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<document id="AD046D827110AD538DB91ED17479D222" ID-DOI="10.5281/zenodo.173802" ID-GBIF-Dataset="0a6c9168-e24b-4d02-955f-29d133c8890a" ID-ISSN="1175­5326" ID-Zenodo-Dep="173802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459784372853" checkinUser="plazi" docAuthor="Bousquet, Yves" docDate="2006" docId="B74587BDFF84FFCED36C3E789C32E497" docLanguage="en" docName="zt01308p029.pdf" docOrigin="Zootaxa 1308" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Semiardistomis puncticollis Dejean 1831" docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="4B7CFFC5FF8FFFC1D264384E9E5CE171" masterDocTitle="Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico" masterLastPageNumber="29" masterPageNumber="1" pageNumber="12" updateTime="1698218318931" updateUser="plazi">
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<mods:title id="B543491E47CD75C6C5F9AA79504EECFA">Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico</mods:title>
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<treatment id="B74587BDFF84FFCED36C3E789C32E497" ID-DOI="http://doi.org/10.5281/zenodo.6262687" ID-GBIF-Taxon="119348758" ID-Zenodo-Dep="6262687" LSID="urn:lsid:plazi:treatment:B74587BDFF84FFCED36C3E789C32E497" httpUri="http://treatment.plazi.org/id/B74587BDFF84FFCED36C3E789C32E497" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="77F66520FF84FFCAD36C3E789FE0E702" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="3F5336ABFF84FFCAD36C3E789D4DE721" blockId="11.[264,785,1590,1651]" box="[264,785,1590,1616]" pageId="11" pageNumber="12">
<heading id="641B81C7FF84FFCAD36C3E789D4DE721" bold="true" box="[264,785,1590,1616]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="F8EC4D28FF84FFCAD36C3E789D4DE721" ID-CoL="4WG3W" authority="Dejean, 1831" authorityName="Dejean" authorityYear="1831" box="[264,785,1590,1616]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF84FFCAD36C3E789D4DE721" bold="true" box="[264,785,1590,1616]" pageId="11" pageNumber="12">
<emphasis id="0D98EAB9FF84FFCAD36C3E789C09E721" bold="true" box="[264,597,1590,1616]" italics="true" pageId="11" pageNumber="12">Semiardistomis puncticollis</emphasis>
(
<bibRefCitation id="5B7D4B5AFF84FFCAD0003E789D55E721" author="Dejean" box="[612,777,1590,1616]" pageId="11" pageNumber="27" refString="Dejean, P. F. M. A. (1831) Species general des Coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris, 883 pp." type="book" year="1831">Dejean, 1831</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3F5336ABFF84FFCAD36C3E179FE0E702" blockId="11.[264,785,1590,1651]" box="[264,444,1625,1651]" pageId="11" pageNumber="12">
(
<figureCitation id="A7D72A2EFF84FFCAD3743E179F00E702" box="[272,348,1625,1651]" captionStart="FIGURES 4 – 8" captionStartId="12.[264,382,1796,1820]" captionTargetBox="[361,1232,331,1758]" captionTargetId="figure@12.[335,1252,306,1791]" captionTargetPageId="12" captionText="FIGURES 4 – 8. Median lobe (a) and paremeres (b). 4, Aspidoglossa subangulata; 5, Semiardistomis viridis; 6, S. puncticollis; 7, Ardistomis schaumii; 8, A. obliquatus." httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="11" pageNumber="12">Figs 6</figureCitation>
,
<figureCitation id="A7D72A2EFF84FFCAD30D3E179FD5E702" box="[361,393,1625,1651]" captionStart="FIGURES 11 – 18. 11 – 14" captionStartId="14.[264,382,1651,1675]" captionTargetBox="[372,1224,323,1650]" captionTargetId="figure@14.[356,1238,316,1655]" captionTargetPageId="14" captionText="FIGURES 11 – 18. 11 – 14. Apical stylomere. 11, Aspidoglossa subangulata; 12, Semiardistomis viridis; 13, Ardistomis schaumii; 14, A. obliquatus. 15 – 18. Spermatheca. 15, Aspidoglossa subangulata; 16, Semiardistomis viridis; 17, S. puncticollis; 18, Ardistomis schaumii." httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="11" pageNumber="12">17</figureCitation>
,
<figureCitation id="A7D72A2EFF84FFCAD3F13E179FEFE702" box="[405,435,1625,1651]" captionStart="FIGURE 33" captionStartId="20.[264,368,1030,1054]" captionTargetBox="[338,1249,326,1006]" captionTargetId="figure@20.[338,1249,326,1006]" captionTargetPageId="20" captionText="FIGURE 33. Collection localities for Semiardistomis puncticollis (Dejean)." httpUri="https://zenodo.org/record/173813/files/figure.png" pageId="11" pageNumber="12">33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF84FFCAD36C3EE79A72E631" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="3F5336ABFF84FFCAD36C3EE79DBDE791" blockId="11.[264,1323,1705,1856]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="1FFC1185FF84FFCAD36C3EE79DBDE791" pageId="11" pageNumber="12">
<taxonomicName id="F8EC4D28FF84FFCAD36C3EE79CFBE7B1" ID-CoL="5ZC3Q" authority="Dejean 1831: 508" authorityName="Dejean" authorityPageNumber="508" authorityYear="1831" box="[264,679,1705,1728]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF84FFCAD36C3EE79F87E7B1" box="[264,475,1705,1728]" italics="true" pageId="11" pageNumber="12">Clivina puncticollis</emphasis>
<treatmentCitation id="BE4D10BAFF84FFCAD3803EE79CFBE7B1" author="Dejean" box="[484,679,1705,1728]" page="508" pageId="11" pageNumber="12" year="1831">
<bibRefCitation id="5B7D4B5AFF84FFCAD3803EE79C2CE7B1" author="Dejean" box="[484,624,1705,1728]" pageId="11" pageNumber="27" refString="Dejean, P. F. M. A. (1831) Species general des Coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris, 883 pp." type="book" year="1831">Dejean 1831</bibRefCitation>
: 508
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="E0578809FF84FFCAD0D73EE79CB4E7B1" box="[691,744,1705,1728]" pageId="11" pageNumber="12">Type</typeStatus>
locality: &quot;Amérique septentrionale&quot;, herein restricted to Highlands Hammock State Park, Hardee Co., Florida,
<collectingCountry id="47FB763BFF84FFCAD1FC3E879DBDE791" box="[920,993,1737,1760]" name="United States of America" pageId="11" pageNumber="12">U.S.A.</collectingCountry>
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF84FFCAD36C3EA79D63E651" blockId="11.[264,1323,1705,1856]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="1FFC1185FF84FFCAD36C3EA79D63E651" pageId="11" pageNumber="12">
<taxonomicName id="F8EC4D28FF84FFCAD36C3EA79CB1E671" ID-CoL="6845C" authority="Putzeys (1846: 647)" authorityName="Putzeys" authorityPageNumber="647" authorityYear="1846" box="[264,749,1769,1792]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF84FFCAD36C3EA79FA3E671" box="[264,511,1769,1792]" italics="true" pageId="11" pageNumber="12">Ardistomis puncticollis</emphasis>
:
<treatmentCitation id="BE4D10BAFF84FFCAD06B3EA79CB1E671" author="Putzeys" box="[527,749,1769,1792]" page="647" pageId="11" pageNumber="12" year="1846">
<bibRefCitation id="5B7D4B5AFF84FFCAD06B3EA79CB1E671" author="Putzeys" box="[527,749,1769,1792]" pageId="11" pageNumber="29" refString="Putzeys, J. A. A. H. (1846) Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege, 2, 521 - 663." type="journal article" year="1846">Putzeys (1846: 647)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="BE4D10BAFF84FFCAD09D3EA79D8DE671" author="LeConte" box="[761,977,1769,1792]" page="80" pageId="11" pageNumber="12" year="1857">
<bibRefCitation id="5B7D4B5AFF84FFCAD09D3EA79D8DE671" author="LeConte" box="[761,977,1769,1792]" pageId="11" pageNumber="28" refString="LeConte, J. L. (1857) Synopsis of the species Clivina and allied genera inhabiting the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, [9], 75 - 83." type="journal article" year="1857">LeConte (1857: 80)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="BE4D10BAFF84FFCAD1BB3EA79AEBE671" author="LeConte" box="[991,1207,1769,1792]" page="32" pageId="11" pageNumber="12" year="1879">
<bibRefCitation id="5B7D4B5AFF84FFCAD1BB3EA79AEBE671" author="LeConte" box="[991,1207,1769,1792]" pageId="11" pageNumber="28" refString="LeConte, J. L. (1879) Synoptic tables: Ardistomis, Putz.; Aspidoglossa, Putz.; Clivina, Latr.; Schizogenius, Putz. Bulletin of the Brooklyn Entomological Society, 2, 32 - 34." type="journal article" year="1879">LeConte (1879: 32)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="BE4D10BAFF84FFCAD6A03EA79FF7E651" author="Blatchley" page="63" pageId="11" pageNumber="12" year="1910">
<bibRefCitation id="5B7D4B5AFF84FFCAD6A03EA79FF7E651" author="Blatchley" pageId="11" pageNumber="26" refString="Blatchley, W. S. (1910) An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana - with bibliography and descriptions of new species. The Nature Publishing Co., Indianapolis, 1386 pp." type="book" year="1910">Blatchley (1910: 63)</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="5B7D4B5AFF84FFCAD3D23F479D67E651" author="Downie" box="[438,827,1801,1824]" pageId="11" pageNumber="27" refString="Downie, N. M, &amp; Arnett, RH. Jr. (1996) The beetles of northeastern North America. Volume I: Introduction; suborders Archostemata, Adephaga, and Polyphaga, thru superfamily Cantharoidea. The Sandhill Crane Press, Gainesville, 880 pp." type="book" year="1996">Downie and Arnett (1996: 111, 112)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3F5336ABFF84FFCAD36C3F679A72E631" blockId="11.[264,1323,1705,1856]" box="[264,1070,1833,1856]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="1FFC1185FF84FFCAD36C3F679A72E631" box="[264,1070,1833,1856]" pageId="11" pageNumber="12">
<taxonomicName id="F8EC4D28FF84FFCAD36C3F679CCCE631" ID-CoL="4WG3W" authority="Nichols" authorityName="Nichols" box="[264,656,1833,1856]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF84FFCAD36C3F679C72E631" box="[264,558,1833,1856]" italics="true" pageId="11" pageNumber="12">Semiardistomis puncticollis</emphasis>
:
<bibRefCitation id="5B7D4B5AFF84FFCAD0593F679CCCE631" author="Nichols" box="[573,656,1833,1856]" lastPageId="11" lastPageNumber="29" pageId="11" pageNumber="28" refString="Nichols, S. W. (1988 a) Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Ph. D. thesis, Cornell University, 393 pp." type="book" year="1988" yearSuffix="a">Nichols</bibRefCitation>
</taxonomicName>
(1988a: 187, 192);
<treatmentCitation id="BE4D10BAFF84FFCAD1073F679A76E631" author="Ciegler" box="[867,1066,1833,1856]" page="43" pageId="11" pageNumber="12" year="2000">
<bibRefCitation id="5B7D4B5AFF84FFCAD1073F679A76E631" author="Ciegler" box="[867,1066,1833,1856]" pageId="11" pageNumber="27" refString="Ciegler, J. C. (2000). Ground beetles and wrinkled bark beetles of South Carolina (Coleoptera: Geadephaga: Carabidae and Rhysodidae). Biota of South Carolina. Vol. 1. South Carolina: Clemson University, Clemson, 149 pp." type="book" year="2000">Ciegler (2000: 43)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="6B936623FF83FFCDD36C3F4A9A2AE64F" httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="12" pageNumber="13" targetBox="[361,1232,331,1758]" targetPageId="12">
<paragraph id="3F5336ABFF83FFCDD36C3F4A9A2AE64F" blockId="12.[264,1322,1796,1854]" pageId="12" pageNumber="13">
<emphasis id="0D98EAB9FF83FFCDD36C3F4A9F9EE66D" bold="true" box="[264,450,1796,1820]" pageId="12" pageNumber="13">FIGURES 4–8.</emphasis>
Median lobe (a) and paremeres (b). 4,
<taxonomicName id="F8EC4D28FF83FFCDD1B03F4A9AA7E66A" box="[980,1275,1796,1819]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF83FFCDD1B03F4A9AA7E66A" box="[980,1275,1796,1819]" italics="true" pageId="12" pageNumber="13">Aspidoglossa subangulata</emphasis>
</taxonomicName>
; 5,
<taxonomicName id="F8EC4D28FF83FFCDD36C3F699FAAE64F" box="[264,502,1831,1854]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="0D98EAB9FF83FFCDD36C3F699FAAE64F" box="[264,502,1831,1854]" italics="true" pageId="12" pageNumber="13">Semiardistomis viridis</emphasis>
</taxonomicName>
; 6,
<taxonomicName id="F8EC4D28FF83FFCDD07B3F699CEAE64F" box="[543,694,1831,1854]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF83FFCDD07B3F699CEAE64F" box="[543,694,1831,1854]" italics="true" pageId="12" pageNumber="13">S. puncticollis</emphasis>
</taxonomicName>
; 7,
<taxonomicName id="F8EC4D28FF83FFCDD0BB3F699DE4E64F" box="[735,952,1831,1854]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF83FFCDD0BB3F699DE4E64F" box="[735,952,1831,1854]" italics="true" pageId="12" pageNumber="13">Ardistomis schaumii</emphasis>
</taxonomicName>
; 8,
<taxonomicName id="F8EC4D28FF83FFCDD1853F699A33E64F" box="[993,1135,1831,1854]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="0D98EAB9FF83FFCDD1853F699A33E64F" box="[993,1135,1831,1854]" italics="true" pageId="12" pageNumber="13">A. obliquatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="6B936623FF82FFCCD36C3C0C9B43E528" box="[264,1311,1090,1114]" httpUri="https://zenodo.org/record/173806/files/figure.png" pageId="13" pageNumber="14" targetBox="[332,1271,322,1062]" targetPageId="13">
<paragraph id="3F5336ABFF82FFCCD36C3C0C9B43E528" blockId="13.[264,1311,1090,1114]" box="[264,1311,1090,1114]" pageId="13" pageNumber="14">
<emphasis id="0D98EAB9FF82FFCCD36C3C0C9F9DE52B" bold="true" box="[264,449,1090,1114]" pageId="13" pageNumber="14">FIGURES 9–10.</emphasis>
Sclerotized structures of endophallus. 9,
<taxonomicName id="F8EC4D28FF82FFCCD11F3C0C9A38E528" box="[891,1124,1090,1113]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="0D98EAB9FF82FFCCD11F3C0C9A38E528" box="[891,1124,1090,1113]" italics="true" pageId="13" pageNumber="14">Ardistomis obliquatus</emphasis>
</taxonomicName>
; 10,
<taxonomicName id="F8EC4D28FF82FFCCD6FE3C0C9B44E528" box="[1178,1304,1090,1113]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF82FFCCD6FE3C0C9B44E528" box="[1178,1304,1090,1113]" italics="true" pageId="13" pageNumber="14">A. schaumii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="77F66520FF82FFCCD36C3CC29DF7E4E6" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="3F5336ABFF82FFCCD36C3CC29DF7E4E6" blockId="13.[264,1324,1164,1871]" pageId="13" pageNumber="14">
<emphasis id="0D98EAB9FF82FFCCD36C3CC29FE0E5D7" bold="true" box="[264,444,1164,1190]" pageId="13" pageNumber="14">
<typeStatus id="E0578809FF82FFCCD36C3CC29F19E5D7" box="[264,325,1164,1190]" pageId="13" pageNumber="14">Type</typeStatus>
Material
</emphasis>
. Dejean originally described this species from an unspecified number of specimens. According to
<bibRefCitation id="5B7D4B5AFF82FFCCD02F3CFB9D17E5BE" author="Putzeys" box="[587,843,1205,1231]" pageId="13" pageNumber="29" refString="Putzeys, J. A. A. H. (1846) Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des scaritides. Memoires de la Societe Royale des Sciences de Liege, 2, 521 - 663." type="journal article" year="1846">Putzeys (1846: 647)</bibRefCitation>
, Dejean's collection includes a single specimen under that name. The
<typeStatus id="E0578809FF82FFCCD0FB3C939CA1E586" box="[671,765,1245,1271]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
, a female located in
<collectionCode id="59FDAE6EFF82FFCCD6743C939A39E586" box="[1040,1125,1245,1271]" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="13" pageNumber="14">MHNP</collectionCode>
, is labelled &quot;36 [handwritten]/
<taxonomicName id="F8EC4D28FF82FFCCD3A33D4B9CB8E46E" authority="Dej. Am." authorityName="Dej. Am." box="[455,740,1285,1311]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="puncticollis">puncticollis Dej. Am.</taxonomicName>
bor. Dej. [handwritten]/
<typeStatus id="E0578809FF82FFCCD62C3D4B9AE1E46E" box="[1096,1213,1285,1311]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="F8EC4D28FF82FFCCD6B03D4B9F8EE436" authority="Dej." authorityName="Dej." class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="puncticollis">Clivina puncticollis Dej.</taxonomicName>
des. S.W. Nichols 1984&quot; (
<emphasis id="0D98EAB9FF82FFCCD1553D639D01E437" box="[817,861,1325,1350]" italics="true" pageId="13" pageNumber="14">fide</emphasis>
<bibRefCitation id="5B7D4B5AFF82FFCCD1083D639A7FE436" author="Nichols" box="[876,1059,1325,1351]" lastPageId="13" lastPageNumber="29" pageId="13" pageNumber="28" refString="Nichols, S. W. (1988 a) Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Ph. D. thesis, Cornell University, 393 pp." type="book" year="1988" yearSuffix="a">Nichols 1988a</bibRefCitation>
: 192). The
<typeStatus id="E0578809FF82FFCCD6DB3D639B76E436" box="[1215,1322,1325,1351]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
designation is hereby published. This species was not surveyed by
<bibRefCitation id="5B7D4B5AFF82FFCCD64E3D1B9AB7E41E" author="Lindroth" box="[1066,1259,1365,1391]" pageId="13" pageNumber="28" refString="Lindroth, C. H. (1955) Dejean's types of North American Carabidae (Col.). Opuscula Entomologica, 20, 10 - 34." type="journal article" year="1955">Lindroth (1955)</bibRefCitation>
, who reported on Dejean's
<typeStatus id="E0578809FF82FFCCD0643D339C62E4E6" box="[512,574,1405,1431]" pageId="13" pageNumber="14">types</typeStatus>
of North American
<taxonomicName id="F8EC4D28FF82FFCCD1483D339DFBE4E6" box="[812,935,1405,1431]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF82FFCCD35C3DEA9DDAE496" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="3F5336ABFF82FFCCD35C3DEA9DDAE496" blockId="13.[264,1324,1164,1871]" pageId="13" pageNumber="14">
<emphasis id="0D98EAB9FF82FFCCD35C3DEA9FECE4CF" bold="true" box="[312,432,1444,1470]" pageId="13" pageNumber="14">Diagnosis</emphasis>
. Distinguished from all other Nearctic Ardistomina by the presence of extra supraorbital setae and of discal setae on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF82FFCED35C3DBA9C94E437" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="3F5336ABFF82FFCCD35C3DBA9C0FE656" blockId="13.[264,1324,1164,1871]" pageId="13" pageNumber="14">
<emphasis id="0D98EAB9FF82FFCCD35C3DBA9F95E77F" bold="true" box="[312,457,1524,1550]" pageId="13" pageNumber="14">Description</emphasis>
. Same character states as for
<taxonomicName id="F8EC4D28FF82FFCCD1563DBB9DC1E77F" box="[818,925,1525,1550]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="0D98EAB9FF82FFCCD1563DBB9DC1E77F" box="[818,925,1525,1550]" italics="true" pageId="13" pageNumber="14">S. viridis</emphasis>
</taxonomicName>
except for the followings. Head: Eyes slightly less globulose. Frons with five to seven supraorbital setae on each side. Mentum tooth not cariniform; setae behind tooth distant. Prothorax: Pronotum with more than ten lateral and sublateral setae; disc with setae. Elytra: Striae (except sutural stria) not impressed; striae 2 and 4 with more setae. Lateral margin not prolonged in form of carina from basal extremity to base of stria 3. Male Genitalia: Median lobe more regularly curved in lateral view (
<figureCitation id="A7D72A2EFF82FFCCD3B83EAB9C72E78E" box="[476,558,1765,1791]" captionStart="FIGURES 4 – 8" captionStartId="12.[264,382,1796,1820]" captionTargetBox="[361,1232,331,1758]" captionTargetId="figure@12.[335,1252,306,1791]" captionTargetPageId="12" captionText="FIGURES 4 – 8. Median lobe (a) and paremeres (b). 4, Aspidoglossa subangulata; 5, Semiardistomis viridis; 6, S. puncticollis; 7, Ardistomis schaumii; 8, A. obliquatus." httpUri="https://zenodo.org/record/173805/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
). Female Genitalia: Seminal tube of spermatheca (
<figureCitation id="A7D72A2EFF82FFCCD6A73EAB9B7DE78E" box="[1219,1313,1765,1791]" captionStart="FIGURES 11 – 18. 11 – 14" captionStartId="14.[264,382,1651,1675]" captionTargetBox="[372,1224,323,1650]" captionTargetId="figure@14.[356,1238,316,1655]" captionTargetPageId="14" captionText="FIGURES 11 – 18. 11 – 14. Apical stylomere. 11, Aspidoglossa subangulata; 12, Semiardistomis viridis; 13, Ardistomis schaumii; 14, A. obliquatus. 15 – 18. Spermatheca. 15, Aspidoglossa subangulata; 16, Semiardistomis viridis; 17, S. puncticollis; 18, Ardistomis schaumii." httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
) somewhat widened apically.
</paragraph>
<paragraph id="3F5336ABFF82FFCCD35C3F7B9C37E63E" blockId="13.[264,1324,1164,1871]" box="[312,619,1845,1871]" pageId="13" pageNumber="14">Body length: 4.8–6.5 mm.</paragraph>
<caption id="6B936623FF81FFCFD36C3E3D9AC8E7A1" httpUri="https://zenodo.org/record/173807/files/figure.png" pageId="14" pageNumber="15" targetBox="[372,1224,323,1650]" targetPageId="14">
<paragraph id="3F5336ABFF81FFCFD36C3E3D9AC8E7A1" blockId="14.[264,1322,1651,1744]" pageId="14" pageNumber="15">
<emphasis id="0D98EAB9FF81FFCFD36C3E3D9F8EE7FA" bold="true" box="[264,466,1651,1675]" pageId="14" pageNumber="15">FIGURES 11–18.</emphasis>
11–14. Apical stylomere. 11,
<taxonomicName id="F8EC4D28FF81FFCFD14E3E3A9A14E7FA" box="[810,1096,1652,1675]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF81FFCFD14E3E3A9A14E7FA" box="[810,1096,1652,1675]" italics="true" pageId="14" pageNumber="15">Aspidoglossa subangulata</emphasis>
</taxonomicName>
; 12,
<taxonomicName id="F8EC4D28FF81FFCFD6E33E3A9F10E7DC" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="0D98EAB9FF81FFCFD6E33E3A9F10E7DC" italics="true" pageId="14" pageNumber="15">Semiardistomis viridis</emphasis>
</taxonomicName>
; 13,
<taxonomicName id="F8EC4D28FF81FFCFD3F13ED89C24E7DC" box="[405,632,1686,1709]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF81FFCFD3F13ED89C24E7DC" box="[405,632,1686,1709]" italics="true" pageId="14" pageNumber="15">Ardistomis schaumii</emphasis>
</taxonomicName>
; 14,
<taxonomicName id="F8EC4D28FF81FFCFD0A63ED89D06E7DC" box="[706,858,1686,1709]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="0D98EAB9FF81FFCFD0A63ED89D06E7DC" box="[706,858,1686,1709]" italics="true" pageId="14" pageNumber="15">A. obliquatus</emphasis>
</taxonomicName>
. 15–18. Spermatheca. 15,
<taxonomicName id="F8EC4D28FF81FFCFD6F83ED89FD0E7A1" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF81FFCFD6F83ED89FD0E7A1" italics="true" pageId="14" pageNumber="15">Aspidoglossa subangulata</emphasis>
</taxonomicName>
; 16,
<taxonomicName id="F8EC4D28FF81FFCFD3A63EF79CECE7A1" box="[450,688,1721,1744]" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="0D98EAB9FF81FFCFD3A63EF79CECE7A1" box="[450,688,1721,1744]" italics="true" pageId="14" pageNumber="15">Semiardistomis viridis</emphasis>
</taxonomicName>
; 17,
<taxonomicName id="F8EC4D28FF81FFCFD0823EF79D21E7A1" box="[742,893,1721,1744]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="0D98EAB9FF81FFCFD0823EF79D21E7A1" box="[742,893,1721,1744]" italics="true" pageId="14" pageNumber="15">S. puncticollis</emphasis>
</taxonomicName>
; 18,
<taxonomicName id="F8EC4D28FF81FFCFD1D73EF79AD0E7A1" box="[947,1164,1721,1744]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF81FFCFD1D73EF79AD0E7A1" box="[947,1164,1721,1744]" italics="true" pageId="14" pageNumber="15">Ardistomis schaumii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3F5336ABFF81FFCFD35C3F4D9F29E61C" blockId="14.[264,1323,1795,1901]" pageId="14" pageNumber="15">
<emphasis id="0D98EAB9FF81FFCFD35C3F4D9CD8E66C" bold="true" box="[312,644,1795,1821]" pageId="14" pageNumber="15">Geographical Distribution</emphasis>
. This species ranges from western Missouri to Virginia, south to Florida (not yet recorded from the Keys) and eastern Texas along the Rio Grande (
<figureCitation id="A7D72A2EFF81FFCFD3753F1D9F34E61C" box="[273,360,1875,1901]" captionStart="FIGURE 33" captionStartId="20.[264,368,1030,1054]" captionTargetBox="[338,1249,326,1006]" captionTargetId="figure@20.[338,1249,326,1006]" captionTargetPageId="20" captionText="FIGURE 33. Collection localities for Semiardistomis puncticollis (Dejean)." httpUri="https://zenodo.org/record/173813/files/figure.png" pageId="14" pageNumber="15">Fig. 33</figureCitation>
).
</paragraph>
<caption id="6B936623FF80FFCED36C3C3D9CADE5A1" httpUri="https://zenodo.org/record/173808/files/figure.png" pageId="15" pageNumber="16" targetBox="[269,1319,329,1113]" targetPageId="15">
<paragraph id="3F5336ABFF80FFCED36C3C3D9CADE5A1" blockId="15.[264,1322,1139,1232]" pageId="15" pageNumber="16">
<emphasis id="0D98EAB9FF80FFCED36C3C3D9F8CE5FA" bold="true" box="[264,464,1139,1163]" pageId="15" pageNumber="16">FIGURES 19–22.</emphasis>
19–21. Head (dorsal view). 19,
<taxonomicName id="F8EC4D28FF80FFCED1573C3A9A12E5FA" box="[819,1102,1140,1163]" class="Insecta" family="Carabidae" genus="Aspidoglossa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="subangulata">
<emphasis id="0D98EAB9FF80FFCED1573C3A9A12E5FA" box="[819,1102,1140,1163]" italics="true" pageId="15" pageNumber="16">Aspidoglossa subangulata</emphasis>
</taxonomicName>
; 20,
<taxonomicName id="F8EC4D28FF80FFCED6E33C3A9F10E5DF" class="Insecta" family="Carabidae" genus="Semiardistomis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="0D98EAB9FF80FFCED6E33C3A9F10E5DF" italics="true" pageId="15" pageNumber="16">Semiardistomis viridis</emphasis>
</taxonomicName>
; 21,
<taxonomicName id="F8EC4D28FF80FFCED3E63CD99C07E5DF" box="[386,603,1175,1198]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF80FFCED3E63CD99C07E5DF" box="[386,603,1175,1198]" italics="true" pageId="15" pageNumber="16">Ardistomis schaumii</emphasis>
</taxonomicName>
. 22, Extremity of body (lateral view) of
<taxonomicName id="F8EC4D28FF80FFCED66F3CD99AB8E5DF" box="[1035,1252,1175,1198]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="0D98EAB9FF80FFCED66F3CD99AB8E5DF" box="[1035,1252,1175,1198]" italics="true" pageId="15" pageNumber="16">Ardistomis schaumii</emphasis>
</taxonomicName>
. Scale bar = 0.2 mm (Figs 20–22), 0.5 mm (Fig. 19).
</paragraph>
</caption>
<paragraph id="3F5336ABFF80FFCED35C3D4D9C94E437" blockId="15.[264,1323,1283,1510]" pageId="15" pageNumber="16">
<emphasis id="0D98EAB9FF80FFCED35C3D4D9FC4E46C" bold="true" box="[312,408,1283,1309]" pageId="15" pageNumber="16">Habitat</emphasis>
. Adults of this species were collected on eroding river bank in northern Florida (
<bibRefCitation id="5B7D4B5AFF80FFCED3083D629CE7E437" author="Choate" box="[364,699,1324,1350]" pageId="15" pageNumber="27" refString="Choate, P. M. &amp; Rogers, T. (1976) The occurrence of a troglophilic ground beetle in Florida (Coleoptera: Carabidae). The Coleopterists Bulletin, 30, 364." type="journal article" year="1976">Choate &amp; Rogers 1976: 364</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="77F66520FF80FFCED35C3D1D9C32E497" pageId="15" pageNumber="16" type="discussion">
<paragraph id="3F5336ABFF80FFCED35C3D1D9C32E497" blockId="15.[264,1323,1283,1510]" pageId="15" pageNumber="16">
<emphasis id="0D98EAB9FF80FFCED35C3D1D9FE3E41C" bold="true" box="[312,447,1363,1389]" pageId="15" pageNumber="16">Comments</emphasis>
.
<bibRefCitation id="5B7D4B5AFF80FFCED3AB3D1A9D5AE41F" author="Nichols" box="[463,774,1364,1390]" lastPageId="15" lastPageNumber="29" pageId="15" pageNumber="28" refString="Nichols, S. W. (1988 a) Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Ph. D. thesis, Cornell University, 393 pp." type="book" year="1988" yearSuffix="a">Nichols (1988a: 193, 195)</bibRefCitation>
reported that this species is &quot;brachypterous or fully­winged&quot; and that the species was &quot;relatively common at a blacklight operated just outside the laboratory building at Archbold Biological Station&quot;. I did not investigate the wing condition of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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