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184 lines
18 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="B7B3D42B5970DE611E64CE89713C8C4B" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus oxyrhinus Seeley 1870, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="47" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="46" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044992" LSID="urn:lsid:plazi:treatment:B7B3D42B5970DE611E64CE89713C8C4B" httpUri="http://treatment.plazi.org/id/B7B3D42B5970DE611E64CE89713C8C4B" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection pageId="45" pageNumber="46" type="nomenclature">
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName LSID="B7B3D42B-5970-DE61-1E64-CE89713C8C4B" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus oxyrhinus" order="Pterosauria" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="oxyrhinus" status="(nomen dubium)">Ornithocheirus oxyrhinus Seeley, 1870</taxonomicName>
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<taxonomicNameLabel pageId="45" pageNumber="46">(nomen dubium)</taxonomicNameLabel>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. A-B Ornithocheirus oxyrhinus, holotype CAMSM B 54612 (Albian, Cambridge Greensand), anterior part of the rostrum. A ventral view B respective line drawing. C-E Ornithocheirus scaphorhynchus, holotype CAMSM B 54441 (Albian, Cambridge Greensand), anterior part of the rostrum C ventral view D respective line drawing E line drawing in posterior view. F-I Ornithocheirus tenuirostris, holotype CAMSM B 54584 (Albian, Cambridge Greensand), anterior part of the rostrum F right lateral view G respective line drawing H ventral view I respective line drawing. J-K Ornithocheirus xyphorhynchus, holotype (Albian, Cambridge Greensand), anterior part of the rostrum J lateral view K dorsal view. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. J and K from Seeley 1881." httpUri="https://binary.pensoft.net/fig/22087" pageId="45" pageNumber="46">Fig. 19A-B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="reference_group">
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ptenodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="oxyrhinus" status="(nomen dubium)">
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<normalizedToken originalValue="“Ptenodactylus”">"Ptenodactylus"</normalizedToken>
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oxyrhinus
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</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
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: p. xvi [disclaimed]
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</paragraph>
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="oxyrhinus" status="(nomen dubium)">Ornithocheirus oxyrhinus</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 117
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</paragraph>
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<paragraph pageId="45" pageNumber="46">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="oxyrhinus" status="(nomen dubium)">Ornithocheirus oxyrhinus</taxonomicName>
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Seeley:
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="holotype">
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<paragraph pageId="45" pageNumber="46">Holotype.</paragraph>
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<paragraph pageId="45" pageNumber="46">
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CAMSM B 54612, anterior fragment of an upper jaw (
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. A-B Ornithocheirus oxyrhinus, holotype CAMSM B 54612 (Albian, Cambridge Greensand), anterior part of the rostrum. A ventral view B respective line drawing. C-E Ornithocheirus scaphorhynchus, holotype CAMSM B 54441 (Albian, Cambridge Greensand), anterior part of the rostrum C ventral view D respective line drawing E line drawing in posterior view. F-I Ornithocheirus tenuirostris, holotype CAMSM B 54584 (Albian, Cambridge Greensand), anterior part of the rostrum F right lateral view G respective line drawing H ventral view I respective line drawing. J-K Ornithocheirus xyphorhynchus, holotype (Albian, Cambridge Greensand), anterior part of the rostrum J lateral view K dorsal view. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge, sul - sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. J and K from Seeley 1881." httpUri="https://binary.pensoft.net/fig/22087" pageId="45" pageNumber="46">Fig. 19A-B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="type locality">
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<paragraph pageId="45" pageNumber="46">Type locality.</paragraph>
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<paragraph pageId="45" pageNumber="46">Smithswashing, Coton, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="46" type="type horizon">
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<paragraph pageId="45" pageNumber="46">Type horizon.</paragraph>
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<paragraph pageId="45" pageNumber="46">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
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<paragraph pageId="45" pageNumber="46">Description.</paragraph>
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<paragraph pageId="46" pageNumber="47">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="oxyrhinus">
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<emphasis italics="true" pageId="46" pageNumber="47">
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<pageBreakToken pageId="46" pageNumber="47" start="start">Ornithocheirus</pageBreakToken>
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oxyrhinus
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</emphasis>
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</taxonomicName>
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was described by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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based on CAMSM B 54612, a portion of an upper jaw, with a palatal ridge and some alveoli. However, the specimen is incomplete and lacks any features that justify recognition of a distinct species. We here regard it as a
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<emphasis italics="true" pageId="46" pageNumber="47">nomen dubium</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="46" pageNumber="47" type="remarks">
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<paragraph pageId="46" pageNumber="47">Remarks.</paragraph>
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<paragraph pageId="46" pageNumber="47">
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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classified it as a
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<emphasis italics="true" pageId="46" pageNumber="47">nomen nudum</emphasis>
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; however, it has a proper description (like most of the new species described by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley in 1870</bibRefCitation>
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) so it is not technically a
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<emphasis italics="true" pageId="46" pageNumber="47">nomen nudum</emphasis>
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.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/22087" pageId="46" pageNumber="47" start="Figure 19" startId="F19">
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<paragraph pageId="46" pageNumber="47">
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<emphasis bold="true" pageId="46" pageNumber="47">Figure 19.</emphasis>
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<emphasis bold="true" pageId="46" pageNumber="47">A-B</emphasis>
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="oxyrhinus">
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<emphasis italics="true" pageId="46" pageNumber="47">Ornithocheirus oxyrhinus</emphasis>
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</taxonomicName>
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, holotype CAMSM B 54612 (Albian, Cambridge Greensand), anterior part of the rostrum.
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<emphasis bold="true" pageId="46" pageNumber="47">A</emphasis>
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ventral view
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<emphasis bold="true" pageId="46" pageNumber="47">B</emphasis>
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respective line drawing.
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<emphasis bold="true" pageId="46" pageNumber="47">C-E</emphasis>
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<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="scaphorhynchus">
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<emphasis italics="true" pageId="46" pageNumber="47">Ornithocheirus scaphorhynchus</emphasis>
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</taxonomicName>
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, holotype CAMSM B 54441 (Albian, Cambridge Greensand), anterior part of the rostrum
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<emphasis bold="true" pageId="46" pageNumber="47">C</emphasis>
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ventral view
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<emphasis bold="true" pageId="46" pageNumber="47">D</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="46" pageNumber="47">E</emphasis>
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line drawing in posterior view.
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<emphasis bold="true" pageId="46" pageNumber="47">F-I</emphasis>
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<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="tenuirostris">
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<emphasis italics="true" pageId="46" pageNumber="47">Ornithocheirus tenuirostris</emphasis>
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</taxonomicName>
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, holotype CAMSM B 54584 (Albian, Cambridge Greensand), anterior part of the rostrum
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<emphasis bold="true" pageId="46" pageNumber="47">F</emphasis>
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right lateral view
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<emphasis bold="true" pageId="46" pageNumber="47">G</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="46" pageNumber="47">H</emphasis>
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ventral view
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<emphasis bold="true" pageId="46" pageNumber="47">I</emphasis>
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respective line drawing.
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<emphasis bold="true" pageId="46" pageNumber="47">J-K</emphasis>
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<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="xyphorhynchus">
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<emphasis italics="true" pageId="46" pageNumber="47">Ornithocheirus xyphorhynchus</emphasis>
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</taxonomicName>
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, holotype (Albian, Cambridge Greensand), anterior part of the rostrum
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<emphasis bold="true" pageId="46" pageNumber="47">J</emphasis>
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lateral view
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<emphasis bold="true" pageId="46" pageNumber="47">K</emphasis>
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dorsal view. Abbreviations:
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<emphasis bold="true" pageId="46" pageNumber="47">m</emphasis>
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- maxillae,
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<emphasis bold="true" pageId="46" pageNumber="47">pm</emphasis>
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- premaxillae,
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<emphasis bold="true" pageId="46" pageNumber="47">prid</emphasis>
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- palatal ridge,
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<emphasis bold="true" pageId="46" pageNumber="47">sul</emphasis>
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- sulcus. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. J and K from
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Geological Magazine" pageId="69" pageNumber="70" pagination="13 - 20" publicationUrl="10.1017/S0016756800113007" refId="B80" refString="Seeley, HG, 1881. On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F.G.S. Geological Magazine 8: 13 - 20, 10.1017/S0016756800113007" title="On evidence of two ornithosaurians referable to the genus Ornithocheirus, from the Upper Greensand of Cambridge, preserved in the collection of W. Reed, Esq., F. G. S." url="10.1017/S0016756800113007" volume="8" year="1881">Seeley 1881</bibRefCitation>
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.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |