287 lines
52 KiB
XML
287 lines
52 KiB
XML
<document id="6669C8B2B9E301E1779133DAE3939A34" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFEBD711FF13FA472C67DDEA" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scolopendrinae Leach 1814" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="26" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:mods id="5A5B6FEDCBB2D306307B03833573D4FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="10FD0726BDA4F47E46A1CB433DDC20CC">
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<mods:title id="52B156D13F092BF6C8E10B9E3B765ADD">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:name id="80ECF2C70A40918924E8B14477C2546A" type="personal">
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<mods:namePart id="9FF8A27FA85CB1F941113F019E6484BB">Schileyko, Arkady A.</mods:namePart>
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<mods:nameIdentifier id="1BF0EB2EDFDE5C0869CE1DA49DE0328E" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:name id="3E3CEE31E87CA9DB11E7D5D202C8A06E" type="personal">
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<mods:namePart id="2B70361FD0020F42674DB534E696C665">Vahtera, Varpu</mods:namePart>
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<mods:nameIdentifier id="01A60E15C34A5DF5B76315DD6B6DFBF5" type="email">varpu.vahtera@gmail.com</mods:nameIdentifier>
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<mods:name id="89EC86824F97BFC30D759ABB16AABC84" type="personal">
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</mods:role>
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<mods:namePart id="0A6B70AEC8648C265A913137C98D4C7D">Edgecombe, Gregory D.</mods:namePart>
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<mods:nameIdentifier id="CDABFCF6F46844FB7790692306EB3972" type="ORCID">0000-0002-9591-8011</mods:nameIdentifier>
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<mods:nameIdentifier id="82DBECF7CBBD82D148EB09D7034B51E6" type="email">schileyko1965@gmail.com</mods:nameIdentifier>
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<mods:title id="84FC0956114DB3DCCA67D1C3E89F50A8">Zootaxa</mods:title>
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<mods:part id="82028864C94031CE5CE3687FB6CC8D53">
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<mods:date id="7857F15CC1A64EFA795F8C3389717825">2020</mods:date>
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<mods:number id="983149AD6025FA76AE85C487025141A3">2020-08-10</mods:number>
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<mods:number id="8D350555BA8D7EC0591428915A1505C4">4825</mods:number>
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<mods:number id="5DFEC7696DDA7E8528A3581A6A65E451">1</mods:number>
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<mods:classification id="DCCE4196F24605CCB1AEF918C0A0E67A">journal article</mods:classification>
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<mods:identifier id="6151722AB30362E90A8DD43A1817DCA7" type="DOI">10.11646/zootaxa.4825.1.1</mods:identifier>
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<treatment id="03DE092DFFEBD711FF13FA472C67DDEA" ID-DOI="http://doi.org/10.5281/zenodo.4455415" ID-GBIF-Taxon="177020725" ID-Zenodo-Dep="4455415" LSID="urn:lsid:plazi:treatment:03DE092DFFEBD711FF13FA472C67DDEA" httpUri="http://treatment.plazi.org/id/03DE092DFFEBD711FF13FA472C67DDEA" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
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<subSubSection id="C36DEBB0FFEBD713FF13FA472F78DA1C" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BC8B83BFFEBD713FF13FA472F3AD9DF" blockId="25.[151,639,1446,1473]" box="[151,639,1446,1473]" pageId="25" pageNumber="26">
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<heading id="D0800F57FFEBD713FF13FA472F3AD9DF" bold="true" box="[151,639,1446,1473]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="3">
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<emphasis id="B9036429FFEBD713FF13FA472F3AD9DF" bold="true" box="[151,639,1446,1473]" pageId="25" pageNumber="26">
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Subfamily
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<taxonomicName id="4C77C3B8FFEBD713FE9BFA472F3AD9DF" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[287,639,1446,1473]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subFamily" subFamily="Scolopendrinae">Scolopendrinae Leach, 1814</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC8B83BFFEBD713FF13FA0A2F78DA1C" blockId="25.[151,573,1515,1539]" box="[151,573,1515,1539]" pageId="25" pageNumber="26">Synonyms. Perustigminae Verhoeff, 1941</paragraph>
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</subSubSection>
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<subSubSection id="C36DEBB0FFEBD711FF13F9D02E32DD96" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="8BC8B83BFFEBD711FF13F9D029FCDCCA" blockId="25.[151,1437,1585,2008]" lastBlockId="27.[151,1437,150,501]" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
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<emphasis id="B9036429FFEBD713FF13F9D02C57DA54" bold="true" box="[151,274,1585,1611]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
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Anterior margin of forcipular coxosternite with tooth-plates; forcipular trochantero-prefemur with welldeveloped process. Sternites with well-developed (usually complete in most sternites) paramedian sutures, in most genera without depressions (may be present in the former
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<taxonomicName id="4C77C3B8FFEBD713FCBCF9982EEADA8B" authorityName="Verhoeff" authorityYear="1907" box="[824,943,1657,1684]" pageId="25" pageNumber="40" rank="tribe" tribe="Asanadini">Asanadini</taxonomicName>
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). Elongated spiracles slit-like, triangular in overwhelming majority of species (oval in some
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<taxonomicName id="4C77C3B8FFEBD713FD3AF97C2E2ADAA8" authorityName="Newport" authorityYear="1844" box="[702,879,1693,1719]" class="Chilopoda" family="Scolopendridae" genus="Cormocephalus" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FD3AF97C2E2ADAA8" box="[702,879,1693,1719]" italics="true" pageId="25" pageNumber="26">Cormocephalus</emphasis>
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</taxonomicName>
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) with well-developed atrium divided by a typical three-valved “flap” (
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<figureCitation id="134CA4BEFFEBD713FEFBF9202C97DAC3" box="[383,466,1729,1756]" captionStart="FIGURES 48–54" captionStartId="29.[151,264,1722,1746]" captionTargetBox="[245,1342,197,1680]" captionTargetId="figure-17@29.[228,1359,181,1699]" captionTargetPageId="29" captionText="FIGURES 48–54. Scolopendra mirabilis Porat, 1876 48 Locomotory legs of mid-body LBS laterally; Scolopendra cingulata Latreille, 1829; Ad. Rc 8007) 49 ultimate LBS + prefemora of ultimate legs ventrally; Scolopendra canidens Newport, 1844; Ad. Rc 6222 50 forcipular segment ventrally; Scolopendra subspinipes Leach, 1815; Ad. Rc 8010 51 left pleura of LBS 14–15 laterally; Ad. Rc 8010 52 head + LBS 1 dorsally; Sad. CDT 53 forcipular segment ventrally; Ad. Rc 8010 54 LBS 21 + ultimate legs dorso-laterally; (20)—leg 20, (ap)—apical spines of coxopleural process, (csp)—corner spine of ultimate prefemur, (cst)— forcipular coxosternite, (cx)—coxopleuron, (im)—intersclerite membrane, (o)—ocelli, (p)—single pleurites, (pr)—trochantero-prefemoral process, (pts)—pretarsus, (s)—spiracle covered by flap, (su)—ventral spines of ultimate prefemur, (T1)—tergite 1, (T15)—tergite 15, (t1)—tarsus 1, (t2)—tarsus 2, (tas)—tarsal spur, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402165" httpUri="https://zenodo.org/record/4402165/files/figure.png" pageId="25" pageNumber="26">Fig. 51</figureCitation>
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, fig. 8 of
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<bibRefCitation id="EFE6C5CAFFEBD713FDC5F9202ED7DAC4" author="Waldock, J. & Edgecombe, G. D." box="[577,914,1729,1755]" pageId="25" pageNumber="26" pagination="22 - 36" refId="ref51689" refString="Waldock, J. & Edgecombe, G. D. (2012) A new genus of scolopendrid centipede (Chilopoda: Scolopendromorpha: Scolopendrini) from the central Australian deserts. Zootaxa, 3321 (1), 22 - 36. https: // doi. org / 10.11646 / zootaxa. 3321.1.2" type="journal article" year="2012">Waldock & Edgecombe 2012</bibRefCitation>
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). LBS 7 without spiracles. Legs (
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<figureCitation id="134CA4BEFFEBD713FA8EF9202819DAC4" box="[1290,1372,1729,1755]" captionStart="FIGURES 48–54" captionStartId="29.[151,264,1722,1746]" captionTargetBox="[245,1342,197,1680]" captionTargetId="figure-17@29.[228,1359,181,1699]" captionTargetPageId="29" captionText="FIGURES 48–54. Scolopendra mirabilis Porat, 1876 48 Locomotory legs of mid-body LBS laterally; Scolopendra cingulata Latreille, 1829; Ad. Rc 8007) 49 ultimate LBS + prefemora of ultimate legs ventrally; Scolopendra canidens Newport, 1844; Ad. Rc 6222 50 forcipular segment ventrally; Scolopendra subspinipes Leach, 1815; Ad. Rc 8010 51 left pleura of LBS 14–15 laterally; Ad. Rc 8010 52 head + LBS 1 dorsally; Sad. CDT 53 forcipular segment ventrally; Ad. Rc 8010 54 LBS 21 + ultimate legs dorso-laterally; (20)—leg 20, (ap)—apical spines of coxopleural process, (csp)—corner spine of ultimate prefemur, (cst)— forcipular coxosternite, (cx)—coxopleuron, (im)—intersclerite membrane, (o)—ocelli, (p)—single pleurites, (pr)—trochantero-prefemoral process, (pts)—pretarsus, (s)—spiracle covered by flap, (su)—ventral spines of ultimate prefemur, (T1)—tergite 1, (T15)—tergite 15, (t1)—tarsus 1, (t2)—tarsus 2, (tas)—tarsal spur, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402165" httpUri="https://zenodo.org/record/4402165/files/figure.png" pageId="25" pageNumber="26">Fig. 48</figureCitation>
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) usually with one (leg 1 sometimes with 2, penultimate leg sometimes lacking) tarsal spur(s). These spurs are totally absent in the
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<taxonomicName id="4C77C3B8FFEBD713FEB5F8E82CA7DB3C" authorityName="Newport" authorityYear="1844" box="[305,482,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Cormocephalus" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FEB5F8E82CA7DB3C" box="[305,482,1801,1827]" italics="true" pageId="25" pageNumber="26">Cormocephalus</emphasis>
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</taxonomicName>
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-clade (i.e.
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<taxonomicName id="4C77C3B8FFEBD713FDE0F8E82E50DB3C" authorityName="Newport" authorityYear="1844" box="[612,789,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Cormocephalus" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FDE0F8E82E50DB3C" box="[612,789,1801,1827]" italics="true" pageId="25" pageNumber="26">Cormocephalus</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C77C3B8FFEBD713FCB3F8E82EBBDB3C" authorityName="Kraepelin" authorityYear="1903" box="[823,1022,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Hemiscolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FCB3F8E82EBBDB3C" box="[823,1022,1801,1827]" italics="true" pageId="25" pageNumber="26">Hemiscolopendra</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C77C3B8FFEBD713FBA4F8E829F2DB3C" authorityName="Shelley" authorityYear="2008" box="[1056,1207,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Akymnopellis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FBA4F8E829F2DB3C" box="[1056,1207,1801,1827]" italics="true" pageId="25" pageNumber="26">Akymnopellis</emphasis>
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</taxonomicName>
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+
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<taxonomicName id="4C77C3B8FFEBD713FB5CF8E828D6DB3C" box="[1240,1427,1801,1827]" class="Chilopoda" family="Scolopendridae" genus="Campylostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FB5CF8E828D6DB3C" box="[1240,1427,1801,1827]" italics="true" pageId="25" pageNumber="26">Campylostigmus</emphasis>
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</taxonomicName>
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) and in the genus
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<taxonomicName id="4C77C3B8FFEBD713FEDCF8CC2CFFDB58" authorityName="Meinert" authorityYear="1886" box="[344,442,1837,1863]" class="Chilopoda" family="Scolopendridae" genus="Asanada" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9036429FFEBD713FEDCF8CC2CFFDB58" box="[344,442,1837,1863]" italics="true" pageId="25" pageNumber="26">Asanada</emphasis>
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</taxonomicName>
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the presence of these spurs may even be subject to intraspecific variability (see below). Ultimate legs in most genera of “common” shape (
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<figureCitation id="134CA4BEFFEBD713FD5DF8B02E74DB73" box="[729,817,1873,1900]" captionStart="FIGURES 48–54" captionStartId="29.[151,264,1722,1746]" captionTargetBox="[245,1342,197,1680]" captionTargetId="figure-17@29.[228,1359,181,1699]" captionTargetPageId="29" captionText="FIGURES 48–54. Scolopendra mirabilis Porat, 1876 48 Locomotory legs of mid-body LBS laterally; Scolopendra cingulata Latreille, 1829; Ad. Rc 8007) 49 ultimate LBS + prefemora of ultimate legs ventrally; Scolopendra canidens Newport, 1844; Ad. Rc 6222 50 forcipular segment ventrally; Scolopendra subspinipes Leach, 1815; Ad. Rc 8010 51 left pleura of LBS 14–15 laterally; Ad. Rc 8010 52 head + LBS 1 dorsally; Sad. CDT 53 forcipular segment ventrally; Ad. Rc 8010 54 LBS 21 + ultimate legs dorso-laterally; (20)—leg 20, (ap)—apical spines of coxopleural process, (csp)—corner spine of ultimate prefemur, (cst)— forcipular coxosternite, (cx)—coxopleuron, (im)—intersclerite membrane, (o)—ocelli, (p)—single pleurites, (pr)—trochantero-prefemoral process, (pts)—pretarsus, (s)—spiracle covered by flap, (su)—ventral spines of ultimate prefemur, (T1)—tergite 1, (T15)—tergite 15, (t1)—tarsus 1, (t2)—tarsus 2, (tas)—tarsal spur, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402165" httpUri="https://zenodo.org/record/4402165/files/figure.png" pageId="25" pageNumber="26">Figs 54</figureCitation>
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,
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<figureCitation id="134CA4BEFFEBD713FCBAF8B02E19DB73" box="[830,860,1873,1900]" captionStart="FIGURES 62–67" captionStartId="33.[151,264,1720,1744]" captionTargetBox="[324,1265,194,1683]" captionTargetId="figure-17@33.[310,1277,181,1696]" captionTargetPageId="33" captionText="FIGURES 62–67. Cormocephalus sp.; Sad. Rc 7468 62 head + LBS 1 dorsally 63 head + forcipular segment ventrally; Cormocephalus guildingii Newport, 1845; Holotype MYRI-016-01 OMNH (photo by Dr. Danner J. Mann) 64 LBS 20, 21 and ultimate legs ventrally; Cormocephalus punctatus Porat, 1871; Ad. Rc 7022 65 ultimate legs laterally; Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 66 LBS 21, prefemur + femur of left ultimate leg and much reduced (regenerated) right ultimate leg ventrally; Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 67 head + LBS 1–2 dorsally; (20)—leg 20, (as)—accessory spines, (bp)—basal plates, (cp)—coxopleural process, (cx)—coxopleuron, (f)—ultimate femur, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (ps)—cephalic paramedian sutures, (pts)—pretarsus, (sc)—spine at place of coxopleural process, (stc)—sutures of forcipular coxosternite, (T1)—tergite 1, (tp)—forcipular tooth-plate, (ts)—anterior transverse suture of tergite 1, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402173" httpUri="https://zenodo.org/record/4402173/files/figure.png" pageId="25" pageNumber="26">65</figureCitation>
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,
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<figureCitation id="134CA4BEFFEBD713FCEDF8B02ECDDB74" box="[873,904,1873,1899]" captionStart="FIGURES 68–72" captionStartId="36.[151,264,1757,1781]" captionTargetBox="[172,1416,201,1716]" captionTargetId="figure-19@36.[151,1436,181,1734]" captionTargetPageId="36" captionText="FIGURES 68–72. Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 68 forcipular segment ventrally 69 left coxopleuron and left ultimate leg laterally; Akymnopellis chilensis (Gervais, 1847); Female Rc 7463 70 left side of ultimate LBS and ultimate legs laterally; BMNH, BM 1904.1.0.29.1–3 71 forcipular segment ventrally; Ad. male Rc 8005 72 LBS 21 and prefemora of ultimate legs ventro-laterally; (cp)—coxopleural process, (g)—gonopods, (pf)—normally developed left prefemur, (pr)—trochantero-prefemoral process, (rpf)—abnormally small (regenerated) right prefemur, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402175" httpUri="https://zenodo.org/record/4402175/files/figure.png" pageId="25" pageNumber="26">70</figureCitation>
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), more rarely quasi “pincer-shaped” (i.e. much shortened and enlarged except for pretarsus, for example in
|
||
<taxonomicName id="4C77C3B8FFEBD713FCBFF8942ED8DB90" authorityName="Meinert" authorityYear="1886" box="[827,925,1909,1935]" class="Chilopoda" family="Scolopendridae" genus="Asanada" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFEBD713FCBFF8942ED8DB90" box="[827,925,1909,1935]" italics="true" pageId="25" pageNumber="26">Asanada</emphasis>
|
||
</taxonomicName>
|
||
and the former genus
|
||
<taxonomicName id="4C77C3B8FFEBD713FB1EF894284FDB90" authorityName="Waldock & Edgecombe" authorityYear="2012" box="[1178,1290,1909,1935]" class="Chilopoda" family="Scolopendridae" genus="Kanparka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFEBD713FB1EF894284FDB90" box="[1178,1290,1909,1935]" italics="true" pageId="25" pageNumber="26">Kanparka</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="134CA4BEFFEBD713FA9DF8942829DB90" box="[1305,1388,1909,1935]" captionStart="FIGURES 78–83" captionStartId="42.[151,264,1708,1732]" captionTargetBox="[168,1421,193,1665]" captionTargetId="figure-19@42.[151,1436,181,1684]" captionTargetPageId="42" captionText="FIGURES 78–83. Asanada sokotrana Pocock, 1899; BMNH 1934.2.15.151–154 78 LBS 21 ventrally; Asanada indica Jangi & Dass, 1984; Smaller ad. Rc 7484 79 head + LBS 1 dorsally; Asanada brevicornis Meinert, 1886; BMNH 1889.7.15.21–23 80 apical part of maxilla 2 dorso-laterally; Asanada sokotrana Pocock, 1899; BMNH 1899.7.4.85–86 81 LBS 20–21 and ultimate legs dorsally; Otostigmus (O.) multidens Haase, 1887; Ad. Rc 7510 82 right pleuron of LBS 3 laterally; Otostigmus (O.) multidens nordicus Schileyko, 1995; Ad. Rc 6747 83 left side of ultimate LBS + right ultimate leg medially; (an)—antenna, (cx)—coxopleuron, (f)—ultimate femur, (im)—intersclerite membrane, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pts)—non enlarged ultimate pretarsus, (s)—open spiracle with atrium, (su)—spines of ultimate prefemur, (T1)—tergite 1, (t1)—tarsus 1, (t2)—tarsus 2, (ti)—tibia, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402181" httpUri="https://zenodo.org/record/4402181/files/figure.png" pageId="25" pageNumber="26">Figs 81</figureCitation>
|
||
and
|
||
<figureCitation id="134CA4BEFFEBD713FF13F8782DF6DBAB" box="[151,179,1945,1972]" captionStart="FIGURES 55–61" captionStartId="30.[151,264,1644,1668]" captionTargetBox="[222,1366,197,1607]" captionTargetId="figure-19@30.[207,1381,181,1620]" captionTargetPageId="30" captionText="FIGURES 55–61. Kanparka (=Scolopendra) leki Waldock & Edgecombe, 2012; WAM T79350 55 head + LBS 1 dorsally; WAM T99422 56 posterior part of LBS 21 + ultimate legs ventrally 57 LBS 21 + left ultimate leg laterally; Arthrorhabdus mjobergi Kraepelin, 1916; Live specimen (photo by Mark A. Newton, corrected; available at https://marknewton.smugmug. com/, https://www.wildsouthaustralia.info/ Centipedes/Arthrorhabdus-spp/i-NkWPntT/A) 58 generalview dorsally; Arthrorhabdus formosus Pocock, 1891; ZMB 4942 59 apical part of maxilla 2 dorsally; 60 forcipular segment ventrally; Cormocephalus (Campylostigmus) orientalis Ribaut, 1923; BMNH 61 forcipular segment ventrally; (as)—accessory spines, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (db)—dorsal brush, (f)—ultimate femur, (mp)—medial spines of coxopleural process, (ms)—medial spines of ultimate prefemur, femur, tibia, (o)—ocelli, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—not enlarged ultimate pretarsus, (su)—a few ventral spines of ultimate femur and prefemur, (T1)—tergite 1, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)—ultimate tibia, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402169" httpUri="https://zenodo.org/record/4402169/files/figure.png" pageId="25" pageNumber="26">57</figureCitation>
|
||
respectively) or even truly “pincer-shaped” (
|
||
<taxonomicName id="4C77C3B8FFEBD713FD2FF8782E26DBAC" authorityName="Brandt" authorityYear="1841" box="[683,867,1945,1971]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFEBD713FD2FF8782E26DBAC" box="[683,867,1945,1971]" italics="true" pageId="25" pageNumber="26">Scolopendropsis</emphasis>
|
||
</taxonomicName>
|
||
and some species of
|
||
<taxonomicName id="4C77C3B8FFEBD713FBCBF8782D94DBC7" authority="(Cormocephalus)" class="Chilopoda" family="Scolopendridae" genus="Cormocephalus" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subGenus" subGenus="Cormocephalus">
|
||
<emphasis id="B9036429FFEBD713FBCBF8782845DBAC" box="[1103,1280,1945,1971]" italics="true" pageId="25" pageNumber="26">Cormocephalus</emphasis>
|
||
(
|
||
<emphasis id="B9036429FFEBD713FA94F8782D83DBC8" italics="true" pageId="25" pageNumber="26">Cormocephalus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="134CA4BEFFEBD713FF5AF85C2C70DBC7" box="[222,309,1981,2008]" captionStart="FIGURES 40–47" captionStartId="26.[151,264,1651,1675]" captionTargetBox="[286,1303,195,1616]" captionTargetId="figure-19@26.[273,1315,181,1627]" captionTargetPageId="26" captionText="FIGURES 40–47. Tonkinodentus lestes Schileyko, 1992; Holotype adt Rc 6358 40 forcipular segment ventrally; Sad. Rc 6555 41 ultimate LBS, legs 20 + ultimate legs dorso-laterally; Scolopendropsis duplicata Chagas-Jr, Edgecombe & Minelli, 2008; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr) 42 head, forcipular segment + LBS 1–3 ventrally; Holotype MNRJ 15258 43 ultimate LBS ventrally; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr) 44 posterior half of ultimate LBS + ultimate legs ventrally; Scolopendropsis bahiensis Brandt, 1841; Live specimen (photo by Dr. Adriano Kury) 45 general view dorsally; Scolopendropsis duplicata Chagas-Jr, Edgecombe & Minelli, 2008 (photos by Dr. Amazonas Chagas-Jr); Paratype IBSP 2392 or MNRJ 15306 46 head + LBS 1 dorsally 47 ultimate LBS + ultimate legs dorsally; (20)—leg 20, (cx)—coxopleuron, (cst)—forcipular coxosternite, (dv)—distoventrolateral process of femur, (lsc)—sternal median longitudinal sulcus, (sl)—cephalic median longitudinal suture, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (sc)—spine at place of coxopleural process, (su)—spines of ultimate prefemur, (T1)—tergite 1, (t1)—tarsus 1, (t2)— tarsus 2, (ti)—tibia, (tp)—forcipular tooth-plate, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402163" httpUri="https://zenodo.org/record/4402163/files/figure.png" pageId="25" pageNumber="26">Figs 45</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFEBD713FEC6F85C2C27DBC7" box="[322,354,1981,2008]" captionStart="FIGURES 62–67" captionStartId="33.[151,264,1720,1744]" captionTargetBox="[324,1265,194,1683]" captionTargetId="figure-17@33.[310,1277,181,1696]" captionTargetPageId="33" captionText="FIGURES 62–67. Cormocephalus sp.; Sad. Rc 7468 62 head + LBS 1 dorsally 63 head + forcipular segment ventrally; Cormocephalus guildingii Newport, 1845; Holotype MYRI-016-01 OMNH (photo by Dr. Danner J. Mann) 64 LBS 20, 21 and ultimate legs ventrally; Cormocephalus punctatus Porat, 1871; Ad. Rc 7022 65 ultimate legs laterally; Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 66 LBS 21, prefemur + femur of left ultimate leg and much reduced (regenerated) right ultimate leg ventrally; Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 67 head + LBS 1–2 dorsally; (20)—leg 20, (as)—accessory spines, (bp)—basal plates, (cp)—coxopleural process, (cx)—coxopleuron, (f)—ultimate femur, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (ps)—cephalic paramedian sutures, (pts)—pretarsus, (sc)—spine at place of coxopleural process, (stc)—sutures of forcipular coxosternite, (T1)—tergite 1, (tp)—forcipular tooth-plate, (ts)—anterior transverse suture of tergite 1, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402173" httpUri="https://zenodo.org/record/4402173/files/figure.png" pageId="25" pageNumber="26">64</figureCitation>
|
||
). Ultimate leg prefemur practically always with some chitinized spines (their number may vary considerably;
|
||
<figureCitation id="134CA4BEFFE9D711FEB3FF762CD5DCAE" box="[311,400,151,177]" captionStart="FIGURES 40–47" captionStartId="26.[151,264,1651,1675]" captionTargetBox="[286,1303,195,1616]" captionTargetId="figure-19@26.[273,1315,181,1627]" captionTargetPageId="26" captionText="FIGURES 40–47. Tonkinodentus lestes Schileyko, 1992; Holotype adt Rc 6358 40 forcipular segment ventrally; Sad. Rc 6555 41 ultimate LBS, legs 20 + ultimate legs dorso-laterally; Scolopendropsis duplicata Chagas-Jr, Edgecombe & Minelli, 2008; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr) 42 head, forcipular segment + LBS 1–3 ventrally; Holotype MNRJ 15258 43 ultimate LBS ventrally; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr) 44 posterior half of ultimate LBS + ultimate legs ventrally; Scolopendropsis bahiensis Brandt, 1841; Live specimen (photo by Dr. Adriano Kury) 45 general view dorsally; Scolopendropsis duplicata Chagas-Jr, Edgecombe & Minelli, 2008 (photos by Dr. Amazonas Chagas-Jr); Paratype IBSP 2392 or MNRJ 15306 46 head + LBS 1 dorsally 47 ultimate LBS + ultimate legs dorsally; (20)—leg 20, (cx)—coxopleuron, (cst)—forcipular coxosternite, (dv)—distoventrolateral process of femur, (lsc)—sternal median longitudinal sulcus, (sl)—cephalic median longitudinal suture, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (sc)—spine at place of coxopleural process, (su)—spines of ultimate prefemur, (T1)—tergite 1, (t1)—tarsus 1, (t2)— tarsus 2, (ti)—tibia, (tp)—forcipular tooth-plate, (ust)—ultimate sternite, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402163" httpUri="https://zenodo.org/record/4402163/files/figure.png" pageId="27" pageNumber="28">Figs 44</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFE9D711FE19FF762CF9DCAE" box="[413,444,151,177]" captionStart="FIGURES 48–54" captionStartId="29.[151,264,1722,1746]" captionTargetBox="[245,1342,197,1680]" captionTargetId="figure-17@29.[228,1359,181,1699]" captionTargetPageId="29" captionText="FIGURES 48–54. Scolopendra mirabilis Porat, 1876 48 Locomotory legs of mid-body LBS laterally; Scolopendra cingulata Latreille, 1829; Ad. Rc 8007) 49 ultimate LBS + prefemora of ultimate legs ventrally; Scolopendra canidens Newport, 1844; Ad. Rc 6222 50 forcipular segment ventrally; Scolopendra subspinipes Leach, 1815; Ad. Rc 8010 51 left pleura of LBS 14–15 laterally; Ad. Rc 8010 52 head + LBS 1 dorsally; Sad. CDT 53 forcipular segment ventrally; Ad. Rc 8010 54 LBS 21 + ultimate legs dorso-laterally; (20)—leg 20, (ap)—apical spines of coxopleural process, (csp)—corner spine of ultimate prefemur, (cst)— forcipular coxosternite, (cx)—coxopleuron, (im)—intersclerite membrane, (o)—ocelli, (p)—single pleurites, (pr)—trochantero-prefemoral process, (pts)—pretarsus, (s)—spiracle covered by flap, (su)—ventral spines of ultimate prefemur, (T1)—tergite 1, (T15)—tergite 15, (t1)—tarsus 1, (t2)—tarsus 2, (tas)—tarsal spur, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402165" httpUri="https://zenodo.org/record/4402165/files/figure.png" pageId="27" pageNumber="28">49</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFE9D711FE4EFF772CADDCAE" box="[458,488,150,177]" captionStart="FIGURES 55–61" captionStartId="30.[151,264,1644,1668]" captionTargetBox="[222,1366,197,1607]" captionTargetId="figure-19@30.[207,1381,181,1620]" captionTargetPageId="30" captionText="FIGURES 55–61. Kanparka (=Scolopendra) leki Waldock & Edgecombe, 2012; WAM T79350 55 head + LBS 1 dorsally; WAM T99422 56 posterior part of LBS 21 + ultimate legs ventrally 57 LBS 21 + left ultimate leg laterally; Arthrorhabdus mjobergi Kraepelin, 1916; Live specimen (photo by Mark A. Newton, corrected; available at https://marknewton.smugmug. com/, https://www.wildsouthaustralia.info/ Centipedes/Arthrorhabdus-spp/i-NkWPntT/A) 58 generalview dorsally; Arthrorhabdus formosus Pocock, 1891; ZMB 4942 59 apical part of maxilla 2 dorsally; 60 forcipular segment ventrally; Cormocephalus (Campylostigmus) orientalis Ribaut, 1923; BMNH 61 forcipular segment ventrally; (as)—accessory spines, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (db)—dorsal brush, (f)—ultimate femur, (mp)—medial spines of coxopleural process, (ms)—medial spines of ultimate prefemur, femur, tibia, (o)—ocelli, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—not enlarged ultimate pretarsus, (su)—a few ventral spines of ultimate femur and prefemur, (T1)—tergite 1, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)—ultimate tibia, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402169" httpUri="https://zenodo.org/record/4402169/files/figure.png" pageId="27" pageNumber="28">56</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFE9D711FE72FF762F56DCAE" box="[502,531,151,177]" captionStart="FIGURES 68–72" captionStartId="36.[151,264,1757,1781]" captionTargetBox="[172,1416,201,1716]" captionTargetId="figure-19@36.[151,1436,181,1734]" captionTargetPageId="36" captionText="FIGURES 68–72. Hemiscolopendra marginata (Say, 1821); Sad. Rc 6494 68 forcipular segment ventrally 69 left coxopleuron and left ultimate leg laterally; Akymnopellis chilensis (Gervais, 1847); Female Rc 7463 70 left side of ultimate LBS and ultimate legs laterally; BMNH, BM 1904.1.0.29.1–3 71 forcipular segment ventrally; Ad. male Rc 8005 72 LBS 21 and prefemora of ultimate legs ventro-laterally; (cp)—coxopleural process, (g)—gonopods, (pf)—normally developed left prefemur, (pr)—trochantero-prefemoral process, (rpf)—abnormally small (regenerated) right prefemur, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402175" httpUri="https://zenodo.org/record/4402175/files/figure.png" pageId="27" pageNumber="28">72</figureCitation>
|
||
) plus corner spine (
|
||
<figureCitation id="134CA4BEFFE9D711FD7EFF762E11DCAE" box="[762,852,150,177]" captionStart="FIGURES 48–54" captionStartId="29.[151,264,1722,1746]" captionTargetBox="[245,1342,197,1680]" captionTargetId="figure-17@29.[228,1359,181,1699]" captionTargetPageId="29" captionText="FIGURES 48–54. Scolopendra mirabilis Porat, 1876 48 Locomotory legs of mid-body LBS laterally; Scolopendra cingulata Latreille, 1829; Ad. Rc 8007) 49 ultimate LBS + prefemora of ultimate legs ventrally; Scolopendra canidens Newport, 1844; Ad. Rc 6222 50 forcipular segment ventrally; Scolopendra subspinipes Leach, 1815; Ad. Rc 8010 51 left pleura of LBS 14–15 laterally; Ad. Rc 8010 52 head + LBS 1 dorsally; Sad. CDT 53 forcipular segment ventrally; Ad. Rc 8010 54 LBS 21 + ultimate legs dorso-laterally; (20)—leg 20, (ap)—apical spines of coxopleural process, (csp)—corner spine of ultimate prefemur, (cst)— forcipular coxosternite, (cx)—coxopleuron, (im)—intersclerite membrane, (o)—ocelli, (p)—single pleurites, (pr)—trochantero-prefemoral process, (pts)—pretarsus, (s)—spiracle covered by flap, (su)—ventral spines of ultimate prefemur, (T1)—tergite 1, (T15)—tergite 15, (t1)—tarsus 1, (t2)—tarsus 2, (tas)—tarsal spur, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402165" httpUri="https://zenodo.org/record/4402165/files/figure.png" pageId="27" pageNumber="28">Figs 54</figureCitation>
|
||
,
|
||
<figureCitation id="134CA4BEFFE9D711FCE6FF772EC7DCAE" box="[866,898,150,177]" captionStart="FIGURES 55–61" captionStartId="30.[151,264,1644,1668]" captionTargetBox="[222,1366,197,1607]" captionTargetId="figure-19@30.[207,1381,181,1620]" captionTargetPageId="30" captionText="FIGURES 55–61. Kanparka (=Scolopendra) leki Waldock & Edgecombe, 2012; WAM T79350 55 head + LBS 1 dorsally; WAM T99422 56 posterior part of LBS 21 + ultimate legs ventrally 57 LBS 21 + left ultimate leg laterally; Arthrorhabdus mjobergi Kraepelin, 1916; Live specimen (photo by Mark A. Newton, corrected; available at https://marknewton.smugmug. com/, https://www.wildsouthaustralia.info/ Centipedes/Arthrorhabdus-spp/i-NkWPntT/A) 58 generalview dorsally; Arthrorhabdus formosus Pocock, 1891; ZMB 4942 59 apical part of maxilla 2 dorsally; 60 forcipular segment ventrally; Cormocephalus (Campylostigmus) orientalis Ribaut, 1923; BMNH 61 forcipular segment ventrally; (as)—accessory spines, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (db)—dorsal brush, (f)—ultimate femur, (mp)—medial spines of coxopleural process, (ms)—medial spines of ultimate prefemur, femur, tibia, (o)—ocelli, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—not enlarged ultimate pretarsus, (su)—a few ventral spines of ultimate femur and prefemur, (T1)—tergite 1, (t1)—ultimate tarsus 1, (t2)—ultimate tarsus 2, (ti)—ultimate tibia, (tp)—forcipular tooth-plate, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402169" httpUri="https://zenodo.org/record/4402169/files/figure.png" pageId="27" pageNumber="28">57</figureCitation>
|
||
); claw-shaped pretarsus well-developed.
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FADDFF762CA5DCCA" author="Edgecombe, G. D. & Bonato, L." pageId="27" pageNumber="28" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 397)</bibRefCitation>
|
||
also wrote: “Posteriorly directed spines along plicae of gizzard.”
|
||
</paragraph>
|
||
<caption id="DF08E8B3FFE8D710FF13F992292DDBEC" ID-DOI="http://doi.org/10.5281/zenodo.4402163" ID-Zenodo-Dep="4402163" httpUri="https://zenodo.org/record/4402163/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1651,1675]" targetBox="[286,1303,195,1616]" targetPageId="26">
|
||
<paragraph id="8BC8B83BFFE8D710FF13F992292DDBEC" blockId="26.[151,1437,1651,2035]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9036429FFE8D710FF13F9922C11DA94" bold="true" box="[151,340,1651,1675]" pageId="26" pageNumber="27">FIGURES 40–47.</emphasis>
|
||
<taxonomicName id="4C77C3B8FFE8D710FEDEF9922F9FDA94" authority="Schileyko, 1992" authorityName="Schileyko" authorityYear="1992" box="[346,730,1651,1675]" class="Chilopoda" family="Cryptopidae" genus="Tonkinodentus" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lestes">
|
||
<emphasis id="B9036429FFE8D710FEDEF9922F6EDA94" box="[346,555,1651,1675]" italics="true" pageId="26" pageNumber="27">Tonkinodentus lestes</emphasis>
|
||
<bibRefCitation id="EFE6C5CAFFE8D710FDB6F9922F9FDA94" author="Schileyko" box="[562,730,1651,1675]" pageId="26" pageNumber="27" pagination="5 - 19" refId="ref49894" refString="Schileyko, Ark. A. (1992) Scolopenders of Vietnam and some aspects of the system of Scolopendromorpha (Chilopoda Epimorpha) Part 1. Arthropoda Selecta, 1 (1), 5 - 19." type="journal article" year="1992">Schileyko, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
; Holotype adt Rc 6358
|
||
<emphasis id="B9036429FFE8D710FC4DF9922EA7DA94" bold="true" box="[969,994,1651,1675]" pageId="26" pageNumber="27">40</emphasis>
|
||
forcipular segment ventrally; Sad. Rc 6555
|
||
<emphasis id="B9036429FFE8D710FF13F9762DF5DAB0" bold="true" box="[151,176,1687,1711]" pageId="26" pageNumber="27">41</emphasis>
|
||
ultimate LBS, legs 20 + ultimate legs dorso-laterally;
|
||
<taxonomicName id="4C77C3B8FFE8D710FD62F9762EB0DAB0" authorityName="Chagas Jr., Edgecombe & Minelli" authorityYear="2008" box="[742,1013,1687,1711]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B9036429FFE8D710FD62F9762EB0DAB0" box="[742,1013,1687,1711]" italics="true" pageId="26" pageNumber="27">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
Chagas-Jr,
|
||
<bibRefCitation id="EFE6C5CAFFE8D710FBEBF97628DCDAB0" author="Edgecombe, G. D. & Koch, M." box="[1135,1433,1687,1711]" pageId="26" pageNumber="27" pagination="872 - 901" refId="ref47403" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00220. x" type="journal article" year="2008">Edgecombe & Minelli, 2008</bibRefCitation>
|
||
; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr)
|
||
<emphasis id="B9036429FFE8D710FC1DF95A2EF7DACC" bold="true" box="[921,946,1723,1747]" pageId="26" pageNumber="27">42</emphasis>
|
||
head, forcipular segment + LBS 1–3 ventrally; Holotype MNRJ 15258
|
||
<emphasis id="B9036429FFE8D710FE0BF93E2CEDDAE8" bold="true" box="[399,424,1759,1783]" pageId="26" pageNumber="27">43</emphasis>
|
||
ultimate LBS ventrally; Paratype IBSP 2392 or MNRJ 15306 (photo by Dr. Amazonas Chagas-Jr)
|
||
<emphasis id="B9036429FFE8D710FF13F8E22DF5DB04" bold="true" box="[151,176,1795,1819]" pageId="26" pageNumber="27">44</emphasis>
|
||
posterior half of ultimate LBS + ultimate legs ventrally;
|
||
<taxonomicName id="4C77C3B8FFE8D710FD7CF8E229D1DB04" authority="Brandt, 1841" authorityName="Brandt" authorityYear="1841" box="[760,1172,1795,1819]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bahiensis">
|
||
<emphasis id="B9036429FFE8D710FD7CF8E22940DB04" box="[760,1029,1795,1819]" italics="true" pageId="26" pageNumber="27">Scolopendropsis bahiensis</emphasis>
|
||
Brandt, 1841
|
||
</taxonomicName>
|
||
; Live specimen (photo by Dr. Adriano Kury)
|
||
<emphasis id="B9036429FFE8D710FEDCF8C62C34DB20" bold="true" box="[344,369,1831,1855]" pageId="26" pageNumber="27">45</emphasis>
|
||
general view dorsally;
|
||
<taxonomicName id="4C77C3B8FFE8D710FDDBF8C62E29DB20" authorityName="Chagas Jr., Edgecombe & Minelli" authorityYear="2008" box="[607,876,1831,1855]" class="Chilopoda" family="Scolopendridae" genus="Scolopendropsis" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="duplicata">
|
||
<emphasis id="B9036429FFE8D710FDDBF8C62E29DB20" box="[607,876,1831,1855]" italics="true" pageId="26" pageNumber="27">Scolopendropsis duplicata</emphasis>
|
||
</taxonomicName>
|
||
Chagas-Jr,
|
||
<bibRefCitation id="EFE6C5CAFFE8D710FC67F8C62840DB20" author="Edgecombe, G. D. & Koch, M." box="[995,1285,1831,1855]" pageId="26" pageNumber="27" pagination="872 - 901" refId="ref47403" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901. https: // doi. org / 10.1111 / j. 1096 - 0031.2008.00220. x" type="journal article" year="2008">Edgecombe & Minelli, 2008</bibRefCitation>
|
||
(photos by Dr. Amazonas Chagas-Jr); Paratype IBSP 2392 or MNRJ 15306
|
||
<emphasis id="B9036429FFE8D710FC89F8AA2E63DB7C" bold="true" box="[781,806,1867,1891]" pageId="26" pageNumber="27">46</emphasis>
|
||
head + LBS 1 dorsally
|
||
<emphasis id="B9036429FFE8D710FB9BF8AA297DDB7C" bold="true" box="[1055,1080,1867,1891]" pageId="26" pageNumber="27">47</emphasis>
|
||
ultimate LBS + ultimate legs dorsally; (
|
||
<emphasis id="B9036429FFE8D710FF58F88E2DB0DB98" bold="true" box="[220,245,1903,1927]" pageId="26" pageNumber="27">20</emphasis>
|
||
)—leg 20, (
|
||
<emphasis id="B9036429FFE8D710FEECF88E2CC5DB98" bold="true" box="[360,384,1903,1927]" pageId="26" pageNumber="27">cx</emphasis>
|
||
)—coxopleuron, (
|
||
<emphasis id="B9036429FFE8D710FDB1F88E2F16DB98" bold="true" box="[565,595,1903,1927]" pageId="26" pageNumber="27">cst</emphasis>
|
||
)—forcipular coxosternite, (
|
||
<emphasis id="B9036429FFE8D710FCE9F88E2ECDDB98" bold="true" box="[877,904,1903,1927]" pageId="26" pageNumber="27">dv</emphasis>
|
||
)—distoventrolateral process of femur, (
|
||
<emphasis id="B9036429FFE8D710FA9FF88E2872DB98" bold="true" box="[1307,1335,1903,1927]" pageId="26" pageNumber="27">lsc</emphasis>
|
||
)—sternal median longitudinal sulcus, (
|
||
<emphasis id="B9036429FFE8D710FE3BF8722C95DBB4" bold="true" box="[447,464,1939,1963]" pageId="26" pageNumber="27">sl</emphasis>
|
||
)—cephalic median longitudinal suture, (
|
||
<emphasis id="B9036429FFE8D710FCF7F8722ECFDBB4" bold="true" box="[883,906,1939,1963]" pageId="26" pageNumber="27">pf</emphasis>
|
||
)—ultimate prefemur, (
|
||
<emphasis id="B9036429FFE8D710FBF2F87229CADBB4" bold="true" box="[1142,1167,1939,1963]" pageId="26" pageNumber="27">pr</emphasis>
|
||
)—trochantero-prefemoral process, (
|
||
<emphasis id="B9036429FFE8D710FF7DF8562C4BDBD0" bold="true" box="[249,270,1975,1999]" pageId="26" pageNumber="27">sc</emphasis>
|
||
)—spine at place of coxopleural process, (
|
||
<emphasis id="B9036429FFE8D710FD38F8562F91DBD0" bold="true" box="[700,724,1975,1999]" pageId="26" pageNumber="27">su</emphasis>
|
||
)—spines of ultimate prefemur, (
|
||
<emphasis id="B9036429FFE8D710FBA6F8562905DBD0" bold="true" box="[1058,1088,1975,1999]" pageId="26" pageNumber="27">T1</emphasis>
|
||
)—tergite 1, (
|
||
<emphasis id="B9036429FFE8D710FB4FF85629A5DBD0" bold="true" box="[1227,1248,1975,1999]" pageId="26" pageNumber="27">t1</emphasis>
|
||
)—tarsus 1, (
|
||
<emphasis id="B9036429FFE8D710FAE1F856283EDBD0" bold="true" box="[1381,1403,1975,1999]" pageId="26" pageNumber="27">t2</emphasis>
|
||
)— tarsus 2, (
|
||
<emphasis id="B9036429FFE8D710FF7EF83A2C4CDBEC" bold="true" box="[250,265,2011,2035]" pageId="26" pageNumber="27">ti</emphasis>
|
||
)—tibia, (
|
||
<emphasis id="B9036429FFE8D710FEEAF83A2CC0DBEC" bold="true" box="[366,389,2011,2035]" pageId="26" pageNumber="27">tp</emphasis>
|
||
)—forcipular tooth-plate, (
|
||
<emphasis id="B9036429FFE8D710FD17F83A2FF6DBEC" bold="true" box="[659,691,2011,2035]" pageId="26" pageNumber="27">ust</emphasis>
|
||
)—ultimate sternite, (
|
||
<emphasis id="B9036429FFE8D710FC0AF83A2EE0DBEC" bold="true" box="[910,933,2011,2035]" pageId="26" pageNumber="27">ut</emphasis>
|
||
)—ultimate tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC8B83BFFE9D711FF43FF3E2D97DD02" blockId="27.[151,1437,150,501]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9036429FFE9D711FF43FF3E2CFFDCE6" bold="true" box="[199,442,222,249]" pageId="27" pageNumber="28">Number of subtaxa.</emphasis>
|
||
11 genera (“More than 220 species in 12 genera” sensu
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FBD0FF3E2D80DD02" author="Edgecombe, G. D. & Bonato, L." pageId="27" pageNumber="28" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato 2011: 397</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFE9D711FF43FEC72C34DD79" blockId="27.[151,1437,150,501]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9036429FFE9D711FF43FEC72CFDDD5F" bold="true" box="[199,440,294,320]" pageId="27" pageNumber="28">Sexual dimorphism.</emphasis>
|
||
Present in a few species of both
|
||
<taxonomicName id="4C77C3B8FFE9D711FCB6FEC62E85DD5E" authorityName="Linnaeus" authorityYear="1758" box="[818,960,295,321]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE9D711FCB6FEC62E85DD5E" box="[818,960,295,321]" italics="true" pageId="27" pageNumber="28">Scolopendra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C77C3B8FFE9D711FC7CFEC629CADD5E" authorityName="Shelley" authorityYear="2008" box="[1016,1167,295,321]" class="Chilopoda" family="Scolopendridae" genus="Akymnopellis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE9D711FC7CFEC629CADD5E" box="[1016,1167,295,321]" italics="true" pageId="27" pageNumber="28">Akymnopellis</emphasis>
|
||
</taxonomicName>
|
||
and, likely, in
|
||
<taxonomicName id="4C77C3B8FFE9D711FABEFEC62DA5DD7A" authorityName="Schileyko" authorityYear="1992" class="Chilopoda" family="Cryptopidae" genus="Tonkinodentus" kingdom="Animalia" order="Scolopendromorpha" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9036429FFE9D711FABEFEC62DA5DD7A" italics="true" pageId="27" pageNumber="28">Tonkinodentus</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
<paragraph id="8BC8B83BFFE9D711FF43FE8E2E32DD96" blockId="27.[151,1437,150,501]" box="[199,887,367,393]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9036429FFE9D711FF43FE8E2C5FDD96" bold="true" box="[199,282,367,393]" pageId="27" pageNumber="28">Range.</emphasis>
|
||
All tropical, subtropical and warm temperate regions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36DEBB0FFE9D711FF43FE722C67DDEA" pageId="27" pageNumber="28" type="discussion">
|
||
<paragraph id="8BC8B83BFFE9D711FF43FE722C67DDEA" blockId="27.[151,1437,150,501]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9036429FFE9D711FF43FE722C7EDDB2" bold="true" box="[199,315,403,429]" pageId="27" pageNumber="28">Remarks.</emphasis>
|
||
Treated as a subfamily in
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FDECFE722EB4DDB2" author="Edgecombe, G. D. & Bonato, L." box="[616,1009,403,429]" pageId="27" pageNumber="28" refId="ref47498" refString="Edgecombe, G. D. & Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe & Bonato (2011: 397)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FC79FE72284CDDB2" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[1021,1289,403,429]" pageId="27" pageNumber="28" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
|
||
Vahtera
|
||
<emphasis id="B9036429FFE9D711FBDEFE7229D6DDB2" box="[1114,1171,403,429]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
(2012a: 4
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FA92FE7228DDDDB2" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[1302,1432,403,429]" pageId="27" pageNumber="28" pagination="225 - 248" refId="ref51336" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, 225 - 248. https: // doi. org / 10.1007 / s 00435 - 012 - 0157 - 0" type="journal article" year="2012">2012b: 238</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FF13FE562C49DDCE" author="Vahtera, V. & Edgecombe, G. D. & Giribet, G." box="[151,268,438,465]" pageId="27" pageNumber="28" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">2013: 579</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FEA4FE562F58DDCE" author="Schileyko, A." box="[288,541,439,465]" pageId="27" pageNumber="28" pagination="151 - 192" refId="ref50143" refString="Schileyko, A. (2014) A contribution to the centipede fauna of Venezuela (Chilopoda: Scolopendromorpha). Zootaxa, 3821 (1), 151 - 192. https: // doi. org / 10.11646 / zootaxa. 3821.2.1" type="journal article" year="2014">Schileyko (2014: 174)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FDADFE562ECEDDCE" author="Schileyko, A. & Stoev, P." box="[553,907,438,465]" pageId="27" pageNumber="28" pagination="247 - 280" refId="ref50391" refString="Schileyko, A. & Stoev, P. (2016) Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda). Zootaxa, 4147 (3), 247 - 280. https: // doi. org / 10.11646 / zootaxa. 4147.3.3" type="journal article" year="2016">Schileyko & Stoev (2016: 252)</bibRefCitation>
|
||
, Schileyko (2018: 69),
|
||
<bibRefCitation id="EFE6C5CAFFE9D711FB16FE562C5BDDEA" author="Schileyko, A. & Solovyeva, E." pageId="27" pageNumber="28" pagination="133 - 155" refId="ref50503" refString="Schileyko, A. & Solovyeva, E. (2019) On the taxonomic position of the enigmatic genus Tonkinodentus Schileyko, 1992 (Chilopoda: Scolopendromorpha): the first molecular data. ZooKeys, 840, 133 - 155. https: // doi. org / 10.3897 / zookeys. 840.33635" type="journal article" year="2019">Schileyko & Solovyeva (2019: 138)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |