber="103" reason="3">
<emphasis id="B92C313DD6430651FF4EFA2AC463FA3B" bold="true" box="[189,335,1481,1507]" pageId="102" pageNumber="103">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EFA1AC05BF9CA" blockId="102.[189,1399,1528,1555]" box="[189,1399,1528,1555]" pageId="102" pageNumber="103">
SIZE. Length
<quantity id="4CA040CAD6430651FEA4FA1BC4F4F9CB" box="[343,472,1528,1555]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="7.3" metricValueMin="5.7" pageId="102" pageNumber="103" unit="mm" value="65.0" valueMax="73.0" valueMin="57.0">5773 mm</quantity>
. Diameter 3.54.0 mm. 6162 podous rings; no podous rings in fornt of telson.
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF9D8C4BBF9A1" blockId="102.[189,1399,1595,1657]" pageId="102" pageNumber="103">COLOUR. After 36 years in alcohol faded to light (greyish) brown; however, a broad mid-dorsal pale stripe still evident.</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF942C4CDF964" blockId="102.[189,481,1697,1724]" box="[189,481,1697,1724]" pageId="102" pageNumber="103">SUPRALABRAL SETAE. 56.</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF907C034F925" blockId="102.[189,1304,1763,1790]" box="[189,1304,1763,1790]" pageId="102" pageNumber="103">MANDIBULAR STIPES. Distal margin slightly concave, posterior-distal corner almost right-angled.</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF8C5C05BF898" blockId="102.[189,1399,1830,1856]" box="[189,1399,1830,1856]" pageId="102" pageNumber="103">
ANAL VALVES (
<figureCitation id="1363F1AAD6430651FE90F8C5C493F898" box="[355,447,1830,1856]" captionStart="Fig" captionStartId="9.[191,234,1714,1740]" captionTargetBox="[189,1398,504,1672]" captionTargetId="figure-242@9.[189,1398,504,1672]" captionTargetPageId="9" captionText="Fig. 4. Telsons of odontopygids. A. Spinotarsus axeli sp. nov., holotype, ♂ (NHMD 621761), telson and last six body rings, lateral view. B. Xystopyge bentemarieae sp. nov., paratype, ♂ (NHMD 621767), telson, lateral view, showing two setae per anal valve as characteristic of the genus Xystopyge. CD. Aquattuor udzungwensis Enghoff, 2015, telson and last body ring (from Enghoff &amp; Frederiksen 2015), showing three setae per anal valve as characteristic of most odontopygids; setae in this species placed on ravelins on raised valve margins. C. Lateral view, also showing the rectangular limbus flaps characteristic of the genus Aquattuor. D. Posterior view. Stars indicate the dorsal anal valve spines characteristic of family Odontopygidae; white arrows indicate anal valve setae. Scale bars: A=1 mm; BD=0.1 mm. AB: Photo S.G. Selvantharan." figureDoi="http://doi.org/10.5281/zenodo.6359081" httpUri="https://zenodo.org/record/6359081/files/figure.png" pageId="102" pageNumber="103">Fig. 4B</figureCitation>
). Each with a distinct dorsal spine and 2 setae on raised margin; no ventral spine.
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF88AC6BCF85A" blockId="102.[189,912,1896,1923]" box="[189,912,1896,1923]" pageId="102" pageNumber="103">
LIMBUS (
<figureCitation id="1363F1AAD6430651FED1F88BC4A3F85B" box="[290,399,1896,1923]" captionStart="Fig" captionStartId="104.[189,232,1750,1776]" captionTargetBox="[189,1398,287,1717]" captionTargetId="figure-86@104.[189,1398,287,1717]" captionTargetPageId="104" captionText="Fig. 62. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766). A. Gonopod sternum (sternum 8). B. Sternum 9. C. Midbody dorsal limbus. DF. First pair of legs. D. Anterior view. E. Lateral view. F. Ventral view. Abbreviations: APS=mesapical prefemoral sensilla; CXS =coxosternal setae; LPS =lateral prefemoral sensilla. Scale bars: A= 0.05 mm; B, E=0.1 mm; C=0.01 mm; D, F =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359251" httpUri="https://zenodo.org/record/6359251/files/figure.png" pageId="102" pageNumber="103">Fig. 62C</figureCitation>
). Margin with short, smooth rounded lobes.
</paragraph>
<paragraph id="8BE7ED2FD6430651FF4EF848C739F830" blockId="102.[189,1399,1962,2024]" pageId="102" pageNumber="103">LEGS. With postfemoral and tibial pads from leg-pair 5, except for last few leg-pairs; size of pads decreasing towards posterior.</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFEEAC694FEB7" blockId="103.[189,1398,265,367]" pageId="103" pageNumber="104">
FIRST PAIR OF LEGS (
<figureCitation id="1363F1AAD6420650FE5BFEEAC71BFEFC" box="[424,567,265,292]" captionStart="Fig" captionStartId="104.[189,232,1750,1776]" captionTargetBox="[189,1398,287,1717]" captionTargetId="figure-86@104.[189,1398,287,1717]" captionTargetPageId="104" captionText="Fig. 62. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766). A. Gonopod sternum (sternum 8). B. Sternum 9. C. Midbody dorsal limbus. DF. First pair of legs. D. Anterior view. E. Lateral view. F. Ventral view. Abbreviations: APS=mesapical prefemoral sensilla; CXS =coxosternal setae; LPS =lateral prefemoral sensilla. Scale bars: A= 0.05 mm; B, E=0.1 mm; C=0.01 mm; D, F =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359251" httpUri="https://zenodo.org/record/6359251/files/figure.png" pageId="103" pageNumber="104">Fig. 62DF</figureCitation>
). Prefemoral lobes short, triangular in ventral view. Six to eight long coxosternal setae (
<emphasis id="B92C313DD6420650FE6DFECCC4FEFE91" box="[414,466,303,329]" italics="true" pageId="103" pageNumber="104">CXS</emphasis>
) adjacent to lateral side of prefemoral process; prefemur with a few peglike mesapical sensilla (
<emphasis id="B92C313DD6420650FE50FEB6C4FAFEB7" box="[419,470,341,367]" italics="true" pageId="103" pageNumber="104">APS</emphasis>
) and ca 12 peglike lateral sensilla (
<emphasis id="B92C313DD6420650FC85FEB6C68BFEB7" box="[886,935,341,367]" italics="true" pageId="103" pageNumber="104">LPS</emphasis>
).
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFE7AC187FE6B" blockId="103.[189,1195,409,435]" box="[189,1195,409,435]" pageId="103" pageNumber="104">
GONOPOD STERNUM (STERNUM 8) (
<figureCitation id="1363F1AAD6420650FDC9FE7AC78FFE6B" box="[570,675,409,435]" captionStart="Fig" captionStartId="104.[189,232,1750,1776]" captionTargetBox="[189,1398,287,1717]" captionTargetId="figure-86@104.[189,1398,287,1717]" captionTargetPageId="104" captionText="Fig. 62. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766). A. Gonopod sternum (sternum 8). B. Sternum 9. C. Midbody dorsal limbus. DF. First pair of legs. D. Anterior view. E. Lateral view. F. Ventral view. Abbreviations: APS=mesapical prefemoral sensilla; CXS =coxosternal setae; LPS =lateral prefemoral sensilla. Scale bars: A= 0.05 mm; B, E=0.1 mm; C=0.01 mm; D, F =0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6359251" httpUri="https://zenodo.org/record/6359251/files/figure.png" pageId="103" pageNumber="104">Fig. 62A</figureCitation>
). Tongue-shaped, ca twice as long as broad.
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFE3EC6DFFE20" blockId="103.[189,1011,477,504]" box="[189,1011,477,504]" pageId="103" pageNumber="104">STERNUM 9. Pentagonal with parallel lateral margins (house-shaped).</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFDC2C4E6FCC6" blockId="103.[188,1399,545,799]" pageId="103" pageNumber="104">
GONOPOD COXA (
<figureCitation id="1363F1AAD6420650FE88FDC2C4FDFDE4" box="[379,465,545,572]" captionStart="Fig" captionStartId="105.[189,232,1857,1883]" captionTargetBox="[263,1324,266,1829]" captionTargetId="figure-90@105.[263,1324,266,1829]" captionTargetPageId="105" captionText="Fig. 63. Xystopyge bentemarieae sp. nov., holotype, ♂ (NHMD 621766), left gonopod coxa. A. Anterior view. B. Posterior view. C. Mesal view. D. Apical view. Abbreviations: MP=metaplica; mpo =metaplical ridged process; mpp =metaplical posterior process; mpv =metaplical vertical lobe; mtp =metaplical triangular process; PP =proplica; PPL = proplical lobe; rx =metaplical subrectangular process; sx =slender metaplical process. Scale bars: AC=0.2 mm; D =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6359254" httpUri="https://zenodo.org/record/6359254/files/figure.png" pageId="103" pageNumber="104">Fig. 63</figureCitation>
). Proplica (
<emphasis id="B92C313DD6420650FDA4FDC1C757FDE3" box="[599,635,546,571]" italics="true" pageId="103" pageNumber="104">PP</emphasis>
) simple, proplical lobe (
<emphasis id="B92C313DD6420650FC6AFDC1C6E1FDE3" box="[921,973,546,571]" italics="true" pageId="103" pageNumber="104">PPL</emphasis>
) relatively slender, clearly visible in anterior view. Metaplica (
<emphasis id="B92C313DD6420650FE1DFDABC735FDB9" box="[494,537,584,609]" italics="true" pageId="103" pageNumber="104">MP</emphasis>
) at level of proplical lobe produced mesad into triangular ridged process (
<emphasis id="B92C313DD6420650FF34FD8EC5D6FD5E" box="[199,250,621,646]" italics="true" pageId="103" pageNumber="104">mpo</emphasis>
), at lateral end of
<emphasis id="B92C313DD6420650FE2BFD8EC727FD5E" box="[472,523,621,646]" italics="true" pageId="103" pageNumber="104">mpo</emphasis>
a large vertical lobe (
<emphasis id="B92C313DD6420650FCE3FD8EC66DFD5E" box="[784,833,621,646]" italics="true" pageId="103" pageNumber="104">mpv</emphasis>
), distally produced into complex structure with four subhorizontal processes: 1. A large, subrectangular meso-posteriad process (
<emphasis id="B92C313DD6420650FB54FD70C1F6FD74" box="[1191,1242,659,684]" italics="true" pageId="103" pageNumber="104">mpp</emphasis>
), 2. A long, triangular mesad process (
<emphasis id="B92C313DD6420650FDF0FD5AC703FD0A" box="[515,559,697,722]" italics="true" pageId="103" pageNumber="104">mtp</emphasis>
), 3. A very slender process (
<emphasis id="B92C313DD6420650FC52FD5AC696FD0A" box="[929,954,697,722]" italics="true" pageId="103" pageNumber="104">sx</emphasis>
) process originating next to
<emphasis id="B92C313DD6420650FAD4FD5AC07FFD0A" box="[1319,1363,697,722]" italics="true" pageId="103" pageNumber="104">mtp</emphasis>
4. A subrectangular anteriad process (
<emphasis id="B92C313DD6420650FD90FD3CC750FD20" box="[611,636,735,760]" italics="true" pageId="103" pageNumber="104">rx</emphasis>
) (
<emphasis id="B92C313DD6420650FD6BFD3CC79DFD20" box="[664,689,735,760]" italics="true" pageId="103" pageNumber="104">sx</emphasis>
and
<emphasis id="B92C313DD6420650FD1DFD3CC62BFD20" box="[750,775,735,760]" italics="true" pageId="103" pageNumber="104">rx</emphasis>
corresponding to the similarly labelled processes in
<taxonomicName id="4C5896ACD6420650FF4CFCE6C476FCC6" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[191,346,772,798]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6420650FF4CFCE6C476FCC6" box="[191,346,772,798]" italics="true" pageId="103" pageNumber="104">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6420650FEACFCE6C497FCC7" box="[351,443,773,799]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFCABC6F1FB9E" blockId="103.[188,1399,840,1094]" pageId="103" pageNumber="104">
GONOPOD TELOPODITE (
<figureCitation id="1363F1AAD6420650FE34FCABC70EFCBB" box="[455,546,840,867]" captionStart="Fig" captionStartId="106.[189,232,1860,1886]" captionTargetBox="[236,1352,265,1827]" captionTargetId="figure-83@106.[236,1352,265,1827]" captionTargetPageId="106" captionText="Fig. 64. Xystopyge bentemarieae sp. nov. AD. Holotype, ♂ (NHMD 621766), left gonopod telopodite. A. Anterior view. B. Posterior view. C. Dorsal view. D. Ventral view. EH. Paratype, ♂ (NHMD 621767), isolated solenomere. Abbreviations: sdl =solenomeral distal microstriate branch; sdp =solenomeral distal spinelike branch; SLM =solenomere; th =telomeral hook; TM =telomere. Scale bars: A E= 0.1 mm; FH =0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6359256" httpUri="https://zenodo.org/record/6359256/files/figure.png" pageId="103" pageNumber="104">Fig. 64</figureCitation>
). Arculus 135°. Torsotope not very well delimited. Solenomere (
<emphasis id="B92C313DD6420650FAC6FCAAC041FCBB" box="[1333,1389,841,867]" italics="true" pageId="103" pageNumber="104">SLM</emphasis>
) at rest concealed within gutter formed by telomere; ca as long as telomere, simple, without a basal spine, towards end divided into a long, striate branch (
<emphasis id="B92C313DD6420650FCCFFC77C673FC76" box="[828,863,916,942]" italics="true" pageId="103" pageNumber="104">sdl</emphasis>
) with the opening of the efferent canal, and a longer, slender, smooth, curved spinelike branch (
<emphasis id="B92C313DD6420650FCFAFC59C61EFC0C" box="[777,818,954,980]" italics="true" pageId="103" pageNumber="104">sdp</emphasis>
), the latter with a short basal accessory spinelike branch. Telomere (
<emphasis id="B92C313DD6420650FE6AFC03C4EEFC21" box="[409,450,992,1017]" italics="true" pageId="103" pageNumber="104">TM</emphasis>
) overall consisting of a ribbon describing a full circle and at the same time folded lengthwise forming a concavity along the inner side of the circle, with a very stout, smooth hook (
<emphasis id="B92C313DD6420650FAA5FBE6C041FBC7" box="[1366,1389,1029,1055]" italics="true" pageId="103" pageNumber="104">th</emphasis>
) ca at mid-length; at the end with a transverse spinelike process (
<emphasis id="B92C313DD6420650FC5EFBCFC6E0FB9D" box="[941,972,1068,1093]" italics="true" pageId="103" pageNumber="104">ttp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6420650FF4EFB8CC4CFFB39" pageId="103" pageNumber="104" type="distribution">
<paragraph id="8BE7ED2FD6420650FF4EFB8CC4C2FB51" blockId="103.[189,494,1135,1162]" box="[189,494,1135,1162]" pageId="103" pageNumber="104">
<heading id="D0AF5A43D6420650FF4EFB8CC4C2FB51" bold="true" box="[189,494,1135,1162]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="B92C313DD6420650FF4EFB8CC4C2FB51" bold="true" box="[189,494,1135,1162]" pageId="103" pageNumber="104">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFB42C4CFFB39" blockId="103.[189,1399,1184,1249]" pageId="103" pageNumber="104">
Only known from lowland rain forest at
<quantity id="4CA040CAD6420650FD51FB43C604FB63" box="[674,808,1184,1211]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="103" pageNumber="104" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</quantity>
a.s.l. at the
<typeStatus id="54E3538DD6420650FC48FB42C6C2FB63" box="[955,1006,1185,1211]" pageId="103" pageNumber="104">type</typeStatus>
locality, Kanga FR in the Kanga Mts (part of Nguru Mts).
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6420650FF4EFAE8C05EFA1F" pageId="103" pageNumber="104" type="discussion">
<paragraph id="8BE7ED2FD6420650FF4EFAE8C403FAFD" blockId="103.[189,303,1291,1317]" box="[189,303,1291,1317]" pageId="103" pageNumber="104">
<heading id="D0AF5A43D6420650FF4EFAE8C403FAFD" bold="true" box="[189,303,1291,1317]" fontSize="11" level="3" pageId="103" pageNumber="104" reason="3">
<emphasis id="B92C313DD6420650FF4EFAE8C403FAFD" bold="true" box="[189,303,1291,1317]" pageId="103" pageNumber="104">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EFADFC05EFA1F" blockId="103.[189,1399,1340,1480]" pageId="103" pageNumber="104">
Although
<taxonomicName id="4C5896ACD6420650FEC7FADEC4C5FA8E" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[308,489,1340,1366]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="bentemariae" status="sp. nov.">
<emphasis id="B92C313DD6420650FEC7FADEC4C5FA8E" box="[308,489,1340,1366]" italics="true" pageId="103" pageNumber="104">X. bentemariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6420650FE02FADFC761FA8E" box="[497,589,1340,1366]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
is here diagnosed vis-à-vis
<taxonomicName id="4C5896ACD6420650FC6AFADEC132FA8E" authorityName="VandenSpiegel &amp; Pierrard" authorityYear="2004" box="[921,1054,1340,1366]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="martella">
<emphasis id="B92C313DD6420650FC6AFADEC132FA8E" box="[921,1054,1340,1366]" italics="true" pageId="103" pageNumber="104">X. martella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5896ACD6420650FBA9FADEC1D8FA8E" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1114,1268,1340,1366]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="doggartae" status="sp. nov.">
<emphasis id="B92C313DD6420650FBA9FADEC1D8FA8E" box="[1114,1268,1340,1366]" italics="true" pageId="103" pageNumber="104">X. doggartae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6420650FB08FADFC075FA8E" box="[1275,1369,1340,1366]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
, it also shows some striking similarities with
<taxonomicName id="4C5896ACD6420650FD5DFA81C6E3FAA4" authority="Attems, 1910" authorityName="Attems" authorityYear="1910" box="[686,975,1378,1404]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B92C313DD6420650FD5DFA81C60BFAA4" box="[686,807,1378,1404]" italics="true" pageId="103" pageNumber="104">X. robusta</emphasis>
<bibRefCitation id="EFC990DED6420650FCDEFA81C6E3FAA4" author="Attems C. G." box="[813,975,1378,1404]" pageId="103" pageNumber="104" pagination="73 - 115" refId="ref46201" refString="Attems C. G. 1910. Myriopoden von Madagaskar, den Comoren und den Inseln Ostafrikas. In: VoeltzkowA. (ed.) Reise in Ostafrika in den Jahren 1903 - 1905, dritter Band: 73 - 115. E. Schweizerbart'sche Verlagsbuchhandlung, Nagele and Dr Sproesser, Stuttgart." type="book chapter" year="1910">Attems, 1910</bibRefCitation>
</taxonomicName>
. This species has been illustrated by
<bibRefCitation id="EFC990DED6420650FF4EFA6BC444FA7A" author="Attems C. G." box="[189,360,1416,1442]" pageId="103" pageNumber="104" pagination="73 - 115" refId="ref46201" refString="Attems C. G. 1910. Myriopoden von Madagaskar, den Comoren und den Inseln Ostafrikas. In: VoeltzkowA. (ed.) Reise in Ostafrika in den Jahren 1903 - 1905, dritter Band: 73 - 115. E. Schweizerbart'sche Verlagsbuchhandlung, Nagele and Dr Sproesser, Stuttgart." type="book chapter" year="1910">Attems (1910)</bibRefCitation>
and
<bibRefCitation id="EFC990DED6420650FE56FA64C61AFA7A" author="VandenSpiegel D. &amp; Pierrard G." box="[421,822,1415,1442]" pageId="103" pageNumber="104" pagination="31 - 46" refId="ref49620" refString="VandenSpiegel D. &amp; Pierrard G. 2004. Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Journal of Afrotropical Zoology 1: 31 - 46." type="journal article" year="2004">VandenSpiegel &amp; Pierrard (2004)</bibRefCitation>
, but the opportunity is here taken to show some SEM images (
<figureCitation id="1363F1AAD6420650FE96FA4EC496FA10" box="[357,442,1453,1480]" captionStart="Fig" captionStartId="107.[189,232,1891,1917]" captionTargetBox="[262,1326,265,1859]" captionTargetId="figure-85@107.[262,1326,265,1859]" captionTargetPageId="107" captionText="Fig. 65. Xystopyge robusta Attems, 1910, ♂, from Morogoro (HU-HR-124), right gonopod. AD. Coxa. A. Anterior view. B. Posterior view. C. Mesal view. D. Lateral view. EJ. Telopodite. E. Anterior view. F. Posterior view. G. Mesal view. H. Ventral view. I. Lateral view. J. Tip of solenomere. Abbreviations: sx =slender metaplical process;th = telomeral hook;TM =telomere. Scale bars:AI =0.2 mm; J =0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6359260" httpUri="https://zenodo.org/record/6359260/files/figure.png" pageId="103" pageNumber="104">Fig. 65</figureCitation>
) of the gonopods of a male of
<taxonomicName id="4C5896ACD6420650FCD1FA4DC6B7FA1F" authorityName="Attems" authorityYear="1910" box="[802,923,1453,1479]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B92C313DD6420650FCD1FA4DC6B7FA1F" box="[802,923,1453,1479]" italics="true" pageId="103" pageNumber="104">X. robusta</emphasis>
</taxonomicName>
collected close to the Uluguru Mts, viz:
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6420650FF4EFA11C6EEF9A6" pageId="103" pageNumber="104" type="materials_examined">
<paragraph id="8BE7ED2FD6420650FF4EFA11C6EEF9A6" blockId="103.[189,1398,1521,1662]" pageId="103" pageNumber="104">
<materialsCitation id="3B30E772D6420650FF4EFA11C6EEF9A6" ID-GBIF-Occurrence="3712026372" collectedFrom="hand collecting from ground and from plants" collectingDate="1989-03-17" collectionCode="HNHM" collectorName="Mahunka &amp; Zicsi" country="Tanzania" location="foot of Uluguru Mts" pageId="103" pageNumber="104" specimenCode="HNHM diplo-03051" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="syntype">
<collectingCountry id="F34FADBFD6420650FF4EFA11C47EF9D4" box="[189,338,1522,1548]" name="Tanzania" pageId="103" pageNumber="104">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D6420650FE98FA11C4A8F9D4" box="[363,388,1522,1548]" pageId="103" pageNumber="104" type="male"></specimenCount>
;
<collectingRegion id="499C23CDD6420650FE62FA11C744F9D3" box="[401,616,1521,1548]" country="Tanzania" name="Morogoro" pageId="103" pageNumber="104">Morogoro Region</collectingRegion>
,
<collectingRegion id="499C23CDD6420650FD87FA11C7C2F9D4" box="[628,750,1522,1548]" country="Tanzania" name="Morogoro" pageId="103" pageNumber="104">Morogoro</collectingRegion>
,
<location id="8E87BBF4D6420650FD09FA12C6C7F9D4" LSID="urn:lsid:plazi:treatment:03F15C39D6430650FDA6FD87C186F831:8E87BBF4D6420650FD09FA12C6C7F9D4" box="[762,1003,1521,1548]" country="Tanzania" name="foot of Uluguru Mts" pageId="103" pageNumber="104" stateProvince="Morogoro Region">foot of Uluguru Mts</location>
,
<locationDeviation id="9A55F40DD6420650FC0BFA12C060F9D4" box="[1016,1356,1521,1548]" pageId="103" pageNumber="104">above the university campus</locationDeviation>
;
<date id="FFE6CBEFD6420650FAAAFA11C415F9E9" pageId="103" pageNumber="104" value="1989-03-17">
<collectingDate id="EFA23207D6420650FAAAFA11C415F9E9" pageId="103" pageNumber="104" value="1989-03-17">17 Mar. 1989</collectingDate>
</date>
;
<collectedFrom id="3844AED2D6420650FEB6F9F4C67BF9E9" box="[325,855,1559,1585]" pageId="103" pageNumber="104">hand collecting from ground and from plants</collectedFrom>
;
<collectorName id="26AD88F9D6420650FC91F9F4C6FDF9E9" box="[866,977,1559,1585]" pageId="103" pageNumber="104">Mahunka</collectorName>
and
<collectorName id="26AD88F9D6420650FBF9F9F4C169F9E9" box="[1034,1093,1559,1585]" pageId="103" pageNumber="104">Zicsi</collectorName>
leg.;
<specimenCode id="DBFE4554D6420650FB75F9FBC05AF9EA" box="[1158,1398,1559,1586]" pageId="103" pageNumber="104">HNHM diplo-03051</specimenCode>
(Compared side by side with a syntype of
<taxonomicName id="4C5896ACD6420650FD25F9DDC678F98F" authorityName="Attems" authorityYear="1910" box="[726,852,1597,1623]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B92C313DD6420650FD25F9DDC678F98F" box="[726,852,1597,1623]" italics="true" pageId="103" pageNumber="104">X. robusta</emphasis>
</taxonomicName>
(NHMW 9114, Insel Pemba, Chake Chake, Voeltkow leg. and don.) the gonopods are completely identical.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D6420650FF4EF944C186F831" pageId="103" pageNumber="104" type="discussion">
<paragraph id="8BE7ED2FD6420650FF4EF944C17FF919" blockId="103.[189,1188,1703,1843]" box="[189,1107,1703,1730]" pageId="103" pageNumber="104">
Specific similarities between
<taxonomicName id="4C5896ACD6420650FDE5F94BC7BCF919" authorityName="Attems" authorityYear="1910" box="[534,656,1703,1729]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B92C313DD6420650FDE5F94BC7BCF919" box="[534,656,1703,1729]" italics="true" pageId="103" pageNumber="104">X. robusta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5896ACD6420650FD3AF94BC6A7F919" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[713,907,1703,1729]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="bentemarieae" status="sp. nov.">
<emphasis id="B92C313DD6420650FD3AF94BC6A7F919" box="[713,907,1703,1729]" italics="true" pageId="103" pageNumber="104">X. bentemarieae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6420650FC60F94BC6C2F91A" box="[915,1006,1704,1730]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
include:
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF92EC188F93F" blockId="103.[189,1188,1703,1843]" box="[189,1188,1741,1767]" pageId="103" pageNumber="104">
- the very complicated distal part of the metaplica, especially the slender process
<emphasis id="B92C313DD6420650FB78F92DC188F93F" box="[1163,1188,1742,1767]" italics="true" pageId="103" pageNumber="104">sx</emphasis>
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF910C73DF8D5" blockId="103.[189,1188,1703,1843]" box="[189,529,1779,1805]" pageId="103" pageNumber="104">- the simple telomere (TM)</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF8FAC672F8EB" blockId="103.[189,1188,1703,1843]" box="[189,862,1817,1843]" pageId="103" pageNumber="104">
- the hooklike moderately curved telomeral process (
<emphasis id="B92C313DD6420650FCCDF8FAC679F8EB" box="[830,853,1817,1843]" italics="true" pageId="103" pageNumber="104">th</emphasis>
)
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF8BEC480F8AF" blockId="103.[189,1398,1885,2025]" box="[189,428,1885,1911]" pageId="103" pageNumber="104">Differences include:</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF860C673F844" blockId="103.[189,1398,1885,2025]" box="[189,863,1923,1949]" pageId="103" pageNumber="104">
- details of the apical metaplical processes other than
<emphasis id="B92C313DD6420650FCB5F860C673F844" box="[838,863,1923,1948]" italics="true" pageId="103" pageNumber="104">sx</emphasis>
</paragraph>
<paragraph id="8BE7ED2FD6420650FF4EF84AC186F831" blockId="103.[189,1398,1885,2025]" pageId="103" pageNumber="104">
- the solenomere (
<emphasis id="B92C313DD6420650FE5BF84BC4FDF81A" box="[424,465,1960,1986]" italics="true" pageId="103" pageNumber="104">slm</emphasis>
) which in
<taxonomicName id="4C5896ACD6420650FDA0F84AC7E1F81A" authorityName="Attems" authorityYear="1910" box="[595,717,1960,1986]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B92C313DD6420650FDA0F84AC7E1F81A" box="[595,717,1960,1986]" italics="true" pageId="103" pageNumber="104">X. robusta</emphasis>
</taxonomicName>
ends in a striate lamella with a terminal hook, vs divided into a striate branch (
<emphasis id="B92C313DD6420650FE2FF82DC4D2F830" box="[476,510,1998,2024]" italics="true" pageId="103" pageNumber="104">sdl</emphasis>
) and a spinelike process (
<emphasis id="B92C313DD6420650FCC3F82DC675F830" box="[816,857,1998,2024]" italics="true" pageId="103" pageNumber="104">sdp</emphasis>
) in
<taxonomicName id="4C5896ACD6420650FC7BF82CC164F830" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[904,1096,1998,2024]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="bentemarieae" status="sp. nov.">
<emphasis id="B92C313DD6420650FC7BF82CC164F830" box="[904,1096,1998,2024]" italics="true" pageId="103" pageNumber="104">X. bentemarieae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6420650FBBCF82CC186F831" box="[1103,1194,1999,2025]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>