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<document id="A9797CB2740987684B5673E3950556C4" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFFD3FFCF3C2C60AAFB4C710C" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trechepaphiopsis unisetosa 2004" docType="treatment" docVersion="2" lastPageNumber="411" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="411" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2DB3C6F0F99AD4B998094FCF9A123A85">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:role id="EDFDBC7F0482232B78A6094EE9012551">
<mods:roleTerm id="02FA9282D65D0A9DB1CD2DF831923376">Author</mods:roleTerm>
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<mods:namePart id="6055FD181352723D5AA3E32E60DA1088">Deuve, Thierry</mods:namePart>
<mods:affiliation id="5481003C0A694B721C65CE42B46EF234">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National d’Histoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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<mods:namePart id="492D6E7C832BD35DAED3260397C25C4A">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="43E19E283515E89D81A9C72253C044F7">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="70A1BB35FA12FC519075D543FD8EA468" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="36B9BE9FB8A8504568A1422B2C3A3C09">Liang, Hongbin</mods:namePart>
<mods:affiliation id="B2D3D847EF7AE2200A33267954C9AA7B">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:title id="344D6AE1B591EDC2213CACB08B6E8A85">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="9BFAA0BB7F0CD84962F87F75FE0E7F6A">
<mods:date id="A54357FC54CA8BD779B25DBED89FB6EC">2016</mods:date>
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<mods:number id="075A0B84BCCDD296172D9406A88DD24C">2016-10-14</mods:number>
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<mods:detail id="9BA2BE7B97403F5EF4FA6B31C8EDF199" type="volume">
<mods:number id="DEB617157104CEFC0E23D8865F7E7405">63</mods:number>
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<treatment id="03D6879DFFD3FFCF3C2C60AAFB4C710C" LSID="urn:lsid:plazi:treatment:03D6879DFFD3FFCF3C2C60AAFB4C710C" httpUri="http://treatment.plazi.org/id/03D6879DFFD3FFCF3C2C60AAFB4C710C" lastPageNumber="411" pageId="70" pageNumber="411">
<subSubSection id="C3656500FFD3FFCF3C2C60AAFD5277A3" box="[123,668,219,243]" pageId="70" pageNumber="411" type="nomenclature">
<paragraph id="8BC0368BFFD3FFCF3C2C60AAFD5277A3" blockId="70.[123,668,219,273]" box="[123,668,219,243]" pageId="70" pageNumber="411">
<heading id="D08881E7FFD3FFCF3C2C60AAFD5277A3" box="[123,668,219,243]" fontSize="10" level="2" pageId="70" pageNumber="411" reason="4">
<emphasis id="B90BEA99FFD3FFCF3C2C60AAFD5277A3" bold="true" box="[123,668,219,243]" pageId="70" pageNumber="411">
<taxonomicName id="4C7F4D08FFD3FFCF3C2C60AAFD8077A3" ID-CoL="8MH87" authority="(Deuve), 2004" authorityName="" authorityYear="2004" baseAuthorityName="Deuve" baseAuthorityYear="2004" box="[123,590,219,243]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD3FFCF3C2C60AAFE6577A3" bold="true" box="[123,427,219,243]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis unisetosa</emphasis>
(Deuve), 2004
</taxonomicName>
(sp. 2)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD3FFCF3C2C608AFE4F7641" box="[123,385,251,273]" pageId="70" pageNumber="411" type="description">
<paragraph id="8BC0368BFFD3FFCF3C2C608AFE4F7641" blockId="70.[123,668,219,273]" box="[123,385,251,273]" pageId="70" pageNumber="411">
(
<figureCitation id="13442A0EFFD3FFCF3CD5608AFF1D7641" box="[130,211,251,273]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Figs. 27</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3C8A608DFECC7641" box="[221,258,252,273]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="70" pageNumber="411">37a</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3D5B608AFEFD7641" box="[268,307,251,273]" captionStart="FIGURE" captionStartId="105.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1606]" captionTargetId="figure-15@105.[120,1176,216,1607]" captionTargetPageId="105" captionText="FIGURE 43. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Luzhang Township, Piana Road 0.1 km E of Fengxue Yakou, elevation 3150 m; habitat in which specimens of Trechus luzhangensis sp. nov. were collected. b. Lushui County, Luzhang Township, 100 m S of Fengxue Yakou on east side of pass, elevation 3150 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve) were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155372" httpUri="https://zenodo.org/record/13155372/files/figure.png" pageId="70" pageNumber="411">43b</figureCitation>
,
<figureCitation id="13442A0EFFD3FFCF3D6A608AFEB57641" box="[317,379,251,273]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="70" pageNumber="411">46–48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD3FFCF3C2C6156FD68762A" pageId="70" pageNumber="411" type="reference_group">
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<treatmentCitationGroup id="AB6F11A5FFD3FFCF3C2C6156FD20760B" pageId="70" pageNumber="411">
<treatmentCitation id="0ADE109AFFD3FFCF3C2C6156FE2F766C" author="DEUVE, T." box="[123,481,295,317]" page="220" pageId="70" pageNumber="411" year="2004">
<taxonomicName id="4C7F4D08FFD3FFCF3C2C6156FE2F766C" ID-CoL="8LP6C" authority="Deuve, 2004: 220" authorityName="Deuve" authorityPageNumber="220" authorityYear="2004" box="[123,481,295,317]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosus">
<emphasis id="B90BEA99FFD3FFCF3C2C6156FEE1766D" box="[123,303,295,317]" italics="true" pageId="70" pageNumber="411">Trechus unisetosus</emphasis>
<materialsCitation id="3B173CD6FFD3FFCF3D636159FE2F766C" box="[308,481,295,317]" country="China" county="Lushui County" elevation="2650" location="Fengxue Yakou" municipality="Gaoligong Shan" pageId="70" pageNumber="411" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<bibRefCitation id="EFEE4B7AFFD3FFCF3D636159FE2F766C" author="DEUVE, T." box="[308,481,295,317]" pageId="70" pageNumber="411" pagination="215 - 234" refId="ref58945" refString="DEUVE, T. 2004. Nouveaux Trechidae (Trechinae et Bembidiinae) d'Afrique et d'Asie (Coleoptera, Caraboidea). Coleopteres 10: 215 - 234." type="journal article" year="2004">Deuve, 2004: 220</bibRefCitation>
</materialsCitation>
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</treatmentCitation>
.
<typeStatus id="54C48829FFD3FFCF3DBD6156FD88766D" box="[490,582,295,317]" pageId="70" pageNumber="411" type="holotype">Holotype</typeStatus>
, a female, in SCAU. Type locality:
<collectingCountry id="F368761BFFD3FFCF3FC06156FC1D766D" box="[919,979,295,317]" name="China" pageId="70" pageNumber="411">China</collectingCountry>
,
<collectingRegion id="49BBF869FFD3FFCF3F8B6159FBE6766D" box="[988,1064,296,317]" country="China" name="Yunnan" pageId="70" pageNumber="411">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BA4ACF1FFD3FFCF38676156FF13760C" pageId="70" pageNumber="411">Gaoligong Shan</collectingMunicipality>
,
<collectingCounty id="62A14E07FFD3FFCF3CB06137FEB7760B" box="[231,377,326,348]" pageId="70" pageNumber="411">Lushui County</collectingCounty>
,
<location id="8EA06050FFD3FFCF3DD56137FDD7760C" LSID="urn:lsid:plazi:treatment:03D6879DFFD3FFCF3C2C60AAFB4C710C:8EA06050FFD3FFCF3DD56137FDD7760C" box="[386,537,326,348]" country="China" county="Lushui County" municipality="Gaoligong Shan" name="Fengxue Yakou" pageId="70" pageNumber="411" stateProvince="Yunnan">Fengxue Yakou</location>
[Pass],
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<elevation id="0052D1B8FFD3FFCF3E316137FD25760B" box="[614,747,326,348]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="70" pageNumber="411" unit="m" value="2650.0" valueMax="2700.0" valueMin="2600.0">2600-2700 m</elevation>
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.
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</paragraph>
<paragraph id="8BC0368BFFD3FFCF3C2C6114FD68762A" blockId="70.[123,1173,295,379]" box="[123,678,357,379]" pageId="70" pageNumber="411">
<taxonomicName id="4C7F4D08FFD3FFCF3C2C6114FE12762A" ID-CoL="8MH87" authority="(Deuve)" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" box="[123,476,357,379]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa" status="comb. nov.">
<emphasis id="B90BEA99FFD3FFCF3C2C6114FE4B762B" box="[123,389,357,379]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis unisetosa</emphasis>
(Deuve)
</taxonomicName>
,
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<emphasis id="B90BEA99FFD3FFCF3DB16114FDD87629" bold="true" box="[486,534,357,378]" pageId="70" pageNumber="411">NEW</emphasis>
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<emphasis id="B90BEA99FFD3FFCF3E4B6114FD68762A" bold="true" box="[540,678,357,379]" pageId="70" pageNumber="411">COMBINATION</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD3FFCF3CFC61FEFE9875BD" pageId="70" pageNumber="411" type="diagnosis">
<paragraph id="8BC0368BFFD3FFCF3CFC61FEFE9875BD" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
<smallCapsWord id="8D26A057FFD3FFCF3CFC61FEFEEE76F5" baselines="417,416" box="[171,288,399,423]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3CFC61FEFEEE76F5" bold="true" box="[171,288,399,423]" pageId="70" pageNumber="411">DIAGNOSIS</emphasis>
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.— Adults of this species (
<figureCitation id="13442A0EFFD3FFCF3E1061FEFD6D76F7" box="[583,675,399,423]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
<quantity id="4C879B6EFFD3FFCF380061C1FF0176B8" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="70" pageNumber="411" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.1 to 3.5 mm</quantity>
), apterous; body color light brown, dorsum slightly iridescent; eyes small but convex; tempora very convex, sparsely pubescent; pronotum transverse (ratio PW/PL = 1.35), with basal angles obtuse and rounded; elytra convex, with median 2 or 3 striae deeply impressed, lateral striae faintly impressed to effaced; recurrent stria terminated anteriorly in presumed location of interval 6; only one discal setiferous pore present, inserted at anterior one-third next to stria 3; preapical seta present, inserted next to stria 2; median lobe of male aedeagus (
<figureCitation id="13442A0EFFD3FFCF3FDB6202FC2775DB" box="[908,1001,627,651]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27b</figureCitation>
) highly distinctive, with ventral margin of shaft sinuous and apex short and thin in lateral view, with broad apical projects in ventral view; endophallus without sclerites, but with a long tubular sleeve covered with long, slender spines.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FFD3FFCF3CFC6284FB4C710C" pageId="70" pageNumber="411" type="discussion">
<paragraph id="8BC0368BFFD3FFCF3CFC6284FF2B74C0" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
<smallCapsWord id="8D26A057FFD3FFCF3CFC6284FEE6745C" baselines="775,775" box="[171,296,757,781]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3CFC6284FEE6745C" bold="true" box="[171,296,757,781]" pageId="70" pageNumber="411">COMMENTS</emphasis>
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.— Among males of all the species of
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<emphasis id="B90BEA99FFD3FFCF3EEB6284FCB9745D" box="[700,887,757,781]" italics="true" pageId="70" pageNumber="411">Trechepaphiopsis</emphasis>
</taxonomicName>
, those of
<taxonomicName id="4C7F4D08FFD3FFCF3F896287FB93745D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" box="[990,1117,758,781]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD3FFCF3F896287FB93745D" box="[990,1117,758,781]" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
have an endophlallus most like that typical for members of the
<taxonomicName id="4C7F4D08FFD3FFCF3EA06367FCB5747E" authorityName="Ueno" authorityYear="1953" box="[759,891,790,814]" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD3FFCF3EA06367FCB5747E" box="[759,891,790,814]" italics="true" pageId="70" pageNumber="411">Epaphiopsis</emphasis>
</taxonomicName>
complex of genera, especially of certain species of
<taxonomicName id="4C7F4D08FFD3FFCF3DF36347FDC7741E" authorityName="Samouelle" authorityYear="1819" box="[420,521,822,846]" class="Insecta" family="Carabidae" genus="Epaphius" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FFD3FFCF3DF36347FDC7741E" box="[420,521,822,846]" italics="true" pageId="70" pageNumber="411">Epaphius</emphasis>
</taxonomicName>
described from
<collectingCountry id="F368761BFFD3FFCF3EE96347FD30741E" box="[702,766,822,846]" name="China" pageId="70" pageNumber="411">China</collectingCountry>
(
<bibRefCitation id="EFEE4B7AFFD3FFCF3F5A6347FC6E741E" author="JEANNEL, R." box="[781,928,822,846]" pageId="70" pageNumber="411" pagination="171 - 207" refId="ref59689" refString="JEANNEL, R. 1962. Les Trechini de l'Extreme-Orient. Revue Francaise d'Entomologie 29: 171 - 207." type="journal article" year="1962">Jeannel 1962</bibRefCitation>
,
<bibRefCitation id="EFEE4B7AFFD3FFCF3FFA6346FB8C741E" author="DEUVE, T." box="[941,1090,822,846]" pageId="70" pageNumber="411" pagination="171 - 184" refId="ref58851" refString="DEUVE, T. 1992 b. Contribution a la connaissance des Trechidae asiatiques (Coleoptera). Bulletin de la Societe Entomologique de France 97: 171 - 184." type="journal article" year="1992">Deuve 1992b</bibRefCitation>
). These also have a long tubular sleeve covered with long spines, symmetrically arranged in ventral view (
<figureCitation id="13442A0EFFD3FFCF3CD46306FF1374C0" box="[131,221,887,912]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27c</figureCitation>
)
</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC63E9FE6473C4" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3CFC63E9FE6A74FE" bold="true" box="[171,420,920,944]" pageId="70" pageNumber="411">
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<smallCapsWord id="8D26A057FFD3FFCF3D4763EAFE6A74FE" baselines="937" box="[272,420,923,942]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="70" pageNumber="411">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in mixed broadleaf evergreen/ deciduous forest at elevations ranging from
<quantity id="4C879B6EFFD3FFCF3ECC63C9FC8A7480" box="[667,836,952,977]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="70" pageNumber="411" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">2460 to 2470 m</quantity>
and collected using both pitfall traps and sifting of leaf litter from the forest floor (
<figureCitation id="13442A0EFFD3FFCF3EFE63A8FCC874A1" box="[681,774,985,1009]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="70" pageNumber="411">Fig. 37a</figureCitation>
). Specimens of
<taxonomicName id="4C7F4D08FFD3FFCF3FE363A8FF0A7341" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechepaphiosis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="uniporosa" status="sp. nov.">
<emphasis id="B90BEA99FFD3FFCF3FE363A8FF0A7341" italics="true" pageId="70" pageNumber="411">Trechepaphiosis uniporosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD3FFCF3C9C638BFED67341" box="[203,280,1018,1041]" pageId="70" pageNumber="411" rank="species">sp. nov</taxonomicNameLabel>
and
<taxonomicName id="4C7F4D08FFD3FFCF3D1A638BFDBC7342" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[333,626,1018,1042]" class="Insecta" family="Carabidae" genus="Trechepaphiama" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="goaligong" status="sp. nov.">
<emphasis id="B90BEA99FFD3FFCF3D1A638BFDBC7342" box="[333,626,1018,1042]" italics="true" pageId="70" pageNumber="411">Trechepaphiama goaligong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FFD3FFCF3E2D638BFD097341" box="[634,711,1018,1041]" pageId="70" pageNumber="411" rank="species">sp. nov</taxonomicNameLabel>
were collected in the same sifted leaf litter samples and are therefore syntopic with
<taxonomicName id="4C7F4D08FFD3FFCF3E7A646AFD627362" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" box="[557,684,1051,1074]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD3FFCF3E7A646AFD627362" box="[557,684,1051,1074]" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
at this site. Additional specimens of
<taxonomicName id="4C7F4D08FFD3FFCF386E646AFF697303" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" baseAuthorityName="Deuve &amp; Kavanaugh &amp; Liang" baseAuthorityYear="2004" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="411" phylum="Arthropoda" rank="species" species="unisetosa">
<emphasis id="B90BEA99FFD3FFCF386E646AFF697303" italics="true" pageId="70" pageNumber="411">T. unisetosa</emphasis>
</taxonomicName>
were collected in pitfall traps set in
<taxonomicName id="4C7F4D08FFD3FFCF3E61644AFD1A7303" box="[566,724,1083,1107]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="70" pageNumber="411" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FFD3FFCF3E61644AFD1A7303" box="[566,724,1083,1107]" italics="true" pageId="70" pageNumber="411">Rhododendron</emphasis>
</taxonomicName>
and bamboo thickets on the east flank of the summit ridge just south of the Fengxue Yakou at
<quantity id="4C879B6EFFD3FFCF3EE9642AFCDE7323" box="[702,784,1115,1139]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="70" pageNumber="411" unit="m" value="3150.0">3150 m</quantity>
(
<figureCitation id="13442A0EFFD3FFCF3F77642AFCB17323" box="[800,895,1115,1139]" captionStart="FIGURE" captionStartId="105.[171,183,1620,1639]" captionTargetBox="[120,1176,216,1606]" captionTargetId="figure-15@105.[120,1176,216,1607]" captionTargetPageId="105" captionText="FIGURE 43. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Luzhang Township, Piana Road 0.1 km E of Fengxue Yakou, elevation 3150 m; habitat in which specimens of Trechus luzhangensis sp. nov. were collected. b. Lushui County, Luzhang Township, 100 m S of Fengxue Yakou on east side of pass, elevation 3150 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve) were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155372" httpUri="https://zenodo.org/record/13155372/files/figure.png" pageId="70" pageNumber="411">Fig. 43b</figureCitation>
), a site slightly above the
<typeStatus id="54C48829FFD3FFCF3C2C640DFF6773C4" box="[123,169,1148,1172]" pageId="70" pageNumber="411">type</typeStatus>
locality for the species.
</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC64EDFF0E72E9" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3CFC64EDFDB173E3" bold="true" box="[171,639,1180,1204]" pageId="70" pageNumber="411">
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<smallCapsWord id="8D26A057FFD3FFCF3D0864D1FE3D73E3" baselines="1198" box="[351,499,1184,1203]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="70" pageNumber="411">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FFD3FFCF3DAA64D1FD8273E3" baselines="1198" box="[509,588,1184,1203]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="70" pageNumber="411">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FFD3FFCF3E0264D1FDB173E3" baselines="1198" box="[597,639,1184,1203]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="70" pageNumber="411">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FFD3FFCF3EDD64EDFCC073E3" baselines="1198,1198" box="[650,782,1180,1204]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3EDD64EDFCC073E3" bold="true" box="[650,782,1180,1204]" pageId="70" pageNumber="411">GAOLIGONG</emphasis>
</smallCapsWord>
<materialsCitation id="3B173CD6FFD3FFCF3F4F64EDFD5C7385" location="We" pageId="70" pageNumber="411" specimenCount="51" specimenCount-female="16" specimenCount-male="10" stateProvince="Shan">
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<collectingRegion id="49BBF869FFD3FFCF3F4F64EDFC9D73E3" box="[792,851,1180,1204]" country="Myanmar" name="Shan" pageId="70" pageNumber="411">SHAN</collectingRegion>
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.—
<figureCitation id="13442A0EFFD3FFCF3F2864EDFC1073E4" box="[895,990,1180,1204]" captionStart="FIGURE" captionStartId="71.[171,183,1575,1594]" captionTargetBox="[120,1176,216,1557]" captionTargetId="figure-14@71.[120,1176,216,1557]" captionTargetPageId="71" captionText="FIGURE 27. Trechepaphiopsis unisetosa (Deuve); a. Dorsal habitus (CASENT1024491). b-c. Median lobe of aedeagus of male (CASENT1001903); b. left lateral aspect; c. ventral aspect. d. Map of locality records (red circles) for T. unisetosa in the Gaoligong Shan region. Scale lines a-c = 0.5 mm, d = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155334" httpUri="https://zenodo.org/record/13155334/files/figure.png" pageId="70" pageNumber="411">Fig. 27d</figureCitation>
.
<location id="8EA06050FFD3FFCF3FBC64ECFBC073E4" LSID="urn:lsid:plazi:treatment:03D6879DFFD3FFCF3C2C60AAFB4C710C:8EA06050FFD3FFCF3FBC64ECFBC073E4" box="[1003,1038,1181,1204]" name="We" pageId="70" pageNumber="411" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FFD3FFCF3C8464CCFEA87385" box="[211,358,1213,1237]" count="25" pageId="70" pageNumber="411" type="generic">25 specimens</specimenCount>
(
<specimenCount id="9D79FD02FFD3FFCF3D2064CCFE147385" box="[375,474,1213,1237]" count="10" pageId="70" pageNumber="411" type="male">10 males</specimenCount>
and
<specimenCount id="9D79FD02FFD3FFCF3E4464CCFD427385" box="[531,652,1213,1237]" count="16" pageId="70" pageNumber="411" type="female">16 females</specimenCount>
)
</materialsCitation>
<materialsCitation id="3B173CD6FFD3FFCF3ECB64CCFF7272E9" collectingDate="1998-10-15" collectingDateMax="2005-05-21" collectingDateMin="1998-10-15" collectionCode="CAS, IOZ, MNHN" collectorName="J.P. DUMBACHER &amp; C. E. Griswold &amp; K. J. Guo &amp; Lushui &amp; Changyan He &amp; C. Ferraris &amp; C. L. Long" county="Lushui County" elevation="100" latitude="25.99363" location="Fengxue Yakou" longLatPrecision="1" longitude="97.66651" municipality="Luzhang Township" pageId="70" pageNumber="411" specimenCount="25" specimenCount-female="15" specimenCount-male="10" stateProvince="SHAN">
from the following localities:
<emphasis id="B90BEA99FFD3FFCF3FB264CCFB5B7385" bold="true" box="[997,1173,1213,1237]" italics="true" pageId="70" pageNumber="411">
<collectingCounty id="62A14E07FFD3FFCF3FB264CCFB407385" box="[997,1166,1213,1237]" pageId="70" pageNumber="411">Lushui County</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA4ACF1FFD3FFCF3C2C64AFFE9373A6" box="[123,349,1246,1270]" pageId="70" pageNumber="411">Luzhang Township</collectingMunicipality>
(
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<elevation id="0052D1B8FFD3FFCF3D2D64AFFE0073A5" box="[378,462,1246,1270]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="411" unit="m" value="100.0">100 m</elevation>
</quantity>
S of
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on east side of pass,
<quantity id="4C879B6EFFD3FFCF387A64ACFB5F73A5" box="[1069,1169,1245,1269]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="70" pageNumber="411" unit="m" value="3150.0">
<elevation id="0052D1B8FFD3FFCF387A64ACFB5F73A5" box="[1069,1169,1245,1269]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="70" pageNumber="411" unit="m" value="3150.0">3150 m</elevation>
</quantity>
,
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/
<geoCoordinate id="EE4B504CFFD3FFCF3D51648FFE4B7246" box="[262,389,1278,1302]" degrees="97.68381" direction="east" orientation="longitude" pageId="70" pageNumber="411" precision="1" value="97.68381">E97.68381°</geoCoordinate>
,
<date id="FFC1104BFFD3FFCF3DC7648FFD8D7246" box="[400,579,1278,1303]" pageId="70" pageNumber="411" value="2005-05-11" valueMax="2005-05-21" valueMin="2005-05-11">
<collectingDate id="EF85E9A3FFD3FFCF3DC7648FFD8D7246" box="[400,579,1278,1303]" pageId="70" pageNumber="411" value="2005-05-11" valueMax="2005-05-21" valueMin="2005-05-11">11-21 May 2005</collectingDate>
</date>
,
<collectorName id="268A535DFFD3FFCF3E1A648EFD4F7246" box="[589,641,1279,1302]" name="J.P. DUMBACHER" pageId="70" pageNumber="411">D.H.</collectorName>
<taxonomicName id="4C7F4D08FFD3FFCF3ED0648FFCCD7246" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[647,771,1278,1302]" pageId="70" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
,
<collectorName id="268A535DFFD3FFCF3F59648FFC687246" box="[782,934,1278,1302]" pageId="70" pageNumber="411">C.E. Griswold</collectorName>
&amp;
<collectorName id="268A535DFFD3FFCF3F9F648EFBE97246" box="[968,1063,1278,1302]" pageId="70" pageNumber="411">K.J. Guo</collectorName>
collectors [
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and
<specimenCount id="9D79FD02FFD3FFCF3CAA656EFE927267" box="[253,348,1311,1335]" count="1" pageId="70" pageNumber="411" type="female">1 female</specimenCount>
;
<collectionCode id="ED6EAE4EFFD3FFCF3D31656EFE527267" box="[358,412,1311,1335]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="70" pageNumber="411" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFFD3FFCF3DF0656EFE147267" box="[423,474,1311,1335]" pageId="70" pageNumber="411">IOZ</collectionCode>
]);
<location id="8EA06050FFD3FFCF3DA5656EFD7F7267" LSID="urn:lsid:plazi:treatment:03D6879DFFD3FFCF3C2C60AAFB4C710C:8EA06050FFD3FFCF3DA5656EFD7F7267" box="[498,689,1311,1335]" county="Lushui County" latitude="25.99363" longLatPrecision="1" longitude="97.66651" municipality="Luzhang Township" name="Pianma Township" pageId="70" pageNumber="411" stateProvince="SHAN">Pianma Township</location>
(
<locationDeviation id="9A722FA9FFD3FFCF3EE9656EFC7A7267" box="[702,948,1311,1335]" location="Pianma" pageId="70" pageNumber="411">
<quantity id="4C879B6EFFD3FFCF3EE9656EFCC67267" box="[702,776,1311,1335]" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="70" pageNumber="411" unit="km" value="9.3">9.3 km</quantity>
ENE of Pianma
</locationDeviation>
along road to
<collectorName id="268A535DFFD3FFCF381C656EFB5B7267" box="[1099,1173,1311,1335]" pageId="70" pageNumber="411">Lushui</collectorName>
at
<collectorName id="268A535DFFD3FFCF3CC1654EFEE37207" box="[150,301,1343,1367]" pageId="70" pageNumber="411">Changyan He</collectorName>
,
<quantity id="4C879B6EFFD3FFCF3D6E654EFE027207" box="[313,460,1343,1367]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="70" pageNumber="411" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">
<elevation id="0052D1B8FFD3FFCF3D6E654EFE027207" box="[313,460,1343,1367]" metricMagnitude="3" metricUnit="m" metricValue="2.465" metricValueMax="2.4699999999999998" metricValueMin="2.46" pageId="70" pageNumber="411" unit="m" value="2465.0" valueMax="2470.0" valueMin="2460.0">2460-2470 m</elevation>
</quantity>
,
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<collectorName id="268A535DFFD3FFCF3C2C6511FED97228" box="[123,279,1376,1400]" pageId="70" pageNumber="411">C.E. Griswold</collectorName>
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<collectorName id="268A535DFFD3FFCF3D746511FE587228" box="[291,406,1376,1400]" pageId="70" pageNumber="411">C. Ferraris</collectorName>
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<collectorName id="268A535DFFD3FFCF3DEE6511FDE67228" box="[441,552,1376,1400]" pageId="70" pageNumber="411">C.L. Long</collectorName>
collectors [
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<collectionCode id="ED6EAE4EFFD3FFCF385E6511FBAE7227" box="[1033,1120,1376,1399]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="70" pageNumber="411">MNHN</collectionCode>
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<collectingDate id="EF85E9A3FFD3FFCF38256511FEC572C8" pageId="70" pageNumber="411" value="2005-05-12" valueMax="2005-05-21" valueMin="2005-05-12">12- 21 May 2005</collectingDate>
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<collectorName id="268A535DFFD3FFCF3D8265F0FDA072C8" box="[469,622,1408,1433]" pageId="70" pageNumber="411">C.E. Griswold</collectorName>
&amp;
<collectorName id="268A535DFFD3FFCF3EC765F0FD2172C9" box="[656,751,1409,1433]" pageId="70" pageNumber="411">K.J. Guo</collectorName>
collectors [
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and
<specimenCount id="9D79FD02FFD3FFCF3FB165F0FB9E72C8" box="[998,1104,1408,1433]" count="2" pageId="70" pageNumber="411" type="female">2 females</specimenCount>
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<collectionCode id="ED6EAE4EFFD3FFCF3C2C65D0FF6072E9" box="[123,174,1441,1465]" pageId="70" pageNumber="411">IOZ</collectionCode>
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</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC65B0FF73714B" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">Members of this species have been collected only in the southcentral part of the study area, on both western and eastern slopes of the Gaoligongshan, which are in Core Areas 4 and 5, respectively.</paragraph>
<paragraph id="8BC0368BFFD3FFCF3CFC6652FB4C710C" blockId="70.[123,1174,399,1628]" pageId="70" pageNumber="411">
<emphasis id="B90BEA99FFD3FFCF3CFC6652FD97716A" bold="true" box="[171,601,1571,1595]" pageId="70" pageNumber="411">
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<smallCapsWord id="8D26A057FFD3FFCF3D4E6656FE0E716A" baselines="1589" box="[281,448,1575,1594]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="70" pageNumber="411">GEOGRAPHICAL</smallCapsWord>
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.— This species currently is known only from Lushui County in the southcentral part of the Gaoligong Shan region, western
<collectingRegion id="49BBF869FFD3FFCF3F226635FBFE710C" box="[885,1072,1604,1628]" country="China" name="Yunnan" pageId="70" pageNumber="411">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFFD3FFCF386C6635FBB0710C" box="[1083,1150,1604,1628]" name="China" pageId="70" pageNumber="411">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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