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<document id="F827C74B9632072C8AEFF49CD4A5728A" ID-DOI="10.11646/zootaxa.5492.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13219142" ID-ZooBank="91164DD5-609C-4415-A659-87A28A010F6D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722840936672" checkinUser="plazi" docAuthor="Ermilov, Sergey G., Rybalov, Leonid B. &amp; Kuzmicheva, Evgeniya A." docDate="2024" docId="03834F1FFFD8FFB8FF3FF947D8250C85" docLanguage="en" docName="zootaxa.5492.1.2.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scheloribates (Topobates) tredecim Ermilov, Rybalov &amp; Kuzmicheva, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="189" masterDocId="FFBA3767FFD2FFB5FFA8FFF1DD1C0944" masterDocTitle="New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia" masterLastPageNumber="190" masterPageNumber="176" pageNumber="186" updateTime="1722841352648" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BCB52BA86354F4BAF49495A3E17804EE">New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia</mods:title>
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<mods:namePart id="D2ED53C43B54BFC4A68E74A35807BB08">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="6CD5C55A8BCD44DA6B3117B40D78FE4F">Tyumen State University, Institute of Environmental and Agricultural Biology (X-BIO), Tyumen, Russia</mods:affiliation>
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<mods:namePart id="7DFA217BE18B584F3D2C5142EFFC3C8B">Rybalov, Leonid B.</mods:namePart>
<mods:affiliation id="DB743069966791927CF5B749AF4C0406">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Soil Zoology and General Entomology, Moscow, Russia</mods:affiliation>
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<mods:namePart id="D4C0E8252296FAC3047DCE3904290AAB">Kuzmicheva, Evgeniya A.</mods:namePart>
<mods:affiliation id="3050DA136E074C961314B40F5D91C1BE">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Historical Ecology, Moscow, Russia</mods:affiliation>
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<paragraph id="8B95FE09FFD8FFBFFF3FF947DFBF0F95" blockId="10.[151,675,1718,1781]" box="[151,675,1718,1745]" pageId="10" pageNumber="186">
<heading id="D0DD4965FFD8FFBFFF3FF947DFBF0F95" bold="true" box="[151,675,1718,1745]" fontSize="11" level="1" pageId="10" pageNumber="186" reason="1">
<emphasis id="B95E221BFFD8FFBFFF3FF947DFBF0F95" bold="true" box="[151,675,1718,1745]" pageId="10" pageNumber="186">
<taxonomicName id="4C2A858AFFD8FFBFFF3FF947DF210F95" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[151,573,1718,1745]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="186" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFD8FFBFFF3FF947DC260F95" bold="true" box="[151,314,1718,1745]" italics="true" pageId="10" pageNumber="186">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD8FFBFFEE3F947DCD80F95" bold="true" box="[331,452,1718,1745]" italics="true" pageId="10" pageNumber="186">Topobates</emphasis>
)
<emphasis id="B95E221BFFD8FFBFFE7DF947DF210F95" bold="true" box="[469,573,1718,1745]" italics="true" pageId="10" pageNumber="186">tredecim</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD8FFBFFDEDF946DFBF0F95" box="[581,675,1719,1745]" pageId="10" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C330AD82FFD8FFBFFF3FF92BDC170FB1" box="[151,267,1754,1781]" pageId="10" pageNumber="186" type="description">
<paragraph id="8B95FE09FFD8FFBFFF3FF92BDC170FB1" blockId="10.[151,675,1718,1781]" box="[151,267,1754,1781]" pageId="10" pageNumber="186">
(
<figureCitation id="1311E28CFFD8FFBFFF37F92BDDF40FB1" box="[159,232,1754,1781]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="10" pageNumber="186">Figs 7</figureCitation>
,
<figureCitation id="1311E28CFFD8FFBFFF5CF92ADC1E0FB1" box="[244,258,1755,1781]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="10" pageNumber="186">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD8FFBEFF3FF8D3DEE00EA5" lastPageId="11" lastPageNumber="187" pageId="10" pageNumber="186" type="materials_examined">
<paragraph id="8B95FE09FFD8FFBFFF3FF8D3DC2A0E88" blockId="10.[151,1437,1826,1996]" pageId="10" pageNumber="186">
<emphasis id="B95E221BFFD8FFBFFF3FF8D3DC1A0E78" bold="true" box="[151,262,1826,1852]" pageId="10" pageNumber="186">Material.</emphasis>
<materialsCitation id="3B42F454FFD8FFBFFEB8F8D3DC2A0E88" collectingDate="2014-11-15" collectorName="African &amp; E. A. Kuzmicheva" country="Ethiopia" county="Southeastern" elevation="2570" latitude="6.7216945" location="Bale Mountains National Park" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" pageId="10" pageNumber="186" specimenCount="14" specimenCount-female="2" specimenCount-male="5" stateProvince="Oromiya" typeStatus="holotype">
<typeStatus id="549140ABFFD8FFBFFEB8F8D3DC640E78" box="[272,376,1826,1852]" pageId="10" pageNumber="186" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D2C3580FFD8FFBFFDB5F8D2DFCD0E79" box="[541,721,1827,1853]" count="7" pageId="10" pageNumber="186" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
(
<specimenCount id="9D2C3580FFD8FFBFFD4BF8D3DE470E78" box="[739,859,1826,1852]" count="5" pageId="10" pageNumber="186" type="male">five males</specimenCount>
and
<specimenCount id="9D2C3580FFD8FFBFFC3FF8D2D9340E78" box="[919,1064,1826,1853]" count="2" pageId="10" pageNumber="186" type="female">two females</specimenCount>
):
<collectingCounty id="62F48685FFD8FFBFFB95F8D3D9D20E78" box="[1085,1230,1826,1852]" pageId="10" pageNumber="186">Southeastern</collectingCounty>
<collectingCountry id="F33DBE99FFD8FFBFFB7FF8D3D8250E78" box="[1239,1337,1826,1852]" name="Ethiopia" pageId="10" pageNumber="186">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD8FFBFFAEFF8D3DDF70E24" country="Ethiopia" name="Oromiya" pageId="10" pageNumber="186">Oromia Region</collectingRegion>
,
<collectingMunicipality id="6BF16473FFD8FFBFFF5DF8B7DC770E25" box="[245,363,1862,1889]" pageId="10" pageNumber="186">Bale Zone</collectingMunicipality>
,
<location id="8EF5A8D2FFD8FFBFFEDEF8B7DFD60E24" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD8FFBFFEDEF8B7DFD60E24" box="[374,714,1862,1888]" country="Ethiopia" county="Southeastern" latitude="6.7216945" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" name="Bale Mountains National Park" pageId="10" pageNumber="186" stateProvince="Oromiya">Bale Mountains National Park</location>
,
<geoCoordinate id="EE1E98CEFFD8FFBFFD7DF8B9DE780E27" box="[725,868,1864,1891]" degrees="6" direction="north" minutes="43" orientation="latitude" pageId="10" pageNumber="186" precision="1" seconds="18.1" value="6.7216945">6°4318.1″N</geoCoordinate>
,
<geoCoordinate id="EE1E98CEFFD8FFBFFCC7F8B9D9140E27" box="[879,1032,1864,1891]" degrees="39" direction="east" minutes="44" orientation="longitude" pageId="10" pageNumber="186" precision="1" seconds="12.1" value="39.736694">39°4412.1″E</geoCoordinate>
,
<elevation id="0007193AFFD8FFBFFBBBF8B7D9BF0E24" box="[1043,1187,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="10" pageNumber="186" unit="m" value="2570.0">
<quantity id="4CD253ECFFD8FFBFFBBBF8B7D97B0E25" box="[1043,1127,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="10" pageNumber="186" unit="m" value="2570.0">2570 m</quantity>
a.s.l.
</elevation>
,
<location id="8EF5A8D2FFD8FFBFFB07F8B6D8410E25" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD8FFBFFB07F8B6D8410E25" box="[1199,1373,1863,1889]" country="Ethiopia" county="Southeastern" latitude="6.7216945" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" name="Harenna Forest" pageId="10" pageNumber="186" stateProvince="Oromiya">Harenna Forest</location>
, litter in bamboo belt (vegetation represented by dense bamboo thickets of
<taxonomicName id="4C2A858AFFD8FFBFFC0BF89BD9470EC0" box="[931,1115,1898,1924]" class="Liliopsida" family="Poaceae" genus="Oldeania" kingdom="Plantae" order="Poales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="B95E221BFFD8FFBFFC0BF89BD9470EC0" box="[931,1115,1898,1924]" italics="true" pageId="10" pageNumber="186">Oldeania alpina</emphasis>
</taxonomicName>
, the
<collectorName id="26DF9BDFFFD8FFBFFB39F89BD9FB0EC0" box="[1169,1255,1898,1924]" pageId="10" pageNumber="186">African</collectorName>
alpine bamboo,
<emphasis id="B95E221BFFD8FFBFFF3FF87EDF8F0EEC" box="[151,659,1934,1960]" italics="true" pageId="10" pageNumber="186">
<taxonomicName id="4C2A858AFFD8FFBFFF3FF87EDCC10EEC" authority=", Myrsine" authorityName="Myrsine" box="[151,477,1934,1960]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="abyssinica">Hagenia abyssinica, Myrsine</taxonomicName>
melanophloeos,
</emphasis>
and
<taxonomicName id="4C2A858AFFD8FFBFFD60F87FDEBF0EEC" box="[712,931,1934,1960]" class="Magnoliopsida" family="Araliaceae" genus="Schefflera" kingdom="Plantae" order="Apiales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="volkensii">
<emphasis id="B95E221BFFD8FFBFFD60F87FDEBF0EEC" box="[712,931,1934,1960]" italics="true" pageId="10" pageNumber="186">Schefflera volkensii</emphasis>
</taxonomicName>
trees accompany bamboo),
<date id="FF94D8C9FFD8FFBFFB72F87FD8460EED" box="[1242,1370,1934,1961]" pageId="10" pageNumber="186" value="2014-11-15">
<collectingDate id="EFD02121FFD8FFBFFB72F87FD8460EED" box="[1242,1370,1934,1961]" pageId="10" pageNumber="186" value="2014-11-15">15.XI.2014</collectingDate>
</date>
(
<collectorName id="26DF9BDFFFD8FFBFFAC1F87EDC360E88" pageId="10" pageNumber="186">E.A. Kuzmicheva</collectorName>
).
</materialsCitation>
</paragraph>
<caption id="DF55AE81FFD9FFBEFF3FF908DE370E72" ID-DOI="http://doi.org/10.5281/zenodo.13219162" ID-Zenodo-Dep="13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="11" pageNumber="187" startId="11.[151,250,1785,1809]" targetBox="[179,1408,196,1746]" targetPageId="11" targetType="figure">
<paragraph id="8B95FE09FFD9FFBEFF3FF908DE370E72" blockId="11.[151,1436,1785,1846]" pageId="11" pageNumber="187">
<emphasis id="B95E221BFFD9FFBEFF3FF908DC0F0E55" bold="true" box="[151,275,1785,1809]" pageId="11" pageNumber="187">FIGURE 7.</emphasis>
<taxonomicName id="4C2A858AFFD9FFBEFEB1F908DFC50E55" authority="Ermilov" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[281,729,1785,1809]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="187" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFD9FFBEFEB1F908DCBF0E55" box="[281,419,1785,1809]" italics="true" pageId="11" pageNumber="187">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD9FFBEFE1AF908DF050E55" box="[434,537,1785,1809]" italics="true" pageId="11" pageNumber="187">Topobates</emphasis>
)
<emphasis id="B95E221BFFD9FFBEFD80F908DF9C0E55" box="[552,640,1785,1809]" italics="true" pageId="11" pageNumber="187">tredecim</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD9FFBEFD77F908DE330E55" bold="true" box="[735,815,1785,1809]" pageId="11" pageNumber="187">
<taxonomicNameLabel id="A26D9F60FFD9FFBEFD77F908DE330E55" box="[735,815,1785,1809]" pageId="11" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<paragraph id="8B95FE09FFD9FFBEFF6FF88EDEE00EA5" blockId="11.[151,1436,1919,2018]" pageId="11" pageNumber="187">
The
<typeStatus id="549140ABFFD9FFBEFF52F88EDC400EDD" box="[250,348,1919,1945]" pageId="11" pageNumber="187" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
<collectingCountry id="F33DBE99FFD9FFBEFA84F871D8840EDE" box="[1324,1432,1920,1946]" name="Germany" pageId="11" pageNumber="187">Germany</collectingCountry>
;
<materialsCitation id="3B42F454FFD9FFBEFF3FF855D8710EF9" box="[151,1389,1955,1982]" country="Russia" location="University Museum of Zoology" pageId="11" pageNumber="187" specimenCount="7" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D2C3580FFD9FFBEFF3FF855DC500EFA" box="[151,332,1956,1982]" count="7" pageId="11" pageNumber="187" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
are in the collection of the
<collectingRegion id="49EE30EBFFD9FFBEFD3AF855DE330EFA" box="[658,815,1956,1982]" country="Russia" name="Tyumen'" pageId="11" pageNumber="187">Tyumen State</collectingRegion>
<location id="8EF5A8D2FFD9FFBEFC90F852D9B90EF9" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD9FFBEFC90F852D9B90EF9" box="[824,1189,1955,1982]" country="Russia" name="University Museum of Zoology" pageId="11" pageNumber="187" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="49EE30EBFFD9FFBEFB1AF855D80C0EFA" box="[1202,1296,1956,1982]" country="Russia" name="Tyumen'" pageId="11" pageNumber="187">Tyumen</collectingRegion>
,
<collectingCountry id="F33DBE99FFD9FFBEFAB7F852D8710EF9" box="[1311,1389,1955,1981]" name="Russia" pageId="11" pageNumber="187">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in a solution of 70% ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<caption id="DF55AE81FFDEFFB9FF3FF96AD9650FBF" ID-DOI="http://doi.org/10.5281/zenodo.13219166" ID-Zenodo-Dep="13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="12" pageNumber="188" startId="12.[151,250,1691,1715]" targetBox="[176,1420,190,1654]" targetPageId="12" targetType="figure">
<paragraph id="8B95FE09FFDEFFB9FF3FF96AD9650FBF" blockId="12.[151,1436,1691,1788]" pageId="12" pageNumber="188">
<emphasis id="B95E221BFFDEFFB9FF3FF96ADC0B0FF7" bold="true" box="[151,279,1691,1715]" pageId="12" pageNumber="188">FIGURE 8.</emphasis>
<taxonomicName id="4C2A858AFFDEFFB9FE8BF96ADFEC0FF7" authority="Ermilov" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[291,752,1691,1715]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="188" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFDEFFB9FE8BF96ADCB10FF7" box="[291,429,1691,1715]" italics="true" pageId="12" pageNumber="188">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFDEFFB9FE68F96ADF3B0FF7" box="[448,551,1691,1715]" italics="true" pageId="12" pageNumber="188">Topobates</emphasis>
)
<emphasis id="B95E221BFFDEFFB9FD92F96ADF8E0FF7" box="[570,658,1691,1715]" italics="true" pageId="12" pageNumber="188">tredecim</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFDEFFB9FD53F96ADE4C0FF7" bold="true" box="[763,848,1691,1715]" pageId="12" pageNumber="188">
<taxonomicNameLabel id="A26D9F60FFDEFFB9FD53F96ADE4C0FF7" box="[763,848,1691,1715]" pageId="12" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG).
</paragraph>
</caption>
<subSubSection id="C330AD82FFDEFFB9FF6FF8B4D8690EAB" pageId="12" pageNumber="188" type="diagnosis">
<paragraph id="8B95FE09FFDEFFB9FF6FF8B4D8690EAB" blockId="12.[151,1436,1861,2032]" pageId="12" pageNumber="188">
<emphasis id="B95E221BFFDEFFB9FF6FF8B4DC5E0E1B" bold="true" box="[199,322,1861,1887]" pageId="12" pageNumber="188">Diagnosis.</emphasis>
Body length: 390450.
<taxonomicName id="4C2A858AFFDEFFB9FDE3F8B7DFDA0E24" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[587,710,1862,1888]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Rostrum protruding, rounded. Prolamella and translamella absent. Rostral, lamellar and interlamellar setae long, setiform, barbed;
<emphasis id="B95E221BFFDEFFB9FB99F89BD95B0EC7" box="[1073,1095,1898,1923]" italics="true" pageId="12" pageNumber="188">in</emphasis>
˃(=)
<emphasis id="B95E221BFFDEFFB9FBD1F898D9910EC7" box="[1145,1165,1897,1923]" italics="true" pageId="12" pageNumber="188">le</emphasis>
˃
<emphasis id="B95E221BFFDEFFB9FB35F89BD9A90EC7" box="[1181,1205,1898,1923]" italics="true" pageId="12" pageNumber="188">ro</emphasis>
; bothridial seta long, lanceolate, barbed. Thirteen pairs of notogastral setae short, needleform. Four pairs of sacculi. Circumpedal carina long. Anal and adanal setae medium-sized, setiform, thin, roughened. Ventrobasal tubercle of leg tibiae I, II well observable; femur II broadly rounded ventrodistally; tarsus I with 19 setae (
<emphasis id="B95E221BFFDEFFB9FC4FF824DEE20EAB" box="[999,1022,2005,2031]" italics="true" pageId="12" pageNumber="188">l”</emphasis>
absent;
<emphasis id="B95E221BFFDEFFB9FBF4F827D96E0EB4" box="[1116,1138,2006,2032]" italics="true" pageId="12" pageNumber="188">v</emphasis>
present). Tridactylous.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFF67D8190C49" pageId="13" pageNumber="189" type="description">
<paragraph id="8B95FE09FFDFFFB8FF6FFF67D9A209BD" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFF67DCAE09F5" bold="true" box="[199,434,150,177]" pageId="13" pageNumber="189">Description of adult</emphasis>
.
<emphasis id="B95E221BFFDFFFB8FE16FF66DF7C09F4" box="[446,608,151,176]" italics="true" pageId="13" pageNumber="189">Measurements</emphasis>
. Body length: 420 (
<typeStatus id="549140ABFFDFFFB8FC91FF66DEBD09F5" box="[825,929,151,177]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 390420 (male
<typeStatus id="549140ABFFDFFFB8FBF6FF66D9D209F5" box="[1118,1230,151,177]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 420450 (female
<typeStatus id="549140ABFFDFFFB8FF3FFF4ADC140991" box="[151,264,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
); body width (level of pteromorph): 240 (
<typeStatus id="549140ABFFDFFFB8FD48FF4ADE540991" box="[736,840,187,213]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 225240 (male
<typeStatus id="549140ABFFDFFFB8FBA5FF4AD9610991" box="[1037,1149,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 255 (female
<typeStatus id="549140ABFFDFFFB8FA88FF4AD88D0991" box="[1312,1425,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
); width of ventral plate: 225 (
<typeStatus id="549140ABFFDFFFB8FE67FF2EDF2B09BD" box="[463,567,223,249]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 195225 (male
<typeStatus id="549140ABFFDFFFB8FD52FF2EDE7609BD" box="[762,874,223,249]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 225240 (female
<typeStatus id="549140ABFFDFFFB8FBEAFF2ED9AF09BD" box="[1090,1203,223,249]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFEF2DC2B0821" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFEF2DC590858" box="[199,325,259,284]" italics="true" pageId="13" pageNumber="189">Integument</emphasis>
. Body color light brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens; ×1000); podosomal region and lateral part of prodorsum partially with dense microgranulate cerotegument.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFE9ED9ED0B5D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFE9EDC5808CD" box="[199,324,367,393]" italics="true" pageId="13" pageNumber="189">Prodorsum</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FEFAFE9EDCD308CD" box="[338,463,367,393]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7A, C</figureCitation>
). Rostrum protruding, rounded. Lamella about 1/2 length of prodorsum; prolamella and translamella absent; sublamella and lateral keel-shaped ridge distinct; sublamellar porose area (7) rounded. Rostral (7175), lamellar (6771) and interlamellar (7579) setae setiform, barbed; bothridial seta (5664) with lanceolate, barbed head, directed posterolaterad; bothridial stalk and head similar in length or stalk slightly longer than head; exobothridial seta (3034) setiform, thin, roughened. Dorsosejugal porose area oval, poorly visible.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFDD2D9750BED" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<taxonomicName id="4C2A858AFFDFFFB8FF6FFDD2DC5F0B78" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,547,572]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="B95E221BFFDFFFB8FF6FFDD2DC5F0B78" box="[199,323,547,572]" italics="true" pageId="13" pageNumber="189">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDFFFB8FEF3FDD2DCF70B79" box="[347,491,547,573]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7A, C</figureCitation>
). Anterior notogastral margin convex medially.
<taxonomicName id="4C2A858AFFDFFFB8FB9DFDD2D9AC0B79" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[1077,1200,547,573]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Pteromorph medium-sized, broadly rounded laterally, its anterior margin straight (perpendicular to longitudinal axis of body in dorsal aspect). Thirteen pairs of notogastral setae (6) needleform. Four pairs of saccules with small opening and drop-like channel. Opisthonotal gland opening and all lyrifissures distinct.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFD42DDF70A7D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<taxonomicName id="4C2A858AFFDFFFB8FF6FFD42DC4A0B89" authorityName="Krefft" authorityYear="1871" box="[199,342,691,717]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="13" pageNumber="189" phylum="Nematoda" rank="genus">
<emphasis id="B95E221BFFDFFFB8FF6FFD42DC4A0B89" box="[199,342,691,717]" italics="true" pageId="13" pageNumber="189">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDFFFB8FECFFD42DCF60B89" box="[359,490,691,717]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="13" pageNumber="189">Figs 8AC</figureCitation>
). Subcapitulum size: 105112 × 7582; subcapitular setae (
<emphasis id="B95E221BFFDFFFB8FB3CFD42D9BE0B88" box="[1172,1186,691,716]" italics="true" pageId="13" pageNumber="189">a</emphasis>
: 1922;
<emphasis id="B95E221BFFDFFFB8FAA2FD42D8020B88" box="[1290,1310,691,716]" italics="true" pageId="13" pageNumber="189">m</emphasis>
: 1519;
<emphasis id="B95E221BFFDFFFB8FA2EFD42D8880B89" box="[1414,1428,691,717]" italics="true" pageId="13" pageNumber="189">h</emphasis>
: 2630) setiform, roughened;
<emphasis id="B95E221BFFDFFFB8FE73FD26DCF30BB4" box="[475,495,727,752]" italics="true" pageId="13" pageNumber="189">m</emphasis>
thinner than
<emphasis id="B95E221BFFDFFFB8FD29FD26DF930BB4" box="[641,655,727,752]" italics="true" pageId="13" pageNumber="189">a</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FD6BFD26DFCD0BB5" box="[707,721,727,753]" italics="true" pageId="13" pageNumber="189">h</emphasis>
; both adoral setae (1315) setiform, barbed. Palp length: 6064; postpalpal seta (7) spiniform, roughened. Chelicera length: 112116; setae (
<emphasis id="B95E221BFFDFFFB8FBA6FD0AD92A0A51" box="[1038,1078,763,789]" italics="true" pageId="13" pageNumber="189">cha</emphasis>
: 3437;
<emphasis id="B95E221BFFDFFFB8FB0AFD0AD9D60A51" box="[1186,1226,763,789]" italics="true" pageId="13" pageNumber="189">chb</emphasis>
: 2224) setiform, barbed.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFCB2DCE40A8D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFCB2DF960A18" box="[199,650,835,861]" italics="true" pageId="13" pageNumber="189">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FD30FCB2DE080A19" box="[664,788,835,861]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7B, C</figureCitation>
). Epimeral formula: 3133; all setae (
<emphasis id="B95E221BFFDFFFB8FB7BFCB2D9EC0A19" box="[1235,1264,835,861]" italics="true" pageId="13" pageNumber="189">1a</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FB55FCB2D8040A19" box="[1277,1304,835,861]" italics="true" pageId="13" pageNumber="189">2a</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FA8EFCB2D85D0A19" box="[1318,1345,835,861]" italics="true" pageId="13" pageNumber="189">3a</emphasis>
: 1113;
<emphasis id="B95E221BFFDFFFB8FF3FFC96DDA80AC5" box="[151,180,871,897]" italics="true" pageId="13" pageNumber="189">1b</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FF6DFC96DDFD0AC5" box="[197,225,871,897]" italics="true" pageId="13" pageNumber="189">3b</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FF5BFC96DC110AC5" box="[243,269,871,897]" italics="true" pageId="13" pageNumber="189">3c</emphasis>
: 2628; others: 1922) setiform, thin, roughened. Humeral porose areas
<emphasis id="B95E221BFFDFFFB8FBC0FC96D9910AC5" box="[1128,1165,871,897]" italics="true" pageId="13" pageNumber="189">Am</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FB64FC96D9F70AC5" box="[1228,1259,871,897]" italics="true" pageId="13" pageNumber="189">Ah</emphasis>
poorly visible, elongate oval. Pedotectum II rounded laterally in ventral aspect. Discidium broadly rounded. Circumpedal carina long, directed to pedotectum II.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFC22DC150D71" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFC22DC8F0AA8" box="[199,403,979,1005]" italics="true" pageId="13" pageNumber="189">Anogenital region</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FE0CFC22DF390AA9" box="[420,549,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7B, C</figureCitation>
). Genital (
<emphasis id="B95E221BFFDFFFB8FD37FC22DFB10AA8" box="[671,685,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FD05FC14DFA90AB7" attach="left" box="[685,693,997,1011]" fontSize="6" pageId="13" pageNumber="189">1</subScript>
: 1922;
<emphasis id="B95E221BFFDFFFB8FCB5FC22DE370AA8" box="[797,811,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FC83FC14DE2F0AB7" attach="left" box="[811,819,997,1011]" fontSize="6" pageId="13" pageNumber="189">2</subScript>
<emphasis id="B95E221BFFDFFFB8FCE9FC22DE530AA8" box="[833,847,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FCE7FC14DE4B0AB7" attach="left" box="[847,855,997,1011]" fontSize="6" pageId="13" pageNumber="189">4</subScript>
: 1113), aggenital (22), anal (22), and adanal (22) setae setiform, thin, roughened. Adanal lyrifissure distinct, close and parallel to anal plate. Marginal porose area band-like.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFBCED8190C49" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFBCEDDE00D1C" box="[199,252,1087,1112]" italics="true" pageId="13" pageNumber="189">Legs</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FEA4FBCEDC950D1D" box="[268,393,1087,1113]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="13" pageNumber="189">Figs 8DG</figureCitation>
). Median claw thick; lateral claws thin, with small tubercle distoventrally; all claws slightly barbed on dorsal side. Ventrobasal tubercle of tibiae I, II well observable. Femur II broadly rounded ventrodistally. Proximoventral porose area on tarsi IIV and distoventral porose area on tibiae IIV present but poorly visible; dorsoparaxial porose area on femora IIV and on trochanters III, IV distinct. Formulas of leg setation and solenidia: I (153419) [122], II (152415) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="C6A8CBB2FFDFFFB8FDB2FB02DF720C49" box="[538,622,1267,1293]" captionStart="TABLE 1" captionStartId="4.[151,239,1592,1616]" captionText="TABLE 1. Leg setation and solenidia of adult Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., S. (S.) monosetosus Ermilov sp. nov., S. (S.) triangulus Ermilov sp. nov., and S. (Topobates) tredecim Ermilov sp. nov." pageId="13" pageNumber="189">Table 1</tableCitation>
. Seta
<emphasis id="B95E221BFFDFFFB8FD1AFB02DFA10C48" box="[690,701,1267,1292]" italics="true" pageId="13" pageNumber="189">s</emphasis>
on tarsus I eupathidial, located between
<emphasis id="B95E221BFFDFFFB8FB2EFB02D9BA0C49" box="[1158,1190,1267,1293]" italics="true" pageId="13" pageNumber="189">(u)</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FB75FB02D9E20C49" box="[1245,1278,1267,1293]" italics="true" pageId="13" pageNumber="189">(a)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFAE6DFFC0CD9" pageId="13" pageNumber="189" type="discussion">
<paragraph id="8B95FE09FFDFFFB8FF6FFAE6DFFC0CD9" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFAE6DC270C75" bold="true" box="[199,315,1303,1329]" pageId="13" pageNumber="189">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C2A858AFFDFFFB8FD3DFAE6D9490C75" authority="(Ramsay, 1966)" baseAuthorityName="Ramsay" baseAuthorityYear="1966" box="[661,1109,1303,1329]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="scheloribatoides" subGenus="Topobates">
<emphasis id="B95E221BFFDFFFB8FD3DFAE6DFB60C75" box="[661,682,1303,1329]" italics="true" pageId="13" pageNumber="189">S.</emphasis>
(
<emphasis id="B95E221BFFDFFFB8FD13FAE6DFCC0C74" box="[699,720,1303,1328]" italics="true" pageId="13" pageNumber="189">T.</emphasis>
)
<emphasis id="B95E221BFFDFFFB8FD49FAE6DE860C75" box="[737,922,1303,1329]" italics="true" pageId="13" pageNumber="189">scheloribatoides</emphasis>
(
<bibRefCitation id="EFBB83F8FFDFFFB8FC04FAE6D9500C75" author="Ramsay, G. W." box="[940,1100,1303,1329]" pageId="13" pageNumber="189" pagination="416 - 425" refId="ref8289" refString="Ramsay, G. W. (1966) Two new oribatid mites (Acari) from a New Zealand pasture. New Zealand Journal of Science, 9, 416 - 425." type="journal article" year="1966">Ramsay, 1966</bibRefCitation>
)
</taxonomicName>
(=
<taxonomicName id="4C2A858AFFDFFFB8FBDEFAE6DDCC0C11" authority="Hammer, 1967" authorityName="Hammer" authorityYear="1967" class="Arachnida" family="Scheloribatidae" genus="Setobates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B95E221BFFDFFFB8FBDEFAE6D8300C74" box="[1142,1324,1303,1329]" italics="true" pageId="13" pageNumber="189">Setobates minor</emphasis>
<bibRefCitation id="EFBB83F8FFDFFFB8FA9CFAE6DDCC0C11" author="Hammer, M." pageId="13" pageNumber="189" pagination="1 - 60" refId="ref8004" refString="Hammer, M. (1967) Investigations on the oribatid fauna of New Zealand. Part II. Det Kongelige Danske Videnskabernes Selskab Biologiske Skrifter, 15, 1 - 60." type="journal article" year="1967">Hammer, 1967</bibRefCitation>
</taxonomicName>
) from
<collectingCountry id="F33DBE99FFDFFFB8FE8BFACADCA70C11" box="[291,443,1339,1365]" name="New Zealand" pageId="13" pageNumber="189">New Zealand</collectingCountry>
in the presence of lanceolate bothridial seta, 13 pairs of minute notogastral setae and tridactylous legs, but differs from the latter by the absence (versus presence) of the prolamella, distinctly longer aggenital, anal and adanal setae, and narrower body.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFA56D8250C85" box="[199,1337,1447,1473]" pageId="13" pageNumber="189" type="etymology">
<paragraph id="8B95FE09FFDFFFB8FF6FFA56D8250C85" blockId="13.[151,1437,150,1473]" box="[199,1337,1447,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFA56DC520C85" bold="true" box="[199,334,1447,1473]" pageId="13" pageNumber="189">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2A858AFFDFFFB8FD91FA56DF860C85" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[569,666,1447,1473]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="tredecim">
<emphasis id="B95E221BFFDFFFB8FD91FA56DF860C85" box="[569,666,1447,1473]" italics="true" pageId="13" pageNumber="189">tredecim</emphasis>
</taxonomicName>
refers to the presence of 13 pairs of setae on the notogaster.
</paragraph>
</subSubSection>
</treatment>
</document>