Add updates up until 2024-08-05 07:11:16

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<document id="8AC8C65257F7BFC7A5814600D210299B" ID-DOI="10.11646/zootaxa.5492.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13219142" ID-ZooBank="91164DD5-609C-4415-A659-87A28A010F6D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722840936672" checkinUser="plazi" docAuthor="Ermilov, Sergey G., Rybalov, Leonid B. &amp; Kuzmicheva, Evgeniya A." docDate="2024" docId="03834F1FFFD3FFB0FF3FFA57D97A0805" docLanguage="en" docName="zootaxa.5492.1.2.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scheloribates (Scheloribates) flexibilis Ermilov 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="181" masterDocId="FFBA3767FFD2FFB5FFA8FFF1DD1C0944" masterDocTitle="New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia" masterLastPageNumber="190" masterPageNumber="176" pageNumber="177" updateTime="1722841352648" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EE65A8AF347D18C80DD4D77216FA64C7">New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia</mods:title>
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<mods:namePart id="A7E7D7B6B2F2BE35C8762FC9B1A53F8E">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="54BC139EC0F3DF13FF8ECB6876746BF4">Tyumen State University, Institute of Environmental and Agricultural Biology (X-BIO), Tyumen, Russia</mods:affiliation>
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<mods:namePart id="A61CF71A74DC0EF9D6D5D745F7E18F7A">Rybalov, Leonid B.</mods:namePart>
<mods:affiliation id="81093A38564D05EB686EB2E4283ACA6A">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Soil Zoology and General Entomology, Moscow, Russia</mods:affiliation>
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<mods:namePart id="3DBD6B9A62822EDF942FB1BEB33D3C31">Kuzmicheva, Evgeniya A.</mods:namePart>
<mods:affiliation id="28163A537F90155EBD2E6EE5FA8EA1C5">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Historical Ecology, Moscow, Russia</mods:affiliation>
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<mods:date id="5EEF8B76DC3725B78D4C7CBB26112092">2024</mods:date>
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<treatment id="03834F1FFFD3FFB0FF3FFA57D97A0805" ID-DOI="http://doi.org/10.5281/zenodo.13219148" ID-Zenodo-Dep="13219148" LSID="urn:lsid:plazi:treatment:03834F1FFFD3FFB0FF3FFA57D97A0805" httpUri="http://treatment.plazi.org/id/03834F1FFFD3FFB0FF3FFA57D97A0805" lastPageId="5" lastPageNumber="181" pageId="1" pageNumber="177">
<subSubSection id="C330AD82FFD3FFB4FF3FFA57DE260C85" box="[151,826,1446,1473]" pageId="1" pageNumber="177" type="nomenclature">
<paragraph id="8B95FE09FFD3FFB4FF3FFA57DE260C85" blockId="1.[151,826,1446,1509]" box="[151,826,1446,1473]" pageId="1" pageNumber="177">
<heading id="D0DD4965FFD3FFB4FF3FFA57DE260C85" bold="true" box="[151,826,1446,1473]" fontSize="11" level="1" pageId="1" pageNumber="177" reason="1">
<emphasis id="B95E221BFFD3FFB4FF3FFA57DE260C85" bold="true" box="[151,826,1446,1473]" pageId="1" pageNumber="177">
<taxonomicName id="4C2A858AFFD3FFB4FF3FFA57DFCA0C84" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[151,726,1446,1473]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="177" phylum="Arthropoda" rank="species" species="flexibilis" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD3FFB4FF3FFA57DC260C85" bold="true" box="[151,314,1446,1473]" italics="true" pageId="1" pageNumber="177">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD3FFB4FEE3FA57DCF20C85" bold="true" box="[331,494,1446,1473]" italics="true" pageId="1" pageNumber="177">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD3FFB4FE57FA57DF7B0C85" bold="true" box="[511,615,1446,1473]" italics="true" pageId="1" pageNumber="177">flexibilis</emphasis>
Ermilov
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD3FFB4FD75FA56DE260C85" box="[733,826,1447,1473]" pageId="1" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD3FFB4FF3FFA3ADC100CA1" box="[151,268,1483,1509]" pageId="1" pageNumber="177" type="description">
<paragraph id="8B95FE09FFD3FFB4FF3FFA3ADC100CA1" blockId="1.[151,826,1446,1509]" box="[151,268,1483,1509]" pageId="1" pageNumber="177">
(
<figureCitation id="1311E28CFFD3FFB4FF37FA3ADDFA0CA1" box="[159,230,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1877]" captionTargetBox="[185,1421,194,1812]" captionTargetId="figure-19@2.[151,1436,179,1827]" captionTargetPageId="2" captionText="FIGURE 1. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="1" pageNumber="177">Figs 1</figureCitation>
,
<figureCitation id="1311E28CFFD3FFB4FF5CFA3ADC1F0CA1" box="[244,259,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1838,1862]" captionTargetBox="[164,1423,194,1801]" captionTargetId="figure-28@3.[151,1436,181,1814]" captionTargetPageId="3" captionText="FIGURE 2. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (AC), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219146" httpUri="https://zenodo.org/record/13219146/files/figure.png" pageId="1" pageNumber="177">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD3FFB4FF3FF9E3D91A0EFD" pageId="1" pageNumber="177" type="materials_examined">
<paragraph id="8B95FE09FFD3FFB4FF3FF9E3DF6B0E09" blockId="1.[151,1437,1554,1977]" pageId="1" pageNumber="177">
<emphasis id="B95E221BFFD3FFB4FF3FF9E3DC100F68" bold="true" box="[151,268,1554,1580]" pageId="1" pageNumber="177">Material.</emphasis>
<typeStatus id="549140ABFFD3FFB4FEB1F9E2DC9B0F69" box="[281,391,1555,1581]" pageId="1" pageNumber="177" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D2C3580FFD3FFB4FD94F9E2DFE20F69" box="[572,766,1555,1581]" count="7" pageId="1" pageNumber="177" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
(
<specimenCount id="9D2C3580FFD3FFB4FCBAF9E2DE8E0F69" box="[786,914,1555,1581]" count="2" pageId="1" pageNumber="177" type="male">two males</specimenCount>
and
<specimenCount id="9D2C3580FFD3FFB4FC7DF9E2D96F0F69" box="[981,1139,1555,1581]" count="5" pageId="1" pageNumber="177" type="female">five females</specimenCount>
): Southeastern
<collectingCountry id="F33DBE99FFD3FFB4FA99F9E2D8840F69" box="[1329,1432,1555,1581]" name="Ethiopia" pageId="1" pageNumber="177">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD3FFB4FF3FF9C6DC4F0F15" box="[151,339,1591,1617]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="177">Oromia Region</collectingRegion>
, Bale Zone, Bale Mountains National Park,
<geoCoordinate id="EE1E98CEFFD3FFB4FCF5F9C9DEF10F17" box="[861,1005,1592,1619]" degrees="6" direction="north" minutes="48" orientation="latitude" pageId="1" pageNumber="177" precision="1" seconds="40.7" value="6.8113055">6°4840.7″N</geoCoordinate>
,
<geoCoordinate id="EE1E98CEFFD3FFB4FC53F9C9D98A0F17" box="[1019,1174,1592,1619]" degrees="39" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="177" precision="1" seconds="07.7" value="39.85214">39°5107.7″E</geoCoordinate>
,
<quantity id="4CD253ECFFD3FFB4FB0DF9C7D9E20F15" box="[1189,1278,1590,1617]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="1" pageNumber="177" unit="m" value="4050.0">4050 m</quantity>
a.s.l., Sanetti Plateau, Afroalpine community, predominance of cushion-shaped shrubs of
<taxonomicName id="4C2A858AFFD3FFB4FB89F9AAD8230F30" box="[1057,1343,1627,1653]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B95E221BFFD3FFB4FB89F9AAD8230F30" box="[1057,1343,1627,1653]" italics="true" pageId="1" pageNumber="177">Helichrysum citrispinum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A858AFFD3FFB4FA29F9AADC050FDD" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="splendidum">
<emphasis id="B95E221BFFD3FFB4FA29F9AADC050FDD" italics="true" pageId="1" pageNumber="177">H. splendidum</emphasis>
</taxonomicName>
, co-dominance of
<taxonomicName id="4C2A858AFFD3FFB4FE5BF98EDFA60FDD" box="[499,698,1663,1689]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="richardii">
<emphasis id="B95E221BFFD3FFB4FE5BF98EDFA60FDD" box="[499,698,1663,1689]" italics="true" pageId="1" pageNumber="177">Festuca richardii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFD3FFB4FD61F98EDEFD0FDD" box="[713,993,1663,1689]" class="Liliopsida" family="Poaceae" genus="Pentaschistis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="pictigluma">
<emphasis id="B95E221BFFD3FFB4FD61F98EDEFD0FDD" box="[713,993,1663,1689]" italics="true" pageId="1" pageNumber="177">Pentaschistis pictigluma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFD3FFB4FC59F98ED9FD0FDC" box="[1009,1249,1663,1689]" class="Liliopsida" family="Poaceae" genus="Agrostis" kingdom="Plantae" order="Poales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="quinqueseta">
<emphasis id="B95E221BFFD3FFB4FC59F98ED9FD0FDC" box="[1009,1249,1663,1689]" italics="true" pageId="1" pageNumber="177">Agrostis quinqueseta</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A858AFFD3FFB4FA8AF98EDC0C0FF9" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="haumannii">
<emphasis id="B95E221BFFD3FFB4FA8AF98EDC0C0FF9" italics="true" pageId="1" pageNumber="177">Alchemilla haumannii</emphasis>
</taxonomicName>
, litter,
<date id="FF94D8C9FFD3FFB4FEC4F952DCF10FF9" box="[364,493,1699,1725]" pageId="1" pageNumber="177" value="2014-11-16">16.XI.2014</date>
(E. Kuzmicheva).
<specimenCount id="9D2C3580FFD3FFB4FD79F952DE940FF9" box="[721,904,1699,1725]" count="8" pageId="1" pageNumber="177" type="generic" typeStatus="paratype">Eight paratypes</specimenCount>
(
<specimenCount id="9D2C3580FFD3FFB4FC32F952D9050FF9" box="[922,1049,1699,1725]" count="4" pageId="1" pageNumber="177" type="male">four males</specimenCount>
and
<specimenCount id="9D2C3580FFD3FFB4FBF2F952D9EF0FF9" box="[1114,1267,1699,1725]" count="4" pageId="1" pageNumber="177" type="female">four females</specimenCount>
): Southeastern
<collectingCountry id="F33DBE99FFD3FFB4FF3FF936DDE60FA5" box="[151,250,1735,1761]" name="Ethiopia" pageId="1" pageNumber="177">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD3FFB4FEAFF936DCA70FA5" box="[263,443,1735,1761]" country="Ethiopia" name="Oromiya" pageId="1" pageNumber="177">Oromia Region</collectingRegion>
, Bale Zone, Bale Mountains National Park,
<geoCoordinate id="EE1E98CEFFD3FFB4FC12F939D9560FA7" box="[954,1098,1736,1763]" degrees="6" direction="north" minutes="43" orientation="latitude" pageId="1" pageNumber="177" precision="1" seconds="18.1" value="6.7216945">6°4318.1″N</geoCoordinate>
,
<geoCoordinate id="EE1E98CEFFD3FFB4FBF0F939D9E80FA7" box="[1112,1268,1736,1763]" degrees="39" direction="east" minutes="44" orientation="longitude" pageId="1" pageNumber="177" precision="1" seconds="12.1" value="39.736694">39°4412.1″E</geoCoordinate>
,
<quantity id="4CD253ECFFD3FFB4FAA8F937D8440FA5" box="[1280,1368,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="1" pageNumber="177" unit="m" value="2570.0">2570 m</quantity>
a.s.l., Harenna Forest, litter in bamboo belt (vegetation represented by dense bamboo thickets of
<taxonomicName id="4C2A858AFFD3FFB4FB18F91AD8760E41" box="[1200,1386,1771,1797]" class="Liliopsida" family="Poaceae" genus="Oldeania" kingdom="Plantae" order="Poales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="B95E221BFFD3FFB4FB18F91AD8760E41" box="[1200,1386,1771,1797]" italics="true" pageId="1" pageNumber="177">Oldeania alpina</emphasis>
</taxonomicName>
, the African alpine bamboo,
<emphasis id="B95E221BFFD3FFB4FE18F8FEDEA50E6D" box="[432,953,1807,1833]" italics="true" pageId="1" pageNumber="177">
<taxonomicName id="4C2A858AFFD3FFB4FE18F8FEDFE20E6C" authority=", Myrsine" authorityName="Myrsine" box="[432,766,1807,1833]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="abyssinica">Hagenia abyssinica, Myrsine</taxonomicName>
melanophloeos,
</emphasis>
and
<taxonomicName id="4C2A858AFFD3FFB4FC5CF8FED9C90E6D" box="[1012,1237,1807,1833]" class="Magnoliopsida" family="Araliaceae" genus="Schefflera" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="177" phylum="Tracheophyta" rank="species" species="volkensii">
<emphasis id="B95E221BFFD3FFB4FC5CF8FED9C90E6D" box="[1012,1237,1807,1833]" italics="true" pageId="1" pageNumber="177">Schefflera volkensii</emphasis>
</taxonomicName>
trees accompany bamboo),
<date id="FF94D8C9FFD3FFB4FEA2F8C3DC960E09" box="[266,394,1842,1869]" pageId="1" pageNumber="177" value="2014-11-15">15.XI.2014</date>
(E.A. Kuzmicheva).
</paragraph>
<paragraph id="8B95FE09FFD3FFB4FF6FF8A6D91A0EFD" blockId="1.[151,1437,1554,1977]" pageId="1" pageNumber="177">
The
<typeStatus id="549140ABFFD3FFB4FF51F8A6DC400E35" box="[249,348,1879,1905]" pageId="1" pageNumber="177" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
<collectingCountry id="F33DBE99FFD3FFB4FA83F8A6D8840E35" box="[1323,1432,1879,1905]" name="Germany" pageId="1" pageNumber="177">Germany</collectingCountry>
;
<materialsCitation id="3B42F454FFD3FFB4FF3FF88BD8760ED1" box="[151,1386,1914,1941]" country="Russia" location="University Museum of Zoology" pageId="1" pageNumber="177" specimenCount="15" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D2C3580FFD3FFB4FF3FF88BDC370ED1" box="[151,299,1914,1941]" count="15" pageId="1" pageNumber="177" type="generic" typeStatus="paratype">15 paratypes</specimenCount>
are in the collection of the
<collectingRegion id="49EE30EBFFD3FFB4FDD3F88ADE010ED1" box="[635,797,1915,1941]" country="Russia" name="Tyumen'" pageId="1" pageNumber="177">Tyumen State</collectingRegion>
<location id="8EF5A8D2FFD3FFB4FC8FF88AD9800ED1" LSID="urn:lsid:plazi:treatment:03834F1FFFD3FFB0FF3FFA57D97A0805:8EF5A8D2FFD3FFB4FC8FF88AD9800ED1" box="[807,1180,1915,1941]" country="Russia" name="University Museum of Zoology" pageId="1" pageNumber="177" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="49EE30EBFFD3FFB4FB03F88AD8100ED1" box="[1195,1292,1915,1941]" country="Russia" name="Tyumen'" pageId="1" pageNumber="177">Tyumen</collectingRegion>
,
<collectingCountry id="F33DBE99FFD3FFB4FAB3F88AD8760ED1" box="[1307,1386,1915,1941]" name="Russia" pageId="1" pageNumber="177">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in a solution of 70% ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<caption id="DF55AE81FFD0FFB7FF3FF8CCD9120E3D" ID-DOI="http://doi.org/10.5281/zenodo.13219144" ID-Zenodo-Dep="13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="2" pageNumber="178" startId="2.[151,250,1853,1877]" targetBox="[185,1421,194,1812]" targetPageId="2" targetType="figure">
<paragraph id="8B95FE09FFD0FFB7FF3FF8CCD9120E3D" blockId="2.[151,1436,1853,1913]" pageId="2" pageNumber="178">
<emphasis id="B95E221BFFD0FFB7FF3FF8CCDC090E11" bold="true" box="[151,277,1853,1877]" pageId="2" pageNumber="178">FIGURE 1.</emphasis>
<taxonomicName id="4C2A858AFFD0FFB7FEB4F8CCDE1E0E11" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[284,770,1853,1877]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="178" phylum="Arthropoda" rank="species" species="flexibilis" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD0FFB7FEB4F8CCDCBA0E11" box="[284,422,1853,1877]" italics="true" pageId="2" pageNumber="178">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD0FFB7FE1EF8CCDF5C0E11" box="[438,576,1853,1877]" italics="true" pageId="2" pageNumber="178">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD0FFB7FDF8F8CCDFBB0E11" box="[592,679,1853,1877]" italics="true" pageId="2" pageNumber="178">flexibilis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD0FFB7FCA2F8CCDE400E11" bold="true" box="[778,860,1853,1877]" pageId="2" pageNumber="178">
<taxonomicNameLabel id="A26D9F60FFD0FFB7FCA2F8CCDE400E11" box="[778,860,1853,1877]" pageId="2" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<caption id="DF55AE81FFD1FFB6FF3FF8DFD9180ECA" ID-DOI="http://doi.org/10.5281/zenodo.13219146" ID-Zenodo-Dep="13219146" httpUri="https://zenodo.org/record/13219146/files/figure.png" pageId="3" pageNumber="179" startId="3.[151,250,1838,1862]" targetBox="[164,1423,194,1801]" targetPageId="3" targetType="figure">
<paragraph id="8B95FE09FFD1FFB6FF3FF8DFD9180ECA" blockId="3.[151,1436,1838,1934]" pageId="3" pageNumber="179">
<emphasis id="B95E221BFFD1FFB6FF3FF8DFDC090E02" bold="true" box="[151,277,1838,1862]" pageId="3" pageNumber="179">FIGURE 2.</emphasis>
<taxonomicName id="4C2A858AFFD1FFB6FEB5F8DFDE190E02" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[285,773,1838,1862]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="179" phylum="Arthropoda" rank="species" species="flexibilis" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD1FFB6FEB5F8DFDCBB0E02" box="[285,423,1838,1862]" italics="true" pageId="3" pageNumber="179">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD1FFB6FE10F8DFDF5E0E02" box="[440,578,1838,1862]" italics="true" pageId="3" pageNumber="179">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD1FFB6FDFBF8DFDFB60E02" box="[595,682,1838,1862]" italics="true" pageId="3" pageNumber="179">flexibilis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD1FFB6FCA5F8DFDE7C0E02" bold="true" box="[781,864,1838,1862]" pageId="3" pageNumber="179">
<taxonomicNameLabel id="A26D9F60FFD1FFB6FCA5F8DFDE7C0E02" box="[781,864,1838,1862]" pageId="3" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (AC), 50 μm (DG).
</paragraph>
</caption>
<subSubSection id="C330AD82FFD6FFB1FF6FFF67DF690821" pageId="4" pageNumber="180" type="diagnosis">
<paragraph id="8B95FE09FFD6FFB1FF6FFF67DF690821" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFF67DC5E09F4" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="180">Diagnosis.</emphasis>
Body length: 480555.
<taxonomicName id="4C2A858AFFD6FFB1FDCBFF66DFC209F5" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[611,734,151,177]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Rostrum rounded. Prolamella complete; translamella represented by two short lines near lamellae. Rostral, lamellar and interlamellar setae long, setiform, barbed;
<emphasis id="B95E221BFFD6FFB1FF5AFF2EDC1409BC" box="[242,264,223,248]" italics="true" pageId="4" pageNumber="180">in</emphasis>
˃
<emphasis id="B95E221BFFD6FFB1FEB0FF2EDC3009BD" box="[280,300,223,249]" italics="true" pageId="4" pageNumber="180">le</emphasis>
˃
<emphasis id="B95E221BFFD6FFB1FE94FF2EDC4809BC" box="[316,340,223,248]" italics="true" pageId="4" pageNumber="180">ro</emphasis>
; bothridial seta long, lanceolate, barbed. All notogastral setae medium-sized, flexible, roughened. Circumpedal carina long. Anal and adanal setae medium-sized, setiform to flexible, thin, roughened. Ventrobasal tubercle of leg tibiae I, II well observable; femur II broadly rounded ventrodistally; tarsus I with 18 setae (
<emphasis id="B95E221BFFD6FFB1FA91FED6D84C0805" box="[1337,1360,295,321]" italics="true" pageId="4" pageNumber="180">l”</emphasis>
and
<emphasis id="B95E221BFFD6FFB1FA2EFED6D8800805" box="[1414,1436,295,321]" italics="true" pageId="4" pageNumber="180">v</emphasis>
absent); genu II with two setae (
<emphasis id="B95E221BFFD6FFB1FDA8FEBADF0A0821" box="[512,534,331,357]" italics="true" pageId="4" pageNumber="180">v</emphasis>
absent).
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD6FFB1FF6FFE9FD8080F4D" pageId="4" pageNumber="180" type="description">
<paragraph id="8B95FE09FFD6FFB1FF6FFE9FD85D0895" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFE9FDCDE08CD" bold="true" box="[199,450,366,393]" pageId="4" pageNumber="180">Description of adult</emphasis>
.
<emphasis id="B95E221BFFD6FFB1FE7FFE9EDF6508CC" box="[471,633,367,392]" italics="true" pageId="4" pageNumber="180">Measurements</emphasis>
. Body length: 555 (
<typeStatus id="549140ABFFD6FFB1FCDBFE9EDEC708CD" box="[883,987,367,393]" pageId="4" pageNumber="180" type="holotype">holotype</typeStatus>
), 480525 (male
<typeStatus id="549140ABFFD6FFB1FB19FE9ED83D08CD" box="[1201,1313,367,393]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
), 525555 (female
<typeStatus id="549140ABFFD6FFB1FF5BFE62DC7808E9" box="[243,356,403,429]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
); body width (level of pteromorph): 360 (
<typeStatus id="549140ABFFD6FFB1FC9AFE62DE8608E9" box="[818,922,403,429]" pageId="4" pageNumber="180" type="holotype">holotype</typeStatus>
), 315330 (male
<typeStatus id="549140ABFFD6FFB1FBF4FE62D9D008E9" box="[1116,1228,403,429]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
), 330360 (female
<typeStatus id="549140ABFFD6FFB1FF3FFE46DC140895" box="[151,264,439,465]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
); width of ventral plate: 330 (
<typeStatus id="549140ABFFD6FFB1FDFAFE46DFA60895" box="[594,698,439,465]" pageId="4" pageNumber="180" type="holotype">holotype</typeStatus>
), 285300 (male
<typeStatus id="549140ABFFD6FFB1FCD5FE46DEF10895" box="[893,1005,439,465]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
), 300330 (female
<typeStatus id="549140ABFFD6FFB1FB6DFE46D82A0895" box="[1221,1334,439,465]" pageId="4" pageNumber="180" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFE2ADC2B0B79" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFE2ADC5908B0" box="[199,325,475,500]" italics="true" pageId="4" pageNumber="180">Integument</emphasis>
. Body color light brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens; ×1000); podosomal region and lateral part of prodorsum partially with dense microgranulate cerotegument.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFDB6DCB10A51" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFDB6DC580B25" box="[199,324,583,609]" italics="true" pageId="4" pageNumber="180">Prodorsum</emphasis>
(
<figureCitation id="1311E28CFFD6FFB1FEFFFDB6DCC30B25" box="[343,479,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1877]" captionTargetBox="[185,1421,194,1812]" captionTargetId="figure-19@2.[151,1436,179,1827]" captionTargetPageId="2" captionText="FIGURE 1. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="4" pageNumber="180">Figs 1A, C</figureCitation>
). Rostrum rounded. Lamella about 1/2 length of prodorsum; prolamella complete (reaching insertion of
<emphasis id="B95E221BFFD6FFB1FE3DFD9ADCB10BC0" box="[405,429,619,644]" italics="true" pageId="4" pageNumber="180">ro</emphasis>
); translamella represented by two short lines near lamellae; sublamella and lateral keel-shaped ridge distinct; sublamellar porose area (911 × 7) oval. Rostral (7175), lamellar (101112) and interlamellar (118131) setae setiform, barbed; bothridial seta (7790) with lanceolate, barbed head, directed posterolaterad; bothridial stalk slightly longer than head; exobothridial seta (3037) setiform, thin, roughened. Dorsosejugal porose area oval, poorly visible.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFCEED9B00AE1" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<taxonomicName id="4C2A858AFFD6FFB1FF6FFCEEDC5F0A7C" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,799,824]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B95E221BFFD6FFB1FF6FFCEEDC5F0A7C" box="[199,323,799,824]" italics="true" pageId="4" pageNumber="180">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFD6FFB1FEF3FCEEDCF70A7D" box="[347,491,799,825]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1877]" captionTargetBox="[185,1421,194,1812]" captionTargetId="figure-19@2.[151,1436,179,1827]" captionTargetPageId="2" captionText="FIGURE 1. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="4" pageNumber="180">Figs 1A, C</figureCitation>
). Anterior notogastral margin convex medially.
<taxonomicName id="4C2A858AFFD6FFB1FB9DFCEED9AC0A7D" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[1077,1200,799,825]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Pteromorph medium-sized, broadly rounded laterally, its anterior margin straight (perpendicular to longitudinal axis of body in dorsal aspect). Ten pairs of notogastral setae (3741) flexible, roughened. Four pairs of saccules with small opening and drop-like channel. Opisthonotal gland opening and all lyrifissures distinct.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFC5EDDF70D71" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<taxonomicName id="4C2A858AFFD6FFB1FF6FFC5EDC4A0A8D" authorityName="Krefft" authorityYear="1871" box="[199,342,943,969]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="180" phylum="Nematoda" rank="genus">
<emphasis id="B95E221BFFD6FFB1FF6FFC5EDC4A0A8D" box="[199,342,943,969]" italics="true" pageId="4" pageNumber="180">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFD6FFB1FECFFC5EDCF60A8D" box="[359,490,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1838,1862]" captionTargetBox="[164,1423,194,1801]" captionTargetId="figure-28@3.[151,1436,181,1814]" captionTargetPageId="3" captionText="FIGURE 2. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (AC), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219146" httpUri="https://zenodo.org/record/13219146/files/figure.png" pageId="4" pageNumber="180">Figs 2AC</figureCitation>
). Subcapitulum size: 116120 × 8286; subcapitular setae (
<emphasis id="B95E221BFFD6FFB1FB3CFC5ED9BE0A8C" box="[1172,1186,943,968]" italics="true" pageId="4" pageNumber="180">a</emphasis>
: 2226;
<emphasis id="B95E221BFFD6FFB1FAA2FC5ED8020A8C" box="[1290,1310,943,968]" italics="true" pageId="4" pageNumber="180">m</emphasis>
: 1922;
<emphasis id="B95E221BFFD6FFB1FA2EFC5ED8880A8D" box="[1414,1428,943,969]" italics="true" pageId="4" pageNumber="180">h</emphasis>
: 3437) setiform, roughened;
<emphasis id="B95E221BFFD6FFB1FE73FC22DCF30AA8" box="[475,495,979,1004]" italics="true" pageId="4" pageNumber="180">m</emphasis>
thinner than
<emphasis id="B95E221BFFD6FFB1FD29FC22DF930AA8" box="[641,655,979,1004]" italics="true" pageId="4" pageNumber="180">a</emphasis>
and
<emphasis id="B95E221BFFD6FFB1FD6BFC22DFCD0AA9" box="[707,721,979,1005]" italics="true" pageId="4" pageNumber="180">h</emphasis>
; both adoral setae (1315) setiform, barbed. Palp length: 7175; postpalpal seta (5) spiniform, roughened. Chelicera length: 123131; setae (
<emphasis id="B95E221BFFD6FFB1FBA6FC06D92A0D55" box="[1038,1078,1015,1041]" italics="true" pageId="4" pageNumber="180">cha</emphasis>
: 4145;
<emphasis id="B95E221BFFD6FFB1FB0AFC06D9D60D55" box="[1186,1226,1015,1041]" italics="true" pageId="4" pageNumber="180">chb</emphasis>
: 2630) setiform, barbed.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFBCEDC270D81" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFBCEDF890D1C" box="[199,661,1087,1113]" italics="true" pageId="4" pageNumber="180">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="1311E28CFFD6FFB1FD0EFBCEDE3B0D1D" box="[678,807,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1877]" captionTargetBox="[185,1421,194,1812]" captionTargetId="figure-19@2.[151,1436,179,1827]" captionTargetPageId="2" captionText="FIGURE 1. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="4" pageNumber="180">Figs 1B, C</figureCitation>
). Epimeral formula: 3133; all setae (
<emphasis id="B95E221BFFD6FFB1FB5FFBCED80F0D1D" box="[1271,1299,1087,1113]" italics="true" pageId="4" pageNumber="180">1b</emphasis>
,
<emphasis id="B95E221BFFD6FFB1FA8AFBCED8220D1D" box="[1314,1342,1087,1113]" italics="true" pageId="4" pageNumber="180">3b</emphasis>
,
<emphasis id="B95E221BFFD6FFB1FAE6FBCED8740D1D" box="[1358,1384,1087,1113]" italics="true" pageId="4" pageNumber="180">3c</emphasis>
,
<emphasis id="B95E221BFFD6FFB1FAD0FBCED8890D1D" box="[1400,1429,1087,1113]" italics="true" pageId="4" pageNumber="180">4a</emphasis>
: 2832; others: 1922) setiform, thin, roughened. Humeral porose areas
<emphasis id="B95E221BFFD6FFB1FC62FB92DEF30D39" box="[970,1007,1123,1149]" italics="true" pageId="4" pageNumber="180">Am</emphasis>
and
<emphasis id="B95E221BFFD6FFB1FB81FB92D9540D39" box="[1065,1096,1123,1149]" italics="true" pageId="4" pageNumber="180">Ah</emphasis>
elongate oval, poorly visible. Pedotectum II rounded laterally in ventral aspect. Discidium broadly rounded. Circumpedal carina long, directed to pedotectum II.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFB3EDE020C75" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFB3EDC8D0DAC" box="[199,401,1231,1257]" italics="true" pageId="4" pageNumber="180">Anogenital region</emphasis>
(
<figureCitation id="1311E28CFFD6FFB1FE08FB3EDF010DAD" box="[416,541,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1853,1877]" captionTargetBox="[185,1421,194,1812]" captionTargetId="figure-19@2.[151,1436,179,1827]" captionTargetPageId="2" captionText="FIGURE 1. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219144" httpUri="https://zenodo.org/record/13219144/files/figure.png" pageId="4" pageNumber="180">Figs 1B, C</figureCitation>
). Genital (
<emphasis id="B95E221BFFD6FFB1FD3CFB3EDFBE0DAC" box="[660,674,1231,1256]" italics="true" pageId="4" pageNumber="180">g</emphasis>
<subScript id="17AEFC4CFFD6FFB1FD0AFB10DFB60DAB" attach="left" box="[674,682,1249,1263]" fontSize="6" pageId="4" pageNumber="180">1</subScript>
: 2426;
<emphasis id="B95E221BFFD6FFB1FCA5FB3EDE070DAC" box="[781,795,1231,1256]" italics="true" pageId="4" pageNumber="180">g</emphasis>
<subScript id="17AEFC4CFFD6FFB1FCB3FB10DE3F0DAB" attach="left" box="[795,803,1249,1263]" fontSize="6" pageId="4" pageNumber="180">2</subScript>
<emphasis id="B95E221BFFD6FFB1FC99FB3EDE230DAC" box="[817,831,1231,1256]" italics="true" pageId="4" pageNumber="180">g</emphasis>
<subScript id="17AEFC4CFFD6FFB1FC97FB10DE5B0DAB" attach="left" box="[831,839,1249,1263]" fontSize="6" pageId="4" pageNumber="180">4</subScript>
: 1719), aggenital (3032), anal (2830), and adanal (3032) setae setiform (anal and adanal setae sometimes flexible), thin, roughened. Adanal lyrifissure distinct, close and parallel to anal plate. Marginal porose area band-like.
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF6FFACAD8080F4D" blockId="4.[151,1437,150,1545]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF6FFACADDE00C10" box="[199,252,1339,1364]" italics="true" pageId="4" pageNumber="180">Legs</emphasis>
(
<figureCitation id="1311E28CFFD6FFB1FEA4FACADC950C11" box="[268,393,1339,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1838,1862]" captionTargetBox="[164,1423,194,1801]" captionTargetId="figure-28@3.[151,1436,181,1814]" captionTargetPageId="3" captionText="FIGURE 2. Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (AC), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219146" httpUri="https://zenodo.org/record/13219146/files/figure.png" pageId="4" pageNumber="180">Figs 2DG</figureCitation>
). Median claw thick; lateral claws thin, with small tubercle distoventrally; all claws slightly barbed on dorsal side. Ventrobasal tubercle of tibiae I, II well observable. Femur II broadly rounded ventrodistally. Proximoventral porose area on tarsi IIV, distoventral porose area on tibiae IIV, dorsoparaxial porose area on femora IIV and on trochanters III, IV distinct. Formulas of leg setation and solenidia: I (153418) [122], II (152415) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="C6A8CBB2FFD6FFB1FE89FA1EDC690F4D" box="[289,373,1519,1545]" captionStart="TABLE 1" captionStartId="4.[151,239,1592,1616]" captionText="TABLE 1. Leg setation and solenidia of adult Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., S. (S.) monosetosus Ermilov sp. nov., S. (S.) triangulus Ermilov sp. nov., and S. (Topobates) tredecim Ermilov sp. nov." pageId="4" pageNumber="180">Table 1</tableCitation>
. Seta
<emphasis id="B95E221BFFD6FFB1FE11FA1EDCD80F4C" box="[441,452,1519,1544]" italics="true" pageId="4" pageNumber="180">s</emphasis>
on tarsus I setiform (not eupathidial), barbed, located between
<emphasis id="B95E221BFFD6FFB1FB21FA1ED9B60F4D" box="[1161,1194,1519,1545]" italics="true" pageId="4" pageNumber="180">(a)</emphasis>
and
<emphasis id="B95E221BFFD6FFB1FB48FA1ED8110F4D" box="[1248,1293,1519,1545]" italics="true" pageId="4" pageNumber="180">(pv)</emphasis>
.
</paragraph>
</subSubSection>
<caption id="DF55AE81FFD6FFB1FF3FF9C9D9AB0F31" pageId="4" pageNumber="180" startId="4.[151,239,1592,1616]" targetType="table">
<paragraph id="8B95FE09FFD6FFB1FF3FF9C9D9AB0F31" blockId="4.[151,1436,1592,1653]" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF3FF9C9DC140F14" bold="true" box="[151,264,1592,1616]" pageId="4" pageNumber="180">TABLE 1.</emphasis>
Leg setation and solenidia of adult
<taxonomicName id="4C2A858AFFD6FFB1FDD0F9C9D9710F14" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[632,1133,1592,1617]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="flexibilis" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD6FFB1FDD0F9C9DE150F14" box="[632,777,1592,1616]" italics="true" pageId="4" pageNumber="180">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FCBFF9C9DEB40F14" box="[791,936,1592,1616]" italics="true" pageId="4" pageNumber="180">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FC1DF9C9D90D0F14" box="[949,1041,1592,1616]" italics="true" pageId="4" pageNumber="180">flexibilis</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD6FFB1FBDAF9C8D9DA0F15" bold="true" box="[1138,1222,1593,1617]" pageId="4" pageNumber="180">
<taxonomicNameLabel id="A26D9F60FFD6FFB1FBDAF9C8D9DA0F15" box="[1138,1222,1593,1617]" pageId="4" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2A858AFFD6FFB1FB79F9C8DDF20F30" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="monosetosus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD6FFB1FB79F9C8D9F90F14" box="[1233,1253,1593,1616]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FB5AF9C8D81B0F14" box="[1266,1287,1593,1616]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FABCF9C8D8800F14" box="[1300,1436,1593,1616]" italics="true" pageId="4" pageNumber="180">monosetosus</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD6FFB1FF5DF9ACDC570F31" bold="true" box="[245,331,1629,1653]" pageId="4" pageNumber="180">
<taxonomicNameLabel id="A26D9F60FFD6FFB1FF5DF9ACDC570F31" box="[245,331,1629,1653]" pageId="4" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C2A858AFFD6FFB1FEF0F9ACDF760F30" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[344,618,1628,1653]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="triangulus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD6FFB1FEF0F9ACDC700F30" box="[344,364,1629,1652]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FED3F9ACDC8C0F30" box="[379,400,1629,1652]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FE37F9ADDF110F30" box="[415,525,1628,1652]" italics="true" pageId="4" pageNumber="180">triangulus</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD6FFB1FDD9F9ACDFDB0F31" bold="true" box="[625,711,1629,1653]" pageId="4" pageNumber="180">
<taxonomicNameLabel id="A26D9F60FFD6FFB1FDD9F9ACDFDB0F31" box="[625,711,1629,1653]" pageId="4" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C2A858AFFD6FFB1FCA9F9ACD9470F30" authority="Ermilov" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[769,1115,1628,1653]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFD6FFB1FCA9F9ACDE090F30" box="[769,789,1629,1652]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FC8DF9ADDE8E0F30" box="[805,914,1628,1652]" italics="true" pageId="4" pageNumber="180">Topobates</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FC09F9ADDEE10F30" box="[929,1021,1628,1652]" italics="true" pageId="4" pageNumber="180">tredecim</emphasis>
Ermilov
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD6FFB1FBCAF9ACD9AB0F31" box="[1122,1207,1629,1653]" pageId="4" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B95FE09FFD6FFB1FF37F977D9850E7C" pageId="4" pageNumber="180">
<table id="F92A0CA9FFD6004AFF37F977D8660E7D" box="[159,1402,1670,1849]" gridcols="6" gridrows="5" pageId="4" pageNumber="180">
<tr id="351AFC4BFFD6004AFF37F977D8660FDF" box="[159,1402,1670,1691]" gridrow="0" pageId="4" pageNumber="180">
<th id="76CB9537FFD6004AFF37F977DDD80FDF" box="[159,196,1670,1691]" gridcol="0" gridrow="0" pageId="4" pageNumber="180">Leg</th>
<th id="76CB9537FFD6004AFF54F977DC350FDF" box="[252,297,1670,1691]" gridcol="1" gridrow="0" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF54F977DC0D0FDF" box="[252,273,1670,1691]" italics="true" pageId="4" pageNumber="180">Tr</emphasis>
</th>
<th id="76CB9537FFD6004AFEFDF977DCC40FDF" box="[341,472,1670,1691]" gridcol="2" gridrow="0" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FEFDF977DC720FDF" box="[341,366,1670,1691]" italics="true" pageId="4" pageNumber="180">Fe</emphasis>
</th>
<th id="76CB9537FFD6004AFD80F977DF930FDF" box="[552,655,1670,1691]" gridcol="3" gridrow="0" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD80F977DF580FDF" box="[552,580,1670,1691]" italics="true" pageId="4" pageNumber="180">Ge</emphasis>
</th>
<th id="76CB9537FFD6004AFD68F977DE230FDF" box="[704,831,1670,1691]" gridcol="4" gridrow="0" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD68F977DFCF0FDF" box="[704,723,1670,1691]" italics="true" pageId="4" pageNumber="180">Ti</emphasis>
</th>
<th id="76CB9537FFD6004AFC39F977D8660FDF" box="[913,1402,1670,1691]" gridcol="5" gridrow="0" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FC39F977DEB40FDF" box="[913,936,1670,1691]" italics="true" pageId="4" pageNumber="180">Ta</emphasis>
</th>
</tr>
<tr id="351AFC4BFFD6004AFF37F95CD8660F8C" box="[159,1402,1709,1736]" gridrow="1" pageId="4" pageNumber="180">
<th id="76CB9537FFD6004AFF37F95CDDD80F8C" box="[159,196,1709,1736]" gridcol="0" gridrow="1" pageId="4" pageNumber="180">I</th>
<td id="76CB9537FFD6004AFF54F95CDC350F8C" box="[252,297,1709,1736]" gridcol="1" gridrow="1" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF54F95CDC130F86" box="[252,271,1709,1730]" italics="true" pageId="4" pageNumber="180">v</emphasis>
</td>
<td id="76CB9537FFD6004AFEFDF95CDCC40F8C" box="[341,472,1709,1736]" gridcol="2" gridrow="1" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FEFDF95CDCC40F86" box="[341,472,1709,1730]" italics="true" pageId="4" pageNumber="180">d, (l), bv”, v”</emphasis>
</td>
<td id="76CB9537FFD6004AFD80F95CDF930F8C" box="[552,655,1709,1736]" gridcol="3" gridrow="1" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD80F95CDF7F0F86" box="[552,611,1709,1730]" italics="true" pageId="4" pageNumber="180">(l), v,</emphasis>
σ
</td>
<td id="76CB9537FFD6004AFD68F95CDE230F8C" box="[704,831,1709,1736]" gridcol="4" gridrow="1" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD68F95CDE1F0F86" box="[704,771,1709,1730]" italics="true" pageId="4" pageNumber="180">(l), (v),</emphasis>
φ 1, φ 2
</td>
<td id="76CB9537FFD6004AFC39F95CD8660F8C" box="[913,1402,1709,1736]" gridcol="5" gridrow="1" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FC39F95CD8380F86" box="[913,1316,1709,1731]" italics="true" pageId="4" pageNumber="180">(ft), (tc), (it), (p), (u), (a), s, (pv), v*, (pl),</emphasis>
ɛ, ω 1, ω 2
</td>
</tr>
<tr id="351AFC4BFFD6004AFF37F925D8660FB4" box="[159,1402,1748,1776]" gridrow="2" pageId="4" pageNumber="180">
<th id="76CB9537FFD6004AFF37F925DDD80FB4" box="[159,196,1748,1776]" gridcol="0" gridrow="2" pageId="4" pageNumber="180">II</th>
<td id="76CB9537FFD6004AFF54F925DC350FB4" box="[252,297,1748,1776]" gridcol="1" gridrow="2" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF54F924DC130FAE" box="[252,271,1749,1770]" italics="true" pageId="4" pageNumber="180">v</emphasis>
</td>
<td id="76CB9537FFD6004AFEFDF925DCC40FB4" box="[341,472,1748,1776]" gridcol="2" gridrow="2" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FEFDF925DCC40FAE" box="[341,472,1748,1770]" italics="true" pageId="4" pageNumber="180">d, (l), bv”, v”</emphasis>
</td>
<td id="76CB9537FFD6004AFD80F925DF930FB4" box="[552,655,1748,1776]" gridcol="3" gridrow="2" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD80F925DF600FAE" box="[552,636,1748,1770]" italics="true" pageId="4" pageNumber="180">(l), v**,</emphasis>
σ
</td>
<td id="76CB9537FFD6004AFD68F925DE230FB4" box="[704,831,1748,1776]" gridcol="4" gridrow="2" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD68F925DE1F0FAE" box="[704,771,1748,1770]" italics="true" pageId="4" pageNumber="180">(l), (v),</emphasis>
φ
</td>
<td id="76CB9537FFD6004AFC39F925D8660FB4" box="[913,1402,1748,1776]" gridcol="5" gridrow="2" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FC39F925D9D70FAE" box="[913,1227,1748,1770]" italics="true" pageId="4" pageNumber="180">(ft), (tc), (it), (p), (u), (a), s, (pv),</emphasis>
ω, ω 1 2
</td>
</tr>
<tr id="351AFC4BFFD6004AFF37F90DD8660E55" box="[159,1402,1788,1809]" gridrow="3" pageId="4" pageNumber="180">
<th id="76CB9537FFD6004AFF37F90DDDD80E55" box="[159,196,1788,1809]" gridcol="0" gridrow="3" pageId="4" pageNumber="180">III</th>
<td id="76CB9537FFD6004AFF54F90DDC350E55" box="[252,297,1788,1809]" gridcol="1" gridrow="3" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF54F90DDC350E55" box="[252,297,1788,1809]" italics="true" pageId="4" pageNumber="180">l, v</emphasis>
</td>
<td id="76CB9537FFD6004AFEFDF90DDCC40E55" box="[341,472,1788,1809]" gridcol="2" gridrow="3" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FEFDF90DDCB90E55" box="[341,421,1788,1809]" italics="true" pageId="4" pageNumber="180">d, l, ev</emphasis>
</td>
<td id="76CB9537FFD6004AFD80F90DDF930E55" box="[552,655,1788,1809]" gridcol="3" gridrow="3" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD80F90DDF210E55" box="[552,573,1788,1809]" italics="true" pageId="4" pageNumber="180">l,</emphasis>
σ
</td>
<td id="76CB9537FFD6004AFD68F90DDE230E55" box="[704,831,1788,1809]" gridcol="4" gridrow="3" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD68F90DDFE70E55" box="[704,763,1788,1809]" italics="true" pageId="4" pageNumber="180">l, (v),</emphasis>
φ
</td>
<td id="76CB9537FFD6004AFC39F90DD8660E55" box="[913,1402,1788,1809]" gridcol="5" gridrow="3" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FC39F90DD9D90E55" box="[913,1221,1788,1809]" italics="true" pageId="4" pageNumber="180">(ft), (tc), (it), (p), (u), (a), s, (pv)</emphasis>
</td>
</tr>
<tr id="351AFC4BFFD6004AFF37F8D2D8660E7D" box="[159,1402,1827,1849]" gridrow="4" pageId="4" pageNumber="180">
<th id="76CB9537FFD6004AFF37F8D2DDD80E7D" box="[159,196,1827,1849]" gridcol="0" gridrow="4" pageId="4" pageNumber="180">IV</th>
<td id="76CB9537FFD6004AFF54F8D2DC350E7D" box="[252,297,1827,1849]" gridcol="1" gridrow="4" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FF54F8D2DC130E7C" box="[252,271,1827,1848]" italics="true" pageId="4" pageNumber="180">v</emphasis>
</td>
<td id="76CB9537FFD6004AFEFDF8D2DCC40E7D" box="[341,472,1827,1849]" gridcol="2" gridrow="4" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FEFDF8D2DC960E7C" box="[341,394,1827,1848]" italics="true" pageId="4" pageNumber="180">d, ev</emphasis>
</td>
<td id="76CB9537FFD6004AFD80F8D2DF930E7D" box="[552,655,1827,1849]" gridcol="3" gridrow="4" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD80F8D2DF530E7C" box="[552,591,1827,1848]" italics="true" pageId="4" pageNumber="180">d, l</emphasis>
</td>
<td id="76CB9537FFD6004AFD68F8D2DE230E7D" box="[704,831,1827,1849]" gridcol="4" gridrow="4" pageId="4" pageNumber="180">
<emphasis id="B95E221BFFD6FFB1FD68F8D2DFE70E7C" box="[704,763,1827,1848]" italics="true" pageId="4" pageNumber="180">l, (v),</emphasis>
φ
</td>
<td id="76CB9537FFD6004AFC39F8D2D8660E7D" box="[913,1402,1827,1849]" gridcol="5" gridrow="4" pageId="4" pageNumber="180">ft”, (tc), (p), (u), (a), s, (pv)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B95FE09FFD6FFB1FF3FF8B8DCBE0E8E" blockId="4.[151,1436,1864,1994]" pageId="4" pageNumber="180">
<tableNote id="76CCFF87FFD6FFB1FF3FF8B8DCBE0E8E" pageId="4" pageNumber="180" targetBox="[159,1402,1670,1849]" targetPageId="4">
Note: Roman letters refer to normal setae, Greek letters to solenidia (except ɛ = famulus); single quotation mark (
<emphasis id="B95E221BFFD6FFB1FB71F8B8D9FD0E1A" box="[1241,1249,1865,1886]" italics="true" pageId="4" pageNumber="180"></emphasis>
) designates seta on the anterior and double quotation mark (
<emphasis id="B95E221BFFD6FFB1FDBEF89CDF3F0EC6" box="[534,547,1901,1922]" italics="true" pageId="4" pageNumber="180"></emphasis>
) seta on the posterior side of a given leg segment; parentheses refer to a pair of setae.
<emphasis id="B95E221BFFD6FFB1FAE7F89CD8470EC6" box="[1359,1371,1901,1922]" italics="true" pageId="4" pageNumber="180">*</emphasis>
—Seta
<emphasis id="B95E221BFFD6FFB1FF3FF860DDB60EE2" box="[151,170,1937,1958]" italics="true" pageId="4" pageNumber="180">v</emphasis>
on tarsus I absent in
<taxonomicName id="4C2A858AFFD6FFB1FEDDF860DF140EE1" authorityName="Ermilov" authorityYear="2024" box="[373,520,1936,1958]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="flexibilis" subGenus="Scheloribates">
<emphasis id="B95E221BFFD6FFB1FEDDF860DC940EE2" box="[373,392,1937,1958]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FE3DF860DCB40EE2" box="[405,424,1937,1958]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FE1DF861DF140EE1" box="[437,520,1936,1957]" italics="true" pageId="4" pageNumber="180">flexibilis</emphasis>
</taxonomicName>
(versus present in the other species);
<emphasis id="B95E221BFFD6FFB1FCD9F860DE820EE2" box="[881,926,1937,1958]" italics="true" pageId="4" pageNumber="180">**—</emphasis>
seta
<emphasis id="B95E221BFFD6FFB1FC62F860DEC10EE2" box="[970,989,1937,1958]" italics="true" pageId="4" pageNumber="180">v</emphasis>
on genu II present in
<taxonomicName id="4C2A858AFFD6FFB1FB06F860D84D0EE1" authorityName="Ermilov" authorityYear="2024" box="[1198,1361,1936,1958]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="180" phylum="Arthropoda" rank="species" species="triangulus" subGenus="Scheloribates">
<emphasis id="B95E221BFFD6FFB1FB06F860D9DD0EE2" box="[1198,1217,1937,1958]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
(
<emphasis id="B95E221BFFD6FFB1FB66F860D9FD0EE2" box="[1230,1249,1937,1958]" italics="true" pageId="4" pageNumber="180">S.</emphasis>
)
<emphasis id="B95E221BFFD6FFB1FB46F861D84D0EE1" box="[1262,1361,1936,1957]" italics="true" pageId="4" pageNumber="180">triangulus</emphasis>
</taxonomicName>
(versus absent in the other species);
</tableNote>
</paragraph>
<subSubSection id="C330AD82FFD7FFB0FF6FFF66D9030859" pageId="5" pageNumber="181" type="discussion">
<paragraph id="8B95FE09FFD7FFB0FF6FFF66D9030859" blockId="5.[151,1436,151,321]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFF66DC2709F5" bold="true" box="[199,315,151,177]" pageId="5" pageNumber="181">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C2A858AFFD7FFB0FD39FF66D9C809F5" authority="Feider, Vasiliu &amp; Calugar, 1973" authorityName="Feider, Vasiliu &amp; Calugar" authorityYear="1973" box="[657,1236,151,177]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="181" phylum="Arthropoda" rank="species" species="longisetosus" subGenus="Scheloribates">
<emphasis id="B95E221BFFD7FFB0FD39FF66DFBA09F5" box="[657,678,151,177]" italics="true" pageId="5" pageNumber="181">S.</emphasis>
(
<emphasis id="B95E221BFFD7FFB0FD1FFF66DFD009F5" box="[695,716,151,177]" italics="true" pageId="5" pageNumber="181">S.</emphasis>
)
<emphasis id="B95E221BFFD7FFB0FD75FF66DE7B09F5" box="[733,871,151,177]" italics="true" pageId="5" pageNumber="181">longisetosus</emphasis>
<bibRefCitation id="EFBB83F8FFD7FFB0FCC7FF66D9C809F5" author="Feider, Z. &amp; Calugar, M. &amp; Vasiliu, N." box="[879,1236,151,177]" pageId="5" pageNumber="181" pagination="25 - 32" refId="ref7925" refString="Feider, Z., Calugar, M. &amp; Vasiliu, N. (1973) Fauna Oribatidelor (Acari) din litiera perdedelor de protectie de la Valul lui Traian. Studii si Cercetari de Biologie, Seria Zoologie, 25, 25 - 32." type="journal article" year="1973">Feider, Vasiliu &amp; Călugăr, 1973</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F33DBE99FFD7FFB0FAB2FF66D86309F5" box="[1306,1407,151,177]" name="Romania" pageId="5" pageNumber="181">Romania</collectingCountry>
in the presence of lanceolate bothridial seta and medium-sized, flexible notogastral setae, but differs from the latter by the presence of medium-sized (versus short) aggenital, anal and adanal setae, and the absence (versus presence) of needleform apex on the bothridial head and anterodorsal tooth on the leg tibia II.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD7FFB0FF6FFED6D97A0805" box="[199,1126,295,321]" pageId="5" pageNumber="181" type="etymology">
<paragraph id="8B95FE09FFD7FFB0FF6FFED6D97A0805" blockId="5.[151,1436,151,321]" box="[199,1126,295,321]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFED6DC520805" bold="true" box="[199,334,295,321]" pageId="5" pageNumber="181">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2A858AFFD7FFB0FD91FED6DF850805" authorityName="Ermilov" authorityYear="2024" box="[569,665,295,321]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="181" phylum="Arthropoda" rank="species" species="flexibilis">
<emphasis id="B95E221BFFD7FFB0FD91FED6DF850805" box="[569,665,295,321]" italics="true" pageId="5" pageNumber="181">flexibilis</emphasis>
</taxonomicName>
refers to flexible setae on the notogaster.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,352 @@
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<paragraph id="8B95FE09FFD5FFB2FF3FF8BCDE4C0E23" blockId="7.[151,848,1869,1932]" box="[151,848,1869,1896]" pageId="7" pageNumber="183">
<heading id="D0DD4965FFD5FFB2FF3FF8BCDE4C0E23" bold="true" box="[151,848,1869,1896]" fontSize="11" level="1" pageId="7" pageNumber="183" reason="1">
<emphasis id="B95E221BFFD5FFB2FF3FF8BCDE4C0E23" bold="true" box="[151,848,1869,1896]" pageId="7" pageNumber="183">
<taxonomicName id="4C2A858AFFD5FFB2FF3FF8BCDFF70E23" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[151,747,1869,1896]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="triangulus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD5FFB2FF3FF8BCDC260E2C" bold="true" box="[151,314,1869,1896]" italics="true" pageId="7" pageNumber="183">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD5FFB2FEE3F8BCDCF20E2C" bold="true" box="[331,494,1869,1896]" italics="true" pageId="7" pageNumber="183">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD5FFB2FE57F8BCDF610E2C" bold="true" box="[511,637,1869,1896]" italics="true" pageId="7" pageNumber="183">triangulus</emphasis>
Ermilov
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD5FFB2FD5AF8BCDE4C0E23" box="[754,848,1869,1895]" pageId="7" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C330AD82FFD5FFB2FF3FF880DC170ECF" box="[151,267,1905,1932]" pageId="7" pageNumber="183" type="description">
<paragraph id="8B95FE09FFD5FFB2FF3FF880DC170ECF" blockId="7.[151,848,1869,1932]" box="[151,267,1905,1932]" pageId="7" pageNumber="183">
(
<figureCitation id="1311E28CFFD5FFB2FF37F880DDF40EC8" box="[159,232,1905,1932]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="7" pageNumber="183">Figs 5</figureCitation>
,
<figureCitation id="1311E28CFFD5FFB2FF5CF880DC1F0ECF" box="[244,259,1905,1931]" captionStart="FIGURE 6" captionStartId="10.[151,250,1576,1600]" captionTargetBox="[164,1423,194,1539]" captionTargetId="figure-19@10.[151,1436,181,1552]" captionTargetPageId="10" captionText="FIGURE 6. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219160" httpUri="https://zenodo.org/record/13219160/files/figure.png" pageId="7" pageNumber="183">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD5FFBDFF3FF848D91A0821" lastPageId="8" lastPageNumber="184" pageId="7" pageNumber="183" type="materials_examined">
<paragraph id="8B95FE09FFD5FFBDFF3FF848DF3009BD" blockId="7.[151,1436,1977,2042]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="184" pageId="7" pageNumber="183">
<emphasis id="B95E221BFFD5FFB2FF3FF848DC140E97" bold="true" box="[151,264,1977,2003]" pageId="7" pageNumber="183">Material.</emphasis>
<typeStatus id="549140ABFFD5FFB2FEBBF848DC600E97" box="[275,380,1977,2003]" pageId="7" pageNumber="183" type="holotype">Holotype</typeStatus>
(male) and
<specimenCount id="9D2C3580FFD5FFB2FDBBF848DFA70E90" box="[531,699,1977,2004]" count="4" pageId="7" pageNumber="183" type="generic" typeStatus="paratype">four paratypes</specimenCount>
(
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and
<specimenCount id="9D2C3580FFD5FFB2FCD3F848D9020E97" box="[891,1054,1977,2003]" count="3" pageId="7" pageNumber="183" type="female">three females</specimenCount>
): Southeastern
<collectingCountry id="F33DBE99FFD5FFB2FB7BF848D82A0E97" box="[1235,1334,1977,2003]" name="Ethiopia" pageId="7" pageNumber="183">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD5FFB2FAEEF848DDF00EB3" country="Ethiopia" name="Oromiya" pageId="7" pageNumber="183">Oromia Region</collectingRegion>
, Bale Zone, Bale Mountains National Park,
<geoCoordinate id="EE1E98CEFFD5FFB2FD5EF82EDE9B0EBE" box="[758,903,2015,2042]" degrees="6" direction="north" minutes="48" orientation="latitude" pageId="7" pageNumber="183" precision="1" seconds="40.7" value="6.8113055">6°4840.7″N</geoCoordinate>
,
<geoCoordinate id="EE1E98CEFFD5FFB2FC3EF82ED92D0EBE" box="[918,1073,2015,2042]" degrees="39" direction="east" minutes="51" orientation="longitude" pageId="7" pageNumber="183" precision="1" seconds="07.7" value="39.85214">39°5107.7″E</geoCoordinate>
,
<quantity id="4CD253ECFFD5FFB2FBE8F82CD9850EBC" box="[1088,1177,2013,2040]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="7" pageNumber="183" unit="m" value="4050.0">4050 m</quantity>
a.s.l., Sanetti Plateau, Afroalpine community, predominance of cushion-shaped shrubs of
<taxonomicName id="4C2A858AFFDAFFBDFC34FF66D9AA09F4" box="[924,1206,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B95E221BFFDAFFBDFC34FF66D9AA09F4" box="[924,1206,151,177]" italics="true" pageId="8" pageNumber="184">Helichrysum citrispinum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A858AFFDAFFBDFB47FF66D88909F5" box="[1263,1429,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="splendidum">
<emphasis id="B95E221BFFDAFFBDFB47FF66D88909F5" box="[1263,1429,151,177]" italics="true" pageId="8" pageNumber="184">H. splendidum</emphasis>
</taxonomicName>
, co-dominance of
<taxonomicName id="4C2A858AFFDAFFBDFECAFF4ADF360991" box="[354,554,187,213]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="richardii">
<emphasis id="B95E221BFFDAFFBDFECAFF4ADF360991" box="[354,554,187,213]" italics="true" pageId="8" pageNumber="184">Festuca richardii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFDAFFBDFD93FF4ADE4F0991" box="[571,851,187,213]" class="Liliopsida" family="Poaceae" genus="Pentaschistis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="pictigluma">
<emphasis id="B95E221BFFDAFFBDFD93FF4ADE4F0991" box="[571,851,187,213]" italics="true" pageId="8" pageNumber="184">Pentaschistis pictigluma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFDAFFBDFCCCFF4AD9490990" box="[868,1109,187,213]" class="Liliopsida" family="Poaceae" genus="Agrostis" kingdom="Plantae" order="Poales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="quinqueseta">
<emphasis id="B95E221BFFDAFFBDFCCCFF4AD9490990" box="[868,1109,187,213]" italics="true" pageId="8" pageNumber="184">Agrostis quinqueseta</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A858AFFDAFFBDFB30FF4AD8890991" box="[1176,1429,187,213]" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="184" phylum="Tracheophyta" rank="species" species="haumannii">
<emphasis id="B95E221BFFDAFFBDFB30FF4AD8890991" box="[1176,1429,187,213]" italics="true" pageId="8" pageNumber="184">Alchemilla haumannii</emphasis>
</taxonomicName>
, litter,
<date id="FF94D8C9FFDAFFBDFF73FF2EDC4009BD" box="[219,348,223,249]" pageId="8" pageNumber="184" value="2014-11-16">16.XI.2014</date>
(E. Kuzmicheva).
</paragraph>
<paragraph id="8B95FE09FFDAFFBDFF6FFEF2D91A0821" blockId="8.[151,1436,151,357]" pageId="8" pageNumber="184">
The
<typeStatus id="549140ABFFDAFFBDFF51FEF2DC400859" box="[249,348,259,285]" pageId="8" pageNumber="184" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
<collectingCountry id="F33DBE99FFDAFFBDFA83FEF2D8840859" box="[1323,1432,259,285]" name="Germany" pageId="8" pageNumber="184">Germany</collectingCountry>
;
<materialsCitation id="3B42F454FFDAFFBDFF3FFED6D8770805" box="[151,1387,295,321]" country="Russia" location="University Museum of Zoology" pageId="8" pageNumber="184" specimenCount="4" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D2C3580FFDAFFBDFF3FFED6DC210805" box="[151,317,295,321]" count="4" pageId="8" pageNumber="184" type="generic" typeStatus="paratype">four paratypes</specimenCount>
are in the collection of the
<collectingRegion id="49EE30EBFFDAFFBDFD2CFED6DE390805" box="[644,805,295,321]" country="Russia" name="Tyumen'" pageId="8" pageNumber="184">Tyumen State</collectingRegion>
<location id="8EF5A8D2FFDAFFBDFC86FED6D9BC0805" LSID="urn:lsid:plazi:treatment:03834F1FFFD5FFBCFF3FF8BCD8550E41:8EF5A8D2FFDAFFBDFC86FED6D9BC0805" box="[814,1184,295,321]" country="Russia" name="University Museum of Zoology" pageId="8" pageNumber="184" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="49EE30EBFFDAFFBDFB05FED6D8120805" box="[1197,1294,295,321]" country="Russia" name="Tyumen'" pageId="8" pageNumber="184">Tyumen</collectingRegion>
,
<collectingCountry id="F33DBE99FFDAFFBDFAB4FED6D8770805" box="[1308,1387,295,321]" name="Russia" pageId="8" pageNumber="184">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in a solution of 70% ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
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<paragraph id="8B95FE09FFDAFFBDFF3FF890D9120EDA" blockId="8.[151,1436,1889,1950]" pageId="8" pageNumber="184">
<emphasis id="B95E221BFFDAFFBDFF3FF890DC0F0E3D" bold="true" box="[151,275,1889,1913]" pageId="8" pageNumber="184">FIGURE 5.</emphasis>
<taxonomicName id="4C2A858AFFDAFFBDFEB2F890DE110E3D" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[282,781,1889,1913]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="triangulus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFDAFFBDFEB2F890DCB80E3D" box="[282,420,1889,1913]" italics="true" pageId="8" pageNumber="184">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFDAFFBDFE1BF890DF210E3D" box="[435,573,1889,1913]" italics="true" pageId="8" pageNumber="184">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFDAFFBDFDE4F890DFA80E3D" box="[588,692,1889,1913]" italics="true" pageId="8" pageNumber="184">triangulus</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFDAFFBDFCBCF890DE790E3D" bold="true" box="[788,869,1889,1913]" pageId="8" pageNumber="184">
<taxonomicNameLabel id="A26D9F60FFDAFFBDFCBCF890DE790E3D" box="[788,869,1889,1913]" pageId="8" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm.
</paragraph>
</caption>
<subSubSection id="C330AD82FFDBFFBCFF6FFF67D9D10821" pageId="9" pageNumber="185" type="diagnosis">
<paragraph id="8B95FE09FFDBFFBCFF6FFF67D9D10821" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFF67DC5E09F4" bold="true" box="[199,322,150,176]" pageId="9" pageNumber="185">Diagnosis.</emphasis>
Body large, length: 9001005.
<taxonomicName id="4C2A858AFFDBFFBCFD6DFF66DE5C09F5" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[709,832,151,177]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
comparatively broad. Rostrum slightly protruding, rounded. Prolamella developed partially (only mediodistal part present); translamella represented by two short lines near lamellae. Rostral, lamellar and interlamellar setae long, setiform, barbed;
<emphasis id="B95E221BFFDBFFBCFBE3FF2ED97D09BC" box="[1099,1121,223,248]" italics="true" pageId="9" pageNumber="185">in</emphasis>
˃
<emphasis id="B95E221BFFDBFFBCFBD8FF2ED99809BD" box="[1136,1156,223,249]" italics="true" pageId="9" pageNumber="185">le</emphasis>
˃
<emphasis id="B95E221BFFDBFFBCFB3CFF2ED9B009BC" box="[1172,1196,223,248]" italics="true" pageId="9" pageNumber="185">ro</emphasis>
; bothridial seta long, lanceolate, barbed. All notogastral setae represented by alveoli. Circumpedal carina long. Anal and adanal setae medium-sized, setiform, thin, roughened. Ventrobasal tubercle of leg tibiae I, II absent; femora IIIV triangular ventrodistally; tarsus I with 19 setae (
<emphasis id="B95E221BFFDBFFBCFD95FEBADF480821" box="[573,596,331,357]" italics="true" pageId="9" pageNumber="185">l”</emphasis>
absent;
<emphasis id="B95E221BFFDBFFBCFD19FEBADFDB0821" box="[689,711,331,357]" italics="true" pageId="9" pageNumber="185">v</emphasis>
present); genu II with three setae (
<emphasis id="B95E221BFFDBFFBCFBE6FEBAD9780821" box="[1102,1124,331,357]" italics="true" pageId="9" pageNumber="185">v</emphasis>
present).
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDBFFBCFF6FFE9FD8080F69" pageId="9" pageNumber="185" type="description">
<paragraph id="8B95FE09FFDBFFBCFF6FFE9FD9670895" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFE9FDCA608CD" bold="true" box="[199,442,366,393]" pageId="9" pageNumber="185">Description of adult</emphasis>
.
<emphasis id="B95E221BFFDBFFBCFE63FE9EDF7108CC" box="[459,621,367,392]" italics="true" pageId="9" pageNumber="185">Measurements</emphasis>
. Body length: 945 (
<typeStatus id="549140ABFFDBFFBCFCF0FE9EDEDC08CD" box="[856,960,367,393]" pageId="9" pageNumber="185" type="holotype">holotype</typeStatus>
), 900 (male
<typeStatus id="549140ABFFDBFFBCFBFBFE9ED9A408CD" box="[1107,1208,367,393]" pageId="9" pageNumber="185" type="paratype">paratype</typeStatus>
), 9901005 (female
<typeStatus id="549140ABFFDBFFBCFF3FFE62DC1408E9" box="[151,264,403,429]" pageId="9" pageNumber="185" type="paratype">paratypes</typeStatus>
); body width (level of pteromorph): 750 (
<typeStatus id="549140ABFFDBFFBCFD4DFE62DE5108E9" box="[741,845,403,429]" pageId="9" pageNumber="185" type="holotype">holotype</typeStatus>
), 750 (male
<typeStatus id="549140ABFFDBFFBCFC75FE62D95E08E9" box="[989,1090,403,429]" pageId="9" pageNumber="185" type="paratype">paratype</typeStatus>
), 780825 (female
<typeStatus id="549140ABFFDBFFBCFA88FE62D88D08E9" box="[1312,1425,403,429]" pageId="9" pageNumber="185" type="paratype">paratypes</typeStatus>
); width of ventral plate: 675 (
<typeStatus id="549140ABFFDBFFBCFE67FE46DF2B0895" box="[463,567,439,465]" pageId="9" pageNumber="185" type="holotype">holotype</typeStatus>
), 660 (male
<typeStatus id="549140ABFFDBFFBCFD6AFE46DE3B0895" box="[706,807,439,465]" pageId="9" pageNumber="185" type="paratype">paratype</typeStatus>
), 690720 (female
<typeStatus id="549140ABFFDBFFBCFC57FE46D96C0895" box="[1023,1136,439,465]" pageId="9" pageNumber="185" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFE2ADCF40B79" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFE2ADC5908B0" box="[199,325,475,500]" italics="true" pageId="9" pageNumber="185">Integument</emphasis>
. Body color brown to dark brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens; ×1000); podosomal region and lateral part of prodorsum partially with dense microgranulate cerotegument.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFDB6DE020A51" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFDB6DC580B25" box="[199,324,583,609]" italics="true" pageId="9" pageNumber="185">Prodorsum</emphasis>
(
<figureCitation id="1311E28CFFDBFFBCFEF0FDB6DCFE0B25" box="[344,482,582,609]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="9" pageNumber="185">Figs 5A, C</figureCitation>
). Rostrum slightly protruding, rounded. Lamella about 1/2 length of prodorsum; mediodistal part of prolamella developed only; translamella represented by two short lines near lamellae; sublamella and lateral keel-shaped ridge distinct; sublamellar porose area (1519 × 911) oval. Rostral (112131), lamellar (221232) and interlamellar (273281) setae setiform, barbed; bothridial seta (123135) with lanceolate, barbed head, directed posterolaterad; bothridial stalk slightly longer than head; exobothridial seta (4156) setiform, thin, roughened. Dorsosejugal porose area oval, poorly visible.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFCEEDDED0A8D" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<taxonomicName id="4C2A858AFFDBFFBCFF6FFCEEDC5F0A7C" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,799,824]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis id="B95E221BFFDBFFBCFF6FFCEEDC5F0A7C" box="[199,323,799,824]" italics="true" pageId="9" pageNumber="185">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDBFFBCFEFCFCEEDCCB0A7D" box="[340,471,798,825]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="9" pageNumber="185">Figs 5A, C</figureCitation>
). Anterior notogastral margin nearly straight or slightly convex medially.
<taxonomicName id="4C2A858AFFDBFFBCFA89FCEED8800A7D" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[1313,1436,799,825]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
comparatively broad. Pteromorph large, broadly rounded laterally, its anterior margin straight (perpendicular to longitudinal axis of body in dorsal aspect). Ten pairs of notogastral setae represented by alveoli. Four pairs of saccules with small opening and drop-like channel. Opisthonotal gland opening and all lyrifissures (except
<emphasis id="B95E221BFFDBFFBCFAD5FC7AD88F0AE0" box="[1405,1427,907,932]" italics="true" pageId="9" pageNumber="185">ia</emphasis>
) distinct.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFC22DF650D1D" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<taxonomicName id="4C2A858AFFDBFFBCFF6FFC22DC4A0AA9" authorityName="Krefft" authorityYear="1871" box="[199,342,979,1005]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="185" phylum="Nematoda" rank="genus">
<emphasis id="B95E221BFFDBFFBCFF6FFC22DC4A0AA9" box="[199,342,979,1005]" italics="true" pageId="9" pageNumber="185">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDBFFBCFEC6FC22DCD30AA9" box="[366,463,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="9" pageNumber="185">Fig. 5B</figureCitation>
). Generally, similar to
<taxonomicName id="4C2A858AFFDBFFBCFD5AFC22DEB20AA9" authorityName="Ermilov" authorityYear="2024" box="[754,942,979,1005]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="flexibilis" subGenus="Scheloribates">
<emphasis id="B95E221BFFDBFFBCFD5AFC22DE1B0AA9" box="[754,775,979,1005]" italics="true" pageId="9" pageNumber="185">S.</emphasis>
(
<emphasis id="B95E221BFFDBFFBCFC88FC22DE290AA9" box="[800,821,979,1005]" italics="true" pageId="9" pageNumber="185">S.</emphasis>
)
<emphasis id="B95E221BFFDBFFBCFCE6FC22DEB20AA9" box="[846,942,979,1005]" italics="true" pageId="9" pageNumber="185">flexibilis</emphasis>
</taxonomicName>
. Subcapitulum size: 176184 × 127135; subcapitular setae (
<emphasis id="B95E221BFFDBFFBCFEDEFC06DC980D54" box="[374,388,1015,1040]" italics="true" pageId="9" pageNumber="185">a</emphasis>
,
<emphasis id="B95E221BFFDBFFBCFE3DFC06DCB50D54" box="[405,425,1015,1040]" italics="true" pageId="9" pageNumber="185">m</emphasis>
: 3741;
<emphasis id="B95E221BFFDBFFBCFDBBFC06DF3D0D55" box="[531,545,1015,1041]" italics="true" pageId="9" pageNumber="185">h</emphasis>
: 7175) setiform, roughened;
<emphasis id="B95E221BFFDBFFBCFC2BFC06DE8B0D54" box="[899,919,1015,1040]" italics="true" pageId="9" pageNumber="185">m</emphasis>
thinner than
<emphasis id="B95E221BFFDBFFBCFB9DFC06D95F0D54" box="[1077,1091,1015,1040]" italics="true" pageId="9" pageNumber="185">a</emphasis>
and
<emphasis id="B95E221BFFDBFFBCFB28FC06D9920D55" box="[1152,1166,1015,1041]" italics="true" pageId="9" pageNumber="185">h</emphasis>
; both adoral setae (19) setiform, barbed. Palp length: 112116; postpalpal seta (9) spiniform, roughened. Chelicera length: 187206; setae (
<emphasis id="B95E221BFFDBFFBCFF09FBCEDDD50D1D" box="[161,201,1087,1113]" italics="true" pageId="9" pageNumber="185">cha</emphasis>
: 6467;
<emphasis id="B95E221BFFDBFFBCFE84FBCEDC480D1D" box="[300,340,1087,1113]" italics="true" pageId="9" pageNumber="185">chb</emphasis>
: 4549) setiform, barbed.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFB92DC270DAD" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFB92DF960D38" box="[199,650,1123,1149]" italics="true" pageId="9" pageNumber="185">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="1311E28CFFDBFFBCFD30FB92DE0F0D39" box="[664,787,1122,1149]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="9" pageNumber="185">Figs 5B, C</figureCitation>
). Epimeral formula: 3133; all setae (
<emphasis id="B95E221BFFDBFFBCFB7BFB92D9F30D39" box="[1235,1263,1123,1149]" italics="true" pageId="9" pageNumber="185">1a</emphasis>
,
<emphasis id="B95E221BFFDBFFBCFB54FB92D80B0D39" box="[1276,1303,1123,1149]" italics="true" pageId="9" pageNumber="185">2a</emphasis>
,
<emphasis id="B95E221BFFDBFFBCFA8DFB92D85D0D39" box="[1317,1345,1123,1149]" italics="true" pageId="9" pageNumber="185">3a</emphasis>
: 2637;
<emphasis id="B95E221BFFDBFFBCFF3FFB76DDA80DE5" box="[151,180,1159,1185]" italics="true" pageId="9" pageNumber="185">1b</emphasis>
,
<emphasis id="B95E221BFFDBFFBCFF6DFB76DDFD0DE5" box="[197,225,1159,1185]" italics="true" pageId="9" pageNumber="185">3b</emphasis>
,
<emphasis id="B95E221BFFDBFFBCFF5AFB76DC100DE5" box="[242,268,1159,1185]" italics="true" pageId="9" pageNumber="185">3c</emphasis>
: 5671; others: 3445) setiform, thin, roughened. Humeral porose areas
<emphasis id="B95E221BFFDBFFBCFBC9FB76D99A0DE5" box="[1121,1158,1159,1185]" italics="true" pageId="9" pageNumber="185">Am</emphasis>
and
<emphasis id="B95E221BFFDBFFBCFB6CFB76D9FF0DE5" box="[1220,1251,1159,1185]" italics="true" pageId="9" pageNumber="185">Ah</emphasis>
not observable. Pedotectum II rounded laterally in ventral aspect. Discidium broadly rounded. Circumpedal carina long, directed to pedotectum II.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFB02DC220C11" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFB02DC8D0C48" box="[199,401,1267,1293]" italics="true" pageId="9" pageNumber="185">Anogenital region</emphasis>
(
<figureCitation id="1311E28CFFDBFFBCFE08FB02DF010C49" box="[416,541,1266,1293]" captionStart="FIGURE 5" captionStartId="8.[151,250,1889,1913]" captionTargetBox="[164,1423,394,1853]" captionTargetId="figure-106@8.[151,1436,381,1866]" captionTargetPageId="8" captionText="FIGURE 5. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219156" httpUri="https://zenodo.org/record/13219156/files/figure.png" pageId="9" pageNumber="185">Figs 5B, C</figureCitation>
). Genital (
<emphasis id="B95E221BFFDBFFBCFD3CFB02DFBE0C48" box="[660,674,1267,1292]" italics="true" pageId="9" pageNumber="185">g</emphasis>
<subScript id="17AEFC4CFFDBFFBCFD0AFAF4DFB60C57" attach="left" box="[674,682,1285,1299]" fontSize="6" pageId="9" pageNumber="185">1</subScript>
: 4145;
<emphasis id="B95E221BFFDBFFBCFCA5FB02DE070C48" box="[781,795,1267,1292]" italics="true" pageId="9" pageNumber="185">g</emphasis>
<subScript id="17AEFC4CFFDBFFBCFCB3FAF4DE3F0C57" attach="left" box="[795,803,1285,1299]" fontSize="6" pageId="9" pageNumber="185">2</subScript>
<emphasis id="B95E221BFFDBFFBCFC99FB02DE230C48" box="[817,831,1267,1292]" italics="true" pageId="9" pageNumber="185">g</emphasis>
<subScript id="17AEFC4CFFDBFFBCFC97FAF4DE5B0C57" attach="left" box="[831,839,1285,1299]" fontSize="6" pageId="9" pageNumber="185">4</subScript>
: 3437), aggenital (2637), anal (4149), and adanal (6782) setae setiform, thin, roughened.Adanal lyrifissure distinct, close and parallel to anal plate. Marginal porose area band-like.
</paragraph>
<paragraph id="8B95FE09FFDBFFBCFF6FFAAED8080F69" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FFAAEDDE00C3C" box="[199,252,1375,1400]" italics="true" pageId="9" pageNumber="185">Legs</emphasis>
(
<figureCitation id="1311E28CFFDBFFBCFEBAFAAEDC870C3D" box="[274,411,1375,1401]" captionStart="FIGURE 6" captionStartId="10.[151,250,1576,1600]" captionTargetBox="[164,1423,194,1539]" captionTargetId="figure-19@10.[151,1436,181,1552]" captionTargetPageId="10" captionText="FIGURE 6. Scheloribates (Scheloribates) triangulus Ermilov sp. nov., adult: A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219160" httpUri="https://zenodo.org/record/13219160/files/figure.png" pageId="9" pageNumber="185">Figs 6AD</figureCitation>
). Median claw thick; lateral claws thin, with small tubercle distoventrally; all claws slightly barbed on dorsal side. Ventrobasal tubercle of tibiae I, II absent. Femora IIIV triangular ventrodistally. Proximoventral porose area on tarsi IIV, distoventral porose area on tibiae IIV, dorsoparaxial porose area on femora IIV and on trochanters III, IV distinct. Formulas of leg setation and solenidia: I (153419) [122], II (153415) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="C6A8CBB2FFDBFFBCFE89F9E2DC690F69" box="[289,373,1555,1581]" captionStart="TABLE 1" captionStartId="4.[151,239,1592,1616]" captionText="TABLE 1. Leg setation and solenidia of adult Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., S. (S.) monosetosus Ermilov sp. nov., S. (S.) triangulus Ermilov sp. nov., and S. (Topobates) tredecim Ermilov sp. nov." pageId="9" pageNumber="185">Table 1</tableCitation>
. Seta
<emphasis id="B95E221BFFDBFFBCFE11F9E2DCD80F68" box="[441,452,1555,1580]" italics="true" pageId="9" pageNumber="185">s</emphasis>
on tarsus I setiform (not eupathidial), barbed, located between
<emphasis id="B95E221BFFDBFFBCFB21F9E2D9B60F69" box="[1161,1194,1555,1581]" italics="true" pageId="9" pageNumber="185">(a)</emphasis>
and
<emphasis id="B95E221BFFDBFFBCFB48F9E2D8110F69" box="[1248,1293,1555,1581]" italics="true" pageId="9" pageNumber="185">(pv)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDBFFBCFF6FF9C6DE2D0FA5" pageId="9" pageNumber="185" type="discussion">
<paragraph id="8B95FE09FFDBFFBCFF6FF9C6DE2D0FA5" blockId="9.[151,1437,150,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FF9C6DC270F15" bold="true" box="[199,315,1591,1617]" pageId="9" pageNumber="185">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C2A858AFFDBFFBCFD25F9C6DE7A0F14" authorityName="Ermilov" authorityYear="2024" box="[653,870,1591,1617]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="monosetosus" subGenus="Scheloribates">
<emphasis id="B95E221BFFDBFFBCFD25F9C6DFBE0F15" box="[653,674,1591,1617]" italics="true" pageId="9" pageNumber="185">S.</emphasis>
(
<emphasis id="B95E221BFFDBFFBCFD1AF9C6DFDB0F15" box="[690,711,1591,1617]" italics="true" pageId="9" pageNumber="185">S.</emphasis>
)
<emphasis id="B95E221BFFDBFFBCFD7FF9C6DE7A0F14" box="[727,870,1591,1616]" italics="true" pageId="9" pageNumber="185">monosetosus</emphasis>
</taxonomicName>
from
<collectingCountry id="F33DBE99FFDBFFBCFC03F9C6D9160F15" box="[939,1034,1591,1617]" name="Ethiopia" pageId="9" pageNumber="185">Ethiopia</collectingCountry>
(see above) in the presence of large body size, lanceolate bothridial seta and medium-sized anal and adanal setae, but differs from the latter by having triangular ventrodistal part of the leg femora IIIV (versus rounded), the presence (versus of absence) of seta
<emphasis id="B95E221BFFDBFFBCFACEF98ED8650FDD" box="[1382,1401,1663,1689]" italics="true" pageId="9" pageNumber="185">v</emphasis>
on the leg genu II, the length of the prolamella (only mediodistal part developed versus complete), and the presence of the notogastral seta
<emphasis id="B95E221BFFDBFFBCFEDEF936DC980FA4" box="[374,388,1735,1760]" italics="true" pageId="9" pageNumber="185">p</emphasis>
<subScript id="17AEFC4CFFDBFFBCFE2CF928DC900FA3" attach="left" box="[388,396,1753,1767]" fontSize="6" pageId="9" pageNumber="185">1</subScript>
as alveolus (versus needleform seta).
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDBFFBCFF6FF91AD8550E41" box="[199,1353,1771,1797]" pageId="9" pageNumber="185" type="etymology">
<paragraph id="8B95FE09FFDBFFBCFF6FF91AD8550E41" blockId="9.[151,1437,150,1797]" box="[199,1353,1771,1797]" pageId="9" pageNumber="185">
<emphasis id="B95E221BFFDBFFBCFF6FF91ADC520E41" bold="true" box="[199,334,1771,1797]" pageId="9" pageNumber="185">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2A858AFFDBFFBCFD91F91ADFB00E41" authorityName="Ermilov" authorityYear="2024" box="[569,684,1771,1797]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B95E221BFFDBFFBCFD91F91ADFB00E41" box="[569,684,1771,1797]" italics="true" pageId="9" pageNumber="185">triangulus</emphasis>
</taxonomicName>
refers to triangular ventrodistal process on the leg femur II.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,372 @@
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<mods:title id="0992027B2549F260AC5CEF12C5972DBF">New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia</mods:title>
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<mods:namePart id="575F10BC36175FBCE3A33648A8078DC4">Ermilov, Sergey G.</mods:namePart>
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<mods:affiliation id="4C8A39023DF98BA4132D99CF5CD48748">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Soil Zoology and General Entomology, Moscow, Russia</mods:affiliation>
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<mods:affiliation id="A947309F60B4518A93B1767A2A94E625">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Laboratory of Historical Ecology, Moscow, Russia</mods:affiliation>
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<treatment id="03834F1FFFD7FFB2FF3FFE63D83A0FB8" ID-DOI="http://doi.org/10.5281/zenodo.13219152" ID-Zenodo-Dep="13219152" LSID="urn:lsid:plazi:treatment:03834F1FFFD7FFB2FF3FFE63D83A0FB8" httpUri="http://treatment.plazi.org/id/03834F1FFFD7FFB2FF3FFE63D83A0FB8" lastPageId="7" lastPageNumber="183" pageId="5" pageNumber="181">
<subSubSection id="C330AD82FFD7FFB0FF3FFE63DE7108E9" box="[151,877,402,429]" pageId="5" pageNumber="181" type="nomenclature">
<paragraph id="8B95FE09FFD7FFB0FF3FFE63DE7108E9" blockId="5.[151,877,402,465]" box="[151,877,402,429]" pageId="5" pageNumber="181">
<heading id="D0DD4965FFD7FFB0FF3FFE63DE7108E9" bold="true" box="[151,877,402,429]" fontSize="11" level="1" pageId="5" pageNumber="181" reason="1">
<emphasis id="B95E221BFFD7FFB0FF3FFE63DE7108E9" bold="true" box="[151,877,402,429]" pageId="5" pageNumber="181">
<taxonomicName id="4C2A858AFFD7FFB0FF3FFE63DE1508E8" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[151,777,402,429]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="181" phylum="Arthropoda" rank="species" species="monosetosus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD7FFB0FF3FFE63DC2608E9" bold="true" box="[151,314,402,429]" italics="true" pageId="5" pageNumber="181">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD7FFB0FEE3FE63DCF208E9" bold="true" box="[331,494,402,429]" italics="true" pageId="5" pageNumber="181">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD7FFB0FE57FE62DF8608E9" bold="true" box="[511,666,403,429]" italics="true" pageId="5" pageNumber="181">monosetosus</emphasis>
Ermilov
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD7FFB0FCA7FE62DE7108E9" box="[783,877,403,429]" pageId="5" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C330AD82FFD7FFB0FF3FFE46DC170895" box="[151,267,439,465]" pageId="5" pageNumber="181" type="description">
<paragraph id="8B95FE09FFD7FFB0FF3FFE46DC170895" blockId="5.[151,877,402,465]" box="[151,267,439,465]" pageId="5" pageNumber="181">
(
<figureCitation id="1311E28CFFD7FFB0FF37FE46DDFB0895" box="[159,231,439,465]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Figs 3</figureCitation>
,
<figureCitation id="1311E28CFFD7FFB0FF5CFE46DC1F0895" box="[244,259,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1475,1499]" captionTargetBox="[164,1423,194,1438]" captionTargetId="figure-28@7.[151,1436,181,1451]" captionTargetPageId="7" captionText="FIGURE 4. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult:A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219154" httpUri="https://zenodo.org/record/13219154/files/figure.png" pageId="5" pageNumber="181">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD7FFB0FF3FFE0FDEE00A51" pageId="5" pageNumber="181" type="materials_examined">
<paragraph id="8B95FE09FFD7FFB0FF3FFE0FDCF90BED" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF3FFE0FDC1A0B5C" bold="true" box="[151,262,510,536]" pageId="5" pageNumber="181">Material.</emphasis>
<typeStatus id="549140ABFFD7FFB0FEB9FE0EDC650B5D" box="[273,377,511,537]" pageId="5" pageNumber="181" type="holotype">Holotype</typeStatus>
(female) and
<specimenCount id="9D2C3580FFD7FFB0FDB7FE0EDFDC0B5D" box="[543,704,511,537]" count="5" pageId="5" pageNumber="181" type="generic" typeStatus="paratype">five paratypes</specimenCount>
(
<specimenCount id="9D2C3580FFD7FFB0FD7AFE0EDE550B5D" box="[722,841,511,537]" count="2" pageId="5" pageNumber="181" type="male">two males</specimenCount>
and
<specimenCount id="9D2C3580FFD7FFB0FC2FFE0ED93A0B5D" box="[903,1062,511,537]" count="3" pageId="5" pageNumber="181" type="female">three females</specimenCount>
): Southeastern
<collectingCountry id="F33DBE99FFD7FFB0FB7EFE0ED8240B5D" box="[1238,1336,511,537]" name="Ethiopia" pageId="5" pageNumber="181">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD7FFB0FAEFFE0EDDF70B79" country="Ethiopia" name="Oromiya" pageId="5" pageNumber="181">Oromia Region</collectingRegion>
, Bale Zone, Bale Mountains National Park,
<geoCoordinate id="EE1E98CEFFD7FFB0FD5FFDD5DE9A0B7B" box="[759,902,548,575]" degrees="6" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="181" precision="1" seconds="40.7" value="6.8113055">6°4840.7″N</geoCoordinate>
,
<geoCoordinate id="EE1E98CEFFD7FFB0FC3EFDD5D9330B7B" box="[918,1071,548,575]" degrees="39" direction="east" minutes="51" orientation="longitude" pageId="5" pageNumber="181" precision="1" seconds="07.7" value="39.85214">39°5107.7″E</geoCoordinate>
,
<quantity id="4CD253ECFFD7FFB0FB97FDD3D9840B79" box="[1087,1176,546,573]" metricMagnitude="3" metricUnit="m" metricValue="4.05" pageId="5" pageNumber="181" unit="m" value="4050.0">4050 m</quantity>
a.s.l., Sanetti Plateau, Afroalpine community, predominance of cushion-shaped shrubs of
<taxonomicName id="4C2A858AFFD7FFB0FC2AFDB6D98A0B24" box="[898,1174,583,609]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="citrispinum">
<emphasis id="B95E221BFFD7FFB0FC2AFDB6D98A0B24" box="[898,1174,583,609]" italics="true" pageId="5" pageNumber="181">Helichrysum citrispinum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A858AFFD7FFB0FB62FDB6D8700B25" box="[1226,1388,583,609]" class="Magnoliopsida" family="Asteraceae" genus="Helichrysum" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="splendidum">
<emphasis id="B95E221BFFD7FFB0FB62FDB6D8700B25" box="[1226,1388,583,609]" italics="true" pageId="5" pageNumber="181">H. splendidum</emphasis>
</taxonomicName>
, co-dominance of
<taxonomicName id="4C2A858AFFD7FFB0FE91FD9ADCE10BC1" box="[313,509,619,645]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="richardii">
<emphasis id="B95E221BFFD7FFB0FE91FD9ADCE10BC1" box="[313,509,619,645]" italics="true" pageId="5" pageNumber="181">Festuca richardii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFD7FFB0FDA4FD9ADE3C0BC1" box="[524,800,619,645]" class="Liliopsida" family="Poaceae" genus="Pentaschistis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="pictigluma">
<emphasis id="B95E221BFFD7FFB0FDA4FD9ADE3C0BC1" box="[524,800,619,645]" italics="true" pageId="5" pageNumber="181">Pentaschistis pictigluma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2A858AFFD7FFB0FC86FD9AD9050BC0" box="[814,1049,619,645]" class="Liliopsida" family="Poaceae" genus="Agrostis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="quinqueseta">
<emphasis id="B95E221BFFD7FFB0FC86FD9AD9050BC0" box="[814,1049,619,645]" italics="true" pageId="5" pageNumber="181">Agrostis quinqueseta</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2A858AFFD7FFB0FBF1FD9AD84F0BC1" box="[1113,1363,619,645]" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" pageId="5" pageNumber="181" phylum="Tracheophyta" rank="species" species="haumannii">
<emphasis id="B95E221BFFD7FFB0FBF1FD9AD84F0BC1" box="[1113,1363,619,645]" italics="true" pageId="5" pageNumber="181">Alchemilla haumannii</emphasis>
</taxonomicName>
, litter,
<date id="FF94D8C9FFD7FFB0FF3FFD7EDC0A0BED" box="[151,278,655,681]" pageId="5" pageNumber="181" value="2014-11-16">16.XI.2014</date>
(E. Kuzmicheva).
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FFD42DEE00A51" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
The
<typeStatus id="549140ABFFD7FFB0FF52FD42DC400B89" box="[250,348,691,717]" pageId="5" pageNumber="181" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
<collectingCountry id="F33DBE99FFD7FFB0FA84FD42D8840B89" box="[1324,1432,691,717]" name="Germany" pageId="5" pageNumber="181">Germany</collectingCountry>
;
<materialsCitation id="3B42F454FFD7FFB0FF3FFD26D8770BB5" box="[151,1387,727,753]" country="Russia" location="University Museum of Zoology" pageId="5" pageNumber="181" specimenCount="5" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D2C3580FFD7FFB0FF3FFD26DC250BB5" box="[151,313,727,753]" count="5" pageId="5" pageNumber="181" type="generic" typeStatus="paratype">five paratypes</specimenCount>
are in the collection of the
<collectingRegion id="49EE30EBFFD7FFB0FD20FD26DE3A0BB5" box="[648,806,727,753]" country="Russia" name="Tyumen'" pageId="5" pageNumber="181">Tyumen State</collectingRegion>
<location id="8EF5A8D2FFD7FFB0FC99FD26D9BD0BB5" LSID="urn:lsid:plazi:treatment:03834F1FFFD7FFB2FF3FFE63D83A0FB8:8EF5A8D2FFD7FFB0FC99FD26D9BD0BB5" box="[817,1185,727,753]" country="Russia" name="University Museum of Zoology" pageId="5" pageNumber="181" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="49EE30EBFFD7FFB0FB18FD26D8120BB5" box="[1200,1294,727,753]" country="Russia" name="Tyumen'" pageId="5" pageNumber="181">Tyumen</collectingRegion>
,
<collectingCountry id="F33DBE99FFD7FFB0FAB5FD26D8770BB5" box="[1309,1387,727,753]" name="Russia" pageId="5" pageNumber="181">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in a solution of 70% ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD7FFB0FF6FFCEFDE440AA9" pageId="5" pageNumber="181" type="diagnosis">
<paragraph id="8B95FE09FFD7FFB0FF6FFCEFDE440AA9" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFCEFDC5E0A7C" bold="true" box="[199,322,798,824]" pageId="5" pageNumber="181">Diagnosis.</emphasis>
Body large, length: 885960.
<taxonomicName id="4C2A858AFFD7FFB0FD38FCEEDE170A7D" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[656,779,799,825]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
broad. Rostrum protruding, rounded. Prolamella complete; translamella represented by two short lines near lamellae. Rostral, lamellar and interlamellar setae long, setiform, barbed;
<emphasis id="B95E221BFFD7FFB0FF5EFC96DC100AC4" box="[246,268,871,896]" italics="true" pageId="5" pageNumber="181">in</emphasis>
˃
<emphasis id="B95E221BFFD7FFB0FEB3FC96DC330AC5" box="[283,303,871,897]" italics="true" pageId="5" pageNumber="181">le</emphasis>
˃
<emphasis id="B95E221BFFD7FFB0FE97FC96DC4B0AC4" box="[319,343,871,896]" italics="true" pageId="5" pageNumber="181">ro</emphasis>
; bothridial seta long, lanceolate, barbed. One pair of notogastral setae (
<emphasis id="B95E221BFFD7FFB0FB09FC96D9B30AC4" box="[1185,1199,871,896]" italics="true" pageId="5" pageNumber="181">p</emphasis>
<subScript id="17AEFC4CFFD7FFB0FB07FC88D9AB0AC3" attach="left" box="[1199,1207,889,903]" fontSize="6" pageId="5" pageNumber="181">1</subScript>
) short, needleform; other setae represented by alveoli. Circumpedal carina long. Anal and adanal setae medium-sized, setiform, thin, roughened. Ventrobasal tubercle of leg tibiae I, II absent; femur II broadly rounded ventrodistally; tarsus I with 19 setae (
<emphasis id="B95E221BFFD7FFB0FF77FC22DDEA0AA9" box="[223,246,979,1005]" italics="true" pageId="5" pageNumber="181">l”</emphasis>
absent;
<emphasis id="B95E221BFFD7FFB0FEFCFC22DC760AA9" box="[340,362,979,1005]" italics="true" pageId="5" pageNumber="181">v</emphasis>
present); genu II with two setae (
<emphasis id="B95E221BFFD7FFB0FD4BFC22DFE50AA9" box="[739,761,979,1005]" italics="true" pageId="5" pageNumber="181">v</emphasis>
absent).
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD7FFB3FF6FFC07D8080EEC" lastPageId="6" lastPageNumber="182" pageId="5" pageNumber="181" type="description">
<paragraph id="8B95FE09FFD7FFB0FF6FFC07DFFF0D1D" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFC07DCA90D55" bold="true" box="[199,437,1014,1041]" pageId="5" pageNumber="181">Description of adult</emphasis>
.
<emphasis id="B95E221BFFD7FFB0FE6CFC06DF7A0D54" box="[452,614,1015,1040]" italics="true" pageId="5" pageNumber="181">Measurements</emphasis>
. Body length: 885 (
<typeStatus id="549140ABFFD7FFB0FCEEFC06DEB20D55" box="[838,942,1015,1041]" pageId="5" pageNumber="181" type="holotype">holotype</typeStatus>
), 885960 (
<typeStatus id="549140ABFFD7FFB0FB9BFC06D9B90D55" box="[1075,1189,1015,1041]" pageId="5" pageNumber="181" type="paratype">paratypes</typeStatus>
); body width (level of pteromorph): 780 (
<typeStatus id="549140ABFFD7FFB0FEC1FBEADCCD0D71" box="[361,465,1051,1077]" pageId="5" pageNumber="181" type="holotype">holotype</typeStatus>
), 705810 (
<typeStatus id="549140ABFFD7FFB0FDFCFBEADFDA0D71" box="[596,710,1051,1077]" pageId="5" pageNumber="181" type="paratype">paratypes</typeStatus>
); width of ventral plate: 690 (
<typeStatus id="549140ABFFD7FFB0FBA4FBEAD9680D71" box="[1036,1140,1051,1077]" pageId="5" pageNumber="181" type="holotype">holotype</typeStatus>
), 615690 (
<typeStatus id="549140ABFFD7FFB0FB5FFBEAD8750D71" box="[1271,1385,1051,1077]" pageId="5" pageNumber="181" type="paratype">paratypes</typeStatus>
). No differences between males and females in body size.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FFB92DCF40D81" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFB92DC590D38" box="[199,325,1123,1148]" italics="true" pageId="5" pageNumber="181">Integument</emphasis>
. Body color brown to dark brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens; ×1000); podosomal region and lateral part of prodorsum partially with dense microgranulate cerotegument.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FFB3EDF860CD9" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FFB3EDC580DAD" box="[199,324,1231,1257]" italics="true" pageId="5" pageNumber="181">Prodorsum</emphasis>
(
<figureCitation id="1311E28CFFD7FFB0FEFEFB3EDCC60DAD" box="[342,474,1231,1257]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Figs 3A, C</figureCitation>
). Rostrum protruding, rounded. Lamella about 1/2 length of prodorsum; prolamella complete (reaching insertion of
<emphasis id="B95E221BFFD7FFB0FE50FB02DF0C0C48" box="[504,528,1267,1292]" italics="true" pageId="5" pageNumber="181">ro</emphasis>
); translamella represented by two short lines near lamellae; sublamella and lateral keel-shaped ridge distinct; sublamellar porose area (1315) rounded. Rostral (131142), lamellar (210214) and interlamellar (281307) setae setiform, barbed; bothridial seta (120124) with lanceolate, barbed head, directed posterolaterad; bothridial stalk slightly longer than head; exobothridial seta (2230) setiform, thin, roughened. Dorsosejugal porose area oval, poorly visible.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FFA56DC6D0F15" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<taxonomicName id="4C2A858AFFD7FFB0FF6FFA56DC5F0C84" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,1447,1472]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B95E221BFFD7FFB0FF6FFA56DC5F0C84" box="[199,323,1447,1472]" italics="true" pageId="5" pageNumber="181">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFD7FFB0FEFDFA56DCC40C85" box="[341,472,1447,1473]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Figs 3A, C</figureCitation>
). Anterior notogastral margin convex medially.
<taxonomicName id="4C2A858AFFD7FFB0FC54FA56D96B0C85" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[1020,1143,1447,1473]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
broad. Pteromorph large, broadly rounded laterally, its anterior margin straight (perpendicular to longitudinal axis of body in dorsal aspect) or slightly convex. Nine pairs of notogastral setae represented by alveoli and one pair (
<emphasis id="B95E221BFFD7FFB0FB9BFA1ED95D0F4C" box="[1075,1089,1519,1544]" italics="true" pageId="5" pageNumber="181">p</emphasis>
<subScript id="17AEFC4CFFD7FFB0FBE9F9F0D9550F4B" attach="left" box="[1089,1097,1537,1551]" fontSize="6" pageId="5" pageNumber="181">1</subScript>
) by needleform (3437) setae. Four pairs of saccules with small opening and drop-like channel. Opisthonotal gland opening and all lyrifissures (except
<emphasis id="B95E221BFFD7FFB0FF59F9C6DC1B0F14" box="[241,263,1591,1616]" italics="true" pageId="5" pageNumber="181">ia</emphasis>
) distinct.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FF9AADFA40FA5" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<taxonomicName id="4C2A858AFFD7FFB0FF6FF9AADC4A0F31" authorityName="Krefft" authorityYear="1871" box="[199,342,1627,1653]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="181" phylum="Nematoda" rank="genus">
<emphasis id="B95E221BFFD7FFB0FF6FF9AADC4A0F31" box="[199,342,1627,1653]" italics="true" pageId="5" pageNumber="181">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFD7FFB0FEC6F9AADCD20F31" box="[366,462,1627,1653]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Fig. 3B</figureCitation>
). Generally, similar to
<taxonomicName id="4C2A858AFFD7FFB0FD5AF9AADEB20F31" authorityName="Ermilov" authorityYear="2024" box="[754,942,1627,1653]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="181" phylum="Arthropoda" rank="species" species="flexibilis" subGenus="Scheloribates">
<emphasis id="B95E221BFFD7FFB0FD5AF9AADE1B0F31" box="[754,775,1627,1653]" italics="true" pageId="5" pageNumber="181">S.</emphasis>
(
<emphasis id="B95E221BFFD7FFB0FC88F9AADE290F31" box="[800,821,1627,1653]" italics="true" pageId="5" pageNumber="181">S.</emphasis>
)
<emphasis id="B95E221BFFD7FFB0FCE6F9AADEB20F31" box="[846,942,1627,1653]" italics="true" pageId="5" pageNumber="181">flexibilis</emphasis>
</taxonomicName>
. Subcapitulum size: 169184 × 131139; subcapitular setae (
<emphasis id="B95E221BFFD7FFB0FEDAF98EDC9C0FDC" box="[370,384,1663,1688]" italics="true" pageId="5" pageNumber="181">a</emphasis>
: 3441;
<emphasis id="B95E221BFFD7FFB0FE4EF98EDCE60FDC" box="[486,506,1663,1688]" italics="true" pageId="5" pageNumber="181">m</emphasis>
: 2834;
<emphasis id="B95E221BFFD7FFB0FDC9F98EDF730FDD" box="[609,623,1663,1689]" italics="true" pageId="5" pageNumber="181">h</emphasis>
: 6475) setiform, roughened;
<emphasis id="B95E221BFFD7FFB0FC62F98EDEC20FDC" box="[970,990,1663,1688]" italics="true" pageId="5" pageNumber="181">m</emphasis>
thinner than
<emphasis id="B95E221BFFD7FFB0FBDEF98ED9980FDC" box="[1142,1156,1663,1688]" italics="true" pageId="5" pageNumber="181">a</emphasis>
and
<emphasis id="B95E221BFFD7FFB0FB16F98ED9D00FDD" box="[1214,1228,1663,1689]" italics="true" pageId="5" pageNumber="181">h</emphasis>
; both adoral setae (19) setiform, barbed. Palp length: 112116; postpalpal seta (9) spiniform, roughened. Chelicera length: 184191; setae (
<emphasis id="B95E221BFFD7FFB0FF77F936DC1B0FA5" box="[223,263,1735,1761]" italics="true" pageId="5" pageNumber="181">cha</emphasis>
: 6064;
<emphasis id="B95E221BFFD7FFB0FEC3F936DC8F0FA5" box="[363,403,1735,1761]" italics="true" pageId="5" pageNumber="181">chb</emphasis>
: 4145) setiform, barbed.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FF91ADCA50E35" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FF91ADF960E40" box="[199,650,1771,1797]" italics="true" pageId="5" pageNumber="181">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="1311E28CFFD7FFB0FD30F91ADE0F0E41" box="[664,787,1771,1797]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Figs 3B, C</figureCitation>
). Epimeral formula: 3133; all setae (
<emphasis id="B95E221BFFD7FFB0FB7BF91AD9F30E41" box="[1235,1263,1771,1797]" italics="true" pageId="5" pageNumber="181">1a</emphasis>
,
<emphasis id="B95E221BFFD7FFB0FB54F91AD80B0E41" box="[1276,1303,1771,1797]" italics="true" pageId="5" pageNumber="181">2a</emphasis>
,
<emphasis id="B95E221BFFD7FFB0FA8DF91AD85D0E41" box="[1317,1345,1771,1797]" italics="true" pageId="5" pageNumber="181">3a</emphasis>
: 2634;
<emphasis id="B95E221BFFD7FFB0FF3FF8FEDDA80E6D" box="[151,180,1807,1833]" italics="true" pageId="5" pageNumber="181">1b</emphasis>
,
<emphasis id="B95E221BFFD7FFB0FF6DF8FEDDFD0E6D" box="[197,225,1807,1833]" italics="true" pageId="5" pageNumber="181">3b</emphasis>
,
<emphasis id="B95E221BFFD7FFB0FF5AF8FEDC100E6D" box="[242,268,1807,1833]" italics="true" pageId="5" pageNumber="181">3c</emphasis>
: 6075; others: 3449) setiform, thin, roughened. Humeral porose areas
<emphasis id="B95E221BFFD7FFB0FBC9F8FED99A0E6D" box="[1121,1158,1807,1833]" italics="true" pageId="5" pageNumber="181">Am</emphasis>
and
<emphasis id="B95E221BFFD7FFB0FB6CF8FED9FF0E6D" box="[1220,1251,1807,1833]" italics="true" pageId="5" pageNumber="181">Ah</emphasis>
not observable. Pedotectum II rounded laterally in ventral aspect. Discidium triangular, with rounded top. Circumpedal carina long, directed to pedotectum II.
</paragraph>
<paragraph id="8B95FE09FFD7FFB0FF6FF88AD9A70E99" blockId="5.[151,1437,510,2013]" pageId="5" pageNumber="181">
<emphasis id="B95E221BFFD7FFB0FF6FF88ADC8D0ED0" box="[199,401,1915,1941]" italics="true" pageId="5" pageNumber="181">Anogenital region</emphasis>
(
<figureCitation id="1311E28CFFD7FFB0FE08F88ADF010ED1" box="[416,541,1915,1941]" captionStart="FIGURE 3" captionStartId="6.[151,250,1611,1635]" captionTargetBox="[158,1426,194,1574]" captionTargetId="figure-19@6.[151,1436,181,1587]" captionTargetPageId="6" captionText="FIGURE 3. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="5" pageNumber="181">Figs 3B, C</figureCitation>
). Genital (
<emphasis id="B95E221BFFD7FFB0FD3CF88ADFBE0ED0" box="[660,674,1915,1940]" italics="true" pageId="5" pageNumber="181">g</emphasis>
<subScript id="17AEFC4CFFD7FFB0FD0AF87CDFB60EDF" attach="left" box="[674,682,1933,1947]" fontSize="6" pageId="5" pageNumber="181">1</subScript>
: 3449;
<emphasis id="B95E221BFFD7FFB0FCA5F88ADE070ED0" box="[781,795,1915,1940]" italics="true" pageId="5" pageNumber="181">g</emphasis>
<subScript id="17AEFC4CFFD7FFB0FCB3F87CDE3F0EDF" attach="left" box="[795,803,1933,1947]" fontSize="6" pageId="5" pageNumber="181">2</subScript>
<emphasis id="B95E221BFFD7FFB0FC99F88ADE230ED0" box="[817,831,1915,1940]" italics="true" pageId="5" pageNumber="181">g</emphasis>
<subScript id="17AEFC4CFFD7FFB0FC97F87CDE5B0EDF" attach="left" box="[831,839,1933,1947]" fontSize="6" pageId="5" pageNumber="181">4</subScript>
: 2234), aggenital (2234), anal (4156), and adanal (5664) setae setiform, thin, roughened; both aggenital setae absent in the
<typeStatus id="549140ABFFD7FFB0FC53F86ED97D0EFD" box="[1019,1121,1951,1977]" pageId="5" pageNumber="181" type="holotype">holotype</typeStatus>
. Adanal lyrifissure distinct, close and parallel to anal plate. Marginal porose area band-like, sometimes shortly interrupted.
</paragraph>
<caption id="DF55AE81FFD4FFB3FF3FF9BAD8700FC3" ID-DOI="http://doi.org/10.5281/zenodo.13219150" ID-Zenodo-Dep="13219150" httpUri="https://zenodo.org/record/13219150/files/figure.png" pageId="6" pageNumber="182" startId="6.[151,250,1611,1635]" targetBox="[158,1426,194,1574]" targetPageId="6" targetType="figure">
<paragraph id="8B95FE09FFD4FFB3FF3FF9BAD8700FC3" blockId="6.[151,1436,1611,1671]" pageId="6" pageNumber="182">
<emphasis id="B95E221BFFD4FFB3FF3FF9BADC090F27" bold="true" box="[151,277,1611,1635]" pageId="6" pageNumber="182">FIGURE 3.</emphasis>
<taxonomicName id="4C2A858AFFD4FFB3FEB6F9BADE2E0F27" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[286,818,1611,1635]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="182" phylum="Arthropoda" rank="species" species="monosetosus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD4FFB3FEB6F9BADCB40F27" box="[286,424,1611,1635]" italics="true" pageId="6" pageNumber="182">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD4FFB3FE12F9BADF580F27" box="[442,580,1611,1635]" italics="true" pageId="6" pageNumber="182">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD4FFB3FDFDF9BDDFCA0F27" box="[597,726,1612,1635]" italics="true" pageId="6" pageNumber="182">monosetosus</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD4FFB3FC93F9BADE920F27" bold="true" box="[827,910,1611,1635]" pageId="6" pageNumber="182">
<taxonomicNameLabel id="A26D9F60FFD4FFB3FC93F9BADE920F27" box="[827,910,1611,1635]" pageId="6" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view (legs omitted); B—ventral view (legs and part of right pteromorph omitted); C—right lateral view (gnathosoma and legs omitted). Scale bar 200 μm.
</paragraph>
</caption>
<paragraph id="8B95FE09FFD4FFB3FF6FF92BD8080EEC" blockId="6.[151,1437,1753,1960]" pageId="6" pageNumber="182">
<emphasis id="B95E221BFFD4FFB3FF6FF92BDDE00FB7" box="[199,252,1754,1779]" italics="true" pageId="6" pageNumber="182">Legs</emphasis>
(
<figureCitation id="1311E28CFFD4FFB3FEBAF928DC870FB7" box="[274,411,1753,1779]" captionStart="FIGURE 4" captionStartId="7.[151,250,1475,1499]" captionTargetBox="[164,1423,194,1438]" captionTargetId="figure-28@7.[151,1436,181,1451]" captionTargetPageId="7" captionText="FIGURE 4. Scheloribates (Scheloribates) monosetosus Ermilov sp. nov., adult:A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219154" httpUri="https://zenodo.org/record/13219154/files/figure.png" pageId="6" pageNumber="182">Figs 4AD</figureCitation>
). Median claw thick; lateral claws thin, with small tubercle distoventrally; all claws slightly barbed on dorsal side. Ventrobasal tubercle of tibiae I, II absent. Femur II broadly rounded ventrodistally. Proximoventral porose area on tarsi IIV, distoventral porose area on tibiae IIV, dorsoparaxial porose area on femora IIV and on trochanters III, IV distinct. Formulas of leg setation and solenidia: I (153419) [122], II (152415) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="C6A8CBB2FFD4FFB3FE89F87CDC690EE3" box="[289,373,1933,1959]" captionStart="TABLE 1" captionStartId="4.[151,239,1592,1616]" captionText="TABLE 1. Leg setation and solenidia of adult Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., S. (S.) monosetosus Ermilov sp. nov., S. (S.) triangulus Ermilov sp. nov., and S. (Topobates) tredecim Ermilov sp. nov." pageId="6" pageNumber="182">Table 1</tableCitation>
. Seta
<emphasis id="B95E221BFFD4FFB3FE11F87FDCD80EE3" box="[441,452,1934,1959]" italics="true" pageId="6" pageNumber="182">s</emphasis>
on tarsus I setiform (not eupathidial), barbed, located between
<emphasis id="B95E221BFFD4FFB3FB21F87CD9B60EE3" box="[1161,1194,1933,1959]" italics="true" pageId="6" pageNumber="182">(a)</emphasis>
and
<emphasis id="B95E221BFFD4FFB3FB48F87CD8110EE3" box="[1248,1293,1933,1959]" italics="true" pageId="6" pageNumber="182">(pv)</emphasis>
.
</paragraph>
</subSubSection>
<caption id="DF55AE81FFD5FFB2FF3FFA32D9750CBB" ID-DOI="http://doi.org/10.5281/zenodo.13219154" ID-Zenodo-Dep="13219154" httpUri="https://zenodo.org/record/13219154/files/figure.png" pageId="7" pageNumber="183" startId="7.[151,250,1475,1499]" targetBox="[164,1423,194,1438]" targetPageId="7" targetType="figure">
<paragraph id="8B95FE09FFD5FFB2FF3FFA32D9750CBB" blockId="7.[151,1436,1475,1535]" pageId="7" pageNumber="183">
<emphasis id="B95E221BFFD5FFB2FF3FFA32DC0E0C9F" bold="true" box="[151,274,1475,1499]" pageId="7" pageNumber="183">FIGURE 4.</emphasis>
<taxonomicName id="4C2A858AFFD5FFB2FEB0FA32DE3D0C9F" authority="Ermilov" authorityName="Ermilov" authorityYear="2024" box="[280,801,1475,1499]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="monosetosus" status="sp. nov." subGenus="Scheloribates">
<emphasis id="B95E221BFFD5FFB2FEB0FA32DCBE0C9F" box="[280,418,1475,1499]" italics="true" pageId="7" pageNumber="183">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD5FFB2FE07FA32DF250C9F" box="[431,569,1475,1499]" italics="true" pageId="7" pageNumber="183">Scheloribates</emphasis>
)
<emphasis id="B95E221BFFD5FFB2FDEFFA35DFD40C9F" box="[583,712,1476,1499]" italics="true" pageId="7" pageNumber="183">monosetosus</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD5FFB2FC8FFA32DE6A0C9F" bold="true" box="[807,886,1475,1499]" pageId="7" pageNumber="183">
<taxonomicNameLabel id="A26D9F60FFD5FFB2FC8FFA32DE6A0C9F" box="[807,886,1475,1499]" pageId="7" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—leg I, right, antiaxial view; B—leg II, right, antiaxial view; C—leg III, left, antiaxial view; D—leg IV, left, antiaxial view. Scale bar 100 μm.
</paragraph>
</caption>
<subSubSection id="C330AD82FFD5FFB2FF6FF9A0D9D30F9C" pageId="7" pageNumber="183" type="discussion">
<paragraph id="8B95FE09FFD5FFB2FF6FF9A0D9D30F9C" blockId="7.[151,1436,1617,1788]" pageId="7" pageNumber="183">
<emphasis id="B95E221BFFD5FFB2FF6FF9A0DC270F2F" bold="true" box="[199,315,1617,1643]" pageId="7" pageNumber="183">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C2A858AFFD5FFB2FD33F9A3D9060F2F" authority="Balogh, 1962" authorityName="Balogh" authorityYear="1962" box="[667,1050,1617,1644]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="concentricus" subGenus="Scheloribates">
<emphasis id="B95E221BFFD5FFB2FD33F9A3DFAC0F28" box="[667,688,1618,1644]" italics="true" pageId="7" pageNumber="183">S.</emphasis>
(
<emphasis id="B95E221BFFD5FFB2FD6AF9A3DFCB0F28" box="[706,727,1618,1644]" italics="true" pageId="7" pageNumber="183">S.</emphasis>
)
<emphasis id="B95E221BFFD5FFB2FD41F9A3DE640F2F" box="[745,888,1618,1643]" italics="true" pageId="7" pageNumber="183">concentricus</emphasis>
<bibRefCitation id="EFBB83F8FFD5FFB2FC2AF9A0D9060F2F" author="Balogh, J." box="[898,1050,1617,1643]" pageId="7" pageNumber="183" pagination="90 - 131" refId="ref7413" refString="Balogh, J. (1962) Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXV. Acari Oribates. Annales du Musee Royal de l'Afrique Centrale Tervuren Belgium, Zoologie, 110, 90 - 131." type="journal article" year="1962">Balogh, 1962</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F33DBE99FFD5FFB2FBCAF9A0D9DB0F2F" box="[1122,1223,1617,1643]" name="Tanzania" pageId="7" pageNumber="183">Tanzania</collectingCountry>
in the presence of large body size, one pair of remaining notogastral setae (
<emphasis id="B95E221BFFD5FFB2FCB2F987DE340FCB" box="[794,808,1654,1679]" italics="true" pageId="7" pageNumber="183">p</emphasis>
<subScript id="17AEFC4CFFD5FFB2FC80F979DE2C0FD2" attach="left" box="[808,816,1672,1686]" fontSize="6" pageId="7" pageNumber="183">1</subScript>
), protruding rostrum, and bothridial seta with elongate head, but differs from the latter by the presence of medium-sized (versus short) anal and adanal setae, short (versus medium-sized) translamellar lines near lamellae and lanceolate (versus fusiform) bothridial seta.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFD5FFB2FF6FF910D83A0FB8" box="[199,1318,1761,1788]" pageId="7" pageNumber="183" type="etymology">
<paragraph id="8B95FE09FFD5FFB2FF6FF910D83A0FB8" blockId="7.[151,1436,1617,1788]" box="[199,1318,1761,1788]" pageId="7" pageNumber="183">
<emphasis id="B95E221BFFD5FFB2FF6FF910DC520FBF" bold="true" box="[199,334,1761,1787]" pageId="7" pageNumber="183">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2A858AFFD5FFB2FD91F913DFD40FBF" authorityName="Ermilov" authorityYear="2024" box="[569,712,1762,1787]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="monosetosus">
<emphasis id="B95E221BFFD5FFB2FD91F913DFD40FBF" box="[569,712,1762,1787]" italics="true" pageId="7" pageNumber="183">monosetosus</emphasis>
</taxonomicName>
refers to the presence of one pair of notogastral setae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,366 @@
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<mods:title id="BCB52BA86354F4BAF49495A3E17804EE">New Scheloribates (Acari, Oribatida, Scheloribatidae) from Ethiopia</mods:title>
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<paragraph id="8B95FE09FFD8FFBFFF3FF947DFBF0F95" blockId="10.[151,675,1718,1781]" box="[151,675,1718,1745]" pageId="10" pageNumber="186">
<heading id="D0DD4965FFD8FFBFFF3FF947DFBF0F95" bold="true" box="[151,675,1718,1745]" fontSize="11" level="1" pageId="10" pageNumber="186" reason="1">
<emphasis id="B95E221BFFD8FFBFFF3FF947DFBF0F95" bold="true" box="[151,675,1718,1745]" pageId="10" pageNumber="186">
<taxonomicName id="4C2A858AFFD8FFBFFF3FF947DF210F95" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[151,573,1718,1745]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="186" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFD8FFBFFF3FF947DC260F95" bold="true" box="[151,314,1718,1745]" italics="true" pageId="10" pageNumber="186">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD8FFBFFEE3F947DCD80F95" bold="true" box="[331,452,1718,1745]" italics="true" pageId="10" pageNumber="186">Topobates</emphasis>
)
<emphasis id="B95E221BFFD8FFBFFE7DF947DF210F95" bold="true" box="[469,573,1718,1745]" italics="true" pageId="10" pageNumber="186">tredecim</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D9F60FFD8FFBFFDEDF946DFBF0F95" box="[581,675,1719,1745]" pageId="10" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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(
<figureCitation id="1311E28CFFD8FFBFFF37F92BDDF40FB1" box="[159,232,1754,1781]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="10" pageNumber="186">Figs 7</figureCitation>
,
<figureCitation id="1311E28CFFD8FFBFFF5CF92ADC1E0FB1" box="[244,258,1755,1781]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="10" pageNumber="186">8</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8B95FE09FFD8FFBFFF3FF8D3DC2A0E88" blockId="10.[151,1437,1826,1996]" pageId="10" pageNumber="186">
<emphasis id="B95E221BFFD8FFBFFF3FF8D3DC1A0E78" bold="true" box="[151,262,1826,1852]" pageId="10" pageNumber="186">Material.</emphasis>
<materialsCitation id="3B42F454FFD8FFBFFEB8F8D3DC2A0E88" collectingDate="2014-11-15" collectorName="African &amp; E. A. Kuzmicheva" country="Ethiopia" county="Southeastern" elevation="2570" latitude="6.7216945" location="Bale Mountains National Park" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" pageId="10" pageNumber="186" specimenCount="14" specimenCount-female="2" specimenCount-male="5" stateProvince="Oromiya" typeStatus="holotype">
<typeStatus id="549140ABFFD8FFBFFEB8F8D3DC640E78" box="[272,376,1826,1852]" pageId="10" pageNumber="186" type="holotype">Holotype</typeStatus>
(female) and
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(
<specimenCount id="9D2C3580FFD8FFBFFD4BF8D3DE470E78" box="[739,859,1826,1852]" count="5" pageId="10" pageNumber="186" type="male">five males</specimenCount>
and
<specimenCount id="9D2C3580FFD8FFBFFC3FF8D2D9340E78" box="[919,1064,1826,1853]" count="2" pageId="10" pageNumber="186" type="female">two females</specimenCount>
):
<collectingCounty id="62F48685FFD8FFBFFB95F8D3D9D20E78" box="[1085,1230,1826,1852]" pageId="10" pageNumber="186">Southeastern</collectingCounty>
<collectingCountry id="F33DBE99FFD8FFBFFB7FF8D3D8250E78" box="[1239,1337,1826,1852]" name="Ethiopia" pageId="10" pageNumber="186">Ethiopia</collectingCountry>
,
<collectingRegion id="49EE30EBFFD8FFBFFAEFF8D3DDF70E24" country="Ethiopia" name="Oromiya" pageId="10" pageNumber="186">Oromia Region</collectingRegion>
,
<collectingMunicipality id="6BF16473FFD8FFBFFF5DF8B7DC770E25" box="[245,363,1862,1889]" pageId="10" pageNumber="186">Bale Zone</collectingMunicipality>
,
<location id="8EF5A8D2FFD8FFBFFEDEF8B7DFD60E24" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD8FFBFFEDEF8B7DFD60E24" box="[374,714,1862,1888]" country="Ethiopia" county="Southeastern" latitude="6.7216945" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" name="Bale Mountains National Park" pageId="10" pageNumber="186" stateProvince="Oromiya">Bale Mountains National Park</location>
,
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,
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,
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<quantity id="4CD253ECFFD8FFBFFBBBF8B7D97B0E25" box="[1043,1127,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="2.57" pageId="10" pageNumber="186" unit="m" value="2570.0">2570 m</quantity>
a.s.l.
</elevation>
,
<location id="8EF5A8D2FFD8FFBFFB07F8B6D8410E25" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD8FFBFFB07F8B6D8410E25" box="[1199,1373,1863,1889]" country="Ethiopia" county="Southeastern" latitude="6.7216945" longLatPrecision="1" longitude="39.736694" municipality="Bale Zone" name="Harenna Forest" pageId="10" pageNumber="186" stateProvince="Oromiya">Harenna Forest</location>
, litter in bamboo belt (vegetation represented by dense bamboo thickets of
<taxonomicName id="4C2A858AFFD8FFBFFC0BF89BD9470EC0" box="[931,1115,1898,1924]" class="Liliopsida" family="Poaceae" genus="Oldeania" kingdom="Plantae" order="Poales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="alpina">
<emphasis id="B95E221BFFD8FFBFFC0BF89BD9470EC0" box="[931,1115,1898,1924]" italics="true" pageId="10" pageNumber="186">Oldeania alpina</emphasis>
</taxonomicName>
, the
<collectorName id="26DF9BDFFFD8FFBFFB39F89BD9FB0EC0" box="[1169,1255,1898,1924]" pageId="10" pageNumber="186">African</collectorName>
alpine bamboo,
<emphasis id="B95E221BFFD8FFBFFF3FF87EDF8F0EEC" box="[151,659,1934,1960]" italics="true" pageId="10" pageNumber="186">
<taxonomicName id="4C2A858AFFD8FFBFFF3FF87EDCC10EEC" authority=", Myrsine" authorityName="Myrsine" box="[151,477,1934,1960]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="abyssinica">Hagenia abyssinica, Myrsine</taxonomicName>
melanophloeos,
</emphasis>
and
<taxonomicName id="4C2A858AFFD8FFBFFD60F87FDEBF0EEC" box="[712,931,1934,1960]" class="Magnoliopsida" family="Araliaceae" genus="Schefflera" kingdom="Plantae" order="Apiales" pageId="10" pageNumber="186" phylum="Tracheophyta" rank="species" species="volkensii">
<emphasis id="B95E221BFFD8FFBFFD60F87FDEBF0EEC" box="[712,931,1934,1960]" italics="true" pageId="10" pageNumber="186">Schefflera volkensii</emphasis>
</taxonomicName>
trees accompany bamboo),
<date id="FF94D8C9FFD8FFBFFB72F87FD8460EED" box="[1242,1370,1934,1961]" pageId="10" pageNumber="186" value="2014-11-15">
<collectingDate id="EFD02121FFD8FFBFFB72F87FD8460EED" box="[1242,1370,1934,1961]" pageId="10" pageNumber="186" value="2014-11-15">15.XI.2014</collectingDate>
</date>
(
<collectorName id="26DF9BDFFFD8FFBFFAC1F87EDC360E88" pageId="10" pageNumber="186">E.A. Kuzmicheva</collectorName>
).
</materialsCitation>
</paragraph>
<caption id="DF55AE81FFD9FFBEFF3FF908DE370E72" ID-DOI="http://doi.org/10.5281/zenodo.13219162" ID-Zenodo-Dep="13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="11" pageNumber="187" startId="11.[151,250,1785,1809]" targetBox="[179,1408,196,1746]" targetPageId="11" targetType="figure">
<paragraph id="8B95FE09FFD9FFBEFF3FF908DE370E72" blockId="11.[151,1436,1785,1846]" pageId="11" pageNumber="187">
<emphasis id="B95E221BFFD9FFBEFF3FF908DC0F0E55" bold="true" box="[151,275,1785,1809]" pageId="11" pageNumber="187">FIGURE 7.</emphasis>
<taxonomicName id="4C2A858AFFD9FFBEFEB1F908DFC50E55" authority="Ermilov" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[281,729,1785,1809]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="187" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFD9FFBEFEB1F908DCBF0E55" box="[281,419,1785,1809]" italics="true" pageId="11" pageNumber="187">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFD9FFBEFE1AF908DF050E55" box="[434,537,1785,1809]" italics="true" pageId="11" pageNumber="187">Topobates</emphasis>
)
<emphasis id="B95E221BFFD9FFBEFD80F908DF9C0E55" box="[552,640,1785,1809]" italics="true" pageId="11" pageNumber="187">tredecim</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFD9FFBEFD77F908DE330E55" bold="true" box="[735,815,1785,1809]" pageId="11" pageNumber="187">
<taxonomicNameLabel id="A26D9F60FFD9FFBEFD77F908DE330E55" box="[735,815,1785,1809]" pageId="11" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<paragraph id="8B95FE09FFD9FFBEFF6FF88EDEE00EA5" blockId="11.[151,1436,1919,2018]" pageId="11" pageNumber="187">
The
<typeStatus id="549140ABFFD9FFBEFF52F88EDC400EDD" box="[250,348,1919,1945]" pageId="11" pageNumber="187" type="holotype">holotype</typeStatus>
is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz,
<collectingCountry id="F33DBE99FFD9FFBEFA84F871D8840EDE" box="[1324,1432,1920,1946]" name="Germany" pageId="11" pageNumber="187">Germany</collectingCountry>
;
<materialsCitation id="3B42F454FFD9FFBEFF3FF855D8710EF9" box="[151,1389,1955,1982]" country="Russia" location="University Museum of Zoology" pageId="11" pageNumber="187" specimenCount="7" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount id="9D2C3580FFD9FFBEFF3FF855DC500EFA" box="[151,332,1956,1982]" count="7" pageId="11" pageNumber="187" type="generic" typeStatus="paratype">seven paratypes</specimenCount>
are in the collection of the
<collectingRegion id="49EE30EBFFD9FFBEFD3AF855DE330EFA" box="[658,815,1956,1982]" country="Russia" name="Tyumen'" pageId="11" pageNumber="187">Tyumen State</collectingRegion>
<location id="8EF5A8D2FFD9FFBEFC90F852D9B90EF9" LSID="urn:lsid:plazi:treatment:03834F1FFFD8FFB8FF3FF947D8250C85:8EF5A8D2FFD9FFBEFC90F852D9B90EF9" box="[824,1189,1955,1982]" country="Russia" name="University Museum of Zoology" pageId="11" pageNumber="187" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion id="49EE30EBFFD9FFBEFB1AF855D80C0EFA" box="[1202,1296,1956,1982]" country="Russia" name="Tyumen'" pageId="11" pageNumber="187">Tyumen</collectingRegion>
,
<collectingCountry id="F33DBE99FFD9FFBEFAB7F852D8710EF9" box="[1311,1389,1955,1981]" name="Russia" pageId="11" pageNumber="187">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in a solution of 70% ethanol with a drop of glycerol.
</paragraph>
</subSubSection>
<caption id="DF55AE81FFDEFFB9FF3FF96AD9650FBF" ID-DOI="http://doi.org/10.5281/zenodo.13219166" ID-Zenodo-Dep="13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="12" pageNumber="188" startId="12.[151,250,1691,1715]" targetBox="[176,1420,190,1654]" targetPageId="12" targetType="figure">
<paragraph id="8B95FE09FFDEFFB9FF3FF96AD9650FBF" blockId="12.[151,1436,1691,1788]" pageId="12" pageNumber="188">
<emphasis id="B95E221BFFDEFFB9FF3FF96ADC0B0FF7" bold="true" box="[151,279,1691,1715]" pageId="12" pageNumber="188">FIGURE 8.</emphasis>
<taxonomicName id="4C2A858AFFDEFFB9FE8BF96ADFEC0FF7" authority="Ermilov" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[291,752,1691,1715]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="188" phylum="Arthropoda" rank="species" species="tredecim" status="sp. nov." subGenus="Topobates">
<emphasis id="B95E221BFFDEFFB9FE8BF96ADCB10FF7" box="[291,429,1691,1715]" italics="true" pageId="12" pageNumber="188">Scheloribates</emphasis>
(
<emphasis id="B95E221BFFDEFFB9FE68F96ADF3B0FF7" box="[448,551,1691,1715]" italics="true" pageId="12" pageNumber="188">Topobates</emphasis>
)
<emphasis id="B95E221BFFDEFFB9FD92F96ADF8E0FF7" box="[570,658,1691,1715]" italics="true" pageId="12" pageNumber="188">tredecim</emphasis>
Ermilov
</taxonomicName>
<emphasis id="B95E221BFFDEFFB9FD53F96ADE4C0FF7" bold="true" box="[763,848,1691,1715]" pageId="12" pageNumber="188">
<taxonomicNameLabel id="A26D9F60FFDEFFB9FD53F96ADE4C0FF7" box="[763,848,1691,1715]" pageId="12" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG).
</paragraph>
</caption>
<subSubSection id="C330AD82FFDEFFB9FF6FF8B4D8690EAB" pageId="12" pageNumber="188" type="diagnosis">
<paragraph id="8B95FE09FFDEFFB9FF6FF8B4D8690EAB" blockId="12.[151,1436,1861,2032]" pageId="12" pageNumber="188">
<emphasis id="B95E221BFFDEFFB9FF6FF8B4DC5E0E1B" bold="true" box="[199,322,1861,1887]" pageId="12" pageNumber="188">Diagnosis.</emphasis>
Body length: 390450.
<taxonomicName id="4C2A858AFFDEFFB9FDE3F8B7DFDA0E24" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[587,710,1862,1888]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Rostrum protruding, rounded. Prolamella and translamella absent. Rostral, lamellar and interlamellar setae long, setiform, barbed;
<emphasis id="B95E221BFFDEFFB9FB99F89BD95B0EC7" box="[1073,1095,1898,1923]" italics="true" pageId="12" pageNumber="188">in</emphasis>
˃(=)
<emphasis id="B95E221BFFDEFFB9FBD1F898D9910EC7" box="[1145,1165,1897,1923]" italics="true" pageId="12" pageNumber="188">le</emphasis>
˃
<emphasis id="B95E221BFFDEFFB9FB35F89BD9A90EC7" box="[1181,1205,1898,1923]" italics="true" pageId="12" pageNumber="188">ro</emphasis>
; bothridial seta long, lanceolate, barbed. Thirteen pairs of notogastral setae short, needleform. Four pairs of sacculi. Circumpedal carina long. Anal and adanal setae medium-sized, setiform, thin, roughened. Ventrobasal tubercle of leg tibiae I, II well observable; femur II broadly rounded ventrodistally; tarsus I with 19 setae (
<emphasis id="B95E221BFFDEFFB9FC4FF824DEE20EAB" box="[999,1022,2005,2031]" italics="true" pageId="12" pageNumber="188">l”</emphasis>
absent;
<emphasis id="B95E221BFFDEFFB9FBF4F827D96E0EB4" box="[1116,1138,2006,2032]" italics="true" pageId="12" pageNumber="188">v</emphasis>
present). Tridactylous.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFF67D8190C49" pageId="13" pageNumber="189" type="description">
<paragraph id="8B95FE09FFDFFFB8FF6FFF67D9A209BD" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFF67DCAE09F5" bold="true" box="[199,434,150,177]" pageId="13" pageNumber="189">Description of adult</emphasis>
.
<emphasis id="B95E221BFFDFFFB8FE16FF66DF7C09F4" box="[446,608,151,176]" italics="true" pageId="13" pageNumber="189">Measurements</emphasis>
. Body length: 420 (
<typeStatus id="549140ABFFDFFFB8FC91FF66DEBD09F5" box="[825,929,151,177]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 390420 (male
<typeStatus id="549140ABFFDFFFB8FBF6FF66D9D209F5" box="[1118,1230,151,177]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 420450 (female
<typeStatus id="549140ABFFDFFFB8FF3FFF4ADC140991" box="[151,264,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
); body width (level of pteromorph): 240 (
<typeStatus id="549140ABFFDFFFB8FD48FF4ADE540991" box="[736,840,187,213]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 225240 (male
<typeStatus id="549140ABFFDFFFB8FBA5FF4AD9610991" box="[1037,1149,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 255 (female
<typeStatus id="549140ABFFDFFFB8FA88FF4AD88D0991" box="[1312,1425,187,213]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
); width of ventral plate: 225 (
<typeStatus id="549140ABFFDFFFB8FE67FF2EDF2B09BD" box="[463,567,223,249]" pageId="13" pageNumber="189" type="holotype">holotype</typeStatus>
), 195225 (male
<typeStatus id="549140ABFFDFFFB8FD52FF2EDE7609BD" box="[762,874,223,249]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
), 225240 (female
<typeStatus id="549140ABFFDFFFB8FBEAFF2ED9AF09BD" box="[1090,1203,223,249]" pageId="13" pageNumber="189" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFEF2DC2B0821" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFEF2DC590858" box="[199,325,259,284]" italics="true" pageId="13" pageNumber="189">Integument</emphasis>
. Body color light brown. Surface densely microfoveolate (visible only under high magnification in dissected specimens; ×1000); podosomal region and lateral part of prodorsum partially with dense microgranulate cerotegument.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFE9ED9ED0B5D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFE9EDC5808CD" box="[199,324,367,393]" italics="true" pageId="13" pageNumber="189">Prodorsum</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FEFAFE9EDCD308CD" box="[338,463,367,393]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7A, C</figureCitation>
). Rostrum protruding, rounded. Lamella about 1/2 length of prodorsum; prolamella and translamella absent; sublamella and lateral keel-shaped ridge distinct; sublamellar porose area (7) rounded. Rostral (7175), lamellar (6771) and interlamellar (7579) setae setiform, barbed; bothridial seta (5664) with lanceolate, barbed head, directed posterolaterad; bothridial stalk and head similar in length or stalk slightly longer than head; exobothridial seta (3034) setiform, thin, roughened. Dorsosejugal porose area oval, poorly visible.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFDD2D9750BED" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<taxonomicName id="4C2A858AFFDFFFB8FF6FFDD2DC5F0B78" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[199,323,547,572]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="B95E221BFFDFFFB8FF6FFDD2DC5F0B78" box="[199,323,547,572]" italics="true" pageId="13" pageNumber="189">Notogaster</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDFFFB8FEF3FDD2DCF70B79" box="[347,491,547,573]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7A, C</figureCitation>
). Anterior notogastral margin convex medially.
<taxonomicName id="4C2A858AFFDFFFB8FB9DFDD2D9AC0B79" authorityName="Fernandez-Triana &amp; Ward" authorityYear="2020" box="[1077,1200,547,573]" class="Insecta" family="Braconidae" genus="Notogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="189" phylum="Arthropoda" rank="genus">Notogaster</taxonomicName>
distinctly elongate. Pteromorph medium-sized, broadly rounded laterally, its anterior margin straight (perpendicular to longitudinal axis of body in dorsal aspect). Thirteen pairs of notogastral setae (6) needleform. Four pairs of saccules with small opening and drop-like channel. Opisthonotal gland opening and all lyrifissures distinct.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFD42DDF70A7D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<taxonomicName id="4C2A858AFFDFFFB8FF6FFD42DC4A0B89" authorityName="Krefft" authorityYear="1871" box="[199,342,691,717]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="13" pageNumber="189" phylum="Nematoda" rank="genus">
<emphasis id="B95E221BFFDFFFB8FF6FFD42DC4A0B89" box="[199,342,691,717]" italics="true" pageId="13" pageNumber="189">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1311E28CFFDFFFB8FECFFD42DCF60B89" box="[359,490,691,717]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="13" pageNumber="189">Figs 8AC</figureCitation>
). Subcapitulum size: 105112 × 7582; subcapitular setae (
<emphasis id="B95E221BFFDFFFB8FB3CFD42D9BE0B88" box="[1172,1186,691,716]" italics="true" pageId="13" pageNumber="189">a</emphasis>
: 1922;
<emphasis id="B95E221BFFDFFFB8FAA2FD42D8020B88" box="[1290,1310,691,716]" italics="true" pageId="13" pageNumber="189">m</emphasis>
: 1519;
<emphasis id="B95E221BFFDFFFB8FA2EFD42D8880B89" box="[1414,1428,691,717]" italics="true" pageId="13" pageNumber="189">h</emphasis>
: 2630) setiform, roughened;
<emphasis id="B95E221BFFDFFFB8FE73FD26DCF30BB4" box="[475,495,727,752]" italics="true" pageId="13" pageNumber="189">m</emphasis>
thinner than
<emphasis id="B95E221BFFDFFFB8FD29FD26DF930BB4" box="[641,655,727,752]" italics="true" pageId="13" pageNumber="189">a</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FD6BFD26DFCD0BB5" box="[707,721,727,753]" italics="true" pageId="13" pageNumber="189">h</emphasis>
; both adoral setae (1315) setiform, barbed. Palp length: 6064; postpalpal seta (7) spiniform, roughened. Chelicera length: 112116; setae (
<emphasis id="B95E221BFFDFFFB8FBA6FD0AD92A0A51" box="[1038,1078,763,789]" italics="true" pageId="13" pageNumber="189">cha</emphasis>
: 3437;
<emphasis id="B95E221BFFDFFFB8FB0AFD0AD9D60A51" box="[1186,1226,763,789]" italics="true" pageId="13" pageNumber="189">chb</emphasis>
: 2224) setiform, barbed.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFCB2DCE40A8D" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFCB2DF960A18" box="[199,650,835,861]" italics="true" pageId="13" pageNumber="189">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FD30FCB2DE080A19" box="[664,788,835,861]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7B, C</figureCitation>
). Epimeral formula: 3133; all setae (
<emphasis id="B95E221BFFDFFFB8FB7BFCB2D9EC0A19" box="[1235,1264,835,861]" italics="true" pageId="13" pageNumber="189">1a</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FB55FCB2D8040A19" box="[1277,1304,835,861]" italics="true" pageId="13" pageNumber="189">2a</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FA8EFCB2D85D0A19" box="[1318,1345,835,861]" italics="true" pageId="13" pageNumber="189">3a</emphasis>
: 1113;
<emphasis id="B95E221BFFDFFFB8FF3FFC96DDA80AC5" box="[151,180,871,897]" italics="true" pageId="13" pageNumber="189">1b</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FF6DFC96DDFD0AC5" box="[197,225,871,897]" italics="true" pageId="13" pageNumber="189">3b</emphasis>
,
<emphasis id="B95E221BFFDFFFB8FF5BFC96DC110AC5" box="[243,269,871,897]" italics="true" pageId="13" pageNumber="189">3c</emphasis>
: 2628; others: 1922) setiform, thin, roughened. Humeral porose areas
<emphasis id="B95E221BFFDFFFB8FBC0FC96D9910AC5" box="[1128,1165,871,897]" italics="true" pageId="13" pageNumber="189">Am</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FB64FC96D9F70AC5" box="[1228,1259,871,897]" italics="true" pageId="13" pageNumber="189">Ah</emphasis>
poorly visible, elongate oval. Pedotectum II rounded laterally in ventral aspect. Discidium broadly rounded. Circumpedal carina long, directed to pedotectum II.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFC22DC150D71" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFC22DC8F0AA8" box="[199,403,979,1005]" italics="true" pageId="13" pageNumber="189">Anogenital region</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FE0CFC22DF390AA9" box="[420,549,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1785,1809]" captionTargetBox="[179,1408,196,1746]" captionTargetId="figure-28@11.[164,1423,181,1761]" captionTargetPageId="11" captionText="FIGURE 7. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—dorsal view (legs omitted); B—ventral view (legs omitted); C—right lateral view (legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219162" httpUri="https://zenodo.org/record/13219162/files/figure.png" pageId="13" pageNumber="189">Figs 7B, C</figureCitation>
). Genital (
<emphasis id="B95E221BFFDFFFB8FD37FC22DFB10AA8" box="[671,685,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FD05FC14DFA90AB7" attach="left" box="[685,693,997,1011]" fontSize="6" pageId="13" pageNumber="189">1</subScript>
: 1922;
<emphasis id="B95E221BFFDFFFB8FCB5FC22DE370AA8" box="[797,811,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FC83FC14DE2F0AB7" attach="left" box="[811,819,997,1011]" fontSize="6" pageId="13" pageNumber="189">2</subScript>
<emphasis id="B95E221BFFDFFFB8FCE9FC22DE530AA8" box="[833,847,979,1004]" italics="true" pageId="13" pageNumber="189">g</emphasis>
<subScript id="17AEFC4CFFDFFFB8FCE7FC14DE4B0AB7" attach="left" box="[847,855,997,1011]" fontSize="6" pageId="13" pageNumber="189">4</subScript>
: 1113), aggenital (22), anal (22), and adanal (22) setae setiform, thin, roughened. Adanal lyrifissure distinct, close and parallel to anal plate. Marginal porose area band-like.
</paragraph>
<paragraph id="8B95FE09FFDFFFB8FF6FFBCED8190C49" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFBCEDDE00D1C" box="[199,252,1087,1112]" italics="true" pageId="13" pageNumber="189">Legs</emphasis>
(
<figureCitation id="1311E28CFFDFFFB8FEA4FBCEDC950D1D" box="[268,393,1087,1113]" captionStart="FIGURE 8" captionStartId="12.[151,250,1691,1715]" captionTargetBox="[176,1420,190,1654]" captionTargetId="figure-19@12.[164,1423,181,1667]" captionTargetPageId="12" captionText="FIGURE 8. Scheloribates (Topobates) tredecim Ermilov sp. nov., adult: A—subcapitulum, ventral view; B—palp, right, antiaxial view; C—chelicera, left, paraxial view; D—leg I, right, antiaxial view; E—leg II, left, paraxial view; F—leg III, left, antiaxial view; G—leg IV, left, antiaxial view. Scale bars 20 μm (A, C), 10 μm (B), 50 μm (DG)." figureDoi="http://doi.org/10.5281/zenodo.13219166" httpUri="https://zenodo.org/record/13219166/files/figure.png" pageId="13" pageNumber="189">Figs 8DG</figureCitation>
). Median claw thick; lateral claws thin, with small tubercle distoventrally; all claws slightly barbed on dorsal side. Ventrobasal tubercle of tibiae I, II well observable. Femur II broadly rounded ventrodistally. Proximoventral porose area on tarsi IIV and distoventral porose area on tibiae IIV present but poorly visible; dorsoparaxial porose area on femora IIV and on trochanters III, IV distinct. Formulas of leg setation and solenidia: I (153419) [122], II (152415) [112], III (231315) [110], IV (122312) [010]; homology of setae and solenidia indicated in
<tableCitation id="C6A8CBB2FFDFFFB8FDB2FB02DF720C49" box="[538,622,1267,1293]" captionStart="TABLE 1" captionStartId="4.[151,239,1592,1616]" captionText="TABLE 1. Leg setation and solenidia of adult Scheloribates (Scheloribates) flexibilis Ermilov sp. nov., S. (S.) monosetosus Ermilov sp. nov., S. (S.) triangulus Ermilov sp. nov., and S. (Topobates) tredecim Ermilov sp. nov." pageId="13" pageNumber="189">Table 1</tableCitation>
. Seta
<emphasis id="B95E221BFFDFFFB8FD1AFB02DFA10C48" box="[690,701,1267,1292]" italics="true" pageId="13" pageNumber="189">s</emphasis>
on tarsus I eupathidial, located between
<emphasis id="B95E221BFFDFFFB8FB2EFB02D9BA0C49" box="[1158,1190,1267,1293]" italics="true" pageId="13" pageNumber="189">(u)</emphasis>
and
<emphasis id="B95E221BFFDFFFB8FB75FB02D9E20C49" box="[1245,1278,1267,1293]" italics="true" pageId="13" pageNumber="189">(a)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFAE6DFFC0CD9" pageId="13" pageNumber="189" type="discussion">
<paragraph id="8B95FE09FFDFFFB8FF6FFAE6DFFC0CD9" blockId="13.[151,1437,150,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFAE6DC270C75" bold="true" box="[199,315,1303,1329]" pageId="13" pageNumber="189">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="4C2A858AFFDFFFB8FD3DFAE6D9490C75" authority="(Ramsay, 1966)" baseAuthorityName="Ramsay" baseAuthorityYear="1966" box="[661,1109,1303,1329]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="scheloribatoides" subGenus="Topobates">
<emphasis id="B95E221BFFDFFFB8FD3DFAE6DFB60C75" box="[661,682,1303,1329]" italics="true" pageId="13" pageNumber="189">S.</emphasis>
(
<emphasis id="B95E221BFFDFFFB8FD13FAE6DFCC0C74" box="[699,720,1303,1328]" italics="true" pageId="13" pageNumber="189">T.</emphasis>
)
<emphasis id="B95E221BFFDFFFB8FD49FAE6DE860C75" box="[737,922,1303,1329]" italics="true" pageId="13" pageNumber="189">scheloribatoides</emphasis>
(
<bibRefCitation id="EFBB83F8FFDFFFB8FC04FAE6D9500C75" author="Ramsay, G. W." box="[940,1100,1303,1329]" pageId="13" pageNumber="189" pagination="416 - 425" refId="ref8289" refString="Ramsay, G. W. (1966) Two new oribatid mites (Acari) from a New Zealand pasture. New Zealand Journal of Science, 9, 416 - 425." type="journal article" year="1966">Ramsay, 1966</bibRefCitation>
)
</taxonomicName>
(=
<taxonomicName id="4C2A858AFFDFFFB8FBDEFAE6DDCC0C11" authority="Hammer, 1967" authorityName="Hammer" authorityYear="1967" class="Arachnida" family="Scheloribatidae" genus="Setobates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B95E221BFFDFFFB8FBDEFAE6D8300C74" box="[1142,1324,1303,1329]" italics="true" pageId="13" pageNumber="189">Setobates minor</emphasis>
<bibRefCitation id="EFBB83F8FFDFFFB8FA9CFAE6DDCC0C11" author="Hammer, M." pageId="13" pageNumber="189" pagination="1 - 60" refId="ref8004" refString="Hammer, M. (1967) Investigations on the oribatid fauna of New Zealand. Part II. Det Kongelige Danske Videnskabernes Selskab Biologiske Skrifter, 15, 1 - 60." type="journal article" year="1967">Hammer, 1967</bibRefCitation>
</taxonomicName>
) from
<collectingCountry id="F33DBE99FFDFFFB8FE8BFACADCA70C11" box="[291,443,1339,1365]" name="New Zealand" pageId="13" pageNumber="189">New Zealand</collectingCountry>
in the presence of lanceolate bothridial seta, 13 pairs of minute notogastral setae and tridactylous legs, but differs from the latter by the absence (versus presence) of the prolamella, distinctly longer aggenital, anal and adanal setae, and narrower body.
</paragraph>
</subSubSection>
<subSubSection id="C330AD82FFDFFFB8FF6FFA56D8250C85" box="[199,1337,1447,1473]" pageId="13" pageNumber="189" type="etymology">
<paragraph id="8B95FE09FFDFFFB8FF6FFA56D8250C85" blockId="13.[151,1437,150,1473]" box="[199,1337,1447,1473]" pageId="13" pageNumber="189">
<emphasis id="B95E221BFFDFFFB8FF6FFA56DC520C85" bold="true" box="[199,334,1447,1473]" pageId="13" pageNumber="189">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2A858AFFDFFFB8FD91FA56DF860C85" authorityName="Ermilov &amp; Rybalov &amp; Kuzmicheva" authorityYear="2024" box="[569,666,1447,1473]" class="Arachnida" family="Scheloribatidae" genus="Scheloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="189" phylum="Arthropoda" rank="species" species="tredecim">
<emphasis id="B95E221BFFDFFFB8FD91FA56DF860C85" box="[569,666,1447,1473]" italics="true" pageId="13" pageNumber="189">tredecim</emphasis>
</taxonomicName>
refers to the presence of 13 pairs of setae on the notogaster.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,718 @@
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<mods:title id="50D869BA6B274B5813F53C06321E2AD0">Snailfishes of the genus Careproctus (Perciformes: Liparidae) with a reduced pelvic disk: three new species and new records from the western North Pacific with comments on their phenotypic diversity</mods:title>
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<mods:namePart id="59B1072793988CA1D1FDF0960B5932FE">Kai, Yoshiaki</mods:namePart>
<mods:affiliation id="9C4297BC14B40A5F77929B5FECC46ADA">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:affiliation id="D4AB0F17E6353E9C6AD15D88534DAD97">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:namePart id="9A5334AA631A12A8320E6E7E6EC63177">Mori, Toshiaki</mods:namePart>
<mods:affiliation id="77525B4CB8CAECEF40AA024E5FF6F7B7">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:affiliation id="AD5D84275AE4EC406C5A00F866D8B551">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, ul. Palchevskogo 17, Vladivostok 690041, Russia</mods:affiliation>
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<mods:affiliation id="31983C4EAC0448DF4EFEA20715FA50EE">Demersal Fish Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, National Research and Development Agency, 25 - 259 Shimomekurakubo, Same, Hachinohe, Aomori 031 - 0841, Japan</mods:affiliation>
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<subSubSection id="C3376569FF86386B8BCDAB6CD218FDDC" pageId="3" pageNumber="194" type="nomenclature">
<paragraph id="8B9236E2FF86386B8BCDAB6CD37EFD80" blockId="3.[151,975,549,683]" box="[151,841,549,576]" pageId="3" pageNumber="194">
<heading id="D0DA818EFF86386B8BCDAB6CD37EFD80" bold="true" box="[151,841,549,576]" fontSize="11" level="1" pageId="3" pageNumber="194" reason="1">
<emphasis id="B959EAF0FF86386B8BCDAB6CD37EFD80" bold="true" box="[151,841,549,576]" pageId="3" pageNumber="194">
<taxonomicName id="4C2D4D61FF86386B8BCDAB6CD182FDFF" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[151,437,549,576]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF86386B8BCDAB6CD182FDFF" bold="true" box="[151,437,549,576]" italics="true" pageId="3" pageNumber="194">Careproctus barbatulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A578BFF86386B8AE7AB6CD22DFD80" box="[445,538,549,575]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
Kai, Matsuzaki &amp; Mori
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<paragraph id="8B9236E2FF86386B8BCDAB03D218FDDC" blockId="3.[151,975,549,683]" box="[151,559,585,612]" pageId="3" pageNumber="194">New Japanese name: Isaribi-bikunin</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386B8BCDAB24D3F8FD14" pageId="3" pageNumber="194" type="description">
<paragraph id="8B9236E2FF86386B8BCDAB24D194FD38" blockId="3.[151,975,549,683]" box="[151,419,621,647]" pageId="3" pageNumber="194">
<figureCitation id="13162A67FF86386B8BCDAB24D0D4FD38" box="[151,227,621,647]" captionStart="FIGURE 1" captionStartId="4.[151,250,1585,1609]" captionTargetBox="[209,1378,306,1558]" captionTargetId="figure-82@4.[209,1378,306,1558]" captionTargetPageId="4" captionText="FIGURE 1. Careproctus barbatulus sp. nov., FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="3" pageNumber="194">Figs. 1</figureCitation>
,
<figureCitation id="13162A67FF86386B8BABAB24D172FD38" box="[241,325,621,647]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="3" pageNumber="194">2A, 2B</figureCitation>
;
<tableCitation id="C6AF0359FF86386B8A0BAB24D194FD38" box="[337,419,621,647]" captionStart="TABLE 1" captionStartId="7.[151,239,404,428]" captionText="TABLE 1. Counts and measurements of Careproctus barbatulus sp. nov., C. longibarbatus sp. nov., and C. spinulosus sp. nov." pageId="3" pageNumber="194">Table 1</tableCitation>
</paragraph>
<paragraph id="8B9236E2FF86386B8BCDABD8D3F8FD14" blockId="3.[151,975,549,683]" box="[151,975,657,683]" pageId="3" pageNumber="194">
<uri id="FFBC3AE0FF86386B8BCDABD8D3F8FD14" box="[151,975,657,683]" pageId="3" pageNumber="194">
urn:lsid:zoobank.org:act:
<uuid id="FF8B0C37FF86386B8AF6ABD8D3F8FD14" box="[428,975,657,683]" pageId="3" pageNumber="194">3DD80080-00E2-4BE3-B2A8-DB813B690CE2</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386B8BCDAB90D430FA20" pageId="3" pageNumber="194" type="materials_examined">
<paragraph id="8B9236E2FF86386B8BCDAB90D469FCA8" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<materialsCitation id="3B453CBFFF86386B8BCDAB90D469FCA8" collectingDate="2019-08" collectionCode="FAKU, V" country="Japan" elevation="650" location="Rausu" municipality="Houyu-maru" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="B959EAF0FF86386B8BCDAB90D13CFD4C" bold="true" box="[151,267,729,755]" pageId="3" pageNumber="194">
<typeStatus id="54968840FF86386B8BCDAB90D131FD4C" box="[151,262,729,755]" pageId="3" pageNumber="194" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED3CAE27FF86386B8A49AB90D16AFD4C" box="[275,349,729,755]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8A3FAB90D20BFD4B" box="[357,572,729,756]" metricMagnitude="5" metricUnit="m" metricValue="1.4724015769999999" pageId="3" pageNumber="194" unit="mm" value="1.472401577E8">147240, 157.7 mm</quantity>
SL, female, off
<location id="8EF26039FF86386B89ACAB93D308FD4B" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B89ACAB93D308FD4B" box="[758,831,730,756]" country="Japan" municipality="Houyu-maru" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8816AB90D38AFD4C" box="[844,957,729,755]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8893AB93D43BFD4B" box="[969,1036,730,756]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B8F1AAB90D53EFD4C" box="[1088,1289,729,756]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B8F1AAB90D4F7FD4B" box="[1088,1216,729,756]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
, F/
<collectionCode id="ED3CAE27FF86386B8E77AB90D577FD4C" box="[1325,1344,729,755]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8E12AB93D0E5FCA8" italics="true" pageId="3" pageNumber="194">
<collectingMunicipality id="6BF6AC98FF86386B8E12AB93D0E5FCA8" pageId="3" pageNumber="194">Houyu-maru</collectingMunicipality>
</emphasis>
,
<date id="FF931022FF86386B8B85ABB4D16FFCA8" box="[223,344,765,791]" pageId="3" pageNumber="194" value="2019-08">
<collectingDate id="EFD7E9CAFF86386B8B85ABB4D16FFCA8" box="[223,344,765,791]" pageId="3" pageNumber="194" value="2019-08">Aug. 2019</collectingDate>
</date>
, coll. by Koji Matsuzaki, Shigeki Fujimoto, and Shigetada Fujimoto.
</materialsCitation>
</paragraph>
<paragraph id="8B9236E2FF86386B8B9DAA68D228FAE8" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<materialsCitation id="3B453CBFFF86386B8B9DAA68D21DFCE0" collectionCode="HUMZ, FAKU" country="Japan" elevation="698" latitude="44.033" location="Rausu" longLatPrecision="68" longitude="145.383" municipality="Hokkaido" pageId="3" pageNumber="194" specimenCode="HUMZ 235308, FAKU 146599" specimenCount="1" typeStatus="paratype">
<emphasis id="B959EAF0FF86386B8B9DAA68D171FC84" bold="true" box="[199,326,801,827]" pageId="3" pageNumber="194">
<typeStatus id="54968840FF86386B8B9DAA68D176FC84" box="[199,321,801,827]" pageId="3" pageNumber="194" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DB8B9E99FF86386B8A09AA6BD230FC83" box="[339,519,801,828]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="3" pageNumber="194">HUMZ 235308</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B8916AA68D2CBFC83" box="[588,764,801,828]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 146599</specimenCode>
),
<quantity id="4CD59B07FF86386B884CAA68D3B9FC83" box="[790,910,801,828]" metricMagnitude="-1" metricUnit="m" metricValue="1.5419999999999998" pageId="3" pageNumber="194" unit="mm" value="154.2">154.2 mm</quantity>
SL, male,
<geoCoordinate id="EE195025FF86386B8F40AA68D4B0FC84" box="[1050,1159,801,827]" degrees="44.033" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="44.033">44.033ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF86386B8FC3AA68D527FC83" box="[1177,1296,801,828]" degrees="145.383" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="145.383">145.383ºE</geoCoordinate>
, off
<location id="8EF26039FF86386B8E15AA6BD5AFFC83" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8E15AA6BD5AFFC83" box="[1359,1432,802,828]" country="Japan" latitude="44.033" longLatPrecision="68" longitude="145.383" municipality="Hokkaido" name="Rausu" pageId="3" pageNumber="194">Rausu</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8BCDAA0CD13FFCE0" box="[151,264,837,863]" pageId="3" pageNumber="194">Hokkaido</collectingMunicipality>
,
<collectingCountry id="F33A7672FF86386B8A4EAA0FD160FCDF" box="[276,343,838,864]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8A38AA0CD21DFCE0" box="[354,554,837,864]" metricMagnitude="2" metricUnit="m" metricValue="6.975" metricValueMax="8.25" metricValueMin="5.7" pageId="3" pageNumber="194" unit="m" value="697.5" valueMax="825.0" valueMin="570.0">
<quantity id="4CD59B07FF86386B8A38AA0CD1D5FCDF" box="[354,482,837,864]" metricMagnitude="2" metricUnit="m" metricValue="6.975" metricValueMax="8.25" metricValueMin="5.7" pageId="3" pageNumber="194" unit="m" value="697.5" valueMax="825.0" valueMin="570.0">570825 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8917AA0CD141FC3C" collectingDate="2018-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8917AA0CD257FCE0" box="[589,608,837,863]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8932AA0FD2C0FCE0" box="[616,759,838,863]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8932AA0FD2C0FCE0" box="[616,759,838,863]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B885EAA0CD349FCE0" box="[772,894,837,863]" pageId="3" pageNumber="194" value="2018-08">
<collectingDate id="EFD7E9CAFF86386B885EAA0CD349FCE0" box="[772,894,837,863]" pageId="3" pageNumber="194" value="2018-08">Aug. 2018</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B88BFAA0CD4A1FCE0" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B88BFAA0CD4A1FCE0" box="[997,1174,837,863]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8FF8AA0CD55EFCE0" box="[1186,1385,837,863]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8BCDAA20D141FC3C" box="[151,374,873,899]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8AD8AA20D551FC3C" box="[386,1382,873,900]" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8AD8AA20D1FBFC3C" box="[386,460,873,899]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8A89AA20D291FC3B" box="[467,678,873,900]" metricMagnitude="5" metricUnit="m" metricValue="1.466331903" pageId="3" pageNumber="194" unit="mm" value="1.466331903E8">146633, 190.3 mm</quantity>
SL, female, off
<collectorName id="26D85334FF86386B8806AA23D392FC3B" box="[860,933,874,900]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B88F5AA20D417FC3C" box="[943,1056,873,899]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8F76AA23D458FC3B" box="[1068,1135,874,900]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B8FFAAA20D551FC3C" box="[1184,1382,873,900]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B8FFAAA20D528FC3B" box="[1184,1311,873,900]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8ED3AA20D4E8FC18" collectingDate="2014-09-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8ED3AA20D5ABFC3C" box="[1417,1436,873,899]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8BCDAAC7D111FC18" box="[151,294,910,935]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8BCDAAC7D111FC18" box="[151,294,910,935]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8A6DAAC4D1F2FC18" box="[311,453,909,935]" pageId="3" pageNumber="194" value="2014-09-08">
<collectingDate id="EFD7E9CAFF86386B8A6DAAC4D1F2FC18" box="[311,453,909,935]" pageId="3" pageNumber="194" value="2014-09-08">8 Sep. 2014</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8969AAC4D2D1FC18" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8969AAC4D2D1FC18" box="[563,742,909,935]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B89AEAAC4D38AFC18" box="[756,957,909,935]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B88A7AAC4D4E8FC18" box="[1021,1247,909,935]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8FB7AAC4D3F3FC74" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8FB7AAC4D500FC18" box="[1261,1335,909,935]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8E1BAAC4D13DFC73" metricMagnitude="5" metricUnit="m" metricValue="1.466342208" pageId="3" pageNumber="194" unit="mm" value="1.466342208E8">146634, 220.8 mm</quantity>
SL, male, off
<collectorName id="26D85334FF86386B8AEAAAFBD1CEFC73" box="[432,505,946,972]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B895CAAF8D240FC74" box="[518,631,945,971]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B89D9AAFBD2F1FC73" box="[643,710,946,972]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B89A1AAF8D3F3FC74" box="[763,964,945,972]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B89A1AAF8D34CFC73" box="[763,891,945,972]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B88B2AAF8D318FC50" collectingDate="2014-09-10" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B88B2AAF8D3CCFC74" box="[1000,1019,945,971]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8F5EAAFBD4A4FC74" box="[1028,1171,946,971]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8F5EAAFBD4A4FC74" box="[1028,1171,946,971]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8FF8AAF8D50CFC74" box="[1186,1339,945,971]" pageId="3" pageNumber="194" value="2014-09-10">
<collectingDate id="EFD7E9CAFF86386B8FF8AAF8D50CFC74" box="[1186,1339,945,971]" pageId="3" pageNumber="194" value="2014-09-10">10 Sep. 2014</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8BCDAA9CD170FC50" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8BCDAA9CD170FC50" box="[151,327,981,1007]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8A0BAA9CD220FC50" box="[337,535,981,1007]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B890AAA9CD318FC50" box="[592,815,981,1007]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8860AA9CD1FEFB88" collectionCode="FAKU" country="Japan" county="Shigetada Fujimoto" location="Rausu" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCode="FAKU 146807-146810, 4" specimenCount="3" specimenCount-female="3" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8860AA9CD475FC50" box="[826,1090,981,1007]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 146807146810</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8F17AA9CD46CFC50" box="[1101,1115,981,1007]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">4</specimenCode>
(
<specimenCount id="9D2BFD6BFF86386B8F31AA9CD4EEFC50" box="[1131,1241,981,1007]" count="3" pageId="3" pageNumber="194" type="female">3 females</specimenCount>
, 1 sex unknown),
<quantity id="4CD59B07FF86386B8BCDAAB0D171FBAB" box="[151,326,1017,1044]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.861" metricValueMin="0.939" pageId="3" pageNumber="194" unit="mm" value="140.0" valueMax="186.1" valueMin="93.9">93.9186.1 mm</quantity>
SL, off
<location id="8EF26039FF86386B8AC4AAB3D1D0FBAB" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8AC4AAB3D1D0FBAB" box="[414,487,1018,1044]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8AAAAAB0D256FBAC" box="[496,609,1017,1043]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8931AAB3D299FBAB" box="[619,686,1018,1044]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, coll. by
<location id="8EF26039FF86386B8854AAB0D38AFBAC" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8854AAB0D38AFBAC" box="[782,957,1017,1043]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B889CAAB0D4BCFBAC" box="[966,1163,1017,1043]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8F98AAB0D5AAFBAC" box="[1218,1437,1017,1043]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
(no further collection data)
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8A8FAD54D4B5FB88" box="[469,1154,1053,1080]" collectingDate="2019-08" collectionCode="FAKU" country="Japan" elevation="650" location="Rausu" municipality="Houyu-maru" pageId="3" pageNumber="194" specimenCode="FAKU 147238" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8A8FAD54D24AFB88" box="[469,637,1053,1079]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147238</specimenCode>
, male,
<quantity id="4CD59B07FF86386B8994AD54D377FB87" box="[718,832,1053,1080]" metricMagnitude="-1" metricUnit="m" metricValue="1.607" pageId="3" pageNumber="194" unit="mm" value="160.7">160.7 mm</quantity>
SL, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8FD4AD54D16CFBC0" collectionCode="FAKU" country="Japan" elevation="645" latitude="44.024" location="Rausu" longLatPrecision="68" longitude="145.308" pageId="3" pageNumber="194" specimenCode="FAKU 147910-147913, 147915, 148026" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8FD4AD54D5AFFB88" box="[1166,1432,1053,1079]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147910147913</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8BCDAD08D0D9FBE3" box="[151,238,1089,1116]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">147915</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8BA0AD08D166FBE4" box="[250,337,1089,1115]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">148026</specimenCode>
,
<specimenCount id="9D2BFD6BFF86386B8A07AD08D183FBE4" box="[349,436,1089,1115]" count="3" pageId="3" pageNumber="194" type="male">3 males</specimenCount>
and
<specimenCount id="9D2BFD6BFF86386B8AB6AD08D26BFBE4" box="[492,604,1089,1115]" count="3" pageId="3" pageNumber="194" type="female">3 females</specimenCount>
,
<quantity id="4CD59B07FF86386B8932AD08D31FFBE3" box="[616,808,1089,1116]" metricMagnitude="-1" metricUnit="m" metricValue="1.6634999999999998" metricValueMax="1.7759999999999998" metricValueMin="1.551" pageId="3" pageNumber="194" unit="mm" value="166.35" valueMax="177.6" valueMin="155.1">155.1177.6 mm</quantity>
SL,
<geoCoordinate id="EE195025FF86386B8805AD08D3FBFBE4" box="[863,972,1089,1115]" degrees="44.024" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="44.024">44.024ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF86386B8882AD08D478FBE3" box="[984,1103,1089,1116]" degrees="145.308" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="145.308">145.308ºE</geoCoordinate>
, off
<location id="8EF26039FF86386B8FD9AD0BD4FBFBE3" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8FD9AD0BD4FBFBE3" box="[1155,1228,1090,1116]" country="Japan" latitude="44.024" longLatPrecision="68" longitude="145.308" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8F82AD08D57EFBE4" box="[1240,1353,1089,1115]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8E0FAD0BD5AFFBE3" box="[1365,1432,1090,1116]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8BCDAD2CD16CFBC0" box="[151,347,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">
<quantity id="4CD59B07FF86386B8BCDAD2CD123FB3F" box="[151,276,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">540750 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8A26AD2CD5AFFBC0" box="[380,1432,1125,1152]" collectingDate="2020-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8A26AD2CD1B8FBC0" box="[380,399,1125,1151]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8ACEAD2FD214FBC0" box="[404,547,1126,1151]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8ACEAD2FD214FBC0" box="[404,547,1126,1151]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8974AD2CD291FBC0" box="[558,678,1125,1151]" pageId="3" pageNumber="194" value="2020-08">
<collectingDate id="EFD7E9CAFF86386B8974AD2CD291FBC0" box="[558,678,1125,1151]" pageId="3" pageNumber="194" value="2020-08">Aug. 2020</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B885DAD2CD381FBC0" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B885DAD2CD381FBC0" box="[775,950,1125,1151]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B88E5AD2CD4B4FBC0" box="[959,1155,1125,1151]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8FE0AD2CD5AFFBC0" box="[1210,1432,1125,1151]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<collectionCode id="ED3CAE27FF86386B8BCDADC0D0D6FB1C" box="[151,225,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8BB3ADC0D189FB1B" box="[233,446,1161,1188]" metricMagnitude="5" metricUnit="m" metricValue="1.4887219919999999" pageId="3" pageNumber="194" unit="mm" value="1.488721992E8">148872, 199.2 mm</quantity>
SL, male,
<materialsCitation id="3B453CBFFF86386B8960ADC3D571FB1C" box="[570,1350,1161,1188]" collectionCode="HUMZ, FAKU" country="Japan" pageId="3" pageNumber="194" specimenCode="HUMZ 235309, FAKU 148880" specimenCount="2" specimenCount-female="2" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8960ADC3D2DFFB1B" box="[570,744,1161,1188]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="3" pageNumber="194">HUMZ 235309</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B8878ADC0D3FBFB1C" box="[802,972,1161,1187]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 148880</specimenCode>
),
<specimenCount id="9D2BFD6BFF86386B88BAADC0D467FB1C" box="[992,1104,1161,1187]" count="2" pageId="3" pageNumber="194" type="female">2 females</specimenCount>
, 169.0
<quantity id="4CD59B07FF86386B8FF0ADC0D52CFB1B" box="[1194,1307,1161,1188]" metricMagnitude="-1" metricUnit="m" metricValue="1.918" pageId="3" pageNumber="194" unit="mm" value="191.8">191.8 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation id="3B453CBFFF86386B8E08ADC0D430FB78" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8E08ADC0D5ABFB1C" box="[1362,1436,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8BCDADE4D158FB77" box="[151,367,1197,1224]" metricMagnitude="5" metricUnit="m" metricValue="1.489801734" pageId="3" pageNumber="194" unit="mm" value="1.489801734E8">148980, 173.4 mm</quantity>
SL, male, off
<collectorName id="26D85334FF86386B8942ADE7D256FB77" box="[536,609,1198,1224]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B8934ADE4D2E8FB78" box="[622,735,1197,1223]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B89B6ADE7D318FB77" box="[748,815,1198,1224]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8866ADE4D430FB78" box="[828,1031,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B8866ADE4D38AFB77" box="[828,957,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8F76ADE4D392FB54" collectingDate="2021-07" collectingDateMax="2021-08-31" collectingDateMin="2021-07" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8F76ADE4D408FB78" box="[1068,1087,1197,1223]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8F12ADE7D4E0FB78" box="[1096,1239,1198,1223]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8F12ADE7D4E0FB78" box="[1096,1239,1198,1223]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8FBDADE4D5AFFB78" box="[1255,1432,1197,1223]" pageId="3" pageNumber="194" value="2021-07" valueMax="2021-08-31" valueMin="2021-07-00">
<collectingDate id="EFD7E9CAFF86386B8FBDADE4D5AFFB78" box="[1255,1432,1197,1223]" pageId="3" pageNumber="194" value="2021-07" valueMax="2021-08-31" valueMin="2021-07-00">Jul.Aug. 2021</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8BA2AD98D19BFB54" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8BA2AD98D19BFB54" box="[248,428,1233,1259]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8AE3AD98D2B4FB54" box="[441,643,1233,1259]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8999AD98D392FB54" box="[707,933,1233,1259]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B88E9AD98D219FAB0" collectingDate="2019-08" collectionCode="OS, FAKU" country="Japan" elevation="650" location="Rausu" municipality="Houyu-maru" pageId="3" pageNumber="194" specimenCode="OS 26590, FAKU 147239" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B88E9AD98D410FB53" box="[947,1063,1233,1260]" collectionCode="OS" country="USA" name="Oregon State University" pageId="3" pageNumber="194">OS 26590</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B8F3FAD98D526FB54" box="[1125,1297,1233,1259]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147239</specimenCode>
),
<quantity id="4CD59B07FF86386B8E72AD98D5AAFB53" box="[1320,1437,1233,1260]" metricMagnitude="-1" metricUnit="m" metricValue="1.538" pageId="3" pageNumber="194" unit="mm" value="153.8">153.8 mm</quantity>
SL, female, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8963ADBCD281FA8C" collectionCode="OS, FAKU" country="Japan" elevation="645" latitude="44.024" location="Rausu" longLatPrecision="68" longitude="145.308" pageId="3" pageNumber="194" specimenCode="OS 26591, FAKU 147914" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8963ADBCD29CFAAF" box="[569,683,1269,1296]" collectionCode="OS" country="USA" name="Oregon State University" pageId="3" pageNumber="194">OS 26591</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B89BFADBCD3B8FAB0" box="[741,911,1269,1295]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147914</specimenCode>
),
<quantity id="4CD59B07FF86386B88F9ADBCD421FAAF" box="[931,1046,1269,1296]" metricMagnitude="-1" metricUnit="m" metricValue="1.2429999999999999" pageId="3" pageNumber="194" unit="mm" value="124.3">124.3 mm</quantity>
SL, female,
<geoCoordinate id="EE195025FF86386B8FF2ADBCD522FAB0" box="[1192,1301,1269,1295]" degrees="44.024" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="44.024">44.024ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF86386B8E7AADBCD5AFFAAF" box="[1312,1432,1269,1296]" degrees="145.308" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="145.308">145.308ºE</geoCoordinate>
, off
<location id="8EF26039FF86386B8B98AC53D13CFA8B" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8B98AC53D13CFA8B" box="[194,267,1306,1332]" country="Japan" latitude="44.024" longLatPrecision="68" longitude="145.308" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8A43AC50D1BDFA8C" box="[281,394,1305,1331]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8AC2AC53D1ECFA8B" box="[408,475,1306,1332]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8AB3AC50D281FA8C" box="[489,694,1305,1332]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">
<quantity id="4CD59B07FF86386B8AB3AC50D25CFA8B" box="[489,619,1305,1332]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">540750 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8986AC50D22BFAE8" collectingDate="2020-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8986AC50D2D8FA8C" box="[732,751,1305,1331]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B89A0AC53D3BEFA8C" box="[762,905,1306,1331]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B89A0AC53D3BEFA8C" box="[762,905,1306,1331]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B88C3AC50D422FA8C" box="[921,1045,1305,1331]" pageId="3" pageNumber="194" value="2020-08">
<collectingDate id="EFD7E9CAFF86386B88C3AC50D422FA8C" box="[921,1045,1305,1331]" pageId="3" pageNumber="194" value="2020-08">Aug. 2020</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8FDFAC50D50EFA8C" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8FDFAC50D50EFA8C" box="[1157,1337,1305,1331]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8E1DAC50D136FAE8" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8A61AC74D22BFAE8" box="[315,540,1341,1367]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9236E2FF86386B8B9DAC28D430FA20" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DAC28D1B7FAC4" bold="true" box="[199,384,1377,1403]" pageId="3" pageNumber="194">Other material.</emphasis>
<materialsCitation id="3B453CBFFF86386B8ADCAC28D4F5FAC4" box="[390,1218,1377,1404]" collectionCode="FAKU" country="Japan" elevation="700" location="Rausu" pageId="3" pageNumber="194" specimenCode="FAKU 147740" specimenCount="1" stateProvince="Hokkaido">
<specimenCode id="DB8B9E99FF86386B8ADCAC28D219FAC4" box="[390,558,1377,1403]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147740</specimenCode>
, C&amp;S, off
<location id="8EF26039FF86386B89F9AC2BD2DBFAC3" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B89F9AC2BD2DBFAC3" box="[675,748,1378,1404]" country="Japan" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B89ACAC28D350FAC4" box="[758,871,1377,1403]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B882BAC2BD383FAC3" box="[881,948,1378,1404]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B88BEAC28D49EFAC4" box="[996,1193,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="194" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">
<quantity id="4CD59B07FF86386B88BEAC28D455FAC3" box="[996,1122,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="194" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600800 m</quantity>
depth
</elevation>
, F
</materialsCitation>
/
<materialsCitation id="3B453CBFFF86386B8F91AC28D434FA20" collectingDate="2020-07-20" collectionCode="V" country="Japan" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF86386B8F91AC28D4E9FAC4" box="[1227,1246,1377,1403]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8FBEAC2BD544FAC4" box="[1252,1395,1378,1403]" italics="true" pageId="3" pageNumber="194">Houyu-maru</emphasis>
,
<date id="FF931022FF86386B8EDAAC28D136FA20" pageId="3" pageNumber="194" value="2020-07-20">
<collectingDate id="EFD7E9CAFF86386B8EDAAC28D136FA20" pageId="3" pageNumber="194" value="2020-07-20">20 Jul. 2020</collectingDate>
</date>
, coll. by Koji Matsuzaki, Shigeki Fujimoto, and Shigetada Fujimoto
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386B8B9DACE0D3E1F9EC" pageId="3" pageNumber="194" type="diagnosis">
<paragraph id="8B9236E2FF86386B8B9DACE0D3E1F9EC" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DACE0D175FA7C" bold="true" box="[199,322,1449,1475]" pageId="3" pageNumber="194">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2D4D61FF86386B8ABAACE3D25DFA7B" authorityName="Kroyer" authorityYear="1862" box="[480,618,1450,1476]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF86386B8ABAACE3D25DFA7B" box="[480,618,1450,1476]" italics="true" pageId="3" pageNumber="194">Careproctus</emphasis>
</taxonomicName>
with the following combination of characters: vertebrae 5558; dorsal-fin rays 5053; anal-fin rays 4548; pectoral-fin rays 2734; pectoral fin with shallow notch, lower lobe short, extending slightly posterior to anus; pelvic disk rounded, greatly reduced; cephalic pores 2-6-7-2, chin pores paired in separate pits; gill slit moderately large, extending ventrally to pectoral-fin rays 410; teeth simple or slightly shouldered; anus located below mid-orbit; peritoneum and stomach black.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386D8B9DAF14D145F960" lastPageId="5" lastPageNumber="196" pageId="3" pageNumber="194" type="description">
<paragraph id="8B9236E2FF86386B8B9DAF14D5B0F9C8" blockId="3.[151,1437,729,2016]" box="[199,1415,1629,1656]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DAF14D16EF9C8" bold="true" box="[199,345,1629,1655]" pageId="3" pageNumber="194">Description.</emphasis>
Measurements shown in
<tableCitation id="C6AF0359FF86386B892CAF14D2FDF9C8" box="[630,714,1629,1655]" captionStart="TABLE 1" captionStartId="7.[151,239,404,428]" captionText="TABLE 1. Counts and measurements of Careproctus barbatulus sp. nov., C. longibarbatus sp. nov., and C. spinulosus sp. nov." pageId="3" pageNumber="194">Table 1</tableCitation>
.
<typeStatus id="54968840FF86386B898DAF17D30EF9C7" box="[727,825,1630,1656]" pageId="3" pageNumber="194" type="paratype">Paratype</typeStatus>
data given in parentheses if different from
<typeStatus id="54968840FF86386B8E44AF14D5B3F9C8" box="[1310,1412,1629,1655]" pageId="3" pageNumber="194" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B9236E2FF86386C8B9DAFC8D2DCFEA2" blockId="3.[151,1437,729,2016]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="195" pageId="3" pageNumber="194">
Body moderately deep, compressed, tapering posteriorly (
<figureCitation id="13162A67FF86386B883CAFC8D39CF924" box="[870,939,1665,1691]" captionStart="FIGURE 1" captionStartId="4.[151,250,1585,1609]" captionTargetBox="[209,1378,306,1558]" captionTargetId="figure-82@4.[209,1378,306,1558]" captionTargetPageId="4" captionText="FIGURE 1. Careproctus barbatulus sp. nov., FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="3" pageNumber="194">Fig. 1</figureCitation>
). Skin relatively thick, covered by cactus-like prickles; gelatinous tissue under skin moderately developed. Head large, moderately compressed, dorsal profile rounded from nape to snout. Snout rounded, not protruding anterior to mouth. Mouth moderately large, terminal, slightly oblique (or horizontal); maxilla extending to mid orbit; oral cleft extending to anterior rim of orbit. Premaxillary teeth simple or weakly shouldered, arranged in oblique rows of 10 (610) teeth per row forming broad band; inner teeth larger, recurved. Mandibular teeth simple or weakly shouldered, arranged in oblique rows of 9 (49) teeth per row forming broad band; inner teeth larger, recurved. Narrow diastema at symphysis of upper and lower jaws. Orbit of moderate size, rounded; pupil rounded. Nostril single, with short tube, level with mid-orbit. Cephalic sensory pores of moderate size, rims thickened: nasal pores 2, maxillary pores 6, preoperculomandibular pores 7, suprabranchial pores 2; cephalic pore pattern 2-6-7-2. Chin pores paired in separate pits, well separated. Coronal pore absent. Gill slit moderately large, upper margin level with mid-orbit, extending ventrally to pectoral-fin ray 5 (410). Gill rakers 79, short, with tiny spines on tips (not counted in
<typeStatus id="54968840FF81386C8F68A9F2D4AEFF6A" box="[1074,1177,187,213]" pageId="4" pageNumber="195" type="holotype">holotype</typeStatus>
). Opercular flap blunt, pointed posteriorly; tip level with ventral rim of orbit. Pyloric caeca 15 (1422), thick, moderately long, covering half of stomach, on center-left side of visceral cavity.
</paragraph>
<caption id="DF52666AFF81386C8BCDAF78D3E6F92E" ID-DOI="http://doi.org/10.5281/zenodo.13219212" ID-Zenodo-Dep="13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="4" pageNumber="195" startId="4.[151,250,1585,1609]" targetBox="[209,1378,306,1558]" targetPageId="4" targetType="figure">
<paragraph id="8B9236E2FF81386C8BCDAF78D3E6F92E" blockId="4.[151,1437,1585,1681]" pageId="4" pageNumber="195">
<emphasis id="B959EAF0FF81386C8BCDAF78D123F9F6" bold="true" box="[151,276,1585,1609]" pageId="4" pageNumber="195">FIGURE 1.</emphasis>
<taxonomicName id="4C2D4D61FF81386C8A46AF78D239F9F6" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[284,526,1585,1609]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF81386C8A46AF78D239F9F6" box="[284,526,1585,1609]" italics="true" pageId="4" pageNumber="195">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF81386C894CAF78D25FF9F6" bold="true" box="[534,616,1585,1609]" pageId="4" pageNumber="195">
<taxonomicNameLabel id="A26A578BFF81386C894CAF78D25FF9F6" box="[534,616,1585,1609]" pageId="4" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively.
</paragraph>
</caption>
<paragraph id="8B9236E2FF81386C8B9DAFAAD38BF846" blockId="4.[151,1437,1763,2042]" pageId="4" pageNumber="195">
Vertebrae 57 (5558), precaudal 10 (910) and caudal 47 (4649). Pleural ribs minute, thin, in 1 pair (1 or 2 pairs) on abdominal vertebra 9 (810). Dorsal-fin rays 51 (5053), not exserted. Anteriormost dorsal-fin pterygiophore inserted between neural spines 3 and 4 (or 4 and 5), bearing a single ray (sometimes rayless). Anal-fin rays 46 (4548), not exserted. Two (1 or 2) anal-fin pterygiophores anterior to 1
<superScript id="7C589BAAFF81386C88BDAE06D3C5F8E2" attach="left" box="[999,1010,1871,1885]" fontSize="6" pageId="4" pageNumber="195">st</superScript>
haemal spine, each bearing a single ray. Membrane of posterior dorsal- and anal-fin rays attached about equidistant to caudal fin. Caudal fin slightly rounded. Principal caudal-fin rays 7 (79), 3 (34) and 4 (45) rays on upper and lower hypurals, respectively, dorsal procurrent rays 1 (01), ventral procurrent rays 0 (01). Hypural plates fused with terminal vertebral centrum and parhypural, upper and lower hypural plate separated by distinct slit.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DA9DED19BFE36" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="196">
Pectoral fin moderately notched, with 30 (2734) rays; upper lobe with 23 (2025) rays, extending just beyond (or slightly anterior to) level of anal fin origin; lower lobe short, with 7 (710) rays, 4
<superScript id="7C589BAAFF80386D8F1EA9F3D466FF77" attach="left" box="[1092,1105,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
(5
<superScript id="7C589BAAFF80386D8F37A9F3D44DFF77" attach="none" box="[1133,1146,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
7
<superScript id="7C589BAAFF80386D8FCCA9F3D494FF77" attach="none" box="[1174,1187,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
) ray from ventralmost longest, extending slightly posterior to anus. Rays in notch more widely spaced than rays of lobes. Tip of pectoral-fin rays of upper lobe slightly protruding, rays of lower lobe strongly exserted. Uppermost pectoral-fin base level with mouth corner. Lowermost pectoral-fin base slightly anterior to (or below) anterior rim of orbit. Pelvic disk rounded, greatly reduced (sometimes modified as lobe). Anus much closer to pelvic disk than to anal fin origin, located below mid-orbit.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DA8DAD555FDDE" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="196">
Pectoral girdle (based on FAKU 146634 and 147740:
<figureCitation id="13162A67FF80386D8843A8DAD3A7FE12" box="[793,912,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="5" pageNumber="196">Fig. 2A, B</figureCitation>
): Proximal pectoral radials 3 or 4; upper radials 1 or 2, hour-grass shaped with two notches, equally spaced, radials 2 or 3 (lowermost within upper radials) with one notch, radial 4 rounded squarish, widely separated from upper radials. Interradial fenestrae 2 or 3, extending between scapula and radials 13; fenestra 1 roundish, fenestrae 2 and 3 roundish or elliptical, sometimes with notch. Scapula broadly Y-shaped, with robust helve. Coracoid diamond-shaped or a roundish triangle, with short thin helve. Distal radials present at base of pectoral-fin rays 2 to 2127, absent from base of ventralmost 78 rays.
</paragraph>
<caption id="DF52666AFF80386D8BCDADCED16EFA90" ID-DOI="http://doi.org/10.5281/zenodo.13219214" ID-Zenodo-Dep="13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="5" pageNumber="196" startId="5.[151,250,1159,1183]" targetBox="[157,1432,673,1131]" targetPageId="5" targetType="figure">
<paragraph id="8B9236E2FF80386D8BCDADCED16EFA90" blockId="5.[151,1436,1159,1327]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8BCDADCED124FB20" bold="true" box="[151,275,1159,1183]" pageId="5" pageNumber="196">FIGURE 2.</emphasis>
Pectoral girdle in five species of
<taxonomicName id="4C2D4D61FF80386D8932ADC1D2D2FB20" authorityName="Kroyer" authorityYear="1862" box="[616,741,1160,1183]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF80386D8932ADC1D2D2FB20" box="[616,741,1160,1183]" italics="true" pageId="5" pageNumber="196">Careproctus</emphasis>
</taxonomicName>
with a reduced pelvic disk.
<taxonomicName id="4C2D4D61FF80386D8F59ADC1D4C2FB20" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[1027,1269,1159,1183]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8F59ADC1D4C2FB20" box="[1027,1269,1159,1183]" italics="true" pageId="5" pageNumber="196">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8FA1ADCED57BFB20" bold="true" box="[1275,1356,1159,1183]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8FA1ADCED57BFB20" box="[1275,1356,1159,1183]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 146634, paratype, 220.8 mm SL (A);
<taxonomicName id="4C2D4D61FF80386D8954ADE5D2C9FB7C" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[526,766,1195,1219]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8954ADE5D2C9FB7C" box="[526,766,1195,1219]" italics="true" pageId="5" pageNumber="196">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8859ADE2D364FB7C" bold="true" box="[771,851,1195,1219]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8859ADE2D364FB7C" box="[771,851,1195,1219]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 147740, non-type (B);
<taxonomicName id="4C2D4D61FF80386D8FD6ADE5D5ABFB7C" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[1164,1436,1195,1219]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="longibarbatus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8FD6ADE5D5ABFB7C" box="[1164,1436,1195,1219]" italics="true" pageId="5" pageNumber="196">Careproctus longibarbatus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8BCDAD86D0DDFB58" bold="true" box="[151,234,1231,1255]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8BCDAD86D0DDFB58" box="[151,234,1231,1255]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 148477, paratype, 223.0 mm SL (C);
<taxonomicName id="4C2D4D61FF80386D8989AD99D368FB58" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[723,863,1231,1255]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8989AD99D2D3FB58" box="[723,740,1232,1255]" italics="true" pageId="5" pageNumber="196">C</emphasis>
.
<emphasis id="B959EAF0FF80386D89AEAD86D368FB58" box="[756,863,1231,1255]" italics="true" pageId="5" pageNumber="196">spinulosus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8832AD86D38CFB58" bold="true" box="[872,955,1231,1255]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8832AD86D38CFB58" box="[872,955,1231,1255]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 146632, paratype, 199.8 mm SL (D);
<taxonomicName id="4C2D4D61FF80386D8BCDADBDD178FAB3" authority=", FAKU" authorityName="FAKU" box="[151,335,1267,1292]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF80386D8BCDADBDD0CAFAB4" box="[151,253,1267,1291]" italics="true" pageId="5" pageNumber="196">C. mederi</emphasis>
, FAKU
</taxonomicName>
148478, non-type, 143.6 mm SL (E);
<taxonomicName id="4C2D4D61FF80386D89BBADBDD384FAB4" authority=", FAKU S" authorityName="FAKU S" box="[737,947,1267,1292]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF80386D89BBADBDD37BFAB4" box="[737,844,1267,1291]" italics="true" pageId="5" pageNumber="196">C. gilberti</emphasis>
, FAKU S
</taxonomicName>
4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage.
</paragraph>
</caption>
<paragraph id="8B9236E2FF80386D8B9DACC8D158F994" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
Coloration (
<figureCitation id="13162A67FF80386D8A0AACC8D1A3FA23" box="[336,404,1409,1436]" captionStart="FIGURE 1" captionStartId="4.[151,250,1585,1609]" captionTargetBox="[209,1378,306,1558]" captionTargetId="figure-82@4.[209,1378,306,1558]" captionTargetPageId="4" captionText="FIGURE 1. Careproctus barbatulus sp. nov., FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="5" pageNumber="196">Fig. 1</figureCitation>
): When fresh (based on FAKU 148026), head and body pale to dark reddish; tip of nostril dark brown; postorbital region and belly somewhat paler. Iris dark silvery. Dorsal, anal, and caudal fins pale to dark reddish; pectoral fin dark red, basal part paler. Pelvic disk white. When preserved, head and body white; upper and lower jaws mottled with black; nostril margined with black. Eyes black. Oral and branchial cavities pale; peritoneum and stomach black.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DAF7CD145F960" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8B9DAF7CD144F9F0" bold="true" box="[199,371,1589,1615]" pageId="5" pageNumber="196">Reproduction.</emphasis>
The
<typeStatus id="54968840FF80386D8AF4AF7CD227F9F0" box="[430,528,1589,1615]" pageId="5" pageNumber="196" type="holotype">holotype</typeStatus>
(FAKU 147240) was an immature female. FAKU 147911 (
<quantity id="4CD59B07FF80386D8FF5AF7FD514F9EF" box="[1199,1315,1590,1616]" metricMagnitude="-1" metricUnit="m" metricValue="1.708" pageId="5" pageNumber="196" unit="mm" value="170.8">170.8 mm</quantity>
SL) was a spent female with yolked (
<quantity id="4CD59B07FF80386D8A9AAF13D252F9CB" box="[448,613,1626,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.605" metricValueMax="2.8" metricValueMin="2.41" pageId="5" pageNumber="196" unit="mm" value="2.605" valueMax="2.8" valueMin="2.41">2.412.80 mm</quantity>
diameter) and small immature (less than
<quantity id="4CD59B07FF80386D8F6EAF10D4AFF9CB" box="[1076,1176,1625,1652]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="196" unit="mm" value="0.5">0.50 mm</quantity>
diameter) eggs. FAKU 148880 (169.0 mm SL and
<quantity id="4CD59B07FF80386D8A90AF37D20BF927" box="[458,572,1662,1688]" metricMagnitude="-1" metricUnit="m" metricValue="1.918" pageId="5" pageNumber="196" unit="mm" value="191.8">191.8 mm</quantity>
SL) were also spent females with yolked (
<quantity id="4CD59B07FF80386D8F43AF37D489F927" box="[1049,1214,1662,1688]" metricMagnitude="-3" metricUnit="m" metricValue="3.225" metricValueMax="3.48" metricValueMin="2.97" pageId="5" pageNumber="196" unit="mm" value="3.225" valueMax="3.48" valueMin="2.97">2.973.48 mm</quantity>
diameter and
<quantity id="4CD59B07FF80386D8E07AF37D0CCF903" metricMagnitude="-3" metricUnit="m" metricValue="3.0949999999999998" metricValueMax="3.38" metricValueMin="2.81" pageId="5" pageNumber="196" unit="mm" value="3.0949999999999998" valueMax="3.38" valueMin="2.81">2.81 3.38 mm</quantity>
diameter, respectively) and small immature (less than
<quantity id="4CD59B07FF80386D8805AFE8D3F4F903" box="[863,963,1697,1724]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="196" unit="mm" value="0.5">0.50 mm</quantity>
diameter in both specimens) eggs. Yolked eggs were rounded.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF80386D8B9DAFA0D138F897" pageId="5" pageNumber="196" type="distribution">
<paragraph id="8B9236E2FF80386D8B9DAFA0D138F897" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8B9DAFA0D156F8BC" bold="true" box="[199,353,1769,1795]" pageId="5" pageNumber="196">Distribution.</emphasis>
Known only from the southern Sea of Okhotsk, off Rausu,
<collectingRegion id="49E9F800FF80386D88B4AFA0D468F8BC" box="[1006,1119,1769,1795]" country="Japan" name="Hokkaido" pageId="5" pageNumber="196">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF80386D8F33AFA3D49BF8BB" box="[1129,1196,1770,1796]" name="Japan" pageId="5" pageNumber="196">Japan</collectingCountry>
, at depths of
<quantity id="4CD59B07FF80386D8E60AFA0D09AF897" metricMagnitude="2" metricUnit="m" metricValue="6.625" metricValueMax="8.25" metricValueMin="5.0" pageId="5" pageNumber="196" unit="m" value="662.5" valueMax="825.0" valueMin="500.0">500825 m</quantity>
(
<figureCitation id="13162A67FF80386D8BE6AE44D136F897" box="[188,257,1805,1832]" captionStart="FIGURE 3" captionStartId="6.[151,250,729,753]" captionTargetBox="[175,1410,188,699]" captionTargetId="figure-20@6.[171,1416,181,706]" captionTargetPageId="6" captionText="FIGURE 3. Distributional records of species of Careproctus with a reduced pelvic disk based on the present and previous studies (see text)." figureDoi="http://doi.org/10.5281/zenodo.13219216" httpUri="https://zenodo.org/record/13219216/files/figure.png" pageId="5" pageNumber="196">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF52666AFF83386E8BCDAB90D17EFCA9" ID-DOI="http://doi.org/10.5281/zenodo.13219216" ID-Zenodo-Dep="13219216" httpUri="https://zenodo.org/record/13219216/files/figure.png" pageId="6" pageNumber="197" startId="6.[151,250,729,753]" targetBox="[175,1410,188,699]" targetPageId="6" targetType="figure">
<paragraph id="8B9236E2FF83386E8BCDAB90D17EFCA9" blockId="6.[151,1436,729,790]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8BCDAB90D122FD4D" bold="true" box="[151,277,729,754]" pageId="6" pageNumber="197">FIGURE 3.</emphasis>
Distributional records of species of
<taxonomicName id="4C2D4D61FF83386E89CCAB93D324FD4E" authorityName="Kroyer" authorityYear="1862" box="[662,787,730,753]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF83386E89CCAB93D324FD4E" box="[662,787,730,753]" italics="true" pageId="6" pageNumber="197">Careproctus</emphasis>
</taxonomicName>
with a reduced pelvic disk based on the present and previous studies (see text).
</paragraph>
</caption>
<subSubSection id="C3376569FF83386E8B9DAA21D38FFC19" pageId="6" pageNumber="197" type="etymology">
<paragraph id="8B9236E2FF83386E8B9DAA21D38FFC19" blockId="6.[151,1436,872,1402]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8B9DAA21D179FC3D" bold="true" box="[199,334,872,898]" pageId="6" pageNumber="197">Etymology.</emphasis>
The specific name
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<emphasis id="B959EAF0FF83386E8969AA21D29BFC3D" box="[563,684,872,898]" italics="true" pageId="6" pageNumber="197">barbatulus</emphasis>
</taxonomicName>
” derived from the diminutive form of Latin “barba” (barbel), refers to the short lower lobe of the pectoral fin, reminiscent of a short barbel.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF83386E8B9DAAF9D1D6FAC5" pageId="6" pageNumber="197" type="discussion">
<paragraph id="8B9236E2FF83386E8B9DAAF9D1D6FAC5" blockId="6.[151,1436,872,1402]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8B9DAAF9D10CFC75" bold="true" box="[199,315,944,970]" pageId="6" pageNumber="197">Remarks.</emphasis>
Within the
<taxonomicName id="4C2D4D61FF83386E8A98AAF9D201FC75" box="[450,566,944,970]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF83386E8A98AAF9D201FC75" box="[450,566,944,970]" italics="true" pageId="6" pageNumber="197">C. gilberti</emphasis>
</taxonomicName>
species group,
<taxonomicName id="4C2D4D61FF83386E89B2AAF9D3B4FC75" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[744,899,944,970]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF83386E89B2AAF9D3B4FC75" box="[744,899,944,970]" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF83386E88D1AAF9D3D1FC75" bold="true" box="[907,998,944,970]" pageId="6" pageNumber="197">
<taxonomicNameLabel id="A26A578BFF83386E88D1AAF9D3D1FC75" box="[907,998,944,970]" pageId="6" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares some morphological characters with
<taxonomicName id="4C2D4D61FF83386E8B88AA9DD173FC51" box="[210,324,980,1006]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF83386E8B88AA9DD173FC51" box="[210,324,980,1006]" italics="true" pageId="6" pageNumber="197">C. mederi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D61FF83386E8A25AA9DD230FC51" authorityName="Gilbert" authorityYear="1896" box="[383,519,980,1006]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF83386E8A25AA9DD230FC51" box="[383,519,980,1006]" italics="true" pageId="6" pageNumber="197">C. ostentum</emphasis>
</taxonomicName>
, including a short lower lobe of the pectoral fin, extending slightly posterior to the anus, and relatively thick skin with strong prickles. The new species can be distinguished from the latter two species by the gill slit length (extending to pectoral-fin ray 410 vs.
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<emphasis id="B959EAF0FF83386E8F52AD55D440FB89" box="[1032,1143,1052,1078]" italics="true" pageId="6" pageNumber="197">C. mederi</emphasis>
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and
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<taxonomicName id="4C2D4D61FF83386E8E56AD55D5A4FB89" authorityName="Gilbert" authorityYear="1896" box="[1292,1427,1052,1078]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF83386E8E56AD55D5A4FB89" box="[1292,1427,1052,1078]" italics="true" pageId="6" pageNumber="197">C. ostentum</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFBC4B13FF83386E8BC5AD09D18EFBE5" author="Mecklenburg, C. W. &amp; Mecklenburg, T. A. &amp; Thorsteinson, L. K." box="[159,441,1088,1114]" pageId="6" pageNumber="197" refId="ref18345" refString="Mecklenburg, C. W., Mecklenburg, T. A. &amp; Thorsteinson, L. K. (2002) Fishes of Alaska. American Fisheries Society, Bethesda, Maryland, 1037 pp." type="book" year="2002">
Mecklenburg
<emphasis id="B959EAF0FF83386E8A67AD08D141FBE5" box="[317,374,1088,1114]" italics="true" pageId="6" pageNumber="197">et al.</emphasis>
2002
</bibRefCitation>
; this study) and coloration of the peritoneum (dark or black vs. pale with black dots). In addition,
<taxonomicName id="4C2D4D61FF83386E8A5AAD2DD1E8FBC1" box="[256,479,1124,1150]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF83386E8A5AAD2DD1E8FBC1" box="[256,479,1124,1150]" italics="true" pageId="6" pageNumber="197">Careproctus mederi</emphasis>
</taxonomicName>
differs from the new species by having the lower end of the pectoral girdle protruding anteriorly (vs. not protruding, see description below). Although
<taxonomicName id="4C2D4D61FF83386E882FADC1D5AFFB1D" authority="Gilbert &amp; Burke, 1912" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[885,1432,1160,1186]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="acanthodes">
<emphasis id="B959EAF0FF83386E882FADC1D4BFFB1D" box="[885,1160,1160,1186]" italics="true" pageId="6" pageNumber="197">Careproctus acanthodes</emphasis>
<bibRefCitation id="EFBC4B13FF83386E8FCBADC1D5AFFB1D" author="Gilbert, C. H. &amp; Burke, C. V." box="[1169,1432,1160,1186]" pageId="6" pageNumber="197" pagination="31 - 96" refId="ref17636" refString="Gilbert, C. H. &amp; Burke, C. V. (1912) Fishes from Bering Sea and Kamchatka. Bulletin of the Bureau of Fisheries, 30, 31 - 96." type="journal article" year="1912">Gilbert &amp; Burke, 1912</bibRefCitation>
</taxonomicName>
, a member of the
<taxonomicName id="4C2D4D61FF83386E8A39ADE5D1DBFB79" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[355,492,1196,1222]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="rastrinus">
<emphasis id="B959EAF0FF83386E8A39ADE5D1DBFB79" box="[355,492,1196,1222]" italics="true" pageId="6" pageNumber="197">C. rastrinus</emphasis>
</taxonomicName>
species group, has the shortest pectoral fin lower lobe among the species group, the present new species clearly differs from
<taxonomicName id="4C2D4D61FF83386E89D5AD99D306FB55" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[655,817,1232,1258]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="acanthodes">
<emphasis id="B959EAF0FF83386E89D5AD99D306FB55" box="[655,817,1232,1258]" italics="true" pageId="6" pageNumber="197">C. acanthodes</emphasis>
</taxonomicName>
in having a smaller pelvic disk (0.40.9% SL vs. 4.2 7.6% SL) (
<bibRefCitation id="EFBC4B13FF83386E8A48ADBDD1F7FAB0" author="Orr, J. W. &amp; Kai, Y. &amp; Nakabo, T." box="[274,448,1268,1295]" pageId="6" pageNumber="197" pagination="301 - 348" refId="ref18697" refString="Orr, J. W., Kai, Y. &amp; Nakabo, T. (2015) Snailfishes of the Careproctus rastrinus complex (Liparidae): redescriptions of seven species in the North Pacific Ocean region, with the description of a new species from the Beaufort Sea. Zootaxa, 4018 (3), 301 - 348. https: // doi. org / 10.11646 / zootaxa. 4018.3.1" type="journal article" year="2015">
Orr
<emphasis id="B959EAF0FF83386E8A19ADBCD143FAB1" box="[323,372,1268,1294]" italics="true" pageId="6" pageNumber="197">et al</emphasis>
. 2015
</bibRefCitation>
). Among the species of
<taxonomicName id="4C2D4D61FF83386E8994ADBDD368FAB1" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[718,863,1268,1294]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF83386E8994ADBDD368FAB1" box="[718,863,1268,1294]" italics="true" pageId="6" pageNumber="197">Elassodiscus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2D4D61FF83386E8837ADBDD5ABFAB1" authority="Gilbert &amp; Burke, 1912" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[877,1436,1268,1294]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="tremebundus">
<emphasis id="B959EAF0FF83386E8837ADBDD4A1FAB1" box="[877,1174,1268,1294]" italics="true" pageId="6" pageNumber="197">Elassodiscus tremebundus</emphasis>
<bibRefCitation id="EFBC4B13FF83386E8FC7ADBDD5ABFAB1" author="Gilbert, C. H. &amp; Burke, C. V." box="[1181,1436,1268,1294]" pageId="6" pageNumber="197" pagination="31 - 96" refId="ref17636" refString="Gilbert, C. H. &amp; Burke, C. V. (1912) Fishes from Bering Sea and Kamchatka. Bulletin of the Bureau of Fisheries, 30, 31 - 96." type="journal article" year="1912">Gilbert &amp; Burke, 1912</bibRefCitation>
</taxonomicName>
also has a relatively short pectoral fin lower lobe. However, together with the diagnostic characters of the genus,
<taxonomicName id="4C2D4D61FF83386E8BCDAC74D17EFAE9" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[151,329,1340,1366]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="tremebundus">
<emphasis id="B959EAF0FF83386E8BCDAC74D17EFAE9" box="[151,329,1340,1366]" italics="true" pageId="6" pageNumber="197">E. tremebundus</emphasis>
</taxonomicName>
is further distinguishable from
<taxonomicName id="4C2D4D61FF83386E89EFAC75D366FAE9" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[693,849,1340,1366]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus">
<emphasis id="B959EAF0FF83386E89EFAC75D366FAE9" box="[693,849,1340,1366]" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
in having a single suprabranchial pore (vs.
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<taxonomicName id="4C2D4D61FF83386E8ED9AC75D127FAC5" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus">
<emphasis id="B959EAF0FF83386E8ED9AC75D127FAC5" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFBC4B13FF83386E8A73AC29D1E1FAC5" author="Kai, Y. &amp; Matsuzaki, K. &amp; Orr, J. W. &amp; Mori, T. &amp; Kamiunten, M." box="[297,470,1376,1402]" pageId="6" pageNumber="197" pagination="55 - 66" refId="ref17899" refString="Kai, Y., Matsuzaki, K., Orr, J. W., Mori, T. &amp; Kamiunten, M. (2020) A new species of Elassodiscus (Cottoidei: Liparidae) from the North Pacific with an emended diagnosis of the genus. Ichthyological Research, 68, 55 - 66. https: // doi. org / 10.1007 / s 10228 - 020 - 00764 - 4" type="journal article" year="2020">
Kai
<emphasis id="B959EAF0FF83386E8A00AC28D1BCFAC5" box="[346,395,1376,1402]" italics="true" pageId="6" pageNumber="197">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,842 @@
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<mods:title id="23B43DF8522E053C92E3DDCC0B7C858A">Snailfishes of the genus Careproctus (Perciformes: Liparidae) with a reduced pelvic disk: three new species and new records from the western North Pacific with comments on their phenotypic diversity</mods:title>
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<mods:namePart id="56B014CC261E66E7222A2D146F2A31B8">Kai, Yoshiaki</mods:namePart>
<mods:affiliation id="C069A57E244235EC01F1CEAED882FF8E">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:namePart id="A040CDA30A89136429A41A1652C51CB2">Matsuzaki, Koji</mods:namePart>
<mods:affiliation id="08C5D4A5FDDE3A4CDDD6F12DF703958C">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:url id="0D292F6288B53A459A556DC215032E9A">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:classification id="67C450214BEE03FBB7D6001BC7EDC5C7">journal article</mods:classification>
<mods:identifier id="CB8593C9037A0A443BF9E4BAE90D8399" type="DOI">10.11646/zootaxa.5492.2.3</mods:identifier>
<mods:identifier id="EBAFC50FA5B82DF715A6909948478BE8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4C6C76C7B0B47463D62F47E7AD5C7E8E" type="Zenodo-Dep">13219210</mods:identifier>
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<heading id="D0DA818EFF8938648BCDAD86D208FB57" bold="true" box="[151,575,1230,1257]" fontSize="11" level="1" pageId="12" pageNumber="203" reason="1">
<taxonomicName id="4C2D4D61FF8938648BCDAD86D208FB57" ID-CoL="R7SQ" authority="Schmidt, 1916" authorityName="Schmidt" authorityYear="1916" box="[151,575,1230,1257]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8938648BCDAD86D208FB57" bold="true" box="[151,575,1230,1257]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648BCDAD86D1B1FB56" bold="true" box="[151,390,1230,1257]" italics="true" pageId="12" pageNumber="203">Careproctus mederi</emphasis>
<bibRefCitation id="EFBC4B13FF8938648AD7AD87D208FB57" author="Schmidt, P. J." box="[397,575,1230,1256]" pageId="12" pageNumber="203" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt, 1916</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9236E2FF8938648BCDADBAD271FAB2" blockId="12.[151,582,1230,1329]" box="[151,582,1267,1293]" pageId="12" pageNumber="203">New Japanese name: Agohige-bikunin</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648BCDAC5ED10DFA8E" box="[151,314,1303,1329]" pageId="12" pageNumber="203" type="description">
<paragraph id="8B9236E2FF8938648BCDAC5ED10DFA8E" blockId="12.[151,582,1230,1329]" box="[151,314,1303,1329]" pageId="12" pageNumber="203">
<figureCitation id="13162A67FF8938648BCDAC5ED0ECFA8E" box="[151,219,1303,1329]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="12" pageNumber="203">Fig. 6</figureCitation>
;
<tableCitation id="C6AF0359FF8938648BBDAC5ED10DFA8E" box="[231,314,1303,1329]" captionStart="TABLE 2" captionStartId="13.[151,239,151,175]" captionText="TABLE 2. Counts and measurements of Careproctus mederi, C. sp. cf. gilberti, and C. gilberti." pageId="12" pageNumber="203">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648BCDAC15D2E8FA0F" pageId="12" pageNumber="203" type="reference_group">
<paragraph id="8B9236E2FF8938648BCDAC15D2E8FA0F" blockId="12.[151,1436,1371,1457]" pageId="12" pageNumber="203">
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<treatmentCitation id="0A8C10F3FF8938648BCDAC15D21CFACC" author="Schmidt, P. J." box="[151,555,1371,1395]" page="628" pageId="12" pageNumber="203" year="1916">
<taxonomicName id="4C2D4D61FF8938648BCDAC15D21CFACC" ID-CoL="R7SQ" authority="Schmidt 1916: 628" authorityName="Schmidt" authorityPageNumber="628" authorityYear="1916" box="[151,555,1371,1395]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8938648BCDAC15D157FACC" box="[151,352,1371,1395]" italics="true" pageId="12" pageNumber="203">Careproctus mederi</emphasis>
<bibRefCitation id="EFBC4B13FF8938648A3DAC12D21CFACC" author="Schmidt, P. J." box="[359,555,1371,1395]" pageId="12" pageNumber="203" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt 1916: 628</bibRefCitation>
</taxonomicName>
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; figs. 7, 8 (original description;
<typeStatus id="54968840FF893864882BAC15D3AAFACB" box="[881,925,1372,1396]" pageId="12" pageNumber="203">type</typeStatus>
locality: Sea of Okhotsk,
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,
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,
<quantity id="4CD59B07FF8938648B9DAC33D13DFA2E" box="[199,266,1402,1426]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="12" pageNumber="203" unit="m" value="322.0">322 m</quantity>
depth);
<treatmentCitation id="0A8C10F3FF8938648A39AC33D21FFA2D" author="Schmidt, P. J." box="[355,552,1402,1426]" page="215" pageId="12" pageNumber="203" year="1950">
<bibRefCitation id="EFBC4B13FF8938648A39AC33D21FFA2D" author="Schmidt, P. J." box="[355,552,1402,1426]" pageId="12" pageNumber="203" pagination="1 - 370" refId="ref19112" refString="Schmidt, P. J. (1950) Fishes of the Sea of Okhotsk. Trudy Tichookeanskogo Komiteta, AkademiJa Nauk S. S. S. R., 6, 1 - 370." type="journal article" year="1950">Schmidt 1950: 215</bibRefCitation>
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(description; off Iony Island, northern Sea of Okhotsk);
<treatmentCitation id="0A8C10F3FF8938648F20AC33D555FA2D" author="Matsubara, K." box="[1146,1378,1402,1426]" page="1193" pageId="12" pageNumber="203" year="1955">
<bibRefCitation id="EFBC4B13FF8938648F20AC33D555FA2D" author="Matsubara, K." box="[1146,1378,1402,1426]" pageId="12" pageNumber="203" refId="ref18245" refString="Matsubara, K. (1955) Fish morphology and hierarchy. Ishizaki-Shoten, Tokyo, 1605 pp, 135 pls." type="book" year="1955">Matsubara 1955: 1193</bibRefCitation>
</treatmentCitation>
(key, apparently based on
<bibRefCitation id="EFBC4B13FF8938648AC2ACD1D21EFA0F" author="Schmidt, P. J." box="[408,553,1432,1456]" pageId="12" pageNumber="203" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt 1916</bibRefCitation>
; Sea of Okhotsk).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648BCDAC97D4C2F9DA" pageId="12" pageNumber="203" type="materials_examined">
<paragraph id="8B9236E2FF8938648BCDAC97D4C2F9DA" blockId="12.[151,1437,1502,1889]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648BCDAC97D1BAFA47" bold="true" box="[151,397,1502,1528]" pageId="12" pageNumber="203">Materials examined.</emphasis>
<materialsCitation id="3B453CBFFF8938648ACFAC96D16CF9A2" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="698" latitude="44.0373" location="Hokkaido" longLatPrecision="40" longitude="145.392" pageId="12" pageNumber="203" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF8938648ACFAC96D1E8FA46" box="[405,479,1503,1529]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="12" pageNumber="203">FAKU</collectionCode>
<quantity id="4CD59B07FF8938648ABDAC97D28AFA46" box="[487,701,1502,1529]" metricMagnitude="5" metricUnit="m" metricValue="1.465721023" pageId="12" pageNumber="203" unit="mm" value="1.465721023E8">146572, 102.3 mm</quantity>
SL, female, off
<collectorName id="26D85334FF8938648822AC96D3F6FA46" box="[888,961,1503,1529]" pageId="12" pageNumber="203">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF8938648897AC96D409FA46" box="[973,1086,1503,1529]" country="Japan" name="Hokkaido" pageId="12" pageNumber="203">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8938648F11AC96D4B9FA46" box="[1099,1166,1503,1529]" name="Japan" pageId="12" pageNumber="203">Japan</collectingCountry>
,
<geoCoordinate id="EE195025FF8938648FC3AC96D523FA46" box="[1177,1300,1503,1529]" degrees="44.0373" direction="north" orientation="latitude" pageId="12" pageNumber="203" precision="5" value="44.0373">44.0373ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8938648E7AAC97D5AFFA46" box="[1312,1432,1502,1529]" degrees="145.392" direction="east" orientation="longitude" pageId="12" pageNumber="203" precision="55" value="145.392">145.392ºE</geoCoordinate>
,
<elevation id="0000D1D1FF8938648BCDAF4BD16CF9A2" box="[151,347,1538,1565]" metricMagnitude="2" metricUnit="m" metricValue="6.975" metricValueMax="8.25" metricValueMin="5.7" pageId="12" pageNumber="203" unit="m" value="697.5" valueMax="825.0" valueMin="570.0">
<quantity id="4CD59B07FF8938648BCDAF4BD122F9A2" box="[151,277,1538,1565]" metricMagnitude="2" metricUnit="m" metricValue="6.975" metricValueMax="8.25" metricValueMin="5.7" pageId="12" pageNumber="203" unit="m" value="697.5" valueMax="825.0" valueMin="570.0">570825 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8938648A26AF4AD5ADF9A2" box="[380,1434,1539,1565]" collectingDate="2018-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="12" pageNumber="203" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF8938648A26AF4AD1B8F9A2" box="[380,399,1539,1565]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="203" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8938648ACFAF4AD213F9A3" box="[405,548,1539,1564]" italics="true" pageId="12" pageNumber="203">
<collectorName id="26D85334FF8938648ACFAF4AD213F9A3" box="[405,548,1539,1564]" pageId="12" pageNumber="203">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF8938648974AF4AD291F9A2" box="[558,678,1539,1565]" pageId="12" pageNumber="203" value="2018-08">
<collectingDate id="EFD7E9CAFF8938648974AF4AD291F9A2" box="[558,678,1539,1565]" pageId="12" pageNumber="203" value="2018-08">Aug. 2018</collectingDate>
</date>
, coll. by
<location id="8EF26039FF893864885DAF4AD381F9A2" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF893864885DAF4AD381F9A2" box="[775,950,1539,1565]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="12" pageNumber="203" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF89386488E5AF4AD4B3F9A2" box="[959,1156,1539,1565]" pageId="12" pageNumber="203">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF8938648FE1AF4AD5ADF9A2" box="[1211,1434,1539,1565]" pageId="12" pageNumber="203">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
.
<materialsCitation id="3B453CBFFF8938648BCDAF6ED3AAF9FE" box="[151,925,1574,1601]" collectionCode="FAKU" country="Japan" location="Rausu" pageId="12" pageNumber="203" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF8938648BCDAF6ED0D6F9FE" box="[151,225,1575,1601]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="12" pageNumber="203">FAKU</collectionCode>
<quantity id="4CD59B07FF8938648BB1AF6FD1F3F9FE" box="[235,452,1574,1601]" metricMagnitude="5" metricUnit="m" metricValue="1.484751436" pageId="12" pageNumber="203" unit="mm" value="1.484751436E8">148475, 143.6 mm</quantity>
SL, female, off
<location id="8EF26039FF89386489DFAF6ED2F9F9FE" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF89386489DFAF6ED2F9F9FE" box="[645,718,1575,1601]" country="Japan" name="Rausu" pageId="12" pageNumber="203" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8938648981AF6ED37BF9FE" box="[731,844,1575,1601]" country="Japan" name="Hokkaido" pageId="12" pageNumber="203">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8938648800AF6ED3AAF9FE" box="[858,925,1575,1601]" name="Japan" pageId="12" pageNumber="203">Japan</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF89386488F6AF6ED3CBF9DA" collectionCode="FAKU" collectorName="Pacific" country="Japan" elevation="532" latitude="42.7213" location="Hokkaido" longLatPrecision="6" longitude="144.3489" pageId="12" pageNumber="203" specimenCode="FAKU 149138-149139" specimenCount="1" stateProvince="Hokkaido">
<specimenCode id="DB8B9E99FF89386488F6AF6ED48FF9FE" box="[940,1208,1575,1601]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="12" pageNumber="203">FAKU 149138149139</specimenCode>
,
<quantity id="4CD59B07FF8938648F9CAF6FD55CF9FE" box="[1222,1387,1574,1601]" metricMagnitude="-2" metricUnit="m" metricValue="6.615" metricValueMax="7.609999999999999" metricValueMin="5.62" pageId="12" pageNumber="203" unit="mm" value="66.15" valueMax="76.1" valueMin="56.2">56.276.1 mm</quantity>
SL, female and male,
<collectorName id="26D85334FF8938648A04AF02D19BF9DA" box="[350,428,1611,1637]" pageId="12" pageNumber="203">Pacific</collectorName>
off
<collectingRegion id="49E9F800FF8938648A80AF02D27CF9DA" box="[474,587,1611,1637]" country="Japan" name="Hokkaido" pageId="12" pageNumber="203">Hokkaido</collectingRegion>
,
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,
<geoCoordinate id="EE195025FF8938648986AF02D355F9DA" box="[732,866,1611,1637]" degrees="144.3489" direction="east" orientation="longitude" pageId="12" pageNumber="203" precision="5" value="144.3489">144.3489ºE</geoCoordinate>
,
<elevation id="0000D1D1FF8938648837AF03D3CBF9DA" box="[877,1020,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="12" pageNumber="203" unit="m" value="532.0">
<quantity id="4CD59B07FF8938648837AF03D383F9DA" box="[877,948,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="12" pageNumber="203" unit="m" value="532.0">532 m</quantity>
depth
</elevation>
</materialsCitation>
,
<materialsCitation id="3B453CBFFF8938648F5DAF02D4D9F9DA" box="[1031,1262,1611,1637]" collectionCode="R, V" country="Japan" location="Wakataka-maru" pageId="12" pageNumber="203" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF8938648F5DAF02D42EF9DA" box="[1031,1049,1611,1637]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="203">R</collectionCode>
/
<collectionCode id="ED3CAE27FF8938648F78AF02D402F9DA" box="[1058,1077,1611,1637]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="203" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8938648F66AF02D4D9F9DA" box="[1084,1262,1611,1637]" italics="true" pageId="12" pageNumber="203">
<location id="8EF26039FF8938648F66AF02D4D9F9DA" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF8938648F66AF02D4D9F9DA" box="[1084,1262,1611,1637]" country="Japan" name="Wakataka-maru" pageId="12" pageNumber="203" stateProvince="Hokkaido">Wakataka-maru</location>
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648B9DAF27D563F882" pageId="12" pageNumber="203" type="diagnosis">
<paragraph id="8B9236E2FF8938648B9DAF27D563F882" blockId="12.[151,1437,1502,1889]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DAF27D175F937" bold="true" box="[199,322,1646,1672]" pageId="12" pageNumber="203">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2D4D61FF8938648ABAAF26D25DF936" authorityName="Kroyer" authorityYear="1862" box="[480,618,1647,1673]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF8938648ABAAF26D25DF936" box="[480,618,1647,1673]" italics="true" pageId="12" pageNumber="203">Careproctus</emphasis>
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with the following combination of characters: vertebrae 5763; dorsal-fin rays 5256; anal-fin rays 4752; pectoral-fin rays 2732; pectoral fin deeply and broadly notched, lower lobe short, reaching to midway between its base and anus; lower part of pectoral girdle protruding anteroventrally below mandible; pelvic disk rounded, greatly reduced; cephalic pores 2-6-7-2, chin pores paired in separate pits; gill slit moderately large, extending ventrally to pectoral-fin rays 611; teeth simple or slightly shouldered; anus located below posterior rim of orbit; peritoneum pale with black dots, stomach black (
<bibRefCitation id="EFBC4B13FF8938648F58AE6AD494F882" author="Schmidt, P. J." box="[1026,1187,1827,1853]" pageId="12" pageNumber="203" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt 1916</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938678B9DAE0FD40AFEDA" lastPageId="15" lastPageNumber="206" pageId="12" pageNumber="203" type="description">
<paragraph id="8B9236E2FF8938648B9DAE0FD452F8DE" blockId="12.[151,1437,1502,1889]" box="[199,1125,1862,1889]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DAE0FD25CF8DF" bold="true" box="[199,619,1862,1889]" pageId="12" pageNumber="203">Description of Japanese specimens.</emphasis>
Counts and measurements shown in
<tableCitation id="C6AF0359FF8938648F51AE0ED456F8DE" box="[1035,1121,1863,1889]" captionStart="TABLE 2" captionStartId="13.[151,239,151,175]" captionText="TABLE 2. Counts and measurements of Careproctus mederi, C. sp. cf. gilberti, and C. gilberti." pageId="12" pageNumber="203">Table 2</tableCitation>
.
</paragraph>
<caption id="DF52666AFF8838658BCDA9DED4A5FF10" box="[151,1170,151,176]" pageId="13" pageNumber="204" startId="13.[151,239,151,175]" targetType="table">
<paragraph id="8B9236E2FF8838658BCDA9DED4A5FF10" blockId="13.[151,1170,151,176]" box="[151,1170,151,176]" pageId="13" pageNumber="204">
<emphasis id="B959EAF0FF8838658BCDA9DED13DFF10" bold="true" box="[151,266,151,175]" pageId="13" pageNumber="204">TABLE 2.</emphasis>
Counts and measurements of
<taxonomicName id="4C2D4D61FF8838658916A9D1D317FF10" authorityName="Schmidt" authorityYear="1916" box="[588,800,151,175]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8838658916A9D1D317FF10" box="[588,800,151,175]" italics="true" pageId="13" pageNumber="204">Careproctus mederi</emphasis>
</taxonomicName>
,
<emphasis id="B959EAF0FF8838658874A9D1D370FF10" box="[814,839,152,175]" italics="true" pageId="13" pageNumber="204">C.</emphasis>
sp.
<taxonomicName id="4C2D4D61FF8838658828A9DED3D4FF10" box="[882,995,151,175]" family="Liparidae" genus="Careproctus" isUncertain="true" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="gilberti">
cf.
<emphasis id="B959EAF0FF88386588CEA9DED3D4FF10" box="[916,995,151,175]" italics="true" pageId="13" pageNumber="204">gilberti</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2D4D61FF8838658F47A9D1D4BCFF10" box="[1053,1163,151,175]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF8838658F47A9D1D4BCFF10" box="[1053,1163,151,175]" italics="true" pageId="13" pageNumber="204">C. gilberti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9236E2FF8838658964A98BD5BCF991" pageId="13" pageNumber="204">
<table id="F92DC442FF88C7978BC5A98BD5BCF991" box="[159,1419,194,1582]" gridcols="5" gridrows="33" pageId="13" pageNumber="204">
<tr id="351D34A0FF88C7978BC5A98BD5BCFF67" box="[159,1419,194,216]" gridrow="0" pageId="13" pageNumber="204" rowspan-0="1" rowspan-2="1">
<th id="76CC5DDCFF88C7978964A98BD2CCFF67" box="[574,763,194,216]" gridcol="1" gridrow="0" pageId="13" pageNumber="204">
<taxonomicName id="4C2D4D61FF8838658964A98BD2CCFF67" authorityName="Schmidt" authorityYear="1916" box="[574,763,194,216]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="mederi">Careproctus mederi</taxonomicName>
</th>
<th id="76CC5DDCFF88C7978F77A98BD483FF67" box="[1069,1204,194,216]" gridcol="3" gridrow="0" pageId="13" pageNumber="204">
<taxonomicName id="4C2D4D61FF8838658F77A98AD494FF67" authorityName="Kroyer" authorityYear="1862" box="[1069,1187,195,216]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF8838658F77A98AD494FF67" box="[1069,1187,195,216]" italics="true" pageId="13" pageNumber="204">Careproctus</emphasis>
</taxonomicName>
</th>
<th id="76CC5DDCFF88C7978FB0A98BD5BCFF67" box="[1258,1419,194,216]" gridcol="4" gridrow="0" pageId="13" pageNumber="204">
<taxonomicName id="4C2D4D61FF8838658FB0A98AD557FF67" authority="Japanese" authorityName="Japanese" box="[1258,1376,195,216]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF8838658FB0A98AD557FF67" box="[1258,1376,195,216]" italics="true" pageId="13" pageNumber="204">Careproctus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="351D34A0FF88C7978BC5A9A4D5BCFEBC" box="[159,1419,237,259]" gridrow="1" pageId="13" pageNumber="204" rowspan-0="1">
<td id="76CC5DDCFF88C7978964A9A4D2CCFEBC" box="[574,763,237,259]" gridcol="1" gridrow="1" pageId="13" pageNumber="204">Japanese</td>
<td id="76CC5DDCFF88C7978860A9A4D3C6FEBC" box="[826,1009,237,259]" gridcol="2" gridrow="1" pageId="13" pageNumber="204">Syntypes</td>
<td id="76CC5DDCFF88C7978F77A9A4D483FEBC" box="[1069,1204,237,259]" gridcol="3" gridrow="1" pageId="13" pageNumber="204">
sp.
<taxonomicName id="4C2D4D61FF8838658F14A9A4D483FEBD" box="[1102,1204,237,259]" family="Liparidae" genus="Careproctus" isUncertain="true" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="gilberti">
cf.
<emphasis id="B959EAF0FF8838658F37A9A4D483FEBD" box="[1133,1204,237,258]" italics="true" pageId="13" pageNumber="204">gilberti</emphasis>
</taxonomicName>
</td>
<td id="76CC5DDCFF88C7978FB0A9A4D5BCFEBC" box="[1258,1419,237,259]" gridcol="4" gridrow="1" pageId="13" pageNumber="204">
<taxonomicName id="4C2D4D61FF8838658FB0A9A4D506FEBD" box="[1258,1329,237,258]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF8838658FB0A9A4D506FEBD" box="[1258,1329,237,258]" italics="true" pageId="13" pageNumber="204">gilberti</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="351D34A0FF88C7978BC5A851D5BCFE91" box="[159,1419,280,302]" gridrow="2" pageId="13" pageNumber="204" rowspan-0="1" rowspan-4="1">
<td id="76CC5DDCFF88C7978964A851D2CCFE91" box="[574,763,280,302]" gridcol="1" gridrow="2" pageId="13" pageNumber="204">specimens</td>
<td id="76CC5DDCFF88C7978860A851D3C6FE91" box="[826,1009,280,302]" gridcol="2" gridrow="2" pageId="13" pageNumber="204">
<emphasis id="B959EAF0FF8838658860A851D371FE92" box="[826,838,280,301]" italics="true" pageId="13" pageNumber="204">n</emphasis>
= 4
</td>
<td id="76CC5DDCFF88C7978F77A851D483FE91" box="[1069,1204,280,302]" gridcol="3" gridrow="2" pageId="13" pageNumber="204">FAKU</td>
</tr>
<tr id="351D34A0FF88C7978BC5A80BD5BCFEE7" box="[159,1419,322,344]" gridrow="3" pageId="13" pageNumber="204" rowspan-0="1">
<td id="76CC5DDCFF88C7978964A80BD2CCFEE7" box="[574,763,322,344]" gridcol="1" gridrow="3" pageId="13" pageNumber="204">
<emphasis id="B959EAF0FF8838658964A80AD27DFEE7" box="[574,586,323,344]" italics="true" pageId="13" pageNumber="204">n</emphasis>
= 4
</td>
<td id="76CC5DDCFF88C7978860A80BD3C6FEE7" box="[826,1009,322,344]" gridcol="2" gridrow="3" pageId="13" pageNumber="204">
(
<bibRefCitation id="EFBC4B13FF883865881AA80BD3FEFEE7" author="Schmidt, P. J." box="[832,969,322,344]" pageId="13" pageNumber="204" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt 1916</bibRefCitation>
)
</td>
<td id="76CC5DDCFF88C7978F77A80BD483FEE7" box="[1069,1204,322,344]" gridcol="3" gridrow="3" pageId="13" pageNumber="204">147753***</td>
<td id="76CC5DDCFF88C7978FB0A80BD5BCFEE7" box="[1258,1419,322,344]" gridcol="4" gridrow="3" pageId="13" pageNumber="204">
<emphasis id="B959EAF0FF8838658FB0A80AD4C1FEE7" box="[1258,1270,323,344]" italics="true" pageId="13" pageNumber="204">n</emphasis>
= 7****
</td>
</tr>
<tr id="351D34A0FF88C7978BC5A824D5BCFE3C" box="[159,1419,365,387]" gridrow="4" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5A824D1C2FE3C" box="[159,501,365,387]" gridcol="0" gridrow="4" pageId="13" pageNumber="204">Standard length (mm)</th>
<td id="76CC5DDCFF88C7978964A824D2CCFE3C" box="[574,763,365,387]" gridcol="1" gridrow="4" pageId="13" pageNumber="204">56.2143.6</td>
<td id="76CC5DDCFF88C7978860A824D3C6FE3C" box="[826,1009,365,387]" gridcol="2" gridrow="4" pageId="13" pageNumber="204">108152</td>
<td id="76CC5DDCFF88C7978F77A824D483FE3C" box="[1069,1204,365,387]" gridcol="3" gridrow="4" pageId="13" pageNumber="204">55.1</td>
<td id="76CC5DDCFF88C7978FB0A824D5BCFE3C" box="[1258,1419,365,387]" gridcol="4" gridrow="4" pageId="13" pageNumber="204">104.4224.8</td>
</tr>
<tr id="351D34A0FF88C7978BC5A8D1D5BCFE12" box="[159,1419,408,429]" gridrow="5" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5A8D1D5BCFE12" box="[159,1419,408,429]" colspan="5" colspanRight="4" gridcol="0" gridrow="5" pageId="13" pageNumber="204">Counts</th>
</tr>
<tr id="351D34A0FF88C7978BC5A88BD5BCFE67" box="[159,1419,450,472]" gridrow="6" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5A88BD1C2FE67" box="[159,501,450,472]" gridcol="0" gridrow="6" pageId="13" pageNumber="204">Dorsal-fin rays</th>
<td id="76CC5DDCFF88C7978964A88BD2CCFE67" box="[574,763,450,472]" gridcol="1" gridrow="6" pageId="13" pageNumber="204">5556</td>
<td id="76CC5DDCFF88C7978860A88BD3C6FE67" box="[826,1009,450,472]" gridcol="2" gridrow="6" pageId="13" pageNumber="204">5253*</td>
<td id="76CC5DDCFF88C7978F77A88BD483FE67" box="[1069,1204,450,472]" gridcol="3" gridrow="6" pageId="13" pageNumber="204">57</td>
<td id="76CC5DDCFF88C7978FB0A88BD5BCFE67" box="[1258,1419,450,472]" gridcol="4" gridrow="6" pageId="13" pageNumber="204">5155</td>
</tr>
<tr id="351D34A0FF88C7978BC5A8A4D5BCFDBC" box="[159,1419,493,515]" gridrow="7" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5A8A4D1C2FDBC" box="[159,501,493,515]" gridcol="0" gridrow="7" pageId="13" pageNumber="204">Anal-fin rays</th>
<td id="76CC5DDCFF88C7978964A8A4D2CCFDBC" box="[574,763,493,515]" gridcol="1" gridrow="7" pageId="13" pageNumber="204">5052</td>
<td id="76CC5DDCFF88C7978860A8A4D3C6FDBC" box="[826,1009,493,515]" gridcol="2" gridrow="7" pageId="13" pageNumber="204">4749*</td>
<td id="76CC5DDCFF88C7978F77A8A4D483FDBC" box="[1069,1204,493,515]" gridcol="3" gridrow="7" pageId="13" pageNumber="204">53</td>
<td id="76CC5DDCFF88C7978FB0A8A4D5BCFDBC" box="[1258,1419,493,515]" gridcol="4" gridrow="7" pageId="13" pageNumber="204">4648</td>
</tr>
<tr id="351D34A0FF88C7978BC5AB51D5BCFD91" box="[159,1419,536,558]" gridrow="8" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AB51D1C2FD91" box="[159,501,536,558]" gridcol="0" gridrow="8" pageId="13" pageNumber="204">Pectoral-fin rays</th>
<td id="76CC5DDCFF88C7978964AB51D2CCFD91" box="[574,763,536,558]" gridcol="1" gridrow="8" pageId="13" pageNumber="204">3032</td>
<td id="76CC5DDCFF88C7978860AB51D3C6FD91" box="[826,1009,536,558]" gridcol="2" gridrow="8" pageId="13" pageNumber="204">2729</td>
<td id="76CC5DDCFF88C7978F77AB51D483FD91" box="[1069,1204,536,558]" gridcol="3" gridrow="8" pageId="13" pageNumber="204">33</td>
<td id="76CC5DDCFF88C7978FB0AB51D5BCFD91" box="[1258,1419,536,558]" gridcol="4" gridrow="8" pageId="13" pageNumber="204">3035</td>
</tr>
<tr id="351D34A0FF88C7978BC5AB0BD5BCFDE7" box="[159,1419,578,600]" gridrow="9" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AB0BD1C2FDE7" box="[159,501,578,600]" gridcol="0" gridrow="9" pageId="13" pageNumber="204">Caudal-fin rays</th>
<td id="76CC5DDCFF88C7978964AB0BD2CCFDE7" box="[574,763,578,600]" gridcol="1" gridrow="9" pageId="13" pageNumber="204">0+4/45+01</td>
<td id="76CC5DDCFF88C7978860AB0BD3C6FDE7" box="[826,1009,578,600]" gridcol="2" gridrow="9" pageId="13" pageNumber="204">0+4/5+0*</td>
<td id="76CC5DDCFF88C7978F77AB0BD483FDE7" box="[1069,1204,578,600]" gridcol="3" gridrow="9" pageId="13" pageNumber="204">1+4/4+1</td>
<td id="76CC5DDCFF88C7978FB0AB0BD5BCFDE7" box="[1258,1419,578,600]" gridcol="4" gridrow="9" pageId="13" pageNumber="204">0+45/5+0</td>
</tr>
<tr id="351D34A0FF88C7978BC5AB24D5BCFD3C" box="[159,1419,621,643]" gridrow="10" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AB24D1C2FD3C" box="[159,501,621,643]" gridcol="0" gridrow="10" pageId="13" pageNumber="204">Precaudal vertebrae</th>
<td id="76CC5DDCFF88C7978964AB24D2CCFD3C" box="[574,763,621,643]" gridcol="1" gridrow="10" pageId="13" pageNumber="204">910</td>
<td id="76CC5DDCFF88C7978860AB24D3C6FD3C" box="[826,1009,621,643]" gridcol="2" gridrow="10" pageId="13" pageNumber="204">9*</td>
<td id="76CC5DDCFF88C7978F77AB24D483FD3C" box="[1069,1204,621,643]" gridcol="3" gridrow="10" pageId="13" pageNumber="204">9</td>
<td id="76CC5DDCFF88C7978FB0AB24D5BCFD3C" box="[1258,1419,621,643]" gridcol="4" gridrow="10" pageId="13" pageNumber="204">10</td>
</tr>
<tr id="351D34A0FF88C7978BC5ABD1D5BCFD11" box="[159,1419,664,686]" gridrow="11" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ABD1D1C2FD11" box="[159,501,664,686]" gridcol="0" gridrow="11" pageId="13" pageNumber="204">Caudal vertebrae</th>
<td id="76CC5DDCFF88C7978964ABD1D2CCFD11" box="[574,763,664,686]" gridcol="1" gridrow="11" pageId="13" pageNumber="204">5153</td>
<td id="76CC5DDCFF88C7978860ABD1D3C6FD11" box="[826,1009,664,686]" gridcol="2" gridrow="11" pageId="13" pageNumber="204">4850*</td>
<td id="76CC5DDCFF88C7978F77ABD1D483FD11" box="[1069,1204,664,686]" gridcol="3" gridrow="11" pageId="13" pageNumber="204">54</td>
<td id="76CC5DDCFF88C7978FB0ABD1D5BCFD11" box="[1258,1419,664,686]" gridcol="4" gridrow="11" pageId="13" pageNumber="204">4748</td>
</tr>
<tr id="351D34A0FF88C7978BC5AB8BD5BCFD67" box="[159,1419,706,728]" gridrow="12" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AB8BD1C2FD67" box="[159,501,706,728]" gridcol="0" gridrow="12" pageId="13" pageNumber="204">Total vertebrae</th>
<td id="76CC5DDCFF88C7978964AB8BD2CCFD67" box="[574,763,706,728]" gridcol="1" gridrow="12" pageId="13" pageNumber="204">6163</td>
<td id="76CC5DDCFF88C7978860AB8BD3C6FD67" box="[826,1009,706,728]" gridcol="2" gridrow="12" pageId="13" pageNumber="204">5759*</td>
<td id="76CC5DDCFF88C7978F77AB8BD483FD67" box="[1069,1204,706,728]" gridcol="3" gridrow="12" pageId="13" pageNumber="204">63</td>
<td id="76CC5DDCFF88C7978FB0AB8BD5BCFD67" box="[1258,1419,706,728]" gridcol="4" gridrow="12" pageId="13" pageNumber="204">5758</td>
</tr>
<tr id="351D34A0FF88C7978BC5ABA4D5BCFCBC" box="[159,1419,749,771]" gridrow="13" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ABA4D1C2FCBC" box="[159,501,749,771]" gridcol="0" gridrow="13" pageId="13" pageNumber="204">Pyloric caeca</th>
<td id="76CC5DDCFF88C7978964ABA4D2CCFCBC" box="[574,763,749,771]" gridcol="1" gridrow="13" pageId="13" pageNumber="204">912</td>
<td id="76CC5DDCFF88C7978860ABA4D3C6FCBC" box="[826,1009,749,771]" gridcol="2" gridrow="13" pageId="13" pageNumber="204">15</td>
<td id="76CC5DDCFF88C7978F77ABA4D483FCBC" box="[1069,1204,749,771]" gridcol="3" gridrow="13" pageId="13" pageNumber="204">16</td>
<td id="76CC5DDCFF88C7978FB0ABA4D5BCFCBC" box="[1258,1419,749,771]" gridcol="4" gridrow="13" pageId="13" pageNumber="204">610</td>
</tr>
<tr id="351D34A0FF88C7978BC5AA51D5BCFC91" box="[159,1419,792,814]" gridrow="14" pageId="13" pageNumber="204" rowspan-3="1">
<th id="76CC5DDCFF88C7978BC5AA51D1C2FC91" box="[159,501,792,814]" gridcol="0" gridrow="14" pageId="13" pageNumber="204">In % of standard length</th>
<td id="76CC5DDCFF88C7978964AA51D2CCFC91" box="[574,763,792,814]" gridcol="1" gridrow="14" pageId="13" pageNumber="204">means (ranges)</td>
<td id="76CC5DDCFF88C7978860AA51D3C6FC91" box="[826,1009,792,814]" gridcol="2" gridrow="14" pageId="13" pageNumber="204">means (ranges)</td>
<td id="76CC5DDCFF88C7978FB0AA51D5BCFC91" box="[1258,1419,792,814]" gridcol="4" gridrow="14" pageId="13" pageNumber="204">means (ranges)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AA0BD5BCFCE7" box="[159,1419,834,856]" gridrow="15" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AA0BD1C2FCE7" box="[159,501,834,856]" gridcol="0" gridrow="15" pageId="13" pageNumber="204">Head length</th>
<td id="76CC5DDCFF88C7978964AA0BD2CCFCE7" box="[574,763,834,856]" gridcol="1" gridrow="15" pageId="13" pageNumber="204">27.6 (24.829.4)</td>
<td id="76CC5DDCFF88C7978860AA0BD3C6FCE7" box="[826,1009,834,856]" gridcol="2" gridrow="15" pageId="13" pageNumber="204">26.0 (23.827.3)</td>
<td id="76CC5DDCFF88C7978F77AA0BD483FCE7" box="[1069,1204,834,856]" gridcol="3" gridrow="15" pageId="13" pageNumber="204">29.3</td>
<td id="76CC5DDCFF88C7978FB0AA0BD5BCFCE7" box="[1258,1419,834,856]" gridcol="4" gridrow="15" pageId="13" pageNumber="204">27.7 (25.928.7)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AA24D5BCFC3C" box="[159,1419,877,899]" gridrow="16" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AA24D1C2FC3C" box="[159,501,877,899]" gridcol="0" gridrow="16" pageId="13" pageNumber="204">Snout length</th>
<td id="76CC5DDCFF88C7978964AA24D2CCFC3C" box="[574,763,877,899]" gridcol="1" gridrow="16" pageId="13" pageNumber="204">9.2 (8.210.1)</td>
<td id="76CC5DDCFF88C7978860AA24D3C6FC3C" box="[826,1009,877,899]" gridcol="2" gridrow="16" pageId="13" pageNumber="204">7.4 (6.88.0)</td>
<td id="76CC5DDCFF88C7978F77AA24D483FC3C" box="[1069,1204,877,899]" gridcol="3" gridrow="16" pageId="13" pageNumber="204">9.8</td>
<td id="76CC5DDCFF88C7978FB0AA24D5BCFC3C" box="[1258,1419,877,899]" gridcol="4" gridrow="16" pageId="13" pageNumber="204">8.7 (7.29.5)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AAD1D5BCFC11" box="[159,1419,920,942]" gridrow="17" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AAD1D1C2FC11" box="[159,501,920,942]" gridcol="0" gridrow="17" pageId="13" pageNumber="204">Orbit length</th>
<td id="76CC5DDCFF88C7978964AAD1D2CCFC11" box="[574,763,920,942]" gridcol="1" gridrow="17" pageId="13" pageNumber="204">7.9 (6.69.0)</td>
<td id="76CC5DDCFF88C7978860AAD1D3C6FC11" box="[826,1009,920,942]" gridcol="2" gridrow="17" pageId="13" pageNumber="204">7.0 (6.28.0)</td>
<td id="76CC5DDCFF88C7978F77AAD1D483FC11" box="[1069,1204,920,942]" gridcol="3" gridrow="17" pageId="13" pageNumber="204">6.9</td>
<td id="76CC5DDCFF88C7978FB0AAD1D5BCFC11" box="[1258,1419,920,942]" gridcol="4" gridrow="17" pageId="13" pageNumber="204">7.5 (6.58.8)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AA8BD5BCFC67" box="[159,1419,962,984]" gridrow="18" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AA8BD1C2FC67" box="[159,501,962,984]" gridcol="0" gridrow="18" pageId="13" pageNumber="204">Interorbital width</th>
<td id="76CC5DDCFF88C7978964AA8BD2CCFC67" box="[574,763,962,984]" gridcol="1" gridrow="18" pageId="13" pageNumber="204">10.8 (10.312.0)</td>
<td id="76CC5DDCFF88C7978860AA8BD3C6FC67" box="[826,1009,962,984]" gridcol="2" gridrow="18" pageId="13" pageNumber="204">13.6 (12.514.9)</td>
<td id="76CC5DDCFF88C7978F77AA8BD483FC67" box="[1069,1204,962,984]" gridcol="3" gridrow="18" pageId="13" pageNumber="204">11.3</td>
<td id="76CC5DDCFF88C7978FB0AA8BD5BCFC67" box="[1258,1419,962,984]" gridcol="4" gridrow="18" pageId="13" pageNumber="204">9.1 (7.910.4)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AAA4D5BCFBBC" box="[159,1419,1005,1027]" gridrow="19" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AAA4D1C2FBBC" box="[159,501,1005,1027]" gridcol="0" gridrow="19" pageId="13" pageNumber="204">Maxilla length</th>
<td id="76CC5DDCFF88C7978964AAA4D2CCFBBC" box="[574,763,1005,1027]" gridcol="1" gridrow="19" pageId="13" pageNumber="204">13.7 (12.415.1)</td>
<td id="76CC5DDCFF88C7978860AAA4D3C6FBBC" box="[826,1009,1005,1027]" gridcol="2" gridrow="19" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AAA4D483FBBC" box="[1069,1204,1005,1027]" gridcol="3" gridrow="19" pageId="13" pageNumber="204">16.0</td>
<td id="76CC5DDCFF88C7978FB0AAA4D5BCFBBC" box="[1258,1419,1005,1027]" gridcol="4" gridrow="19" pageId="13" pageNumber="204">14.1 (13.215.4)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AD51D5BCFB91" box="[159,1419,1048,1070]" gridrow="20" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AD51D1C2FB91" box="[159,501,1048,1070]" gridcol="0" gridrow="20" pageId="13" pageNumber="204">Gill slit length</th>
<td id="76CC5DDCFF88C7978964AD51D2CCFB91" box="[574,763,1048,1070]" gridcol="1" gridrow="20" pageId="13" pageNumber="204">12.6 (10.614.5)</td>
<td id="76CC5DDCFF88C7978860AD51D3C6FB91" box="[826,1009,1048,1070]" gridcol="2" gridrow="20" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AD51D483FB91" box="[1069,1204,1048,1070]" gridcol="3" gridrow="20" pageId="13" pageNumber="204">8.9</td>
<td id="76CC5DDCFF88C7978FB0AD51D5BCFB91" box="[1258,1419,1048,1070]" gridcol="4" gridrow="20" pageId="13" pageNumber="204">10.5 (9.611.5)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AD0BD5BCFBE7" box="[159,1419,1090,1112]" gridrow="21" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AD0BD1C2FBE7" box="[159,501,1090,1112]" gridcol="0" gridrow="21" pageId="13" pageNumber="204">Body depth at pelvic disk</th>
<td id="76CC5DDCFF88C7978964AD0BD2CCFBE7" box="[574,763,1090,1112]" gridcol="1" gridrow="21" pageId="13" pageNumber="204">22.6 (18.829.4)</td>
<td id="76CC5DDCFF88C7978860AD0BD3C6FBE7" box="[826,1009,1090,1112]" gridcol="2" gridrow="21" pageId="13" pageNumber="204">30.9 (27.834.0)**</td>
<td id="76CC5DDCFF88C7978F77AD0BD483FBE7" box="[1069,1204,1090,1112]" gridcol="3" gridrow="21" pageId="13" pageNumber="204">16.9</td>
<td id="76CC5DDCFF88C7978FB0AD0BD5BCFBE7" box="[1258,1419,1090,1112]" gridcol="4" gridrow="21" pageId="13" pageNumber="204">21.5 (18.823.5)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AD24D5BCFB3C" box="[159,1419,1133,1155]" gridrow="22" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AD24D1C2FB3C" box="[159,501,1133,1155]" gridcol="0" gridrow="22" pageId="13" pageNumber="204">Pectoral fin length</th>
<td id="76CC5DDCFF88C7978964AD24D2CCFB3C" box="[574,763,1133,1155]" gridcol="1" gridrow="22" pageId="13" pageNumber="204">14.9 (12.318.1)</td>
<td id="76CC5DDCFF88C7978860AD24D3C6FB3C" box="[826,1009,1133,1155]" gridcol="2" gridrow="22" pageId="13" pageNumber="204">15.6 (14.016.5)</td>
<td id="76CC5DDCFF88C7978F77AD24D483FB3C" box="[1069,1204,1133,1155]" gridcol="3" gridrow="22" pageId="13" pageNumber="204">13.5</td>
<td id="76CC5DDCFF88C7978FB0AD24D5BCFB3C" box="[1258,1419,1133,1155]" gridcol="4" gridrow="22" pageId="13" pageNumber="204">14.2 (12.815.6)</td>
</tr>
<tr id="351D34A0FF88C7978BC5ADD1D5BCFB11" box="[159,1419,1176,1198]" gridrow="23" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ADD1D1C2FB11" box="[159,501,1176,1198]" gridcol="0" gridrow="23" pageId="13" pageNumber="204">Pectoral-fin lower lobe length</th>
<td id="76CC5DDCFF88C7978964ADD1D2CCFB11" box="[574,763,1176,1198]" gridcol="1" gridrow="23" pageId="13" pageNumber="204">5.4 (4.55.7)</td>
<td id="76CC5DDCFF88C7978860ADD1D3C6FB11" box="[826,1009,1176,1198]" gridcol="2" gridrow="23" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77ADD1D483FB11" box="[1069,1204,1176,1198]" gridcol="3" gridrow="23" pageId="13" pageNumber="204">14.4</td>
<td id="76CC5DDCFF88C7978FB0ADD1D5BCFB11" box="[1258,1419,1176,1198]" gridcol="4" gridrow="23" pageId="13" pageNumber="204">23.7 (20.427.0)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AD8BD5BCFB67" box="[159,1419,1218,1240]" gridrow="24" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AD8BD1C2FB67" box="[159,501,1218,1240]" gridcol="0" gridrow="24" pageId="13" pageNumber="204">Pectoral-fin notch-ray length</th>
<td id="76CC5DDCFF88C7978964AD8BD2CCFB67" box="[574,763,1218,1240]" gridcol="1" gridrow="24" pageId="13" pageNumber="204">1.8 (1.52.1)</td>
<td id="76CC5DDCFF88C7978860AD8BD3C6FB67" box="[826,1009,1218,1240]" gridcol="2" gridrow="24" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AD8BD483FB67" box="[1069,1204,1218,1240]" gridcol="3" gridrow="24" pageId="13" pageNumber="204">3.8</td>
<td id="76CC5DDCFF88C7978FB0AD8BD5BCFB67" box="[1258,1419,1218,1240]" gridcol="4" gridrow="24" pageId="13" pageNumber="204">4.2 (2.76.3)</td>
</tr>
<tr id="351D34A0FF88C7978BC5ADA4D5BCFABC" box="[159,1419,1261,1283]" gridrow="25" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ADA4D1C2FABC" box="[159,501,1261,1283]" gridcol="0" gridrow="25" pageId="13" pageNumber="204">Predorsal length</th>
<td id="76CC5DDCFF88C7978964ADA4D2CCFABC" box="[574,763,1261,1283]" gridcol="1" gridrow="25" pageId="13" pageNumber="204">28.2 (24.032.4)</td>
<td id="76CC5DDCFF88C7978860ADA4D3C6FABC" box="[826,1009,1261,1283]" gridcol="2" gridrow="25" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77ADA4D483FABC" box="[1069,1204,1261,1283]" gridcol="3" gridrow="25" pageId="13" pageNumber="204">25.5</td>
<td id="76CC5DDCFF88C7978FB0ADA4D5BCFABC" box="[1258,1419,1261,1283]" gridcol="4" gridrow="25" pageId="13" pageNumber="204">27.7 (26.030.1)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AC51D5BCFA91" box="[159,1419,1304,1326]" gridrow="26" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AC51D1C2FA91" box="[159,501,1304,1326]" gridcol="0" gridrow="26" pageId="13" pageNumber="204">Preanal length</th>
<td id="76CC5DDCFF88C7978964AC51D2CCFA91" box="[574,763,1304,1326]" gridcol="1" gridrow="26" pageId="13" pageNumber="204">36.8 (34.042.9)</td>
<td id="76CC5DDCFF88C7978860AC51D3C6FA91" box="[826,1009,1304,1326]" gridcol="2" gridrow="26" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AC51D483FA91" box="[1069,1204,1304,1326]" gridcol="3" gridrow="26" pageId="13" pageNumber="204">33.7</td>
<td id="76CC5DDCFF88C7978FB0AC51D5BCFA91" box="[1258,1419,1304,1326]" gridcol="4" gridrow="26" pageId="13" pageNumber="204">36.8 (34.639.1)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AC0BD5BCFAE7" box="[159,1419,1346,1368]" gridrow="27" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AC0BD1C2FAE7" box="[159,501,1346,1368]" gridcol="0" gridrow="27" pageId="13" pageNumber="204">Snout to anus length</th>
<td id="76CC5DDCFF88C7978964AC0BD2CCFAE7" box="[574,763,1346,1368]" gridcol="1" gridrow="27" pageId="13" pageNumber="204">14.7 (10.918.2)</td>
<td id="76CC5DDCFF88C7978860AC0BD3C6FAE7" box="[826,1009,1346,1368]" gridcol="2" gridrow="27" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AC0BD483FAE7" box="[1069,1204,1346,1368]" gridcol="3" gridrow="27" pageId="13" pageNumber="204">17.8</td>
<td id="76CC5DDCFF88C7978FB0AC0BD5BCFAE7" box="[1258,1419,1346,1368]" gridcol="4" gridrow="27" pageId="13" pageNumber="204">19.4 (17.024.8)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AC24D5BCFA3C" box="[159,1419,1389,1411]" gridrow="28" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AC24D1C2FA3C" box="[159,501,1389,1411]" gridcol="0" gridrow="28" pageId="13" pageNumber="204">Pelvic disk length</th>
<td id="76CC5DDCFF88C7978964AC24D2CCFA3C" box="[574,763,1389,1411]" gridcol="1" gridrow="28" pageId="13" pageNumber="204">1.0 (0.71.4)</td>
<td id="76CC5DDCFF88C7978860AC24D3C6FA3C" box="[826,1009,1389,1411]" gridcol="2" gridrow="28" pageId="13" pageNumber="204">1.1 (1.01.2)</td>
<td id="76CC5DDCFF88C7978F77AC24D483FA3C" box="[1069,1204,1389,1411]" gridcol="3" gridrow="28" pageId="13" pageNumber="204">1.7</td>
<td id="76CC5DDCFF88C7978FB0AC24D5BCFA3C" box="[1258,1419,1389,1411]" gridcol="4" gridrow="28" pageId="13" pageNumber="204">2.3 (1.82.7)</td>
</tr>
<tr id="351D34A0FF88C7978BC5ACD1D5BCFA11" box="[159,1419,1432,1454]" gridrow="29" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ACD1D1C2FA11" box="[159,501,1432,1454]" gridcol="0" gridrow="29" pageId="13" pageNumber="204">Caudal fin length</th>
<td id="76CC5DDCFF88C7978964ACD1D2CCFA11" box="[574,763,1432,1454]" gridcol="1" gridrow="29" pageId="13" pageNumber="204">11.2 (12.213.2)</td>
<td id="76CC5DDCFF88C7978860ACD1D3C6FA11" box="[826,1009,1432,1454]" gridcol="2" gridrow="29" pageId="13" pageNumber="204">11.3 (10.512.5)</td>
<td id="76CC5DDCFF88C7978F77ACD1D483FA11" box="[1069,1204,1432,1454]" gridcol="3" gridrow="29" pageId="13" pageNumber="204">13.6</td>
<td id="76CC5DDCFF88C7978FB0ACD1D5BCFA11" box="[1258,1419,1432,1454]" gridcol="4" gridrow="29" pageId="13" pageNumber="204">11.7 (9.6 12.7)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AC8BD5BCFA67" box="[159,1419,1474,1496]" gridrow="30" pageId="13" pageNumber="204" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76CC5DDCFF88C7978BC5AC8BD1C2FA67" box="[159,501,1474,1496]" gridcol="0" gridrow="30" pageId="13" pageNumber="204">In % of caudal in</th>
</tr>
<tr id="351D34A0FF88C7978BC5ACA4D5BCF9BC" box="[159,1419,1517,1539]" gridrow="31" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5ACA4D1C2F9BC" box="[159,501,1517,1539]" gridcol="0" gridrow="31" pageId="13" pageNumber="204">Dorsal-fin connection to caudal fin</th>
<td id="76CC5DDCFF88C7978964ACA4D2CCF9BC" box="[574,763,1517,1539]" gridcol="1" gridrow="31" pageId="13" pageNumber="204">24.3 (15.631.5)</td>
<td id="76CC5DDCFF88C7978860ACA4D3C6F9BC" box="[826,1009,1517,1539]" gridcol="2" gridrow="31" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77ACA4D483F9BC" box="[1069,1204,1517,1539]" gridcol="3" gridrow="31" pageId="13" pageNumber="204">39.5</td>
<td id="76CC5DDCFF88C7978FB0ACA4D5BCF9BC" box="[1258,1419,1517,1539]" gridcol="4" gridrow="31" pageId="13" pageNumber="204">33.0 (25.938.6)</td>
</tr>
<tr id="351D34A0FF88C7978BC5AF51D5BCF991" box="[159,1419,1560,1582]" gridrow="32" pageId="13" pageNumber="204">
<th id="76CC5DDCFF88C7978BC5AF51D1C2F991" box="[159,501,1560,1582]" gridcol="0" gridrow="32" pageId="13" pageNumber="204">Anal-fin connection to caudal fin</th>
<td id="76CC5DDCFF88C7978964AF51D2CCF991" box="[574,763,1560,1582]" gridcol="1" gridrow="32" pageId="13" pageNumber="204">30.0 (22.939.2)</td>
<td id="76CC5DDCFF88C7978860AF51D3C6F991" box="[826,1009,1560,1582]" gridcol="2" gridrow="32" pageId="13" pageNumber="204">-</td>
<td id="76CC5DDCFF88C7978F77AF51D483F991" box="[1069,1204,1560,1582]" gridcol="3" gridrow="32" pageId="13" pageNumber="204">48.0</td>
<td id="76CC5DDCFF88C7978FB0AF51D5BCF991" box="[1258,1419,1560,1582]" gridcol="4" gridrow="32" pageId="13" pageNumber="204">36.6 (32.542.6)</td>
</tr>
</table>
</paragraph>
<tableNote id="76CB376CFF8838658BCDAF76D36CF97E" pageId="13" pageNumber="204" targetBox="[159,1419,194,1582]" targetPageId="13">
<paragraph id="8B9236E2FF8838658BCDAF76D200F9EA" blockId="13.[151,859,1599,1729]" box="[151,567,1599,1621]" pageId="13" pageNumber="204">* Data taken in this study from radiographs</paragraph>
<paragraph id="8B9236E2FF8838658BCDAF2AD1C0F9C6" blockId="13.[151,859,1599,1729]" box="[151,503,1635,1657]" pageId="13" pageNumber="204">** Body depth at origin of dorsal fin</paragraph>
<paragraph id="8B9236E2FF8838658BCDAFCED1B8F923" blockId="13.[151,859,1599,1729]" box="[151,399,1671,1693]" pageId="13" pageNumber="204">*** Collected from Japan</paragraph>
<paragraph id="8B9236E2FF8838658BCDAFE2D36CF97E" blockId="13.[151,859,1599,1729]" box="[151,859,1707,1729]" pageId="13" pageNumber="204">**** Collected from the Bering Sea, Aleutian Islands, and Gulf of Alaska</paragraph>
</tableNote>
<paragraph id="8B9236E2FF8838668B9DAE5AD28FFE36" blockId="13.[151,1436,1811,2018]" lastBlockId="14.[151,1437,151,393]" lastPageId="14" lastPageNumber="205" pageId="13" pageNumber="204">
Body moderately deep, compressed, tapering posteriorly (
<figureCitation id="13162A67FF8838658813AE5AD395F892" box="[841,930,1811,1837]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="13" pageNumber="204">Fig. 6A</figureCitation>
). Skin relatively thick, covered by cactus-like prickles; gelatinous tissue under skin moderately developed. Head moderately large, dorsal profile gradually sloping from nape to snout. Snout rounded, not protruding anterior to mouth. Mouth moderately large, terminal, horizontal; maxilla extending to mid orbit; oral cleft extending to anterior rim of orbit. Premaxillary teeth sharp, simple, arranged in oblique rows of 58 teeth per row forming relatively narrow band, inner teeth enlarged. Mandibular teeth sharp, simple, arranged in oblique rows of 48 teeth per row forming narrow band, inner teeth not enlarged. Diastema at symphysis of upper jaws V-shaped, wider anteriorly, that of lower jaws narrow (
<figureCitation id="13162A67FF8B38668F72A9DED4B6FF0E" box="[1064,1153,151,177]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="14" pageNumber="205">Fig. 6C</figureCitation>
). Orbit of moderate size, rounded; pupil rounded. Nostril single, with short tube, level with mid-orbit. Cephalic sensory pores of moderate size: nasal pores 2, maxillary pores 6, preoperculomandibular pores 7, suprabranchial pores 2; cephalic pore pattern 2-6-7-2. Chin pores paired in separate pits, well separated. Coronal pore absent. Gill slit moderately large, upper margin level with mid-orbit, extending ventrally to pectoral-fin rays 611. Gill rakers 9 or 10, short, with tiny spines on tips. Opercular flap rounded; tip level with ventral rim of orbit. Pyloric caeca 912, thick, covering half of stomach, on center-left side of visceral cavity.
</paragraph>
<caption id="DF52666AFF8B38668BCDADC2D546FB54" ID-DOI="http://doi.org/10.5281/zenodo.13219224" ID-Zenodo-Dep="13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="14" pageNumber="205" startId="14.[151,250,1163,1187]" targetBox="[190,1397,417,1140]" targetPageId="14" targetType="figure">
<paragraph id="8B9236E2FF8B38668BCDADC2D546FB54" blockId="14.[151,1436,1163,1259]" pageId="14" pageNumber="205">
<emphasis id="B959EAF0FF8B38668BCDADC2D125FB1C" bold="true" box="[151,274,1163,1187]" pageId="14" pageNumber="205">FIGURE 6.</emphasis>
<taxonomicName id="4C2D4D61FF8B38668A43ADC2D206FB1C" authority=", FAKU" authorityName="FAKU" box="[281,561,1163,1187]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="205" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8B38668A43ADC2D1D6FB1C" box="[281,481,1163,1187]" italics="true" pageId="14" pageNumber="205">Careproctus mederi</emphasis>
, FAKU
</taxonomicName>
146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively.
</paragraph>
</caption>
<paragraph id="8B9236E2FF8B38668B9DAC74D272F9EC" blockId="14.[151,1437,1341,2015]" pageId="14" pageNumber="205">
Vertebrae 6163, precaudal 910 and caudal 5153. Pleural ribs in 1 pair on abdominal vertebrae 9 or absent. Dorsal-fin rays 5556, not exserted. Anteriormost dorsal-fin pterygiophore inserted between neural spines 3 and 4 or 4 and 5, bearing a single ray. Anterior rayless pterygiophore present or absent. Anal-fin rays 5052, not exserted. One or two anal-fin pterygiophore anterior to 1
<superScript id="7C589BAAFF8B386689E9ACE0D289FA08" attach="left" box="[691,702,1449,1463]" fontSize="6" pageId="14" pageNumber="205">st</superScript>
haemal spine, each bearing a single ray. Membrane of posterior dorsal- and anal-fin rays attached about equidistant to caudal fin. Caudal fin slightly rounded or truncated. Principal caudal-fin rays 89, 4 and 45 rays on upper and lower hypurals, respectively, dorsal procurrent rays absent, ventral procurrent rays absent or 1. Hypural plates fused with terminal vertebral centrum and parhypural, upper and lower hypural plate separated by narrow slit.
</paragraph>
<paragraph id="8B9236E2FF8B38668B9DAF14D1BDF8F0" blockId="14.[151,1437,1341,2015]" pageId="14" pageNumber="205">
Pectoral fin deeply and broadly notched, with 3032 rays; upper lobe with 1519 rays, reaching to anal-fin ray 14; basal part of 57 rays in notch adhering to body skin, tips free: lower lobe short, with 8 rays, 4
<superScript id="7C589BAAFF8B38668FB3AFC8D4C1F930" attach="left" box="[1257,1270,1665,1679]" fontSize="6" pageId="14" pageNumber="205">th</superScript>
or 5
<superScript id="7C589BAAFF8B38668E72AFC8D502F930" attach="left" box="[1320,1333,1665,1679]" fontSize="6" pageId="14" pageNumber="205">th</superScript>
ray from ventralmost longest, reaching to midway between its base and anus. Rays in notch more widely spaced than rays of lobes. Uppermost pectoral-fin base level with corner of mouth. Lowermost pectoral-fin base below region between snout and anterior rim of orbit, lower part of pectoral girdle protruding anteroventrally below mandible. Pelvic disk rounded, minute, located below mid orbit. Anus much closer to pelvic disk than to anal fin origin, located below posterior rim of orbit.
</paragraph>
<paragraph id="8B9236E2FF8B38678B9DAE10D24CFF6A" blockId="14.[151,1437,1341,2015]" lastBlockId="15.[151,1437,151,1725]" lastPageId="15" lastPageNumber="206" pageId="14" pageNumber="205">
Pectoral girdle (based on FAKU 148478:
<figureCitation id="13162A67FF8B386689CFAE10D2DCF8CC" box="[661,747,1881,1907]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="14" pageNumber="205">Fig. 2E</figureCitation>
): Proximal pectoral radials 4; upper radials 1 and 2 hour-grass shaped with two notches; radial 3 rounded; radials 13 closely positioned to each other; radial 4 rounded squarish, widely separated from radials 13. Interradial fenestrae 3, extending between scapula and radials 12, and below radial 2; fenestra 1 roundish, fenestrae 2 and 3 elliptical, latter small and slit-like. Scapula broadly Y-shaped, with robust helve. Coracoid diamond-shaped, with short thin helve. Distal radials present at base of pectoral-fin rays 2 to 24, absent from base of ventralmost 7 rays.
</paragraph>
<paragraph id="8B9236E2FF8A38678B9DA996D562FEA2" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">Coloration: Fresh coloration unknown. When preserved, head, body, and fins white. Distal part of dorsal and anal fins dark. Eyes black. Oral and branchial cavities pale; peritoneum pale with black dots, stomach black.</paragraph>
<paragraph id="8B9236E2FF8A38678B9DA86FD40AFEDA" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">
<emphasis id="B959EAF0FF8A38678B9DA86FD144FEFF" bold="true" box="[199,371,294,320]" pageId="15" pageNumber="206">Reproduction.</emphasis>
FAKU 146572 (
<quantity id="4CD59B07FF8A3867896EA86ED29EFEFE" box="[564,681,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.023" pageId="15" pageNumber="206" unit="mm" value="102.3">102.3 mm</quantity>
SL) was a female with elliptical yolked eggs, large-axis diameter ranging from 5.666.00 mm and short-axis diameter from
<quantity id="4CD59B07FF8A3867887CA802D3FEFEDA" box="[806,969,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.39" metricValueMin="2.98" pageId="15" pageNumber="206" unit="mm" value="3.185" valueMax="3.39" valueMin="2.98">2.983.39 mm</quantity>
(
<figureCitation id="13162A67FF8A38678882A802D406FEDA" box="[984,1073,331,357]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="15" pageNumber="206">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8A38678B9DA827D0D2FD82" pageId="15" pageNumber="206" type="distribution">
<paragraph id="8B9236E2FF8A38678B9DA827D0D2FD82" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">
<emphasis id="B959EAF0FF8A38678B9DA827D156FE37" bold="true" box="[199,353,366,392]" pageId="15" pageNumber="206">Distribution.</emphasis>
The
<typeStatus id="54968840FF8A38678AC1A826D1CAFE36" box="[411,509,367,393]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
were collected from near Iony Is., northern Sea of Okhotsk. Additional specimens from the same locality were reported by
<bibRefCitation id="EFBC4B13FF8A38678924A8DAD303FE12" author="Schmidt, P. J." box="[638,820,402,429]" pageId="15" pageNumber="206" pagination="1 - 370" refId="ref19112" refString="Schmidt, P. J. (1950) Fishes of the Sea of Okhotsk. Trudy Tichookeanskogo Komiteta, AkademiJa Nauk S. S. S. R., 6, 1 - 370." type="journal article" year="1950">Schmidt (1950)</bibRefCitation>
. Since then, no other records are known, except for
<bibRefCitation id="EFBC4B13FF8A38678BCDA8FED101FE6E" author="Kido, K." box="[151,310,439,465]" pageId="15" pageNumber="206" pagination="125 - 256" refId="ref18070" refString="Kido, K. (1988) Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memories of the Faculty of Fisheries, Hokkaido University, 35, 125 - 256." type="journal article" year="1988">Kidos (1988)</bibRefCitation>
specimens, which are described herein as
<taxonomicName id="4C2D4D61FF8A3867887DA8FED3D0FE6E" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[807,999,439,465]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="longibarbatus" status="sp. nov.">
<emphasis id="B959EAF0FF8A3867887DA8FED3D0FE6E" box="[807,999,439,465]" italics="true" pageId="15" pageNumber="206">C. longibarbatus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF8A386788ABA8FED479FE6E" bold="true" box="[1009,1102,439,465]" pageId="15" pageNumber="206">
<taxonomicNameLabel id="A26A578BFF8A386788ABA8FED479FE6E" box="[1009,1102,439,465]" pageId="15" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The present specimens were collected from Rausu,
<collectingRegion id="49E9F800FF8A38678AC8A892D234FE4A" box="[402,515,475,501]" country="Japan" name="Hokkaido" pageId="15" pageNumber="206">Hokkaido</collectingRegion>
, southern Sea of Okhotsk, and Kushiro, Pacific coast of
<collectingRegion id="49E9F800FF8A38678F28A892D4D4FE4A" box="[1138,1251,475,501]" country="Japan" name="Hokkaido" pageId="15" pageNumber="206">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8A38678FB4A892D506FE4A" box="[1262,1329,475,501]" name="Japan" pageId="15" pageNumber="206">Japan</collectingCountry>
, the latter representing the southernmost record of the species. Known from depths between
<quantity id="4CD59B07FF8A38678F6DA8B7D480FDA6" box="[1079,1207,510,537]" metricMagnitude="2" metricUnit="m" metricValue="5.135" metricValueMax="8.25" metricValueMin="2.02" pageId="15" pageNumber="206" unit="m" value="513.5" valueMax="825.0" valueMin="202.0">202825 m</quantity>
(
<bibRefCitation id="EFBC4B13FF8A38678F9FA8B6D550FDA6" author="Schmidt, P. J." box="[1221,1383,510,537]" pageId="15" pageNumber="206" pagination="1 - 370" refId="ref19112" refString="Schmidt, P. J. (1950) Fishes of the Sea of Okhotsk. Trudy Tichookeanskogo Komiteta, AkademiJa Nauk S. S. S. R., 6, 1 - 370." type="journal article" year="1950">Schmidt 1950</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8A38678B9DAB0ED53BF902" pageId="15" pageNumber="206" type="discussion">
<paragraph id="8B9236E2FF8A38678B9DAB0ED525FC52" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">
<emphasis id="B959EAF0FF8A38678B9DAB0ED10CFDDE" bold="true" box="[199,315,583,609]" pageId="15" pageNumber="206">Remarks.</emphasis>
The
<typeStatus id="54968840FF8A38678A20AB0ED1EBFDDE" box="[378,476,583,609]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C2D4D61FF8A3867895DAB0ED24EFDDE" box="[519,633,583,609]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A3867895DAB0ED24EFDDE" box="[519,633,583,609]" italics="true" pageId="15" pageNumber="206">C. mederi</emphasis>
</taxonomicName>
(ZIN 18958) are characterized by the following characters: vertebrae 5759; dorsal-fin rays 5253; anal-fin rays 4749; pectoral fin deeply and broadly notched, lower lobe short; pelvic disk minute, 1.01.2% of SL; cephalic pores 2-6-7-2, chin pores paired in separate pits; gill slit moderately large, extending ventrally to pectoral-fin rays 1012; body completely covered with cactus-like prickles (
<bibRefCitation id="EFBC4B13FF8A38678FAFABFAD5A0FD72" author="Schmidt, P. J." box="[1269,1431,691,717]" pageId="15" pageNumber="206" pagination="611 - 630" refId="ref19060" refString="Schmidt, P. J. (1916) Ichthyological notes. 1. On some new and little known Cottidae of North Pacific. 2. On a new cyclogasterid fish with a rudimentary ventral disk. Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk, 20, 611 - 630." type="journal article" year="1916">Schmidt 1916</bibRefCitation>
; this study) (
<figureCitation id="13162A67FF8A38678A43AB9ED1A6FD4E" box="[281,401,727,753]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="15" pageNumber="206">Fig. 6DF</figureCitation>
). The present specimens from the southern Sea of Okhotsk agreed with the
<typeStatus id="54968840FF8A38678F97AB9ED518FD4E" box="[1229,1327,727,753]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
in having a small pelvic disk 0.71.4% of SL, a short pectoral fin lower lobe, not reaching the anus, and the body covered with cactus-like prickles. In addition, both the
<typeStatus id="54968840FF8A386789F6AA56D339FC86" box="[684,782,799,825]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
and present specimens had the pectoral girdle protruding anteroventrally below the mandible (
<figureCitation id="13162A67FF8A38678919AA0AD2BCFCE2" box="[579,651,835,861]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="15" pageNumber="206">Fig. 6</figureCitation>
). One of the
<typeStatus id="54968840FF8A38678877AA0AD3B8FCE2" box="[813,911,835,861]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
(150.0 mm SL) was a female with elliptical yolked eggs, of similar size and shape to those of FAKU 146572 (
<figureCitation id="13162A67FF8A38678821AA2ED3E4FC3E" box="[891,979,871,897]" captionStart="FIGURE 6" captionStartId="14.[151,250,1163,1187]" captionTargetBox="[190,1397,417,1140]" captionTargetId="figure-132@14.[190,1397,417,1140]" captionTargetPageId="14" captionText="FIGURE 6. Careproctus mederi, FAKU 146572, 102.3 mm SL, (preserved) (A); diastema at symphysis of upper jaw of FAKU 148478, 143.6 mm SL (B); eggs of FAKU 146572 (C); one of four syntypes of ZIN 18958, 150.0 mm SL, (preserved) (D), eggs (E), and ventral view of head region (F). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219224" httpUri="https://zenodo.org/record/13219224/files/figure.png" pageId="15" pageNumber="206">Fig. 6E</figureCitation>
). Owing to these similarities, the present specimens were identified here as
<taxonomicName id="4C2D4D61FF8A3867894FAAC2D2B4FC1A" box="[533,643,907,933]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A3867894FAAC2D2B4FC1A" box="[533,643,907,933]" italics="true" pageId="15" pageNumber="206">C. mederi</emphasis>
</taxonomicName>
, despite their counts of vertebrae, dorsal- and anal-fin rays, and pyloric caeca being somewhat different from those of the
<typeStatus id="54968840FF8A386789BEAAE6D37DFC76" box="[740,842,943,969]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
. Such are considered here as geographic variations (
<tableCitation id="C6AF0359FF8A38678BC5AA9AD0C1FC52" box="[159,246,979,1005]" captionStart="TABLE 2" captionStartId="13.[151,239,151,175]" captionText="TABLE 2. Counts and measurements of Careproctus mederi, C. sp. cf. gilberti, and C. gilberti." pageId="15" pageNumber="206">Table 2</tableCitation>
). The new Japanese name &quot;Agohige-bikunin&quot; is proposed here on the basis of FAKU 146572.
</paragraph>
<paragraph id="8B9236E2FF8A38678B9DAABED253FAC6" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">
The pectoral girdle morphology, protruding anteroventrally, in
<taxonomicName id="4C2D4D61FF8A386788D4AABED3CAFBAE" box="[910,1021,1015,1041]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A386788D4AABED3CAFBAE" box="[910,1021,1015,1041]" italics="true" pageId="15" pageNumber="206">C. mederi</emphasis>
</taxonomicName>
was also seen in
<taxonomicName id="4C2D4D61FF8A38678F9EAABED57DFBAF" authorityName="Gilbert" authorityYear="1896" box="[1220,1354,1015,1041]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF8A38678F9EAABED57DFBAF" box="[1220,1354,1015,1041]" italics="true" pageId="15" pageNumber="206">C. ostentum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D61FF8A38678ED9AABED12FFB8A" authorityName="Imamura &amp; Nobetsu" authorityYear="2002" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="parvidiscus">
<emphasis id="B959EAF0FF8A38678ED9AABED12FFB8A" italics="true" pageId="15" pageNumber="206">C. parvidiscus</emphasis>
</taxonomicName>
(
<figureCitation id="13162A67FF8A38678A73AD52D193FB8A" box="[297,420,1051,1077]" captionStart="FIGURE 7" captionStartId="16.[151,250,1098,1122]" captionTargetBox="[189,1398,181,1074]" captionTargetId="figure-20@16.[189,1398,181,1074]" captionTargetPageId="16" captionText="FIGURE 7. Careproctus gilberti and related species. Careproctus ostentum, UW 150870, 72.1 mm SL, (preserved) (A); C. parvidiscus, HUMZ 164407, holotype, 167.7 mm SL, (preserved) (B); Careproctus sp. cf. gilberti, FAKU 147753, 55.1 mm SL, (preserved) (C); C. gilberti, OS 8684, 69.2 mm SL, (preserved) (D), and ventral view of head region of OS 8684 (E)." figureDoi="http://doi.org/10.5281/zenodo.13219226" httpUri="https://zenodo.org/record/13219226/files/figure.png" pageId="15" pageNumber="206">Fig. 7A, B</figureCitation>
).
<taxonomicName id="4C2D4D61FF8A38678AE0AD52D2ADFB8A" box="[442,666,1051,1077]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A38678AE0AD52D2ADFB8A" box="[442,666,1051,1077]" italics="true" pageId="15" pageNumber="206">Careproctus mederi</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C2D4D61FF8A38678828AD52D3CEFB8B" authorityName="Gilbert" authorityYear="1896" box="[882,1017,1051,1077]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF8A38678828AD52D3CEFB8B" box="[882,1017,1051,1077]" italics="true" pageId="15" pageNumber="206">C. ostentum</emphasis>
</taxonomicName>
, which has 5156 dorsal- and 4751 anal-fin rays, 813 pyloric caeca, a small pelvic disk, ca. 1.6% of SL, a moderately large gill slit, extending ventrally to pectoral-fin rays 816, a deep and broadly notched pectoral fin, black stomach, pale peritoneum with black dots, and elliptical eggs (
<bibRefCitation id="EFBC4B13FF8A38678AEFADCED27DFB1E" author="Gilbert, C. H." box="[437,586,1159,1185]" pageId="15" pageNumber="206" pagination="393 - 476" refId="ref17520" refString="Gilbert, C. H. (1896) The ichthyological collections of the steamer Albatross during the years 1890 and 1891. United States Commission of Fish and Fisheries, Report of the Commissioner, 19 (Art. 6), 393 - 476, pls. 20 - 35." type="journal article" year="1896">Gilbert 1896</bibRefCitation>
;
<bibRefCitation id="EFBC4B13FF8A3867890DADCED2E8FB1E" author="Burke, C. V." box="[599,735,1159,1185]" pageId="15" pageNumber="206" pagination="1 - 204" refId="ref17139" refString="Burke, C. V. (1930) Revision of the fishes of the family Liparidae. Bulletin of the United States National Museum, 150, i - xii + 1 - 204. https: // doi. org / 10.5479 / si. 03629236.150.1" type="journal article" year="1930">Burke 1930</bibRefCitation>
;
<bibRefCitation id="EFBC4B13FF8A386789B7ADCED43FFB1E" author="Mecklenburg, C. W. &amp; Mecklenburg, T. A. &amp; Thorsteinson, L. K." box="[749,1032,1159,1185]" pageId="15" pageNumber="206" refId="ref18345" refString="Mecklenburg, C. W., Mecklenburg, T. A. &amp; Thorsteinson, L. K. (2002) Fishes of Alaska. American Fisheries Society, Bethesda, Maryland, 1037 pp." type="book" year="2002">
Mecklenburg
<emphasis id="B959EAF0FF8A386788D0ADCED3F3FB1E" box="[906,964,1159,1185]" italics="true" pageId="15" pageNumber="206">et al.</emphasis>
2002
</bibRefCitation>
; this study). However,
<taxonomicName id="4C2D4D61FF8A38678E4BADCED5B6FB1E" box="[1297,1409,1159,1185]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A38678E4BADCED5B6FB1E" box="[1297,1409,1159,1185]" italics="true" pageId="15" pageNumber="206">C. mederi</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C2D4D61FF8A38678A2EADE2D1CCFB7B" authorityName="Gilbert" authorityYear="1896" box="[372,507,1195,1221]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF8A38678A2EADE2D1CCFB7B" box="[372,507,1195,1221]" italics="true" pageId="15" pageNumber="206">C. ostentum</emphasis>
</taxonomicName>
by its lighter body coloration (dark in
<taxonomicName id="4C2D4D61FF8A386788E3ADE2D477FB7B" authorityName="Gilbert" authorityYear="1896" box="[953,1088,1195,1221]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF8A386788E3ADE2D477FB7B" box="[953,1088,1195,1221]" italics="true" pageId="15" pageNumber="206">C. ostentum</emphasis>
</taxonomicName>
:
<figureCitation id="13162A67FF8A38678F0AADE2D49EFB7A" box="[1104,1193,1195,1221]" captionStart="FIGURE 7" captionStartId="16.[151,250,1098,1122]" captionTargetBox="[189,1398,181,1074]" captionTargetId="figure-20@16.[189,1398,181,1074]" captionTargetPageId="16" captionText="FIGURE 7. Careproctus gilberti and related species. Careproctus ostentum, UW 150870, 72.1 mm SL, (preserved) (A); C. parvidiscus, HUMZ 164407, holotype, 167.7 mm SL, (preserved) (B); Careproctus sp. cf. gilberti, FAKU 147753, 55.1 mm SL, (preserved) (C); C. gilberti, OS 8684, 69.2 mm SL, (preserved) (D), and ventral view of head region of OS 8684 (E)." figureDoi="http://doi.org/10.5281/zenodo.13219226" httpUri="https://zenodo.org/record/13219226/files/figure.png" pageId="15" pageNumber="206">Fig. 7A</figureCitation>
) and COI haplotypes (
<figureCitation id="13162A67FF8A38678BC5AD86D0D6FB56" box="[159,225,1231,1257]" captionStart="FIGURE 8" captionStartId="17.[151,250,1694,1718]" captionTargetBox="[160,1430,1289,1664]" captionTargetId="figure-531@17.[153,1434,1281,1671]" captionTargetPageId="17" captionText="FIGURE 8. (A): Neighbor-joining (NJ) tree of the Careproctus gilberti, C. rastrinus, and C. reinhardti species groups based on sequence variations in the mitochondrial COI (474 bp) gene. Support values (≥50% ML bootstrap probability) indicated along branches. Each node labeled with collection registration number (sequences determined here) or INSDC accession number (sequences determined in previous studies, in italics). (B): A minimum spanning network of the C. gilberti species group. Each circle represents a unique haplotype; area of each circle proportional to number of individuals; each bar on internal nodes represents a single nucleotide change. In addition to the sequences determined here, the following sequences from INSDC were used. Careproctus ostentum: MH715573, KU053754; C. gilberti: MH630293, KU053753, MH715534MH715569, MH882481, JQ354029, MH630294." figureDoi="http://doi.org/10.5281/zenodo.13219230" httpUri="https://zenodo.org/record/13219230/files/figure.png" pageId="15" pageNumber="206">Fig. 8</figureCitation>
), although further morphological and genetic information may support the synonymization of the two species. For the present, however, the two species are conservatively recognized as valid. In contrast,
<taxonomicName id="4C2D4D61FF8A38678FF1ADBAD57AFAB2" authorityName="Imamura &amp; Nobetsu" authorityYear="2002" box="[1195,1357,1267,1293]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="parvidiscus">
<emphasis id="B959EAF0FF8A38678FF1ADBAD57AFAB2" box="[1195,1357,1267,1293]" italics="true" pageId="15" pageNumber="206">C. parvidiscus</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="4C2D4D61FF8A38678A91AC5ED20BFA8E" box="[459,572,1303,1329]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A38678A91AC5ED20BFA8E" box="[459,572,1303,1329]" italics="true" pageId="15" pageNumber="206">C. mederi</emphasis>
</taxonomicName>
in having 50 dorsal- and 44 anal-fin rays, and extremely deciduous cactus-like prickles on the body (vs. 5256 dorsal- and 4752 anal-fin rays, and the body with strong cactus-like prickles) (
<bibRefCitation id="EFBC4B13FF8A38678BC5AC16D1EBFAC6" author="Imamura, H. &amp; Nobetsu, T." box="[159,476,1375,1401]" pageId="15" pageNumber="206" pagination="156 - 158" refId="ref17712" refString="Imamura, H. &amp; Nobetsu, T. (2002) Careproctus parvidiscus, a new species of liparid fish (Teleostei: Scorpaeniformes) collected from the southern Okhotsk Sea, Japan. Ichthyological Research, 49, 156 - 158. https: // doi. org / 10.1007 / s 102280200020" type="journal article" year="2002">Imamura and Nobetsu 2002</bibRefCitation>
; this study).
</paragraph>
<paragraph id="8B9236E2FF8A38678B9DACCBD53BF902" blockId="15.[151,1437,151,1725]" pageId="15" pageNumber="206">
<emphasis id="B959EAF0FF8A38678B9DACCBD1D4FA23" bold="true" box="[199,483,1410,1436]" pageId="15" pageNumber="206">Comparative materials.</emphasis>
<taxonomicName id="4C2D4D61FF8A38678AB1ACCAD2FCFA22" box="[491,715,1411,1437]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF8A38678AB1ACCAD2FCFA22" box="[491,715,1411,1437]" italics="true" pageId="15" pageNumber="206">Careproctus mederi</emphasis>
</taxonomicName>
:
<materialsCitation id="3B453CBFFF8A38678980ACCAD4B2FA7E" collectionCode="ZIN" elevation="322" latitude="56.3" location="Iony Island" longLatPrecision="63" longitude="145.067" municipality="northern Sea" pageId="15" pageNumber="206" specimenCode="ZIN 18958" specimenCount="4" typeStatus="syntype">
<specimenCode id="DB8B9E99FF8A38678980ACCAD360FA22" box="[730,855,1410,1437]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="15" pageNumber="206" type="Museum">ZIN 18958</specimenCode>
(X-rays only),
<specimenCount id="9D2BFD6BFF8A38678F5CACCAD4A3FA22" box="[1030,1172,1411,1437]" count="4" pageId="15" pageNumber="206" type="generic">4 specimens</specimenCount>
,
<typeStatus id="54968840FF8A38678FC5ACCAD533FA22" box="[1183,1284,1411,1437]" pageId="15" pageNumber="206" type="syntype">syntypes</typeStatus>
, 108.0152.0 mm SL,
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,
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, off
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,
<collectingMunicipality id="6BF6AC98FF8A3867899AACEED365FA7E" box="[704,850,1447,1473]" pageId="15" pageNumber="206">northern Sea</collectingMunicipality>
of Okhotsk,
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<quantity id="4CD59B07FF8A386788B5ACEED40DFA7E" box="[1007,1082,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="3.22" pageId="15" pageNumber="206" unit="m" value="322.0">322 m</quantity>
depth
</elevation>
</materialsCitation>
,
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, coll. by G.
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<collectionCode id="ED3CAE27FF8A38678BE4AC82D0E3FA5A" box="[190,212,1483,1509]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="206">R</collectionCode>
.
<location id="8EF26039FF8A38678BBEAC82D104FA5A" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF8A38678BBEAC82D104FA5A" box="[228,307,1483,1509]" name="Meder." pageId="15" pageNumber="206">Meder.</location>
<taxonomicName id="4C2D4D61FF8A38678A64AC82D20EFA5B" authorityName="Gilbert" authorityYear="1896" box="[318,569,1483,1509]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF8A38678A64AC82D20EFA5B" box="[318,569,1483,1509]" italics="true" pageId="15" pageNumber="206">Careproctus ostentum</emphasis>
</taxonomicName>
</materialsCitation>
:
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<specimenCode id="DB8B9E99FF8A38678917AC82D2CDFA5A" box="[589,762,1482,1509]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="15" pageNumber="206">FAKU 120225</specimenCode>
,
<specimenCode id="DB8B9E99FF8A38678850AC82D3F5FA5A" box="[778,962,1483,1509]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="15" pageNumber="206">120227120229</specimenCode>
,
<specimenCount id="9D2BFD6BFF8A38678889AC82D471FA5A" box="[979,1094,1483,1509]" count="4" pageId="15" pageNumber="206" type="female">4 females</specimenCount>
,
<quantity id="4CD59B07FF8A38678F0CAC82D4C9FA5A" box="[1110,1278,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.755" metricValueMax="8.09" metricValueMin="7.42" pageId="15" pageNumber="206" unit="mm" value="77.55" valueMax="80.9" valueMin="74.2">74.280.9 mm</quantity>
SL,
<collectorName id="26D85334FF8A38678E66AC82D083F9B6" pageId="15" pageNumber="206">Aleutian Is.</collectorName>
,
<geoCoordinate id="EE195025FF8A38678B99ACA7D107F9B6" box="[195,304,1518,1545]" degrees="52.550" direction="north" orientation="latitude" pageId="15" pageNumber="206" precision="55" value="52.55">52.550ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8A38678A64ACA7D182F9B6" box="[318,437,1518,1545]" degrees="179.950" direction="east" orientation="longitude" pageId="15" pageNumber="206" precision="55" value="179.95">179.950ºE</geoCoordinate>
,
<elevation id="0000D1D1FF8A38678A99ACA6D26FF9B6" box="[451,600,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="15" pageNumber="206" unit="m" value="279.0">
<quantity id="4CD59B07FF8A38678A99ACA6D23BF9B6" box="[451,524,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="15" pageNumber="206" unit="m" value="279.0">279 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8A3867893FACA6D253F992" collectionCode="HUMZ" collectorName="Aleutian Is." elevation="610" latitude="59.328" longLatPrecision="62" longitude="-178.112" pageId="15" pageNumber="206" specimenCode="HUMZ 82565-82567" specimenCount="2" specimenCount-male="2" typeStatus="syntype">
<specimenCode id="DB8B9E99FF8A3867893FACA6D369F9B6" box="[613,862,1518,1545]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="15" pageNumber="206">HUMZ 8256582567</specimenCode>
, 48.0
<quantity id="4CD59B07FF8A386788F6ACA6D425F9B6" box="[940,1042,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="6.659999999999999" pageId="15" pageNumber="206" unit="mm" value="66.6">66.6 mm</quantity>
SL,
<specimenCount id="9D2BFD6BFF8A38678F16ACA6D492F9B6" box="[1100,1189,1519,1545]" count="2" pageId="15" pageNumber="206" type="male">2 males</specimenCount>
and female,
<collectorName id="26D85334FF8A38678E66ACA6D083F992" pageId="15" pageNumber="206">Aleutian Is.</collectorName>
,
<geoCoordinate id="EE195025FF8A38678B9EAF5BD106F992" box="[196,305,1554,1581]" degrees="59.328" direction="north" orientation="latitude" pageId="15" pageNumber="206" precision="55" value="59.328">59.328ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8A38678A1AAF5AD189F992" box="[320,446,1555,1581]" degrees="178.112" direction="west" orientation="longitude" pageId="15" pageNumber="206" precision="55" value="-178.112">178.112ºW</geoCoordinate>
,
<elevation id="0000D1D1FF8A38678A97AF5AD253F992" box="[461,612,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="15" pageNumber="206" unit="m" value="610.0">
<quantity id="4CD59B07FF8A38678A97AF5AD22FF992" box="[461,536,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="15" pageNumber="206" unit="m" value="610.0">610 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8A38678929AF5AD315F9EE" collectionCode="USNM" country="United States of America" elevation="199" latitude="53.564" location="Unalaska Is." longLatPrecision="64" longitude="-167.781" pageId="15" pageNumber="206" specimenCode="USNM 48619" specimenCount="1" typeStatus="lectotype">
<specimenCode id="DB8B9E99FF8A38678929AF5AD322F992" box="[627,789,1555,1581]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="206" type="Museum">USNM 48619</specimenCode>
(photo and X-ray photo),
<typeStatus id="54968840FF8A38678F15AF5AD48EF992" box="[1103,1209,1555,1581]" pageId="15" pageNumber="206" type="lectotype">lectotype</typeStatus>
, damaged, north of
<location id="8EF26039FF8A38678BCDAF7ED110F9EE" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF8A38678BCDAF7ED110F9EE" box="[151,295,1591,1617]" country="United States of America" latitude="53.564" longLatPrecision="64" longitude="-167.781" name="Unalaska Is." pageId="15" pageNumber="206">Unalaska Is.</location>
,
<collectingCountry id="F33A7672FF8A38678A6DAF7ED145F9EE" box="[311,370,1591,1617]" name="United States of America" pageId="15" pageNumber="206">USA</collectingCountry>
,
<geoCoordinate id="EE195025FF8A38678ADBAF7FD1D9F9EE" box="[385,494,1590,1617]" degrees="53.564" direction="north" orientation="latitude" pageId="15" pageNumber="206" precision="55" value="53.564">53.564ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8A38678AA7AF7ED24BF9EE" box="[509,636,1591,1617]" degrees="167.781" direction="west" orientation="longitude" pageId="15" pageNumber="206" precision="55" value="-167.781">167.781ºW</geoCoordinate>
,
<elevation id="0000D1D1FF8A386789D1AF7ED315F9EE" box="[651,802,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" pageId="15" pageNumber="206" unit="m" value="199.0">
<quantity id="4CD59B07FF8A386789D1AF7ED2E2F9EE" box="[651,725,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" pageId="15" pageNumber="206" unit="m" value="199.0">199 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8A3867886BAF7ED259F9CA" collectionCode="UW" collectorName="Bering Sea" country="United States of America" elevation="332" latitude="59.398" longLatPrecision="62" longitude="-177.67" pageId="15" pageNumber="206" specimenCode="UW 119726" specimenCount="1" typeStatus="lectotype">
<specimenCode id="DB8B9E99FF8A3867886BAF7ED3F7F9EE" box="[817,960,1591,1617]" collectionCode="UW" country="USA" name="University of Washington Fish Collection" pageId="15" pageNumber="206" type="Museum">UW 119726</specimenCode>
,
<quantity id="4CD59B07FF8A38678895AF7ED400F9EE" box="[975,1079,1591,1617]" metricMagnitude="-2" metricUnit="m" metricValue="9.77" pageId="15" pageNumber="206" unit="mm" value="97.7">97.7 mm</quantity>
SL, sex undetermined,
<collectorName id="26D85334FF8A38678E0AAF7ED0F5F9CA" pageId="15" pageNumber="206">Bering Sea</collectorName>
,
<geoCoordinate id="EE195025FF8A38678B8BAF13D10AF9CA" box="[209,317,1626,1653]" degrees="59.398" direction="north" orientation="latitude" pageId="15" pageNumber="206" precision="55" value="59.398">59.398ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8A38678A16AF12D1FDF9CA" box="[332,458,1627,1653]" degrees="177.670" direction="west" orientation="longitude" pageId="15" pageNumber="206" precision="55" value="-177.67">177.670ºW</geoCoordinate>
,
<elevation id="0000D1D1FF8A38678A82AF12D259F9CA" box="[472,622,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="3.32" pageId="15" pageNumber="206" unit="m" value="332.0">
<quantity id="4CD59B07FF8A38678A82AF12D215F9CA" box="[472,546,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="3.32" pageId="15" pageNumber="206" unit="m" value="332.0">332 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8A38678926AF12D21DF926" collectionCode="UW" collectorName="Aleutian Is." country="United States of America" latitude="51.717" longLatPrecision="65" longitude="177.294" pageId="15" pageNumber="206" specimenCode="UW 150870" specimenCount="1" typeStatus="lectotype">
<specimenCode id="DB8B9E99FF8A38678926AF12D33CF9CA" box="[636,779,1626,1653]" collectionCode="UW" country="USA" name="University of Washington Fish Collection" pageId="15" pageNumber="206" type="Museum">UW 150870</specimenCode>
,
<quantity id="4CD59B07FF8A38678843AF12D3B7F9CA" box="[793,896,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="7.209999999999999" pageId="15" pageNumber="206" unit="mm" value="72.1">72.1 mm</quantity>
SL, sex undetermined,
<collectorName id="26D85334FF8A38678FCFAF12D52BF9CA" box="[1173,1308,1627,1653]" pageId="15" pageNumber="206">Aleutian Is.</collectorName>
,
<geoCoordinate id="EE195025FF8A38678E71AF13D5AFF9CA" box="[1323,1432,1626,1653]" degrees="51.717" direction="north" orientation="latitude" pageId="15" pageNumber="206" precision="55" value="51.717">51.717ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8A38678BCDAF36D127F926" box="[151,272,1663,1689]" degrees="177.294" direction="east" orientation="longitude" pageId="15" pageNumber="206" precision="55" value="177.294">177.294ºE</geoCoordinate>
.
<taxonomicName id="4C2D4D61FF8A38678A43AF36D21DF926" authorityName="Imamura &amp; Nobetsu" authorityYear="2002" box="[281,554,1663,1689]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="15" pageNumber="206" phylum="Chordata" rank="species" species="parvidiscus">
<emphasis id="B959EAF0FF8A38678A43AF36D21DF926" box="[281,554,1663,1689]" italics="true" pageId="15" pageNumber="206">Careproctus parvidiscus</emphasis>
</taxonomicName>
</materialsCitation>
:
<materialsCitation id="3B453CBFFF8A3867896DAF36D2F9F902" collectionCode="HUMZ" collectorName="Shiretoko Point" country="Japan" elevation="550" location="southern Sea" pageId="15" pageNumber="206" specimenCode="HUMZ 164407" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<specimenCode id="DB8B9E99FF8A3867896DAF36D2D0F926" box="[567,743,1663,1689]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="15" pageNumber="206">HUMZ 164407</specimenCode>
,
<typeStatus id="54968840FF8A386789AAAF36D362F926" box="[752,853,1663,1689]" pageId="15" pageNumber="206" type="holotype">holotype</typeStatus>
,
<quantity id="4CD59B07FF8A38678805AF36D3E7F926" box="[863,976,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="1.677" pageId="15" pageNumber="206" unit="mm" value="167.7">167.7 mm</quantity>
SL, male, off
<collectorName id="26D85334FF8A38678F36AF36D52AF926" box="[1132,1309,1663,1689]" pageId="15" pageNumber="206">Shiretoko Point</collectorName>
,
<collectingRegion id="49E9F800FF8A38678E7DAF36D5AFF926" box="[1319,1432,1663,1689]" country="Japan" name="Hokkaido" pageId="15" pageNumber="206">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8A38678BCDAFEAD0EDF902" box="[151,218,1699,1725]" name="Japan" pageId="15" pageNumber="206">Japan</collectingCountry>
,
<location id="8EF26039FF8A38678BBFAFEAD143F902" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF8A38678BBFAFEAD143F902" box="[229,372,1699,1725]" country="Japan" name="southern Sea" pageId="15" pageNumber="206" stateProvince="Hokkaido">southern Sea</location>
of Okhotsk,
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<quantity id="4CD59B07FF8A3867895DAFEAD2B1F902" box="[519,646,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="15" pageNumber="206" unit="m" value="550.0" valueMax="700.0" valueMin="400.0">400700 m</quantity>
depth
</elevation>
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, bottom gill net,
<date id="FF931022FF8A386788D6AFEAD413F902" box="[908,1060,1699,1725]" pageId="15" pageNumber="206" value="1997-05-12">12 May 1997</date>
, coll. by
<materialsCitation id="3B453CBFFF8A38678FD0AFEAD53FF902" box="[1162,1288,1699,1725]" collectionCode="T" country="Japan" location="Nobetsu" pageId="15" pageNumber="206" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<collectionCode id="ED3CAE27FF8A38678FD0AFEAD4ABF902" box="[1162,1180,1699,1725]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="206">T</collectionCode>
.
<location id="8EF26039FF8A38678FFDAFEAD53FF902" LSID="urn:lsid:plazi:treatment:038487F4FF8938678BCDAD86D53BF902:8EF26039FF8A38678FFDAFEAD53FF902" box="[1191,1288,1699,1725]" country="Japan" name="Nobetsu" pageId="15" pageNumber="206" stateProvince="Hokkaido">Nobetsu</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,589 @@
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<mods:title id="23AB164C708ABD869DE2A410459A1C7E">Snailfishes of the genus Careproctus (Perciformes: Liparidae) with a reduced pelvic disk: three new species and new records from the western North Pacific with comments on their phenotypic diversity</mods:title>
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<mods:namePart id="530DE95E874D6940B3A7103A0EDE42F9">Kai, Yoshiaki</mods:namePart>
<mods:affiliation id="E57AFE617202FDF2F8D1BF9129890E8F">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:namePart id="173F3638E0327585154F5CF0958CC8B6">Matsuzaki, Koji</mods:namePart>
<mods:affiliation id="FB600D1240E9074BDB77579A5096F064">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="C31868507834DC7E740E38AA47A27595">Mori, Toshiaki</mods:namePart>
<mods:affiliation id="2687AD23AA6F24F14D21584BF84940BE">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="42E13B931F3800880171113B7B5B6DB9">Pitruk, Dmitry L.</mods:namePart>
<mods:affiliation id="82617CC2DB04CC24F6F3C74205D2F64E">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, ul. Palchevskogo 17, Vladivostok 690041, Russia</mods:affiliation>
</mods:name>
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<mods:namePart id="878E95E6DA4EF1F4418B10F9207859D2">Misawa, Ryo</mods:namePart>
<mods:affiliation id="1DDDFB240BEDC6B92042968871212ACB">Demersal Fish Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, National Research and Development Agency, 25 - 259 Shimomekurakubo, Same, Hachinohe, Aomori 031 - 0841, Japan</mods:affiliation>
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<mods:namePart id="2A169756887E7FC1957688C616C7AA5F">Tashiro, Fumihito</mods:namePart>
<mods:affiliation id="87359EF51FB954C150A4FECAB4A7CCEA">Fisheries Science Center, The Hokkaido University Museum, 3 - 1 - 1 Minato-cho, Hakodate, Hokkaido 041 - 8611, Japan</mods:affiliation>
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<mods:date id="45492313DB4D3B4AE7CCAFA20CFCE98A">2024</mods:date>
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<taxonomicName id="4C2D4D61FF8F38628BCDAB52D183FD8A" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[151,436,538,565]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF8F38628BCDAB52D183FD8A" bold="true" box="[151,436,538,565]" italics="true" pageId="10" pageNumber="201">Careproctus spinulosus</emphasis>
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Kai, Matsuzaki &amp; Mori
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<paragraph id="8B9236E2FF8F38628BCDAB76D269FDE6" blockId="10.[151,979,538,673]" box="[151,606,575,601]" pageId="10" pageNumber="201">New Japanese name: Tomoshibi-bikunin</paragraph>
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<subSubSection id="C3376569FF8F38628BCDAB2AD3E4FD1E" pageId="10" pageNumber="201" type="description">
<paragraph id="8B9236E2FF8F38628BCDAB2AD15DFDC2" blockId="10.[151,979,538,673]" box="[151,362,610,637]" pageId="10" pageNumber="201">
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,
<figureCitation id="13162A67FF8F38628BA1AB2BD13CFDC2" box="[251,267,610,637]" captionStart="FIGURE 5" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[242,1344,705,1916]" captionTargetId="figure-284@11.[242,1344,705,1916]" captionTargetPageId="11" captionText="FIGURE 5. Careproctus spinulosus sp. nov., FAKU 146600, holotype, 201.0 mm SL, fresh condition (A), preserved condition (B), and ventral view of head region (C); ventral view of head region of FAKU 148864, paratype, 112.2 mm SL (D). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219222" httpUri="https://zenodo.org/record/13219222/files/figure.png" pageId="10" pageNumber="201">5</figureCitation>
;
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</paragraph>
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.
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depth
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, F/
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<emphasis id="B959EAF0FF8F38628A4FABBAD193FCB3" box="[277,420,755,780]" italics="true" pageId="10" pageNumber="201">Houyu-maru</emphasis>
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<collectingDate id="EFD7E9CAFF8F38628AEBABBAD21DFCB2" box="[433,554,755,781]" pageId="10" pageNumber="201" value="2018-08">Aug. 2018</collectingDate>
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, coll. by Koji Matsuzaki, Shigeki Fujimoto, and Shigetada Fujimoto.
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<typeStatus id="54968840FF8F38628B9DAA5ED176FC8E" box="[199,321,791,817]" pageId="10" pageNumber="201" type="paratype">Paratypes</typeStatus>
.
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,
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,
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,
<specimenCode id="DB8B9E99FF8F38628985AA5FD32BFC8E" box="[735,796,790,817]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">105.1</specimenCode>
<quantity id="4CD59B07FF8F38628871AA5FD394FC8E" box="[811,931,790,817]" metricMagnitude="-1" metricUnit="m" metricValue="1.906" pageId="10" pageNumber="201" unit="mm" value="190.6">190.6 mm</quantity>
SL,
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and
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</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628A52AA72D5AFFCEA" box="[264,1432,826,853]" collectionCode="FAKU" country="Japan" elevation="650" location="Rausu" municipality="Hokkaido" pageId="10" pageNumber="201" specimenCode="FAKU 146631-146632, 170.6" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628A52AA72D225FCEA" box="[264,530,827,853]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 146631146632</specimenCode>
,
<specimenCode id="DB8B9E99FF8F38628947AA72D26BFCEA" box="[541,604,827,853]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">170.6</specimenCode>
<quantity id="4CD59B07FF8F38628931AA72D2EBFCEA" box="[619,732,827,853]" metricMagnitude="-1" metricUnit="m" metricValue="1.998" pageId="10" pageNumber="201" unit="mm" value="199.8">199.8 mm</quantity>
SL,
<specimenCount id="9D2BFD6BFF8F38628848AA72D3B6FCEA" box="[786,897,827,853]" count="2" pageId="10" pageNumber="201" type="female">2 females</specimenCount>
, off
<location id="8EF26039FF8F386288E9AA72D3CBFCEA" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F386288E9AA72D3CBFCEA" box="[947,1020,827,853]" country="Japan" municipality="Hokkaido" name="Rausu" pageId="10" pageNumber="201">Rausu</location>
,
<collectingMunicipality id="6BF6AC98FF8F38628F5DAA72D44FFCEA" box="[1031,1144,827,853]" pageId="10" pageNumber="201">Hokkaido</collectingMunicipality>
,
<collectingCountry id="F33A7672FF8F38628FD9AA72D4F1FCEA" box="[1155,1222,827,853]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<elevation id="0000D1D1FF8F38628F8BAA73D5AFFCEA" box="[1233,1432,826,853]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="10" pageNumber="201" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF8F38628F8BAA73D567FCEA" box="[1233,1360,826,853]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="10" pageNumber="201" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8F38628BF5AA16D576FCC6" box="[175,1345,863,889]" collectingDate="2014-09-10" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628BF5AA16D0F5FCC6" box="[175,194,863,889]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628B94AA16D16AFCC7" box="[206,349,863,888]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628B94AA16D16AFCC7" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628B94AA16D16AFCC7" box="[206,349,863,888]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F38628A2BAA16D226FCC6" box="[369,529,863,889]" pageId="10" pageNumber="201" value="2014-09-10">
<collectingDate id="EFD7E9CAFF8F38628A2BAA16D226FCC6" box="[369,529,863,889]" pageId="10" pageNumber="201" value="2014-09-10">10 Sep. 2014</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F386289DDAA16D30AFCC6" box="[647,829,863,889]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F38628817AA16D42FFCC6" box="[845,1048,863,889]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628F07AA16D576FCC6" box="[1117,1345,863,889]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628E08AA16D4F9FC22" collectionCode="FAKU" collectorName="Rausu" country="Japan" latitude="44.023" location="Hokkaido" longLatPrecision="68" longitude="145.353" pageId="10" pageNumber="201" specimenCode="FAKU 146900" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628E08AA16D0D9FC22" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 146900</specimenCode>
, male and female, off
<collectorName id="26D85334FF8F38628AB3AACAD205FC22" box="[489,562,899,925]" pageId="10" pageNumber="201">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF8F38628967AACAD299FC22" box="[573,686,899,925]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F386289E0AACAD2CAFC22" box="[698,765,899,925]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<geoCoordinate id="EE195025FF8F38628852AACAD343FC22" box="[776,884,899,925]" degrees="44.023" direction="north" orientation="latitude" pageId="10" pageNumber="201" precision="55" value="44.023">44.023ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8F386288DAAACBD3C0FC22" box="[896,1015,898,925]" degrees="145.353" direction="east" orientation="longitude" pageId="10" pageNumber="201" precision="55" value="145.353">145.353ºE</geoCoordinate>
, depth unknown, F
</materialsCitation>
/
<materialsCitation id="3B453CBFFF8F38628F8CAACAD427FC7E" collectingDate="2019-08-06" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628F8CAACAD4DEFC22" box="[1238,1257,899,925]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628FABAACAD5B7FC23" box="[1265,1408,899,924]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628FABAACAD5B7FC23" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628FABAACAD5B7FC23" box="[1265,1408,899,924]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F38628ED4AACAD127FC7E" pageId="10" pageNumber="201" value="2019-08-06">
<collectingDate id="EFD7E9CAFF8F38628ED4AACAD127FC7E" pageId="10" pageNumber="201" value="2019-08-06">6 Aug. 2019</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F38628A2CAAEED211FC7E" box="[374,550,935,961]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F3862896BAAEED2C0FC7E" box="[561,759,935,961]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F3862886BAAEED427FC7E" box="[817,1040,935,961]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628F41AAEED21BFC5A" collectionCode="FAKU" country="Japan" location="Rausu" pageId="10" pageNumber="201" specimenCode="FAKU 147539" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628F41AAEED4F4FC7E" box="[1051,1219,934,961]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 147539</specimenCode>
, female, off
<location id="8EF26039FF8F38628E15AAEED5AFFC7E" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628E15AAEED5AFFC7E" box="[1359,1432,935,961]" country="Japan" name="Rausu" pageId="10" pageNumber="201" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8F38628BCDAA82D13FFC5A" box="[151,264,971,997]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F38628A4EAA82D160FC5A" box="[276,343,971,997]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
, depth unknown, F
</materialsCitation>
/
<materialsCitation id="3B453CBFFF8F3862896FAA82D142FBB6" collectingDate="2020-04-14" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F3862896FAA82D27FFC5A" box="[565,584,971,997]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628915AA82D2E9FC5B" box="[591,734,971,996]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628915AA82D2E9FC5B" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628915AA82D2E9FC5B" box="[591,734,971,996]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F386289B6AA82D3B6FC5A" box="[748,897,971,997]" pageId="10" pageNumber="201" value="2020-04-14">
<collectingDate id="EFD7E9CAFF8F386289B6AA82D3B6FC5A" box="[748,897,971,997]" pageId="10" pageNumber="201" value="2020-04-14">14 Apr. 2020</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F386288BDAA82D4AFFC5A" box="[999,1176,971,997]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F38628FF8AA82D55EFC5A" box="[1186,1385,971,997]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628BCDAAA6D142FBB6" box="[151,373,1007,1033]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628ADAAAA6D16BFB92" collectionCode="FAKU" country="Japan" elevation="700" location="Rausu" pageId="10" pageNumber="201" specimenCode="FAKU 147741-147742, 156.2" specimenCount="2" specimenCount-female="2" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628ADAAAA6D2BFFBB6" box="[384,648,1007,1033]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 147741147742</specimenCode>
,
<specimenCode id="DB8B9E99FF8F386289C8AAA7D2E5FBB6" box="[658,722,1006,1033]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">156.2</specimenCode>
+ mm SL,
<quantity id="4CD59B07FF8F3862881CAAA6D380FBB6" box="[838,951,1007,1033]" metricMagnitude="-1" metricUnit="m" metricValue="1.663" pageId="10" pageNumber="201" unit="mm" value="166.3">166.3 mm</quantity>
SL,
<specimenCount id="9D2BFD6BFF8F386288B0AAA6D46FFBB6" box="[1002,1112,1007,1033]" count="2" pageId="10" pageNumber="201" type="female">2 females</specimenCount>
, off
<location id="8EF26039FF8F38628FDDAAA6D4E7FBB6" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628FDDAAA6D4E7FBB6" box="[1159,1232,1007,1033]" country="Japan" name="Rausu" pageId="10" pageNumber="201" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8F38628F80AAA6D57CFBB6" box="[1242,1355,1007,1033]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F38628E0FAAA6D5AFFBB6" box="[1365,1432,1007,1033]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<elevation id="0000D1D1FF8F38628BCDAD5AD16BFB92" box="[151,348,1043,1069]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="10" pageNumber="201" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">
<quantity id="4CD59B07FF8F38628BCDAD5AD122FB92" box="[151,277,1043,1069]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="10" pageNumber="201" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8F38628A24AD5AD136FBEE" collectingDate="2020-07-20" collectionCode="V" collectorName="Koji Matsuzaki, Y &amp; Kai &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628A24AD5AD1A6FB92" box="[382,401,1043,1069]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628AC2AD5AD210FB93" box="[408,551,1043,1068]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628AC2AD5AD210FB93" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628AC2AD5AD210FB93" box="[408,551,1043,1068]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F3862896EAD5AD2F7FB92" box="[564,704,1043,1069]" pageId="10" pageNumber="201" value="2020-07-20">
<collectingDate id="EFD7E9CAFF8F3862896EAD5AD2F7FB92" box="[564,704,1043,1069]" pageId="10" pageNumber="201" value="2020-07-20">20 Jul. 2020</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F38628879AD5AD3C7FB92" box="[803,1008,1043,1069]" pageId="10" pageNumber="201">Koji Matsuzaki, Y</collectorName>
.
<collectorName id="26D85334FF8F386288A0AD5AD412FB92" box="[1018,1061,1043,1069]" pageId="10" pageNumber="201">Kai</collectorName>
,
<collectorName id="26D85334FF8F38628F75AD5AD4C2FB92" box="[1071,1269,1043,1069]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628E74AD5AD136FBEE" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628A55AD7ED0EFFBCA" collectionCode="FAKU" country="Japan" elevation="645" latitude="44.024" location="Rausu" longLatPrecision="68" longitude="145.308" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628A55AD7ED16EFBEE" box="[271,345,1079,1105]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU</collectionCode>
<quantity id="4CD59B07FF8F38628A38AD7ED20DFBEE" box="[354,570,1079,1105]" metricMagnitude="5" metricUnit="m" metricValue="1.479092004" pageId="10" pageNumber="201" unit="mm" value="1.479092004E8">147909, 200.4 mm</quantity>
SL, female,
<geoCoordinate id="EE195025FF8F38628995AD7ED30BFBEE" box="[719,828,1079,1105]" degrees="44.024" direction="north" orientation="latitude" pageId="10" pageNumber="201" precision="55" value="44.024">44.024ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF8F38628813AD7FD3F6FBEE" box="[841,961,1078,1105]" degrees="145.308" direction="east" orientation="longitude" pageId="10" pageNumber="201" precision="55" value="145.308">145.308ºE</geoCoordinate>
, off
<location id="8EF26039FF8F386288ADAD7ED477FBEE" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F386288ADAD7ED477FBEE" box="[1015,1088,1079,1105]" country="Japan" latitude="44.024" longLatPrecision="68" longitude="145.308" name="Rausu" pageId="10" pageNumber="201" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8F38628F17AD7ED489FBEE" box="[1101,1214,1079,1105]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F38628F96AD7ED538FBEE" box="[1228,1295,1079,1105]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<elevation id="0000D1D1FF8F38628E41AD7FD0EFFBCA" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="10" pageNumber="201" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">
<quantity id="4CD59B07FF8F38628E41AD7FD5ABFBEE" box="[1307,1436,1078,1105]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="10" pageNumber="201" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">540750 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8F38628BA6AD12D50AFBCA" box="[252,1341,1115,1141]" collectingDate="2020-08" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628BA6AD12D138FBCA" box="[252,271,1115,1141]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628A42AD12D190FBCB" box="[280,423,1115,1140]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628A42AD12D190FBCB" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628A42AD12D190FBCB" box="[280,423,1115,1140]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F38628AEFAD12D207FBCA" box="[437,560,1115,1141]" pageId="10" pageNumber="201" value="2020-08">
<collectingDate id="EFD7E9CAFF8F38628AEFAD12D207FBCA" box="[437,560,1115,1141]" pageId="10" pageNumber="201" value="2020-08">Aug. 2020</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F386289C3AD12D37BFBCA" box="[665,844,1115,1141]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F38628802AD12D428FBCA" box="[856,1055,1115,1141]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628F06AD12D50AFBCA" box="[1116,1341,1115,1141]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628E10AD12D4F2FB26" collectionCode="HUMZ, FAKU" country="Japan" elevation="650" location="Rausu" pageId="10" pageNumber="201" specimenCode="HUMZ 235310, FAKU 148864" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628E10AD12D0DCFB26" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="10" pageNumber="201">HUMZ 235310</specimenCode>
(
<specimenCode id="DB8B9E99FF8F38628BA1AD36D19FFB26" box="[251,424,1151,1177]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 148864</specimenCode>
),
<quantity id="4CD59B07FF8F38628AE7AD36D207FB26" box="[445,560,1151,1177]" metricMagnitude="-1" metricUnit="m" metricValue="1.122" pageId="10" pageNumber="201" unit="mm" value="112.2">112.2 mm</quantity>
SL, male, off
<location id="8EF26039FF8F38628982AD36D316FB26" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628982AD36D316FB26" box="[728,801,1151,1177]" country="Japan" name="Rausu" pageId="10" pageNumber="201" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8F38628877AD36D3A9FB26" box="[813,926,1151,1177]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F386288F6AD36D3D8FB26" box="[940,1007,1151,1177]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<elevation id="0000D1D1FF8F386288A1AD37D4F2FB26" box="[1019,1221,1150,1177]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="10" pageNumber="201" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF8F386288A1AD37D44BFB26" box="[1019,1148,1150,1177]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="10" pageNumber="201" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8F38628FB0AD36D3CBFB02" collectingDate="2021-07" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F38628FB0AD36D4CAFB26" box="[1258,1277,1151,1177]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F38628E5CAD36D5A2FB27" box="[1286,1429,1151,1176]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F38628E5CAD36D5A2FB27" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628E5CAD36D5A2FB27" box="[1286,1429,1151,1176]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F38628BCDADEAD137FB02" box="[151,256,1187,1213]" pageId="10" pageNumber="201" value="2021-07">
<collectingDate id="EFD7E9CAFF8F38628BCDADEAD137FB02" box="[151,256,1187,1213]" pageId="10" pageNumber="201" value="2021-07">Jul. 2021</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F38628A3FADEAD222FB02" box="[357,533,1187,1213]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F38628945ADEAD2D2FB02" box="[543,741,1187,1213]" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628844ADEAD3CBFB02" box="[798,1020,1187,1213]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF8F38628F5DADEAD2F3FB5E" collectionCode="FAKU" country="Japan" elevation="1000" location="Rausu" pageId="10" pageNumber="201" specimenCode="FAKU 148884" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF8F38628F5DADEAD498FB02" box="[1031,1199,1187,1213]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="10" pageNumber="201">FAKU 148884</specimenCode>
, male,
<quantity id="4CD59B07FF8F38628FA7ADEAD558FB02" box="[1277,1391,1187,1213]" metricMagnitude="-1" metricUnit="m" metricValue="1.988" pageId="10" pageNumber="201" unit="mm" value="198.8">198.8 mm</quantity>
SL, off
<location id="8EF26039FF8F38628B9AAD8ED13EFB5E" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F38628B9AAD8ED13EFB5E" box="[192,265,1223,1249]" country="Japan" name="Rausu" pageId="10" pageNumber="201" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF8F38628A4CAD8ED1B0FB5E" box="[278,391,1223,1249]" country="Japan" name="Hokkaido" pageId="10" pageNumber="201">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8F38628ACFAD8ED1EFFB5E" box="[405,472,1223,1249]" name="Japan" pageId="10" pageNumber="201">Japan</collectingCountry>
,
<elevation id="0000D1D1FF8F38628ABEAD8ED2F3FB5E" box="[484,708,1223,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="201" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">
<quantity id="4CD59B07FF8F38628ABEAD8ED24DFB5E" box="[484,634,1223,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="201" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">8001,200 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF8F386289B3AD8ED22BFABA" collectingDate="2021-05" collectionCode="V" collectorName="Koji Matsuzaki &amp; Shigeki Fujimoto &amp; Shigetada Fujimoto" country="Japan" location="Houyu-maru" pageId="10" pageNumber="201" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF8F386289B3AD8ED2CBFB5E" box="[745,764,1223,1249]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="201" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF8F3862885FAD8ED3A3FB5F" box="[773,916,1223,1248]" italics="true" pageId="10" pageNumber="201">
<location id="8EF26039FF8F3862885FAD8ED3A3FB5F" LSID="urn:lsid:plazi:treatment:038487F4FF8F38648BCDAB52D188FBC2:8EF26039FF8F3862885FAD8ED3A3FB5F" box="[773,916,1223,1248]" country="Japan" name="Houyu-maru" pageId="10" pageNumber="201" stateProvince="Hokkaido">Houyu-maru</location>
</emphasis>
,
<date id="FF931022FF8F386288FEAD8ED42BFB5E" box="[932,1052,1223,1249]" pageId="10" pageNumber="201" value="2021-05">
<collectingDate id="EFD7E9CAFF8F386288FEAD8ED42BFB5E" box="[932,1052,1223,1249]" pageId="10" pageNumber="201" value="2021-05">May 2021</collectingDate>
</date>
, coll. by
<collectorName id="26D85334FF8F38628FDDAD8ED50DFB5E" box="[1159,1338,1223,1249]" pageId="10" pageNumber="201">Koji Matsuzaki</collectorName>
,
<collectorName id="26D85334FF8F38628E1DAD8ED136FABA" pageId="10" pageNumber="201">Shigeki Fujimoto</collectorName>
, and
<collectorName id="26D85334FF8F38628A61ADA2D22BFABA" box="[315,540,1259,1285]" pageId="10" pageNumber="201">Shigetada Fujimoto</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8F38628B9DAC47D1C1FA62" pageId="10" pageNumber="201" type="diagnosis">
<paragraph id="8B9236E2FF8F38628B9DAC47D1C1FA62" blockId="10.[151,1437,718,2041]" pageId="10" pageNumber="201">
<emphasis id="B959EAF0FF8F38628B9DAC47D175FA97" bold="true" box="[199,322,1294,1320]" pageId="10" pageNumber="201">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2D4D61FF8F38628A8DAC46D256FA96" authorityName="Kroyer" authorityYear="1862" box="[471,609,1295,1321]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="10" pageNumber="201" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF8F38628A8DAC46D256FA96" box="[471,609,1295,1321]" italics="true" pageId="10" pageNumber="201">Careproctus</emphasis>
</taxonomicName>
with the following combination of characters: vertebrae 5558; dorsal-fin rays 5052; anal-fin rays 4348; pectoral-fin rays 3136; pectoral fin deeply notched, lower lobe long, same length as or slightly longer than upper lobe, extending to two-thirds between anus and anal fin origin; pelvic disk rounded, greatly reduced; cephalic pores 2-6-7-2, chin pores paired in separate pits; gill slit large, extending ventrally to pectoral-fin rays 611; teeth simple, shouldered or weakly trilobed; anus located below posterior rim of orbit; peritoneum and stomach black.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8F38648B9DACAFD4D5FD82" lastPageId="12" lastPageNumber="203" pageId="10" pageNumber="201" type="description">
<paragraph id="8B9236E2FF8F38628B9DACAFD5B0F9BE" blockId="10.[151,1437,718,2041]" box="[199,1415,1510,1537]" pageId="10" pageNumber="201">
<emphasis id="B959EAF0FF8F38628B9DACAFD16EF9BF" bold="true" box="[199,345,1510,1536]" pageId="10" pageNumber="201">Description.</emphasis>
Measurements shown in
<tableCitation id="C6AF0359FF8F3862892CACAED2FDF9BE" box="[630,714,1511,1537]" captionStart="TABLE 1" captionStartId="7.[151,239,404,428]" captionText="TABLE 1. Counts and measurements of Careproctus barbatulus sp. nov., C. longibarbatus sp. nov., and C. spinulosus sp. nov." pageId="10" pageNumber="201">Table 1</tableCitation>
.
<typeStatus id="54968840FF8F3862898DACAED30EF9BE" box="[727,825,1511,1537]" pageId="10" pageNumber="201" type="paratype">Paratype</typeStatus>
data given in parentheses if different from
<typeStatus id="54968840FF8F38628E44ACAED5B3F9BE" box="[1310,1412,1511,1537]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B9236E2FF8F38628B9DAF42D3ECF846" blockId="10.[151,1437,718,2041]" pageId="10" pageNumber="201">Body deep anteriorly, moderately compressed, tapering posteriorly. Skin thick, completely covered by cactus-like prickles; gelatinous tissue under skin well developed. Head large, robust, broadly rounded from nape to snout. Snout rounded, not protruding anterior to mouth. Mouth moderately large, terminal, horizontal; maxilla extending to mid orbit (or slightly shorter); oral cleft extending to anterior rim of orbit. Premaxillary teeth blunt, simple, shouldered or weakly trilobed, arranged in oblique rows of 6 (46) teeth per row forming relatively narrow band, inner teeth not enlarged. Mandibular teeth blunt, simple, shouldered or weakly trilobed, arranged in oblique rows of 6 (36) teeth per row forming narrow band, inner teeth not enlarged. Narrow diastema at symphysis of upper and lower jaws. Orbit of moderate size, rounded; pupil small and rounded. Nostril single, with short tube, level with mid-orbit (or dorsal part of orbit). Cephalic sensory pores of moderate size: nasal pores 2, maxillary pores 6, preoperculomandibular pores 7, suprabranchial pores 2; cephalic pore pattern 2-6-7-2. Chin pores paired in separate pits, well separated. Coronal pore absent. Gill slit moderately large, upper margin level with mid-orbit, extending ventrally to pectoral-fin ray 6 (611). Gill rakers 10, short, with tiny spines on tips (based on FAKU 146632 and FAKU 147742). Opercular flap rounded; tip at level with ventral rim of orbit. Pyloric caeca 20 (1423), thick and long, covering two-thirds of stomach, on center-left side of visceral cavity.</paragraph>
<paragraph id="8B9236E2FF8E38638B9DA9DED3BEFE12" blockId="11.[151,1437,150,681]" pageId="11" pageNumber="202">
Vertebrae 55 (5558), precaudal 9 (9 or 10) and caudal 46 (4648). Pleural ribs in 2 pairs (1 or 2 pairs), on abdominal vertebrae 8 and 9 (or 9 and 10). Dorsal-fin rays 50 (5052), not exserted. Anteriormost dorsal-fin pterygiophore rayless, inserted between neural spines 3 and 4 (or 2 and 3). Anal-fin rays 45 (4348), not exserted. Two (two or three) anal-fin pterygiophores anterior to 1
<superScript id="7C589BAAFF8E38638840A84AD312FEAE" attach="left" box="[794,805,259,273]" fontSize="6" pageId="11" pageNumber="202">st</superScript>
haemal spine, each bearing a single ray. Membrane of posterior dorsal- and anal-fin rays attached about equidistant to caudal fin. Caudal fin slightly rounded. Principal caudal-fin rays 8 (89), 4(34) and 4 (45) rays on upper and lower hypurals, respectively, dorsal procurrent rays 1 (rarely absent), ventral procurrent rays absent (or 1). Hypural plates fused with terminal vertebral centrum and parhypural, upper and lower hypural plate separated by narrow slit.
</paragraph>
<paragraph id="8B9236E2FF8E38638B9DA8FED387FD16" blockId="11.[151,1437,150,681]" pageId="11" pageNumber="202">
Pectoral fin deeply notched, with 34 (3136) rays; upper lobe with 21 (2126) rays, just reaching to (or not reaching) vertical of anal fin origin; lower lobe elongate, with 13 (913) rays, 6
<superScript id="7C589BAAFF8E386388ADA893D433FE57" attach="left" box="[1015,1028,474,488]" fontSize="6" pageId="11" pageNumber="202">th</superScript>
(or 5
<superScript id="7C589BAAFF8E38638F66A893D47EFE57" attach="none" box="[1084,1097,474,488]" fontSize="6" pageId="11" pageNumber="202">th</superScript>
) ray from ventralmost longest, longer than upper lobe, extending to two-thirds between anus and anal-fin origin. Rays in notch more widely spaced than rays of lobes. Tip of pectoral-fin rays of upper lobe slightly protruding, rays of lower lobe strongly exserted. Uppermost pectoral-fin base below level of ventral rim of orbit, level with lower jaw tip. Lowermost pectoral-fin base below anterior half of orbit. Pelvic disk rounded, minute and greatly reduced (
<figureCitation id="13162A67FF8E38638F1CAB22D497FD3A" box="[1094,1184,618,645]" captionStart="FIGURE 5" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[242,1344,705,1916]" captionTargetId="figure-284@11.[242,1344,705,1916]" captionTargetPageId="11" captionText="FIGURE 5. Careproctus spinulosus sp. nov., FAKU 146600, holotype, 201.0 mm SL, fresh condition (A), preserved condition (B), and ventral view of head region (C); ventral view of head region of FAKU 148864, paratype, 112.2 mm SL (D). Dotted red line and arrow indicates position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219222" httpUri="https://zenodo.org/record/13219222/files/figure.png" pageId="11" pageNumber="202">Fig. 5C</figureCitation>
). Anus much closer to pelvic disk than to anal fin origin, located below posterior rim of orbit.
</paragraph>
<caption id="DF52666AFF8E38638BCDAEDAD368F84C" ID-DOI="http://doi.org/10.5281/zenodo.13219222" ID-Zenodo-Dep="13219222" httpUri="https://zenodo.org/record/13219222/files/figure.png" pageId="11" pageNumber="202" startId="11.[151,250,1939,1963]" targetBox="[242,1344,705,1916]" targetPageId="11" targetType="figure">
<paragraph id="8B9236E2FF8E38638BCDAEDAD368F84C" blockId="11.[151,1437,1939,2035]" pageId="11" pageNumber="202">
<emphasis id="B959EAF0FF8E38638BCDAEDAD124F814" bold="true" box="[151,275,1939,1963]" pageId="11" pageNumber="202">FIGURE 5.</emphasis>
<taxonomicName id="4C2D4D61FF8E38638A43AEDDD230F814" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[281,519,1939,1963]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="11" pageNumber="202" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF8E38638A43AEDDD230F814" box="[281,519,1939,1963]" italics="true" pageId="11" pageNumber="202">Careproctus spinulosus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF8E38638956AEDAD26AF814" bold="true" box="[524,605,1939,1963]" pageId="11" pageNumber="202">
<taxonomicNameLabel id="A26A578BFF8E38638956AEDAD26AF814" box="[524,605,1939,1963]" pageId="11" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 146600, holotype, 201.0 mm SL, fresh condition (A), preserved condition (B), and ventral view of head region (C); ventral view of head region of FAKU 148864, paratype, 112.2 mm SL (D). Dotted red line and arrow indicates position of anus and pelvic disk, respectively.
</paragraph>
</caption>
<paragraph id="8B9236E2FF8938648B9DA9DED52BFEFE" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
Pectoral girdle (based on FAKU 146632:
<figureCitation id="13162A67FF89386489CEA9DED2DAFF0E" box="[660,749,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="12" pageNumber="203">Fig. 2D</figureCitation>
): Proximal pectoral radials 3; upper radials 1 and 2 hour-glass shaped, with two notches, placed close to each other; radial 3 rounded squarish, widely separated from upper two radials. Interradial fenestrae 3, extending between scapula and radials 12, and below radial 2; fenestra 1 roundish, fenestrae 2 and 3 elliptical. Scapula broadly Y-shaped, with robust helve. Coracoid diamond-shaped, with short thin helve. Distal radials present at base of pectoral-fin rays 2 to 24, absent from base of ventralmost 9 rays.
</paragraph>
<paragraph id="8B9236E2FF8938648B9DA802D299FE6E" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">Coloration: When fresh (based on FAKU 146600), head and body dark reddish, belly somewhat paler. Iris dark, margined with silver. Dorsal, anal, and caudal fins pale red, mottled with dark red; pectoral fin dark red, basal part paler. Pelvic disk white. When preserved, head, body, and fins white. Eyes black. Oral and branchial cavities pale; peritoneum and stomach dark brown (or black).</paragraph>
<paragraph id="8B9236E2FF8938648B9DA893D4D5FD82" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DA893D144FE4B" bold="true" box="[199,371,474,500]" pageId="12" pageNumber="203">Reproduction.</emphasis>
The
<typeStatus id="54968840FF8938648AE8A892D223FE4A" box="[434,532,475,501]" pageId="12" pageNumber="203" type="holotype">holotype</typeStatus>
(201.0 mm SL) was an immature female. The ovary was whitish, with small rounded eggs less than
<quantity id="4CD59B07FF8938648ACDA8B7D1CDFDA6" box="[407,506,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="203" unit="mm" value="0.5">0.50 mm</quantity>
diameter. FAKU 146591 (
<quantity id="4CD59B07FF8938648847A8B6D3B8FDA6" box="[797,911,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.906" pageId="12" pageNumber="203" unit="mm" value="190.6">190.6 mm</quantity>
SL) was a spent female with yolked (
<quantity id="4CD59B07FF8938648E71A8B6D0F4FD82" metricMagnitude="-3" metricUnit="m" metricValue="3.105" metricValueMax="3.3" metricValueMin="2.91" pageId="12" pageNumber="203" unit="mm" value="3.105" valueMax="3.3" valueMin="2.91">2.913.30 mm</quantity>
diameter) and small immature (less than
<quantity id="4CD59B07FF89386489C9AB6BD2C0FD82" box="[659,759,546,573]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="203" unit="mm" value="0.5">0.50 mm</quantity>
diameter) eggs. Yolked eggs were rounded.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648B9DAB0FD162FD3A" pageId="12" pageNumber="203" type="distribution">
<paragraph id="8B9236E2FF8938648B9DAB0FD162FD3A" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DAB0FD156FDDF" bold="true" box="[199,353,582,608]" pageId="12" pageNumber="203">Distribution.</emphasis>
Known only from the southern Sea of Okhotsk, off Rausu,
<collectingRegion id="49E9F800FF8938648F50AB0ED44CFDDE" box="[1034,1147,583,609]" country="Japan" name="Hokkaido" pageId="12" pageNumber="203">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF8938648FD2AB0ED4FCFDDE" box="[1160,1227,583,609]" name="Japan" pageId="12" pageNumber="203">Japan</collectingCountry>
, at depths of
<quantity id="4CD59B07FF8938648E3EAB0FD0C4FD3A" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="12" pageNumber="203" unit="m" value="850.0" valueMax="1200.0" valueMin="500.0">500 1,200 m</quantity>
(
<figureCitation id="13162A67FF8938648A58AB22D170FD3A" box="[258,327,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,729,753]" captionTargetBox="[175,1410,188,699]" captionTargetId="figure-20@6.[171,1416,181,706]" captionTargetPageId="6" captionText="FIGURE 3. Distributional records of species of Careproctus with a reduced pelvic disk based on the present and previous studies (see text)." figureDoi="http://doi.org/10.5281/zenodo.13219216" httpUri="https://zenodo.org/record/13219216/files/figure.png" pageId="12" pageNumber="203">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648B9DABC6D374FD72" pageId="12" pageNumber="203" type="etymology">
<paragraph id="8B9236E2FF8938648B9DABC6D374FD72" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DABC6D179FD16" bold="true" box="[199,334,655,681]" pageId="12" pageNumber="203">Etymology.</emphasis>
The specific name
<taxonomicName id="4C2D4D61FF8938648975ABC6D285FD16" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[559,690,655,681]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="spinulosus">
<emphasis id="B959EAF0FF8938648966ABC6D285FD16" box="[572,690,655,681]" italics="true" pageId="12" pageNumber="203">spinulosus</emphasis>
</taxonomicName>
” is derived from the diminutive form of Latin “spina” (spine), in reference to the cactus-like prickles covering the entire body.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF8938648B9DAB9ED188FBC2" pageId="12" pageNumber="203" type="discussion">
<paragraph id="8B9236E2FF8938648B9DAB9ED0CDFC3E" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
<emphasis id="B959EAF0FF8938648B9DAB9ED10CFD4E" bold="true" box="[199,315,727,753]" pageId="12" pageNumber="203">Remarks.</emphasis>
Among the species with a reduced pelvic disk,
<taxonomicName id="4C2D4D61FF893864881CAB9ED3EBFD4E" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[838,988,727,753]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF893864881CAB9ED3EBFD4E" box="[838,988,727,753]" italics="true" pageId="12" pageNumber="203">C. spinulosus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF89386488B8AB9ED40DFD4E" bold="true" box="[994,1082,727,753]" pageId="12" pageNumber="203">
<taxonomicNameLabel id="A26A578BFF89386488B8AB9ED40DFD4E" box="[994,1082,727,753]" pageId="12" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares a lower pectoral-fin lobe apparently longer than the upper lobe with
<taxonomicName id="4C2D4D61FF8938648925ABB2D2C5FCAA" box="[639,754,763,789]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF8938648925ABB2D2C5FCAA" box="[639,754,763,789]" italics="true" pageId="12" pageNumber="203">C. gilberti</emphasis>
</taxonomicName>
, but can be distinguished from the latter in having a smaller gill opening, extending ventrally to pectoral-fin rays 511 (vs. 1416), and the body completely covered by cactus-like prickles (vs. smooth, without cactus-like prickles) (
<bibRefCitation id="EFBC4B13FF893864885FAA0AD429FCE2" author="Mecklenburg, C. W. &amp; Mecklenburg, T. A. &amp; Thorsteinson, L. K." box="[773,1054,835,861]" pageId="12" pageNumber="203" refId="ref18345" refString="Mecklenburg, C. W., Mecklenburg, T. A. &amp; Thorsteinson, L. K. (2002) Fishes of Alaska. American Fisheries Society, Bethesda, Maryland, 1037 pp." type="book" year="2002">
Mecklenburg
<emphasis id="B959EAF0FF89386488F9AA0AD3E3FCE2" box="[931,980,835,861]" italics="true" pageId="12" pageNumber="203">et al</emphasis>
. 2002
</bibRefCitation>
; see also Remarks under
<emphasis id="B959EAF0FF8938648E62AA0AD565FCE2" box="[1336,1362,835,861]" italics="true" pageId="12" pageNumber="203">C.</emphasis>
sp.
<taxonomicName id="4C2D4D61FF8938648E25AA0AD0DEFC3E" family="Liparidae" genus="Careproctus" isUncertain="true" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="gilberti">
cf.
<emphasis id="B959EAF0FF8938648BCDAA2ED0DEFC3E" box="[151,233,871,897]" italics="true" pageId="12" pageNumber="203">gilberti</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9236E2FF8938648B9DAAC2D188FBC2" blockId="12.[151,1437,151,1149]" pageId="12" pageNumber="203">
The
<taxonomicName id="4C2D4D61FF8938648BA2AAC2D14AFC1B" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[248,381,907,933]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="rastrinus">
<emphasis id="B959EAF0FF8938648BA2AAC2D14AFC1B" box="[248,381,907,933]" italics="true" pageId="12" pageNumber="203">C. rastrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D61FF8938648AEBAAC2D208FC1A" authorityName="Kroyer" authorityYear="1862" box="[433,575,907,933]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="reinhardti">
<emphasis id="B959EAF0FF8938648AEBAAC2D208FC1A" box="[433,575,907,933]" italics="true" pageId="12" pageNumber="203">C. reinhardti</emphasis>
</taxonomicName>
species groups are also characterized as having a long pectoral fin lower lobe, except for
<taxonomicName id="4C2D4D61FF8938648A53AAE6D269FC76" authority="(Orr et al. 2015)" baseAuthorityName="Orr" baseAuthorityYear="2015" box="[265,606,942,969]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="acanthodes">
<emphasis id="B959EAF0FF8938648A53AAE6D190FC76" box="[265,423,943,969]" italics="true" pageId="12" pageNumber="203">C. acanthodes</emphasis>
(
<bibRefCitation id="EFBC4B13FF8938648AE9AAE6D261FC76" author="Orr, J. W. &amp; Kai, Y. &amp; Nakabo, T." box="[435,598,942,969]" pageId="12" pageNumber="203" pagination="301 - 348" refId="ref18697" refString="Orr, J. W., Kai, Y. &amp; Nakabo, T. (2015) Snailfishes of the Careproctus rastrinus complex (Liparidae): redescriptions of seven species in the North Pacific Ocean region, with the description of a new species from the Beaufort Sea. Zootaxa, 4018 (3), 301 - 348. https: // doi. org / 10.11646 / zootaxa. 4018.3.1" type="journal article" year="2015">
Orr
<emphasis id="B959EAF0FF8938648ABAAAE6D239FC76" box="[480,526,943,969]" italics="true" pageId="12" pageNumber="203">et al</emphasis>
. 2015
</bibRefCitation>
)
</taxonomicName>
.As stated above, the members of both species groups have a relatively larger pelvic disk (2.29.6% SL) than
<taxonomicName id="4C2D4D61FF8938648AA3AA9AD2A7FC52" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[505,656,979,1005]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF8938648AA3AA9AD2A7FC52" box="[505,656,979,1005]" italics="true" pageId="12" pageNumber="203">C. spinulosus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF89386489CCAA9AD2D8FC52" bold="true" box="[662,751,979,1005]" pageId="12" pageNumber="203">
<taxonomicNameLabel id="A26A578BFF89386489CCAA9AD2D8FC52" box="[662,751,979,1005]" pageId="12" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(0.41.4% SL). The new species can be further distinguished from the former in having a black peritoneum (vs. light or gray) (
<bibRefCitation id="EFBC4B13FF8938648822AABED412FBAE" author="Orr, J. W. &amp; Kai, Y. &amp; Nakabo, T." box="[888,1061,1014,1041]" pageId="12" pageNumber="203" pagination="301 - 348" refId="ref18697" refString="Orr, J. W., Kai, Y. &amp; Nakabo, T. (2015) Snailfishes of the Careproctus rastrinus complex (Liparidae): redescriptions of seven species in the North Pacific Ocean region, with the description of a new species from the Beaufort Sea. Zootaxa, 4018 (3), 301 - 348. https: // doi. org / 10.11646 / zootaxa. 4018.3.1" type="journal article" year="2015">
Orr
<emphasis id="B959EAF0FF89386488F2AABED3EDFBAE" box="[936,986,1015,1041]" italics="true" pageId="12" pageNumber="203">et al</emphasis>
. 2015
</bibRefCitation>
).
<taxonomicName id="4C2D4D61FF8938648F62AABED0EFFB8A" authority="(Gilbert, 1915)" baseAuthorityName="Gilbert" baseAuthorityYear="1915" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="caudatus">
<emphasis id="B959EAF0FF8938648F62AABED502FBAE" box="[1080,1333,1015,1041]" italics="true" pageId="12" pageNumber="203">Elassodiscus caudatus</emphasis>
(
<bibRefCitation id="EFBC4B13FF8938648E1EAABED0E7FB8A" author="Gilbert, C. H." pageId="12" pageNumber="203" pagination="305 - 380" refId="ref17573" refString="Gilbert, C. H. (1915) Fishes collected by the United States Fisheries steamer &quot; Albatross &quot; in southern California in 1904. Proceedings of the United States National Museum, 48 (2075), 305 - 380, pls. 14 - 22. https: // doi. org / 10.5479 / si. 00963801.48 - 2075.305" type="journal article" year="1915">Gilbert, 1915</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C2D4D61FF8938648A57AD52D41BFB8A" authority="Kai, Matsuzaki, Orr, Mori &amp; Kamiunten, 2020" authorityName="Kai, Matsuzaki, Orr, Mori &amp; Kamiunten" authorityYear="2020" box="[269,1068,1051,1077]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="nyctereutes">
<emphasis id="B959EAF0FF8938648A57AD52D216FB8B" box="[269,545,1051,1077]" italics="true" pageId="12" pageNumber="203">Elassodiscus nyctereutes</emphasis>
<bibRefCitation id="EFBC4B13FF8938648972AD52D41BFB8A" author="Matsuzaki, K. &amp; Mori, T. &amp; Kamiunten, M. &amp; Yanagimoto, T. &amp; Kai, Y." box="[552,1068,1051,1077]" pageId="12" pageNumber="203" pagination="399 - 407" refId="ref18267" refString="Matsuzaki, K., Mori, T., Kamiunten, M., Yanagimoto, T. &amp; Kai, Y. (2020) A new species of Careproctus (Cottoidei: Liparidae) from the Sea of Okhotsk and a redescription of the blacktip snailfish Careproctus zachirus. Ichthyological Research, 67, 399 - 407. https: // doi. org / 10.1007 / s 10228 - 020 - 00734 - w" type="journal article" year="2020">Kai, Matsuzaki, Orr, Mori &amp; Kamiunten, 2020</bibRefCitation>
</taxonomicName>
mostly resemble the new species in having a long pectoral fin lower lobe, but clearly differ in having trilobed teeth (vs. simple, shouldered or weakly trilobed) (
<bibRefCitation id="EFBC4B13FF8938648A5CAD2AD184FBC2" author="Kai, Y. &amp; Matsuzaki, K. &amp; Orr, J. W. &amp; Mori, T. &amp; Kamiunten, M." box="[262,435,1123,1149]" pageId="12" pageNumber="203" pagination="55 - 66" refId="ref17899" refString="Kai, Y., Matsuzaki, K., Orr, J. W., Mori, T. &amp; Kamiunten, M. (2020) A new species of Elassodiscus (Cottoidei: Liparidae) from the North Pacific with an emended diagnosis of the genus. Ichthyological Research, 68, 55 - 66. https: // doi. org / 10.1007 / s 10228 - 020 - 00764 - 4" type="journal article" year="2020">
Kai
<emphasis id="B959EAF0FF8938648A6DAD2AD15EFBC2" box="[311,361,1123,1149]" italics="true" pageId="12" pageNumber="203">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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@ -0,0 +1,364 @@
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<emphasis id="B94DF959BD16CC6496826EB06A54E2D9" bold="true" box="[151,346,403,430]" italics="true" pageId="3" pageNumber="282">Mesophylax isly</emphasis>
Ibrahimi, Taybi &amp; Mabrouki
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<taxonomicNameLabel id="A27E4422BD16CC6494C16EB0683FE2DA" box="[724,817,403,429]" pageId="3" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B86254BBD16CC6496826E946A7DE2A6" blockId="3.[151,817,403,465]" box="[151,371,439,465]" pageId="3" pageNumber="282">
(
<figureCitation id="130239CEBD16CC64968A6E946A26E2A6" box="[159,296,439,465]" captionStart="FIGURES 26" captionStartId="4.[151,264,1821,1845]" captionTargetBox="[260,1326,184,1798]" captionTargetId="figure-25@4.[258,1329,181,1798]" captionTargetPageId="4" captionText="FIGURES 26. Male and female genitalia of Mesophylax isly sp. nov. 24, male genitalia: 2, left lateral; 3, ventrocaudal; 4, dorsal; 5, 6, female genitalia: 5, left lateral; 6, ventral. Scale bar = 1 mm. ae = aedeagus; af = axillary fossa (paired); ia = intermediate appendage (paired); ifa = inferior appendage (paired); il-ifa = inner lobe of an inferior appendage; ll = lateral lobe (paired) of vulvar scale; lsh = lateral shoulder of segment IX (paired); ml = mesal lobe of vulvar scale; pa = paramere (paired); sa = superior appendage (paired); sIX = segment IX; sp = supragenital plate; sVIII = segment VIII; sX = segment X; vs = vulvar scale." pageId="3" pageNumber="282">Figures 26</figureCitation>
,
<figureCitation id="130239CEBD16CC6497216E946A62E2A6" box="[308,364,439,465]" captionStart="FIGURES 711" captionStartId="5.[151,264,1299,1323]" captionTargetBox="[164,1423,181,1275]" captionTargetId="figure-20@5.[164,1423,181,1275]" captionTargetPageId="5" captionText="FIGURES 711. Photos of genitalia of Mesophylax aspersus aspersus Rambur 1842, Mesophylax impunctatus aduncus Navás 1923 and Mesophylax isly sp. nov. 7, M. a. aspersus from Guenfouda, Morocco, male, left lateral; 8, M. i. aduncus from Ballgjaj, Albania, male, left lateral; 9, M. isly sp. nov., male, left lateral; 10, M. isly sp. nov., female, left lateral; 11, Parameres (paired) and aedeagus of M. isly sp. nov., left lateral. Scale bar = 1 mm." pageId="3" pageNumber="282">711</figureCitation>
)
</paragraph>
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<emphasis id="B94DF959BD16CC6496826EDC6A48E16E" bold="true" box="[151,326,511,537]" pageId="3" pageNumber="282">Type material.</emphasis>
<materialsCitation id="3B512F16BD16CC6497586EDC69DCE14A" collectingDate="2018-06-03" collectorName="Y. Mabrouki &amp; A. F. Taybi." country="Morocco" county="Isly River" elevation="454" latitude="34.6094" location="Eastern Region" longLatPrecision="6" longitude="-1.8602" municipality="Aval Oued" pageId="3" pageNumber="282" specimenCount="1" typeStatus="holotype">
<emphasis id="B94DF959BD16CC6497586EDC6AB4E16E" bold="true" box="[333,442,511,537]" pageId="3" pageNumber="282">
<typeStatus id="54829BE9BD16CC6497586EDC6AB4E16E" box="[333,442,511,537]" pageId="3" pageNumber="282" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingCountry id="F32E65DBBD16CC64940F6D23698CE16D" box="[538,642,512,538]" name="Morocco" pageId="3" pageNumber="282">Morocco</collectingCountry>
,
<location id="8EE67390BD16CC64949B6D236830E16E" LSID="urn:lsid:plazi:treatment:0390945DBD16CC6196826EB06FB3E38E:8EE67390BD16CC64949B6D236830E16E" box="[654,830,511,538]" country="Morocco" county="Isly River" latitude="34.6094" longLatPrecision="6" longitude="-1.8602" municipality="Aval Oued" name="Eastern Region" pageId="3" pageNumber="282">Eastern Region</location>
,
<collectingMunicipality id="6BE2BF31BD16CC64955D6EDC68CDE16E" box="[840,963,511,537]" pageId="3" pageNumber="282">Aval Oued</collectingMunicipality>
,
<collectingCounty id="62E75DC7BD16CC6495DA6EDC6F4FE16E" box="[975,1089,511,537]" pageId="3" pageNumber="282">Isly River</collectingCounty>
,
<geoCoordinate id="EE0D438CBD16CC6492586EDC6FC4E16E" box="[1101,1226,511,537]" degrees="34.6094" direction="north" orientation="latitude" pageId="3" pageNumber="282" precision="5" value="34.6094">34.6094°N</geoCoordinate>
,
<geoCoordinate id="EE0D438CBD16CC6492C36EDC6E47E16E" box="[1238,1353,511,537]" degrees="1.8602" direction="west" orientation="longitude" pageId="3" pageNumber="282" precision="5" value="-1.8602">1.8602°W</geoCoordinate>
,
<elevation id="0014C278BD16CC6493406EDC6BC3E14A" metricMagnitude="2" metricUnit="m" metricValue="4.54" pageId="3" pageNumber="282" unit="m" value="454.0">
<quantity id="4CC188AEBD16CC6493406EDC6E92E16D" box="[1365,1436,511,538]" metricMagnitude="2" metricUnit="m" metricValue="4.54" pageId="3" pageNumber="282" unit="m" value="454.0">454 m</quantity>
a.s.l.
</elevation>
,
<date id="FF87038BBD16CC6496CC6D006A5DE14A" box="[217,339,547,573]" pageId="3" pageNumber="282" value="2018-06-03">
<collectingDate id="EFC3FA63BD16CC6496CC6D006A5DE14A" box="[217,339,547,573]" pageId="3" pageNumber="282" value="2018-06-03">03.vi.2018</collectingDate>
</date>
, leg.
<collectorName id="26CC409DBD16CC6497986D076914E14A" box="[397,538,547,574]" pageId="3" pageNumber="282">Y. Mabrouki</collectorName>
and
<collectorName id="26CC409DBD16CC64945A6D0069DCE14A" box="[591,722,547,574]" pageId="3" pageNumber="282">A. F. Taybi.</collectorName>
</materialsCitation>
<materialsCitation id="3B512F16BD16CC6494CD6D006BF3E116" collectingDate="2018-06-03" collectorName="Y. Mabrouki &amp; A. F. Taybi." country="Morocco" county="Isly River" elevation="454" latitude="34.6094" location="Eastern Region" longLatPrecision="6" longitude="-1.8602" municipality="Aval Oued" pageId="3" pageNumber="282" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B94DF959BD16CC6494CD6D00685EE14A" bold="true" box="[728,848,547,573]" pageId="3" pageNumber="282">
<typeStatus id="54829BE9BD16CC6494CD6D00685EE14A" box="[728,848,547,573]" pageId="3" pageNumber="282" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D3FEEC2BD16CC6495756D0068B7E14A" box="[864,953,547,573]" count="3" pageId="3" pageNumber="282" type="male">3 males</specimenCount>
,
<specimenCount id="9D3FEEC2BD16CC6495D16D006F38E14A" box="[964,1078,547,573]" count="2" pageId="3" pageNumber="282" type="female">2 females</specimenCount>
): with the same label data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32376C0BD16CC6496D26D486804E1F2" box="[199,778,619,646]" pageId="3" pageNumber="282" type="distribution">
<paragraph id="8B86254BBD16CC6496D26D486804E1F2" blockId="3.[151,1437,511,1978]" box="[199,778,619,646]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D26D486A6FE1F2" bold="true" box="[199,353,619,645]" pageId="3" pageNumber="282">Distribution.</emphasis>
<collectingCountry id="F32E65DBBD16CC6497726D4F6AC1E1F1" box="[359,463,620,646]" name="Morocco" pageId="3" pageNumber="282">Morocco</collectingCountry>
, Eastern Region, Isly River.
</paragraph>
</subSubSection>
<subSubSection id="C32376C0BD16CC6496D26DAC69ECE79E" pageId="3" pageNumber="282" type="diagnosis">
<paragraph id="8B86254BBD16CC6496D26DAC6981E09A" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D26DAC6A4CE1DE" bold="true" box="[199,322,655,681]" pageId="3" pageNumber="282">Diagnosis.</emphasis>
The male of the new species is most similar to that of
<taxonomicName id="4C395EC8BD16CC6495B56DAC6E10E1DE" authorityName="Malicky" authorityYear="1988" box="[928,1310,655,681]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="subSpecies" species="aspersus" subSpecies="hoggarensis">
<emphasis id="B94DF959BD16CC6495B56DAC6E10E1DE" box="[928,1310,655,681]" italics="true" pageId="3" pageNumber="282">Mesophylax aspersus hoggarensis</emphasis>
</taxonomicName>
and differs mainly in exhibiting (1) high, broad inferior appendages, higher than intermediate appendages, each with generally straight inner and outer margins in lateral view; (2) apically longer than basally inferior appendages in ventral view; (3) apically placed inner lobes of inferior appendages in ventro-caudal view; (4) thick parameres in ventro-caudal view; (5) shallow axillary fossa of inferior appendages. The
<emphasis id="B94DF959BD16CC6495596C036F17E04E" box="[844,1049,799,825]" italics="true" pageId="3" pageNumber="282">M. a. hoggarensis</emphasis>
male has (1) low, narrow inferior appendages, lower than intermediate appendages, each with sinuate inner and outer margins in lateral view; (2) inferior appendages basally longer than apically in ventrocaudal view; (3) mesally placed inner lobes of inferior appendages in ventrocaudal view; (4) slender parameres in ventrocaudal view; (5) deep axillary fossae of inferior appendages. In addition, males of
<taxonomicName id="4C395EC8BD16CC64940D6C93696BE0BE" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[536,613,943,969]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD16CC64940D6C93696BE0BE" box="[536,613,943,969]" italics="true" pageId="3" pageNumber="282">M. isly</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E4422BD16CC6494796C8C69C9E0BD" box="[620,711,943,970]" pageId="3" pageNumber="282" rank="species">
<emphasis id="B94DF959BD16CC6494796C8C69CFE0BE" bold="true" box="[620,705,943,969]" pageId="3" pageNumber="282">sp. nov</emphasis>
.
</taxonomicNameLabel>
are distinguished from those of
<emphasis id="B94DF959BD16CC6492216C936FF0E0BE" box="[1076,1278,943,969]" italics="true" pageId="3" pageNumber="282">M. a. hoggarensis</emphasis>
by their more curved and shorter parameres in lateral view.
</paragraph>
<paragraph id="8B86254BBD16CC6496D26CD469ECE79E" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
The female of
<taxonomicName id="4C395EC8BD16CC6497636CD46921E766" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[374,559,1015,1041]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD16CC6497636CD46921E766" box="[374,559,1015,1041]" italics="true" pageId="3" pageNumber="282">Mesophylax isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD16CC64942C6CD46998E766" bold="true" box="[569,662,1015,1041]" pageId="3" pageNumber="282">
<taxonomicNameLabel id="A27E4422BD16CC64942C6CD46998E766" box="[569,662,1015,1041]" pageId="3" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to that of
<emphasis id="B94DF959BD16CC6495DA6CDB6F91E766" box="[975,1183,1015,1041]" italics="true" pageId="3" pageNumber="282">M. a. hoggarensis</emphasis>
from which it differs by exhibiting (1) a downward-arching, apically blunt segment X in lateral view; (2) a short, apically rounded supragenital plate in lateral view; (3) an apically bipartite mesal lobe of the vulvar scale in ventral view; and (4) apically rounded, small lateral shoulders of segment IX in lateral view. The female of
<emphasis id="B94DF959BD16CC6492726B476E3CE70A" box="[1127,1330,1123,1149]" italics="true" pageId="3" pageNumber="282">M. a. hoggarensis</emphasis>
has (1) a mesad-arching, apically tapering segment X in lateral view; (2) a long, rounded supragenital plate in lateral view; (3) a unipartite, apically slightly convex mesal lobe of the vulvar scale in ventral view; and (4) apically acuminate, large lateral shoulders of segment IX in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C32376C0BD16CC6196D26BD06FB3E38E" lastPageId="6" lastPageNumber="285" pageId="3" pageNumber="282" type="description">
<paragraph id="8B86254BBD16CC6496D26BD069D0E60E" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D26BD06A57E67A" bold="true" box="[199,345,1267,1293]" pageId="3" pageNumber="282">Description.</emphasis>
<emphasis id="B94DF959BD16CC64974A6BD0694AE67A" box="[351,580,1267,1293]" italics="true" pageId="3" pageNumber="282">General appearance</emphasis>
. Habitus brown, sclerites and tergites dark brown; cephalic and thoracic setal areas pale; cephalic, thoracic, and abdominal setation blonde; legs brown to fawn, with darker areas; wings brown with darker nuances, with fine blonde setae. Length of each forewing
<quantity id="4CC188AEBD16CC6495B86A186F1CE621" box="[941,1042,1339,1366]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="282" unit="mm" value="14.5">14.5 mm</quantity>
(
<typeStatus id="54829BE9BD16CC6492346A186F87E622" box="[1057,1161,1339,1365]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
), 14.516.0 mm (n = 4). Male spur formula 0,2,2; female spur formula 1,2,2.
</paragraph>
<paragraph id="8B86254BBD16CC6496D26AA06A69E5EE" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D26AA06A7FE6EA" box="[199,369,1411,1437]" italics="true" pageId="3" pageNumber="282">Male genitalia.</emphasis>
Segment IX fused with basal parts of inferior appendages, with narrow acuminate dorsal portion and longer ventral portion, laterally even longer, convex anteriorly, in lateral view. Superior appendages apically bipartite with round ends in lateral view. Intermediate appendages wide basally, tapering apically, downward arching in lateral view. Free parts of inferior appendages broad, triangular, acute apically, each with posterior edge notched basally in lateral view. Inner lobe of free part of each inferior appendage visible in lateral view as small subapical triangle; in ventral view broadly quadratic, blunt dorsoapically, elongate, acute ventroapically. Phallic apparatus consisting of simple aedeagus and pair of parameres; in ventral view parameres broader basally, acute apices as high as tip of aedeagus.
</paragraph>
<paragraph id="8B86254BBD16CC6496D26980680DE43A" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D269806A8BE5CA" box="[199,389,1699,1725]" italics="true" pageId="3" pageNumber="282">Female genitalia</emphasis>
. Segment IX setation abundant, more abundant in the dorsal half and along the lateral margins. Segment X triangular in lateral view, blunt apically in dorsal view; in ventral view wide along basal half and parallelsided apically. Supragenital plate apically round in lateral view, shorter than segment X; vulvar scale even shorter, elongate, subtriangular. Lateral shoulders of segment IX membranous, with sclerotized edges, long ventrally, short dorsally, generally triangular, turned mesad posteriorly.
</paragraph>
<paragraph id="8B86254BBD16CC6496D2687B6AF0E406" blockId="3.[151,1437,511,1978]" box="[199,510,1879,1905]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D2687B6A8CE406" box="[199,386,1879,1905]" italics="true" pageId="3" pageNumber="282">Larva and pupa.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B86254BBD16CC6496D2685869B4E4CE" blockId="3.[151,1437,511,1978]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496D268586A4BE4E2" box="[199,325,1915,1941]" italics="true" pageId="3" pageNumber="282">Etymology.</emphasis>
The species epithet
<taxonomicName id="4C395EC8BD16CC64943368586952E4E2" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[550,604,1915,1941]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="isly">“isly</taxonomicName>
” refers to the Isly River, where the new species was discovered, and, in the Amazigh language, it translates to “new groom.”
</paragraph>
<caption id="DF4675C3BD11CC639682683E6BDFE49D" pageId="4" pageNumber="283" startId="4.[151,264,1821,1845]" targetBox="[260,1326,184,1798]" targetPageId="4" targetType="figure">
<paragraph id="8B86254BBD11CC639682683E6BDFE49D" blockId="4.[151,1437,1821,2026]" pageId="4" pageNumber="283">
<emphasis id="B94DF959BD11CC639682683E6A33E442" bold="true" box="[151,317,1821,1845]" pageId="4" pageNumber="283">FIGURES 26.</emphasis>
Male and female genitalia of
<taxonomicName id="4C395EC8BD11CC639468683D682DE441" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[637,803,1822,1846]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="283" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD11CC639468683D682DE441" box="[637,803,1822,1846]" italics="true" pageId="4" pageNumber="283">Mesophylax isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD11CC639538683D688EE441" bold="true" box="[813,896,1822,1846]" pageId="4" pageNumber="283">
<taxonomicNameLabel id="A27E4422BD11CC639538683D688EE441" box="[813,896,1822,1846]" pageId="4" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
24, male genitalia: 2, left lateral; 3, ventrocaudal; 4, dorsal; 5, 6, female genitalia: 5, left lateral; 6, ventral. Scale bar = 1 mm. ae = aedeagus; af = axillary fossa (paired); ia = intermediate appendage (paired); ifa = inferior appendage (paired); il-ifa = inner lobe of an inferior appendage; ll = lateral lobe (paired) of vulvar scale; lsh = lateral shoulder of segment IX (paired); ml = mesal lobe of vulvar scale; pa = paramere (paired); sa = superior appendage (paired); sIX = segment IX; sp = supragenital plate; sVIII = segment VIII; sX = segment X; vs = vulvar scale.
</paragraph>
</caption>
<caption id="DF4675C3BD10CC6296826A306860E6E0" pageId="5" pageNumber="284" startId="5.[151,264,1299,1323]" targetBox="[164,1423,181,1275]" targetPageId="5" targetType="figure">
<paragraph id="8B86254BBD10CC6296826A306860E6E0" blockId="5.[151,1437,1299,1432]" pageId="5" pageNumber="284">
<emphasis id="B94DF959BD10CC6296826A306A48E65B" bold="true" box="[151,326,1299,1324]" pageId="5" pageNumber="284">FIGURES 711.</emphasis>
Photos of genitalia of
<taxonomicName id="4C395EC8BD10CC62943E6A3768F8E65B" authority="Rambur 1842" authorityName="Rambur" authorityYear="1842" box="[555,1014,1300,1324]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="aspersus" subSpecies="aspersus">
<emphasis id="B94DF959BD10CC62943E6A37686DE65C" box="[555,867,1300,1324]" italics="true" pageId="5" pageNumber="284">Mesophylax aspersus aspersus</emphasis>
Rambur 1842
</taxonomicName>
,
<taxonomicName id="4C395EC8BD10CC6292156A376BC4E627" authority="Navas 1923" authorityName="Navas" authorityYear="1923" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="impunctatus" subSpecies="aduncus">
<emphasis id="B94DF959BD10CC6292156A376E58E65B" box="[1024,1366,1300,1324]" italics="true" pageId="5" pageNumber="284">Mesophylax impunctatus aduncus</emphasis>
Navás 1923
</taxonomicName>
and
<taxonomicName id="4C395EC8BD10CC6297176A1B6AA6E627" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[258,424,1336,1360]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD10CC6297176A1B6AA6E627" box="[258,424,1336,1360]" italics="true" pageId="5" pageNumber="284">Mesophylax isly</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E4422BD10CC6297A76A1B6909E638" box="[434,519,1336,1360]" pageId="5" pageNumber="284" rank="species">
<emphasis id="B94DF959BD10CC6297A76A1B690FE627" bold="true" box="[434,513,1336,1360]" pageId="5" pageNumber="284">sp. nov</emphasis>
.
</taxonomicNameLabel>
7,
<emphasis id="B94DF959BD10CC62943B6A1B69C6E638" box="[558,712,1336,1359]" italics="true" pageId="5" pageNumber="284">
M. a.
<taxonomicName id="4C395EC8BD10CC62947A6A1B69C6E638" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[623,712,1336,1359]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="aspersus">aspersus</taxonomicName>
</emphasis>
from Guenfouda, Morocco, male, left lateral; 8,
<emphasis id="B94DF959BD10CC6292C46A1B6E6FE627" box="[1233,1377,1336,1360]" italics="true" pageId="5" pageNumber="284">M. i. aduncus</emphasis>
from Ballgjaj, Albania, male, left lateral; 9,
<taxonomicName id="4C395EC8BD10CC6294096A7F696FE603" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[540,609,1372,1396]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD10CC6294096A7F696FE603" box="[540,609,1372,1396]" italics="true" pageId="5" pageNumber="284">M. isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD10CC6294726A7F69B6E603" bold="true" box="[615,696,1372,1396]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A27E4422BD10CC6294726A7F69B6E603" box="[615,696,1372,1396]" pageId="5" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, left lateral; 10,
<taxonomicName id="4C395EC8BD10CC62958B6A7F68EDE603" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[926,995,1372,1396]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD10CC62958B6A7F68EDE603" box="[926,995,1372,1396]" italics="true" pageId="5" pageNumber="284">M. isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD10CC6295FC6A7F6F34E603" bold="true" box="[1001,1082,1372,1396]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A27E4422BD10CC6295FC6A7F6F34E603" box="[1001,1082,1372,1396]" pageId="5" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, left lateral; 11, Parameres (paired) and aedeagus of
<taxonomicName id="4C395EC8BD10CC6297826AA36AD2E6EF" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[407,476,1408,1432]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD10CC6297826AA36AD2E6EF" box="[407,476,1408,1432]" italics="true" pageId="5" pageNumber="284">M. isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD10CC6297F76AA3693DE6EF" bold="true" box="[482,563,1408,1432]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A27E4422BD10CC6297F76AA3693DE6EF" box="[482,563,1408,1432]" pageId="5" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, left lateral. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8B86254BBD10CC6296D26AC16F86E5C7" blockId="5.[151,1436,1506,2042]" pageId="5" pageNumber="284">
<emphasis id="B94DF959BD10CC6296D26AC16A2AE68B" box="[199,292,1506,1532]" italics="true" pageId="5" pageNumber="284">Habitat.</emphasis>
The new species was found in one of the permanent tributaries of the Isly River (south of Oujda City), in a location known as Tayret. The river exhibits irregular flow patterns closely tied to rainfall, experiencing floods from autumn to spring and severe summer low flows nearly year-round. The intense sunlight fosters the growth of perilithon, filamentous algae, and green algae (
<taxonomicName id="4C395EC8BD10CC6294BC696D682AE51F" authorityName="Agardh" authorityYear="1824" box="[681,804,1614,1640]" class="Charophyceae" family="Characeae" kingdom="Plantae" order="Charales" pageId="5" pageNumber="284" phylum="Charophyta" rank="family">Characeae</taxonomicName>
). The sediment in the springs consists of blocks, stones, pebbles, sand, and occasionally mud. The vegetation along the banks is sparse, predominantly comprising reeds (
<taxonomicName id="4C395EC8BD10CC6296B469B56A4EE5C7" box="[161,320,1686,1712]" class="Liliopsida" family="Poaceae" genus="Arundo" kingdom="Plantae" order="Poales" pageId="5" pageNumber="284" phylum="Tracheophyta" rank="species" species="donax">
<emphasis id="B94DF959BD10CC6296B469B56A4EE5C7" box="[161,320,1686,1712]" italics="true" pageId="5" pageNumber="284">Arundo donax</emphasis>
</taxonomicName>
L.), juncus (
<emphasis id="B94DF959BD10CC6297C569B46960E5C7" box="[464,622,1687,1712]" italics="true" pageId="5" pageNumber="284">Juncus acutus</emphasis>
L.), and occasionally typha (
<taxonomicName id="4C395EC8BD10CC6295A069B56F57E5C7" box="[949,1113,1686,1712]" class="Charophyceae" family="Characeae" genus="Typha" kingdom="Plantae" order="Charales" pageId="5" pageNumber="284" phylum="Charophyta" rank="species" species="latifolia">
<emphasis id="B94DF959BD10CC6295A069B56F57E5C7" box="[949,1113,1686,1712]" italics="true" pageId="5" pageNumber="284">Typha latifolia</emphasis>
</taxonomicName>
L.).
</paragraph>
<paragraph id="8B86254BBD10CC6296D269996FC6E4DB" blockId="5.[151,1436,1506,2042]" pageId="5" pageNumber="284">
The habitat of the new species is populated by various benthic macroinvertebrates, including dragonflies such as
<taxonomicName id="4C395EC8BD10CC6296AC69FD696AE58F" authority="(Brulle 1832)" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[185,612,1758,1784]" class="Insecta" family="Libellulidae" genus="Crocothemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="erythraea">
<emphasis id="B94DF959BD10CC6296AC69FD6AB0E58F" box="[185,446,1758,1784]" italics="true" pageId="5" pageNumber="284">Crocothemis erythraea</emphasis>
(Brullé 1832)
</taxonomicName>
,
<taxonomicName id="4C395EC8BD10CC62946669FD6F8AE58F" authority="(Palisotde Beauvois 1807)" baseAuthorityName="Palisotde Beauvois" baseAuthorityYear="1807" box="[627,1156,1758,1784]" class="Insecta" family="Libellulidae" genus="Trithemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="B94DF959BD10CC62946669FD6845E58F" box="[627,843,1758,1784]" italics="true" pageId="5" pageNumber="284">Trithemis annulata</emphasis>
(Palisotde Beauvois 1807)
</taxonomicName>
,
<taxonomicName id="4C395EC8BD10CC62928669FD6A5FE46A" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Libellulidae" genus="Orthetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="B94DF959BD10CC62928669FD6E92E58F" box="[1171,1436,1758,1784]" italics="true" pageId="5" pageNumber="284">Orthetrum cancellatum</emphasis>
(Linnaeus 1758)
</taxonomicName>
(in both larval and adult stages); aquatic
<taxonomicName id="4C395EC8BD10CC62953C68216891E46B" box="[809,927,1794,1820]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
such as
<taxonomicName id="4C395EC8BD10CC62921668206E96E46B" authority="Fabricius 1798" authorityName="Fabricius" authorityYear="1798" box="[1027,1432,1794,1820]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B94DF959BD10CC62921668206FEBE46B" box="[1027,1253,1794,1820]" italics="true" pageId="5" pageNumber="284">Naucoris maculatus</emphasis>
Fabricius 1798
</taxonomicName>
, and
<taxonomicName id="4C395EC8BD10CC6296DD6804694AE436" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" box="[200,580,1830,1857]" class="Insecta" family="Notonectidae" genus="Notonecta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="glauca">
<emphasis id="B94DF959BD10CC6296DD68046A9EE437" box="[200,400,1830,1856]" italics="true" pageId="5" pageNumber="284">Notonecta glauca</emphasis>
Linnaeus 1758
</taxonomicName>
; aquatic coleopterans including
<taxonomicName id="4C395EC8BD10CC6295A668056E21E437" authority="(Forster 1771)" baseAuthorityName="Forster" baseAuthorityYear="1771" box="[947,1327,1830,1856]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="nebulosus">
<emphasis id="B94DF959BD10CC6295A668056F8EE437" box="[947,1152,1830,1856]" italics="true" pageId="5" pageNumber="284">Agabus nebulosus</emphasis>
(Forster 1771)
</taxonomicName>
; mayflies like
<taxonomicName id="4C395EC8BD10CC6296DE6869695FE413" authority="(Linnaeus 1761)" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[203,593,1866,1892]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="dipterum">
<emphasis id="B94DF959BD10CC6296DE68696A85E413" box="[203,395,1866,1892]" italics="true" pageId="5" pageNumber="284">Cloeon dipterum</emphasis>
(Linnaeus 1761)
</taxonomicName>
and
<taxonomicName id="4C395EC8BD10CC62949B68696F2DE413" authority="(Burmeister 1839)" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[654,1059,1866,1892]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="B94DF959BD10CC62949B6869684BE413" box="[654,837,1866,1892]" italics="true" pageId="5" pageNumber="284">Caenis luctuosa</emphasis>
(Burmeister 1839)
</taxonomicName>
, as well as dipteran larvae (e.g.,
<taxonomicName id="4C395EC8BD10CC629682684D6A33E4FF" box="[151,317,1902,1928]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
). Additionally, two aquatic vertebrates have been observed in the habitat:
<taxonomicName id="4C395EC8BD10CC6292B3684D6A36E4DB" authority="(Hartert 1913)" baseAuthorityName="Hartert" baseAuthorityYear="1913" class="Amphibia" family="Ranidae" genus="Pelophylax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="saharicus">
<emphasis id="B94DF959BD10CC6292B3684D6E92E4FF" box="[1190,1436,1902,1928]" italics="true" pageId="5" pageNumber="284">Pelophylax saharicus</emphasis>
(Hartert 1913)
</taxonomicName>
and
<taxonomicName id="4C395EC8BD10CC62977A68B06813E4DB" authority="(Schweigger 1812)" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[367,797,1938,1964]" class="Testudines" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="leprosa">
<emphasis id="B94DF959BD10CC62977A68B06930E4DB" box="[367,574,1938,1964]" italics="true" pageId="5" pageNumber="284">Mauremys leprosa</emphasis>
(Schweigger 1812)
</taxonomicName>
, both widely distributed in the region.
</paragraph>
<paragraph id="8B86254BBD10CC6196D268956FB3E38E" blockId="5.[151,1436,1506,2042]" lastBlockId="6.[151,1436,151,249]" lastPageId="6" lastPageNumber="285" pageId="5" pageNumber="284">
<taxonomicName id="4C395EC8BD10CC6296D268956A75E4A7" authorityName="Taybi &amp; Mabrouki &amp; Ibrahimi" authorityYear="2024" box="[199,379,1974,2000]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="284" phylum="Arthropoda" rank="species" species="isly" status="sp. nov.">
<emphasis id="B94DF959BD10CC6296D268956A75E4A7" box="[199,379,1974,2000]" italics="true" pageId="5" pageNumber="284">Mesophylax isly</emphasis>
</taxonomicName>
<emphasis id="B94DF959BD10CC62979468956AD4E4A7" bold="true" box="[385,474,1974,2000]" pageId="5" pageNumber="284">
<taxonomicNameLabel id="A27E4422BD10CC62979468956AD4E4A7" box="[385,474,1974,2000]" pageId="5" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to require habitats with good water quality. The analysis of the physicochemical parameters of the water (ammonium and BOD
<subScript id="17BD270EBD10CC6294BF68CF69BCE48D" attach="left" box="[682,690,2028,2042]" fontSize="6" pageId="5" pageNumber="284">5</subScript>
) showed a good environmental quality of the water, according to the Moroccan surface water guidelines of good quality (
<tableCitation id="C6BB10F0BD13CC6195106FB46857E3C6" box="[773,857,151,177]" captionStart="TABLE 1" captionStartId="6.[151,239,296,320]" captionText="TABLE 1. Mean (ranges) of values of the physical and chemical water parameters measured at the type locality of Mesophylax isly sp. nov. M (mean value), MSWG (Moroccan surface water guidelines of good quality)." pageId="6" pageNumber="285">Table 1</tableCitation>
). However, because it is located in an arid rural and agricultural environment, the habitat is disturbed by human activities, particularly water abstraction in summer, as is the case throughout the eastern region of
<collectingCountry id="F32E65DBBD13CC6194956FFC69EBE38E" box="[640,741,223,249]" name="Morocco" pageId="6" pageNumber="285">Morocco</collectingCountry>
(
<bibRefCitation id="EFA858BABD13CC6194E66FFC68EAE38E" author="Mabrouki, Y. &amp; Taybi, A. F. &amp; Chavanon, G. &amp; Millan, A. &amp; Berrahou, A." box="[755,996,223,249]" pageId="6" pageNumber="285" pagination="185 - 211" refId="ref4949" refString="Mabrouki, Y., Taybi, A. F., Chavanon, G., Millan, A., &amp; Berrahou, A. (2018) New and interesting data on the distribution of aquatic beetles from Morocco (Coleoptera, Polyphaga). Arxius de Miscellania Zoologica, 16, 185 - 211. https: // doi. org / 10.32800 / amz. 2018.16.0185" type="journal article" year="2018">
Mabrouki
<emphasis id="B94DF959BD13CC61957F6FFC68ADE38E" box="[874,931,223,249]" italics="true" pageId="6" pageNumber="285">et al.</emphasis>
2018
</bibRefCitation>
,
<bibRefCitation id="EFA858BABD13CC6195FA6FFC6FBFE38E" author="Taybi, A. F. &amp; Mabrouki, Y. &amp; Chavanon, G. &amp; Millan, A. &amp; Berrahou, A." box="[1007,1201,223,249]" pageId="6" pageNumber="285" pagination="139 - 163" refId="ref5662" refString="Taybi, A. F., Mabrouki, Y., Chavanon, G., Millan, A. &amp; Berrahou, A. (2018) New data on the distribution of aquatic bugs (Hemiptera) from Morocco with notes on their chorology. Zootaxa, 4459 (1), 139 - 163. https: // doi. org / 10.11646 / zootaxa. 4459.1.6" type="journal article" year="2018">
Taybi
<emphasis id="B94DF959BD13CC6192206FFC6F63E38E" box="[1077,1133,223,249]" italics="true" pageId="6" pageNumber="285">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,106 @@
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<mods:title id="C5A654331B9781AC86B11A7F05950A29">Description of Mesophylax isly sp. nov. (Trichoptera, Limnephilidae) from Morocco, and new genus records from the Balkans</mods:title>
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<paragraph id="8B86254BBD16CC6496826FB469C5E3C6" blockId="3.[151,715,151,213]" box="[151,715,151,178]" pageId="3" pageNumber="282">
<heading id="D0CE9227BD16CC6496826FB469C5E3C6" bold="true" box="[151,715,151,178]" fontSize="11" level="1" pageId="3" pageNumber="282" reason="1">
<taxonomicName id="4C395EC8BD16CC6496826FB469C5E3C6" ID-CoL="5JNB7" authority="Navas 1923" authorityName="Navas" authorityYear="1923" box="[151,715,151,178]" class="Insecta" family="Limnephilidae" genus="Mesophylax" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="subSpecies" species="impunctatus" subSpecies="aduncus">
<emphasis id="B94DF959BD16CC6496826FB469C5E3C6" bold="true" box="[151,715,151,178]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496826FB4693BE3C5" bold="true" box="[151,565,151,178]" italics="true" pageId="3" pageNumber="282">Mesophylax impunctatus aduncus</emphasis>
Navas 1923
</emphasis>
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<subSubSection id="C32376C0BD16CC6496826F986A06E3A2" box="[151,264,187,213]" pageId="3" pageNumber="282" type="description">
<paragraph id="8B86254BBD16CC6496826F986A06E3A2" blockId="3.[151,715,151,213]" box="[151,264,187,213]" pageId="3" pageNumber="282">
(
<figureCitation id="130239CEBD16CC64968A6F986BF1E3A2" box="[159,255,187,213]" captionStart="FIGURES 711" captionStartId="5.[151,264,1299,1323]" captionTargetBox="[164,1423,181,1275]" captionTargetId="figure-20@5.[164,1423,181,1275]" captionTargetPageId="5" captionText="FIGURES 711. Photos of genitalia of Mesophylax aspersus aspersus Rambur 1842, Mesophylax impunctatus aduncus Navás 1923 and Mesophylax isly sp. nov. 7, M. a. aspersus from Guenfouda, Morocco, male, left lateral; 8, M. i. aduncus from Ballgjaj, Albania, male, left lateral; 9, M. isly sp. nov., male, left lateral; 10, M. isly sp. nov., female, left lateral; 11, Parameres (paired) and aedeagus of M. isly sp. nov., left lateral. Scale bar = 1 mm." pageId="3" pageNumber="282">Figure 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32376C0BD16CC6496826E206AD6E236" pageId="3" pageNumber="282" type="materials_examined">
<paragraph id="8B86254BBD16CC6496826E206AD6E236" blockId="3.[151,1436,259,322]" pageId="3" pageNumber="282">
<emphasis id="B94DF959BD16CC6496826E206A86E26A" bold="true" box="[151,392,259,285]" pageId="3" pageNumber="282">Material examined:</emphasis>
<materialsCitation id="3B512F16BD16CC6497866E206AD6E236" collectingDate="2012-06-21" collectorName="H. Ibrahimi." country="Albania" county="Fushe Lure" latitude="41.794" location="Malla River" longLatPrecision="69" longitude="20.228" municipality="Ballgjaj" pageId="3" pageNumber="282" specimenCount="1" specimenCount-male="1" stateProvince="Diber">
<specimenCount id="9D3FEEC2BD16CC6497866E206AE8E26A" box="[403,486,259,285]" count="1" pageId="3" pageNumber="282" type="male">1 male</specimenCount>
,
<collectingCountry id="F32E65DBBD16CC6497E16E20695EE26A" box="[500,592,259,285]" name="Albania" pageId="3" pageNumber="282">Albania</collectingCountry>
,
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<collectingDate id="EFC3FA63BD16CC6496826E046A1FE236" box="[151,273,295,321]" pageId="3" pageNumber="282" value="2012-06-21">21.vi.2012</collectingDate>
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, leg.
<collectorName id="26CC409DBD16CC6497586E0B6AD6E236" box="[333,472,295,322]" pageId="3" pageNumber="282">H. Ibrahimi.</collectorName>
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View file

@ -0,0 +1,130 @@
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<emphasis id="B94DF959BD17CC659682680A6907E433" bold="true" box="[151,521,1833,1860]" italics="true" pageId="2" pageNumber="281">Mesophylax aspersus aspersus</emphasis>
Rambur 1842
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(
<figureCitation id="130239CEBD17CC65968A686E6A0EE410" box="[159,256,1869,1895]" captionStart="FIGURES 711" captionStartId="5.[151,264,1299,1323]" captionTargetBox="[164,1423,181,1275]" captionTargetId="figure-20@5.[164,1423,181,1275]" captionTargetPageId="5" captionText="FIGURES 711. Photos of genitalia of Mesophylax aspersus aspersus Rambur 1842, Mesophylax impunctatus aduncus Navás 1923 and Mesophylax isly sp. nov. 7, M. a. aspersus from Guenfouda, Morocco, male, left lateral; 8, M. i. aduncus from Ballgjaj, Albania, male, left lateral; 9, M. isly sp. nov., male, left lateral; 10, M. isly sp. nov., female, left lateral; 11, Parameres (paired) and aedeagus of M. isly sp. nov., left lateral. Scale bar = 1 mm." pageId="2" pageNumber="281">Figure 7</figureCitation>
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<emphasis id="B94DF959BD17CC65968268B66A8CE4D8" bold="true" box="[151,386,1941,1967]" pageId="2" pageNumber="281">Material examined:</emphasis>
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,
<collectingCountry id="F32E65DBBD17CC6597F468B56935E4C7" box="[481,571,1942,1968]" name="Kosovo" pageId="2" pageNumber="281">Kosovo</collectingCountry>
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i
<collectingMunicipality id="6BE2BF31BD17CC65948068B6686EE4D8" box="[661,864,1941,1967]" pageId="2" pageNumber="281">Bardhë watershed</collectingMunicipality>
, upstream area of the
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, leg.
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,
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,
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</date>
, leg.
<collectorName id="26CC409DBD17CC6594DE68FE684EE480" box="[715,832,2013,2039]" pageId="2" pageNumber="281">F.A. Taybi</collectorName>
&amp;
<collectorName id="26CC409DBD17CC65957168FD68F6E480" box="[868,1016,2013,2040]" pageId="2" pageNumber="281">Y. Mabrouki.</collectorName>
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View file

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<mods:part id="8BCBC449CE84D71D6732127F30087F93">
<mods:date id="2FC64A978D622BDD81888C20496B6A59">2024</mods:date>
<mods:detail id="FB5725C5519526CD35767F8891F8C9DC" type="pubDate">
<mods:number id="4648507CFB94EDED89CC67FD4446AAD7">2024-08-05</mods:number>
</mods:detail>
<mods:detail id="83A6A799D911CBD7F15E9D466F3491E8" type="volume">
<mods:number id="E38360C078B0A53DA4A4C205A4305417">5492</mods:number>
</mods:detail>
<mods:detail id="84AF8548C8A3513A4308D990D87A81AB" type="issue">
<mods:number id="C0E79D9509FE95C71D958A157F6EBE31">2</mods:number>
</mods:detail>
<mods:extent id="1E5B545FBCE8B6B650F31F08E21C21C1" unit="page">
<mods:start id="39979C854C111AF7E0EE97333EA768E4">151</mods:start>
<mods:end id="8B20CF54E80F892A5BEF68E5482D49BA">175</mods:end>
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<mods:location id="44534C73B579E1A9291CAE113646BE26">
<mods:url id="9F685FDE2AEF0987881A196CB14B7437">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
<mods:classification id="E695958E3E68EF4A758D6CC28EF6DF0A">journal article</mods:classification>
<mods:identifier id="26B52DC7CB722F4C1BE32825811493D9" type="DOI">10.11646/zootaxa.5492.2.1</mods:identifier>
<mods:identifier id="F9FC18316E34E302340CD0A88E4AB7FB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="178BB720E96D93A6BE0B7CBE6B3B4605" type="Zenodo-Dep">13219174</mods:identifier>
<mods:identifier id="5CF9AFC00FBB1DF8C5AB4B8C9080D94E" type="ZooBank">4F1EFAD5-6C53-4A2C-8694-AD45B222A9EF</mods:identifier>
</mods:mods>
<treatment id="0393572EFFA8223DFF15FE651715FC6F" LSID="urn:lsid:plazi:treatment:0393572EFFA8223DFF15FE651715FC6F" httpUri="http://treatment.plazi.org/id/0393572EFFA8223DFF15FE651715FC6F" lastPageId="22" lastPageNumber="173" pageId="20" pageNumber="171">
<subSubSection id="C320B5B3FFA8223FFF15FE65165BFE6E" box="[151,520,430,457]" pageId="20" pageNumber="171" type="nomenclature">
<paragraph id="8B85E638FFA8223FFF15FE65165BFE6E" blockId="20.[151,520,430,493]" box="[151,520,430,457]" pageId="20" pageNumber="171">
<heading id="D0CD5154FFA8223FFF15FE65165BFE6E" bold="true" box="[151,520,430,457]" fontSize="11" level="1" pageId="20" pageNumber="171" reason="1">
<taxonomicName id="4C3A9DBBFFA8223FFF15FE65165BFE6E" ID-CoL="3T6DX" authority="Verrill, 1900" authorityName="Verrill" authorityYear="1900" box="[151,520,430,457]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="171" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFA8223FFF15FE65165BFE6E" bold="true" box="[151,520,430,457]" pageId="20" pageNumber="171">
<emphasis id="B94E3A2AFFA8223FFF15FE651536FE6E" bold="true" box="[151,357,430,457]" italics="true" pageId="20" pageNumber="171">Leodice unifrons</emphasis>
<bibRefCitation id="EFAB9BC9FFA8223FFEEFFE65165BFE6E" author="Verrill, A. E." box="[365,520,430,456]" pageId="20" pageNumber="171" pagination="595 - 671" refId="ref14403" refString="Verrill, A. E. (1900) Additions to the Turbellaria, Nemertina, and Annelida of the Bermudas, with revisions of some New England genera and species. Transactions of the Connecticut Academy of Arts and Sciences, 10 (2), 595 - 671. https: // doi. org / 10.5962 / bhl. part. 7035" type="journal article" year="1900">Verrill, 1900</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFA8223FFF15FE1814B8FE4B" box="[151,235,467,493]" pageId="20" pageNumber="171" type="description">
<paragraph id="8B85E638FFA8223FFF15FE1814B8FE4B" blockId="20.[151,520,430,493]" box="[151,235,467,493]" pageId="20" pageNumber="171">
(
<figureCitation id="1301FABDFFA8223FFF1DFE1814B0FE4B" box="[159,227,467,493]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFA8223FFF15FDD81674FDCF" pageId="20" pageNumber="171" type="reference_group">
<paragraph id="8B85E638FFA8223FFF15FDD815A8FD8D" blockId="20.[151,788,531,617]" box="[151,507,531,555]" pageId="20" pageNumber="171">
<treatmentCitationGroup id="AB2AC116FFA8223FFF15FDD815A8FD8D" box="[151,507,531,555]" pageId="20" pageNumber="171">
<treatmentCitation id="0A9BC029FFA8223FFF15FDD815A8FD8D" author="Verrill, A. E." box="[151,507,531,555]" page="644" pageId="20" pageNumber="171" year="1900">
<taxonomicName id="4C3A9DBBFFA8223FFF15FDD815A8FD8D" ID-CoL="3T6DX" authority="Verrill, 1900: 644" authorityName="Verrill" authorityPageNumber="644" authorityYear="1900" box="[151,507,531,555]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="171" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFA8223FFF15FDD81512FD8D" box="[151,321,531,555]" italics="true" pageId="20" pageNumber="171">Leodice unifrons</emphasis>
<bibRefCitation id="EFAB9BC9FFA8223FFECAFDD815A8FD8D" author="Verrill, A. E." box="[328,507,531,555]" pageId="20" pageNumber="171" pagination="595 - 671" refId="ref14403" refString="Verrill, A. E. (1900) Additions to the Turbellaria, Nemertina, and Annelida of the Bermudas, with revisions of some New England genera and species. Transactions of the Connecticut Academy of Arts and Sciences, 10 (2), 595 - 671. https: // doi. org / 10.5962 / bhl. part. 7035" type="journal article" year="1900">Verrill, 1900: 644</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFA8223FFF15FDF91747FDEC" blockId="20.[151,788,531,617]" box="[151,788,562,586]" pageId="20" pageNumber="171">
<treatmentCitationGroup id="AB2AC116FFA8223FFF15FDF91747FDEC" box="[151,788,562,586]" pageId="20" pageNumber="171">
<emphasis id="B94E3A2AFFA8223FFF15FDF9153AFDEC" box="[151,361,562,586]" italics="true" pageId="20" pageNumber="171">
<taxonomicName id="4C3A9DBBFFA8223FFF15FDF91576FDEF" ID-CoL="3CFDB" authorityName="sensu Hartman" authorityYear="1948" box="[151,293,562,585]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="171" phylum="Annelida" rank="species" species="vittata">Eunice vittata</taxonomicName>
sensu
</emphasis>
<treatmentCitation id="0A9BC029FFA8223FFEF2FDF91670FDEC" author="Hartman, O." box="[368,547,562,586]" page="9" pageId="20" pageNumber="171" year="1942">
<bibRefCitation id="EFAB9BC9FFA8223FFEF2FDF91670FDEC" author="Hartman, O." box="[368,547,562,586]" pageId="20" pageNumber="171" pagination="1 - 98" refId="ref14026" refString="Hartman, O. (1942) A review of the types of polychaetous annelids at the Peabody Museum of Natural History, Yale University. Bulletin of the Bingham Oceanographic Collections, 8, 1 - 98." type="journal article" year="1942">Hartman, 1942: 9</bibRefCitation>
</treatmentCitation>
, not Delle Chiaje, 1929
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFA8223FFF15FD9A1674FDCF" blockId="20.[151,788,531,617]" box="[151,551,592,617]" pageId="20" pageNumber="171">
<treatmentCitationGroup id="AB2AC116FFA8223FFF15FD9A1674FDCF" box="[151,551,592,617]" pageId="20" pageNumber="171">
<taxonomicName id="4C3A9DBBFFA8223FFF15FD9A1564FDCF" ID-CoL="3CFD2" baseAuthorityName="Verrill" baseAuthorityYear="1900" box="[151,311,593,617]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="171" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFA8223FFF15FD9A1564FDCF" box="[151,311,593,617]" italics="true" pageId="20" pageNumber="171">Eunice unifrons</emphasis>
</taxonomicName>
<treatmentCitation id="0A9BC029FFA8223FFED0FD9B1670FDCF" author="Fauchald, K." box="[338,547,592,617]" page="330" pageId="20" pageNumber="171" year="1992">
<bibRefCitation id="EFAB9BC9FFA8223FFED0FD9B1670FDCF" author="Fauchald, K." box="[338,547,592,617]" pageId="20" pageNumber="171" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald, 1992: 330</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFA8223FFF15FD5911D8FC9D" pageId="20" pageNumber="171" type="materials_examined">
<paragraph id="8B85E638FFA8223FFF15FD5911D8FC9D" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">
<emphasis id="B94E3A2AFFA8223FFF15FD5915D6FD0A" bold="true" box="[151,389,658,684]" pageId="20" pageNumber="171">Material examined.</emphasis>
<materialsCitation id="3B52EC65FFA8223FFE12FD5811D4FC9D" collectionCode="POLY-UFPB" location="Paraiba State" pageId="20" pageNumber="171" specimenCount="9" stateProvince="Paraiba">
NON-TYPE material
<emphasis id="B94E3A2AFFA8223FFD0CFD5A176DFD0A" bold="true" box="[654,830,657,684]" pageId="20" pageNumber="171">
<collectingRegion id="49FE28DAFFA8223FFD0CFD5A176AFD0A" box="[654,825,657,684]" country="Brazil" name="Paraiba" pageId="20" pageNumber="171">Paraíba State</collectingRegion>
:
</emphasis>
<specimenCount id="9D3C2DB1FFA8223FFCCBFD5917ACFD0A" box="[841,1023,658,684]" count="9" pageId="20" pageNumber="171" type="generic">Nine specimens</specimenCount>
analyzed. One complete specimen, station 1 (
<collectionCode id="ED2B7EFDFFA8223FFE85FD7D15F1FD76" box="[263,418,694,720]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2008); one incomplete specimen, station 4 (
<collectionCode id="ED2B7EFDFFA8223FFC1BFD7D1067FD76" box="[921,1076,694,720]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2009); one complete specimen, station 35 (
<collectionCode id="ED2B7EFDFFA8223FFE8DFD1115F9FD53" box="[271,426,730,757]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2015); one incomplete specimen, station 37 (
<collectionCode id="ED2B7EFDFFA8223FFC19FD111066FD52" box="[923,1077,730,756]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
466); one incomplete specimen, station 39 (
<collectionCode id="ED2B7EFDFFA8223FFE92FD3515F9FCBE" box="[272,426,766,792]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2014); one incomplete specimen, station 60 (
<collectionCode id="ED2B7EFDFFA8223FFC22FD351069FCBE" box="[928,1082,766,792]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2010); one complete specimen, station 62 (
<collectionCode id="ED2B7EFDFFA8223FFE91FCEA15FDFC9D" box="[275,430,801,827]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2011); two incomplete specimens, station 66 (
<collectionCode id="ED2B7EFDFFA8223FFC3AFCEA1002FC9D" box="[952,1105,801,827]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2012;
<collectionCode id="ED2B7EFDFFA8223FFB23FCEA1168FC9D" box="[1185,1339,801,827]" pageId="20" pageNumber="171">POLY-UFPB</collectionCode>
2013)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFA8223DFF45FC8E15D6FD9B" lastPageId="22" lastPageNumber="173" pageId="20" pageNumber="171" type="description">
<paragraph id="8B85E638FFA8223FFF45FC8E11CAFC25" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">
<emphasis id="B94E3A2AFFA8223FFF45FC8E150AFCF9" bold="true" box="[199,345,837,863]" pageId="20" pageNumber="171">Description.</emphasis>
Six incomplete specimens with 4164 chaetigers and three complete specimens with 8098 chaetigers. Total length of complete specimens
<quantity id="4CC24BDDFFA8223FFD29FCA21774FC25" box="[683,807,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.4" metricValueMin="1.2" pageId="20" pageNumber="171" unit="mm" value="18.0" valueMax="24.0" valueMin="12.0">1224 mm</quantity>
; L10 1,52,5 mm; W10 without parapodia 2.0
<quantity id="4CC24BDDFFA8223FFABEFCA211C6FC25" box="[1340,1429,873,900]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="171" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
<paragraph id="8B85E638FFA8223FFF45FC471719FB63" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">
Prostomium bilobed, with a shallow median groove, distinct anteriorly and ventrally, but not dorsally; truncate, anteriorly steep with conspicuous ventral buccal lips, distinctly shorter and narrower than peristomium, (
<figureCitation id="1301FABDFFA8223FFABFFC7B1483FC48" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9A, B, C</figureCitation>
). Prostomial appendages evenly spaced in semicircle, palps anterior and slightly lateral to lateral antennae. Palpophores and ceratophores short and ringshaped, without articulations. Ceratostyles digitiform, tapering, with cylindrical articulations, palpostyles short and digitiform, with cylindrical articulations (
<figureCitation id="1301FABDFFA8223FFB0EFBD01163FB90" box="[1164,1328,1051,1078]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9A, B, C</figureCitation>
). Median antenna slightly longer than lateral antennae; lateral antennae about twice as long as palps; palps reaching posterior end of first peristomial ring with 47 articulations. Lateral antennae reaching third chaetiger to fifth chaetiger with 710 articulations. Median antenna reaching fourth chaetiger to eighth chaetiger with 813 articulations. Pair of eyes present, posterior to palps and lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFA8223FFF45FB051140FAF5" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">
Peristomium cylindrical, laterally longer than medially; dorsal anterior margin covering posterior margin of median antenna ceratophore; with slightly conspicuous ventro-lateral lips. First peristomial ring about ⅔ of total length, separation between rings conspicuously visible on all sides (
<figureCitation id="1301FABDFFA8223FFC1DFADD17A8FA96" box="[927,1019,1302,1328]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9A</figureCitation>
). Peristomial cirri tapering, reaching anterior part of first peristomial ring to almost middle of prostomium, with 34 articulations (
<figureCitation id="1301FABDFFA8223FFB2CFAF21155FAF5" box="[1198,1286,1337,1363]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFA8223FFF45FA9616DBF98C" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">
Maxillary formula: 1+1; (78)+(910); (910)+0; (810)+10, 1+1. MxI with not extended falcal arch, outer edge arched, rectangular, basal inner edge with curvature where most posterior region of MxII fits; MxIII almost same length as MxI, located partially ventral to MxII, anterior end part of distal arch with left MxIV. Left MxIV with decreasingly sized teeth on right portion. MxV wide, with a small tooth. MxVI absent. MxIV left attachment lamellae semi-circular in shape, tapering on right innermost portion; right attachment lamella semi-circular in shape, tapering on left innermost portion (
<figureCitation id="1301FABDFFA8223FFDA2F9DB1628F98C" box="[544,635,1552,1578]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="20" pageNumber="171">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFA8223FFF45F9F8175CF91F" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">Branchiae pectinate present from chaetiger 3 to 3038, in less than 55% of total number of chaetigers. First branchia with one branchial filament shorter than notopodial cirri, maximum number of branchial filaments 24, between chaetigers 20 and 32. Branchiae longer than notopodial cirri. Branchial filaments tapering, longer than notopodial cirri and longer than central axis of branchia.</paragraph>
<paragraph id="8B85E638FFA8223FFF45F909151CF8EE" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">Anterior chaetal lobe conical with short projection resembling a nipple near middle of lobe and posterior to aciculae; median and posterior chaetal lobe conical, with aciculae emerging from dorsal half. All prechaetal lobes low transverse folds. Anterior postchaetal lobes rounded, longer than chaetal lobe; median and posterior low transverse folds.</paragraph>
<paragraph id="8B85E638FFA8223FFF45F89A1757F85D" blockId="20.[151,1437,657,2043]" pageId="20" pageNumber="171">Ventral cirri longer than chaetal lobes in all parapodia, ventral cirri robust and digitiform until chaetigers 3 6, cirri basally inflated from chaetigers 47, inflated base oval with elongated digitiform tips decreasing in size around chaetiger 3033, after which ventral cirri taper to digitiform tips, longer than in anterior parapodia. Anterior notopodial cirri tapering to digitiform tips, with four articulations becoming shorter in median and posterior parapodia; median and posterior with 23 articulations.</paragraph>
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<paragraph id="8B85E638FFA9223EFF1AF88C11D1F86D" blockId="21.[151,1436,1863,1995]" pageId="21" pageNumber="172">
<emphasis id="B94E3A2AFFA9223EFF1AF88C1548F8F9" bold="true" box="[152,283,1863,1887]" pageId="21" pageNumber="172">FIGURE 9.</emphasis>
<taxonomicName id="4C3A9DBBFFA9223EFEA7F88C1580F8F9" authorityName="Verrill" authorityYear="1900" box="[293,467,1863,1887]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="172" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFA9223EFEA7F88C1580F8F9" box="[293,467,1863,1887]" italics="true" pageId="21" pageNumber="172">Leodice unifrons</emphasis>
</taxonomicName>
AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008.
<emphasis id="B94E3A2AFFA9223EFB6FF88C10ACF8F9" bold="true" box="[1261,1279,1863,1887]" pageId="21" pageNumber="172">A</emphasis>
—anterior end, lateral view;
<emphasis id="B94E3A2AFFA9223EFEA1F8A01567F825" bold="true" box="[291,308,1899,1923]" pageId="21" pageNumber="172">B</emphasis>
—anterior and, dorsal view;
<emphasis id="B94E3A2AFFA9223EFDE7F8A01624F825" bold="true" box="[613,631,1899,1923]" pageId="21" pageNumber="172">C</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFA9223EFC30F8A01797F825" bold="true" box="[946,964,1899,1923]" pageId="21" pageNumber="172">D</emphasis>
—maxillae, dorsal view;
<emphasis id="B94E3A2AFFA9223EFB4DF8A010AAF825" bold="true" box="[1231,1273,1899,1923]" pageId="21" pageNumber="172">E—</emphasis>
parapodium 53, pectinate chaetae;
<emphasis id="B94E3A2AFFA9223EFEDEF8441538F801" bold="true" box="[348,363,1935,1959]" pageId="21" pageNumber="172">F</emphasis>
—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook;
<emphasis id="B94E3A2AFFA9223EFAEDF84411D0F801" bold="true" box="[1391,1411,1935,1959]" pageId="21" pageNumber="172">G</emphasis>
— parapodium 40, aciculae;
<emphasis id="B94E3A2AFFA9223EFE1FF87815E2F86D" bold="true" box="[413,433,1971,1995]" pageId="21" pageNumber="172">H</emphasis>
—parapodium 53, compound falcigers chaetae;
<emphasis id="B94E3A2AFFA9223EFC15F87817E9F86D" bold="true" box="[919,954,1971,1995]" pageId="21" pageNumber="172">I—</emphasis>
parapodium 28, compound falcigers chaetae.
</paragraph>
</caption>
<paragraph id="8B85E638FFAA223DFF45FF5C16A0FE53" blockId="22.[151,1437,151,969]" pageId="22" pageNumber="173">
Limbate chaetae marginally serrated. Pectinate chaetae flat with 79 fine teeth and one much longer marginal tooth (
<figureCitation id="1301FABDFFAA223DFF61FF70156EFF73" box="[227,317,187,213]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="22" pageNumber="173">Fig. 9E</figureCitation>
). Compound falcigers bidentate, marginally serrated; anterior with long, narrow article, proximal tooth triangular tapering laterally, of similar length to distal tooth, tapering laterally (
<figureCitation id="1301FABDFFAA223DFBD9FF1410F8FF5F" box="[1115,1195,223,249]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="22" pageNumber="173">Fig. 9I</figureCitation>
); median with article longer and wider than in anterior parapodia, proximal tooth triangular, tapering laterally, larger than distal tooth, tapering laterally (
<figureCitation id="1301FABDFFAA223DFEEBFEEC15B1FEE7" box="[361,482,295,321]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="22" pageNumber="173">Fig. 9F, H</figureCitation>
); posterior article longer and wider than in median parapodia, article with proximal tooth triangular tapering laterally, larger than distal tooth tapering laterally.Aciculae paired, yellow, anterior straight, tapering to blunt tips, one mucronate (
<figureCitation id="1301FABDFFAA223DFDD6FEA41683FE2F" box="[596,720,367,393]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="22" pageNumber="173">Fig. 9F, G</figureCitation>
); median gently curved tapering to blunt tips, one mucronate; posterior tapering to fine tips, one mucronate acicula; last one acicula larger and longer than other in same parapodia. Subacicular hooks yellow, tridentate, first present on chaetiger 2027. Subacicular hooks always single; leaf-shaped guards covering first and second basal teeth (
<figureCitation id="1301FABDFFAA223DFD12FE1016B5FE53" box="[656,742,475,501]" captionStart="FIGURE 9" captionStartId="21.[152,255,1863,1887]" captionTargetBox="[151,1436,181,1839]" captionTargetId="figure-19@21.[151,1436,181,1839]" captionTargetPageId="21" captionText="FIGURE 9. Leodice unifrons AC, POLY-UFPB2012. D, POLY-UFPB2015. EI, POLY-UFPB2008. A—anterior end, lateral view; B—anterior and, dorsal view; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 53, pectinate chaetae; F—parapodium 53, limbate chaetae, aciculae, compound falcigers chaetae and subacicular hook; G— parapodium 40, aciculae; H—parapodium 53, compound falcigers chaetae; I—parapodium 28, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219202" httpUri="https://zenodo.org/record/13219202/files/figure.png" pageId="22" pageNumber="173">Fig. 9F</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFAA223DFF45FE3415D6FD9B" blockId="22.[151,1437,151,969]" pageId="22" pageNumber="173">Pygidium with two pairs of anal cirri, without articulations. Dorsal pygidial cirri about four times longer than ventral pygidial cirri.</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAA223DFF45FD8D15B6FD0F" pageId="22" pageNumber="173" type="distribution">
<paragraph id="8B85E638FFAA223DFF45FD8D15B6FD0F" blockId="22.[151,1437,151,969]" pageId="22" pageNumber="173">
<emphasis id="B94E3A2AFFAA223DFF45FD8D1532FDC6" bold="true" box="[199,353,582,608]" pageId="22" pageNumber="173">Distribution.</emphasis>
<taxonomicName id="4C3A9DBBFFAA223DFEE5FD8C16E4FDC7" authority="Verrill, 1900" authorityName="Verrill" authorityYear="1900" box="[359,695,583,609]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="173" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFAA223DFEE5FD8C1670FDC7" box="[359,547,583,609]" italics="true" pageId="22" pageNumber="173">Leodice unifrons</emphasis>
<bibRefCitation id="EFAB9BC9FFAA223DFDA8FD8C16E4FDC7" author="Verrill, A. E." box="[554,695,583,609]" pageId="22" pageNumber="173" pagination="595 - 671" refId="ref14403" refString="Verrill, A. E. (1900) Additions to the Turbellaria, Nemertina, and Annelida of the Bermudas, with revisions of some New England genera and species. Transactions of the Connecticut Academy of Arts and Sciences, 10 (2), 595 - 671. https: // doi. org / 10.5962 / bhl. part. 7035" type="journal article" year="1900">Verrill, 1900</bibRefCitation>
</taxonomicName>
was originally described from
<collectingCountry id="F32DA6A8FFAA223DFB91FD8C1029FDC7" box="[1043,1146,583,609]" name="Bermuda" pageId="22" pageNumber="173">Bermuda</collectingCountry>
(
<bibRefCitation id="EFAB9BC9FFAA223DFB0AFD8C11C2FDC7" author="Read, G. &amp; Fauchald, K." box="[1160,1425,583,609]" pageId="22" pageNumber="173" refId="ref14264" refString="Read, G. &amp; Fauchald, K. (Eds.) (2023) World Polychaeta Database. Eunice pellucida Kinberg, 1865. Accessed through: World Register of Marine Species. Available from: https: // www. marinespecies. org / aphia. php? p = taxdetails &amp; id = 324708 (accessed 11 May 2023)" type="url" year="2023">Read &amp; Fauchald 2023</bibRefCitation>
). It has been reported in various locations in
<collectingCountry id="F32DA6A8FFAA223DFDF9FDA01691FD23" box="[635,706,619,645]" name="Brazil" pageId="22" pageNumber="173">Brazil</collectingCountry>
, including
<collectingCountry id="F32DA6A8FFAA223DFCBCFDA017F0FD23" box="[830,931,619,645]" name="El Salvador" pageId="22" pageNumber="173">Salvador</collectingCountry>
, Bahia and Rio de Janeiro States, at
<quantity id="4CC24BDDFFAA223DFAB9FDA014FEFD0F" metricMagnitude="2" metricUnit="m" metricValue="6.025" metricValueMax="11.26" metricValueMin="0.79" pageId="22" pageNumber="173" unit="m" value="602.5" valueMax="1126.0" valueMin="79.0">791126 m</quantity>
depth (
<bibRefCitation id="EFAB9BC9FFAA223DFE83FD44158AFD0F" author="Amaral, A. C. Z. &amp; Nallin, S. A. H. &amp; Steiner, T. M. &amp; Forroni, T. O. &amp; Gomes-Filho, D. &amp; Araujo, G. R. &amp; Freitas, R. &amp; Costa, C. A. O. &amp; Ruta, C. &amp; Gomes, K. R. E. &amp; Bonaldo, R. O." box="[257,473,655,681]" pageId="22" pageNumber="173" refId="ref13263" refString="Amaral, A. C. Z., Nallin, S. A. H., Steiner, T. M., Forroni, T. O. &amp; Gomes-Filho, D., Araujo, G. R., Freitas, R., Costa, C. A. O., Ruta, C., Gomes, K. R. E. &amp; Bonaldo, R. O. (2022) Catalogo das especies de Annelida &quot; Polychaeta &quot; do Brasil. Available from: http: // www. ib. unicamp. br / museu _ zoologia / files / lab _ museu _ zoologia / Catalogo _ Polychaeta _ Amaral _ et _ al _ 2022. pdf (accessed 11 May 2023)" type="url" year="2022">
Amaral
<emphasis id="B94E3A2AFFAA223DFEDFFD4415C5FD0F" box="[349,406,655,681]" italics="true" pageId="22" pageNumber="173">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAA223DFF45FD781715FC6F" pageId="22" pageNumber="173" type="discussion">
<paragraph id="8B85E638FFAA223DFF45FD78160BFC27" blockId="22.[151,1437,151,969]" pageId="22" pageNumber="173">
<emphasis id="B94E3A2AFFAA223DFF45FD781568FD6B" bold="true" box="[199,315,691,717]" pageId="22" pageNumber="173">Remarks.</emphasis>
The analyzed specimens share with
<taxonomicName id="4C3A9DBBFFAA223DFD53FD781719FD6B" authorityName="Verrill" authorityYear="1900" box="[721,842,691,717]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="173" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFAA223DFD53FD781719FD6B" box="[721,842,691,717]" italics="true" pageId="22" pageNumber="173">L. unifrons</emphasis>
</taxonomicName>
redescribed by
<bibRefCitation id="EFAB9BC9FFAA223DFC78FD7810E1FD6B" author="Fauchald, K." box="[1018,1202,691,717]" pageId="22" pageNumber="173" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald (1992)</bibRefCitation>
most characteristics, except for: 1. the presence of eyes located posterior to palps and lateral to lateral antennae, not observed in type and
<typeStatus id="5481589AFFAA223DFF4BFD301578FCB3" box="[201,299,763,789]" pageId="22" pageNumber="173" type="topotype">topotype</typeStatus>
specimens; 2. antennae and palps in semi-circular distribution, while they are horseshoeshaped in type and
<typeStatus id="5481589AFFAA223DFF7FFCD4150CFC9F" box="[253,351,799,825]" pageId="22" pageNumber="173" type="topotype">topotype</typeStatus>
specimens; 3. last branchiae is in chaetigers 3033, while it is in chaetigers
<quantity id="4CC24BDDFFAA223DFB2EFCD5115DFC9F" box="[1196,1294,798,825]" metricMagnitude="0" metricUnit="m" metricValue="1.2573" metricValueMax="1.3716" metricValueMin="1.143" pageId="22" pageNumber="173" unit="in" value="49.5" valueMax="54.0" valueMin="45.0">4554 in</quantity>
the type and
<typeStatus id="5481589AFFAA223DFF15FC8814AAFCFB" box="[151,249,835,861]" pageId="22" pageNumber="173" type="topotype">topotype</typeStatus>
specimens with more than 70 chaetigers; and 4. maximum number of branchial filaments is 24, while it is
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the type and
<typeStatus id="5481589AFFAA223DFEF1FCAC1586FC27" box="[371,469,871,897]" pageId="22" pageNumber="173" type="topotype">topotype</typeStatus>
specimens.
</paragraph>
<paragraph id="8B85E638FFAA223DFF45FC401715FC6F" blockId="22.[151,1437,151,969]" pageId="22" pageNumber="173">
We propose the placement of
<taxonomicName id="4C3A9DBBFFAA223DFD8CFC401730FC03" authority="(Verrill, 1900)" baseAuthorityName="Verrill" baseAuthorityYear="1900" box="[526,867,907,933]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="173" phylum="Annelida" rank="species" species="unifrons">
<emphasis id="B94E3A2AFFAA223DFD8CFC4016EDFC03" box="[526,702,907,933]" italics="true" pageId="22" pageNumber="173">Eunice unifrons</emphasis>
(
<bibRefCitation id="EFAB9BC9FFAA223DFD49FC401709FC03" author="Verrill, A. E." box="[715,858,907,933]" pageId="22" pageNumber="173" pagination="595 - 671" refId="ref14403" refString="Verrill, A. E. (1900) Additions to the Turbellaria, Nemertina, and Annelida of the Bermudas, with revisions of some New England genera and species. Transactions of the Connecticut Academy of Arts and Sciences, 10 (2), 595 - 671. https: // doi. org / 10.5962 / bhl. part. 7035" type="journal article" year="1900">Verrill, 1900</bibRefCitation>
)
</taxonomicName>
back to the genus
<taxonomicName id="4C3A9DBBFFAA223DFBACFC4010D4FC03" authorityName="Lamarck" authorityYear="1818" box="[1070,1159,907,933]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="22" pageNumber="173" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFAA223DFBACFC4010D4FC03" box="[1070,1159,907,933]" italics="true" pageId="22" pageNumber="173">Leodice</emphasis>
</taxonomicName>
, as it was first described, based on the presence of yellow tridentate subacicular hooks.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,317 @@
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<mods:title id="53734E839B36A357170E6E16CE15E7C0">Taxonomy of Leodice Lamarck, 1818 (Annelida, Eunicidae) from Northeast Brazil (Tropical Southwest Atlantic Ocean) with description of a new species</mods:title>
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<mods:namePart id="830FF50C686CC7C74AE96AA109106BD7">Souza, Thaís Kananda Da Silva</mods:namePart>
<mods:affiliation id="D0702A38CF064071D25688BA8F8B0CB5">Programa de Pós-graduação em Zoologia. Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista s / n, São Cristovão, Rio de Janeiro, RJ 20940 - 040, Brazil. Current address. &amp; Laboratório de Biodiversidade de Annelida (LaBiAnne), Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Av. Bartolomeu de Gusmão, 875, São Cristóvão, Rio de Janeiro, RJ 20941 - 160, Brazil.</mods:affiliation>
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<mods:namePart id="5C28AF605F2BAA6CA32793E171F3B1DE">Zapff, Luana</mods:namePart>
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<mods:namePart id="EE684C9FFD3FA1B436A350493A4577CB">Zanol, Joana</mods:namePart>
<mods:affiliation id="B77CD37B225D74249F14B7867C03E144">Laboratório de Biodiversidade de Annelida (LaBiAnne), Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Av. Bartolomeu de Gusmão, 875, São Cristóvão, Rio de Janeiro, RJ 20941 - 160, Brazil.</mods:affiliation>
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<mods:date id="0CE238878840D7E020C6468F9DF5305C">2024</mods:date>
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<mods:number id="CCCBED70E7A9A065ED6BEB53166CD1E1">5492</mods:number>
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<treatment id="0393572EFFAD223FFF15F8D410E2FEC3" ID-DOI="http://doi.org/10.5281/zenodo.13219190" ID-Zenodo-Dep="13219190" LSID="urn:lsid:plazi:treatment:0393572EFFAD223FFF15F8D410E2FEC3" httpUri="http://treatment.plazi.org/id/0393572EFFAD223FFF15F8D410E2FEC3" lastPageId="20" lastPageNumber="171" pageId="17" pageNumber="168">
<subSubSection id="C320B5B3FFAD223AFF15F8D415A4F89F" box="[151,503,1823,1850]" pageId="17" pageNumber="168" type="nomenclature">
<paragraph id="8B85E638FFAD223AFF15F8D415A4F89F" blockId="17.[151,503,1823,1885]" box="[151,503,1823,1850]" pageId="17" pageNumber="168">
<heading id="D0CD5154FFAD223AFF15F8D415A4F89F" bold="true" box="[151,503,1823,1850]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
<taxonomicName id="4C3A9DBBFFAD223AFF15F8D415A4F89F" ID-CoL="3T6DL" authority="(Grube, 1856)" baseAuthorityName="Grube" baseAuthorityYear="1856" box="[151,503,1823,1850]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="rubra">
<emphasis id="B94E3A2AFFAD223AFF15F8D415A4F89F" bold="true" box="[151,503,1823,1850]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF15F8D41512F89C" bold="true" box="[151,321,1823,1850]" italics="true" pageId="17" pageNumber="168">Leodice rubra</emphasis>
(
<bibRefCitation id="EFAB9BC9FFAD223AFED1F8D415BCF89F" author="Grube, A. E." box="[339,495,1823,1849]" pageId="17" pageNumber="168" pagination="44 - 62" refId="ref13957" refString="Grube, A. E. (1856) Annulata Orstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845 - 1848 suscepto legit cl. A. S. Orsted, adjectis speciebus nonnullis a cl. H. Kroyero in itinere ad Americam meridionalem collectis. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i KObenhavn, 1, 44 - 62." type="journal article" year="1856">Grube, 1856</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAD223AFF15F88814B8F8FB" box="[151,235,1859,1885]" pageId="17" pageNumber="168" type="description">
<paragraph id="8B85E638FFAD223AFF15F88814B8F8FB" blockId="17.[151,503,1823,1885]" box="[151,235,1859,1885]" pageId="17" pageNumber="168">
(
<figureCitation id="1301FABDFFAD223AFF1DF88814B1F8FB" box="[159,226,1859,1885]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="17" pageNumber="168">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAD223AFF15F84F164CF85E" pageId="17" pageNumber="168" type="reference_group">
<paragraph id="8B85E638FFAD223AFF15F84F1583F83D" blockId="17.[151,559,1923,2040]" box="[151,464,1923,1948]" pageId="17" pageNumber="168">
<treatmentCitationGroup id="AB2AC116FFAD223AFF15F84F1583F83D" box="[151,464,1923,1948]" pageId="17" pageNumber="168">
<treatmentCitation id="0A9BC029FFAD223AFF15F84F159FF83D" author="Grube, A. E." box="[151,460,1923,1948]" page="59" pageId="17" pageNumber="168" year="1856">
<taxonomicName id="4C3A9DBBFFAD223AFF15F84F159FF83D" ID-CoL="3CFBM" authority="Grube, 1856: 59" authorityName="Grube" authorityPageNumber="59" authorityYear="1856" box="[151,460,1923,1948]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="rubra">
<emphasis id="B94E3A2AFFAD223AFF15F84F154DF83A" box="[151,286,1924,1948]" italics="true" pageId="17" pageNumber="168">Eunice rubra</emphasis>
<bibRefCitation id="EFAB9BC9FFAD223AFEA6F84F159FF83D" author="Grube, A. E." box="[292,460,1923,1948]" pageId="17" pageNumber="168" pagination="44 - 62" refId="ref13957" refString="Grube, A. E. (1856) Annulata Orstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845 - 1848 suscepto legit cl. A. S. Orsted, adjectis speciebus nonnullis a cl. H. Kroyero in itinere ad Americam meridionalem collectis. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i KObenhavn, 1, 44 - 62." type="journal article" year="1856">Grube, 1856: 59</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFAD223AFF15F8681647F81C" blockId="17.[151,559,1923,2040]" box="[151,532,1954,1978]" pageId="17" pageNumber="168">
<treatmentCitationGroup id="AB2AC116FFAD223AFF15F8681647F81C" box="[151,532,1954,1978]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF15F8681576F81C" box="[151,293,1954,1978]" italics="true" pageId="17" pageNumber="168">
<taxonomicName id="4C3A9DBBFFAD223AFF15F8681572F81C" ID-CoL="3CFBM" authorityName="Grube" authorityYear="1856" box="[151,289,1954,1978]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="rubra">Eunice rubra</taxonomicName>
.
</emphasis>
<treatmentCitation id="0A9BC029FFAD223AFEBCF8691643F81C" author="Fauchald, K." box="[318,528,1954,1978]" page="290" pageId="17" pageNumber="168" year="1992">
<bibRefCitation id="EFAB9BC9FFAD223AFEBCF8691643F81C" author="Fauchald, K." box="[318,528,1954,1978]" pageId="17" pageNumber="168" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald, 1992: 290</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFAD223AFF15F809167CF87F" blockId="17.[151,559,1923,2040]" box="[151,559,1985,2009]" pageId="17" pageNumber="168">
<treatmentCitationGroup id="AB2AC116FFAD223AFF15F809167CF87F" box="[151,559,1985,2009]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF15F8091568F87F" box="[151,315,1985,2009]" italics="true" pageId="17" pageNumber="168">
<taxonomicName id="4C3A9DBBFFAD223AFF15F809154EF87F" ID-CoL="3CFBM" authorityName="Grube" authorityYear="1856" box="[151,285,1985,2009]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="rubra">Eunice rubra</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0A9BC029FFAD223AFEB8F80A1678F87F" author="Steiner, T. M. &amp; Nogueira, J. M. M. &amp; Amaral, A. C. Z." box="[314,555,1985,2009]" page="243" pageId="17" pageNumber="168" year="2002">
<bibRefCitation id="EFAB9BC9FFAD223AFEB8F80A1678F87F" author="Steiner, T. M. &amp; Nogueira, J. M. M. &amp; Amaral, A. C. Z." box="[314,555,1985,2009]" pageId="17" pageNumber="168" pagination="243 - 259" refId="ref14328" refString="Steiner, T. M., Nogueira, J. M. M. &amp; Amaral, A. C. Z. (2002) Eunice rubra Grube (Annelida, Polychaeta, Eunicidae), a redescription based on the holotype and Brazilian specimens. Revista Brasileira de Zoologia, 19 (1), 243 - 259. https: // doi. org / 10.1590 / S 0101 - 81752002000500019" type="journal article" year="2002">
Steiner
<emphasis id="B94E3A2AFFAD223AFE0BF80915E6F87F" box="[393,437,1985,2009]" italics="true" pageId="17" pageNumber="168">et al</emphasis>
. 2002: 243
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFAD223AFF15F82B164CF85E" blockId="17.[151,559,1923,2040]" box="[151,543,2016,2040]" pageId="17" pageNumber="168">
<treatmentCitationGroup id="AB2AC116FFAD223AFF15F82B164CF85E" box="[151,543,2016,2040]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF15F82B1516F85E" box="[151,325,2016,2040]" italics="true" pageId="17" pageNumber="168">
<taxonomicName id="4C3A9DBBFFAD223AFF15F82B1574F85E" ID-CoL="3T6DL" baseAuthorityName="Grube" baseAuthorityYear="1856" box="[151,295,2016,2040]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="rubra">Leodice rubra</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0A9BC029FFAD223AFEC6F82B1648F85E" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[324,539,2016,2040]" page="90" pageId="17" pageNumber="168" year="2014">
<bibRefCitation id="EFAB9BC9FFAD223AFEC6F82B1648F85E" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[324,539,2016,2040]" pageId="17" pageNumber="168" pagination="79 - 100" refId="ref14618" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="B94E3A2AFFAD223AFE04F82B15EAF85E" box="[390,441,2016,2040]" italics="true" pageId="17" pageNumber="168">et al.</emphasis>
2014: 90
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF45B6B0FFAE2239FF1AFA5315B8F9BA" ID-DOI="http://doi.org/10.5281/zenodo.13219198" ID-Zenodo-Dep="13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="18" pageNumber="169" startId="18.[152,255,1432,1456]" targetBox="[151,1436,181,1408]" targetPageId="18" targetType="figure">
<paragraph id="8B85E638FFAE2239FF1AFA5315B8F9BA" blockId="18.[151,1436,1432,1564]" pageId="18" pageNumber="169">
<emphasis id="B94E3A2AFFAE2239FF1AFA53154EFA16" bold="true" box="[152,285,1432,1456]" pageId="18" pageNumber="169">FIGURE 8.</emphasis>
<emphasis id="B94E3A2AFFAE2239FEAAFA531597FA16" box="[296,452,1432,1456]" italics="true" pageId="18" pageNumber="169">
<taxonomicName id="4C3A9DBBFFAE2239FEAAFA531593FA16" baseAuthorityName="Grube" baseAuthorityYear="1856" box="[296,448,1432,1456]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="169" phylum="Annelida" rank="species" species="rubra">Leodice rubra</taxonomicName>
.
</emphasis>
POLY-UFPB-2065.
<emphasis id="B94E3A2AFFAE2239FD1DFA5316E2FA16" bold="true" box="[671,689,1432,1456]" pageId="18" pageNumber="169">A</emphasis>
—anterior end, dorsal view;
<emphasis id="B94E3A2AFFAE2239FC65FA531042FA16" bold="true" box="[999,1041,1432,1456]" pageId="18" pageNumber="169">B—</emphasis>
anterior and lateral;
<emphasis id="B94E3A2AFFAE2239FB6EFA5310ADFA16" bold="true" box="[1260,1278,1432,1456]" pageId="18" pageNumber="169">C</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFAE2239FEA1FA771566FA72" bold="true" box="[291,309,1468,1492]" pageId="18" pageNumber="169">D</emphasis>
—maxillae, dorsal view;
<emphasis id="B94E3A2AFFAE2239FDB5FA771632FA72" bold="true" box="[567,609,1468,1492]" pageId="18" pageNumber="169">E—</emphasis>
parapodium 8, compound falcigers chaetae;
<emphasis id="B94E3A2AFFAE2239FBA6FA771060FA72" bold="true" box="[1060,1075,1468,1492]" pageId="18" pageNumber="169">F</emphasis>
—parapodium 52, pectinate chaeta;
<emphasis id="B94E3A2AFFAE2239FF15FA2B14F8FA5E" bold="true" box="[151,171,1504,1528]" pageId="18" pageNumber="169">G</emphasis>
—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri;
<emphasis id="B94E3A2AFFAE2239FB40FA2B1085FA5E" bold="true" box="[1218,1238,1504,1528]" pageId="18" pageNumber="169">H</emphasis>
—parapodium 116, compound falcigers and aciculae.
</paragraph>
</caption>
<subSubSection id="C320B5B3FFAE2239FF15F9A5113BF822" pageId="18" pageNumber="169" type="materials_examined">
<paragraph id="8B85E638FFAE2239FF15F9A5113BF822" blockId="18.[151,1437,1645,2033]" pageId="18" pageNumber="169">
<emphasis id="B94E3A2AFFAE2239FF15F9A515D3F92E" bold="true" box="[151,384,1646,1672]" pageId="18" pageNumber="169">Material examined.</emphasis>
<materialsCitation id="3B52EC65FFAE2239FE04F9A51130F822" collectionCode="POLY-UFPB" location="Paraiba State" pageId="18" pageNumber="169" specimenCount="21" stateProvince="Paraiba">
NON-TYPE material
<emphasis id="B94E3A2AFFAE2239FDF8F9A61776F92E" bold="true" box="[634,805,1645,1672]" pageId="18" pageNumber="169">
<collectingRegion id="49FE28DAFFAE2239FDF8F9A61773F92E" box="[634,800,1645,1672]" country="Brazil" name="Paraiba" pageId="18" pageNumber="169">Paraíba State</collectingRegion>
:
</emphasis>
<specimenCount id="9D3C2DB1FFAE2239FCA9F9A51791F92E" box="[811,962,1646,1672]" count="21" pageId="18" pageNumber="169" type="generic">21 specimens</specimenCount>
analyzed. One complete specimen, station 01 (
<collectionCode id="ED2B7EFDFFAE2239FF40F959150EF90B" box="[194,349,1682,1709]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2055); one complete specimen, station 10 (
<collectionCode id="ED2B7EFDFFAE2239FCCAF95917B1F90A" box="[840,994,1682,1708]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2062); one complete specimen, station 11 (
<collectionCode id="ED2B7EFDFFAE2239FF43F97D150FF976" box="[193,348,1718,1744]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2063); one complete specimen, station 20 (
<collectionCode id="ED2B7EFDFFAE2239FCCAF97D17B1F976" box="[840,994,1718,1744]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2064); one complete specimen, station 36 (
<collectionCode id="ED2B7EFDFFAE2239FF40F911150FF952" box="[194,348,1754,1780]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2060); one complete specimen, station 37 (
<collectionCode id="ED2B7EFDFFAE2239FCCAF91117B7F952" box="[840,996,1754,1780]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2061); one complete specimen, station 41 (
<collectionCode id="ED2B7EFDFFAE2239FF47F9351533F8BF" box="[197,352,1790,1817]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2059); two incomplete specimens, station 43 (
<collectionCode id="ED2B7EFDFFAE2239FCFFF9351044F8BF" box="[893,1047,1790,1817]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2056); one incomplete specimen, station 60 (
<collectionCode id="ED2B7EFDFFAE2239FE8CF8E915FAF89B" box="[270,425,1826,1853]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2057); one complete specimen, station 62 (
<collectionCode id="ED2B7EFDFFAE2239FC03F8E9104FF89B" box="[897,1052,1826,1853]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2058); seven complete specimens, station 67 (
<collectionCode id="ED2B7EFDFFAE2239FE96F88D15FDF8C7" box="[276,430,1862,1889]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2065;
<collectionCode id="ED2B7EFDFFAE2239FE7FF88D16C5F8C6" box="[509,662,1862,1888]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2066); one complete specimen, station 76 (
<collectionCode id="ED2B7EFDFFAE2239FB03F88D114FF8C6" box="[1153,1308,1862,1888]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2067); one complete specimen, station 82 (
<collectionCode id="ED2B7EFDFFAE2239FE7BF8A116C7F822" box="[505,660,1898,1924]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2068); one complete specimen, station 84 (
<collectionCode id="ED2B7EFDFFAE2239FBFEF8A11144F822" box="[1148,1303,1898,1924]" pageId="18" pageNumber="169">POLY-UFPB</collectionCode>
2069)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAE2238FF45F845149EF8EB" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169" type="description">
<paragraph id="8B85E638FFAE2239FF45F84515ADF856" blockId="18.[151,1437,1645,2033]" pageId="18" pageNumber="169">
<emphasis id="B94E3A2AFFAE2239FF45F845150AF80E" bold="true" box="[199,345,1934,1960]" pageId="18" pageNumber="169">Description.</emphasis>
Three incomplete specimens with 2357 chaetigers and 18 complete specimens with 84123 chaetigers; total length of complete specimens ranging from
<quantity id="4CC24BDDFFAE2239FCC4F8791788F86B" box="[838,987,1970,1997]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="7.0" metricValueMin="2.3" pageId="18" pageNumber="169" unit="mm" value="46.5" valueMax="70.0" valueMin="23.0">23 to 70 mm</quantity>
; L10 ranging from 3.5 to 8.0 mm, W10
<quantity id="4CC24BDDFFAE2239FF15F81D1573F857" box="[151,288,2006,2033]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="18" pageNumber="169" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 mm</quantity>
, without parapodia.
</paragraph>
<paragraph id="8B85E638FFAF2238FF45FF5C1678FE0B" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
Prostomium bilobed with deep median groove visible dorsally and ventrally,truncated anteriorly with conspicuous ventral buccal lips, distinctly shorter and narrower than peristomium (
<figureCitation id="1301FABDFFAF2238FC1CFF701065FF73" box="[926,1078,187,213]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8A, B, C</figureCitation>
). Prostomial appendages evenly spaced in semicircle; palps anterior to lateral antennae. Palpophores and ceratophores short, ring-shaped, without articulations. Ceratostyles and palpostyles tapering, with moniliform articulations. Median antennae longest and palps shortest; palps reaching middle of first peristomial ring to posterior of second peristomial ring, with 814 articulations; lateral antennae reaching middle of first peristomial ring to fourth chaetiger, with 1424 articulations; median antenna reaching from first to seventh chaetiger, with 1828 articulations. Pair of eyes present, posterior to palps and lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFAF2238FF45FE7C1114FDC7" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
Peristomium cylindrical, laterally longer than medially; dorsal anterior margin partially covering bases of median and lateral antennae; anterior half narrower than posterior; with conspicuous ventro-lateral lips. First ring about ¾ of total length; separation between rings visible dorsally and ventrally, but not laterally. Second peristomial ring longest dorsally; longer medially than laterally at ventral side and on dorsal side. Peristomial cirri with 35 articulations, basally inflated, tapering distally, reaching anterior edge of first peristomial ring (
<figureCitation id="1301FABDFFAF2238FB42FD8C1169FDC7" box="[1216,1338,583,609]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8A, B</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFAF2238FF45FDA010FAFCB3" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
Maxillary formula: 1+1; (57)+(68); (68)+0; (58)+(711); 1+1. MxI without extended falcal arch, basal inner edge with curvature, where most posterior region of MxII fits; MxII with teeth distributed in more than half of its length; MxIII partially ventral to MxII with anterior region part of distal ventral arch to MxIV; MxIV with teeth decreasing in size in right portion of plate; MxV wide with a small tooth (
<figureCitation id="1301FABDFFAF2238FC4AFD1C1072FD57" box="[968,1057,727,753]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8D</figureCitation>
). MxVI absent. MxIV left lamella semi-circular in shape, tapering to right portion, right fixation lamella semi-circular in shape.
</paragraph>
<paragraph id="8B85E638FFAF2238FF45FCD415F4FC4B" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">Branchiae pectinate, present from chaetigers 57 to 75120, present in more than 65% total number of chaetigers; first branchia with one branchial filament shorter than notopodial cirri, maximum number of branchial filaments 617, reached on chaetiger 1018; branchial filaments longer than notopodial cirri, central axis of branchiae longer than branchial filaments and notopodial cirri; branchial filaments reduced in length and in number in middle of branchial distribution, of 23 branchial filaments; and increase in both length and number in posterior region, to 45 branchial filaments.</paragraph>
<paragraph id="8B85E638FFAF2238FF45FC3C1100FBFF" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">Anterior, median and posterior parapodia with truncated chaetal lobe, aciculae emerging on dorsal half. Prechaetal lobes low transverse folds, shorter than chaetal lobe. Postchaetal lobes almost same length as chaetal lobe in anterior and median parapodia, postchaetal lobes, same length as chaetal lobe in posterior parapodia.</paragraph>
<paragraph id="8B85E638FFAF2238FF45FBA8179DFB4F" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
Notopodial cirri tapering, anterior with 45 articulations, median and posterior with 3. Ventral cirri longer than chaetal lobes in all chaetigers; robust and digitiform until 45 chaetiger, basally inflated from chaetiger 56, tapering to digitiform tips until 2641 chaetiger, posteriorly inflated base decrease in size. Median and posterior parapodia with tapering ventral cirri, longer than anterior ones. (
<figureCitation id="1301FABDFFAF2238FCE5FB041792FB4F" box="[871,961,1231,1257]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8G</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFAF2238FF45FB3817D9F947" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
Limbate chaetae marginally serrated, present in all parapodia. Pectinate chaetae flat, 1115 teeth, with one marginal tooth longer than others, number of teeth increasing in posterior chaetae (
<figureCitation id="1301FABDFFAF2238FBD1FADC10F8FA97" box="[1107,1195,1303,1329]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8F</figureCitation>
). Compound falciger chaetae bidentate in most chaetigers and tridentate in last chaetiger; anterior with article tapering from proximal to distal end, with narrow guard, proximal tooth directed laterally, triangular shorter than distal tooth, laterally or obliquely directed (
<figureCitation id="1301FABDFFAF2238FE17FA4815BDFA3B" box="[405,494,1411,1437]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8E</figureCitation>
); median with articles longer and wider than in anterior parapodia, proximal tooth triangular, laterally directed, larger than distal tooth, laterally directed and more robust than anterior ones; posterior article shorter and narrower than anterior and median, with proximal tooth triangular, laterally directed, larger than distal tooth, laterally directed and more robust than in median parapodia (
<figureCitation id="1301FABDFFAF2238FC71FA241003F9AF" box="[1011,1104,1519,1545]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8H</figureCitation>
); in last parapodia tridentate with small apical tooth directed obliquely upward, a proximal, laterally directed, triangular tooth larger than distal, laterally directed tooth (
<figureCitation id="1301FABDFFAF2238FE24F9FC1653F9F7" box="[422,512,1591,1617]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8H</figureCitation>
). Aciculae yellow, usually paired; anterior ones tapered, smoothly curved blunt or sometimes mucronate tips (
<figureCitation id="1301FABDFFAF2238FE4AF990164CF9D3" box="[456,543,1627,1653]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8E</figureCitation>
); median ones curved bifid tips (
<figureCitation id="1301FABDFFAF2238FC0BF99017B1F9D3" box="[905,994,1627,1653]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8G</figureCitation>
); posterior ones more curved than those in median parapodia, bifid tips (
<figureCitation id="1301FABDFFAF2238FD8EF9B4163AF93F" box="[524,617,1663,1689]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8H</figureCitation>
). Subacicular hooks, yellow, tridentate, present from chaetigers 2026. Hooks present in all middle and posterior chaetigers, usually one per parapodium, some parapodia with two, leaf-shaped guards covering first and second most basal teeth (
<figureCitation id="1301FABDFFAF2238FCA1F90C172EF947" box="[803,893,1735,1761]" captionStart="FIGURE 8" captionStartId="18.[152,255,1432,1456]" captionTargetBox="[151,1436,181,1408]" captionTargetId="figure-23@18.[151,1436,181,1408]" captionTargetPageId="18" captionText="FIGURE 8. Leodice rubra. POLY-UFPB-2065. A—anterior end, dorsal view; B—anterior and lateral; C—anterior end, ventral view; D—maxillae, dorsal view; E—parapodium 8, compound falcigers chaetae; F—parapodium 52, pectinate chaeta; G—parapodium 52, ventral cirri, subacicular hook, compound falcigers, aciculae and notopodial cirri; H—parapodium 116, compound falcigers and aciculae." figureDoi="http://doi.org/10.5281/zenodo.13219198" httpUri="https://zenodo.org/record/13219198/files/figure.png" pageId="19" pageNumber="170">Fig. 8G</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFAF2238FF45F920149EF8EB" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">Pygidium with two pairs of anal cirri. Dorsal pygidial cirri, tapering, regularly articulated with nine articulations. Ventral pygidial cirri not articulated. Dorsal pygidial cirri approximately five times longer than ventral pygidial cirri.</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAF2238FF45F89D1622F87B" pageId="19" pageNumber="170" type="distribution">
<paragraph id="8B85E638FFAF2238FF45F89D1622F87B" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="170">
<emphasis id="B94E3A2AFFAF2238FF45F89D1533F8D7" bold="true" box="[199,352,1878,1905]" pageId="19" pageNumber="170">Distribution.</emphasis>
<taxonomicName id="4C3A9DBBFFAF2238FEE4F89C16FBF8D7" authority="(Grube, 1856)" baseAuthorityName="Grube" baseAuthorityYear="1856" box="[358,680,1878,1905]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="170" phylum="Annelida" rank="species" species="rubra">
<emphasis id="B94E3A2AFFAF2238FEE4F89C1656F8D7" box="[358,517,1879,1905]" italics="true" pageId="19" pageNumber="170">Leodice rubra</emphasis>
(
<bibRefCitation id="EFAB9BC9FFAF2238FD90F89C16CCF8D7" author="Grube, A. E." box="[530,671,1878,1905]" pageId="19" pageNumber="170" pagination="44 - 62" refId="ref13957" refString="Grube, A. E. (1856) Annulata Orstediana. Enumeratio Annulatorum, quae in itinere per Indiam occidentalem et Americam centralem annis 1845 - 1848 suscepto legit cl. A. S. Orsted, adjectis speciebus nonnullis a cl. H. Kroyero in itinere ad Americam meridionalem collectis. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i KObenhavn, 1, 44 - 62." type="journal article" year="1856">Grube, 1856</bibRefCitation>
)
</taxonomicName>
was originally described from
<collectingCountry id="F32DA6A8FFAF2238FC7DF89C10B1F8D7" box="[1023,1250,1879,1905]" name="Saint Thomas Island" pageId="19" pageNumber="170">Saint Thomas Island</collectingCountry>
(Caribbean Sea) (
<bibRefCitation id="EFAB9BC9FFAF2238FF1DF8B0151AF833" author="Fauchald, K." box="[159,329,1915,1941]" pageId="19" pageNumber="170" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald 1992</bibRefCitation>
). It has been reported in diverse locations in South America, along the Brazilian coast in the states of Alagoas, Bahia, Espírito Santo, Paraná, Paraíba, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Sergipe and São Paulo (
<bibRefCitation id="EFAB9BC9FFAF2238FEC4F808164EF87B" author="Amaral, A. C. Z. &amp; Nallin, S. A. H. &amp; Steiner, T. M. &amp; Forroni, T. O. &amp; Gomes-Filho, D. &amp; Araujo, G. R. &amp; Freitas, R. &amp; Costa, C. A. O. &amp; Ruta, C. &amp; Gomes, K. R. E. &amp; Bonaldo, R. O." box="[326,541,1987,2013]" pageId="19" pageNumber="170" refId="ref13263" refString="Amaral, A. C. Z., Nallin, S. A. H., Steiner, T. M., Forroni, T. O. &amp; Gomes-Filho, D., Araujo, G. R., Freitas, R., Costa, C. A. O., Ruta, C., Gomes, K. R. E. &amp; Bonaldo, R. O. (2022) Catalogo das especies de Annelida &quot; Polychaeta &quot; do Brasil. Available from: http: // www. ib. unicamp. br / museu _ zoologia / files / lab _ museu _ zoologia / Catalogo _ Polychaeta _ Amaral _ et _ al _ 2022. pdf (accessed 11 May 2023)" type="url" year="2022">
Amaral
<emphasis id="B94E3A2AFFAF2238FE21F8081588F87B" box="[419,475,1987,2013]" italics="true" pageId="19" pageNumber="170">et al.</emphasis>
2022
</bibRefCitation>
,
<bibRefCitation id="EFAB9BC9FFAF2238FDAAF8081636F87B" author="Amaral, A. C. Z. &amp; Cunha, B. P. &amp; Checon, H. H. &amp; Godoy, A. S. &amp; Silva, C. F. &amp; Corte, G. N. &amp; Nogueira, J. M. M. &amp; Fukuda, M. V. &amp; Steiner, T. M. &amp; Kawauchi, G. Y. &amp; Turra, A. &amp; Denadai, M. R. &amp; Ferreira, L. &amp; Mendonca, J. B. &amp; Tavares, M. &amp; Leite, F. P. P. &amp; Costa, M. F. B. &amp; Siqueira, S. G. L. &amp; Vieira, L. M. &amp; Dias, G. M. &amp; Teixeira, J. A. &amp; Rocha, R. M. &amp; Gusmao, L. C. &amp; Borges, M. &amp; Alitto, R. &amp; Machado, F. M. &amp; Passos, F. D. &amp; Cunha, C. M. &amp; Simone, L. R. L. &amp; Araujo, A. P. G. &amp; Carbayo, F. &amp; Bahia, J. &amp; Bulnes, V. N. &amp; Castello-Branco, C. &amp; Hadju, E. &amp; Vilas-Boas, C. &amp; Garraffoni, A. R. S. &amp; Schockaert, E. &amp; Fonseca, G. &amp; Domenico, M. D. &amp; Curini-Galleti, M. &amp; Sorensen, M. V. &amp; Hochberg, R. &amp; Oliveira, A. J. F. C. &amp; Zampieri, B. D. B. &amp; Chinelatto, R. M. &amp; Migotto, A. E." box="[552,613,1987,2013]" pageId="19" pageNumber="170" refId="ref12897" refString="Amaral, A. C. Z., Cunha, B. P., Checon, H. H., Godoy, A. S., Silva, C. F., Corte, G. N., Nogueira, J. M. M., Fukuda, M. V., Steiner, T. M., Kawauchi, G. Y., Turra, A., Denadai, M. R., Ferreira, L., Mendonca, J. B., Tavares, M., Leite, F. P. P., Costa, M. F. B., Siqueira, S. G. L., Vieira, L. M., Dias, G. M., Teixeira, J. A., Rocha, R. M., Gusmao, L. C., Borges, M., Alitto, R., Machado, F. M., Passos, F. D., Cunha, C. M., Simone, L. R. L., Araujo, A. P. G., Carbayo, F., Bahia, J., Bulnes, V. N., Castello-Branco, C., Hadju, E., Vilas-Boas, C., Garraffoni, A. R. S., Schockaert, E., Fonseca, G., Domenico, M. D., Curini-Galleti, M., Sorensen, M. V., Hochberg, R., Oliveira, A. J. F. C., Zampieri, B. D. B., Chinelatto, R. M. &amp; Migotto, A. E. (2024) The high biodiversity of benthic organisms in a coastal ecosystem revealed by an integrative approach. Biota Neotropica, 24, e 20231583. https: // doi. org / 10.1590 / 1676 - 0611 - BN- 2023 - 1583" type="journal volume" year="2024">2024</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFA8223FFF45FF5C10E2FEC3" pageId="20" pageNumber="171" type="discussion">
<paragraph id="8B85E638FFA8223FFF45FF5C10E2FEC3" blockId="20.[151,1436,151,357]" pageId="20" pageNumber="171">
<emphasis id="B94E3A2AFFA8223FFF45FF5C1568FF17" bold="true" box="[199,315,151,177]" pageId="20" pageNumber="171">Remarks.</emphasis>
The specimens analyzed resemble redescriptions of
<taxonomicName id="4C3A9DBBFFA8223FFC17FF5C17A7FF17" baseAuthorityName="Grube" baseAuthorityYear="1856" box="[917,1012,151,177]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="171" phylum="Annelida" rank="species" species="rubra">
<emphasis id="B94E3A2AFFA8223FFC17FF5C17A7FF17" box="[917,1012,151,177]" italics="true" pageId="20" pageNumber="171">L. rubra</emphasis>
</taxonomicName>
based on
<typeStatus id="5481589AFFA8223FFBE8FF5C10C9FF17" box="[1130,1178,151,177]" pageId="20" pageNumber="171">type</typeStatus>
and non-type material (
<bibRefCitation id="EFAB9BC9FFA8223FFF1DFF70151FFF73" author="Fauchald, K." box="[159,332,187,213]" pageId="20" pageNumber="171" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald 1992</bibRefCitation>
;
<bibRefCitation id="EFAB9BC9FFA8223FFEDEFF701666FF73" author="Steiner, T. M. &amp; Nogueira, J. M. M. &amp; Amaral, A. C. Z." box="[348,565,187,213]" pageId="20" pageNumber="171" pagination="243 - 259" refId="ref14328" refString="Steiner, T. M., Nogueira, J. M. M. &amp; Amaral, A. C. Z. (2002) Eunice rubra Grube (Annelida, Polychaeta, Eunicidae), a redescription based on the holotype and Brazilian specimens. Revista Brasileira de Zoologia, 19 (1), 243 - 259. https: // doi. org / 10.1590 / S 0101 - 81752002000500019" type="journal article" year="2002">
Steiner
<emphasis id="B94E3A2AFFA8223FFE37FF7015A2FF73" box="[437,497,187,213]" italics="true" pageId="20" pageNumber="171">et al.</emphasis>
2002
</bibRefCitation>
) in most of their characteristics, except for maximum number of branchial filaments (
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specimens examined in this study and
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redescriptions), and presence of tridentate compound falciger chaetae in last chaetigers of our specimens, which have not been described previously. However, this latter difference may be due to a larger sampling of parapodia analyzed here, since tridentate falciger compound chaetae were observed only in posterior most chaetiger, which is missing in the
<typeStatus id="5481589AFFA8223FFB94FE801015FEC3" box="[1046,1094,331,357]" pageId="20" pageNumber="171">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,283 @@
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<mods:title id="2CCDE082DDD0881C6FB4D038CB17B1DA">Taxonomy of Leodice Lamarck, 1818 (Annelida, Eunicidae) from Northeast Brazil (Tropical Southwest Atlantic Ocean) with description of a new species</mods:title>
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<emphasis id="B94E3A2AFFB32224FF15FC651695FC6E" bold="true" box="[151,710,942,969]" pageId="15" pageNumber="166">
<taxonomicName id="4C3A9DBBFFB32224FF15FC65166BFC6E" authority="(Kinberg, 1865)" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" baseAuthorityName="Kinberg" baseAuthorityYear="1865" box="[151,568,942,969]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="pellucida" status="comb. nov.">
<emphasis id="B94E3A2AFFB32224FF15FC65153FFC6F" bold="true" box="[151,364,942,969]" italics="true" pageId="15" pageNumber="166">Leodice pellucida</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB32224FEF9FC651663FC6E" author="Kinberg, J. G. H." box="[379,560,942,968]" pageId="15" pageNumber="166" pagination="559 - 574" refId="ref14107" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Koniglich Vetenskaps - Akademiens forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A27D8751FFB32224FDC2FC651695FC6E" box="[576,710,942,968]" pageId="15" pageNumber="166" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C320B5B3FFB32224FF15FC1914B8FC4A" box="[151,235,978,1004]" pageId="15" pageNumber="166" type="description">
<paragraph id="8B85E638FFB32224FF15FC1914B8FC4A" blockId="15.[151,710,942,1004]" box="[151,235,978,1004]" pageId="15" pageNumber="166">
(
<figureCitation id="1301FABDFFB32224FF1DFC1914B0FC4A" box="[159,227,978,1004]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="15" pageNumber="166">Fig. 7</figureCitation>
)
</paragraph>
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<subSubSection id="C320B5B3FFB32224FF15FBDC1772FBEB" pageId="15" pageNumber="166" type="reference_group">
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<treatmentCitationGroup id="AB2AC116FFB32224FF15FBDC1757FB89" box="[151,772,1046,1071]" pageId="15" pageNumber="166">
<treatmentCitation id="0A9BC029FFB32224FF15FBDC165CFB88" author="Kinberg, J. G. H." box="[151,527,1046,1071]" page="562" pageId="15" pageNumber="166" year="1865">
<taxonomicName id="4C3A9DBBFFB32224FF15FBDC165CFB88" ID-CoL="8SZCJ" authority="Kinberg, 1865: 562" authorityName="Kinberg" authorityPageNumber="562" authorityYear="1865" box="[151,527,1046,1071]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="pellucida">
<emphasis id="B94E3A2AFFB32224FF15FBDC1511FB89" box="[151,322,1047,1071]" italics="true" pageId="15" pageNumber="166">Eunice pellucida</emphasis>
<bibRefCitation id="EFAB9BC9FFB32224FECAFBDC165CFB88" author="Kinberg, J. G. H." box="[328,527,1046,1071]" pageId="15" pageNumber="166" pagination="559 - 574" refId="ref14107" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Koniglich Vetenskaps - Akademiens forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg, 1865: 562</bibRefCitation>
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.—
<treatmentCitation id="0A9BC029FFB32224FDAEFBDD1753FB89" author="Fauchald, K." box="[556,768,1046,1071]" page="261" pageId="15" pageNumber="166" year="1992">
<bibRefCitation id="EFAB9BC9FFB32224FDAEFBDD1753FB89" author="Fauchald, K." box="[556,768,1046,1071]" pageId="15" pageNumber="166" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald, 1992: 261</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B85E638FFB32224FF15FBFD1772FBEB" blockId="15.[151,801,1046,1101]" box="[151,801,1077,1101]" pageId="15" pageNumber="166">
<treatmentCitationGroup id="AB2AC116FFB32224FF15FBFD1772FBEB" box="[151,801,1077,1101]" pageId="15" pageNumber="166">
<treatmentCitation id="0A9BC029FFB32224FF15FBFD1676FBEB" author="Hartman, O." box="[151,549,1077,1101]" page="77" pageId="15" pageNumber="166" year="1948">
<taxonomicName id="4C3A9DBBFFB32224FF15FBFD1676FBEB" ID-CoL="3CFDB" authority="sensu Hartman, 1948: 77" authorityName="sensu Hartman" authorityPageNumber="77" authorityYear="1948" box="[151,549,1077,1101]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="vittata">
<emphasis id="B94E3A2AFFB32224FF15FBFD1530FBEB" box="[151,355,1078,1101]" italics="true" pageId="15" pageNumber="166">Eunice vittata sensu</emphasis>
<bibRefCitation id="EFAB9BC9FFB32224FEE8FBFD1676FBEB" author="Hartman, O." box="[362,549,1077,1101]" pageId="15" pageNumber="166" pagination="1 - 137" refId="ref14065" refString="Hartman, O. (1948) The marine annelids erected by Kinberg with notes on some other types in the Swedish State Museum. Arkiv fur Zoologi, 42 A (1), 1 - 137, 18 pls." type="journal article" year="1948">Hartman, 1948:77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Not Delle Chiaje, 1828.
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</paragraph>
</subSubSection>
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<emphasis id="B94E3A2AFFB32224FF15FBB015D0FB33" bold="true" box="[151,387,1147,1173]" pageId="15" pageNumber="166">Material examined.</emphasis>
<materialsCitation id="3B52EC65FFB32224FE0FFBB7115DFAC8" collectionCode="POLY-UFPB" location="Paraiba State" pageId="15" pageNumber="166" specimenCount="23" stateProvince="Paraiba">
NON-TYPE material
<emphasis id="B94E3A2AFFB32224FD05FBB01766FB33" bold="true" box="[647,821,1147,1174]" pageId="15" pageNumber="166">
<collectingRegion id="49FE28DAFFB32224FD05FBB01762FB33" box="[647,817,1147,1174]" country="Brazil" name="Paraiba" pageId="15" pageNumber="166">Paraíba State</collectingRegion>
:
</emphasis>
<specimenCount id="9D3C2DB1FFB32224FCBDFBB71789FB33" box="[831,986,1147,1174]" count="23" pageId="15" pageNumber="166" type="generic">23 specimens</specimenCount>
examined. Two incomplete specimens, station 04 (
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2029); one complete and one incomplete specimen, station 37 (
<collectionCode id="ED2B7EFDFFB32224FB02FB6B1148FB1C" box="[1152,1307,1184,1210]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2033); one complete and one incomplete specimen, station 46 (
<collectionCode id="ED2B7EFDFFB32224FD79FB0F17C4FB78" box="[763,919,1220,1246]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2031); one incomplete specimen, station 55 (
<collectionCode id="ED2B7EFDFFB32224FF1DFB23156AFAA4" box="[159,313,1256,1282]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2030); one incomplete specimen, station 62 (
<collectionCode id="ED2B7EFDFFB32224FCB1FB23179EFAA4" box="[819,973,1256,1282]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2032); one complete and two incomplete specimens, station 86 (
<collectionCode id="ED2B7EFDFFB32224FE21FAC7166FFA80" box="[419,572,1292,1318]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2034;
<collectionCode id="ED2B7EFDFFB32224FD12FAC01779FA80" box="[656,810,1291,1318]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2035); two incomplete specimens, station 87 (
<collectionCode id="ED2B7EFDFFB32224FACFFAC7148CFAEF" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2036;
<collectionCode id="ED2B7EFDFFB32224FEB7FAFB159DFAEC" box="[309,462,1328,1354]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2037); one incomplete specimen, station 88 (
<collectionCode id="ED2B7EFDFFB32224FC5EFAFB1024FAEC" box="[988,1143,1328,1354]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2038); one complete and eight incomplete specimens, station 93 (
<collectionCode id="ED2B7EFDFFB32224FDDAFA9F16A2FAC8" box="[600,753,1364,1390]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2039;
<collectionCode id="ED2B7EFDFFB32224FCC0FA9F1789FAC8" box="[834,986,1364,1390]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2040;
<collectionCode id="ED2B7EFDFFB32224FBABFA9F1090FAC8" box="[1065,1219,1364,1390]" pageId="15" pageNumber="166">POLY-UFPB</collectionCode>
2041)
</materialsCitation>
.
</paragraph>
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<emphasis id="B94E3A2AFFB32224FF45FABC1502FA37" bold="true" box="[199,337,1399,1425]" pageId="15" pageNumber="166">Description</emphasis>
. 19 incomplete individuals with 1983 chaetigers and four complete individuals with 87109 chaetigers, total length of complete individuals ranging from
<quantity id="4CC24BDDFFB32224FCBFFA5717EBFA10" box="[829,952,1436,1462]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="7.7" metricValueMin="2.0" pageId="15" pageNumber="166" unit="mm" value="48.5" valueMax="77.0" valueMin="20.0">2077 mm</quantity>
; L10 ranging from
<quantity id="4CC24BDDFFB32224FB0FFA501153FA10" box="[1165,1280,1435,1462]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="15" pageNumber="166" unit="mm" value="5.75" valueMax="7.5" valueMin="4.0">47.5 mm</quantity>
, W10 without parapodia ranging from
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.
</paragraph>
<paragraph id="8B85E638FFB32224FF45FA281678F95C" blockId="15.[151,1437,1147,2038]" pageId="15" pageNumber="166">
Prostomium bilobed, truncate anteriorly, dorsally inflated, shorter and narrower than peristomium, shallow median groove visible dorsally and ventrally; ventral buccal lips conspicuous (
<figureCitation id="1301FABDFFB32224FB92F9CC10E3F984" box="[1040,1200,1543,1570]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="15" pageNumber="166">Fig. 7A, C, D</figureCitation>
). Palps and antennae evenly spaced, arranged in a horseshoe shape. Palpophores and ceratophores short and ring-shaped, without articulations. Ceratostyles and palpostyles tapering to slender tips, with moniliform regular articulations, drop-shaped at distal end (
<figureCitation id="1301FABDFFB32224FE0EF9B81663F928" box="[396,560,1651,1678]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="15" pageNumber="166">Fig. 7A, C, D</figureCitation>
). Median antennae longest and palps shortest; palps with 611 articulations, reaching posterior edge of first peristomial ring to second peristomial ring; lateral antennae with 1022 articulations, reaching chaetiger 25; median antenna with 1429 articulations, reaching chaetiger 58. Eyes present, posterior to palps and lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFB32224FF45F8C815B1F808" blockId="15.[151,1437,1147,2038]" pageId="15" pageNumber="166">
Peristomium cylindrical, laterally longer than medially; dorsal anterior margin covering base of median antenna; anterior region tapering, following prostomium width closely with conspicuous ventrolateral lips. First peristomial ring about ¾ of peristomium total length, separation between rings visible dorsally and ventrally, not laterally. Peristomial cirri slender, tapering, with 36 cylindrical articulations, almost reaching anterior edge of first peristomial ring (
<figureCitation id="1301FABDFFB32224FEDBF8581586F808" box="[345,469,1939,1966]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="15" pageNumber="166">Fig. 7A, D</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB3223AFF45F87C17F4FF5E" blockId="15.[151,1437,1147,2038]" lastBlockId="17.[151,1437,151,1749]" lastPageId="17" lastPageNumber="168" pageId="15" pageNumber="166">
Maxillary formula: 1+1; (57)+(68); (78)+0; (48)+(811); 1+1. MxI with not extended falcal arch, outer basal edge straight, rectangular, basal inner edge with curve, where posterior region of MxII fits. MxIII partially ventral to MxII, anterior end part of distal arch with left MxIV. Left MxIV with teeth decreasing in size in right portion of plate. MxV wide, with small prominent tooth. MxVI absent. MxIV left fixation lamellae semicircular in shape, tapering to right portion, right semi-circular in shape (
<figureCitation id="1301FABDFFAD223AFCC0FF1517C9FF5E" box="[834,922,222,248]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7B</figureCitation>
).
</paragraph>
<caption id="DF45B6B0FFAC223BFF1AF8BE1105F85F" ID-DOI="http://doi.org/10.5281/zenodo.13219196" ID-Zenodo-Dep="13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="16" pageNumber="167" startId="16.[152,255,1909,1933]" targetBox="[196,1392,181,1885]" targetPageId="16" targetType="figure">
<paragraph id="8B85E638FFAC223BFF1AF8BE1105F85F" blockId="16.[151,1436,1909,2041]" pageId="16" pageNumber="167">
<emphasis id="B94E3A2AFFAC223BFF1AF8BE154EF82B" bold="true" box="[152,285,1909,1933]" pageId="16" pageNumber="167">FIGURE 7.</emphasis>
<taxonomicName id="4C3A9DBBFFAC223BFEAAF8BE15B2F82B" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" baseAuthorityName="Kinberg" baseAuthorityYear="1865" box="[296,481,1909,1933]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="167" phylum="Annelida" rank="species" species="pellucida" status="comb. nov.">
<emphasis id="B94E3A2AFFAC223BFEAAF8BE15B2F82B" box="[296,481,1909,1933]" italics="true" pageId="16" pageNumber="167">Leodice pellucida</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFAC223BFE6EF8BE1636F82B" bold="true" box="[492,613,1909,1933]" pageId="16" pageNumber="167">
<taxonomicNameLabel id="A27D8751FFAC223BFE6EF8BE1636F82B" box="[492,613,1909,1933]" pageId="16" pageNumber="167" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
POLY-UFPB-2023.
<emphasis id="B94E3A2AFFAC223BFCC3F8BE1700F82B" bold="true" box="[833,851,1909,1933]" pageId="16" pageNumber="167">A</emphasis>
—anterior end, dorsal view;
<emphasis id="B94E3A2AFFAC223BFB0AF8BE10CAF82B" bold="true" box="[1160,1177,1909,1933]" pageId="16" pageNumber="167">B</emphasis>
—maxillae, dorsal view;
<emphasis id="B94E3A2AFFAC223BFF15F85214FAF817" bold="true" box="[151,169,1945,1969]" pageId="16" pageNumber="167">C</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFAC223BFE59F85215BEF817" bold="true" box="[475,493,1945,1969]" pageId="16" pageNumber="167">D</emphasis>
—anterior end, lateral view;
<emphasis id="B94E3A2AFFAC223BFC9AF8521711F817" bold="true" box="[792,834,1945,1969]" pageId="16" pageNumber="167">E—</emphasis>
parapodium 105, acicula, pectinate chaetae and compound falcigers;
<emphasis id="B94E3A2AFFAC223BFF7FF8761576F873" bold="true" box="[253,293,1981,2005]" pageId="16" pageNumber="167">F—</emphasis>
parapodium 6, compound falcigers;
<emphasis id="B94E3A2AFFAC223BFD15F87616F8F873" bold="true" box="[663,683,1981,2005]" pageId="16" pageNumber="167">G</emphasis>
—parapodium 80, aciculae, compound falcigers chaetae;
<emphasis id="B94E3A2AFFAC223BFB74F8761170F873" bold="true" box="[1270,1315,1981,2005]" pageId="16" pageNumber="167">H—</emphasis>
parapodium 105, acicula, compound falcigers and subacicular hook;
<emphasis id="B94E3A2AFFAC223BFD50F82A16A6F85F" bold="true" box="[722,757,2017,2041]" pageId="16" pageNumber="167">I—</emphasis>
parapodium 105, compound falcigers and pectinate chaetae.
</paragraph>
</caption>
<paragraph id="8B85E638FFAD223AFF45FEC91050FE0D" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">Branchiae pectinate, present from chaetiger 5 to 2633, in less than 55% of total number of chaetigers. First branchia with one branchial filament, shorter than notopodial cirri, maximum number of branchial filaments 712, reached at chaetigers 1222. Most branchiae longer than notopodial cirri, branchial filaments longer than notopodial cirri and central axis of branchiae; branchiae decreasing in length and number of branchial filaments along mid region of distribution. Branchial central axis slender, tapering in all branchiae.</paragraph>
<paragraph id="8B85E638FFAD223AFF45FE7F1534FD24" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">All chaetal lobes rounded, with aciculae emerging from dorsal half; all pre- and postchaetal lobes as transverse folds, shorter than chaetal lobes. Notopodial cirri basally inflated, tapering to pointed tips, anterior with 46 articulations, median with 34, and posterior with 23. Ventral cirri longer than chaetal lobes in all chaetigers; slender and digitiform until chaetiger 45 after that, cirri basally inflated and tapering to digitiform tips from chaetiger 56. Inflated base oval, decreasing in size around chaetiger 1938, after which digitiform tips are longer than anterior ones.</paragraph>
<paragraph id="8B85E638FFAD223AFF45FD4016A7FBF4" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">
Limbate chaetae, marginally serrated. Pectinate chaetae flat or slightly curved, with 1012 teeth, marginal teeth unequal in length, longer and thicker than other teeth. Median and posterior pectinate chaetae with blade wider and more abruptly tapering, teeth longer and thinner than anterior pectinate chaetae (
<figureCitation id="1301FABDFFAD223AFB19FD19115AFD4A" box="[1179,1289,722,749]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7E, I</figureCitation>
). Compound falcigers chaetae bidentate or tridentate; anterior with article tapering from proximal to distal end, marginally serrated, bidentate, proximal tooth triangular directed laterally, shorter than laterally or obliquely directed distal tooth (
<figureCitation id="1301FABDFFAD223AFF60FCF6156AFCFE" box="[226,313,829,856]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7F</figureCitation>
); median with article longer, more robust and with wider guards than anterior, bidentate, proximal tooth triangular, more robust than in anterior chaetae, both teeth laterally; posterior, articles and guards wider than more anterior, tridentate, proximal tooth larger than distal tooth, directed laterally, more robust than in median chaetae, with minute apically directed oblique tooth, both largest teeth directed laterally (
<figureCitation id="1301FABDFFAD223AFB2EFC621172FC65" box="[1196,1313,937,963]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7E, I</figureCitation>
). Aciculae yellow, usually paired; anterior aciculae tapered, straight or slightly curved, with blunt or sometimes mucronate tips; median aciculae curved, with bifid tips; posterior aciculae more curved than median, one acicula with bifid tip and tapering to thin point (
<figureCitation id="1301FABDFFAD223AFE3DFBDF160AFB88" box="[447,601,1044,1071]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7E, G, H</figureCitation>
). Subacicular hooks yellow tridentate, present from chaetigers 2127, with guards covering proximal and median teeth (
<figureCitation id="1301FABDFFAD223AFD0FFBF316B4FBF4" box="[653,743,1080,1106]" captionStart="FIGURE 7" captionStartId="16.[152,255,1909,1933]" captionTargetBox="[196,1392,181,1885]" captionTargetId="figure-23@16.[196,1392,181,1885]" captionTargetPageId="16" captionText="FIGURE 7. Leodice pellucida comb. nov. POLY-UFPB-2023. A—anterior end, dorsal view; B—maxillae, dorsal view; C—anterior end, ventral view; D—anterior end, lateral view; E—parapodium 105, acicula, pectinate chaetae and compound falcigers; F—parapodium 6, compound falcigers; G—parapodium 80, aciculae, compound falcigers chaetae; H—parapodium 105, acicula, compound falcigers and subacicular hook; I—parapodium 105, compound falcigers and pectinate chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219196" httpUri="https://zenodo.org/record/13219196/files/figure.png" pageId="17" pageNumber="168">Fig. 7H</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFAD223AFF45FB90175AFB3C" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">Pygidium with two pairs of anal cirri. Dorsal cirri digitiform, regularly articulated, with approximately seven segments. Ventral cirri missing, but visible scar present.</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAD223AFF45FB68151BFA8E" pageId="17" pageNumber="168" type="distribution">
<paragraph id="8B85E638FFAD223AFF45FB68151BFA8E" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF45FB681532FB1B" bold="true" box="[199,353,1187,1213]" pageId="17" pageNumber="168">Distribution.</emphasis>
<taxonomicName id="4C3A9DBBFFAD223AFEE4FB6816B5FB18" authority="(Kinberg, 1865)" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" baseAuthorityName="Kinberg" baseAuthorityYear="1865" box="[358,742,1187,1214]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="pellucida" status="comb. nov.">
<emphasis id="B94E3A2AFFAD223AFEE4FB68167EFB1B" box="[358,557,1187,1213]" italics="true" pageId="17" pageNumber="168">Leodice pellucida</emphasis>
(
<bibRefCitation id="EFAB9BC9FFAD223AFDB9FB68168DFB18" author="Kinberg, J. G. H." box="[571,734,1187,1214]" pageId="17" pageNumber="168" pagination="559 - 574" refId="ref14107" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Koniglich Vetenskaps - Akademiens forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
)
</taxonomicName>
<emphasis id="B94E3A2AFFAD223AFD6EFB68173FFB1B" bold="true" box="[748,876,1187,1213]" pageId="17" pageNumber="168">
<taxonomicNameLabel id="A27D8751FFAD223AFD6EFB68173FFB1B" box="[748,876,1187,1213]" pageId="17" pageNumber="168" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
was described from St. Thomas Island
<emphasis id="B94E3A2AFFAD223AFAA2FB69117AFB1A" bold="true" box="[1312,1321,1186,1212]" pageId="17" pageNumber="168">(</emphasis>
Caribbean Sea) and it has also been reported associated with
<emphasis id="B94E3A2AFFAD223AFD66FB0C17BBFB47" box="[740,1000,1223,1249]" italics="true" pageId="17" pageNumber="168">Phragmatoma caudata</emphasis>
in
<collectingRegion id="49FE28DAFFAD223AFB90FB0C1026FB47" box="[1042,1141,1223,1249]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="17" pageNumber="168">Veracruz</collectingRegion>
(Gulf of
<collectingRegion id="49FE28DAFFAD223AFB65FB0C116DFB47" box="[1255,1342,1223,1249]" country="Mexico" name="Mexico" pageId="17" pageNumber="168">Mexico</collectingRegion>
) (León-González and Díaz-Castañeda 2006). This new record expands the distribution of species to
<collectingRegion id="49FE28DAFFAD223AFB4EFB211136FAA2" box="[1228,1381,1258,1284]" country="Brazil" name="Paraiba" pageId="17" pageNumber="168">Paraíba State</collectingRegion>
, SW Atlantic Ocean.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFAD223AFF45FAF9151CF973" pageId="17" pageNumber="168" type="discussion">
<paragraph id="8B85E638FFAD223AFF45FAF91624FA7D" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">
<emphasis id="B94E3A2AFFAD223AFF45FAF91568FAEA" bold="true" box="[199,315,1330,1356]" pageId="17" pageNumber="168">Remarks.</emphasis>
<bibRefCitation id="EFAB9BC9FFAD223AFEC7FAF915A8FAEA" author="Hartman, O." box="[325,507,1330,1356]" pageId="17" pageNumber="168" pagination="1 - 137" refId="ref14065" refString="Hartman, O. (1948) The marine annelids erected by Kinberg with notes on some other types in the Swedish State Museum. Arkiv fur Zoologi, 42 A (1), 1 - 137, 18 pls." type="journal article" year="1948">Hartman (1948)</bibRefCitation>
and
<bibRefCitation id="EFAB9BC9FFAD223AFDB5FAF916A2FAEA" author="Fauchald, K." box="[567,753,1330,1356]" pageId="17" pageNumber="168" pagination="1 - 422" refId="ref13911" refString="Fauchald, K. (1992) A review of the genus Eunice (Eunicidae: Polychaeta) based upon type material. Smithsonian Contributions to Zoology, 523, 1 - 422. https: // doi. org / 10.5479 / si. 00810282.523" type="journal article" year="1992">Fauchald (1992)</bibRefCitation>
redescribed the species based on the
<typeStatus id="5481589AFFAD223AFB25FAF91084FAEA" box="[1191,1239,1330,1356]" pageId="17" pageNumber="168">type</typeStatus>
specimens. Both redescriptions differ in the presence of articulations in notopodial cirri, which are described as present by Hartman and absent by Fauchald.We have observed in our material that this feature is easily lost in poorly preserved specimens. Thus, the contrast among both redescriptions may be due to the state of preservation of the
<typeStatus id="5481589AFFAD223AFB1CFA55109DFA1E" box="[1182,1230,1438,1464]" pageId="17" pageNumber="168">type</typeStatus>
specimens almost 50 years later, when Fauchald examined it.
</paragraph>
<paragraph id="8B85E638FFAD223AFF45FA2F1486F9CC" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">The specimens examined for this study contrast with the redescriptions in having eyes, and tridentate compound falciger chaetae. However, the latter may not have been observed in previous studies because, traditionally, one to few parapodia were examined per specimen, not allowing the full investigation of any variation occurring along the body.</paragraph>
<paragraph id="8B85E638FFAD223AFF45F9BF151CF973" blockId="17.[151,1437,151,1749]" pageId="17" pageNumber="168">
We propose the assignment of
<taxonomicName id="4C3A9DBBFFAD223AFDB0F9BF17F1F928" authority="Kinberg, 1865" authorityName="Kinberg" authorityYear="1865" box="[562,930,1651,1678]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="species" species="pellucida">
<emphasis id="B94E3A2AFFAD223AFDB0F9BF16A1F92B" box="[562,754,1651,1677]" italics="true" pageId="17" pageNumber="168">Eunice pellucida</emphasis>
<bibRefCitation id="EFAB9BC9FFAD223AFD7FF9B817F1F928" author="Kinberg, J. G. H." box="[765,930,1651,1678]" pageId="17" pageNumber="168" pagination="559 - 574" refId="ref14107" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Koniglich Vetenskaps - Akademiens forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
</taxonomicName>
to
<emphasis id="B94E3A2AFFAD223AFC4FF9B8107EF92B" box="[973,1069,1651,1677]" italics="true" pageId="17" pageNumber="168">
<taxonomicName id="4C3A9DBBFFAD223AFC4FF9B8107BF92B" authorityName="Lamarck" authorityYear="1818" box="[973,1064,1651,1677]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="168" phylum="Annelida" rank="genus">Leodice</taxonomicName>
,
</emphasis>
based on the presence of three antennae and a pair of palps regularly articulated, yellow tridentate subacicular hooks, and tridentate compound falciger chaetae.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="A296C8EE6F325FFDD87C0C588262E500" ID-DOI="10.11646/zootaxa.5492.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13219174" ID-ZooBank="4F1EFAD5-6C53-4A2C-8694-AD45B222A9EF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722840951683" checkinUser="plazi" docAuthor="Souza, Thaís Kananda Da Silva, Zapff, Luana, Christoffersen, Martin Lindsey &amp; Zanol, Joana" docDate="2024" docId="0393572EFFB52220FF15F9D91031F9E7" docLanguage="en" docName="zootaxa.5492.1.1.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leodice calcaricola Bergamo, Carrerette, Zanol &amp; Nogueira 2018" docType="treatment" docVersion="2" lastPageNumber="162" masterDocId="FFAA2F56FFBC222BFF82FFCB1453FFA6" masterDocTitle="Taxonomy of Leodice Lamarck, 1818 (Annelida, Eunicidae) from Northeast Brazil (Tropical Southwest Atlantic Ocean) with description of a new species" masterLastPageNumber="175" masterPageNumber="151" pageNumber="160" updateTime="1722841404778" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="04DB53B155432EEDFB2507082FE9ABE0">Taxonomy of Leodice Lamarck, 1818 (Annelida, Eunicidae) from Northeast Brazil (Tropical Southwest Atlantic Ocean) with description of a new species</mods:title>
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<mods:namePart id="EF89F0E385BC73A5D04432289B80BE5A">Souza, Thaís Kananda Da Silva</mods:namePart>
<mods:affiliation id="4348B09104F9577F496A0F0435F3A9E9">Programa de Pós-graduação em Zoologia. Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista s / n, São Cristovão, Rio de Janeiro, RJ 20940 - 040, Brazil. Current address. &amp; Laboratório de Biodiversidade de Annelida (LaBiAnne), Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Av. Bartolomeu de Gusmão, 875, São Cristóvão, Rio de Janeiro, RJ 20941 - 160, Brazil.</mods:affiliation>
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<mods:namePart id="7A786F19572853B4924DB2EE7E40B80E">Zapff, Luana</mods:namePart>
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<mods:namePart id="CDB5ADB0046F519BEAE403DF72AC35C0">Christoffersen, Martin Lindsey</mods:namePart>
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<mods:namePart id="3F9CEE7E7C21CA94DAFFD9550FFC67A5">Zanol, Joana</mods:namePart>
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<heading id="D0CD5154FFB52222FF15F9D9179FF98A" bold="true" box="[151,972,1554,1581]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
<taxonomicName id="4C3A9DBBFFB52222FF15F9D9179FF98A" ID-CoL="3T6CJ" authority="Bergamo, Carrerette, Zanol &amp; Nogueira, 2018" authorityName="Bergamo, Carrerette, Zanol &amp; Nogueira" authorityYear="2018" box="[151,972,1554,1581]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB52222FF15F9D9179FF98A" bold="true" box="[151,972,1554,1581]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF15F9D915D0F98B" bold="true" box="[151,387,1554,1581]" italics="true" pageId="9" pageNumber="160">Leodice calcaricola</emphasis>
<bibRefCitation id="EFAB9BC9FFB52222FE08F9D8179FF98A" author="Bergamo, G. &amp; Carrerette, O. &amp; Zanol, J. &amp; Nogueira, J. M. M." box="[394,972,1554,1581]" pageId="9" pageNumber="160" pagination="376 - 390" refId="ref13400" refString="Bergamo, G., Carrerette, O., Zanol, J. &amp; Nogueira, J. M. M. (2018) Species of Eunicidae (Annelida, Errantia, Eunicida) from rhodolith beds off the States of Paraiba and Pernambuco, northeastern Brazilian coast. Zootaxa, 4521 (3) 376 - 390. https: // doi. org / 10.11646 / zootaxa. 4521.3.4" type="journal article" year="2018">Bergamo, Carrerette, Zanol &amp; Nogueira, 2018</bibRefCitation>
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<paragraph id="8B85E638FFB52222FF15F9FC14B8F9F7" blockId="9.[151,972,1554,1617]" box="[151,235,1591,1617]" pageId="9" pageNumber="160">
(
<figureCitation id="1301FABDFFB52222FF1DF9FC14B0F9F7" box="[159,227,1591,1617]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
)
</paragraph>
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<paragraph id="8B85E638FFB52222FF15F9B01725F935" blockId="9.[151,886,1659,1684]" box="[151,886,1659,1684]" pageId="9" pageNumber="160">
<treatmentCitationGroup id="AB2AC116FFB52222FF15F9B01725F935" box="[151,886,1659,1684]" pageId="9" pageNumber="160">
<treatmentCitation id="0A9BC029FFB52222FF15F9B01721F935" author="Bergamo, G. &amp; Carrerette, O. &amp; Zanol, J. &amp; Nogueira, J. M. M." box="[151,882,1659,1684]" page="378" pageId="9" pageNumber="160" year="2018">
<taxonomicName id="4C3A9DBBFFB52222FF15F9B01721F935" ID-CoL="3T6CJ" authority="Bergamo, Carrerette, Zanol &amp; Nogueira, 2018: 378" authorityName="Bergamo, Carrerette, Zanol &amp; Nogueira" authorityPageNumber="378" authorityYear="2018" box="[151,882,1659,1684]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB52222FF15F9B01532F935" box="[151,353,1659,1683]" italics="true" pageId="9" pageNumber="160">Leodice calcaricola</emphasis>
<bibRefCitation id="EFAB9BC9FFB52222FEE5F9B71721F935" author="Bergamo, G. &amp; Carrerette, O. &amp; Zanol, J. &amp; Nogueira, J. M. M." box="[359,882,1659,1684]" pageId="9" pageNumber="160" pagination="376 - 390" refId="ref13400" refString="Bergamo, G., Carrerette, O., Zanol, J. &amp; Nogueira, J. M. M. (2018) Species of Eunicidae (Annelida, Errantia, Eunicida) from rhodolith beds off the States of Paraiba and Pernambuco, northeastern Brazilian coast. Zootaxa, 4521 (3) 376 - 390. https: // doi. org / 10.11646 / zootaxa. 4521.3.4" type="journal article" year="2018">Bergamo, Carrerette, Zanol &amp; Nogueira, 2018: 378</bibRefCitation>
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<paragraph id="8B85E638FFB52222FF15F90A1673F812" blockId="9.[151,1437,1728,1972]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF15F90A15D4F97D" bold="true" box="[151,391,1729,1755]" pageId="9" pageNumber="160">Material examined.</emphasis>
<materialsCitation id="3B52EC65FFB52222FE17F90A164FF812" collectionCode="POLY-UFPB" location="Paraiba State" pageId="9" pageNumber="160" specimenCount="11" stateProvince="Paraiba">
NON-TYPE material
<emphasis id="B94E3A2AFFB52222FD1BF90B1718F97D" bold="true" box="[665,843,1728,1755]" pageId="9" pageNumber="160">
<collectingRegion id="49FE28DAFFB52222FD1BF90B1715F97D" box="[665,838,1728,1755]" country="Brazil" name="Paraiba" pageId="9" pageNumber="160">Paraíba State</collectingRegion>
:
</emphasis>
<specimenCount id="9D3C2DB1FFB52222FCDBF90A17A4F97D" box="[857,1015,1729,1755]" count="11" pageId="9" pageNumber="160" type="generic">11 specimens</specimenCount>
analyzed. One complete and three incomplete specimens, station 01 (POLY-UFPB-2020); one complete and one incomplete specimen, station 04 (
<collectionCode id="ED2B7EFDFFB52222FF1DF8C2156AF885" box="[159,313,1801,1827]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2021;
<collectionCode id="ED2B7EFDFFB52222FE05F8C21673F885" box="[391,544,1801,1827]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2022); one incomplete specimen, station 27 (
<collectionCode id="ED2B7EFDFFB52222FB94F8C210E2F882" box="[1046,1201,1801,1828]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2025); one complete specimen, station 43 (
<collectionCode id="ED2B7EFDFFB52222FE0BF8E61677F8E1" box="[393,548,1837,1863]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2023); two complete specimens, station 66 (
<collectionCode id="ED2B7EFDFFB52222FB97F8E610FCF8E1" box="[1045,1199,1837,1863]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2024); one complete specimen, station 82 (
<collectionCode id="ED2B7EFDFFB52222FE17F89A167CF8CD" box="[405,559,1873,1899]" pageId="9" pageNumber="160">POLY-UFPB</collectionCode>
2026).
<emphasis id="B94E3A2AFFB52222FD08F89A174FF8CD" bold="true" box="[650,796,1873,1899]" pageId="9" pageNumber="160">Description.</emphasis>
Four incomplete specimens with 3874 chaetigers and seven complete specimens with 4057 chaetigers. Total length of complete specimens
<quantity id="4CC24BDDFFB52222FBF3F8BE10B3F836" box="[1137,1248,1909,1936]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="9" pageNumber="160" unit="mm" value="13.0" valueMax="20.0" valueMin="6.0">620 mm</quantity>
, L10
<quantity id="4CC24BDDFFB52222FAA0F8BE11CBF836" box="[1314,1432,1909,1936]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="9" pageNumber="160" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
, W10 without parapodia
<quantity id="4CC24BDDFFB52222FE25F852164FF812" box="[423,540,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="160" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
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.
</paragraph>
<caption id="DF45B6B0FFB62221FF1AF9EB1738F904" ID-DOI="http://doi.org/10.5281/zenodo.13219188" ID-Zenodo-Dep="13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="10" pageNumber="161" startId="10.[152,255,1568,1592]" targetBox="[151,1436,181,1544]" targetPageId="10" targetType="figure">
<paragraph id="8B85E638FFB62221FF1AF9EB1738F904" blockId="10.[151,1437,1568,1698]" pageId="10" pageNumber="161">
<emphasis id="B94E3A2AFFB62221FF1AF9EB1544F99E" bold="true" box="[152,279,1568,1592]" pageId="10" pageNumber="161">FIGURE 4.</emphasis>
<taxonomicName id="4C3A9DBBFFB62221FE9EF9EB1627F99E" authority="POLY-UFPB" authorityName="POLY-UFPB" box="[284,628,1568,1592]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="161" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB62221FE9EF9EB15B6F99E" box="[284,485,1568,1592]" italics="true" pageId="10" pageNumber="161">Leodice calcaricola</emphasis>
POLY-UFPB
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2020.
<emphasis id="B94E3A2AFFB62221FD38F9EB169FF99E" bold="true" box="[698,716,1568,1592]" pageId="10" pageNumber="161">A</emphasis>
—anterior end, dorsal view;
<emphasis id="B94E3A2AFFB62221FC68F9EB17A8F99E" bold="true" box="[1002,1019,1568,1592]" pageId="10" pageNumber="161">B</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFB62221FAA0F9EB1167F99E" bold="true" box="[1314,1332,1568,1592]" pageId="10" pageNumber="161">C</emphasis>
—anterior end, lateral view;
<emphasis id="B94E3A2AFFB62221FECAF9881527F9FD" bold="true" box="[328,372,1603,1627]" pageId="10" pageNumber="161">D—</emphasis>
parapodium 10, compound falcigers chaetae;
<emphasis id="B94E3A2AFFB62221FCB8F9881718F9FD" bold="true" box="[826,843,1603,1627]" pageId="10" pageNumber="161">E</emphasis>
—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri;
<emphasis id="B94E3A2AFFB62221FD07F9AD16FEF9D8" bold="true" box="[645,685,1638,1662]" pageId="10" pageNumber="161">F—</emphasis>
parapodium 20, aciculae;
<emphasis id="B94E3A2AFFB62221FC33F9AD1796F9D8" bold="true" box="[945,965,1638,1662]" pageId="10" pageNumber="161">G</emphasis>
—parapodium 45, compound falcigers chaetae and pectinae chaetae;
<emphasis id="B94E3A2AFFB62221FEF4F94115D9F904" bold="true" box="[374,394,1674,1698]" pageId="10" pageNumber="161">H</emphasis>
—parapodium 31, compound falcigers chaetae.
</paragraph>
</caption>
<paragraph id="8B85E638FFB62221FF45F922153FF85C" blockId="10.[151,1437,1769,2042]" pageId="10" pageNumber="161">
Prostomium bilobed, truncate anteriorly, with conspicuous ventral buccal lips, dorsally inflated, shorter and narrower than peristomium, with shallow median groove, visible dorsally and ventrally. Prostomial appendages arranged in semi-circle, evenly spaced (
<figureCitation id="1301FABDFFB62221FDE9F8E416BEF8EF" box="[619,749,1839,1865]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="10" pageNumber="161">Fig. 4A, B</figureCitation>
). Palpophores and ceratophores short and ring-shaped, not articulated. Ceratostyles and palpostyles tapering in distal half, with regular moniliform articulations, distally drop-shaped (
<figureCitation id="1301FABDFFB62221FF77F8BD15C1F836" box="[245,402,1910,1936]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="10" pageNumber="161">Fig. 4A, B, C</figureCitation>
). Median antennae longest and palps shortest; palps with 37 articulations, reaching middle of first peristomial ring to second peristomial ring; lateral antennae with 414 articulations, reaching chaetiger 14; median antenna with 819 articulations, reaching chaetiger 39. Pair of eyes present, posterior to palps and lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFB72220FF45FF5C10F2FEBD" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">
Peristomium cylindrical, laterally longer than medially (
<figureCitation id="1301FABDFFB72220FCC4FF5C17B5FF17" box="[838,998,151,177]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4A, B, C</figureCitation>
); dorsal anterior margin covering base of median and lateral antennae. Ventrolateral lips conspicuously inflated. First peristomial ring about ¾ of total peristomium length, separation between rings visible dorsally, ventrally, and laterally. Peristomial cirri tapering to digitiform tips with 24 articulations, almost reaching anterior edge of first peristomial ring.
</paragraph>
<paragraph id="8B85E638FFB72220FF45FEEE1139FE57" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">Maxillary formula 1+1, (57)+(810), (78)+0, (67)+(911), 1+1. MxI forceps shaped with not extended falcal, outer edge straight, rectangular; basal inner edge a curvature where most posterior region of MxII fits. MxIII partially ventral to MxII, anterior end part of distal arch with left MxIV. Left attachment lamellae tapering on left side, right lamella semi-circular in shape. MxIV left with teeth decreasing in size towards the innermost portion. MxV wide, with a small prominent tooth. MxVI absent. MxIV left attachment lamella semi-circular shape, tapering towards the right portion, right MxIV attachment lamella semi-circular in shape, tapering towards left portion.</paragraph>
<paragraph id="8B85E638FFB72220FF45FE301129FDFA" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">
Branchiae pectinate, present in less than 55% of the total number of chaetigers, starting from chaetiger 57 to chaetiger 1224. Branchiae initially with single filament. Maximum number of branchial filaments
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chaetiger 1014. Central axis longer than branchial filaments and both longer than notopodial cirri where best developed.
</paragraph>
<paragraph id="8B85E638FFB72220FF45FDAE174CFD05" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">Chaetal lobes truncated along the body, aciculae emerging dorsally. Prechaetal lobes low transverse folds. Postchaetal lobes rounded, almost as long as chaetal lobe.</paragraph>
<paragraph id="8B85E638FFB72220FF45FD661050FCA8" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">Anterior notopodial cirri tapering distally to pointed tips, with 23 articulations. Ventral cirri longer than chaetal lobes in all parapodia, robust, digitiform until chaetiger 34, from chaetiger 46 basally inflated with tapering to digitiform tips until chaetiger 1630, after this, inflated base decreases in size.</paragraph>
<paragraph id="8B85E638FFB72220FF45FCDC1512FBD4" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">
Limbate chaetae slender, distally tapered, longer than other chaetae. Pectinate chaetae, flat, 810 teeth, one marginal tooth longer and thicker than other teeth (
<figureCitation id="1301FABDFFB72220FD5EFCF0176BFCF3" box="[732,824,827,853]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4G</figureCitation>
). Compound falcigers chaetae bidentate present along the whole body and tridentate from chaetiger 23. Anterior bidentate compound with article tapering from proximal to distal end; triangular proximal tooth directed laterally, distal tooth directed laterally or obliquely, both similar in size (
<figureCitation id="1301FABDFFB72220FF70FC6D151DFC66" box="[242,334,934,960]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4D</figureCitation>
); articles stouter towards the posterior end, with wider guards, triangular proximal tooth directed laterally, larger than distal tooth. Tridentate compound falcigers with small third apical tooth directed obliquely upwards (
<figureCitation id="1301FABDFFB72220FE8AFC261535FBA1" box="[264,358,1005,1031]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4H</figureCitation>
). Aciculae 12 aciculae per parapodia, yellow; anterior ones with mucronate tips, tapering or blunt; median bifid, few slightly curved (
<figureCitation id="1301FABDFFB72220FDE4FBDA168FFB8D" box="[614,732,1041,1067]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4E, F</figureCitation>
); posterior bifid, more curved than those of median parapodia. Subacicular hooks single, yellow, tridentate, present from chaetiger 1422; guards covering proximal and first distal teeth (
<figureCitation id="1301FABDFFB72220FF5FFB931566FBD4" box="[221,309,1112,1138]" captionStart="FIGURE 4" captionStartId="10.[152,255,1568,1592]" captionTargetBox="[151,1436,181,1544]" captionTargetId="figure-23@10.[151,1436,181,1544]" captionTargetPageId="10" captionText="FIGURE 4. Leodice calcaricola POLY-UFPB2020. A—anterior end, dorsal view; B—anterior end, ventral view; C—anterior end, lateral view; D—parapodium 10, compound falcigers chaetae; E—parapodium 23, compound falcigers chaetae, subacicular hook, aciculae, notopodial cirri and ventral cirri; F—parapodium 20, aciculae; G—parapodium 45, compound falcigers chaetae and pectinae chaetae; H—parapodium 31, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219188" httpUri="https://zenodo.org/record/13219188/files/figure.png" pageId="11" pageNumber="162">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB72220FF45FBB01692FB1F" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">Pygidium with two pairs of pygidial cirri; dorsal cirri long, with 67 regular articulations, about six times longer than ventral; ventral pygidial cirri not articulated.</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB72220FF45FB091679FA82" pageId="11" pageNumber="162" type="distribution">
<paragraph id="8B85E638FFB72220FF45FB091679FA82" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">
<emphasis id="B94E3A2AFFB72220FF45FB091532FB7A" bold="true" box="[199,353,1218,1244]" pageId="11" pageNumber="162">Distribution.</emphasis>
<taxonomicName id="4C3A9DBBFFB72220FEEAFB08161BFB7B" box="[360,584,1219,1245]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB72220FEEAFB08161BFB7B" box="[360,584,1219,1245]" italics="true" pageId="11" pageNumber="162">Leodice calcaricola</emphasis>
</taxonomicName>
was recently described from the coast of
<collectingRegion id="49FE28DAFFB72220FB9DFB0810E2FB7B" box="[1055,1201,1219,1245]" country="Brazil" name="Paraiba" pageId="11" pageNumber="162">Paraíba state</collectingRegion>
, based on specimens collected intertidally to depths of
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(
<bibRefCitation id="EFAB9BC9FFB72220FDDDFB2D1714FAA6" author="Bergamo, G. &amp; Carrerette, O. &amp; Zanol, J. &amp; Nogueira, J. M. M." box="[607,839,1254,1281]" pageId="11" pageNumber="162" pagination="376 - 390" refId="ref13400" refString="Bergamo, G., Carrerette, O., Zanol, J. &amp; Nogueira, J. M. M. (2018) Species of Eunicidae (Annelida, Errantia, Eunicida) from rhodolith beds off the States of Paraiba and Pernambuco, northeastern Brazilian coast. Zootaxa, 4521 (3) 376 - 390. https: // doi. org / 10.11646 / zootaxa. 4521.3.4" type="journal article" year="2018">
Bergamo
<emphasis id="B94E3A2AFFB72220FD4FFB2C16AEFAA6" box="[717,765,1254,1280]" italics="true" pageId="11" pageNumber="162">et al</emphasis>
. 2018
</bibRefCitation>
). The present record expands bathymetric distribution of the species to
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deep (
<figureCitation id="1301FABDFFB72220FE55FAC1164AFA82" box="[471,537,1290,1316]" captionStart="FIGURE 1" captionStartId="3.[152,255,1913,1937]" captionTargetBox="[151,1429,195,1889]" captionTargetId="figure-19@3.[151,1436,181,1889]" captionTargetPageId="3" captionText="FIGURE 1. General map of the stations of the Marine Algae project along the Continental Shelf of the State of Paraíba where Leodice species have been recorded." figureDoi="http://doi.org/10.5281/zenodo.13219176" httpUri="https://zenodo.org/record/13219176/files/figure.png" pageId="11" pageNumber="162">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB72220FF45FAE51031F9E7" pageId="11" pageNumber="162" type="discussion">
<paragraph id="8B85E638FFB72220FF45FAE51031F9E7" blockId="11.[151,1437,151,1602]" pageId="11" pageNumber="162">
<emphasis id="B94E3A2AFFB72220FF45FAE51568FAEE" bold="true" box="[199,315,1326,1352]" pageId="11" pageNumber="162">Remarks.</emphasis>
The specimens described here share with
<taxonomicName id="4C3A9DBBFFB72220FCC6FAE61516FACD" authority="Bergamo, Carrerette, Zanol &amp; Nogueira, 2018" authorityName="Bergamo, Carrerette, Zanol &amp; Nogueira" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB72220FCC6FAE61079FAE1" box="[836,1066,1325,1351]" italics="true" pageId="11" pageNumber="162">Leodice calcaricola</emphasis>
<bibRefCitation id="EFAB9BC9FFB72220FBBAFAE51516FACD" author="Bergamo, G. &amp; Carrerette, O. &amp; Zanol, J. &amp; Nogueira, J. M. M." pageId="11" pageNumber="162" pagination="376 - 390" refId="ref13400" refString="Bergamo, G., Carrerette, O., Zanol, J. &amp; Nogueira, J. M. M. (2018) Species of Eunicidae (Annelida, Errantia, Eunicida) from rhodolith beds off the States of Paraiba and Pernambuco, northeastern Brazilian coast. Zootaxa, 4521 (3) 376 - 390. https: // doi. org / 10.11646 / zootaxa. 4521.3.4" type="journal article" year="2018">Bergamo, Carrerette, Zanol &amp; Nogueira, 2018</bibRefCitation>
</taxonomicName>
articulated antennae and moniliform palps, articulated peristomial cirri, pectinate branchiae present in less than 55% of total number of segments, yellow tridentate subacicular hooks, and falcigers compound chaetae bidentate and tridentate along body. The specimens analyzed resemble the original description of
<taxonomicName id="4C3A9DBBFFB72220FB63FA52112CFA14" authorityName="Bergamo, Carrerette, Zanol &amp; Nogueira" authorityYear="2018" box="[1249,1407,1432,1458]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="calcaricola">
<emphasis id="B94E3A2AFFB72220FB63FA52112CFA14" box="[1249,1407,1432,1458]" italics="true" pageId="11" pageNumber="162">L. calcaricola</emphasis>
</taxonomicName>
in most of its characteristics, except for 1. maximum number of branchial filaments, 13 filaments original description versus 14 filaments in our specimens; 2. presence of anadont and heterodont pectinate chaetae in the original description versus only heterodont pectinate chaetae in our specimens; and 3. paired subacicular hooks in some parapodia in the original description versus single subacicular hooks in our specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="8B85E638FFB72220FF15F94116B4F903" blockId="11.[151,743,1674,1738]" box="[151,743,1674,1701]" pageId="11" pageNumber="162">
<heading id="D0CD5154FFB72220FF15F94116B4F903" bold="true" box="[151,743,1674,1701]" fontSize="11" level="1" pageId="11" pageNumber="162" reason="1">
<taxonomicName id="4C3A9DBBFFB72220FF15F94116B4F903" ID-CoL="3T6D9" authority="(Zanol, Paiva &amp; Attolini, 2000)" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[151,743,1674,1701]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="marcusi">
<emphasis id="B94E3A2AFFB72220FF15F94116B4F903" bold="true" box="[151,743,1674,1701]" pageId="11" pageNumber="162">
<emphasis id="B94E3A2AFFB72220FF15F941150CF903" bold="true" box="[151,351,1674,1701]" italics="true" pageId="11" pageNumber="162">Leodice marcusi</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB72220FEECF940168DF903" author="Zanol, J. &amp; Paiva, P. C. &amp; Attolini, F. S." box="[366,734,1675,1701]" pageId="11" pageNumber="162" pagination="449 - 463" refId="ref14731" refString="Zanol, J., Paiva, P. C. &amp; Attolini, F. S. (2000) Eunice and Palola (Eunicidae: Polychaeta) from the eastern Brazilian Coast (13 ° 00 ' - 22 ° 30 ' S). Bulletin of Marine Science, 67 (1), 449 - 463." type="journal article" year="2000">Zanol, Paiva &amp; Attolini, 2000</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB72220FF15F9641558F96F" box="[151,267,1711,1738]" pageId="11" pageNumber="162" type="description">
<paragraph id="8B85E638FFB72220FF15F9641558F96F" blockId="11.[151,743,1674,1738]" box="[151,267,1711,1738]" pageId="11" pageNumber="162">
(
<figureCitation id="1301FABDFFB72220FF1DF96414BBF96C" box="[159,232,1711,1738]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="11" pageNumber="162">Figs 5</figureCitation>
,
<figureCitation id="1301FABDFFB72220FF76F9641550F96F" box="[244,259,1711,1737]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="11" pageNumber="162">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB72220FF15F93B166AF880" pageId="11" pageNumber="162" type="reference_group">
<paragraph id="8B85E638FFB72220FF15F93B16F0F8A1" blockId="11.[151,675,1775,1830]" box="[151,675,1775,1800]" pageId="11" pageNumber="162">
<treatmentCitationGroup id="AB2AC116FFB72220FF15F93B16F0F8A1" box="[151,675,1775,1800]" pageId="11" pageNumber="162">
<treatmentCitation id="0A9BC029FFB72220FF15F93B16CDF8A1" author="Zanol, J. &amp; Paiva, P. C. &amp; Attolini, F. S." box="[151,670,1775,1800]" page="454" pageId="11" pageNumber="162" year="2000">
<taxonomicName id="4C3A9DBBFFB72220FF15F93B16CDF8A1" ID-CoL="3CF8W" authority="Zanol, Paiva &amp; Attolini, 2000: 454" authorityName="Zanol, Paiva &amp; Attolini" authorityPageNumber="454" authorityYear="2000" box="[151,670,1775,1800]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="marcusi">
<emphasis id="B94E3A2AFFB72220FF15F93B1566F8A1" box="[151,309,1776,1799]" italics="true" pageId="11" pageNumber="162">Eunice marcusi</emphasis>
<bibRefCitation id="EFAB9BC9FFB72220FEB9F92416CDF8A1" author="Zanol, J. &amp; Paiva, P. C. &amp; Attolini, F. S." box="[315,670,1775,1800]" pageId="11" pageNumber="162" pagination="449 - 463" refId="ref14731" refString="Zanol, J., Paiva, P. C. &amp; Attolini, F. S. (2000) Eunice and Palola (Eunicidae: Polychaeta) from the eastern Brazilian Coast (13 ° 00 ' - 22 ° 30 ' S). Bulletin of Marine Science, 67 (1), 449 - 463." type="journal article" year="2000">Zanol, Paiva &amp; Attolini, 2000: 454</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B85E638FFB72220FF15F8C5166AF880" blockId="11.[151,675,1775,1830]" box="[151,569,1806,1830]" pageId="11" pageNumber="162">
<treatmentCitationGroup id="AB2AC116FFB72220FF15F8C5166AF880" box="[151,569,1806,1830]" pageId="11" pageNumber="162">
<emphasis id="B94E3A2AFFB72220FF15F8C51515F880" box="[151,326,1806,1830]" italics="true" pageId="11" pageNumber="162">
<taxonomicName id="4C3A9DBBFFB72220FF15F8C51511F880" ID-CoL="3T6D9" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[151,322,1806,1830]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="162" phylum="Annelida" rank="species" species="marcusi">Leodice marcusi</taxonomicName>
.
</emphasis>
<treatmentCitation id="0A9BC029FFB72220FEDDF8C51666F880" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[351,565,1806,1830]" page="90" pageId="11" pageNumber="162" year="2014">
<bibRefCitation id="EFAB9BC9FFB72220FEDDF8C51666F880" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[351,565,1806,1830]" pageId="11" pageNumber="162" pagination="79 - 100" refId="ref14618" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="B94E3A2AFFB72220FE1DF8C41580F880" box="[415,467,1806,1830]" italics="true" pageId="11" pageNumber="162">et al.</emphasis>
2014: 90
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB72226FF15F89B1000FE99" lastPageId="13" lastPageNumber="164" pageId="11" pageNumber="162" type="materials_examined">
<paragraph id="8B85E638FFB72220FF15F89B1168F85C" blockId="11.[151,1437,1872,2043]" pageId="11" pageNumber="162">
<emphasis id="B94E3A2AFFB72220FF15F89B152EF8CC" bold="true" box="[151,381,1872,1898]" pageId="11" pageNumber="162">Material examined.</emphasis>
<materialsCitation id="3B52EC65FFB72220FE07F89B158BF828" collectingDate="1992-02" collectionCode="MNRJP" elevation="44" latitude="-21.690277" longLatPrecision="21" longitude="-40.34611" pageId="11" pageNumber="162" specimenCount="1" typeStatus="holotype">
<emphasis id="B94E3A2AFFB72220FE07F89B1672F8CC" bold="true" box="[389,545,1872,1898]" pageId="11" pageNumber="162">
<typeStatus id="5481589AFFB72220FE07F89B1672F8CC" box="[389,545,1872,1898]" pageId="11" pageNumber="162" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, one complete specimen (
<collectionCode id="ED2B7EFDFFB72220FCC1F89B17CDF8CD" box="[835,926,1872,1899]" pageId="11" pageNumber="162">MNRJP</collectionCode>
003258),
<geoCoordinate id="EE0E80FFFFB72220FB97F89B10CAF8CD" box="[1045,1177,1872,1899]" degrees="21" direction="south" minutes="41" orientation="latitude" pageId="11" pageNumber="162" precision="15" seconds="25" value="-21.690277">21°4125”S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB72220FB24F89B1160F8CC" box="[1190,1331,1872,1898]" degrees="40" direction="west" minutes="20" orientation="longitude" pageId="11" pageNumber="162" precision="15" seconds="46" value="-40.34611">40°2046”W</geoCoordinate>
, January/
<date id="FF84C0F8FFB72220FF15F8BF156CF828" box="[151,319,1908,1934]" pageId="11" pageNumber="162" value="1992-02">
<collectingDate id="EFC03910FFB72220FF15F8BF156CF828" box="[151,319,1908,1934]" pageId="11" pageNumber="162" value="1992-02">February 1992</collectingDate>
</date>
,
<elevation id="0017010BFFB72220FECDF8BF158BF828" box="[335,472,1908,1935]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="162" unit="m" value="44.0">
<quantity id="4CC24BDDFFB72220FECDF8BF15D8F829" box="[335,395,1908,1935]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="162" unit="m" value="44.0">44 m</quantity>
depth
</elevation>
</materialsCitation>
.
<materialsCitation id="3B52EC65FFB72220FE64F8BF165DF814" collectingDate="1992-03" collectionCode="MNRJP" elevation="44" latitude="-21.690277" longLatPrecision="21" longitude="-40.34611" pageId="11" pageNumber="162" specimenCount="1" typeStatus="paratype">
<emphasis id="B94E3A2AFFB72220FE64F8BF1624F828" bold="true" box="[486,631,1908,1934]" pageId="11" pageNumber="162">
<typeStatus id="5481589AFFB72220FE64F8BF1624F828" box="[486,631,1908,1934]" pageId="11" pageNumber="162" type="paratype">PARATYPE</typeStatus>
</emphasis>
one complete specimen (
<collectionCode id="ED2B7EFDFFB72220FC23F8BF17AEF829" box="[929,1021,1908,1935]" pageId="11" pageNumber="162">MNRJP</collectionCode>
003259),
<geoCoordinate id="EE0E80FFFFB72220FBF5F8BF10A8F829" box="[1143,1275,1908,1935]" degrees="21" direction="south" minutes="41" orientation="latitude" pageId="11" pageNumber="162" precision="15" seconds="25" value="-21.690277">21°4125”S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB72220FA89F8BF11CBF828" box="[1291,1432,1908,1934]" degrees="40" direction="west" minutes="20" orientation="longitude" pageId="11" pageNumber="162" precision="15" seconds="46" value="-40.34611">40°2046”W</geoCoordinate>
, February/
<date id="FF84C0F8FFB72220FE80F85315DBF814" box="[258,392,1944,1970]" pageId="11" pageNumber="162" value="1992-03">
<collectingDate id="EFC03910FFB72220FE80F85315DBF814" box="[258,392,1944,1970]" pageId="11" pageNumber="162" value="1992-03">March 1992</collectingDate>
</date>
,
<elevation id="0017010BFFB72220FE10F853165DF814" box="[402,526,1944,1971]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="162" unit="m" value="44.0">
<quantity id="4CC24BDDFFB72220FE10F853159BF815" box="[402,456,1944,1971]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="162" unit="m" value="44.0">44 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B52EC65FFB72220FD9BF85315F5F870" collectingDate="1992-03" collectionCode="MNRJP" elevation="29" latitude="-21.352222" longLatPrecision="21" longitude="-40.58361" pageId="11" pageNumber="162" specimenCount="1" typeStatus="paratype">
six incomplete specimens (
<collectionCode id="ED2B7EFDFFB72220FCBFF85317CAF814" box="[829,921,1944,1970]" pageId="11" pageNumber="162">MNRJP</collectionCode>
003260),
<geoCoordinate id="EE0E80FFFFB72220FB8FF85310C3F814" box="[1037,1168,1944,1970]" degrees="21" direction="south" minutes="21" orientation="latitude" pageId="11" pageNumber="162" precision="15" seconds="08" value="-21.352222">21°2108”S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB72220FB19F853117BF815" box="[1179,1320,1944,1971]" degrees="40" direction="west" minutes="35" orientation="longitude" pageId="11" pageNumber="162" precision="15" seconds="01" value="-40.58361">40°3501”W</geoCoordinate>
, February/
<date id="FF84C0F8FFB72220FF15F877154DF870" box="[151,286,1980,2006]" pageId="11" pageNumber="162" value="1992-03">
<collectingDate id="EFC03910FFB72220FF15F877154DF870" box="[151,286,1980,2006]" pageId="11" pageNumber="162" value="1992-03">March 1992</collectingDate>
</date>
,
<elevation id="0017010BFFB72220FEABF87715F5F870" box="[297,422,1980,2007]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="162" unit="m" value="29.0">
<quantity id="4CC24BDDFFB72220FEABF877150CF871" box="[297,351,1980,2007]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="11" pageNumber="162" unit="m" value="29.0">29 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B52EC65FFB72220FE33F8771545F85C" collectionCode="MNRJP" elevation="82" latitude="-22.878334" longLatPrecision="21" longitude="-41.154167" pageId="11" pageNumber="162" specimenCount="1" typeStatus="paratype">
six incomplete and three complete specimens (
<collectionCode id="ED2B7EFDFFB72220FC2EF877105AF870" box="[940,1033,1980,2006]" pageId="11" pageNumber="162">MNRJP</collectionCode>
003261),
<geoCoordinate id="EE0E80FFFFB72220FBFFF8771152F871" box="[1149,1281,1980,2007]" degrees="22" direction="south" minutes="52" orientation="latitude" pageId="11" pageNumber="162" precision="15" seconds="42" value="-22.878334">22°5242”S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB72220FA89F87711CBF871" box="[1291,1432,1980,2007]" degrees="41" direction="west" minutes="09" orientation="longitude" pageId="11" pageNumber="162" precision="15" seconds="15" value="-41.154167">41°0915”W</geoCoordinate>
,
<elevation id="0017010BFFB72220FF15F82B1545F85C" box="[151,278,2016,2043]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="162" unit="m" value="82.0">
<quantity id="4CC24BDDFFB72220FF15F82B149DF85D" box="[151,206,2016,2043]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="162" unit="m" value="82.0">82 m</quantity>
depth
</elevation>
</materialsCitation>
;
<materialsCitation id="3B52EC65FFB72220FEA3F82A1164F85C" box="[289,1335,2016,2043]" collectingDate="1991-07" collectionCode="MNRJP" elevation="20" latitude="-21.345001" longLatPrecision="21" longitude="-40.594997" pageId="11" pageNumber="162" specimenCount="1" typeStatus="paratype">
one incomplete specimen (
<collectionCode id="ED2B7EFDFFB72220FDC4F82B16F1F85C" box="[582,674,2016,2042]" pageId="11" pageNumber="162">MNRJP</collectionCode>
003262),
<geoCoordinate id="EE0E80FFFFB72220FC95F82B17CFF85C" box="[791,924,2016,2042]" degrees="21" direction="south" minutes="20" orientation="latitude" pageId="11" pageNumber="162" precision="15" seconds="42" value="-21.345001">21°2042”S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB72220FC25F82B1067F85D" box="[935,1076,2016,2043]" degrees="40" direction="west" minutes="35" orientation="longitude" pageId="11" pageNumber="162" precision="15" seconds="42" value="-40.594997">40°3542”W</geoCoordinate>
,
<date id="FF84C0F8FFB72220FBBDF82B10FFF85C" box="[1087,1196,2016,2042]" pageId="11" pageNumber="162" value="1991-07">
<collectingDate id="EFC03910FFB72220FBBDF82B10FFF85C" box="[1087,1196,2016,2042]" pageId="11" pageNumber="162" value="1991-07">July 1991</collectingDate>
</date>
,
<elevation id="0017010BFFB72220FB3AF82B1164F85C" box="[1208,1335,2016,2043]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="162" unit="m" value="20.0">
<quantity id="4CC24BDDFFB72220FB3AF82B10BCF85D" box="[1208,1263,2016,2043]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="162" unit="m" value="20.0">20 m</quantity>
depth
</elevation>
</materialsCitation>
.
</paragraph>
<caption id="DF45B6B0FFB02227FF1AF9D11704F9F0" ID-DOI="http://doi.org/10.5281/zenodo.13219192" ID-Zenodo-Dep="13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="12" pageNumber="163" startId="12.[152,255,1562,1586]" targetBox="[151,1436,181,1538]" targetPageId="12" targetType="figure">
<paragraph id="8B85E638FFB02227FF1AF9D11704F9F0" blockId="12.[151,1436,1562,1622]" pageId="12" pageNumber="163">
<emphasis id="B94E3A2AFFB02227FF1AF9D1154AF994" bold="true" box="[152,281,1562,1586]" pageId="12" pageNumber="163">FIGURE 5.</emphasis>
<taxonomicName id="4C3A9DBBFFB02227FEA2F9D1159BF994" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[288,456,1562,1586]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="163" phylum="Annelida" rank="species" species="marcusi">
<emphasis id="B94E3A2AFFB02227FEA2F9D1159BF994" box="[288,456,1562,1586]" italics="true" pageId="12" pageNumber="163">Leodice marcusi</emphasis>
</taxonomicName>
. A, B, D MNRJP003258. C, MNRJP003261.
<emphasis id="B94E3A2AFFB02227FC3BF9D11798F994" bold="true" box="[953,971,1562,1586]" pageId="12" pageNumber="163">A</emphasis>
—anterior end, dorsal view;
<emphasis id="B94E3A2AFFB02227FB73F9D11151F994" bold="true" box="[1265,1282,1562,1586]" pageId="12" pageNumber="163">B</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFB02227FEA0F9F51567F9F0" bold="true" box="[290,308,1598,1622]" pageId="12" pageNumber="163">C</emphasis>
maxillae, dorsal view;
<emphasis id="B94E3A2AFFB02227FDAAF9F51607F9F0" bold="true" box="[552,596,1598,1622]" pageId="12" pageNumber="163">D—</emphasis>
anterior end, lateral view.
</paragraph>
</caption>
<paragraph id="8B85E638FFB02226FF45F96B1000FE99" blockId="12.[151,1437,1695,2042]" lastBlockId="13.[151,1437,151,319]" lastPageId="13" lastPageNumber="164" pageId="12" pageNumber="163">
<materialsCitation id="3B52EC65FFB02226FF45F96B101DFE99" collectionCode="POLY-UFPB" lastPageId="13" lastPageNumber="164" location="Paraiba State" pageId="12" pageNumber="163" specimenCount="48" stateProvince="Paraiba">
NON-TYPE material
<emphasis id="B94E3A2AFFB02227FE42F9541638F91F" bold="true" box="[448,619,1695,1722]" pageId="12" pageNumber="163">
<collectingRegion id="49FE28DAFFB02227FE42F9541635F91F" box="[448,614,1695,1722]" country="Brazil" name="Paraiba" pageId="12" pageNumber="163">Paraíba State</collectingRegion>
,
</emphasis>
<specimenCount id="9D3C2DB1FFB02227FDF7F96B1741F91C" box="[629,786,1696,1722]" count="48" pageId="12" pageNumber="163" type="generic">48 specimens</specimenCount>
: one complete and two incomplete specimens, station 14 (
<collectionCode id="ED2B7EFDFFB02227FF1DF908156FF97B" box="[159,316,1731,1757]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1991); three incomplete specimens, station 29 (
<collectionCode id="ED2B7EFDFFB02227FCF6F908105CF97B" box="[884,1039,1731,1757]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1988;
<collectionCode id="ED2B7EFDFFB02227FBE1F90810AFF97B" box="[1123,1276,1731,1757]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1989;
<collectionCode id="ED2B7EFDFFB02227FAD2F90814B2F8A7" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1990); two incomplete specimens, station 37 (
<collectionCode id="ED2B7EFDFFB02227FC86F92D17CCF8A6" box="[772,927,1766,1792]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1986;
<collectionCode id="ED2B7EFDFFB02227FC73F92C10DFF8A7" box="[1009,1164,1767,1793]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1987); one incomplete, station 38 (
<collectionCode id="ED2B7EFDFFB02227FE90F8C115FCF882" box="[274,431,1802,1828]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1993); six incomplete specimen, station 39 (
<collectionCode id="ED2B7EFDFFB02227FC1CF8C11069F883" box="[926,1082,1802,1829]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1985); one complete specimen, station 45 (
<collectionCode id="ED2B7EFDFFB02227FEA0F8E515EDF8EE" box="[290,446,1838,1864]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1979); three incomplete specimens, station 46 (
<collectionCode id="ED2B7EFDFFB02227FB81F8E510CCF8EE" box="[1027,1183,1838,1864]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1980); two complete and four incomplete specimens, station 47 (
<collectionCode id="ED2B7EFDFFB02227FD23F89A176FF8CD" box="[673,828,1873,1899]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1976;
<collectionCode id="ED2B7EFDFFB02227FC12F89A1079F8CD" box="[912,1066,1873,1899]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1977;
<collectionCode id="ED2B7EFDFFB02227FBFFF89A114BF8CD" box="[1149,1304,1873,1899]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1978); one incomplete specimen, station 54 (
<collectionCode id="ED2B7EFDFFB02227FD90F8BF16FDF829" box="[530,686,1908,1935]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1975); one incomplete specimen, station 55 (
<collectionCode id="ED2B7EFDFFB02227FB33F8BE111DF829" box="[1201,1358,1909,1935]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1973); one incomplete specimen, station 56 (
<collectionCode id="ED2B7EFDFFB02227FDCCF85316B9F814" box="[590,746,1944,1970]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1974); one incomplete specimen, station 61 (
<collectionCode id="ED2B7EFDFFB02227FB7BF85311DDF814" box="[1273,1422,1944,1970]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1981); one incomplete specimen, station 62 (
<collectionCode id="ED2B7EFDFFB02227FD23F877176EF870" box="[673,829,1980,2006]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1982); one incomplete specimen, station 63 (
<collectionCode id="ED2B7EFDFFB02227FACFF87714B3F85F" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
2000); one complete specimen, station 64 (
<collectionCode id="ED2B7EFDFFB02227FD69F81417DBF85F" box="[747,904,2015,2041]" pageId="12" pageNumber="163">POLY-UFPB</collectionCode>
1983); two incomplete specimens, station 65 (
<collectionCode id="ED2B7EFDFFB12226FF1DFF5C1568FF17" box="[159,315,151,177]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1984); one incomplete specimen, station 68 (
<collectionCode id="ED2B7EFDFFB12226FCEEFF5C105BFF17" box="[876,1032,151,177]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
2001); one incomplete specimen, station 76 (
<collectionCode id="ED2B7EFDFFB12226FE99FF7115E4FF72" box="[283,439,186,212]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1992); one complete and two incomplete specimens, station 80 (
<collectionCode id="ED2B7EFDFFB12226FB33FF71111EFF72" box="[1201,1357,186,212]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1994); one incomplete specimen, station 85 (
<collectionCode id="ED2B7EFDFFB12226FDB9FF161684FF5E" box="[571,727,221,248]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1995); two complete and two incomplete specimens, station 86 (
<collectionCode id="ED2B7EFDFFB12226FF1DFECA1568FEBD" box="[159,315,257,283]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1996); one incomplete specimen, station 86 (
<collectionCode id="ED2B7EFDFFB12226FCB4FECA1781FEBD" box="[822,978,257,283]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1997); one incomplete specimen, station 90 (
<collectionCode id="ED2B7EFDFFB12226FF40FEEE150DFE99" box="[194,350,293,319]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1998); two incomplete specimens, station 93 (
<collectionCode id="ED2B7EFDFFB12226FCE5FEEE1050FE99" box="[871,1027,293,319]" pageId="13" pageNumber="164">POLY-UFPB</collectionCode>
1999)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF45B6B0FFB12226FF1AF88C109FF849" ID-DOI="http://doi.org/10.5281/zenodo.13219194" ID-Zenodo-Dep="13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="13" pageNumber="164" startId="13.[152,255,1863,1887]" targetBox="[151,1436,343,1840]" targetPageId="13" targetType="figure">
<paragraph id="8B85E638FFB12226FF1AF88C109FF849" blockId="13.[151,1436,1863,2032]" pageId="13" pageNumber="164">
<emphasis id="B94E3A2AFFB12226FF1AF88C154BF8F9" bold="true" box="[152,280,1863,1887]" pageId="13" pageNumber="164">FIGURE 6.</emphasis>
<emphasis id="B94E3A2AFFB12226FE9CF88C159FF8F9" box="[286,460,1863,1887]" italics="true" pageId="13" pageNumber="164">
<taxonomicName id="4C3A9DBBFFB12226FE9CF88C159BF8F9" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[286,456,1863,1887]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="164" phylum="Annelida" rank="species" species="marcusi">Leodice marcusi</taxonomicName>
.
</emphasis>
A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261.
<emphasis id="B94E3A2AFFB12226FBEFF88C102CF8F9" bold="true" box="[1133,1151,1863,1887]" pageId="13" pageNumber="164">A</emphasis>
—parapodium 5, compound falcigers;
<emphasis id="B94E3A2AFFB12226FE82F8A71542F822" bold="true" box="[256,273,1900,1924]" pageId="13" pageNumber="164">B</emphasis>
—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri;
<emphasis id="B94E3A2AFFB12226FB4FF8A010AAF825" bold="true" box="[1229,1273,1899,1923]" pageId="13" pageNumber="164">C—</emphasis>
parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae;
<emphasis id="B94E3A2AFFB12226FCFEF85B17FBF80E" bold="true" box="[892,936,1936,1960]" pageId="13" pageNumber="164">D—</emphasis>
parapodium 43, compound falcigers chaetae;
<emphasis id="B94E3A2AFFB12226FAF0F85B11D0F80E" bold="true" box="[1394,1411,1936,1960]" pageId="13" pageNumber="164">E</emphasis>
— parapodium 110, compound falciger chaetae;
<emphasis id="B94E3A2AFFB12226FDECF87F162EF86A" bold="true" box="[622,637,1972,1996]" pageId="13" pageNumber="164">F</emphasis>
—parapodium 44, pectinate chaetae;
<emphasis id="B94E3A2AFFB12226FC7FF8781042F86D" bold="true" box="[1021,1041,1971,1995]" pageId="13" pageNumber="164">G</emphasis>
—parapodium 110, subacicular hooks;
<emphasis id="B94E3A2AFFB12226FF15F81314F8F856" bold="true" box="[151,171,2008,2032]" pageId="13" pageNumber="164">H</emphasis>
—parapodium 3, pectinate chaetae;
<emphasis id="B94E3A2AFFB12226FD9AF8131671F856" bold="true" box="[536,546,2008,2032]" pageId="13" pageNumber="164">I</emphasis>
—parapodium 110, aciculae;
<emphasis id="B94E3A2AFFB12226FCCFF8131709F856" bold="true" box="[845,858,2008,2032]" pageId="13" pageNumber="164">J</emphasis>
—parapodium 67, subacicular hook.
</paragraph>
</caption>
<subSubSection id="C320B5B3FFB22224FF45FF5D1513FE2E" lastPageId="15" lastPageNumber="166" pageId="14" pageNumber="165" type="description">
<paragraph id="8B85E638FFB22225FF45FF5D1082FEC3" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
<emphasis id="B94E3A2AFFB22225FF45FF5D117AFF16" bold="true" box="[199,1321,150,177]" pageId="14" pageNumber="165">
Description based on
<typeStatus id="5481589AFFB22225FE4AFF5C1660FF17" box="[456,563,151,177]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
; variation of
<typeStatus id="5481589AFFB22225FD53FF5C1714FF17" box="[721,839,151,177]" pageId="14" pageNumber="165" type="paratype">paratypes</typeStatus>
in parentheses of (
<typeStatus id="5481589AFFB22225FBA4FF5D10F1FF16" box="[1062,1186,150,176]" pageId="14" pageNumber="165" type="paratype">paratypes</typeStatus>
; non-type).
</emphasis>
Complete
<typeStatus id="5481589AFFB22225FF15FF7114AAFF72" box="[151,249,186,212]" pageId="14" pageNumber="165" type="holotype">holotype</typeStatus>
with 121 chaetigers, L10
<quantity id="4CC24BDDFFB22225FD98FF70160FFF73" box="[538,604,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="165" unit="mm" value="9.0">9 mm</quantity>
, total length
<quantity id="4CC24BDDFFB22225FD69FF71176FFF73" box="[747,828,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="14" pageNumber="165" unit="mm" value="66.0">66 mm</quantity>
, maximum W10 without parapodium
<quantity id="4CC24BDDFFB22225FB60FF701175FF73" box="[1250,1318,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="165" unit="mm" value="3.0">3 mm</quantity>
.
<typeStatus id="5481589AFFB22225FAADFF7011CFFF73" box="[1327,1436,187,213]" pageId="14" pageNumber="165" type="paratype">Paratypes</typeStatus>
14 incomplete specimens with 19108 chaetigers and 3 complete specimens with 100107 chaetigers, total length of complete specimens
<quantity id="4CC24BDDFFB22225FE27FEC91671FEBB" box="[421,546,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="14" pageNumber="165" unit="mm" value="66.5" valueMax="68.0" valueMin="65.0">6568 mm</quantity>
, L10
<quantity id="4CC24BDDFFB22225FDE4FEC91686FEBB" box="[614,725,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="14" pageNumber="165" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
, W10 without parapodium
<quantity id="4CC24BDDFFB22225FB8CFEC91023FEBB" box="[1038,1136,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="14" pageNumber="165" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
. Non-types 48 incomplete specimens with 19108 chaetigers and 12 complete specimens with 82122 chaetigers, total length of the complete specimens
<quantity id="4CC24BDDFFB22225FE91FE8115CFFEC3" box="[275,412,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="6.95" metricValueMax="10.0" metricValueMin="3.9" pageId="14" pageNumber="165" unit="mm" value="69.5" valueMax="100.0" valueMin="39.0">39100 mm</quantity>
, L10
<quantity id="4CC24BDDFFB22225FE5EFE811622FEC3" box="[476,625,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.5" metricValueMin="4.5" pageId="14" pageNumber="165" unit="mm" value="8.0" valueMax="11.5" valueMin="4.5">4.511.5 mm</quantity>
, W10 without parapodium
<quantity id="4CC24BDDFFB22225FC21FE811053FEC3" box="[931,1024,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="14" pageNumber="165" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
in chaetiger 520.
</paragraph>
<paragraph id="8B85E638FFB22225FF45FEA51067FD25" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
Prostomium bilobed with deep median groove, visible dorsally and ventrally, dorsally flat, rounded anteriorly, less deep and distinctly shorter than peristomium (
<figureCitation id="1301FABDFFB22225FD4FFE59173AFE0A" box="[717,873,401,428]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5A, B, D</figureCitation>
); dorsal and ventral buccal lips present, appearing quadrilobed (
<figureCitation id="1301FABDFFB22225FEACFE7E15DBFE76" box="[302,392,437,464]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5C</figureCitation>
). Prostomial appendages evenly spaced in semi-circle, palps anteriorly and slightly laterally to lateral antennae. Palpophores and ceratophores short and ringshaped, without articulations. Ceratostyles and palpostyles tapering to thin tips with regular moniliform articulations. Median antennae longest and palps shortest (
<figureCitation id="1301FABDFFB22225FF1DFDEA14AAFD9A" box="[159,249,545,572]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5A</figureCitation>
); palps to posterior border first peristomial ring, with 9 (59; 68) articulations; lateral antennae to second chaetiger, with 9 (714; 615) articulations; median antenna to fourth (14; 16) chaetiger, with 16 (713; 818) articulations. Pair of eyes present, posterior to palps and lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFB22225FF45FD461535FC90" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
Peristomium cylindrical, laterally longer than medially; dorsal anterior margin covering base of median and lateral antennae; conspicuously inflated ventro-lateral lips. First peristomial ring about ¾ of total length, separation between rings visible dorsally and ventrally, not visible laterally (
<figureCitation id="1301FABDFFB22225FCE8FD1F1057FD49" box="[874,1028,724,751]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5A, B, D</figureCitation>
). Peristomial cirri thin, tapering with 8 (37; 36) regular cylindrical articulations, reaching anterior end of prostomium (anterior end of first peristomial ring) (
<figureCitation id="1301FABDFFB22225FF5FFCD71509FC90" box="[221,346,796,823]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5A, D</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB22225FF45FC8B10ABFC4C" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
Maxillary formula (
<typeStatus id="5481589AFFB22225FE21FC8B1643FCFC" box="[419,528,832,858]" pageId="14" pageNumber="165" type="paratype">paratypes</typeStatus>
and non-types): 1+1; (46)+(46); (68)+0; (47)+(810); 1+1; 1+1. MxI pincer like, not extended falcal arch, straight rectangular outer edge, basal inner edge with curvature where most posterior region of MxII fits. MxIII with part of the distal arch posterior to MxIV and partially ventral to MxII; MxV wide, with a prominent pointed tooth; MxVI present, drop-shaped. Mandibles well calcified, wing shaped. MxIV left attachment lamella tapering on left portion, right attachment lamella semi-circular shaped (
<figureCitation id="1301FABDFFB22225FB10FC0410B8FC4C" box="[1170,1259,975,1002]" captionStart="FIGURE 5" captionStartId="12.[152,255,1562,1586]" captionTargetBox="[151,1436,181,1538]" captionTargetId="figure-23@12.[151,1436,181,1538]" captionTargetPageId="12" captionText="FIGURE 5. Leodice marcusi. A, B, D MNRJP003258. C, MNRJP003261. A—anterior end, dorsal view; B—anterior end, ventral view; Cmaxillae, dorsal view; D—anterior end, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13219192" httpUri="https://zenodo.org/record/13219192/files/figure.png" pageId="14" pageNumber="165">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB22225FF45FC38152BFB3B" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">Branchiae pectinate present from chaetiger 4110 (490; 465). Branchiae present in more than 65% of total number of chaetigers. First branchia with single short branchial filament; 35 branchial filaments from chaetiger 642, branchial filaments reduce in length and number from chaetigers 43110. Branchial filaments and central axis of branchiae shorter than notopodial cirri. Branchiae decrease in length and number of filaments in median and posterior parapodia.</paragraph>
<paragraph id="8B85E638FFB22225FF45FB6C1489FAAE" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">Anterior chaetal lobes rounded, aciculae emerging dorsally; median and posterior chaetal lobe truncate, aciculae emerging dorsally. Prechaetal lobe low transverse fold along the body. Postchaetal lobe rounded, lower than chaetal lobes.</paragraph>
<paragraph id="8B85E638FFB22225FF45FAD91678FA1A" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
Anterior notopodial cirri slightly inflated basally, tapering to thin tips with up to five articulations. Median and posterior notopodial cirri slightly inflated basally, tapering to thin tips with up to two articulations, shorter than anterior. Ventral cirri basally inflated from chaetiger 6 (47) to 41 (3845; 2228) (
<figureCitation id="1301FABDFFB22225FBC8FA9110F1FAD2" box="[1098,1186,1370,1396]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6B</figureCitation>
), decreasing in length along body; anterior longer than chaetal lobe, inflated bases oval, tapering to digitiform tips; median and posterior lightly tapering with digitiform tips.
</paragraph>
<paragraph id="8B85E638FFB22225FF45FA0D1550F814" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">
Limbate chaetae slender, elongated and distally tapering, longer than other chaetae. Pectinate chaetae with one marginal tooth longer, anterior with 1215 teeth (
<figureCitation id="1301FABDFFB22225FD44FA221772F9A5" box="[710,801,1513,1539]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6H</figureCitation>
), posterior with 15 teeth, anterior slightly narrower than posterior. Compound falcigers chaetae bidentate, anterior parapodia with article tapering from proximal to distal end with narrow guards, marginally serrated (
<figureCitation id="1301FABDFFB22225FD15F9FA16A2F9ED" box="[663,753,1585,1611]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6A</figureCitation>
); article and guards increase towards posterior end; posterior guards slightly inflated, marginally serrated; laterally directed proximal tooth and prominent larger directed laterally distal tooth in anterior and median parapodia (
<figureCitation id="1301FABDFFB22225FD1EF9B216A5F935" box="[668,758,1657,1683]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6D</figureCitation>
); proximal tooth and distal tooth similar in length, triangular and laterally directed in posterior parapodia (
<figureCitation id="1301FABDFFB22225FD21F95616ADF911" box="[675,766,1693,1719]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6E</figureCitation>
). Aciculae paired along the whole body; anterior aciculae yellow, paired, straight, smoothly tapering to round tips (
<figureCitation id="1301FABDFFB22225FC9CF90A1724F97D" box="[798,887,1729,1755]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6B</figureCitation>
); median aciculae brown, straight, with rounded tips (
<figureCitation id="1301FABDFFB22225FF53F92E1578F959" box="[209,299,1765,1791]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6C</figureCitation>
); posterior aciculae paired, dark brown, smooth, tapering, with curved tips; last parapodia with one acicula larger than other, tapering to thin tips (
<figureCitation id="1301FABDFFB22225FD25F8C316ABF884" box="[679,760,1800,1826]" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6I</figureCitation>
). Subacicular hooks brown, tapering to narrow necks, large heads, relative to neck, bidentate, proximal tooth larger and more robust than distal tooth, tapering, directed laterally; present from chaetigers 26 (2224; 1925), median and posterior hooks with proximal and distal teeth tapering to blunt tips; distal tooth narrow, almost erect, tapering directed distally; guards partially covering proximal tooth (
<figureCitation id="1301FABDFFB22225FAEDF8BF14A6F814" captionStart="FIGURE 6" captionStartId="13.[152,255,1863,1887]" captionTargetBox="[151,1436,343,1840]" captionTargetId="figure-74@13.[151,1436,343,1840]" captionTargetPageId="13" captionText="FIGURE 6. Leodice marcusi. A, D, E, G, I, J, POLY-UFPB1991. B, C, F, H, MNRJP003261. A—parapodium 5, compound falcigers; B—parapodium 3, notopodial cirri, compound falcigers chaetae, aciculae and ventral cirri; C—parapodium 44, ventral cirri, compound falcigers chaetae, pectinate chaetae and aciculae; D—parapodium 43, compound falcigers chaetae; E— parapodium 110, compound falciger chaetae; F—parapodium 44, pectinate chaetae; G—parapodium 110, subacicular hooks; H—parapodium 3, pectinate chaetae; I—parapodium 110, aciculae; J—parapodium 67, subacicular hook." figureDoi="http://doi.org/10.5281/zenodo.13219194" httpUri="https://zenodo.org/record/13219194/files/figure.png" pageId="14" pageNumber="165">Fig. 6C, G, J</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB22225FF45F877157AF85C" blockId="14.[151,1437,150,2042]" pageId="14" pageNumber="165">Pygidium with two pairs of pygidial cirri basally inflated, tapering to thin tips, with up to six cylindrical regular articulations.</paragraph>
<paragraph id="8B85E638FFB32224FF45FF5D1513FE2E" blockId="15.[151,1437,150,861]" pageId="15" pageNumber="166">
<emphasis id="B94E3A2AFFB32224FF45FF5D1519FF16" bold="true" box="[199,330,150,176]" pageId="15" pageNumber="166">Variations.</emphasis>
<materialsCitation id="3B52EC65FFB32224FED6FF5D156FFE2E" collectionCode="MNRJP" pageId="15" pageNumber="166" specimenCode="MNRJP003260" specimenCount="2" typeStatus="paratype">
The specimens observed varied in number of chaetigers (100121) and length (
<quantity id="4CC24BDDFFB32224FB71FF5D1121FF16" box="[1267,1394,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="15" pageNumber="166" unit="mm" value="66.5" valueMax="68.0" valueMin="65.0">6568 mm</quantity>
) of complete individuals, smallest specimen (
<typeStatus id="5481589AFFB32224FDE6FF711695FF72" box="[612,710,186,212]" pageId="15" pageNumber="166" type="paratype">paratype</typeStatus>
<specimenCode id="DB9C4E43FFB32224FD4EFF71172FFF72" box="[716,892,186,212]" collectionCode="MNRJP" pageId="15" pageNumber="166">MNRJP003260</specimenCode>
) incomplete with 52 chaetiger, measuring
<quantity id="4CC24BDDFFB32224FACDFF7111CEFF73" box="[1359,1437,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="166" unit="mm" value="20.0">20 mm</quantity>
in length and having just one branchial filament. Number of articulations on ceratostyles and notopodial cirri (25) also varied among specimens.
<typeStatus id="5481589AFFB32224FE6CFEC81687FEBA" box="[494,724,258,285]" isOtherTypeStatus="true" pageId="15" pageNumber="166">Non-type specimens</typeStatus>
have more wrinkled and rectangular articulations in prostomial appendages than in type specimens, moniliform articulations becoming drop-shaped distally. Number of teeth in MxII-IV. Starting point of branchiae, inflated base of ventral cirri, and of subacicular hooks are also variable among our specimens
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB32224FF45FE59114BFE52" pageId="15" pageNumber="166" type="distribution">
<paragraph id="8B85E638FFB32224FF45FE59114BFE52" blockId="15.[151,1437,150,861]" pageId="15" pageNumber="166">
<emphasis id="B94E3A2AFFB32224FF45FE591533FE0B" bold="true" box="[199,352,402,429]" pageId="15" pageNumber="166">Distribution.</emphasis>
<taxonomicName id="4C3A9DBBFFB32224FEE8FE591676FE0A" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[362,549,402,428]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="marcusi">
<emphasis id="B94E3A2AFFB32224FEE8FE591676FE0A" box="[362,549,402,428]" italics="true" pageId="15" pageNumber="166">Leodice marcusi</emphasis>
</taxonomicName>
was described from the states of
<collectingRegion id="49FE28DAFFB32224FC29FE591001FE0A" box="[939,1106,402,428]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="166">Rio de Janeiro</collectingRegion>
and
<collectingRegion id="49FE28DAFFB32224FB0FFE591161FE0A" box="[1165,1330,402,428]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="166">Espírito Santo</collectingRegion>
, between 20 and
<quantity id="4CC24BDDFFB32224FF72FE7D157FFE77" box="[240,300,438,465]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="166" unit="m" value="82.0">82 m</quantity>
in depth, in calcareous algae and medium sand bottoms (
<bibRefCitation id="EFAB9BC9FFB32224FC4CFE7D10CDFE76" author="Zanol, J. &amp; Paiva, P. C. &amp; Attolini, F. S." box="[974,1182,438,464]" pageId="15" pageNumber="166" pagination="449 - 463" refId="ref14731" refString="Zanol, J., Paiva, P. C. &amp; Attolini, F. S. (2000) Eunice and Palola (Eunicidae: Polychaeta) from the eastern Brazilian Coast (13 ° 00 ' - 22 ° 30 ' S). Bulletin of Marine Science, 67 (1), 449 - 463." type="journal article" year="2000">
Zanol
<emphasis id="B94E3A2AFFB32224FB99FE7C1004FE76" box="[1051,1111,438,464]" italics="true" pageId="15" pageNumber="166">et al.</emphasis>
2000
</bibRefCitation>
). Here we expand the geographic distribution to northeastern
<collectingCountry id="F32DA6A8FFB32224FDD3FE1116C6FE52" box="[593,661,474,500]" name="Brazil" pageId="15" pageNumber="166">Brazil</collectingCountry>
and the shallowest record of the species to
<quantity id="4CC24BDDFFB32224FBFFFE1110E5FE53" box="[1149,1206,474,501]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="166" unit="m" value="12.0">12 m</quantity>
(
<figureCitation id="1301FABDFFB32224FB47FE111155FE52" box="[1221,1286,474,500]" captionStart="FIGURE 1" captionStartId="3.[152,255,1913,1937]" captionTargetBox="[151,1429,195,1889]" captionTargetId="figure-19@3.[151,1436,181,1889]" captionTargetPageId="3" captionText="FIGURE 1. General map of the stations of the Marine Algae project along the Continental Shelf of the State of Paraíba where Leodice species have been recorded." figureDoi="http://doi.org/10.5281/zenodo.13219176" httpUri="https://zenodo.org/record/13219176/files/figure.png" pageId="15" pageNumber="166">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB32224FF45FE3515B1FCFA" pageId="15" pageNumber="166" type="discussion">
<paragraph id="8B85E638FFB32224FF45FE3517A1FD0E" blockId="15.[151,1437,150,861]" pageId="15" pageNumber="166">
<emphasis id="B94E3A2AFFB32224FF45FE351568FDBE" bold="true" box="[199,315,510,536]" pageId="15" pageNumber="166">Remarks.</emphasis>
Our redescription of
<taxonomicName id="4C3A9DBBFFB32224FDA4FE3416C8FDBE" baseAuthorityName="Zanol, Paiva &amp; Attolini" baseAuthorityYear="2000" box="[550,667,511,536]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="166" phylum="Annelida" rank="species" species="marcusi">
<emphasis id="B94E3A2AFFB32224FDA4FE3416C8FDBE" box="[550,667,511,536]" italics="true" pageId="15" pageNumber="166">L. marcusi</emphasis>
</taxonomicName>
includes information and illustrations missing in original description, such as anterior end ventral view; anterior end lateral view; maxillary apparatus dorsal view. The maxillary apparatus of the
<typeStatus id="5481589AFFB32224FED1FD8D15E6FDC6" box="[339,437,582,608]" pageId="15" pageNumber="166" type="holotype">holotype</typeStatus>
was not dissected. Also, we have observed features in type-material that vary from with the original description. Shape of branchiae is pectinate, and not palmate, as originally described. Color of aciculae and subacicular hooks are dark brown, and not black as originally described.
</paragraph>
<paragraph id="8B85E638FFB32224FF45FD7915B1FCFA" blockId="15.[151,1437,150,861]" pageId="15" pageNumber="166">
The specimens from the State of
<collectingRegion id="49FE28DAFFB32224FDC8FD7916CDFD6A" box="[586,670,690,716]" country="Brazil" name="Paraiba" pageId="15" pageNumber="166">Paraíba</collectingRegion>
agree well with
<typeStatus id="5481589AFFB32224FCE1FD7817C0FD6B" box="[867,915,691,717]" pageId="15" pageNumber="166">type</typeStatus>
material. However, they contrast in shape of antennae and palps articulations, which are more wrinkled in
<typeStatus id="5481589AFFB32224FCE3FD1C107EFD56" box="[865,1069,726,753]" isOtherTypeStatus="true" pageId="15" pageNumber="166">non-type material</typeStatus>
. One potential factor that could account for these differences is the fixation and preservation methods;
<typeStatus id="5481589AFFB32224FC57FD3010EBFCB2" box="[981,1208,762,789]" isOtherTypeStatus="true" pageId="15" pageNumber="166">non-type specimens</typeStatus>
were preserved for many years in formaldehyde solution used to fix the specimens whereas
<typeStatus id="5481589AFFB32224FC4BFCD417AAFC9F" box="[969,1017,799,825]" pageId="15" pageNumber="166">type</typeStatus>
material was washed and transferred to alcohol soon after fixation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="40829D9E537CE129AB865F277281C220">Taxonomy of Leodice Lamarck, 1818 (Annelida, Eunicidae) from Northeast Brazil (Tropical Southwest Atlantic Ocean) with description of a new species</mods:title>
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<heading id="D0CD5154FFB8222FFF15F9FC1642F9F7" bold="true" box="[151,529,1591,1618]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<emphasis id="B94E3A2AFFB8222FFF15F9FC1642F9F7" bold="true" box="[151,529,1591,1618]" pageId="4" pageNumber="155">
Genus
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<emphasis id="B94E3A2AFFB8222FFF6CF9FC151FF9F4" bold="true" box="[238,332,1591,1618]" italics="true" pageId="4" pageNumber="155">Leodice</emphasis>
<bibRefCitation id="EFAB9BC9FFB8222FFED6F9FC1642F9F7" author="Lamarck, J. B. de" box="[340,529,1591,1617]" pageId="4" pageNumber="155" refId="ref14137" refString="Lamarck, J. B. de (1818) Histoire naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une Int. Vol. 5. Deterville Libraire &amp; Verdiere Libraire, Paris, 612 pp." type="book" year="1818">Lamarck, 1818</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8B85E638FFB8222FFF15F9B41713F93F" blockId="4.[151,1437,1663,2014]" box="[151,832,1663,1689]" pageId="4" pageNumber="155">
<emphasis id="B94E3A2AFFB8222FFF15F9B41561F93F" bold="true" box="[151,306,1663,1689]" pageId="4" pageNumber="155">
<typeStatus id="5481589AFFB8222FFF15F9B41481F93F" box="[151,210,1663,1689]" pageId="4" pageNumber="155">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3A9DBBFFB8222FFEBBF9B4163FF93F" ID-CoL="3T6C9" authority="Savigny" authorityName="Savigny" box="[313,620,1663,1689]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="155" phylum="Annelida" rank="species" species="antennata">
<emphasis id="B94E3A2AFFB8222FFEBBF9B4165AF93F" box="[313,521,1663,1689]" italics="true" pageId="4" pageNumber="155">Leodice antennata</emphasis>
Savigny
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in
<bibRefCitation id="EFAB9BC9FFB8222FFD0DF9B4176FF93F" author="Lamarck, J. B. de" box="[655,828,1663,1689]" pageId="4" pageNumber="155" refId="ref14137" refString="Lamarck, J. B. de (1818) Histoire naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une Int. Vol. 5. Deterville Libraire &amp; Verdiere Libraire, Paris, 612 pp." type="book" year="1818">Lamarck, 1818</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB8222FFF45F96810E9F8D7" pageId="4" pageNumber="155" type="diagnosis">
<paragraph id="8B85E638FFB8222FFF45F96810E9F8D7" blockId="4.[151,1437,1663,2014]" pageId="4" pageNumber="155">
<emphasis id="B94E3A2AFFB8222FFF45F96815D1F91B" bold="true" box="[199,386,1699,1725]" pageId="4" pageNumber="155">
Diagnosis
<emphasis id="B94E3A2AFFB8222FFEC3F96815D1F91B" bold="true" box="[321,386,1699,1725]" italics="true" pageId="4" pageNumber="155">sensu</emphasis>
</emphasis>
<bibRefCitation id="EFAB9BC9FFB8222FFE0BF96816F3F91B" author="Zanol, J. &amp; Budaeva, N." box="[393,672,1699,1725]" pageId="4" pageNumber="155" refId="ref14468" refString="Zanol, J. &amp; Budaeva, N. (2021) Eunicidae. In: Purschke, G., Westheide, W. &amp; Boggemann, M. (Eds.), Handbook of Zoology Annelida. Vol. 3. Sedentaria III. Errantia I. De Gruyter, Berlin and Boston, pp. 414 - 452. https: // doi. org / 10.1515 / 9783110291704" type="book" year="2021">Zanol &amp; Budaeva (2021)</bibRefCitation>
. Median, lateral antennae and palps present with regular or irregular articulations. Prostomium anteriorly steep truncate or round. Peristomial cirri present. Maxillae with four or five paired plates and one unpaired plate. Mandibles flat. Limbate chaetae, thin shafted pectinate chaetae, compound bidentate or tridentate falcigers, aciculae, and subacicular hooks present. Aciculae light or dark. Dark aciculae vary in color along body; anterior usually lightest but maintain similar color. Subacicular hooks light or dark, bidentate or tridentate. Lateral black dots between parapodia present or absent (
<emphasis id="B94E3A2AFFB8222FFC27F89317B0F8D7" box="[933,995,1880,1905]" italics="true" pageId="4" pageNumber="155">sensu</emphasis>
<bibRefCitation id="EFAB9BC9FFB8222FFC68F89C10FDF8D7" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[1002,1198,1879,1905]" pageId="4" pageNumber="155" pagination="79 - 100" refId="ref14618" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="B94E3A2AFFB8222FFBB0F8931039F8D7" box="[1074,1130,1879,1905]" italics="true" pageId="4" pageNumber="155">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB8222EFF45F8B01612FEBB" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="discussion">
<paragraph id="8B85E638FFB8222EFF45F8B01612FEBB" blockId="4.[151,1437,1663,2014]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
<emphasis id="B94E3A2AFFB8222FFF45F8B01568F833" bold="true" box="[199,315,1915,1941]" pageId="4" pageNumber="155">Remarks.</emphasis>
The genus
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<emphasis id="B94E3A2AFFB8222FFE3DF8B0164BF833" box="[447,536,1915,1941]" italics="true" pageId="4" pageNumber="155">Leodice</emphasis>
<bibRefCitation id="EFAB9BC9FFB8222FFD9DF8B0169AF833" author="Lamarck, J. B. de" box="[543,713,1915,1941]" pageId="4" pageNumber="155" refId="ref14137" refString="Lamarck, J. B. de (1818) Histoire naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une Int. Vol. 5. Deterville Libraire &amp; Verdiere Libraire, Paris, 612 pp." type="book" year="1818">Lamarck, 1818</bibRefCitation>
</taxonomicName>
was revalidated by
<bibRefCitation id="EFAB9BC9FFB8222FFC2EF8B01023F833" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[940,1136,1915,1942]" pageId="4" pageNumber="155" pagination="79 - 100" refId="ref14618" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="B94E3A2AFFB8222FFC77F8B71075F833" box="[1013,1062,1915,1941]" italics="true" pageId="4" pageNumber="155">et al</emphasis>
. 2014
</bibRefCitation>
, and amended to represent a clade that includes the
<typeStatus id="5481589AFFB8222FFE39F86B15B8F81C" box="[443,491,1952,1978]" pageId="4" pageNumber="155">type</typeStatus>
species,
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<emphasis id="B94E3A2AFFB8222FFDDAF86B16BBF81F" box="[600,744,1952,1977]" italics="true" pageId="4" pageNumber="155">L. antennata</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3A9DBBFFB8222FFD78F8541700F81F" authorityName="Lamarck" authorityYear="1818" box="[762,851,1951,1977]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="155" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB8222FFD78F8541700F81F" box="[762,851,1951,1977]" italics="true" pageId="4" pageNumber="155">Leodice</emphasis>
</taxonomicName>
includes species with five prostomial appendages, peristomial cirri, subacicular hooks and at least one of the following exclusive features: 1. regularly articulated prostomial appendages, 2. light bidentate or tridentate subacicular hooks or, 3. lateral black dots between parapodia. It shares with
<taxonomicName id="4C3A9DBBFFB9222EFEBFFF701678FF73" authority="Cuvier, 1817" authorityName="Cuvier" authorityYear="1817" box="[317,555,187,213]" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="156" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB9222EFEBFFF7015D8FF72" box="[317,395,187,212]" italics="true" pageId="5" pageNumber="156">Eunice</emphasis>
Cuvier, 1817
</taxonomicName>
,
<taxonomicName id="4C3A9DBBFFB9222EFDBBFF70179AFF73" authority="Wesenberg-Lund, 1949" authorityName="Wesenberg-Lund" authorityYear="1949" box="[569,969,187,213]" class="Polychaeta" family="Eunicidae" genus="Euniphysa" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="156" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB9222EFDBBFF7016FCFF73" box="[569,687,187,213]" italics="true" pageId="5" pageNumber="156">Euniphysa</emphasis>
Wesenberg-Lund, 1949
</taxonomicName>
,
<taxonomicName id="4C3A9DBBFFB9222EFC59FF70156BFF5F" authority="Carrera-Parra &amp; Salazar-Vallejo, 1998" authorityName="Carrera-Parra &amp; Salazar-Vallejo" authorityYear="1998" class="Polychaeta" family="Eunicidae" genus="Fauchaldius" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="156" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB9222EFC59FF701034FF73" box="[987,1127,187,213]" italics="true" pageId="5" pageNumber="156">Fauchaldius</emphasis>
<bibRefCitation id="EFAB9BC9FFB9222EFBF4FF70156BFF5F" author="Carrera-Parra, L. F. &amp; Salazar-Vallejo, S. I." pageId="5" pageNumber="156" pagination="145 - 182" refId="ref13473" refString="Carrera-Parra, L. F. &amp; Salazar-Vallejo, S. I. (1998) A new genus and 12 new species of Eunicidae (Polychaeta) from the Caribbean Sea. Journal of the Marine Biological Association of the United Kingdom, 78, 145 - 182. https: // doi. org / 10.1017 / S 0025315400040005" type="journal article" year="1998">Carrera-Parra &amp; Salazar-Vallejo, 1998</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3A9DBBFFB9222EFEFDFF1416C6FF5F" authority="Kinberg, 1865" authorityName="Kinberg" authorityYear="1865" box="[383,661,222,249]" class="Polychaeta" family="Eunicidae" genus="Nicidion" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="156" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB9222EFEFDFF14158CFF5F" box="[383,479,223,249]" italics="true" pageId="5" pageNumber="156">Nicidion</emphasis>
<bibRefCitation id="EFAB9BC9FFB9222EFE69FF1416C6FF5F" author="Kinberg, J. G. H." box="[491,661,222,249]" pageId="5" pageNumber="156" pagination="559 - 574" refId="ref14107" refString="Kinberg, J. G. H. (1865) Annulata nova. Ofversigt af Koniglich Vetenskaps - Akademiens forhandlingar, Stockholm, 21, 559 - 574." type="journal article" year="1865">Kinberg, 1865</bibRefCitation>
</taxonomicName>
, five prostomial appendages, a pair of peristomial cirri, and dark subacicular hooks (
<bibRefCitation id="EFAB9BC9FFB9222EFEEDFEC81666FEBB" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[367,565,259,285]" pageId="5" pageNumber="156" pagination="79 - 100" refId="ref14618" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="B94E3A2AFFB9222EFE3BFEC815A1FEBB" box="[441,498,259,285]" italics="true" pageId="5" pageNumber="156">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
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</mods:mods>
<treatment id="0393572EFFB92222FF15FEA515A8FA67" ID-DOI="http://doi.org/10.5281/zenodo.13219170" ID-Zenodo-Dep="13219170" LSID="urn:lsid:plazi:treatment:0393572EFFB92222FF15FEA515A8FA67" httpUri="http://treatment.plazi.org/id/0393572EFFB92222FF15FEA515A8FA67" lastPageId="9" lastPageNumber="160" pageId="5" pageNumber="156">
<subSubSection id="C320B5B3FFB9222EFF15FEA51659FE2E" box="[151,522,366,393]" pageId="5" pageNumber="156" type="nomenclature">
<paragraph id="8B85E638FFB9222EFF15FEA51659FE2E" blockId="5.[151,522,366,429]" box="[151,522,366,393]" pageId="5" pageNumber="156">
<heading id="D0CD5154FFB9222EFF15FEA51659FE2E" bold="true" box="[151,522,366,393]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
<emphasis id="B94E3A2AFFB9222EFF15FEA51659FE2E" bold="true" box="[151,522,366,393]" pageId="5" pageNumber="156">
<emphasis id="B94E3A2AFFB9222EFF15FEA51520FE2F" bold="true" box="[151,371,366,393]" italics="true" pageId="5" pageNumber="156">
<taxonomicName id="4C3A9DBBFFB9222EFF15FEA5153EFE2F" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[151,365,366,393]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="156" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">Leodice ivanildae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A27D8751FFB9222EFEF8FEA41659FE2E" box="[378,522,366,393]" pageId="5" pageNumber="156" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB9222EFF15FE58153FFE0B" box="[151,364,403,429]" pageId="5" pageNumber="156" type="description">
<paragraph id="8B85E638FFB9222EFF15FE58153FFE0B" blockId="5.[151,522,366,429]" box="[151,364,403,429]" pageId="5" pageNumber="156">
(
<figureCitation id="1301FABDFFB9222EFF1DFE5814BBFE0B" box="[159,232,403,429]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Figs 2</figureCitation>
,
<figureCitation id="1301FABDFFB9222EFF76FE581550FE0B" box="[244,259,403,429]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="5" pageNumber="156">3</figureCitation>
,
<tableCitation id="C6B8D383FFB9222EFE8DFE581530FE0B" box="[271,355,403,429]" captionStart="TABLE 2" captionStartId="8.[151,243,523,549]" captionText="TABLE 2. Comparison of morphological features among Leodice ivanildae sp. nov. and most similar species." pageId="5" pageNumber="156">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB9222EFF15FE11159CFC27" pageId="5" pageNumber="156" type="materials_examined">
<paragraph id="8B85E638FFB9222EFF15FE11159CFC27" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
<emphasis id="B94E3A2AFFB9222EFF15FE111672FE52" bold="true" box="[151,545,474,500]" pageId="5" pageNumber="156">
Material examined.
<typeStatus id="5481589AFFB9222EFE00FE111672FE52" box="[386,545,474,500]" pageId="5" pageNumber="156" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, complete specimen, station 89 (MNRJP-007863).
<materialsCitation id="3B52EC65FFB9222EFBD2FE111598FC27" pageId="5" pageNumber="156" specimenCode="ZUEC-POL 25422, ZUEC-POL 25423" specimenCount="21" typeStatus="paratype">
<emphasis id="B94E3A2AFFB9222EFBD2FE1110A7FE52" bold="true" box="[1104,1268,474,500]" pageId="5" pageNumber="156">
<typeStatus id="5481589AFFB9222EFBD2FE1110A7FE52" box="[1104,1268,474,500]" pageId="5" pageNumber="156" type="paratype">PARATYPES</typeStatus>
</emphasis>
,
<specimenCount id="9D3C2DB1FFB9222EFA82FE1011CBFE53" box="[1280,1432,475,501]" count="21" pageId="5" pageNumber="156" type="generic">21 specimens</specimenCount>
; one complete specimen, station 2 (MNRJP-007865); one incomplete specimen, station 3 (MNRJP-007864); one complete specimen, station 5 (MNRJP-007866); one incomplete specimen, station 11 (MNRJP-007867); one incomplete specimen, station 33 (MNRJP-007868); one incomplete specimen, station 39 (MNRJP-007869); one complete specimen, station 46 (POLY-UFPB-1961); one complete specimen, station 47 (POLY-UFPB-1958); one incomplete specimen, station 48 (POLY-UFPB-1959); one incomplete specimen, station 49 (MNRJP-007871); one complete specimen, station 53 (MZUSP-6104); one complete specimen, station 54 (POLY-UFPB-1957); one incomplete specimen, station 55 (POLY-UFPB-1954); one incomplete specimen, station 57 (MNRJP-007870); one incomplete specimen, station 62 (POLY-UFPB-1962); one incomplete specimen, station 80 (POLY-UFPB-2081); one incomplete specimen, station 45 (POLY-UFPB-1960); one incomplete specimen, station 88 (POLY-UFPB-1968); one incomplete and three complete specimens, station 93 (POLY-UFPB-1972, MZUSP-6103,
<specimenCode id="DB9C4E43FFB9222EFA94FC8814B3FC27" collectionCode="ZUEC-POL" pageId="5" pageNumber="156">ZUEC-POL 25422</specimenCode>
,
<specimenCode id="DB9C4E43FFB9222EFF69FCAC1590FC27" box="[235,451,870,897]" collectionCode="ZUEC-POL" pageId="5" pageNumber="156">ZUEC-POL 25423</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB92222FF45FC4116F5FC9F" lastPageId="9" lastPageNumber="160" pageId="5" pageNumber="156" type="description">
<paragraph id="8B85E638FFB9222EFF45FC41115CFB4F" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
<emphasis id="B94E3A2AFFB9222EFF45FC41150AFC02" bold="true" box="[199,345,906,932]" pageId="5" pageNumber="156">Description.</emphasis>
Complete
<typeStatus id="5481589AFFB9222EFE52FC401661FC03" box="[464,562,907,933]" pageId="5" pageNumber="156" type="holotype">holotype</typeStatus>
with 174 chaetigers, L10
<quantity id="4CC24BDDFFB9222EFCCFFC4017DDFC03" box="[845,910,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="156" unit="mm" value="6.0">6 mm</quantity>
, total length
<quantity id="4CC24BDDFFB9222EFB98FC401024FC03" box="[1050,1143,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="156" unit="mm" value="100.0">100 mm</quantity>
, W10 without parapodium
<quantity id="4CC24BDDFFB9222EFF15FC6414B3FC6F" box="[151,224,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="156" unit="mm" value="2.0">2 mm</quantity>
.
<typeStatus id="5481589AFFB9222EFF6DFC64150FFC6F" box="[239,348,943,969]" pageId="5" pageNumber="156" type="paratype">Paratypes</typeStatus>
10 incomplete specimens with 67149 chaetigers and 11 complete specimens with 143159 chaetigers, total length of complete specimens
<quantity id="4CC24BDDFFB9222EFD15FC18174DFC4B" box="[663,798,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="10.4" metricValueMin="7.6" pageId="5" pageNumber="156" unit="mm" value="90.0" valueMax="104.0" valueMin="76.0">76104 mm</quantity>
, L10
<quantity id="4CC24BDDFFB9222EFCDBFC1817E4FC4B" box="[857,951,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="156" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
. W10 without parapodium
<quantity id="4CC24BDDFFB9222EFB62FC181169FC4B" box="[1248,1338,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="156" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
segment 520. Prostomium bilobed with deep median groove, rounded (truncated) anteriorly, flattened dorsally; shallower, shorter, and narrower than peristomium. Prostomial appendages evenly spaced in semicircle. Palpophores and ceratophores short and ringshaped, without articulations. Ceratostyles and palpostyles tapering to thin tips with regular, moniliform articulations (
<figureCitation id="1301FABDFFB9222EFD9EFBA8162BFBDB" box="[540,632,1123,1149]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2C</figureCitation>
). Median antenna about one third longer than lateral antennae, lateral antennae about twice as long as palps (
<figureCitation id="1301FABDFFB9222EFDC7FB4C168DFB07" box="[581,734,1159,1185]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2A, B, C</figureCitation>
); palps with 8 (69) articulations, reaching anterior peristomial ring (first chaetiger); lateral antennae with 19 (1122) articulations, reaching chaetiger 6 (36); median antenna with 28 (1535) articulations, reaching chaetiger 10 (513). Pair of eyes present, lateral to lateral antennae.
</paragraph>
<paragraph id="8B85E638FFB9222EFF45FB381018FADF" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
Peristomium cylindrical, laterally longer than medially (
<figureCitation id="1301FABDFFB9222EFCC4FB3817B5FAAB" box="[838,998,1267,1293]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2A, B, C</figureCitation>
); dorsal anterior margin covering base of median antenna; with conspicuously inflated ventrolateral lips. First peristomial ring about ¾ of total length, separation between rings visible dorsally and ventrally, not visible laterally (
<figureCitation id="1301FABDFFB9222EFC6FFAF01036FAF3" box="[1005,1125,1339,1365]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2B, C</figureCitation>
). Thin, tapering peristomial cirri with 7 (511) cylindrical articulations, reaching anterior margin of prostomium.
</paragraph>
<paragraph id="8B85E638FFB9222EFF45FA4810F7F9AF" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
Maxillary formula: 1+1; 6 (57)+6(58); 7(68)+0; 5(57)+9(811); 1+1; 1+1 (
<figureCitation id="1301FABDFFB9222EFBD6FA4810E3FA3B" box="[1108,1200,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2D</figureCitation>
). MxI with extended falcal arch, arched outer edge, rectangular; basal inner edge with curvature, where most posterior region of MxII fits; MxIII partially ventral to MxII, part distal arch with left MxIV; MxV with a prominent tooth; MxVI drop-shape. MxIV left attachment lamella tapering on left side, semi-circular in shape on right (
<figureCitation id="1301FABDFFB9222EFBBEFA2410C4F9AF" box="[1084,1175,1519,1545]" captionStart="FIGURE 2" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[167,1419,181,1919]" captionTargetId="figure-23@6.[167,1419,181,1919]" captionTargetPageId="6" captionText="FIGURE 2. Leodice ivanildae sp. nov. AD, MNRJP-007863. A—anterior end, dorsal view; B—anterior end, lateral view; C—anterior end, ventral view; Dmaxillae, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="5" pageNumber="156">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB9222EFF45F9D814ACF947" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
Branchiae pectinate with slender digitiform axis and filaments, present from chaetiger 4 (34) to 46 (3851) (
<figureCitation id="1301FABDFFB9222EFF1DF9FC14AAF9F7" box="[159,249,1591,1617]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="5" pageNumber="156">Fig. 3D</figureCitation>
); first branchiae with two short filaments; maximum number of filaments 7 (513) on chaetiger 15 (923); branchial filaments longer than central axis and both longer than notopodial cirri where best developed; branchial filaments reduce in length and number from middle of branchial distribution. Anterior and median parapodia with rounded chaetal lobes, aciculae emerging on dorsally; median parapodia narrower pointed, with aciculae emerging medially.
</paragraph>
<paragraph id="8B85E638FFB9222EFF45F92010EAF88F" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">Prechaetal lobes low transverse folds along the body. Anterior and median postchaetal lobes longer than chaetal lobes and rounded; posterior postchaetal lobes low transverse folds, shorter than chaetal lobes.</paragraph>
<paragraph id="8B85E638FFB9222EFF45F8F8116BF81F" blockId="5.[151,1437,474,1977]" pageId="5" pageNumber="156">
Notopodial cirri shorter towards posterior region. Anterior notopodial cirri basally inflated, tapering to thin tips with 5 (67) cylindrical articulations (
<figureCitation id="1301FABDFFB9222EFDC5F89C16F1F8D7" box="[583,674,1879,1905]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="5" pageNumber="156">Fig. 3G</figureCitation>
); median with 3 (4) articulations; posterior with 2 (3) articulations. Ventral cirri longer than chaetal lobe along the body, basally inflated from chaetiger 3 (4) to 73 (5573). Inflated base short, tapering to digitiform tips, slightly tapering from median chaetigers to posterior end (
<figureCitation id="1301FABDFFB9222EFB53F8541178F81F" box="[1233,1323,1951,1977]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="5" pageNumber="156">Fig. 3A</figureCitation>
).
</paragraph>
<caption id="DF45B6B0FFBA222DFF1AF85C1681F875" ID-DOI="http://doi.org/10.5281/zenodo.13219182" ID-Zenodo-Dep="13219182" httpUri="https://zenodo.org/record/13219182/files/figure.png" pageId="6" pageNumber="157" startId="6.[152,255,1943,1967]" targetBox="[167,1419,181,1919]" targetPageId="6" targetType="figure">
<paragraph id="8B85E638FFBA222DFF1AF85C1681F875" blockId="6.[151,1436,1943,2004]" pageId="6" pageNumber="157">
<emphasis id="B94E3A2AFFBA222DFF1AF85C1549F809" bold="true" box="[152,282,1943,1967]" pageId="6" pageNumber="157">FIGURE 2.</emphasis>
<taxonomicName id="4C3A9DBBFFBA222DFEA0F85C158BF809" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[290,472,1943,1967]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="157" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">
<emphasis id="B94E3A2AFFBA222DFEA0F85C158BF809" box="[290,472,1943,1967]" italics="true" pageId="6" pageNumber="157">Leodice ivanildae</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFBA222DFE62F8531660F816" bold="true" box="[480,563,1944,1968]" pageId="6" pageNumber="157">
<taxonomicNameLabel id="A27D8751FFBA222DFE62F8531660F816" box="[480,563,1944,1968]" pageId="6" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AD, MNRJP-007863.
<emphasis id="B94E3A2AFFBA222DFCAFF85C170AF809" bold="true" box="[813,857,1943,1967]" pageId="6" pageNumber="157">A—</emphasis>
anterior end, dorsal view;
<emphasis id="B94E3A2AFFBA222DFBEBF8531029F816" bold="true" box="[1129,1146,1944,1968]" pageId="6" pageNumber="157">B</emphasis>
—anterior end, lateral view;
<emphasis id="B94E3A2AFFBA222DFF15F87014FAF875" bold="true" box="[151,169,1979,2003]" pageId="6" pageNumber="157">C</emphasis>
—anterior end, ventral view;
<emphasis id="B94E3A2AFFBA222DFE56F87715B5F872" bold="true" box="[468,486,1980,2004]" pageId="6" pageNumber="157">D</emphasis>
maxillae, dorsal view.
</paragraph>
</caption>
<caption id="DF45B6B0FFBB222CFF1AF943168EF8F2" ID-DOI="http://doi.org/10.5281/zenodo.13219184" ID-Zenodo-Dep="13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="7" pageNumber="158" startId="7.[152,255,1672,1696]" targetBox="[151,1436,181,1648]" targetPageId="7" targetType="figure">
<paragraph id="8B85E638FFBB222CFF1AF943168EF8F2" blockId="7.[151,1437,1672,1876]" pageId="7" pageNumber="158">
<emphasis id="B94E3A2AFFBB222CFF1AF9431545F906" bold="true" box="[152,278,1672,1696]" pageId="7" pageNumber="158">FIGURE 3.</emphasis>
<taxonomicName id="4C3A9DBBFFBB222CFE99F943159DF906" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[283,462,1672,1696]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="7" pageNumber="158" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">
<emphasis id="B94E3A2AFFBB222CFE99F943159DF906" box="[283,462,1672,1696]" italics="true" pageId="7" pageNumber="158">Leodice ivanildae</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFBB222CFE50F9431672F906" bold="true" box="[466,545,1672,1696]" pageId="7" pageNumber="158">
<taxonomicNameLabel id="A27D8751FFBB222CFE50F9431672F906" box="[466,545,1672,1696]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, AD, MNRJP-007863. FG MNRJP-007866.
<emphasis id="B94E3A2AFFBB222CFC78F9431170F906" bold="true" box="[1018,1315,1672,1696]" pageId="7" pageNumber="158">Parapodia and chaetae. A—</emphasis>
parapodium 09, aciculae, compound falcigers and ventral cirri;
<emphasis id="B94E3A2AFFBB222CFD34F9671694F962" bold="true" box="[694,711,1708,1732]" pageId="7" pageNumber="158">B</emphasis>
—parapodium 02, compound falcigers chaetae;
<emphasis id="B94E3A2AFFBB222CFB43F9671080F962" bold="true" box="[1217,1235,1708,1732]" pageId="7" pageNumber="158">C</emphasis>
—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae;
<emphasis id="B94E3A2AFFBB222CFC59F91B17BEF94E" bold="true" box="[987,1005,1744,1768]" pageId="7" pageNumber="158">D</emphasis>
—parapodium 161, pectinate chaetae and aciculae;
<emphasis id="B94E3A2AFFBB222CFF75F93F155BF8AA" bold="true" box="[247,264,1780,1804]" pageId="7" pageNumber="158">E</emphasis>
—parapodium 110, compound falciger chaeta;
<emphasis id="B94E3A2AFFBB222CFD64F93F16A6F8AA" bold="true" box="[742,757,1780,1804]" pageId="7" pageNumber="158">F</emphasis>
—parapodium 99, pectinate chaetae;
<emphasis id="B94E3A2AFFBB222CFBEDF93F10D0F8AA" bold="true" box="[1135,1155,1780,1804]" pageId="7" pageNumber="158">G</emphasis>
parapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri;
<emphasis id="B94E3A2AFFBB222CFB92F8D31077F896" bold="true" box="[1040,1060,1816,1840]" pageId="7" pageNumber="158">H</emphasis>
—parapodium 49, compound falciger chaeta;
<emphasis id="B94E3A2AFFBB222CFF67F8F714BCF8F2" bold="true" box="[229,239,1852,1876]" pageId="7" pageNumber="158">I</emphasis>
—parapodium 161, compound falcigers chaetae.
</paragraph>
</caption>
<caption id="DF45B6B0FFB42223FF15FDC01166FD83" box="[151,1333,523,549]" pageId="8" pageNumber="159" startId="8.[151,243,523,549]" targetType="table">
<paragraph id="8B85E638FFB42223FF15FDC01166FD83" blockId="8.[151,1333,523,549]" box="[151,1333,523,549]" pageId="8" pageNumber="159">
<emphasis id="B94E3A2AFFB42223FF15FDC0155DFD83" bold="true" box="[151,270,523,549]" pageId="8" pageNumber="159">TABLE 2.</emphasis>
Comparison of morphological features among
<taxonomicName id="4C3A9DBBFFB42223FC85FDC01796FD83" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[775,965,523,549]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">
<emphasis id="B94E3A2AFFB42223FC85FDC01796FD83" box="[775,965,523,549]" italics="true" pageId="8" pageNumber="159">Leodice ivanildae</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFB42223FC4EFDC01071FD83" bold="true" box="[972,1058,523,549]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="A27D8751FFB42223FC4EFDC01071FD83" box="[972,1058,523,549]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and most similar species.
</paragraph>
</caption>
<paragraph id="8B85E638FFB42223FD84FDF7120FF936" pageId="8" pageNumber="159">
<table id="F93A1498FFB4DDD4FF20FDF713B5F936" box="[162,2022,572,1680]" gridcols="12" gridrows="32" pageId="8" pageNumber="159">
<tr id="350AE47AFFB4DDD4FF20FDF713B5FDF3" box="[162,2022,572,597]" gridrow="0" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-4="1" rowspan-7="1" rowspan-8="1">
<th id="76DB8D06FFB4DDD4FE44FDF716FEFDF3" box="[454,685,572,597]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="8" pageNumber="159">Branchiae</th>
<th id="76DB8D06FFB4DDD4FCDDFDF71033FDF3" box="[863,1120,572,597]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="8" pageNumber="159">Subacicular hooks</th>
<th id="76DB8D06FFB4DDD4FA51FDF71288FDF3" box="[1491,1755,572,597]" gridcol="9" gridrow="0" pageId="8" pageNumber="159">Aciculae</th>
</tr>
<tr id="350AE47AFFB4DDD4FF20FDAE13B5FD60" box="[162,2022,613,710]" gridrow="1" pageId="8" pageNumber="159">
<th id="76DB8D06FFB4DDD4FF20FDAE155FFD60" box="[162,268,613,710]" gridcol="0" gridrow="1" pageId="8" pageNumber="159">Species</th>
<td id="76DB8D06FFB4DDD4FEA7FDAE15F9FD60" box="[293,426,613,710]" gridcol="1" gridrow="1" pageId="8" pageNumber="159">Articulations of antennae and palps</td>
<td id="76DB8D06FFB4DDD4FE44FDAE1673FD60" box="[454,544,613,710]" gridcol="2" gridrow="1" pageId="8" pageNumber="159">Starting chaetiger</td>
<td id="76DB8D06FFB4DDD4FDB9FDAE16FEFD60" box="[571,685,613,710]" gridcol="3" gridrow="1" pageId="8" pageNumber="159">Number of filaments</td>
<td id="76DB8D06FFB4DDD4FD44FDAE1711FD60" box="[710,834,613,710]" gridcol="4" gridrow="1" pageId="8" pageNumber="159">Notopodial cirri</td>
<td id="76DB8D06FFB4DDD4FCDDFDAE1790FD60" box="[863,963,613,710]" gridcol="5" gridrow="1" pageId="8" pageNumber="159">Starting chaetigers</td>
<td id="76DB8D06FFB4DDD4FC6AFDAE1033FD60" box="[1000,1120,613,710]" gridcol="6" gridrow="1" pageId="8" pageNumber="159">Number per parapodium</td>
<td id="76DB8D06FFB4DDD4FB00FDAE10AAFD60" box="[1154,1273,613,710]" gridcol="7" gridrow="1" pageId="8" pageNumber="159">Compound falcigers</td>
<td id="76DB8D06FFB4DDD4FA98FDAE11EFFD60" box="[1306,1468,613,710]" gridcol="8" gridrow="1" pageId="8" pageNumber="159">Pectinate chaetae</td>
<td id="76DB8D06FFB4DDD4FA51FDAE1288FD60" box="[1491,1755,613,710]" gridcol="9" gridrow="1" pageId="8" pageNumber="159">Shape</td>
<td id="76DB8D06FFB4DDD4F975FDAE133DFD60" box="[1783,1902,613,710]" gridcol="10" gridrow="1" pageId="8" pageNumber="159">Number per parapodium</td>
<td id="76DB8D06FFB4DDD4F809FDAE13B5FD60" box="[1931,2022,613,710]" gridcol="11" gridrow="1" pageId="8" pageNumber="159">Data source</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FD3213B5FCB4" box="[162,2022,761,786]" gridrow="2" pageId="8" pageNumber="159">
<th id="76DB8D06FFB4DDD4FF20FD32155FFCB4" box="[162,268,761,786]" gridcol="0" gridrow="2" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FD3214A3FCB4" authorityName="Lamarck" authorityYear="1818" box="[162,240,761,786]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB42223FF20FD3214A3FCB4" box="[162,240,761,786]" italics="true" pageId="8" pageNumber="159">Leodice</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FD3215F9FCB4" box="[293,426,761,786]" gridcol="1" gridrow="2" pageId="8" pageNumber="159">moniliform</td>
<td id="76DB8D06FFB4DDD4FE44FD321673FCB4" box="[454,544,761,786]" gridcol="2" gridrow="2" pageId="8" pageNumber="159">34</td>
<td id="76DB8D06FFB4DDD4FDB9FD3216FEFCB4" box="[571,685,761,786]" gridcol="3" gridrow="2" pageId="8" pageNumber="159">113</td>
<td id="76DB8D06FFB4DDD4FD44FD321711FCB4" box="[710,834,761,786]" gridcol="4" gridrow="2" pageId="8" pageNumber="159">articulated</td>
<td id="76DB8D06FFB4DDD4FCDDFD321790FCB4" box="[863,963,761,786]" gridcol="5" gridrow="2" pageId="8" pageNumber="159">3143</td>
<td id="76DB8D06FFB4DDD4FC6AFD321033FCB4" box="[1000,1120,761,786]" gridcol="6" gridrow="2" pageId="8" pageNumber="159">12</td>
<td id="76DB8D06FFB4DDD4FB00FD3210AAFCB4" box="[1154,1273,761,786]" gridcol="7" gridrow="2" pageId="8" pageNumber="159">bidentate</td>
<td id="76DB8D06FFB4DDD4FA98FD3211EFFCB4" box="[1306,1468,761,786]" gridcol="8" gridrow="2" pageId="8" pageNumber="159">both marginal</td>
<td id="76DB8D06FFB4DDD4FA51FD321288FCB4" box="[1491,1755,761,786]" gridcol="9" gridrow="2" pageId="8" pageNumber="159">yellow, distally expanded</td>
<td id="76DB8D06FFB4DDD4F975FD32133DFCB4" box="[1783,1902,761,786]" gridcol="10" gridrow="2" pageId="8" pageNumber="159">paired</td>
<td id="76DB8D06FFB4DDD4F809FD3213B5FCB4" box="[1931,2022,761,786]" gridcol="11" gridrow="2" pageId="8" pageNumber="159">present</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FCDD13B5FC89" box="[162,2022,790,815]" gridrow="3" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FCDD155FFC89" box="[162,268,790,815]" gridcol="0" gridrow="3" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FCDD1553FC89" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[162,256,790,815]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="ivanildae">
<emphasis id="B94E3A2AFFB42223FF20FCDD1553FC89" box="[162,256,790,815]" italics="true" pageId="8" pageNumber="159">ivanildae</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FD44FCDD1711FC89" box="[710,834,790,815]" gridcol="4" gridrow="3" pageId="8" pageNumber="159">in all</td>
<td id="76DB8D06FFB4DDD4FA98FCDD11EFFC89" box="[1306,1468,790,815]" gridcol="8" gridrow="3" pageId="8" pageNumber="159">teeth longer</td>
<td id="76DB8D06FFB4DDD4FA51FCDD1288FC89" box="[1491,1755,790,815]" gridcol="9" gridrow="3" pageId="8" pageNumber="159">into round lobes in</td>
<td id="76DB8D06FFB4DDD4F809FCDD13B5FC89" box="[1931,2022,790,815]" gridcol="11" gridrow="3" pageId="8" pageNumber="159">study</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FCFF13B5FCEB" box="[162,2022,820,845]" gridrow="4" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FCFF155FFCEB" box="[162,268,820,845]" gridcol="0" gridrow="4" pageId="8" pageNumber="159">
<emphasis id="B94E3A2AFFB42223FF20FCFF14A1FCEB" bold="true" box="[162,242,820,845]" pageId="8" pageNumber="159">sp. nov.</emphasis>
</th>
<td id="76DB8D06FFB4DDD4FD44FCFF1711FCEB" box="[710,834,820,845]" gridcol="4" gridrow="4" pageId="8" pageNumber="159">chaetigers</td>
<td id="76DB8D06FFB4DDD4FA98FCFF11EFFCEB" box="[1306,1468,820,845]" gridcol="8" gridrow="4" pageId="8" pageNumber="159">than other</td>
<td id="76DB8D06FFB4DDD4FA51FCFF1288FCEB" box="[1491,1755,820,845]" gridcol="9" gridrow="4" pageId="8" pageNumber="159">the anterior parapodia,</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FC9A13B5FCCC" box="[162,2022,849,874]" gridrow="5" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td id="76DB8D06FFB4DDD4FA98FC9A11EFFCCC" box="[1306,1468,849,874]" gridcol="8" gridrow="5" pageId="8" pageNumber="159">teeth, but one</td>
<td id="76DB8D06FFB4DDD4FA51FC9A1288FCCC" box="[1491,1755,849,874]" gridcol="9" gridrow="5" pageId="8" pageNumber="159">becoming slightly curved</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FCA513B5FC21" box="[162,2022,878,903]" gridrow="6" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td id="76DB8D06FFB4DDD4FA98FCA511EFFC21" box="[1306,1468,878,903]" gridcol="8" gridrow="6" pageId="8" pageNumber="159">marginal tooth</td>
<td id="76DB8D06FFB4DDD4FA51FCA51288FC21" box="[1491,1755,878,903]" gridcol="9" gridrow="6" pageId="8" pageNumber="159">and tapering in posterior</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FC4713B5FC03" box="[162,2022,908,933]" gridrow="7" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td id="76DB8D06FFB4DDD4FA98FC4711EFFC03" box="[1306,1468,908,933]" gridcol="8" gridrow="7" pageId="8" pageNumber="159">longer than the</td>
<td id="76DB8D06FFB4DDD4FA51FC471288FC03" box="[1491,1755,908,933]" gridcol="9" gridrow="7" pageId="8" pageNumber="159">chaetigers</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FC6213B5FC64" box="[162,2022,937,962]" gridrow="8" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<td id="76DB8D06FFB4DDD4FA98FC6211EFFC64" box="[1306,1468,937,962]" gridcol="8" gridrow="8" pageId="8" pageNumber="159">other</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FC1A13B5FC4C" box="[162,2022,977,1002]" gridrow="9" pageId="8" pageNumber="159" rowspan-4="1">
<th id="76DB8D06FFB4DDD4FF20FC1A155FFC4C" box="[162,268,977,1002]" gridcol="0" gridrow="9" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FC1A14A3FC4C" authorityName="Lamarck" authorityYear="1818" box="[162,240,977,1002]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB42223FF20FC1A14A3FC4C" box="[162,240,977,1002]" italics="true" pageId="8" pageNumber="159">Leodice</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FC1A15F9FC4C" box="[293,426,977,1002]" gridcol="1" gridrow="9" pageId="8" pageNumber="159">regularly</td>
<td id="76DB8D06FFB4DDD4FE44FC1A1673FC4C" box="[454,544,977,1002]" gridcol="2" gridrow="9" pageId="8" pageNumber="159">4</td>
<td id="76DB8D06FFB4DDD4FDB9FC1A16FEFC4C" box="[571,685,977,1002]" gridcol="3" gridrow="9" pageId="8" pageNumber="159">16</td>
<td id="76DB8D06FFB4DDD4FCDDFC1A1790FC4C" box="[863,963,977,1002]" gridcol="5" gridrow="9" pageId="8" pageNumber="159">40</td>
<td id="76DB8D06FFB4DDD4FC6AFC1A1033FC4C" box="[1000,1120,977,1002]" gridcol="6" gridrow="9" pageId="8" pageNumber="159">single</td>
<td id="76DB8D06FFB4DDD4FB00FC1A10AAFC4C" box="[1154,1273,977,1002]" gridcol="7" gridrow="9" pageId="8" pageNumber="159">bidentate</td>
<td id="76DB8D06FFB4DDD4FA98FC1A11EFFC4C" box="[1306,1468,977,1002]" gridcol="8" gridrow="9" pageId="8" pageNumber="159">one marginal</td>
<td id="76DB8D06FFB4DDD4FA51FC1A1288FC4C" box="[1491,1755,977,1002]" gridcol="9" gridrow="9" pageId="8" pageNumber="159">yellow, distally pointed,</td>
<td id="76DB8D06FFB4DDD4F975FC1A133DFC4C" box="[1783,1902,977,1002]" gridcol="10" gridrow="9" pageId="8" pageNumber="159">23</td>
<td id="76DB8D06FFB4DDD4F809FC1A13B5FC4C" box="[1931,2022,977,1002]" gridcol="11" gridrow="9" pageId="8" pageNumber="159">Fauchald</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FC2513B5FBA1" box="[162,2022,1006,1031]" gridrow="10" pageId="8" pageNumber="159" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FC25155FFBA1" box="[162,268,1006,1031]" gridcol="0" gridrow="10" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FC251555FBA1" baseAuthorityName="Ehlers" baseAuthorityYear="1887" box="[162,262,1006,1031]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="articulata">
<emphasis id="B94E3A2AFFB42223FF20FC251555FBA1" box="[162,262,1006,1031]" italics="true" pageId="8" pageNumber="159">articulata</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FC2515F9FBA1" box="[293,426,1006,1031]" gridcol="1" gridrow="10" pageId="8" pageNumber="159">cylindrical</td>
<td id="76DB8D06FFB4DDD4FD44FC251711FBA1" box="[710,834,1006,1031]" gridcol="4" gridrow="10" pageId="8" pageNumber="159">articulated</td>
<td id="76DB8D06FFB4DDD4FA98FC2511EFFBA1" box="[1306,1468,1006,1031]" gridcol="8" gridrow="10" pageId="8" pageNumber="159">tooth longer</td>
<td id="76DB8D06FFB4DDD4FA51FC251288FBA1" box="[1491,1755,1006,1031]" gridcol="9" gridrow="10" pageId="8" pageNumber="159">straight</td>
<td id="76DB8D06FFB4DDD4F809FC2513B5FBA1" box="[1931,2022,1006,1031]" gridcol="11" gridrow="10" pageId="8" pageNumber="159">1992</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FBC713B5FB83" box="[162,2022,1036,1061]" gridrow="11" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76DB8D06FFB4DDD4FF20FBC7155FFB83" box="[162,268,1036,1061]" gridcol="0" gridrow="11" pageId="8" pageNumber="159">(Ehlers,</th>
<td id="76DB8D06FFB4DDD4FD44FBC71711FB83" box="[710,834,1036,1061]" gridcol="4" gridrow="11" pageId="8" pageNumber="159">in anterior</td>
<td id="76DB8D06FFB4DDD4FA98FBC711EFFB83" box="[1306,1468,1036,1061]" gridcol="8" gridrow="11" pageId="8" pageNumber="159">than the other</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FBE213B5FBE4" box="[162,2022,1065,1090]" gridrow="12" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76DB8D06FFB4DDD4FF20FBE2155FFBE4" box="[162,268,1065,1090]" gridcol="0" gridrow="12" pageId="8" pageNumber="159">1887)</th>
<td id="76DB8D06FFB4DDD4FD44FBE21711FBE4" box="[710,834,1065,1090]" gridcol="4" gridrow="12" pageId="8" pageNumber="159">chaetigers</td>
<td id="76DB8D06FFB4DDD4FA98FBE211EFFBE4" box="[1306,1468,1065,1090]" gridcol="8" gridrow="12" pageId="8" pageNumber="159">teeth</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FB9A13B5FBCC" box="[162,2022,1105,1130]" gridrow="13" pageId="8" pageNumber="159">
<th id="76DB8D06FFB4DDD4FF20FB9A155FFBCC" box="[162,268,1105,1130]" gridcol="0" gridrow="13" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FB9A14A3FBCC" authorityName="Lamarck" authorityYear="1818" box="[162,240,1105,1130]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB42223FF20FB9A14A3FBCC" box="[162,240,1105,1130]" italics="true" pageId="8" pageNumber="159">Leodice</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FB9A15F9FBCC" box="[293,426,1105,1130]" gridcol="1" gridrow="13" pageId="8" pageNumber="159">irregularly</td>
<td id="76DB8D06FFB4DDD4FE44FB9A1673FBCC" box="[454,544,1105,1130]" gridcol="2" gridrow="13" pageId="8" pageNumber="159">4</td>
<td id="76DB8D06FFB4DDD4FDB9FB9A16FEFBCC" box="[571,685,1105,1130]" gridcol="3" gridrow="13" pageId="8" pageNumber="159">16</td>
<td id="76DB8D06FFB4DDD4FD44FB9A1711FBCC" box="[710,834,1105,1130]" gridcol="4" gridrow="13" pageId="8" pageNumber="159">articulated</td>
<td id="76DB8D06FFB4DDD4FCDDFB9A1790FBCC" box="[863,963,1105,1130]" gridcol="5" gridrow="13" pageId="8" pageNumber="159">33</td>
<td id="76DB8D06FFB4DDD4FC6AFB9A1033FBCC" box="[1000,1120,1105,1130]" gridcol="6" gridrow="13" pageId="8" pageNumber="159">12</td>
<td id="76DB8D06FFB4DDD4FB00FB9A10AAFBCC" box="[1154,1273,1105,1130]" gridcol="7" gridrow="13" pageId="8" pageNumber="159">bidentate</td>
<td id="76DB8D06FFB4DDD4FA98FB9A11EFFBCC" box="[1306,1468,1105,1130]" gridcol="8" gridrow="13" pageId="8" pageNumber="159">both marginal</td>
<td id="76DB8D06FFB4DDD4FA51FB9A1288FBCC" box="[1491,1755,1105,1130]" gridcol="9" gridrow="13" pageId="8" pageNumber="159">yellow, flattened and</td>
<td id="76DB8D06FFB4DDD4F975FB9A133DFBCC" box="[1783,1902,1105,1130]" gridcol="10" gridrow="13" pageId="8" pageNumber="159">paired</td>
<td id="76DB8D06FFB4DDD4F809FB9A13B5FBCC" box="[1931,2022,1105,1130]" gridcol="11" gridrow="13" pageId="8" pageNumber="159">Fauchald</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FBA413B5FB2E" box="[162,2022,1135,1160]" gridrow="14" pageId="8" pageNumber="159" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FBA4155FFB2E" box="[162,268,1135,1160]" gridcol="0" gridrow="14" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FBA41554FB2E" baseAuthorityName="Ehlers" baseAuthorityYear="1887" box="[162,263,1135,1160]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="antillensis">
<emphasis id="B94E3A2AFFB42223FF20FBA41554FB2E" box="[162,263,1135,1160]" italics="true" pageId="8" pageNumber="159">antillensis</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FBA415F9FB2E" box="[293,426,1135,1160]" gridcol="1" gridrow="14" pageId="8" pageNumber="159">cylindrical</td>
<td id="76DB8D06FFB4DDD4FD44FBA41711FB2E" box="[710,834,1135,1160]" gridcol="4" gridrow="14" pageId="8" pageNumber="159">in all</td>
<td id="76DB8D06FFB4DDD4FA98FBA411EFFB2E" box="[1306,1468,1135,1160]" gridcol="8" gridrow="14" pageId="8" pageNumber="159">teeth longer</td>
<td id="76DB8D06FFB4DDD4FA51FBA41288FB2E" box="[1491,1755,1135,1160]" gridcol="9" gridrow="14" pageId="8" pageNumber="159">expanded tips</td>
<td id="76DB8D06FFB4DDD4F809FBA413B5FB2E" box="[1931,2022,1135,1160]" gridcol="11" gridrow="14" pageId="8" pageNumber="159">1992</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FB4713B5FB03" box="[162,2022,1164,1189]" gridrow="15" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76DB8D06FFB4DDD4FF20FB47155FFB03" box="[162,268,1164,1189]" gridcol="0" gridrow="15" pageId="8" pageNumber="159">(Ehlers,</th>
<td id="76DB8D06FFB4DDD4FD44FB471711FB03" box="[710,834,1164,1189]" gridcol="4" gridrow="15" pageId="8" pageNumber="159">chaetigers</td>
<td id="76DB8D06FFB4DDD4FA98FB4711EFFB03" box="[1306,1468,1164,1189]" gridcol="8" gridrow="15" pageId="8" pageNumber="159">than other teeth</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FB6213B5FB64" box="[162,2022,1193,1218]" gridrow="16" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76DB8D06FFB4DDD4FF20FB62155FFB64" box="[162,268,1193,1218]" gridcol="0" gridrow="16" pageId="8" pageNumber="159">1887)</th>
</tr>
<tr id="350AE47AFFB4DDD4FF20FB1913B5FB4D" box="[162,2022,1234,1259]" gridrow="17" pageId="8" pageNumber="159">
<th id="76DB8D06FFB4DDD4FF20FB19155FFB4D" box="[162,268,1234,1259]" gridcol="0" gridrow="17" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FB1914A3FB4D" authorityName="Lamarck" authorityYear="1818" box="[162,240,1234,1259]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB42223FF20FB1914A3FB4D" box="[162,240,1234,1259]" italics="true" pageId="8" pageNumber="159">Leodice</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FB1915F9FB4D" box="[293,426,1234,1259]" gridcol="1" gridrow="17" pageId="8" pageNumber="159">regularly</td>
<td id="76DB8D06FFB4DDD4FE44FB191673FB4D" box="[454,544,1234,1259]" gridcol="2" gridrow="17" pageId="8" pageNumber="159">3</td>
<td id="76DB8D06FFB4DDD4FDB9FB1916FEFB4D" box="[571,685,1234,1259]" gridcol="3" gridrow="17" pageId="8" pageNumber="159">118</td>
<td id="76DB8D06FFB4DDD4FD44FB191711FB4D" box="[710,834,1234,1259]" gridcol="4" gridrow="17" pageId="8" pageNumber="159">articulated</td>
<td id="76DB8D06FFB4DDD4FCDDFB191790FB4D" box="[863,963,1234,1259]" gridcol="5" gridrow="17" pageId="8" pageNumber="159">32</td>
<td id="76DB8D06FFB4DDD4FC6AFB191033FB4D" box="[1000,1120,1234,1259]" gridcol="6" gridrow="17" pageId="8" pageNumber="159">13</td>
<td id="76DB8D06FFB4DDD4FB00FB1910AAFB4D" box="[1154,1273,1234,1259]" gridcol="7" gridrow="17" pageId="8" pageNumber="159">bidentate</td>
<td id="76DB8D06FFB4DDD4FA98FB1911EFFB4D" box="[1306,1468,1234,1259]" gridcol="8" gridrow="17" pageId="8" pageNumber="159">both marginal</td>
<td id="76DB8D06FFB4DDD4FA51FB191288FB4D" box="[1491,1755,1234,1259]" gridcol="9" gridrow="17" pageId="8" pageNumber="159">yellow, anterior bluntly</td>
<td id="76DB8D06FFB4DDD4F975FB19133DFB4D" box="[1783,1902,1234,1259]" gridcol="10" gridrow="17" pageId="8" pageNumber="159">paired</td>
<td id="76DB8D06FFB4DDD4F809FB1913B5FB4D" box="[1931,2022,1234,1259]" gridcol="11" gridrow="17" pageId="8" pageNumber="159">Fauchald</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FB2413B5FAAE" box="[162,2022,1263,1288]" gridrow="18" pageId="8" pageNumber="159" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FB24155FFAAE" box="[162,268,1263,1288]" gridcol="0" gridrow="18" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FB241550FAAE" baseAuthorityName="Fauchald" baseAuthorityYear="1970" box="[162,259,1263,1288]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="mexicana">
<emphasis id="B94E3A2AFFB42223FF20FB241550FAAE" box="[162,259,1263,1288]" italics="true" pageId="8" pageNumber="159">mexicana</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FB2415F9FAAE" box="[293,426,1263,1288]" gridcol="1" gridrow="18" pageId="8" pageNumber="159">cylindrical</td>
<td id="76DB8D06FFB4DDD4FD44FB241711FAAE" box="[710,834,1263,1288]" gridcol="4" gridrow="18" pageId="8" pageNumber="159">in anterior</td>
<td id="76DB8D06FFB4DDD4FA98FB2411EFFAAE" box="[1306,1468,1263,1288]" gridcol="8" gridrow="18" pageId="8" pageNumber="159">teeth longer</td>
<td id="76DB8D06FFB4DDD4FA51FB241288FAAE" box="[1491,1755,1263,1288]" gridcol="9" gridrow="18" pageId="8" pageNumber="159">pointed, distally straight,</td>
<td id="76DB8D06FFB4DDD4F809FB2413B5FAAE" box="[1931,2022,1263,1288]" gridcol="11" gridrow="18" pageId="8" pageNumber="159">1992</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FAC713B5FA83" box="[162,2022,1292,1317]" gridrow="19" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FAC7155FFA83" box="[162,268,1292,1317]" gridcol="0" gridrow="19" pageId="8" pageNumber="159">(Fauchald,</th>
<td id="76DB8D06FFB4DDD4FD44FAC71711FA83" box="[710,834,1292,1317]" gridcol="4" gridrow="19" pageId="8" pageNumber="159">and median</td>
<td id="76DB8D06FFB4DDD4FA98FAC711EFFA83" box="[1306,1468,1292,1317]" gridcol="8" gridrow="19" pageId="8" pageNumber="159">than other teeth</td>
<td id="76DB8D06FFB4DDD4FA51FAC71288FA83" box="[1491,1755,1292,1317]" gridcol="9" gridrow="19" pageId="8" pageNumber="159">medially at least</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FAE113B5FAE5" box="[162,2022,1322,1347]" gridrow="20" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76DB8D06FFB4DDD4FF20FAE1155FFAE5" box="[162,268,1322,1347]" gridcol="0" gridrow="20" pageId="8" pageNumber="159">1970)</th>
<td id="76DB8D06FFB4DDD4FD44FAE11711FAE5" box="[710,834,1322,1347]" gridcol="4" gridrow="20" pageId="8" pageNumber="159">chaetigers,</td>
<td id="76DB8D06FFB4DDD4FA51FAE11288FAE5" box="[1491,1755,1322,1347]" gridcol="9" gridrow="20" pageId="8" pageNumber="159">superior most aciculae</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FA8C13B5FAC6" box="[162,2022,1351,1376]" gridrow="21" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FD44FA8C1711FAC6" box="[710,834,1351,1376]" gridcol="4" gridrow="21" pageId="8" pageNumber="159">posterior</td>
<td id="76DB8D06FFB4DDD4FA51FA8C1288FAC6" box="[1491,1755,1351,1376]" gridcol="9" gridrow="21" pageId="8" pageNumber="159">distally expanded into</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FAAF13B5FADB" box="[162,2022,1380,1405]" gridrow="22" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FD44FAAF1711FADB" box="[710,834,1380,1405]" gridcol="4" gridrow="22" pageId="8" pageNumber="159">without</td>
<td id="76DB8D06FFB4DDD4FA51FAAF1288FADB" box="[1491,1755,1380,1405]" gridcol="9" gridrow="22" pageId="8" pageNumber="159">tabs; posterior tapering,</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FA4913B5FA3D" box="[162,2022,1410,1435]" gridrow="23" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FD44FA491711FA3D" box="[710,834,1410,1435]" gridcol="4" gridrow="23" pageId="8" pageNumber="159">articulations</td>
<td id="76DB8D06FFB4DDD4FA51FA491288FA3D" box="[1491,1755,1410,1435]" gridcol="9" gridrow="23" pageId="8" pageNumber="159">distinctly bent dorsally.</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FA6113B5FA65" box="[162,2022,1450,1475]" gridrow="24" pageId="8" pageNumber="159">
<th id="76DB8D06FFB4DDD4FF20FA61155FFA65" box="[162,268,1450,1475]" gridcol="0" gridrow="24" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FA6114A3FA65" authorityName="Lamarck" authorityYear="1818" box="[162,240,1450,1475]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="genus">
<emphasis id="B94E3A2AFFB42223FF20FA6114A3FA65" box="[162,240,1450,1475]" italics="true" pageId="8" pageNumber="159">Leodice</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FA6115F9FA65" box="[293,426,1450,1475]" gridcol="1" gridrow="24" pageId="8" pageNumber="159">regularly</td>
<td id="76DB8D06FFB4DDD4FE44FA611673FA65" box="[454,544,1450,1475]" gridcol="2" gridrow="24" pageId="8" pageNumber="159">3</td>
<td id="76DB8D06FFB4DDD4FDB9FA6116FEFA65" box="[571,685,1450,1475]" gridcol="3" gridrow="24" pageId="8" pageNumber="159">115</td>
<td id="76DB8D06FFB4DDD4FD44FA611711FA65" box="[710,834,1450,1475]" gridcol="4" gridrow="24" pageId="8" pageNumber="159">articulated</td>
<td id="76DB8D06FFB4DDD4FCDDFA611790FA65" box="[863,963,1450,1475]" gridcol="5" gridrow="24" pageId="8" pageNumber="159">31</td>
<td id="76DB8D06FFB4DDD4FC6AFA611033FA65" box="[1000,1120,1450,1475]" gridcol="6" gridrow="24" pageId="8" pageNumber="159">12</td>
<td id="76DB8D06FFB4DDD4FB00FA6110AAFA65" box="[1154,1273,1450,1475]" gridcol="7" gridrow="24" pageId="8" pageNumber="159">bidentate</td>
<td id="76DB8D06FFB4DDD4FA98FA6111EFFA65" box="[1306,1468,1450,1475]" gridcol="8" gridrow="24" pageId="8" pageNumber="159">one marginal</td>
<td id="76DB8D06FFB4DDD4FA51FA611288FA65" box="[1491,1755,1450,1475]" gridcol="9" gridrow="24" pageId="8" pageNumber="159">anterior bluntly tapering</td>
<td id="76DB8D06FFB4DDD4F975FA61133DFA65" box="[1783,1902,1450,1475]" gridcol="10" gridrow="24" pageId="8" pageNumber="159">23</td>
<td id="76DB8D06FFB4DDD4F809FA6113B5FA65" box="[1931,2022,1450,1475]" gridcol="11" gridrow="24" pageId="8" pageNumber="159">Fauchald</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FA0C13B5FA46" box="[162,2022,1479,1504]" gridrow="25" pageId="8" pageNumber="159" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FA0C155FFA46" box="[162,268,1479,1504]" gridcol="0" gridrow="25" pageId="8" pageNumber="159">
<taxonomicName id="4C3A9DBBFFB42223FF20FA0C14ABFA46" baseAuthorityName="Fauchald" baseAuthorityYear="1969" box="[162,248,1479,1504]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="159" phylum="Annelida" rank="species" species="websteri">
<emphasis id="B94E3A2AFFB42223FF20FA0C14ABFA46" box="[162,248,1479,1504]" italics="true" pageId="8" pageNumber="159">websteri</emphasis>
</taxonomicName>
</th>
<td id="76DB8D06FFB4DDD4FEA7FA0C15F9FA46" box="[293,426,1479,1504]" gridcol="1" gridrow="25" pageId="8" pageNumber="159">cylindrical</td>
<td id="76DB8D06FFB4DDD4FD44FA0C1711FA46" box="[710,834,1479,1504]" gridcol="4" gridrow="25" pageId="8" pageNumber="159">in all</td>
<td id="76DB8D06FFB4DDD4FA98FA0C11EFFA46" box="[1306,1468,1479,1504]" gridcol="8" gridrow="25" pageId="8" pageNumber="159">tooth longer</td>
<td id="76DB8D06FFB4DDD4FA51FA0C1288FA46" box="[1491,1755,1479,1504]" gridcol="9" gridrow="25" pageId="8" pageNumber="159">or nearly truncate;</td>
<td id="76DB8D06FFB4DDD4F809FA0C13B5FA46" box="[1931,2022,1479,1504]" gridcol="11" gridrow="25" pageId="8" pageNumber="159">1992</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20FA2E13B5FA58" box="[162,2022,1509,1534]" gridrow="26" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DB8D06FFB4DDD4FF20FA2E155FFA58" box="[162,268,1509,1534]" gridcol="0" gridrow="26" pageId="8" pageNumber="159">(Fauchald,</th>
<td id="76DB8D06FFB4DDD4FD44FA2E1711FA58" box="[710,834,1509,1534]" gridcol="4" gridrow="26" pageId="8" pageNumber="159">chaetigers</td>
<td id="76DB8D06FFB4DDD4FA98FA2E11EFFA58" box="[1306,1468,1509,1534]" gridcol="8" gridrow="26" pageId="8" pageNumber="159">than other teeth</td>
<td id="76DB8D06FFB4DDD4FA51FA2E1288FA58" box="[1491,1755,1509,1534]" gridcol="9" gridrow="26" pageId="8" pageNumber="159">median distally expanded</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20F9C913B5F9BD" box="[162,2022,1538,1563]" gridrow="27" pageId="8" pageNumber="159" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76DB8D06FFB4DDD4FF20F9C9155FF9BD" box="[162,268,1538,1563]" gridcol="0" gridrow="27" pageId="8" pageNumber="159">1969)</th>
<td id="76DB8D06FFB4DDD4FA51F9C91288F9BD" box="[1491,1755,1538,1563]" gridcol="9" gridrow="27" pageId="8" pageNumber="159">to distinct rounded tabs;</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20F9D413B5F99E" box="[162,2022,1567,1592]" gridrow="28" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FA51F9D41288F99E" box="[1491,1755,1567,1592]" gridcol="9" gridrow="28" pageId="8" pageNumber="159">posterior becoming</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20F9F613B5F9F0" box="[162,2022,1597,1622]" gridrow="29" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FA51F9F61288F9F0" box="[1491,1755,1597,1622]" gridcol="9" gridrow="29" pageId="8" pageNumber="159">slender, tapering to fine</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20F99113B5F9D5" box="[162,2022,1626,1651]" gridrow="30" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FA51F9911288F9D5" box="[1491,1755,1626,1651]" gridcol="9" gridrow="30" pageId="8" pageNumber="159">tips, sometimes distinctly</td>
</tr>
<tr id="350AE47AFFB4DDD4FF20F9BC13B5F936" box="[162,2022,1655,1680]" gridrow="31" pageId="8" pageNumber="159" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<td id="76DB8D06FFB4DDD4FA51F9BC1288F936" box="[1491,1755,1655,1680]" gridcol="9" gridrow="31" pageId="8" pageNumber="159">bent dorsally.</td>
</tr>
</table>
</paragraph>
<paragraph id="8B85E638FFB52222FF45FF5C159AFDBF" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
Limbate chaetae, tapering, serrated marginally, longer in posterior parapodia. Pectinate chaetae with curved blades, 1012 teeth, marginal teeth unequal in length and longer than other teeth (
<figureCitation id="1301FABDFFB52222FBAEFF7010F7FF73" box="[1068,1188,187,213]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="9" pageNumber="160">Fig. 3D, F</figureCitation>
). Compound falcigers chaetae bidentate; anterior parapodia, article tapering abruptly from proximal to distal end, proximal tooth laterally directed, shorter than distal tooth, guards slightly inflated and marginally serrated; in median and posterior parapodia articles longer and narrower than in anterior parapodia, proximal tooth tapering, laterally directed, straight, longer, and thicker than laterally directed distal tooth (
<figureCitation id="1301FABDFFB52222FD37FE801767FEC3" box="[693,820,331,357]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="9" pageNumber="160">Fig. 3B, H</figureCitation>
), guards more inflated than in anterior parapodia and coarsely serrated (
<figureCitation id="1301FABDFFB52222FEE4FEA41585FE2F" box="[358,470,367,393]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="9" pageNumber="160">Fig. 3E, I</figureCitation>
). Aciculae yellow, paired; distally expanded in anterior parapodia, becoming slightly curved and tapering in posterior chaetigers (
<figureCitation id="1301FABDFFB52222FD18FE581710FE0B" box="[666,835,403,429]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="9" pageNumber="160">Fig. 3A, C, D</figureCitation>
). Subacicular hooks yellow, bidentate, present from chaetiger 41(3143); paired in some posterior chaetigers; guard leaf-shaped and partially covering proximal tooth; distal tooth thin, narrow, almost erect, and distally directed; proximal tooth larger than distal tooth, tapered, and laterally directed (
<figureCitation id="1301FABDFFB52222FEE1FE3415EFFDBF" box="[355,444,511,537]" captionStart="FIGURE 3" captionStartId="7.[152,255,1672,1696]" captionTargetBox="[151,1436,181,1648]" captionTargetId="figure-19@7.[151,1436,181,1648]" captionTargetPageId="7" captionText="FIGURE 3. Leodice ivanildae sp. nov., AD, MNRJP-007863. FG MNRJP-007866.Parapodia and chaetae. A—parapodium 09, aciculae, compound falcigers and ventral cirri; B—parapodium 02, compound falcigers chaetae; C—parapodium 161, aciculae, subacicular hooks, pectinate chaetae and compound falcigers chaetae; D—parapodium 161, pectinate chaetae and aciculae; E—parapodium 110, compound falciger chaeta; F—parapodium 99, pectinate chaetae; Gparapodium 11, branchiae, notopodial cirri, acicula, pectinate chaetae, compound falcigers chaetae and ventral cirri; H—parapodium 49, compound falciger chaeta; I—parapodium 161, compound falcigers chaetae." figureDoi="http://doi.org/10.5281/zenodo.13219184" httpUri="https://zenodo.org/record/13219184/files/figure.png" pageId="9" pageNumber="160">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8B85E638FFB52222FF45FDE817FBFDC7" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">Pygidium with two pairs of pygidial cirri. Dorsal cirri digitiform, with about 910 articulations; five times longer than ventral cirri; ventral cirri digitiform, without articulations.</paragraph>
<paragraph id="8B85E638FFB52222FF45FDA116F5FC9F" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF45FDA11519FD22" bold="true" box="[199,330,618,644]" pageId="9" pageNumber="160">Variations.</emphasis>
Specimens analyzed showed variation in: 1. total number of chaetigers, body width and length; 2. shape of prostomium, varying from rounded to truncated; 3. length of peristomium cirri, reaching either anterior margin of the prostomium or beyond it; 4. number of teeth of MxIII and MxIV; 5. beginning of branchial distribution; 6. maximum number of branchial filaments; 7. number of articulations of notopodial cirri in anterior, median and posterior parapodia; 8. number of teeth of pectinate chaetae; 9. beginning of subacicular hook distribution; 10. number of articulations of dorsal pygidial cirri.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB52222FF45FC8815B1FC27" pageId="9" pageNumber="160" type="materials_examined">
<paragraph id="8B85E638FFB52222FF45FC8815B1FC27" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
<materialsCitation id="3B52EC65FFB52222FF45FC88158EFC27" country="Brazil" county="State" elevation="17" latitude="-6.483333" location="Southwest Atlantic Ocean" longLatPrecision="1304" longitude="-34.9" municipality="Continental Shelf" pageId="9" pageNumber="160" specimenCount="1" stateProvince="Paraiba" typeStatus="holotype">
<emphasis id="B94E3A2AFFB52222FF45FC88153BFCFA" bold="true" box="[199,360,834,861]" pageId="9" pageNumber="160">
<typeStatus id="5481589AFFB52222FF45FC881551FCFB" box="[199,258,835,861]" pageId="9" pageNumber="160">Type</typeStatus>
locality.
</emphasis>
<location id="8EE5B0E3FFB52222FEF3FC8816CFFCFB" LSID="urn:lsid:plazi:treatment:0393572EFFB92222FF15FEA515A8FA67:8EE5B0E3FFB52222FEF3FC8816CFFCFB" box="[369,668,835,861]" country="Brazil" county="State" latitude="-6.483333" longLatPrecision="1304" longitude="-34.9" municipality="Continental Shelf" name="Southwest Atlantic Ocean" pageId="9" pageNumber="160" stateProvince="Paraiba">Southwest Atlantic Ocean</location>
,
<collectingMunicipality id="6BE17C42FFB52222FD28FC881722FCFB" box="[682,881,835,861]" pageId="9" pageNumber="160">Continental Shelf</collectingMunicipality>
along the coast of the
<collectingCounty id="62E49EB4FFB52222FBFCFC8810E5FCFB" box="[1150,1206,835,861]" pageId="9" pageNumber="160">State</collectingCounty>
of
<collectingRegion id="49FE28DAFFB52222FB63FC881166FCFB" box="[1249,1333,835,861]" country="Brazil" name="Paraiba" pageId="9" pageNumber="160">Paraíba</collectingRegion>
(
<collectingCountry id="F32DA6A8FFB52222FAC4FC8811C3FCFB" box="[1350,1424,835,861]" name="Brazil" pageId="9" pageNumber="160">Brazil</collectingCountry>
),
<geoCoordinate id="EE0E80FFFFB52222FF15FCAD14B2FC27" box="[151,225,870,897]" degrees="6" direction="south" minutes="29" orientation="latitude" pageId="9" pageNumber="160" precision="925" value="-6.483333">6°29S</geoCoordinate>
<geoCoordinate id="EE0E80FFFFB52222FF73FCAD1509FC27" box="[241,346,870,897]" degrees="34" direction="west" minutes="54" orientation="longitude" pageId="9" pageNumber="160" precision="925" value="-34.9">34°54W</geoCoordinate>
,
<quantity id="4CC24BDDFFB52222FEE4FCAC15CCFC27" box="[358,415,871,897]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="160" unit="m" value="17.0">
<elevation id="0017010BFFB52222FEE4FCAC15CCFC27" box="[358,415,871,897]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="160" unit="m" value="17.0">17 m</elevation>
</quantity>
deep
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB52222FF45FC41171FFC6F" pageId="9" pageNumber="160" type="distribution">
<paragraph id="8B85E638FFB52222FF45FC41171FFC6F" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF45FC411532FC02" bold="true" box="[199,353,906,932]" pageId="9" pageNumber="160">Distribution.</emphasis>
Southwest Atlantic Ocean, Continental Shelf along the coast of the State of
<collectingRegion id="49FE28DAFFB52222FB62FC401167FC03" box="[1248,1332,907,933]" country="Brazil" name="Paraiba" pageId="9" pageNumber="160">Paraíba</collectingRegion>
(
<collectingCountry id="F32DA6A8FFB52222FAC4FC4011C3FC03" box="[1350,1424,907,933]" name="Brazil" pageId="9" pageNumber="160">Brazil</collectingCountry>
),
<geoCoordinate id="EE0E80FFFFB52222FF15FC6514B4FC6F" box="[151,231,942,969]" degrees="6" direction="south" minutes="58" orientation="latitude" pageId="9" pageNumber="160" precision="925" value="-6.9666667">6°58S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB52222FF71FC64150AFC6F" box="[243,345,943,969]" degrees="34" direction="west" minutes="46" orientation="longitude" pageId="9" pageNumber="160" precision="925" value="-34.766666">34°46W</geoCoordinate>
to
<geoCoordinate id="EE0E80FFFFB52222FEFCFC64159EFC6F" box="[382,461,943,969]" degrees="7" direction="south" minutes="34" orientation="latitude" pageId="9" pageNumber="160" precision="925" value="-7.5666666">7°34S</geoCoordinate>
,
<geoCoordinate id="EE0E80FFFFB52222FE58FC651612FC6F" box="[474,577,942,969]" degrees="34" direction="west" minutes="45" orientation="longitude" pageId="9" pageNumber="160" precision="925" value="-34.75">34°45W</geoCoordinate>
,
<quantity id="4CC24BDDFFB52222FDCEFC6516FCFC6F" box="[588,687,942,969]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="9" pageNumber="160" unit="m" value="22.5" valueMax="35.0" valueMin="10.0">1035 m</quantity>
deep (
<figureCitation id="1301FABDFFB52222FD78FC641768FC6F" box="[762,827,943,969]" captionStart="FIGURE 1" captionStartId="3.[152,255,1913,1937]" captionTargetBox="[151,1429,195,1889]" captionTargetId="figure-19@3.[151,1436,181,1889]" captionTargetPageId="3" captionText="FIGURE 1. General map of the stations of the Marine Algae project along the Continental Shelf of the State of Paraíba where Leodice species have been recorded." figureDoi="http://doi.org/10.5281/zenodo.13219176" httpUri="https://zenodo.org/record/13219176/files/figure.png" pageId="9" pageNumber="160">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB52222FF45FC18159AFBB7" pageId="9" pageNumber="160" type="etymology">
<paragraph id="8B85E638FFB52222FF45FC18159AFBB7" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF45FC18151DFC4B" bold="true" box="[199,334,979,1005]" pageId="9" pageNumber="160">Etymology.</emphasis>
The specific name is in honor of the mother of the first author, Ivanilda Maria, who supported the completion of the research.
</paragraph>
</subSubSection>
<subSubSection id="C320B5B3FFB52222FF45FBD015A8FA67" pageId="9" pageNumber="160" type="discussion">
<paragraph id="8B85E638FFB52222FF45FBD015A8FA67" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="160">
<emphasis id="B94E3A2AFFB52222FF45FBD01567FB93" bold="true" box="[199,308,1051,1077]" pageId="9" pageNumber="160">Remarks</emphasis>
. Among the species with branchiae restricted to the anterior body region, present in less than 55% of chaetigers, and with yellow bidentate subacicular hooks,
<emphasis id="B94E3A2AFFB52222FCBFFBF4105EFBFF" box="[829,1037,1087,1113]" italics="true" pageId="9" pageNumber="160">
<taxonomicName id="4C3A9DBBFFB52222FCBFFBF4105AFBFF" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[829,1033,1087,1113]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">Leodice ivanildae</taxonomicName>
,
</emphasis>
<emphasis id="B94E3A2AFFB52222FB97FBF4103CFBFF" bold="true" box="[1045,1135,1087,1113]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A27D8751FFB52222FB97FBF4103CFBFF" box="[1045,1135,1087,1113]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C3A9DBBFFB52222FAC6FBF415EFFBDB" authority="(Ehlers, 1887)" baseAuthorityName="Ehlers" baseAuthorityYear="1887" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="antillensis">
<emphasis id="B94E3A2AFFB52222FAC6FBF41559FBDB" italics="true" pageId="9" pageNumber="160">Leodice antillensis</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB52222FE9FFBA815E6FBDB" author="Ehlers, E." box="[285,437,1123,1149]" pageId="9" pageNumber="160" pagination="1 - 1335" refId="ref13717" refString="Ehlers, E. (1887) Reports on the results of dredging, under the direction of L. F. Pourtales, during the years 1868 - 1870, and of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1878 - 79), in the U. S. Coast Survey Steamer &quot; Blake &quot;, XXXI report on the annelids. Memoirs of the Museum of Comparative Zoology at Harvard College, 15, 1 - 1335. https: // doi. org / 10.5962 / bhl. title. 65639" type="journal article" year="1887">Ehlers, 1887</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3A9DBBFFB52222FE4EFBA81703FBDB" authority="(Ehlers, 1887)" baseAuthorityName="Ehlers" baseAuthorityYear="1887" box="[460,848,1123,1149]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="articulata">
<emphasis id="B94E3A2AFFB52222FE4EFBA816CDFBDB" box="[460,670,1123,1149]" italics="true" pageId="9" pageNumber="160">Leodice articulata</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB52222FD33FBA8171AFBDB" author="Ehlers, E." box="[689,841,1123,1149]" pageId="9" pageNumber="160" pagination="1 - 1335" refId="ref13717" refString="Ehlers, E. (1887) Reports on the results of dredging, under the direction of L. F. Pourtales, during the years 1868 - 1870, and of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1878 - 79), in the U. S. Coast Survey Steamer &quot; Blake &quot;, XXXI report on the annelids. Memoirs of the Museum of Comparative Zoology at Harvard College, 15, 1 - 1335. https: // doi. org / 10.5962 / bhl. title. 65639" type="journal article" year="1887">Ehlers, 1887</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3A9DBBFFB52222FCE2FBA81153FBDB" authority="(Fauchald, 1970)" baseAuthorityName="Fauchald" baseAuthorityYear="1970" box="[864,1280,1123,1149]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="mexicana">
<emphasis id="B94E3A2AFFB52222FCE2FBA8107CFBDA" box="[864,1071,1123,1149]" italics="true" pageId="9" pageNumber="160">Leodice mexicana</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB52222FBC0FBA810ABFBDB" author="Fauchald, K." box="[1090,1272,1123,1149]" pageId="9" pageNumber="160" pagination="1 - 335" refId="ref13871" refString="Fauchald, K. (1970) Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico. Allan Hancock Monographs in Marine Biology, 5, 1 - 335." type="journal article" year="1970">Fauchald, 1970</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C3A9DBBFFB52222FAC6FBA815EDFB07" authority="(Fauchald, 1969)" baseAuthorityName="Fauchald" baseAuthorityYear="1969" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="websteri">
<emphasis id="B94E3A2AFFB52222FAC6FBA814A5FB07" italics="true" pageId="9" pageNumber="160">Leodice websteri</emphasis>
(
<bibRefCitation id="EFAB9BC9FFB52222FE86FB4C15E5FB07" author="Fauchald, K." box="[260,438,1159,1185]" pageId="9" pageNumber="160" pagination="1 - 15" refId="ref13824" refString="Fauchald, K. (1969) A revision of six species of the flavus-bidentatus group of Eunice (Eunicidae, Polychaeta). Smithsonian Contributions to Zoology, 6, 1 - 15. https: // doi. org / 10.5479 / si. 00810282.6" type="journal article" year="1969">Fauchald, 1969</bibRefCitation>
)
</taxonomicName>
. It is clearly distinct from last three species, differing at least in the shape of articulations of antennae and palps (
<tableCitation id="C6B8D383FFB52222FE1AFB6015A3FB63" box="[408,496,1195,1221]" captionStart="TABLE 2" captionStartId="8.[151,243,523,549]" captionText="TABLE 2. Comparison of morphological features among Leodice ivanildae sp. nov. and most similar species." pageId="9" pageNumber="160">Table 2</tableCitation>
).
<taxonomicName id="4C3A9DBBFFB52222FD86FB601698FB63" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[516,715,1195,1221]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">
<emphasis id="B94E3A2AFFB52222FD86FB601698FB63" box="[516,715,1195,1221]" italics="true" pageId="9" pageNumber="160">Leodice ivanildae</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFB52222FD50FB601778FB63" bold="true" box="[722,811,1195,1221]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A27D8751FFB52222FD50FB601778FB63" box="[722,811,1195,1221]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C3A9DBBFFB52222FC78FB6010D9FB63" baseAuthorityName="Ehlers" baseAuthorityYear="1887" box="[1018,1162,1195,1221]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="antillensis">
<emphasis id="B94E3A2AFFB52222FC78FB6010D9FB63" box="[1018,1162,1195,1221]" italics="true" pageId="9" pageNumber="160">L. antillensis</emphasis>
</taxonomicName>
. Both species also share articulated notopodial cirri along the whole body, 12 subacicular hooks per parapodium, and pectinate chaetae with both marginal teeth longer than the others. However, in
<taxonomicName id="4C3A9DBBFFB52222FCDFFB3817A2FAAB" baseAuthorityName="Ehlers" baseAuthorityYear="1887" box="[861,1009,1267,1293]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="antillensis">
<emphasis id="B94E3A2AFFB52222FCDFFB3817A2FAAB" box="[861,1009,1267,1293]" italics="true" pageId="9" pageNumber="160">L. antillensis</emphasis>
</taxonomicName>
antennae and palps have cylindrical, irregular articulations, pectinate branchiae are shorter than notopodial cirri and do not decrease in size from the middle of branchial distribution, and pectinate chaetae have flat blades. In
<taxonomicName id="4C3A9DBBFFB52222FC58FAF01033FAF3" authorityName="Souza &amp; Zapff &amp; Christoffersen &amp; Zanol" authorityYear="2024" box="[986,1120,1339,1365]" class="Polychaeta" family="Eunicidae" genus="Leodice" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="160" phylum="Annelida" rank="species" species="ivanildae" status="sp. nov.">
<emphasis id="B94E3A2AFFB52222FC58FAF01033FAF3" box="[986,1120,1339,1365]" italics="true" pageId="9" pageNumber="160">L. ivanildae</emphasis>
</taxonomicName>
<emphasis id="B94E3A2AFFB52222FBE5FAF01093FAF3" bold="true" box="[1127,1216,1339,1365]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="A27D8751FFB52222FBE5FAF01093FAF3" box="[1127,1216,1339,1365]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
palps and antennae articulations are moniliform, pectinate branchiae are longer than notopodial cirri where best developed, decreasing in length from the middle of branchial distribution, and pectinate chaetae have curved blades. Thus, this species is here considered new to science.
</paragraph>
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</treatment>
</document>

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@ -0,0 +1,112 @@
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<emphasis id="B93A930F514BFFF7FF651077FEA57702" bold="true" box="[151,262,1590,1616]" pageId="16" pageNumber="276">Material.</emphasis>
<materialsCitation id="3B264540514BFFF7FEE31077FDFF7727" collectingDate="2022-06-25" collectionCode="ZMUT" country="Iraq" latitude="35.429253" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.110886" pageId="16" pageNumber="276" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D514BFFF7FEE31077FEF67702" box="[273,341,1590,1616]" name="Iraq" pageId="16" pageNumber="276">IRAQ</collectingCountry>
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<collectingRegion id="498A81FF514BFFF7FE9A1077FD037703" box="[360,672,1590,1617]" country="Iraq" name="As Sulaymaniyah" pageId="16" pageNumber="276">Sulaymaniyah Governorate</collectingRegion>
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(
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), Sangaw, Zhallay Darband Vil., nr. Basarah bridge,
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<collectingDate id="EFB49035514BFFF7FE29101BFDFF7727" box="[475,604,1626,1653]" pageId="16" pageNumber="276" value="2022-06-25">25.06.2022</collectingDate>
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<emphasis id="B93A930F514BFFF7FF35103FFEC277CA" bold="true" box="[199,353,1662,1688]" pageId="16" pageNumber="276">Distribution.</emphasis>
<collectingCountry id="F3590F8D514BFFF7FE95103FFE1A77CA" box="[359,441,1662,1688]" name="Croatia" pageId="16" pageNumber="276">Croatia</collectingCountry>
to
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. New family, genus, and species records for
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<emphasis id="B93A930F514BFFF7FF6516D6FE9E71E0" bold="true" box="[151,317,151,178]" italics="true" pageId="16" pageNumber="276">Xysticus edax</emphasis>
(O. Pickard-Cambridge, 1872)
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<emphasis id="B93A930F514BFFF7FF351766FEE47013" bold="true" box="[199,327,295,321]" pageId="16" pageNumber="276">Records in</emphasis>
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<collectingCountry id="F3590F8D514BFFF7FEBC1766FE267013" box="[334,389,295,321]" name="Iraq" pageId="16" pageNumber="276">Iraq</collectingCountry>
.
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<collectingRegion id="498A81FF514BFFF7FE631766FE427013" box="[401,481,295,321]" country="Iraq" name="Dihok" pageId="16" pageNumber="276">Duhok</collectingRegion>
. New record for
</materialsCitation>
<collectingRegion id="498A81FF514BFFF7FD6C1766FCE17013" box="[670,834,295,321]" country="Iraq" name="Slemani" pageId="16" pageNumber="276">Sulaymaniyah</collectingRegion>
.
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<emphasis id="B93A930F514BFFF7FF35170AFEC27037" bold="true" box="[199,353,331,357]" pageId="16" pageNumber="276">Distribution.</emphasis>
<collectingCountry id="F3590F8D514BFFF7FE95170AFE1B7037" box="[359,440,331,357]" name="Turkey" pageId="16" pageNumber="276">Turkey</collectingCountry>
,
<collectingCountry id="F3590F8D514BFFF7FE31170AFDA77037" box="[451,516,331,357]" name="Israel" pageId="16" pageNumber="276">Israel</collectingCountry>
,
<collectingCountry id="F3590F8D514BFFF7FDE2170DFDE37034" box="[528,576,332,358]" name="Iraq" pageId="16" pageNumber="276">Iraq</collectingCountry>
, and
<collectingCountry id="F3590F8D514BFFF7FD88170DFD087034" box="[634,683,332,358]" name="Iran" pageId="16" pageNumber="276">Iran</collectingCountry>
.
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</document>

View file

@ -0,0 +1,217 @@
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Zamani &amp; Marusik
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,
<figureCitation id="137553985151FFEDFED3114EFEC2767B" box="[289,353,1807,1833]" captionStart="FIGURE 8" captionStartId="12.[152,255,1357,1381]" captionTargetBox="[151,1436,181,1333]" captionTargetId="figure-18@12.[151,1436,181,1333]" captionTargetPageId="12" captionText="FIGURE 8. SEM imagesof the male palp of Palpimanus garmiyanus sp. n.A, C, D distal part, ventral, prolateral and retrolateral; B, E, F embolic complex, ventral, retrolateral and prolateral. Scale bars: 0.1 mm. Courtesy of A.A. Fomichev." pageId="10" pageNumber="270">8AF</figureCitation>
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<collectingRegion id="498A81FF5151FFEDFD481116FC577620" box="[698,1012,1879,1906]" country="Iraq" name="As Sulaymaniyah" pageId="10" pageNumber="270">Sulaymaniyah Governorate</collectingRegion>
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, Hanara Vil.,
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<geoCoordinate id="EE7A29DA5151FFEDFEC5113AFE6C76C4" box="[311,463,1915,1942]" degrees="45" direction="east" minutes="14" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="58.8" value="45.24967">45°14'58.8&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFE29113AFDFA76C7" box="[475,601,1915,1941]" pageId="10" pageNumber="270" value="2022-03-24">
<collectingDate id="EFB490355151FFEDFE29113AFDFA76C7" box="[475,601,1915,1941]" pageId="10" pageNumber="270" value="2022-03-24">24.03.2022</collectingDate>
</date>
(
<collectorName id="26BB2ACB5151FFEDFD9A113AFCAA76C7" box="[616,777,1915,1941]" pageId="10" pageNumber="270">F.A. Khudhur</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF3511DEFCD4768F" pageId="10" pageNumber="270" type="etymology">
<paragraph id="8BF14F1D5151FFEDFF3511DEFCD4768F" blockId="10.[151,1437,1879,2013]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF3511DEFEED76EB" bold="true" box="[199,334,1951,1977]" pageId="10" pageNumber="270">Etymology.</emphasis>
The specific epithet refers to Garmiyan, a great warm land plain extending beyond the southwestern side of Qara Dagh mountain chain in
<collectingRegion id="498A81FF5151FFEDFDCE1182FCD0768F" box="[572,883,1987,2013]" country="Iraq" name="As Sulaymaniyah" pageId="10" pageNumber="270">Sulaymaniyah Governorate</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="DF311F955150FFECFF6A1102FA3B762D" pageId="11" pageNumber="271" startId="11.[152,255,1859,1883]" targetBox="[160,1409,194,1835]" targetPageId="11" targetType="figure">
<paragraph id="8BF14F1D5150FFECFF6A1102FA3B762D" blockId="11.[151,1436,1859,1919]" pageId="11" pageNumber="271">
<emphasis id="B93A930F5150FFECFF6A1102FEB87609" bold="true" box="[152,283,1859,1883]" pageId="11" pageNumber="271">FIGURE 7.</emphasis>
Habitus (BC), abdomen and posterior half of cephalothorax (A) and palp (DG) of the male of
<taxonomicName id="4C4E349E5150FFECFAD11102FEAC762D" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" class="Arachnida" family="Palpimanidae" genus="Palpimanus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="271" phylum="Arthropoda" rank="species" species="garmiyanus" status="sp. nov.">
<emphasis id="B93A930F5150FFECFAD11102FEAC762D" italics="true" pageId="11" pageNumber="271">Palpimanus garmiyanus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A2092E745150FFECFEE71126FEED762D" box="[277,334,1895,1919]" pageId="11" pageNumber="271" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, E ventral; B, F dorsal; C lateral; D retrolateral; G prolateral. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<caption id="DF311F955157FFEBFF6A130CFB5374DB" pageId="12" pageNumber="272" startId="12.[152,255,1357,1381]" targetBox="[151,1436,181,1333]" targetPageId="12" targetType="figure">
<paragraph id="8BF14F1D5157FFEBFF6A130CFB5374DB" blockId="12.[151,1437,1357,1417]" pageId="12" pageNumber="272">
<emphasis id="B93A930F5157FFEBFF6A130CFEB57437" bold="true" box="[152,278,1357,1381]" pageId="12" pageNumber="272">FIGURE 8.</emphasis>
SEM images of the male palp of
<taxonomicName id="4C4E349E5157FFEBFDAB130CFCED7437" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[601,846,1357,1381]" class="Arachnida" family="Palpimanidae" genus="Palpimanus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="272" phylum="Arthropoda" rank="species" species="garmiyanus" status="sp. nov.">
<emphasis id="B93A930F5157FFEBFDAB130CFCED7437" box="[601,846,1357,1381]" italics="true" pageId="12" pageNumber="272">Palpimanus garmiyanus</emphasis>
</taxonomicName>
<emphasis id="B93A930F5157FFEBFCA0130CFC2A7437" bold="true" box="[850,905,1357,1381]" pageId="12" pageNumber="272">
<taxonomicNameLabel id="A2092E745157FFEBFCA0130CFC2A7437" box="[850,905,1357,1381]" pageId="12" pageNumber="272" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A, C, D distal part, ventral, prolateral and retrolateral; B, E, F embolic complex, ventral, retrolateral and prolateral. Scale bars: 0.1 mm. Courtesy of A.A. Fomichev.
</paragraph>
</caption>
<subSubSection id="C3541C965157FFEBFF35139AFCCC774B" pageId="12" pageNumber="272" type="diagnosis">
<paragraph id="8BF14F1D5157FFEBFF35139AFCCC774B" blockId="12.[151,1437,1499,2030]" pageId="12" pageNumber="272">
<emphasis id="B93A930F5157FFEBFF35139AFEE174A7" bold="true" box="[199,322,1499,1525]" pageId="12" pageNumber="272">Diagnosis.</emphasis>
The new species differs from the congeners by lacking a tegular apophysis or membranous outgrowth and possessing an unbranched lamella-shaped embolus complex.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965157FFEAFF351062FEA371AB" lastPageId="13" lastPageNumber="273" pageId="12" pageNumber="272" type="description">
<paragraph id="8BF14F1D5157FFEBFF351062FA3477A0" blockId="12.[151,1437,1499,2030]" pageId="12" pageNumber="272">
<emphasis id="B93A930F5157FFEBFF351062FEFA776F" bold="true" box="[199,345,1571,1597]" pageId="12" pageNumber="272">Description.</emphasis>
Male. Habitus as in
<figureCitation id="137553985157FFEBFDC91062FD11776F" box="[571,690,1571,1597]" captionStart="FIGURE 7" captionStartId="11.[152,255,1859,1883]" captionTargetBox="[160,1409,194,1835]" captionTargetId="figure-35@11.[151,1436,181,1835]" captionTargetPageId="11" captionText="FIGURE 7. Habitus (BC), abdomen and posterior half of cephalothorax (A) and palp (DG) of the male of Palpimanus garmiyanus sp. n. A, E ventral; B, F dorsal; C lateral; D retrolateral; G prolateral. Scale bars: 0.2 mm, unless stated otherwise." pageId="12" pageNumber="272">Fig. 7BC</figureCitation>
. Total length 5.80. Carapace 2.80 long, 2.30 wide. Eye sizes: AME: 0.13, ALE: 0.08, PME: 0.06, PLE: 0.08. Carapace, labium, chelicerae, maxillae and sternum dark reddish, coated with white setae. Leg I red, legs IIIV yellowish-brown, without annulations. All metatarsi and tarsi IIIV with distinct ventral hair tufts. Abdomen dark beige, with scattered long dark setae and a ventral scutum, 2 long diagonal and 2 dot-like scutula (
<figureCitation id="137553985157FFEBFE6510F2FE52779F" box="[407,497,1715,1741]" captionStart="FIGURE 7" captionStartId="11.[152,255,1859,1883]" captionTargetBox="[160,1409,194,1835]" captionTargetId="figure-35@11.[151,1436,181,1835]" captionTargetPageId="11" captionText="FIGURE 7. Habitus (BC), abdomen and posterior half of cephalothorax (A) and palp (DG) of the male of Palpimanus garmiyanus sp. n. A, E ventral; B, F dorsal; C lateral; D retrolateral; G prolateral. Scale bars: 0.2 mm, unless stated otherwise." pageId="12" pageNumber="272">Fig. 7A</figureCitation>
). Spinnerets dark beige. Measurements of legs: I: 6.46 (2.04, 1.76, 1.45, 0.61, 0.60), II: 5.49 (1.65, 1.06, 1.27, 0.95, 0.56), III: 4.53 (1.38, 0.86, 1.01, 0.83, 0.45), IV: 6.24 (1.81, 1.11, 1.48, 1.30, 0.54).
</paragraph>
<paragraph id="8BF14F1D5157FFEBFF3510BAFE3976D3" blockId="12.[151,1437,1499,2030]" pageId="12" pageNumber="272">
Palp as in
<figureCitation id="137553985157FFEBFEB710BAFE647647" box="[325,455,1787,1813]" captionStart="FIGURE 7" captionStartId="11.[152,255,1859,1883]" captionTargetBox="[160,1409,194,1835]" captionTargetId="figure-35@11.[151,1436,181,1835]" captionTargetPageId="11" captionText="FIGURE 7. Habitus (BC), abdomen and posterior half of cephalothorax (A) and palp (DG) of the male of Palpimanus garmiyanus sp. n. A, E ventral; B, F dorsal; C lateral; D retrolateral; G prolateral. Scale bars: 0.2 mm, unless stated otherwise." pageId="12" pageNumber="272">Figs 7DG</figureCitation>
,
<figureCitation id="137553985157FFEBFE2410BAFDBA7647" box="[470,537,1787,1813]" captionStart="FIGURE 8" captionStartId="12.[152,255,1357,1381]" captionTargetBox="[151,1436,181,1333]" captionTargetId="figure-18@12.[151,1436,181,1333]" captionTargetPageId="12" captionText="FIGURE 8. SEM imagesof the male palp of Palpimanus garmiyanus sp. n.A, C, D distal part, ventral, prolateral and retrolateral; B, E, F embolic complex, ventral, retrolateral and prolateral. Scale bars: 0.1 mm. Courtesy of A.A. Fomichev." pageId="12" pageNumber="272">8AF</figureCitation>
; tibia slightly swollen, ca. 1.3 times longer than wide, 1.6 times shorter than cymbium; cymbium almost triangular in dorsal view, 2.6 times longer than wide; tegulum ovate, 1.6 times longer than wide, lacking any outgrowth; embolic complex an almost flat lamina, lacking any prongs, as long as tegulum, with a sinusoidal bend.
</paragraph>
<paragraph id="8BF14F1D5157FFEBFF3511CAFE3D76F7" blockId="12.[151,1437,1499,2030]" box="[199,414,1931,1957]" pageId="12" pageNumber="272">Female. Unknown.</paragraph>
<paragraph id="8BF14F1D5157FFEAFF3511EEFEA371AB" blockId="12.[151,1437,1499,2030]" lastBlockId="13.[151,1436,151,285]" lastPageId="13" lastPageNumber="273" pageId="12" pageNumber="272">
<emphasis id="B93A930F5157FFEBFF3511EEFEA5769B" bold="true" box="[199,262,1967,1993]" pageId="12" pageNumber="272">Note.</emphasis>
Previously, the family
<taxonomicName id="4C4E349E5157FFEBFDE411EEFD0D769B" authorityName="Thorell" authorityYear="1869" box="[534,686,1967,1993]" class="Arachnida" family="Palpimanidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="272" phylum="Arthropoda" rank="family">Palpimanidae</taxonomicName>
was known from
<collectingCountry id="F3590F8D5157FFEBFC7111F1FC137698" box="[899,944,1968,1994]" name="Iraq" pageId="12" pageNumber="272">Iraq</collectingCountry>
only by a single record of an unidentified species of
<taxonomicName id="4C4E349E5157FFEBFEF81192FD8B76BF" authority="Dufour, 1820" authorityName="Dufour" authorityYear="1820" box="[266,552,2003,2029]" class="Arachnida" family="Palpimanidae" genus="Palpimanus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B93A930F5157FFEBFEF81192FE3376BF" box="[266,400,2003,2029]" italics="true" pageId="12" pageNumber="272">Palpimanus</emphasis>
Dufour, 1820
</taxonomicName>
from
<collectingRegion id="498A81FF5157FFEBFD9B1192FD1676BF" box="[617,693,2003,2029]" country="Iraq" name="Dihok" pageId="12" pageNumber="272">Duhok</collectingRegion>
(
<bibRefCitation id="EFDF32EC5157FFEBFD301192FC1276BF" author="Fomichev, A. A. &amp; Marusik, Y. M. &amp; Koponen, S." box="[706,945,2003,2030]" pageId="12" pageNumber="272" pagination="329 - 339" refId="ref8361" refString="Fomichev, A. A., Marusik, Y. M. &amp; Koponen, S. (2018) New data on spiders (Arachnida: Araneae) of Iraq. Zoology in the Middle East, 64 (4), 329 - 339. https: // doi. org / 10.1080 / 09397140.2018.1484018" type="journal article" year="2018">
Fomichev
<emphasis id="B93A930F5157FFEBFCCB1195FCCB76BF" box="[825,872,2003,2029]" italics="true" pageId="12" pageNumber="272">et al</emphasis>
. 2018
</bibRefCitation>
). Most likely, the new species described here should be placed in a separate genus. However, due to the lack of a generic revision and the uncertain conformation of the male palp in the
<typeStatus id="54F5F1BF5156FFEAFE6916FAFE687187" box="[411,459,187,213]" pageId="13" pageNumber="273">type</typeStatus>
species,
<taxonomicName id="4C4E349E5156FFEAFDC016FAFC637187" authority="Dufour, 1820" authorityName="Dufour" authorityYear="1820" box="[562,960,187,213]" class="Arachnida" family="Palpimanidae" genus="Palpimanus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="273" phylum="Arthropoda" rank="species" species="gibbulus">
<emphasis id="B93A930F5156FFEAFDC016FAFCBC7187" box="[562,799,187,213]" italics="true" pageId="13" pageNumber="273">Palpimanus gibbulus</emphasis>
Dufour, 1820
</taxonomicName>
, we are unable to establish a new genus at this time.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965156FFEAFF351743FAC8704F" box="[199,1387,258,285]" pageId="13" pageNumber="273" type="distribution">
<paragraph id="8BF14F1D5156FFEAFF351743FAC8704F" blockId="13.[151,1436,151,285]" box="[199,1387,258,285]" pageId="13" pageNumber="273">
<emphasis id="B93A930F5156FFEAFF351743FEC2704E" bold="true" box="[199,353,258,284]" pageId="13" pageNumber="273">Distribution.</emphasis>
Known only from the
<typeStatus id="54F5F1BF5156FFEAFD921742FD33704F" box="[608,656,259,285]" pageId="13" pageNumber="273">type</typeStatus>
locality in
<collectingRegion id="498A81FF5156FFEAFCFD1742FBE5704F" box="[783,1094,259,285]" country="Iraq" name="As Sulaymaniyah" pageId="13" pageNumber="273">Sulaymaniyah Governorate</collectingRegion>
, Iraqi
<collectingRegion id="498A81FF5156FFEAFB7E1742FB59704F" box="[1164,1274,259,285]" country="Iran" name="Kordestan" pageId="13" pageNumber="273">Kurdistan</collectingRegion>
(
<figureCitation id="137553985156FFEAFAF81742FAFE704F" box="[1290,1373,259,285]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="13" pageNumber="273">Fig. 10</figureCitation>
).
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,119 @@
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<subSubSection id="C3541C965151FFEDFF65127EFD7A750B" box="[151,729,1087,1114]" pageId="10" pageNumber="270" type="nomenclature">
<paragraph id="8BF14F1D5151FFEDFF65127EFD7A750B" blockId="10.[151,729,1087,1114]" box="[151,729,1087,1114]" pageId="10" pageNumber="270">
<heading id="D0B9F8715151FFEDFF65127EFD7A750B" bold="true" box="[151,729,1087,1114]" fontSize="11" level="1" pageId="10" pageNumber="270" reason="1">
<taxonomicName id="4C4E349E5151FFEDFF65127EFD7A750B" ID-CoL="597PM" authority="O. Pickard-Cambridge, 1876" authorityName="O. Pickard-Cambridge" authorityYear="1876" box="[151,729,1087,1114]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="urbana">
<emphasis id="B93A930F5151FFEDFF65127EFD7A750B" bold="true" box="[151,729,1087,1114]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF65127EFECB7508" bold="true" box="[151,360,1087,1114]" italics="true" pageId="10" pageNumber="270">Trochosa urbana</emphasis>
O. Pickard-Cambridge, 1876
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF6512C6FDC37594" pageId="10" pageNumber="270" type="materials_examined">
<paragraph id="8BF14F1D5151FFEDFF6512C6FDC37594" blockId="10.[151,1436,1159,1294]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF6512C6FEA575F3" bold="true" box="[151,262,1159,1185]" pageId="10" pageNumber="270">Material.</emphasis>
<materialsCitation id="3B2645405151FFEDFEE112C6FDFF7594" collectingDate="2023-03-25" collectionCode="ZMUT" country="Iraq" latitude="34.8935" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.66389" pageId="10" pageNumber="270" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5151FFEDFEE112C6FEF475F3" box="[275,343,1159,1185]" name="Iraq" pageId="10" pageNumber="270">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5151FFEDFE9912C6FD0675F0" box="[363,677,1159,1186]" italics="true" pageId="10" pageNumber="270">
<collectingRegion id="498A81FF5151FFEDFE9912C6FD0675F0" box="[363,677,1159,1186]" country="Iraq" name="As Sulaymaniyah" pageId="10" pageNumber="270">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945151FFEDFD4812C6FD7E75F3" box="[698,733,1159,1185]" count="1" pageId="10" pageNumber="270" type="male">1♂</specimenCount>
(
<collectionCode id="ED5FD7D85151FFEDFD0312C6FCE575F3" box="[753,838,1159,1185]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="10" pageNumber="270">ZMUT</collectionCode>
), Garmyan, Maydan, Horen Vil., Hawasan stream,
<geoCoordinate id="EE7A29DA5151FFEDFF6512EAFE937594" box="[151,304,1195,1222]" degrees="34" direction="north" minutes="53" orientation="latitude" pageId="10" pageNumber="270" precision="1" seconds="36.6" value="34.8935">34°53'36.6&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5151FFEDFEC512EAFE737594" box="[311,464,1195,1222]" degrees="45" direction="east" minutes="39" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="50.0" value="45.66389">45°39'50.0&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFE2912EAFDFF7594" box="[475,604,1195,1222]" pageId="10" pageNumber="270" value="2023-03-25">
<collectingDate id="EFB490355151FFEDFE2912EAFDFF7594" box="[475,604,1195,1222]" pageId="10" pageNumber="270" value="2023-03-25">25.03.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF35128EFD30745C" pageId="10" pageNumber="270" type="distribution">
<paragraph id="8BF14F1D5151FFEDFF35128EFD30745C" blockId="10.[151,1436,1159,1294]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF35128EFEC275BB" bold="true" box="[199,353,1231,1257]" pageId="10" pageNumber="270">Distribution.</emphasis>
Previously known from North Africa,
<collectingCountry id="F3590F8D5151FFEDFCE3128EFCD375BB" box="[785,880,1231,1257]" name="Ethiopia" pageId="10" pageNumber="270">Ethiopia</collectingCountry>
to
<collectingCountry id="F3590F8D5151FFEDFC61128EFBAD75BB" box="[915,1038,1231,1257]" name="Seychelles" pageId="10" pageNumber="270">Seychelles</collectingCountry>
,
<collectingCountry id="F3590F8D5151FFEDFBEB128EFB0375BB" box="[1049,1184,1231,1257]" name="Madagascar" pageId="10" pageNumber="270">Madagascar</collectingCountry>
(Nosy Be),
<collectingCountry id="F3590F8D5151FFEDFADA128EFACA75BB" box="[1320,1385,1231,1257]" name="Israel" pageId="10" pageNumber="270">Israel</collectingCountry>
, and
<collectingCountry id="F3590F8D5151FFEDFF6512B5FF6B745C" box="[151,200,1268,1294]" name="Iran" pageId="10" pageNumber="270">Iran</collectingCountry>
. New genus and species records for
<collectingCountry id="F3590F8D5151FFEDFDAD12B5FD33745C" box="[607,656,1268,1294]" name="Iraq" pageId="10" pageNumber="270">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,116 @@
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<mods:title id="C3C78D9EF2A5C8CCFE24ECB3DE453A74">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
</mods:titleInfo>
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<mods:namePart id="228665C635164B446220E56043D06AD2">Zamani, Alireza</mods:namePart>
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<mods:namePart id="E998A726AC2574D23137E984D4365FB5">Khudhur, Farhad A.</mods:namePart>
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<mods:affiliation id="E4C20AD563EECCB3105FD62D3E2FB8AC">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="2E716445A94AEA01B3E0D7148F901FF4" type="email">qi.ude.lusvinu@ruhduhk.dahraf</mods:nameIdentifier>
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<mods:namePart id="11B14D462656767CCFD809361DBAF868">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="038BE5B2524F4FFE2006946040303410">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<subSubSection id="C3541C965151FFEDFF6513E6FD827493" box="[151,545,1447,1474]" pageId="10" pageNumber="270" type="nomenclature">
<paragraph id="8BF14F1D5151FFEDFF6513E6FD827493" blockId="10.[151,545,1447,1474]" box="[151,545,1447,1474]" pageId="10" pageNumber="270">
<heading id="D0B9F8715151FFEDFF6513E6FD827493" bold="true" box="[151,545,1447,1474]" fontSize="11" level="1" pageId="10" pageNumber="270" reason="1">
<taxonomicName id="4C4E349E5151FFEDFF6513E6FD827493" ID-CoL="4BGKC" authority="Latreille, 1806" authorityName="Latreille" authorityYear="1806" box="[151,545,1447,1474]" class="Arachnida" family="Oxyopidae" genus="Oxyopes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="B93A930F5151FFEDFF6513E6FD827493" bold="true" box="[151,545,1447,1474]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF6513E6FEC67490" bold="true" box="[151,357,1447,1474]" italics="true" pageId="10" pageNumber="270">Oxyopes lineatus</emphasis>
Latreille, 1806
</emphasis>
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<paragraph id="8BF14F1D5151FFEDFF6513AEFC35777F" blockId="10.[151,1357,1519,1618]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF6513AEFEA5775B" bold="true" box="[151,262,1519,1545]" pageId="10" pageNumber="270">Material.</emphasis>
<materialsCitation id="3B2645405151FFEDFEFF13AEFC30777F" collectingDate="2021-06-13" collectionCode="ZMUT" country="Iraq" county="Records in Iraq." latitude="35.161083" location="Halabja" longLatPrecision="1" longitude="46.11647" municipality="New" pageId="10" pageNumber="270" specimenCount="1" specimenCount-male="1" stateProvince="Baghdad">
<collectingCountry id="F3590F8D5151FFEDFEFF13AEFEF2775B" box="[269,337,1519,1545]" name="Iraq" pageId="10" pageNumber="270">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5151FFEDFE9313AEFDF17758" box="[353,594,1519,1546]" italics="true" pageId="10" pageNumber="270">Halabja Governorate</emphasis>
:
<specimenCount id="9D4884945151FFEDFD9313AEFD27775B" box="[609,644,1519,1545]" count="1" pageId="10" pageNumber="270" type="male">1♂</specimenCount>
(
<collectionCode id="ED5FD7D85151FFEDFD6113AEFD4B775B" box="[659,744,1519,1545]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="10" pageNumber="270">ZMUT</collectionCode>
), Hawar Vil.,
<geoCoordinate id="EE7A29DA5151FFEDFC7113AEFBBF7758" box="[899,1052,1519,1546]" degrees="35" direction="north" minutes="09" orientation="latitude" pageId="10" pageNumber="270" precision="1" seconds="39.9" value="35.161083">35°09'39.9&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5151FFEDFBD113AEFB1F7758" box="[1059,1212,1519,1546]" degrees="46" direction="east" minutes="06" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="59.3" value="46.11647">46°06'59.3&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFB3513AEFAEB775B" box="[1223,1352,1519,1545]" pageId="10" pageNumber="270" value="2021-06-13">
<collectingDate id="EFB490355151FFEDFB3513AEFAEB775B" box="[1223,1352,1519,1545]" pageId="10" pageNumber="270" value="2021-06-13">13.06.2021</collectingDate>
</date>
.
<emphasis id="B93A930F5151FFEDFF351052FE29777F" bold="true" box="[199,394,1555,1581]" pageId="10" pageNumber="270">
<collectingCounty id="629037915151FFEDFF351052FE29777F" box="[199,394,1555,1581]" pageId="10" pageNumber="270">Records in Iraq.</collectingCounty>
</emphasis>
<collectingRegion id="498A81FF5151FFEDFE631052FE56777F" box="[401,501,1555,1581]" country="Iraq" name="Baghdad" pageId="10" pageNumber="270">Baghdad</collectingRegion>
and
<collectingRegion id="498A81FF5151FFEDFDD91052FDD8777F" box="[555,635,1555,1581]" country="Iraq" name="Dihok" pageId="10" pageNumber="270">Dohuk</collectingRegion>
.
<collectingMunicipality id="6B95D5675151FFEDFD741055FD18777C" box="[646,699,1556,1582]" pageId="10" pageNumber="270">New</collectingMunicipality>
record for
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</materialsCitation>
.
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<emphasis id="B93A930F5151FFEDFF351076FEC27703" bold="true" box="[199,353,1591,1617]" pageId="10" pageNumber="270">Distribution.</emphasis>
West Palaearctic.
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View file

@ -0,0 +1,123 @@
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<heading id="D0B9F8715151FFEDFF65155EFDC3726B" bold="true" box="[151,608,799,826]" fontSize="11" level="1" pageId="10" pageNumber="270" reason="1">
<taxonomicName id="4C4E349E5151FFEDFF65155EFDC3726B" ID-CoL="4DJK2" authority="Tongiorgi, 1966" authorityName="Tongiorgi" authorityYear="1966" box="[151,608,799,826]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="aenigmatica">
<emphasis id="B93A930F5151FFEDFF65155EFDC3726B" bold="true" box="[151,608,799,826]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF65155EFE35726B" bold="true" box="[151,406,799,826]" italics="true" pageId="10" pageNumber="270">Pardosa aenigmatica</emphasis>
Tongiorgi, 1966
</emphasis>
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<emphasis id="B93A930F5151FFEDFF651526FEA572D3" bold="true" box="[151,262,871,897]" pageId="10" pageNumber="270">Material.</emphasis>
<materialsCitation id="3B2645405151FFEDFEE61526FEBA72F7" collectingDate="2022-03-24" collectionCode="ZMUT" country="Iraq" latitude="35.35908" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.47439" pageId="10" pageNumber="270" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5151FFEDFEE61526FEFB72D3" box="[276,344,871,897]" name="Iraq" pageId="10" pageNumber="270">IRAQ</collectingCountry>
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<collectingRegion id="498A81FF5151FFEDFE9F1526FD0B72D0" box="[365,680,871,898]" country="Iraq" name="As Sulaymaniyah" pageId="10" pageNumber="270">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945151FFEDFD4C1526FD4272D3" box="[702,737,871,897]" count="1" pageId="10" pageNumber="270" type="male">1♂</specimenCount>
(
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), Qara Dagh, Telzait,
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<geoCoordinate id="EE7A29DA5151FFEDFB0D1526FA3B72D0" box="[1279,1432,871,898]" degrees="45" direction="east" minutes="28" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="27.8" value="45.47439">45°28'27.8&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFF6515CAFEBA72F7" box="[151,281,907,933]" pageId="10" pageNumber="270" value="2022-03-24">
<collectingDate id="EFB490355151FFEDFF6515CAFEBA72F7" box="[151,281,907,933]" pageId="10" pageNumber="270" value="2022-03-24">24.03.2022</collectingDate>
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.
</paragraph>
<paragraph id="8BF14F1D5151FFEDFF3515EEFC92729B" blockId="10.[151,1436,871,1006]" box="[199,817,943,970]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF3515EEFEE4729B" bold="true" box="[199,327,943,969]" pageId="10" pageNumber="270">Records in</emphasis>
<materialsCitation id="3B2645405151FFEDFEBC15EEFD20729B" box="[334,643,943,970]" country="Iraq" location="Erbil" pageId="10" pageNumber="270" specimenCount="1" stateProvince="Hewler">
<emphasis id="B93A930F5151FFEDFEBC15EEFE29729B" bold="true" box="[334,394,943,969]" pageId="10" pageNumber="270">
<collectingCountry id="F3590F8D5151FFEDFEBC15EEFE26729B" box="[334,389,943,969]" name="Iraq" pageId="10" pageNumber="270">Iraq</collectingCountry>
.
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<collectingRegion id="498A81FF5151FFEDFE6315EEFE6E729B" box="[401,461,943,969]" country="Iraq" name="Hewler" pageId="10" pageNumber="270">Erbil</collectingRegion>
. New record for
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<collectingRegion id="498A81FF5151FFEDFD7B15EEFC8E729B" box="[649,813,943,969]" country="Iraq" name="Slemani" pageId="10" pageNumber="270">Sulaymaniyah</collectingRegion>
.
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<paragraph id="8BF14F1D5151FFEDFF351592FE5072BC" blockId="10.[151,1436,871,1006]" box="[199,499,979,1006]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF351592FEC272BF" bold="true" box="[199,353,979,1005]" pageId="10" pageNumber="270">Distribution.</emphasis>
<collectingCountry id="F3590F8D5151FFEDFE951592FE3972BF" box="[359,410,979,1005]" name="Italy" pageId="10" pageNumber="270">Italy</collectingCountry>
to
<collectingCountry id="F3590F8D5151FFEDFE4D1595FE5372BC" box="[447,496,980,1006]" name="Iran" pageId="10" pageNumber="270">Iran</collectingCountry>
.
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View file

@ -0,0 +1,145 @@
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<emphasis id="B93A930F5151FFEDFF65169EFE0E71A8" bold="true" box="[151,429,223,250]" italics="true" pageId="10" pageNumber="270">Alopecosa albofasciata</emphasis>
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<collectingRegion id="498A81FF5151FFEDFE9A1766FD027010" box="[360,673,295,322]" country="Iraq" name="As Sulaymaniyah" pageId="10" pageNumber="270">Sulaymaniyah Governorate</collectingRegion>
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(
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<collectingDate id="EFB490355151FFEDFF65170AFEBA7037" box="[151,281,331,357]" pageId="10" pageNumber="270" value="2022-02-04">02.04.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2645405151FFEDFED6170AFAB67037" box="[292,1301,331,358]" collectingDate="2022-08-03" collectionCode="ZMUT" country="Iraq" latitude="35.62419" location="Hawary Shar Park" longLatPrecision="1" longitude="45.458" municipality="Goyzha Mt." pageId="10" pageNumber="270" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Sulaymaniyah Governorate">
<specimenCount id="9D4884945151FFEDFED6170AFEE97037" box="[292,330,331,357]" count="2" pageId="10" pageNumber="270" type="male">2♂</specimenCount>
<specimenCount id="9D4884945151FFEDFEB8170AFEC97037" box="[330,362,331,357]" count="2" pageId="10" pageNumber="270" type="female">2♀</specimenCount>
(
<collectionCode id="ED5FD7D85151FFEDFE8B170AFE6D7037" box="[377,462,331,357]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="10" pageNumber="270">ZMUT</collectionCode>
),
<collectingMunicipality id="6B95D5675151FFEDFE13170AFDCA7034" box="[481,617,331,358]" pageId="10" pageNumber="270">Goyzha Mt.</collectingMunicipality>
,
<location id="8E9119C65151FFEDFD87170DFCE67037" LSID="urn:lsid:plazi:treatment:03E7FE0B5151FFEDFF65169EFD6270FF:8E9119C65151FFEDFD87170DFCE67037" box="[629,837,331,358]" country="Iraq" latitude="35.62419" longLatPrecision="1" longitude="45.458" municipality="Goyzha Mt." name="Hawary Shar Park" pageId="10" pageNumber="270" stateProvince="Sulaymaniyah Governorate">Hawary Shar Park</location>
,
<geoCoordinate id="EE7A29DA5151FFEDFCA2170AFC4B7034" box="[848,1000,331,358]" degrees="35" direction="north" minutes="37" orientation="latitude" pageId="10" pageNumber="270" precision="1" seconds="27.1" value="35.62419">35°37'27.1&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5151FFEDFC02170AFB2B7034" box="[1008,1160,331,358]" degrees="45" direction="east" minutes="27" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="28.8" value="45.458">45°27'28.8&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFB66170AFAB67037" box="[1172,1301,331,357]" pageId="10" pageNumber="270" value="2022-08-03">
<collectingDate id="EFB490355151FFEDFB66170AFAB67037" box="[1172,1301,331,357]" pageId="10" pageNumber="270" value="2022-08-03">08.03.2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14F1D5151FFEDFF35172EFB9A70DB" blockId="10.[151,1437,295,430]" box="[199,1081,367,394]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF35172EFEE470DB" bold="true" box="[199,327,367,393]" pageId="10" pageNumber="270">Records in</emphasis>
<materialsCitation id="3B2645405151FFEDFEBC172EFDE970DB" box="[334,586,367,393]" country="Iraq" location="Dhi Qar" pageId="10" pageNumber="270" specimenCount="1" stateProvince="Dhi Qar">
<emphasis id="B93A930F5151FFEDFEBC172EFE2970DB" bold="true" box="[334,394,367,393]" pageId="10" pageNumber="270">
<collectingCountry id="F3590F8D5151FFEDFEBC172EFE2670DB" box="[334,389,367,393]" name="Iraq" pageId="10" pageNumber="270">Iraq</collectingCountry>
.
</emphasis>
Basrah,
<collectingRegion id="498A81FF5151FFEDFE1F172EFDE970DB" box="[493,586,367,393]" country="Iraq" name="Dhi Qar" pageId="10" pageNumber="270">Dhi Qar</collectingRegion>
</materialsCitation>
, and
<collectingRegion id="498A81FF5151FFEDFD76172EFD7770DB" box="[644,724,367,393]" country="Iraq" name="Dihok" pageId="10" pageNumber="270">Duhok</collectingRegion>
. New record for
<collectingRegion id="498A81FF5151FFEDFC63172EFB9670DB" box="[913,1077,367,393]" country="Iraq" name="Slemani" pageId="10" pageNumber="270">Sulaymaniyah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF3517D2FD6270FF" box="[199,705,403,430]" pageId="10" pageNumber="270" type="distribution">
<paragraph id="8BF14F1D5151FFEDFF3517D2FD6270FF" blockId="10.[151,1437,295,430]" box="[199,705,403,430]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF3517D2FEC270FF" bold="true" box="[199,353,403,429]" pageId="10" pageNumber="270">Distribution.</emphasis>
Mediterranean to Central Asia.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,126 @@
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<mods:namePart id="40947F34959AB973D05F534875F712CD">Khudhur, Farhad A.</mods:namePart>
<mods:nameIdentifier id="92A4F238363745DF3E16FE748B079AD7" type="ORCID">0000-0001-5267-6334</mods:nameIdentifier>
<mods:affiliation id="7EAC17274330A4FFBC28C57FC4D8C96C">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<mods:affiliation id="340E998E0E0706B5BB24FE196409FFBA">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<treatment id="03E7FE0B5151FFEDFF6517BEFDC9739F" LSID="urn:lsid:plazi:treatment:03E7FE0B5151FFEDFF6517BEFDC9739F" httpUri="http://treatment.plazi.org/id/03E7FE0B5151FFEDFF6517BEFDC9739F" lastPageNumber="270" pageId="10" pageNumber="270">
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<paragraph id="8BF14F1D5151FFEDFF6517BEFDE7734B" blockId="10.[151,580,511,538]" box="[151,580,511,538]" pageId="10" pageNumber="270">
<heading id="D0B9F8715151FFEDFF6517BEFDE7734B" bold="true" box="[151,580,511,538]" fontSize="11" level="1" pageId="10" pageNumber="270" reason="1">
<taxonomicName id="4C4E349E5151FFEDFF6517BEFDE7734B" ID-CoL="GBJT" authority="Mcheidze, 1946" authorityName="Mcheidze" authorityYear="1946" box="[151,580,511,538]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="270" phylum="Arthropoda" rank="species" species="tbilisiensis">
<emphasis id="B93A930F5151FFEDFF6517BEFDE7734B" bold="true" box="[151,580,511,538]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF6517BEFED97348" bold="true" box="[151,378,511,538]" italics="true" pageId="10" pageNumber="270">Arctosa tbilisiensis</emphasis>
Mcheidze, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF651406FC7373FB" pageId="10" pageNumber="270" type="materials_examined">
<paragraph id="8BF14F1D5151FFEDFF651406FEBE73D4" blockId="10.[151,1436,583,718]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF651406FEA57333" bold="true" box="[151,262,583,609]" pageId="10" pageNumber="270">Material.</emphasis>
<materialsCitation id="3B2645405151FFEDFEEB1406FEBB73D4" collectingDate="2023-05-17" collectionCode="ZMUT" country="Iraq" latitude="36.697083" location="Erbil Governorate" longLatPrecision="1" longitude="44.87044" pageId="10" pageNumber="270" specimenCount="1" specimenCount-female="1" stateProvince="Arbil">
<collectingCountry id="F3590F8D5151FFEDFEEB1406FEFE7333" box="[281,349,583,609]" name="Iraq" pageId="10" pageNumber="270">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5151FFEDFE8B1406FDF67330" box="[377,597,583,610]" italics="true" pageId="10" pageNumber="270">
<collectingRegion id="498A81FF5151FFEDFE8B1406FDF67330" box="[377,597,583,610]" country="Iraq" name="Arbil" pageId="10" pageNumber="270">Erbil Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945151FFEDFD821406FD317333" box="[624,658,583,609]" count="1" pageId="10" pageNumber="270" type="female">1♀</specimenCount>
(
<collectionCode id="ED5FD7D85151FFEDFD5F1406FCA17333" box="[685,770,583,609]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="10" pageNumber="270">ZMUT</collectionCode>
), Choman Nawanda Vil.,
<geoCoordinate id="EE7A29DA5151FFEDFBA11406FB4F7330" box="[1107,1260,583,610]" degrees="36" direction="north" minutes="41" orientation="latitude" pageId="10" pageNumber="270" precision="1" seconds="49.5" value="36.697083">36°41'49.5&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5151FFEDFB0D1406FA3B7330" box="[1279,1432,583,610]" degrees="44" direction="east" minutes="52" orientation="longitude" pageId="10" pageNumber="270" precision="1" seconds="13.6" value="44.87044">44°52'13.6&quot;E</geoCoordinate>
,
<date id="FFF069DD5151FFEDFF65142AFEBB73D4" box="[151,280,619,646]" pageId="10" pageNumber="270" value="2023-05-17">
<collectingDate id="EFB490355151FFEDFF65142AFEBB73D4" box="[151,280,619,646]" pageId="10" pageNumber="270" value="2023-05-17">17.05.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14F1D5151FFEDFF3514CEFC7373FB" blockId="10.[151,1436,583,718]" box="[199,976,655,682]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF3514CEFEE473FB" bold="true" box="[199,327,655,681]" pageId="10" pageNumber="270">Records in</emphasis>
<materialsCitation id="3B2645405151FFEDFEBC14CEFDE973FB" box="[334,586,655,681]" country="Iraq" location="Dhi Qar" pageId="10" pageNumber="270" specimenCount="1" stateProvince="Dhi Qar">
<emphasis id="B93A930F5151FFEDFEBC14CEFE2973FB" bold="true" box="[334,394,655,681]" pageId="10" pageNumber="270">
<collectingCountry id="F3590F8D5151FFEDFEBC14CEFE2673FB" box="[334,389,655,681]" name="Iraq" pageId="10" pageNumber="270">Iraq</collectingCountry>
.
</emphasis>
Basrah,
<collectingRegion id="498A81FF5151FFEDFE1F14CEFDE973FB" box="[493,586,655,681]" country="Iraq" name="Dhi Qar" pageId="10" pageNumber="270">Dhi Qar</collectingRegion>
</materialsCitation>
, and
<collectingRegion id="498A81FF5151FFEDFD7614CEFD7773FB" box="[644,724,655,681]" country="Iraq" name="Dihok" pageId="10" pageNumber="270">Duhok</collectingRegion>
. New record for
<collectingRegion id="498A81FF5151FFEDFC6314CEFC6E73FB" box="[913,973,655,681]" country="Iraq" name="Hewler" pageId="10" pageNumber="270">Erbil</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965151FFEDFF3514F2FDC9739F" box="[199,618,691,718]" pageId="10" pageNumber="270" type="distribution">
<paragraph id="8BF14F1D5151FFEDFF3514F2FDC9739F" blockId="10.[151,1436,583,718]" box="[199,618,691,718]" pageId="10" pageNumber="270">
<emphasis id="B93A930F5151FFEDFF3514F2FEC2739F" bold="true" box="[199,353,691,717]" pageId="10" pageNumber="270">Distribution.</emphasis>
<collectingCountry id="F3590F8D5151FFEDFE9514F2FE15739F" box="[359,438,691,717]" name="Greece" pageId="10" pageNumber="270">Greece</collectingCountry>
to
<collectingCountry id="F3590F8D5151FFEDFE2B14F2FDC5739F" box="[473,614,691,717]" name="Afghanistan" pageId="10" pageNumber="270">Afghanistan</collectingCountry>
.
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,113 @@
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<emphasis id="B93A930F5152FFEEFF6510E2FE0677EF" bold="true" box="[151,421,1699,1726]" italics="true" pageId="9" pageNumber="269">Mesiotelus caucasicus</emphasis>
Zamani &amp; Marusik, 2021
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<emphasis id="B93A930F5152FFEEFF6510AAFEA57657" bold="true" box="[151,262,1771,1797]" pageId="9" pageNumber="269">Material.</emphasis>
<materialsCitation id="3B2645405152FFEEFEE210AAFEBA767B" collectingDate="2022-03-14" collectionCode="ZMUT" country="Iraq" latitude="35.450695" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.18203" pageId="9" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5152FFEEFEE210AAFEF77657" box="[272,340,1771,1797]" name="Iraq" pageId="9" pageNumber="269">IRAQ</collectingCountry>
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<collectingRegion id="498A81FF5152FFEEFE9410AAFD3E7654" box="[358,669,1771,1798]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
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(
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), Delezha, Chapachnara,
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<geoCoordinate id="EE7A29DA5152FFEEFB0D10AAFA3B7654" box="[1279,1432,1771,1798]" degrees="45" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="269" precision="1" seconds="55.3" value="45.18203">45°10'55.3&quot;E</geoCoordinate>
,
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.
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<emphasis id="B93A930F5152FFEEFF351172FEC2761F" bold="true" box="[199,353,1843,1869]" pageId="9" pageNumber="269">Distribution.</emphasis>
Previously known from
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<collectingCountry id="F3590F8D5152FFEEFC2F1175FBAD761C" box="[989,1038,1844,1870]" name="Iran" pageId="9" pageNumber="269">Iran</collectingCountry>
. New genus and species records for
<collectingCountry id="F3590F8D5152FFEEFF651119FF6B7620" box="[151,200,1880,1906]" name="Iraq" pageId="9" pageNumber="269">Iraq</collectingCountry>
.
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</document>

View file

@ -0,0 +1,182 @@
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Zamani &amp; Marusik
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<figureCitation id="137553985152FFEEFF65155EFF537268" box="[151,240,799,826]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="9" pageNumber="269">Figs 1D</figureCitation>
,
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<emphasis id="B93A930F5152FFEEFF651526FEE972D3" bold="true" box="[151,330,871,897]" pageId="9" pageNumber="269">Type material.</emphasis>
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<collectingRegion id="498A81FF5152FFEEFD451526FC5272D0" box="[695,1009,871,898]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
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<location id="8E9119C65152FFEEFB821529FB5C72D3" LSID="urn:lsid:plazi:treatment:03E7FE0B5152FFEEFF6514BAFAC8775B:8E9119C65152FFEEFB821529FB5C72D3" box="[1136,1279,871,898]" country="Iraq" latitude="35.49789" longLatPrecision="1" longitude="45.31561" municipality="Bakrajo" name="Hazar Merd" pageId="9" pageNumber="269" stateProvince="As Sulaymaniyah">Hazar Merd</location>
, Dekon Vill.,
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<geoCoordinate id="EE7A29DA5152FFEEFEC515CAFE6C72F4" box="[311,463,907,934]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="269" precision="1" seconds="56.2" value="45.31561">45°18'56.2&quot;E</geoCoordinate>
,
<date id="FFF069DD5152FFEEFE2915CAFDFA72F7" box="[475,601,907,933]" pageId="9" pageNumber="269" value="2022-03-28">
<collectingDate id="EFB490355152FFEEFE2915CAFDFA72F7" box="[475,601,907,933]" pageId="9" pageNumber="269" value="2022-03-28">28.03.2022</collectingDate>
</date>
(
<collectorName id="26BB2ACB5152FFEEFD9A15CAFCAA72F7" box="[616,777,907,933]" pageId="9" pageNumber="269">F.A. Khudhur</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3541C965152FFEEFF3515EEFB3672BF" pageId="9" pageNumber="269" type="etymology">
<paragraph id="8BF14F1D5152FFEEFF3515EEFB3672BF" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
<emphasis id="B93A930F5152FFEEFF3515EEFEED729B" bold="true" box="[199,334,943,969]" pageId="9" pageNumber="269">Etymology.</emphasis>
The specific epithet refers to Hazar Merd ancient archeological cave within Baranan mountain chain in
<collectingRegion id="498A81FF5152FFEEFF0A1592FD8D72BF" box="[248,558,979,1005]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
, near which the
<typeStatus id="54F5F1BF5152FFEEFD171595FCB672BC" box="[741,789,980,1006]" pageId="9" pageNumber="269">type</typeStatus>
locality of this species is situated.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965152FFEEFF3515B6FDD8752F" pageId="9" pageNumber="269" type="diagnosis">
<paragraph id="8BF14F1D5152FFEEFF3515B6FDD8752F" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
<emphasis id="B93A930F5152FFEEFF3515B6FEE17543" bold="true" box="[199,322,1015,1041]" pageId="9" pageNumber="269">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="4C4E349E5152FFEEFD5115B9FC4A7543" authority="Chatzaki, 2010" authorityName="Chatzaki" authorityYear="2010" box="[675,1001,1015,1042]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="269" phylum="Arthropoda" rank="species" species="helicoides">
<emphasis id="B93A930F5152FFEEFD5115B9FD107543" box="[675,691,1016,1041]" italics="true" pageId="9" pageNumber="269">Z</emphasis>
.
<emphasis id="B93A930F5152FFEEFD3015B6FC977543" box="[706,820,1015,1041]" italics="true" pageId="9" pageNumber="269">helicoides</emphasis>
<bibRefCitation id="EFDF32EC5152FFEEFCCF15B6FC4A7543" author="Chatzaki, M." box="[829,1001,1015,1041]" pageId="9" pageNumber="269" pagination="43 - 61" refId="ref8257" refString="Chatzaki, M. (2010) New data on the least known zelotines (Araneae, Gnaphosidae) of Greece and adjacent regions. Zootaxa, 2564 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 2564.1.3" type="journal article" year="2010">Chatzaki, 2010</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3590F8D5152FFEEFBC015B6FB227543" box="[1074,1153,1015,1041]" name="Greece" pageId="9" pageNumber="269">Greece</collectingCountry>
by having large anterior hoods, but can be distinguished by the relatively larger receptacles and converging lateral margins of the epigynal fovea (vs. subparallel), as well as by the receptacle span being wider than the anterior pockets (vs. subequal) (cf.
<figureCitation id="137553985152FFEEFF651222FEAF752F" box="[151,268,1123,1149]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="9" pageNumber="269">Fig. 6BC</figureCitation>
and
<bibRefCitation id="EFDF32EC5152FFEEFEB01222FE4A752F" author="Chatzaki, M." box="[322,489,1123,1149]" pageId="9" pageNumber="269" pagination="43 - 61" refId="ref8257" refString="Chatzaki, M. (2010) New data on the least known zelotines (Araneae, Gnaphosidae) of Greece and adjacent regions. Zootaxa, 2564 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 2564.1.3" type="journal article" year="2010">Chatzaki 2010</bibRefCitation>
: figs 22, 23).
</paragraph>
</subSubSection>
<subSubSection id="C3541C965152FFEEFF3512C6FE2674B7" pageId="9" pageNumber="269" type="description">
<paragraph id="8BF14F1D5152FFEEFF3512C6FAD47463" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
<emphasis id="B93A930F5152FFEEFF3512C6FEFA75F3" bold="true" box="[199,345,1159,1185]" pageId="9" pageNumber="269">Description.</emphasis>
Female. Habitus as in
<figureCitation id="137553985152FFEEFDA412C6FD0975F0" box="[598,682,1159,1186]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="9" pageNumber="269">Fig. 1D</figureCitation>
. Total length 5.70. Carapace 2.44 long, 1.95 wide. Eye sizes: AME: 0.08, ALE: 0.12, PME: 0.09, PLE: 0.08. Carapace, chelicerae, maxillae, labium and sternum black. Legs proximally black and distally light brown, without annulations. Abdomen dark gray, without any pattern. Spinnerets slightly darker than abdomen, uniform in color. Measurements of legs: I: Fe: 1.63, Pa: 1.04 (rest of segments missing), II: 4.65 (1.37, 0.86, 0.95, 0.80, 0.67), III: 4.44 (1.26, 0.76, 0.79, 0.99, 0.64), IV: 6.09 (1.73, 0.94, 1.22, 1.47, 0.73).
</paragraph>
<paragraph id="8BF14F1D5152FFEEFF35137AFF587493" blockId="9.[151,1437,871,1546]" pageId="9" pageNumber="269">
Epigyne as in
<figureCitation id="137553985152FFEEFE9A137AFE427407" box="[360,481,1339,1365]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="9" pageNumber="269">Fig. 6BC</figureCitation>
; epigynal plate ca. 1.3 times longer than wide; fovea (
<emphasis id="B93A930F5152FFEEFBB6137DFBC07407" box="[1092,1123,1340,1365]" italics="true" pageId="9" pageNumber="269">Fo</emphasis>
) ovate, ca. 1.4 times longer than wide; anterior hoods (
<emphasis id="B93A930F5152FFEEFE35131EFE45742B" box="[455,486,1375,1401]" italics="true" pageId="9" pageNumber="269">Ah</emphasis>
) broad, subequal to half width of fovea, spaced by less than their width; receptacles (
<emphasis id="B93A930F5152FFEEFF5313C5FF1C74CF" box="[161,191,1412,1437]" italics="true" pageId="9" pageNumber="269">Re</emphasis>
) large, globular, their diameters equal to width of fovea, spaced by less than one radius; accessorial glands (
<emphasis id="B93A930F5152FFEEFA8613C2FA3074CF" box="[1396,1427,1411,1437]" italics="true" pageId="9" pageNumber="269">Ag</emphasis>
) globular.
</paragraph>
<paragraph id="8BF14F1D5152FFEEFF35138AFE2674B7" blockId="9.[151,1437,871,1546]" box="[199,389,1483,1509]" pageId="9" pageNumber="269">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541C965152FFEEFF3513AEFAC8775B" box="[199,1387,1519,1546]" pageId="9" pageNumber="269" type="distribution">
<paragraph id="8BF14F1D5152FFEEFF3513AEFAC8775B" blockId="9.[151,1437,871,1546]" box="[199,1387,1519,1546]" pageId="9" pageNumber="269">
<emphasis id="B93A930F5152FFEEFF3513AEFEC2775B" bold="true" box="[199,353,1519,1545]" pageId="9" pageNumber="269">Distribution.</emphasis>
Known only from the
<typeStatus id="54F5F1BF5152FFEEFD9213B1FD337758" box="[608,656,1520,1546]" pageId="9" pageNumber="269">type</typeStatus>
locality in
<collectingRegion id="498A81FF5152FFEEFCFD13AEFBE5775B" box="[783,1094,1519,1545]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
, Iraqi
<collectingRegion id="498A81FF5152FFEEFB7E13AEFB59775B" box="[1164,1274,1519,1545]" country="Iran" name="Kordestan" pageId="9" pageNumber="269">Kurdistan</collectingRegion>
(
<figureCitation id="137553985152FFEEFAF813AEFAFE775B" box="[1290,1373,1519,1545]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="9" pageNumber="269">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,137 @@
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<mods:title id="7311FC05D6AA72594C197BF92D22B91A">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<heading id="D0B9F8715152FFEEFF6516D6FD7771E3" bold="true" box="[151,724,151,178]" fontSize="11" level="1" pageId="9" pageNumber="269" reason="1">
<taxonomicName id="4C4E349E5152FFEEFF6516D6FD7771E3" ID-CoL="98YTC" authority="(Kroneberg, 1875)" baseAuthorityName="Kroneberg" baseAuthorityYear="1875" box="[151,724,151,178]" class="Arachnida" family="Gnaphosidae" genus="Marinarozelotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="269" phylum="Arthropoda" rank="species" species="jaxartensis">
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<emphasis id="B93A930F5152FFEEFF6516D6FE4B71E3" bold="true" box="[151,488,151,178]" italics="true" pageId="9" pageNumber="269">Marinarozelotes jaxartensis</emphasis>
(Kroneberg, 1875)
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<emphasis id="B93A930F5152FFEEFF65169EFEA571AB" bold="true" box="[151,262,223,249]" pageId="9" pageNumber="269">Material.</emphasis>
<materialsCitation id="3B2645405152FFEEFEE1169EFE0B704C" collectingDate="2023-04-15" collectionCode="ZMUT" country="Iraq" latitude="35.50089" location="Sulaymaniyah Governorate" longLatPrecision="12" longitude="45.315277" pageId="9" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5152FFEEFEE1169EFEF471AB" box="[275,343,223,249]" name="Iraq" pageId="9" pageNumber="269">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5152FFEEFE9F169EFD0471A8" box="[365,679,223,250]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
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(
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), Bakrajo, Hazar Merd, Dekon Vil.,
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<geoCoordinate id="EE7A29DA5152FFEEFF651742FEB9704C" box="[151,282,259,286]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="9" pageNumber="269" precision="15" seconds="55" value="45.315277">45°18'55&quot;E</geoCoordinate>
,
<date id="FFF069DD5152FFEEFED41742FE0B704C" box="[294,424,259,286]" pageId="9" pageNumber="269" value="2023-04-15">
<collectingDate id="EFB490355152FFEEFED41742FE0B704C" box="[294,424,259,286]" pageId="9" pageNumber="269" value="2023-04-15">15.04.2023</collectingDate>
</date>
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.
</paragraph>
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<emphasis id="B93A930F5152FFEEFF351766FEE47013" bold="true" box="[199,327,295,321]" pageId="9" pageNumber="269">Records in</emphasis>
<materialsCitation id="3B2645405152FFEEFEBC1766FC507013" box="[334,1011,295,322]" country="Iraq" location="Baghdad" pageId="9" pageNumber="269" specimenCount="1" stateProvince="Baghdad">
<emphasis id="B93A930F5152FFEEFEBC1766FE297013" bold="true" box="[334,394,295,321]" pageId="9" pageNumber="269">
<collectingCountry id="F3590F8D5152FFEEFEBC1766FE267013" box="[334,389,295,321]" name="Iraq" pageId="9" pageNumber="269">Iraq</collectingCountry>
.
</emphasis>
<collectingRegion id="498A81FF5152FFEEFE631766FE567013" box="[401,501,295,321]" country="Iraq" name="Baghdad" pageId="9" pageNumber="269">Baghdad</collectingRegion>
and
<collectingRegion id="498A81FF5152FFEEFDD91766FD2B7013" box="[555,648,295,321]" country="Iraq" name="Dhi Qar" pageId="9" pageNumber="269">Dhi Qar</collectingRegion>
. New record for Iraqi Kurdistan
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.
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<emphasis id="B93A930F5152FFEEFF35170AFEC27037" bold="true" box="[199,353,331,357]" pageId="9" pageNumber="269">Distribution.</emphasis>
Northern Africa to Caucasus,
<collectingCountry id="F3590F8D5152FFEEFD4C170AFCAA7037" box="[702,777,331,357]" name="Russia" pageId="9" pageNumber="269">Russia</collectingCountry>
(Europe) to Central Asia, and Middle East. Introduced to
<collectingRegion id="498A81FF5152FFEEFF65172EFF4870DB" box="[151,235,367,393]" country="United States of America" name="Hawaii" pageId="9" pageNumber="269">Hawaii</collectingRegion>
,
<collectingCountry id="F3590F8D5152FFEEFF04172EFE9270DB" box="[246,305,367,393]" name="United States of America" pageId="9" pageNumber="269">USA</collectingCountry>
,
<collectingCountry id="F3590F8D5152FFEEFECE172EFE3770DB" box="[316,404,367,393]" name="Mexico" pageId="9" pageNumber="269">Mexico</collectingCountry>
,
<collectingCountry id="F3590F8D5152FFEEFE52172EFD9370DB" box="[416,560,367,393]" name="South Africa" pageId="9" pageNumber="269">South Africa</collectingCountry>
,
<collectingCountry id="F3590F8D5152FFEEFDCE172EFDDB70DB" box="[572,632,367,393]" name="India" pageId="9" pageNumber="269">India</collectingCountry>
,
<collectingCountry id="F3590F8D5152FFEEFD71172EFD6A70DB" box="[643,713,367,393]" name="China" pageId="9" pageNumber="269">China</collectingCountry>
, and
<collectingCountry id="F3590F8D5152FFEEFCF61731FCE470D8" box="[772,839,368,394]" name="Japan" pageId="9" pageNumber="269">Japan</collectingCountry>
.
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,117 @@
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<emphasis id="B93A930F5152FFEEFF65179AFEDF70A4" bold="true" box="[151,380,475,502]" italics="true" pageId="9" pageNumber="269">Pterotricha strandi</emphasis>
Spassky, 1936
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<emphasis id="B93A930F5152FFEEFF651462FEA5736F" bold="true" box="[151,262,547,573]" pageId="9" pageNumber="269">Material.</emphasis>
<materialsCitation id="3B2645405152FFEEFEE71462FDFF7333" collectingDate="2023-03-19" collectionCode="ZMUT" country="Iraq" latitude="35.429253" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.110886" pageId="9" pageNumber="269" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5152FFEEFEE71462FEFA736F" box="[277,345,547,573]" name="Iraq" pageId="9" pageNumber="269">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5152FFEEFE9D1462FD08736C" box="[367,683,547,574]" country="Iraq" name="As Sulaymaniyah" pageId="9" pageNumber="269">Sulaymaniyah Governorate</collectingRegion>
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:
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(
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), Sangaw, Zhallay Darband Vil., Basarah bridge,
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,
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<collectingDate id="EFB490355152FFEEFE291406FDFF7333" box="[475,604,583,609]" pageId="9" pageNumber="269" value="2023-03-19">19.03.2023</collectingDate>
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.
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<emphasis id="B93A930F5152FFEEFF35142AFEC273D7" bold="true" box="[199,353,619,645]" pageId="9" pageNumber="269">Distribution.</emphasis>
Previously known from
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,
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,
<collectingCountry id="F3590F8D5152FFEEFCA7142AFC4273D7" box="[853,993,619,645]" name="Afghanistan" pageId="9" pageNumber="269">Afghanistan</collectingCountry>
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<collectingCountry id="F3590F8D5152FFEEFBE8142AFBF473D7" box="[1050,1111,619,645]" name="India" pageId="9" pageNumber="269">India</collectingCountry>
. New species record for
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, with the current material representing the westernmost record of the species across the whole known range.
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@ -0,0 +1,118 @@
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<heading id="D0B9F8715153FFEFFF651064FC7C776D" bold="true" box="[151,991,1573,1600]" fontSize="11" level="1" pageId="8" pageNumber="268" reason="1">
<taxonomicName id="4C4E349E5153FFEFFF651064FC7C776D" ID-CoL="98YW4" authority="Zamani, Chatzaki, Esyunin &amp; Marusik, 2021" authorityName="Zamani, Chatzaki, Esyunin &amp; Marusik" authorityYear="2021" box="[151,991,1573,1600]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="qashqai">
<emphasis id="B93A930F5153FFEFFF651064FC7C776D" bold="true" box="[151,991,1573,1600]" pageId="8" pageNumber="268">
<emphasis id="B93A930F5153FFEFFF651064FE077712" bold="true" box="[151,420,1573,1600]" italics="true" pageId="8" pageNumber="268">Haplodrassus qashqai</emphasis>
Zamani, Chatzaki, Esyunin &amp; Marusik, 2021
</emphasis>
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<figureCitation id="137553985153FFEFFF65100BFF4C7736" box="[151,239,1610,1636]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="8" pageNumber="268">Figs 1E</figureCitation>
,
<figureCitation id="137553985153FFEFFF0E100BFEBD7736" box="[252,286,1610,1636]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="8" pageNumber="268">6A</figureCitation>
</paragraph>
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<paragraph id="8BF14F1D5153FFEFFF6510CAFE627798" blockId="8.[151,1437,1675,1810]" pageId="8" pageNumber="268">
<emphasis id="B93A930F5153FFEFFF6510CAFEA577F7" bold="true" box="[151,262,1675,1701]" pageId="8" pageNumber="268">Material.</emphasis>
<materialsCitation id="3B2645405153FFEFFEE110CAFE1E7798" collectingDate="2023-01-05" collectionCode="ZMUT" country="Iraq" latitude="34.77211" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.652195" pageId="8" pageNumber="268" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5153FFEFFEE110CAFEF477F7" box="[275,343,1675,1701]" name="Iraq" pageId="8" pageNumber="268">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5153FFEFFE9E10CAFD0477F7" box="[364,679,1675,1701]" italics="true" pageId="8" pageNumber="268">
<collectingRegion id="498A81FF5153FFEFFE9E10CAFD0477F7" box="[364,679,1675,1701]" country="Iraq" name="As Sulaymaniyah" pageId="8" pageNumber="268">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945153FFEFFD4910CAFD7D77F7" box="[699,734,1675,1701]" count="1" pageId="8" pageNumber="268" type="female">1♀</specimenCount>
(
<collectionCode id="ED5FD7D85153FFEFFD0110CAFCEB77F7" box="[755,840,1675,1701]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="8" pageNumber="268">ZMUT</collectionCode>
), Garmyan, Maydan, Gorgnosh Vil.,
<geoCoordinate id="EE7A29DA5153FFEFFAF110CAFA3F77F7" box="[1283,1436,1675,1701]" degrees="34" direction="north" minutes="46" orientation="latitude" pageId="8" pageNumber="268" precision="1" seconds="19.6" value="34.77211">34°46'19.6&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5153FFEFFF6510EEFE937798" box="[151,304,1711,1738]" degrees="45" direction="east" minutes="39" orientation="longitude" pageId="8" pageNumber="268" precision="1" seconds="07.9" value="45.652195">45°39'07.9&quot;E</geoCoordinate>
,
<date id="FFF069DD5153FFEFFEC910EEFE1E7798" box="[315,445,1711,1738]" pageId="8" pageNumber="268" value="2023-01-05">
<collectingDate id="EFB490355153FFEFFEC910EEFE1E7798" box="[315,445,1711,1738]" pageId="8" pageNumber="268" value="2023-01-05">01.05.2023</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965153FFEFFF351092FBC97640" pageId="8" pageNumber="268" type="distribution">
<paragraph id="8BF14F1D5153FFEFFF351092FBC97640" blockId="8.[151,1437,1675,1810]" pageId="8" pageNumber="268">
<emphasis id="B93A930F5153FFEFFF351092FEC277BF" bold="true" box="[199,353,1747,1773]" pageId="8" pageNumber="268">Distribution.</emphasis>
Previously known only from
<collectingCountry id="F3590F8D5153FFEFFD3D1095FCA377BC" box="[719,768,1748,1774]" name="Iran" pageId="8" pageNumber="268">Iran</collectingCountry>
. New species record for
<collectingCountry id="F3590F8D5153FFEFFBC61095FBC777BC" box="[1076,1124,1748,1774]" name="Iraq" pageId="8" pageNumber="268">Iraq</collectingCountry>
, with the current material representing the northwesternmost record of the species across the whole known range.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,109 @@
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<mods:title id="D5EED2F2E5F7C266597D7C53D44F0796">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:namePart id="882FC04B4A143DFCABC3840E82E2DCCA">Khudhur, Farhad A.</mods:namePart>
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<mods:affiliation id="C6EEA48F20B192FE966DECF076CC4CC3">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<mods:affiliation id="677C774D7FC6CD41592EA5A00E698D95">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<taxonomicName id="4C4E349E5153FFEFFF651114FD24763D" ID-CoL="3JMKS" authority="(C. L. Koch, 1839)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1839" box="[151,647,1877,1904]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="268" phylum="Arthropoda" rank="species" species="signifer">
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<emphasis id="B93A930F5153FFEFFF651114FE01763D" bold="true" box="[151,418,1877,1904]" italics="true" pageId="8" pageNumber="268">Haplodrassus signifer</emphasis>
(C.L. Koch, 1839)
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<emphasis id="B93A930F5153FFEFFF6511D6FEA576E3" bold="true" box="[151,262,1943,1969]" pageId="8" pageNumber="268">Material.</emphasis>
<materialsCitation id="3B2645405153FFEFFEE311D6FDFF7687" collectingDate="2023-03-19" collectionCode="ZMUT" country="Iraq" latitude="35.429253" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.110886" pageId="8" pageNumber="268" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5153FFEFFEE311D6FEF676E3" box="[273,341,1943,1969]" name="Iraq" pageId="8" pageNumber="268">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5153FFEFFE9A11D6FD0376E3" box="[360,672,1943,1969]" country="Iraq" name="As Sulaymaniyah" pageId="8" pageNumber="268">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
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(
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), Sangaw, Zhallay Darband Vil., nr. Basarah bridge,
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<geoCoordinate id="EE7A29DA5153FFEFFEC511FAFE737684" box="[311,464,1979,2006]" degrees="45" direction="east" minutes="06" orientation="longitude" pageId="8" pageNumber="268" precision="1" seconds="39.2" value="45.110886">45°06'39.2&quot;E</geoCoordinate>
,
<date id="FFF069DD5153FFEFFE2911FAFDFF7687" box="[475,604,1979,2005]" pageId="8" pageNumber="268" value="2023-03-19">
<collectingDate id="EFB490355153FFEFFE2911FAFDFF7687" box="[475,604,1979,2005]" pageId="8" pageNumber="268" value="2023-03-19">19.03.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="B93A930F5153FFEFFF35119EFEC276AB" bold="true" box="[199,353,2015,2041]" pageId="8" pageNumber="268">Distribution.</emphasis>
Circum-Holarctic. New species record for
<collectingCountry id="F3590F8D5153FFEFFCB711A1FCD576A8" box="[837,886,2016,2042]" name="Iraq" pageId="8" pageNumber="268">Iraq</collectingCountry>
.
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@ -0,0 +1,104 @@
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Trans-Palaearctic. New genus and species records for Iraq
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View file

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<emphasis id="B93A930F5154FFE8FF3510EDFEC27794" bold="true" box="[199,353,1708,1734]" pageId="15" pageNumber="275">Distribution.</emphasis>
<collectingCountry id="F3590F8D5154FFE8FE9510EDFE157794" box="[359,438,1708,1734]" name="Greece" pageId="15" pageNumber="275">Greece</collectingCountry>
to
<collectingCountry id="F3590F8D5154FFE8FE2810EDFD827794" box="[474,545,1708,1734]" name="China" pageId="15" pageNumber="275">China</collectingCountry>
.
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View file

@ -0,0 +1,118 @@
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<emphasis id="B93A930F5154FFE8FF65114AFEC07677" bold="true" box="[151,355,1802,1829]" italics="true" pageId="15" pageNumber="275">Xysticus acerbus</emphasis>
Thorell, 1872
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<emphasis id="B93A930F5154FFE8FF651131FEA576D8" bold="true" box="[151,262,1904,1930]" pageId="15" pageNumber="275">Material.</emphasis>
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(
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<emphasis id="B93A930F5154FFE8FF3511F9FEC27680" bold="true" box="[199,353,1976,2002]" pageId="15" pageNumber="275">Distribution.</emphasis>
Europe to Central Asia, and
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(Europe to Far East). New species record for
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<subSubSection id="C3541C965154FFE8FF651372FDF6741C" box="[151,597,1331,1358]" pageId="15" pageNumber="275" type="nomenclature">
<paragraph id="8BF14F1D5154FFE8FF651372FDF6741C" blockId="15.[151,597,1331,1358]" box="[151,597,1331,1358]" pageId="15" pageNumber="275">
<heading id="D0B9F8715154FFE8FF651372FDF6741C" bold="true" box="[151,597,1331,1358]" fontSize="11" level="1" pageId="15" pageNumber="275" reason="1">
<taxonomicName id="4C4E349E5154FFE8FF651372FDF6741C" ID-CoL="KV6S" authority="(Roewer, 1955)" baseAuthorityName="Roewer" baseAuthorityYear="1955" box="[151,597,1331,1358]" class="Arachnida" family="Thomisidae" genus="Bassaniodes" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="B93A930F5154FFE8FF651372FDF6741C" bold="true" box="[151,597,1331,1358]" pageId="15" pageNumber="275">
<emphasis id="B93A930F5154FFE8FF651372FE37741C" bold="true" box="[151,404,1331,1358]" italics="true" pageId="15" pageNumber="275">Bassaniodes loeffleri</emphasis>
(Roewer, 1955)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541C965154FFE8FF651334FE0874E6" pageId="15" pageNumber="275" type="materials_examined">
<paragraph id="8BF14F1D5154FFE8FF651334FE0874E6" blockId="15.[151,1436,1397,1496]" pageId="15" pageNumber="275">
<emphasis id="B93A930F5154FFE8FF651334FEA574DD" bold="true" box="[151,262,1397,1423]" pageId="15" pageNumber="275">Material.</emphasis>
<materialsCitation id="3B2645405154FFE8FEE11334FE0B74E6" collectingDate="2023-04-15" collectionCode="ZMUT" country="Iraq" latitude="35.50089" location="Sulaymaniyah Governorate" longLatPrecision="12" longitude="45.315277" pageId="15" pageNumber="275" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5154FFE8FEE11334FEF474DD" box="[275,343,1397,1423]" name="Iraq" pageId="15" pageNumber="275">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5154FFE8FE9F1334FD0474C2" box="[365,679,1397,1424]" italics="true" pageId="15" pageNumber="275">
<collectingRegion id="498A81FF5154FFE8FE9F1334FD0474C2" box="[365,679,1397,1424]" country="Iraq" name="As Sulaymaniyah" pageId="15" pageNumber="275">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945154FFE8FD4E1334FD7C74DD" box="[700,735,1397,1423]" count="1" pageId="15" pageNumber="275" type="female">1♀</specimenCount>
(
<collectionCode id="ED5FD7D85154FFE8FD061334FCEA74DD" box="[756,841,1397,1423]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="15" pageNumber="275">ZMUT</collectionCode>
), Bakrajo, Hazar Merd, Dekon Vil.,
<geoCoordinate id="EE7A29DA5154FFE8FAF11334FA3F74C2" box="[1283,1436,1397,1424]" degrees="35" direction="north" minutes="30" orientation="latitude" pageId="15" pageNumber="275" precision="1" seconds="03.2" value="35.50089">35°30'03.2&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5154FFE8FF6513D8FEB974E6" box="[151,282,1433,1460]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="15" pageNumber="275" precision="15" seconds="55" value="45.315277">45°18'55&quot;E</geoCoordinate>
,
<date id="FFF069DD5154FFE8FED413D8FE0B74E6" box="[294,424,1433,1460]" pageId="15" pageNumber="275" value="2023-04-15">
<collectingDate id="EFB490355154FFE8FED413D8FE0B74E6" box="[294,424,1433,1460]" pageId="15" pageNumber="275" value="2023-04-15">15.04.2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965154FFE8FF3513FCFC11748A" box="[199,946,1469,1496]" pageId="15" pageNumber="275" type="distribution">
<paragraph id="8BF14F1D5154FFE8FF3513FCFC11748A" blockId="15.[151,1436,1397,1496]" box="[199,946,1469,1496]" pageId="15" pageNumber="275">
<emphasis id="B93A930F5154FFE8FF3513FCFEC27485" bold="true" box="[199,353,1469,1495]" pageId="15" pageNumber="275">Distribution.</emphasis>
<collectingCountry id="F3590F8D5154FFE8FE9513FCFE157485" box="[359,438,1469,1495]" name="Greece" pageId="15" pageNumber="275">Greece</collectingCountry>
to Central Asia. New species record for
<collectingCountry id="F3590F8D5154FFE8FC8C13FFFC0C748A" box="[894,943,1470,1496]" name="Iraq" pageId="15" pageNumber="275">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,118 @@
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<mods:title id="F65244FD45955C9736964083B4DB98FF">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:namePart id="0F7A9E65C9423CD5811536AA258DA6CC">Khudhur, Farhad A.</mods:namePart>
<mods:nameIdentifier id="F6D8E0F9FEBF11CB7D46E43EB9E11649" type="ORCID">0000-0001-5267-6334</mods:nameIdentifier>
<mods:affiliation id="DAA4D15F95AE5F10D61152AAD2AF8D20">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="5887A7BF3207E8EF25C5FB18B376E96F" type="email">qi.ude.lusvinu@ruhduhk.dahraf</mods:nameIdentifier>
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<mods:affiliation id="01A3B71271B9CC979D6E8BA5F748C6C4">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<treatment id="03E7FE0B5154FFE8FF6514F3FC5772D3" LSID="urn:lsid:plazi:treatment:03E7FE0B5154FFE8FF6514F3FC5772D3" httpUri="http://treatment.plazi.org/id/03E7FE0B5154FFE8FF6514F3FC5772D3" lastPageNumber="275" pageId="15" pageNumber="275">
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<paragraph id="8BF14F1D5154FFE8FF6514F3FCA2739E" blockId="15.[151,769,690,753]" box="[151,769,690,717]" pageId="15" pageNumber="275">
<heading id="D0B9F8715154FFE8FF6514F3FCA2739E" bold="true" box="[151,769,690,717]" fontSize="11" level="1" pageId="15" pageNumber="275" reason="1">
<taxonomicName id="4C4E349E5154FFE8FF6514F3FCA2739E" ID-CoL="39WM5" authority="Bosmans &amp; Van Keer, 1999" authorityName="Bosmans &amp; Van Keer" authorityYear="1999" box="[151,769,690,717]" class="Arachnida" family="Theridiidae" genus="Enoplognatha" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="275" phylum="Arthropoda" rank="species" species="gemina">
<emphasis id="B93A930F5154FFE8FF6514F3FCA2739E" bold="true" box="[151,769,690,717]" pageId="15" pageNumber="275">
<emphasis id="B93A930F5154FFE8FF6514F3FE06739F" bold="true" box="[151,421,690,717]" italics="true" pageId="15" pageNumber="275">Enoplognatha gemina</emphasis>
Bosmans &amp; Van Keer, 1999
</emphasis>
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<figureCitation id="137553985154FFE8FF651496FF5373A3" box="[151,240,727,753]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="15" pageNumber="275">Figs 1C</figureCitation>
,
<figureCitation id="137553985154FFE8FF0F1496FEE373A3" box="[253,320,727,753]" captionStart="FIGURE 9" captionStartId="13.[152,255,1975,1999]" captionTargetBox="[166,1425,633,1952]" captionTargetId="figure-142@13.[151,1436,633,1952]" captionTargetPageId="13" captionText="FIGURE 9. Male palp of Enoplognatha gemina (AC) and Xysticus acerbus (EF), and intact epigyne of Rhysodromus rikhteri (D). A, D, E ventral; B, F retrolateral; C prolateral. Scale bars: 0.2 mm." pageId="15" pageNumber="275">9AC</figureCitation>
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<emphasis id="B93A930F5154FFE8FF65155FFEA5726A" bold="true" box="[151,262,798,824]" pageId="15" pageNumber="275">Material.</emphasis>
<materialsCitation id="3B2645405154FFE8FEE3155EFEBA720F" collectingDate="2022-03-28" collectionCode="ZMUT" country="Iraq" latitude="35.49789" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.31561" pageId="15" pageNumber="275" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5154FFE8FEE3155EFEF6726B" box="[273,341,799,825]" name="Iraq" pageId="15" pageNumber="275">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5154FFE8FE9A155EFD02726B" box="[360,673,799,825]" italics="true" pageId="15" pageNumber="275">
<collectingRegion id="498A81FF5154FFE8FE9A155EFD02726B" box="[360,673,799,825]" country="Iraq" name="As Sulaymaniyah" pageId="15" pageNumber="275">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
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(
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), Bakrajo, Hazar Merd,
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<geoCoordinate id="EE7A29DA5154FFE8FB0D155FFA3B726B" box="[1279,1432,798,825]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="15" pageNumber="275" precision="1" seconds="56.2" value="45.31561">45°18'56.2&quot;E</geoCoordinate>
,
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<collectingDate id="EFB490355154FFE8FF651502FEBA720F" box="[151,281,835,861]" pageId="15" pageNumber="275" value="2022-03-28">28.03.2022</collectingDate>
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.
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<emphasis id="B93A930F5154FFE8FF351527FEC272D2" bold="true" box="[199,353,870,896]" pageId="15" pageNumber="275">Distribution.</emphasis>
Mediterranean to
<collectingCountry id="F3590F8D5154FFE8FDDF1526FD0F72D3" box="[557,684,871,897]" name="Azerbaijan" pageId="15" pageNumber="275">Azerbaijan</collectingCountry>
. New species record for
<collectingCountry id="F3590F8D5154FFE8FC321526FC5272D3" box="[960,1009,871,897]" name="Iraq" pageId="15" pageNumber="275">Iraq</collectingCountry>
.
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View file

@ -0,0 +1,125 @@
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<emphasis id="B93A930F5154FFE8FF651593FE3172BF" bold="true" box="[151,402,978,1005]" italics="true" pageId="15" pageNumber="275">Steatoda triangulosa</emphasis>
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<emphasis id="B93A930F5154FFE8FF351223FEC2752E" bold="true" box="[199,353,1122,1148]" pageId="15" pageNumber="275">Distribution.</emphasis>
Trans-Palaearctic. Introduced to
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. New genus and species records for
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.
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View file

@ -0,0 +1,169 @@
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,
<location id="8E9119C65154FFE8FBE3172EFB5370DB" LSID="urn:lsid:plazi:treatment:03E7FE0B5154FFE8FF651766FBE7734B:8E9119C65154FFE8FBE3172EFB5370DB" box="[1041,1264,367,393]" country="Iraq" latitude="35.618526" longLatPrecision="1" longitude="45.450527" municipality="Goyzha Mt." name="Hawary Shar Park" pageId="15" pageNumber="275" stateProvince="As Sulaymaniyah">Hawary Shar Park</location>
,
<geoCoordinate id="EE7A29DA5154FFE8FAF1172FFA3F70DB" box="[1283,1436,366,393]" degrees="35" direction="north" minutes="37" orientation="latitude" pageId="15" pageNumber="275" precision="1" seconds="06.7" value="35.618526">35°37'06.7&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5154FFE8FF6517D3FE9370FF" box="[151,304,402,429]" degrees="45" direction="east" minutes="27" orientation="longitude" pageId="15" pageNumber="275" precision="1" seconds="01.9" value="45.450527">45°27'01.9&quot;E</geoCoordinate>
,
<date id="FFF069DD5154FFE8FECE17D2FE1D70FF" box="[316,446,403,429]" pageId="15" pageNumber="275" value="2022-03-19">
<collectingDate id="EFB490355154FFE8FECE17D2FE1D70FF" box="[316,446,403,429]" pageId="15" pageNumber="275" value="2022-03-19">19.03.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2645405154FFE8FE3917D2FA3B70FF" box="[459,1432,402,429]" collectingDate="2022-03-30" collectionCode="ZMUT" country="Iraq" latitude="35.615887" location="Hawary Shar Park" longLatPrecision="1" longitude="45.44639" municipality="Goyzha Mt." pageId="15" pageNumber="275" specimenCount="1" specimenCount-male="1" stateProvince="Sulaymaniyah Governorate">
<specimenCount id="9D4884945154FFE8FE3917D2FE4D70FF" box="[459,494,403,429]" count="1" pageId="15" pageNumber="275" type="male">1♂</specimenCount>
(
<collectionCode id="ED5FD7D85154FFE8FE0C17D2FDF070FF" box="[510,595,403,429]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="15" pageNumber="275">ZMUT</collectionCode>
),
<collectingMunicipality id="6B95D5675154FFE8FD9A17D2FD5270FF" box="[616,753,403,429]" pageId="15" pageNumber="275">Goyzha Mt.</collectingMunicipality>
,
<location id="8E9119C65154FFE8FD0C17D2FC7270FF" LSID="urn:lsid:plazi:treatment:03E7FE0B5154FFE8FF651766FBE7734B:8E9119C65154FFE8FD0C17D2FC7270FF" box="[766,977,403,429]" country="Iraq" latitude="35.615887" longLatPrecision="1" longitude="45.44639" municipality="Goyzha Mt." name="Hawary Shar Park" pageId="15" pageNumber="275" stateProvince="Sulaymaniyah Governorate">Hawary Shar Park</location>
,
<geoCoordinate id="EE7A29DA5154FFE8FC2C17D3FBD470FF" box="[990,1143,402,429]" degrees="35" direction="north" minutes="36" orientation="latitude" pageId="15" pageNumber="275" precision="1" seconds="57.2" value="35.615887">35°36'57.2&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5154FFE8FB8D17D3FABB70FF" box="[1151,1304,402,429]" degrees="45" direction="east" minutes="26" orientation="longitude" pageId="15" pageNumber="275" precision="1" seconds="47.0" value="45.44639">45°26'47.0&quot;E</geoCoordinate>
,
<date id="FFF069DD5154FFE8FAD617D2FA3B70FF" box="[1316,1432,403,429]" pageId="15" pageNumber="275" value="2022-03-30">
<collectingDate id="EFB490355154FFE8FAD617D2FA3B70FF" box="[1316,1432,403,429]" pageId="15" pageNumber="275" value="2022-03-30">30.3.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2645405154FFE8FF6517F6FBDF7083" box="[151,1148,438,465]" collectingDate="2022-04-16" collectionCode="ZMUT" country="Iraq" county="Gapilon" latitude="35.78311" location="Qulla Chakh" longLatPrecision="1" longitude="45.37561" municipality="Balkhe Vil." pageId="15" pageNumber="275" specimenCount="2" specimenCount-male="2" stateProvince="Sulaymaniyah Governorate">
<specimenCount id="9D4884945154FFE8FF6517F6FF197083" box="[151,186,439,465]" count="2" pageId="15" pageNumber="275" type="male">2♂</specimenCount>
(
<collectionCode id="ED5FD7D85154FFE8FF3A17F6FEBE7083" box="[200,285,439,465]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="15" pageNumber="275">ZMUT</collectionCode>
),
<collectingCounty id="629037915154FFE8FEDC17F6FE287083" box="[302,395,439,465]" pageId="15" pageNumber="275">Gapilon</collectingCounty>
,
<collectingMunicipality id="6B95D5675154FFE8FE6717F6FDB57083" box="[405,534,439,465]" pageId="15" pageNumber="275">Balkhe Vil.</collectingMunicipality>
,
<location id="8E9119C65154FFE8FDED17F6FD0C7083" LSID="urn:lsid:plazi:treatment:03E7FE0B5154FFE8FF651766FBE7734B:8E9119C65154FFE8FDED17F6FD0C7083" box="[543,687,439,465]" country="Iraq" county="Gapilon" latitude="35.78311" longLatPrecision="1" longitude="45.37561" municipality="Balkhe Vil." name="Qulla Chakh" pageId="15" pageNumber="275" stateProvince="Sulaymaniyah Governorate">Qulla Chakh</location>
,
<geoCoordinate id="EE7A29DA5154FFE8FD4B17F7FCF17083" box="[697,850,438,465]" degrees="35" direction="north" minutes="46" orientation="latitude" pageId="15" pageNumber="275" precision="1" seconds="59.2" value="35.78311">35°46'59.2&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5154FFE8FCA517F7FC537083" box="[855,1008,438,465]" degrees="45" direction="east" minutes="22" orientation="longitude" pageId="15" pageNumber="275" precision="1" seconds="32.2" value="45.37561">45°22'32.2&quot;E</geoCoordinate>
,
<date id="FFF069DD5154FFE8FC0817F6FBDF7083" box="[1018,1148,439,465]" pageId="15" pageNumber="275" value="2022-04-16">
<collectingDate id="EFB490355154FFE8FC0817F6FBDF7083" box="[1018,1148,439,465]" pageId="15" pageNumber="275" value="2022-04-16">16.04.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2645405154FFE8FB7717F6FD1570A7" collectingDate="2022-03-24" collectionCode="ZMUT" country="Iraq" latitude="35.35908" location="Telzait" longLatPrecision="1" longitude="45.47439" municipality="Qara Dagh" pageId="15" pageNumber="275" specimenCount="1" specimenCount-male="1" stateProvince="Sulaymaniyah Governorate">
<specimenCount id="9D4884945154FFE8FB7717F6FB0B7083" box="[1157,1192,439,465]" count="1" pageId="15" pageNumber="275" type="male">1♂</specimenCount>
(
<collectionCode id="ED5FD7D85154FFE8FB4417F6FAA87083" box="[1206,1291,439,465]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="15" pageNumber="275">ZMUT</collectionCode>
),
<collectingMunicipality id="6B95D5675154FFE8FAEF17F6FA3B7083" box="[1309,1432,439,465]" pageId="15" pageNumber="275">Qara Dagh</collectingMunicipality>
,
<location id="8E9119C65154FFE8FF65179AFF4570A7" LSID="urn:lsid:plazi:treatment:03E7FE0B5154FFE8FF651766FBE7734B:8E9119C65154FFE8FF65179AFF4570A7" box="[151,230,475,501]" country="Iraq" latitude="35.35908" longLatPrecision="1" longitude="45.47439" municipality="Qara Dagh" name="Telzait" pageId="15" pageNumber="275" stateProvince="Sulaymaniyah Governorate">Telzait</location>
,
<geoCoordinate id="EE7A29DA5154FFE8FF03179BFE2A70A7" box="[241,393,474,501]" degrees="35" direction="north" minutes="21" orientation="latitude" pageId="15" pageNumber="275" precision="1" seconds="32.7" value="35.35908">35°21'32.7&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5154FFE8FE63179BFD8A70A7" box="[401,553,474,501]" degrees="45" direction="east" minutes="28" orientation="longitude" pageId="15" pageNumber="275" precision="1" seconds="27.8" value="45.47439">45°28'27.8&quot;E</geoCoordinate>
,
<date id="FFF069DD5154FFE8FDC7179AFD1570A7" box="[565,694,475,501]" pageId="15" pageNumber="275" value="2022-03-24">
<collectingDate id="EFB490355154FFE8FDC7179AFD1570A7" box="[565,694,475,501]" pageId="15" pageNumber="275" value="2022-03-24">24.03.2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965154FFE8FF3517BFFBE7734B" box="[199,1092,510,537]" pageId="15" pageNumber="275" type="distribution">
<paragraph id="8BF14F1D5154FFE8FF3517BFFBE7734B" blockId="15.[151,1436,366,537]" box="[199,1092,510,537]" pageId="15" pageNumber="275">
<emphasis id="B93A930F5154FFE8FF3517BFFEC2734A" bold="true" box="[199,353,510,536]" pageId="15" pageNumber="275">Distribution.</emphasis>
<collectingCountry id="F3590F8D5154FFE8FE9517BEFE14734B" box="[359,439,511,537]" name="Turkey" pageId="15" pageNumber="275">Turkey</collectingCountry>
to
<collectingCountry id="F3590F8D5154FFE8FE2917BEFDDA734B" box="[475,633,511,537]" name="Turkmenistan" pageId="15" pageNumber="275">Turkmenistan</collectingCountry>
. New genus and species records for
<collectingCountry id="F3590F8D5154FFE8FBE217BEFBE2734B" box="[1040,1089,511,537]" name="Iraq" pageId="15" pageNumber="275">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,130 @@
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<mods:title id="4FCF99A032FA93D04B0718F14AFAA91D">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:namePart id="53C0273BCCE1951CDE702229C1E9B8CF">Khudhur, Farhad A.</mods:namePart>
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<mods:affiliation id="9E46BDD0B6785381A1B2C2F72845D5B6">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<paragraph id="8BF14F1D5155FFE9FF651130FCE876D8" blockId="14.[151,843,1904,1931]" box="[151,843,1904,1931]" pageId="14" pageNumber="274">
<heading id="D0B9F8715155FFE9FF651130FCE876D8" bold="true" box="[151,843,1904,1931]" fontSize="11" level="1" pageId="14" pageNumber="274" reason="1">
<taxonomicName id="4C4E349E5155FFE9FF651130FCE876D8" ID-CoL="3D9V8" authority="Moradmand &amp; Jager, 2012" authorityName="Moradmand &amp; Jager" authorityYear="2012" box="[151,843,1904,1931]" class="Arachnida" family="Sparassidae" genus="Eusparassus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="274" phylum="Arthropoda" rank="species" species="mesopotamicus">
<emphasis id="B93A930F5155FFE9FF651130FCE876D8" bold="true" box="[151,843,1904,1931]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF651130FE5176D8" bold="true" box="[151,498,1904,1931]" italics="true" pageId="14" pageNumber="274">Eusparassus mesopotamicus</emphasis>
Moradmand &amp; Jäger, 2012
</emphasis>
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<paragraph id="8BF14F1D5155FFE9FF6511F9FEBE76A4" blockId="14.[151,1436,1976,2038]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF6511F9FEA57680" bold="true" box="[151,262,1976,2002]" pageId="14" pageNumber="274">Material.</emphasis>
<materialsCitation id="3B2645405155FFE9FEFD11F8FEBA76A4" collectingDate="2022-06-10" collectionCode="ZMUT" country="Iraq" latitude="35.792442" location="Kanaru Town" longLatPrecision="1" longitude="45.579887" municipality="Chwarta" pageId="14" pageNumber="274" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5155FFE9FEFD11F8FEF07681" box="[271,339,1977,2003]" name="Iraq" pageId="14" pageNumber="274">IRAQ</collectingCountry>
:
<emphasis id="B93A930F5155FFE9FE9111F9FD397681" box="[355,666,1976,2003]" italics="true" pageId="14" pageNumber="274">
<collectingRegion id="498A81FF5155FFE9FE9111F9FD397681" box="[355,666,1976,2003]" country="Iraq" name="As Sulaymaniyah" pageId="14" pageNumber="274">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
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(
<collectionCode id="ED5FD7D85155FFE9FD2F11F8FC917681" box="[733,818,1977,2003]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="14" pageNumber="274">ZMUT</collectionCode>
),
<collectingMunicipality id="6B95D5675155FFE9FCB511F9FC0A7680" box="[839,937,1976,2002]" pageId="14" pageNumber="274">Chwarta</collectingMunicipality>
,
<location id="8E9119C65155FFE9FC4711F8FBF27681" LSID="urn:lsid:plazi:treatment:03E7FE0B5155FFE8FF651130FD6E7187:8E9119C65155FFE9FC4711F8FBF27681" box="[949,1105,1977,2003]" country="Iraq" latitude="35.792442" longLatPrecision="1" longitude="45.579887" municipality="Chwarta" name="Kanaru Town" pageId="14" pageNumber="274" stateProvince="As Sulaymaniyah">Kanaru Town</location>
,
<geoCoordinate id="EE7A29DA5155FFE9FBAC11F9FB547681" box="[1118,1271,1976,2003]" degrees="35" direction="north" minutes="47" orientation="latitude" pageId="14" pageNumber="274" precision="1" seconds="32.8" value="35.792442">35°47'32.8&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5155FFE9FB0D11F9FA3B7681" box="[1279,1432,1976,2003]" degrees="45" direction="east" minutes="34" orientation="longitude" pageId="14" pageNumber="274" precision="1" seconds="47.6" value="45.579887">45°34'47.6&quot;E</geoCoordinate>
,
<date id="FFF069DD5155FFE9FF65119DFEBA76A4" box="[151,281,2012,2038]" pageId="14" pageNumber="274" value="2022-06-10">
<collectingDate id="EFB490355155FFE9FF65119DFEBA76A4" box="[151,281,2012,2038]" pageId="14" pageNumber="274" value="2022-06-10">06.10.2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
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<emphasis id="B93A930F5154FFE8FF3516D6FEE471E2" bold="true" box="[199,327,150,177]" pageId="15" pageNumber="275">Records in</emphasis>
<materialsCitation id="3B2645405154FFE8FEBD16D6FC6C71E3" box="[335,975,151,177]" country="Iraq" location="Dhi Qar" pageId="15" pageNumber="275" specimenCount="1" stateProvince="Dhi Qar">
<emphasis id="B93A930F5154FFE8FEBD16D6FE2971E3" bold="true" box="[335,394,151,177]" pageId="15" pageNumber="275">
<collectingCountry id="F3590F8D5154FFE8FEBD16D6FE2571E3" box="[335,390,151,177]" name="Iraq" pageId="15" pageNumber="275">Iraq</collectingCountry>
.
</emphasis>
<collectingRegion id="498A81FF5154FFE8FE6316D6FE4F71E3" box="[401,492,151,177]" country="Iraq" name="Dhi Qar" pageId="15" pageNumber="275">Dhi Qar</collectingRegion>
and
<collectingRegion id="498A81FF5154FFE8FDD116D6FDC671E3" box="[547,613,151,177]" country="Iraq" name="An Najaf" pageId="15" pageNumber="275">Najaf</collectingRegion>
. New record for Iraqi Kurdistan
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.
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</subSubSection>
<subSubSection id="C3541C965154FFE8FF3516FBFD6E7187" box="[199,717,186,213]" pageId="15" pageNumber="275" type="distribution">
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<emphasis id="B93A930F5154FFE8FF3516FBFEC27186" bold="true" box="[199,353,186,212]" pageId="15" pageNumber="275">Distribution.</emphasis>
<collectingCountry id="F3590F8D5154FFE8FE9516FAFE347187" box="[359,407,187,213]" name="Iraq" pageId="15" pageNumber="275">Iraq</collectingCountry>
,
<collectingCountry id="F3590F8D5154FFE8FE5116FAFE707187" box="[419,467,187,213]" name="Iran" pageId="15" pageNumber="275">Iran</collectingCountry>
, and probably
<collectingCountry id="F3590F8D5154FFE8FD8516FAFD6A7187" box="[631,713,187,213]" name="Turkey" pageId="15" pageNumber="275">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,123 @@
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<taxonomicName id="4C4E349E5155FFE9FF651285FDEF758C" ID-CoL="6VRKY" authority="(Dalmas, 1920)" baseAuthorityName="Dalmas" baseAuthorityYear="1920" box="[151,588,1220,1247]" class="Arachnida" family="Salticidae" genus="Plexippoides" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="274" phylum="Arthropoda" rank="species" species="gestroi">
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<emphasis id="B93A930F5155FFE9FF651285FE2A758C" bold="true" box="[151,393,1220,1247]" italics="true" pageId="14" pageNumber="274">Plexippoides gestroi</emphasis>
(Dalmas, 1920)
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<emphasis id="B93A930F5155FFE9FF65134DFEA57474" bold="true" box="[151,262,1292,1318]" pageId="14" pageNumber="274">Material.</emphasis>
<materialsCitation id="3B2645405155FFE9FEE3134CFEBA7419" collectingDate="2022-02-04" collectionCode="ZMUT" country="Iraq" latitude="35.49542" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.336193" pageId="14" pageNumber="274" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5155FFE9FEE3134CFEF67475" box="[273,341,1293,1319]" name="Iraq" pageId="14" pageNumber="274">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5155FFE9FE9A134DFD027475" box="[360,673,1292,1319]" country="Iraq" name="As Sulaymaniyah" pageId="14" pageNumber="274">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<specimenCount id="9D4884945155FFE9FD41134CFD757475" box="[691,726,1293,1319]" count="1" pageId="14" pageNumber="274" type="female">1♀</specimenCount>
(
<collectionCode id="ED5FD7D85155FFE9FD1B134CFC9D7475" box="[745,830,1293,1319]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="14" pageNumber="274">ZMUT</collectionCode>
), Bakrajo, Hazar Merd,
<geoCoordinate id="EE7A29DA5155FFE9FBAE134DFB567475" box="[1116,1269,1292,1319]" degrees="35" direction="north" minutes="29" orientation="latitude" pageId="14" pageNumber="274" precision="1" seconds="43.5" value="35.49542">35°29'43.5&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5155FFE9FB0D134DFA3B7475" box="[1279,1432,1292,1319]" degrees="45" direction="east" minutes="20" orientation="longitude" pageId="14" pageNumber="274" precision="1" seconds="10.3" value="45.336193">45°20'10.3&quot;E</geoCoordinate>
,
<date id="FFF069DD5155FFE9FF651370FEBA7419" box="[151,281,1329,1355]" pageId="14" pageNumber="274" value="2022-02-04">
<collectingDate id="EFB490355155FFE9FF651370FEBA7419" box="[151,281,1329,1355]" pageId="14" pageNumber="274" value="2022-02-04">02.04.2022</collectingDate>
</date>
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.
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<paragraph id="8BF14F1D5155FFE9FF351314FCE5743C" blockId="14.[151,1437,1292,1427]" box="[199,838,1364,1391]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF351314FEE4743C" bold="true" box="[199,327,1364,1391]" pageId="14" pageNumber="274">Records in</emphasis>
<materialsCitation id="3B2645405155FFE9FEBC1314FD34743C" box="[334,663,1364,1391]" country="Iraq" location="Duhok" pageId="14" pageNumber="274" specimenCount="1" stateProvince="Dihok">
<emphasis id="B93A930F5155FFE9FEBC1314FE29743D" bold="true" box="[334,394,1365,1391]" pageId="14" pageNumber="274">
<collectingCountry id="F3590F8D5155FFE9FEBC1314FE26743D" box="[334,389,1365,1391]" name="Iraq" pageId="14" pageNumber="274">Iraq</collectingCountry>
.
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<collectingRegion id="498A81FF5155FFE9FE631315FE42743C" box="[401,481,1364,1390]" country="Iraq" name="Dihok" pageId="14" pageNumber="274">Duhok</collectingRegion>
. New record for
</materialsCitation>
<collectingRegion id="498A81FF5155FFE9FD6C1315FCE1743C" box="[670,834,1364,1390]" country="Iraq" name="Slemani" pageId="14" pageNumber="274">Sulaymaniyah</collectingRegion>
.
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<emphasis id="B93A930F5155FFE9FF351339FEC274C0" bold="true" box="[199,353,1400,1426]" pageId="14" pageNumber="274">Distribution.</emphasis>
<collectingCountry id="F3590F8D5155FFE9FE951338FE1574C1" box="[359,438,1401,1427]" name="Greece" pageId="14" pageNumber="274">Greece</collectingCountry>
to
<collectingCountry id="F3590F8D5155FFE9FE281338FDA874C1" box="[474,523,1401,1427]" name="Iran" pageId="14" pageNumber="274">Iran</collectingCountry>
.
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View file

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<emphasis id="B93A930F5155FFE9FF6513A5FDEA74AC" bold="true" box="[151,585,1508,1535]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF6513A5FED474AD" bold="true" box="[151,375,1508,1535]" italics="true" pageId="14" pageNumber="274">Plexippus paykulli</emphasis>
(Audouin, 1826)
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<emphasis id="B93A930F5155FFE9FF65106DFEA57714" bold="true" box="[151,262,1580,1606]" pageId="14" pageNumber="274">Material.</emphasis>
<materialsCitation id="3B2645405155FFE9FEE7106CFE1E7739" collectingDate="2023-07-04" collectionCode="ZMUT" country="Iraq" latitude="35.618526" location="Hawary Shar Park" longLatPrecision="1" longitude="45.450527" municipality="Goyzha Mt." pageId="14" pageNumber="274" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
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:
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<collectingRegion id="498A81FF5155FFE9FE82106DFD0E7715" box="[368,685,1580,1607]" country="Iraq" name="As Sulaymaniyah" pageId="14" pageNumber="274">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
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(
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),
<collectingMunicipality id="6B95D5675155FFE9FC9F106DFC5E7715" box="[877,1021,1580,1607]" pageId="14" pageNumber="274">Goyzha Mt.</collectingMunicipality>
,
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,
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<geoCoordinate id="EE7A29DA5155FFE9FF651011FE937739" box="[151,304,1616,1643]" degrees="45" direction="east" minutes="27" orientation="longitude" pageId="14" pageNumber="274" precision="1" seconds="01.9" value="45.450527">45°27'01.9&quot;E</geoCoordinate>
,
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<collectingDate id="EFB490355155FFE9FEC91010FE1E7739" box="[315,445,1617,1643]" pageId="14" pageNumber="274" value="2023-07-04">07.04.2023</collectingDate>
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.
</paragraph>
<paragraph id="8BF14F1D5155FFE9FF351034FBE877DC" blockId="14.[151,1436,1580,1751]" box="[199,1099,1652,1679]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF351034FEE477DC" bold="true" box="[199,327,1652,1679]" pageId="14" pageNumber="274">Records in</emphasis>
<materialsCitation id="3B2645405155FFE9FEBC1034FD2477DC" box="[334,647,1652,1679]" country="Iraq" location="Baghdad" pageId="14" pageNumber="274" specimenCount="1" stateProvince="Baghdad">
<emphasis id="B93A930F5155FFE9FEBC1034FE2977DD" bold="true" box="[334,394,1653,1679]" pageId="14" pageNumber="274">
<collectingCountry id="F3590F8D5155FFE9FEBC1034FE2677DD" box="[334,389,1653,1679]" name="Iraq" pageId="14" pageNumber="274">Iraq</collectingCountry>
.
</emphasis>
<collectingRegion id="498A81FF5155FFE9FE631035FE5477DC" box="[401,503,1652,1678]" country="Iraq" name="Baghdad" pageId="14" pageNumber="274">Baghdad</collectingRegion>
, Basrah, and
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<collectingRegion id="498A81FF5155FFE9FD7C1035FD7F77DC" box="[654,732,1652,1678]" country="Iraq" name="Diyala" pageId="14" pageNumber="274">Diyala</collectingRegion>
. New record for Iraqi Kurdistan.
</paragraph>
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<paragraph id="8BF14F1D5155FFE9FF3510D9FEA47785" blockId="14.[151,1436,1580,1751]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF3510D9FEC277E0" bold="true" box="[199,353,1688,1714]" pageId="14" pageNumber="274">Distribution.</emphasis>
Africa. Introduced to both Americas, Europe, Middle East,
<collectingCountry id="F3590F8D5155FFE9FBF410D9FBEF77E0" box="[1030,1100,1688,1714]" name="Nepal" pageId="14" pageNumber="274">Nepal</collectingCountry>
, southern Asia,
<collectingCountry id="F3590F8D5155FFE9FB0C10D9FACB77E0" box="[1278,1384,1688,1714]" name="Australia" pageId="14" pageNumber="274">Australia</collectingCountry>
, and Pacific Is.
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,145 @@
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<mods:title id="F2B2521DD23757286CB764D610C64E04">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:affiliation id="D5604D9F959B584C3C2C72E75A726DAC">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<emphasis id="B93A930F5155FFE9FF65143AFECC73C4" bold="true" box="[151,367,635,662]" italics="true" pageId="14" pageNumber="274">Evarcha insularis</emphasis>
(Metzner, 1999)
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<emphasis id="B93A930F5155FFE9FF6514FDFEA57384" bold="true" box="[151,262,700,726]" pageId="14" pageNumber="274">Material.</emphasis>
<materialsCitation id="3B2645405155FFE9FEE314FCFE6773A9" collectingDate="2021-04-05" collectionCode="ZMUT" country="Iraq" latitude="35.76461" location="Piramagroon Mt." longLatPrecision="1" longitude="45.24" municipality="Dukan" pageId="14" pageNumber="274" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5155FFE9FEE314FCFEF67385" box="[273,341,701,727]" name="Iraq" pageId="14" pageNumber="274">IRAQ</collectingCountry>
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<collectingRegion id="498A81FF5155FFE9FE9A14FDFD027385" box="[360,673,700,727]" country="Iraq" name="As Sulaymaniyah" pageId="14" pageNumber="274">Sulaymaniyah Governorate</collectingRegion>
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:
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(
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)
<collectingMunicipality id="6B95D5675155FFE9FCBC14FDFC3F7384" box="[846,924,700,726]" pageId="14" pageNumber="274">Dukan</collectingMunicipality>
,
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, Zewe Vil.,
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,
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;
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(
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),
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, Balkhe Vil.,
<location id="8E9119C65155FFE9FC5F14A1FBE573A8" LSID="urn:lsid:plazi:treatment:03E7FE0B5155FFE9FF65143AFDAC7235:8E9119C65155FFE9FC5F14A1FBE573A8" box="[941,1094,736,762]" country="Iraq" latitude="35.781166" longLatPrecision="1" longitude="45.374638" municipality="Gapilon" name="Qulla Chakh" pageId="14" pageNumber="274" stateProvince="Sulaymaniyah Governorate">Qulla Chakh</location>
,
<geoCoordinate id="EE7A29DA5155FFE9FBAA14A1FB5273A9" box="[1112,1265,736,763]" degrees="35" direction="north" minutes="46" orientation="latitude" pageId="14" pageNumber="274" precision="1" seconds="52.2" value="35.781166">35°46'52.2&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5155FFE9FB0D14A1FA3B73A9" box="[1279,1432,736,763]" degrees="45" direction="east" minutes="22" orientation="longitude" pageId="14" pageNumber="274" precision="1" seconds="28.7" value="45.374638">45°22'28.7&quot;E</geoCoordinate>
,
<date id="FFF069DD5155FFE9FF651544FEBB724D" box="[151,280,773,799]" pageId="14" pageNumber="274" value="2022-04-21">
<collectingDate id="EFB490355155FFE9FF651544FEBB724D" box="[151,280,773,799]" pageId="14" pageNumber="274" value="2022-04-21">21.04.2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14F1D5155FFE9FF351568FCE57210" blockId="14.[151,1436,700,871]" box="[199,838,808,835]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF351568FEE47210" bold="true" box="[199,327,808,835]" pageId="14" pageNumber="274">Records in</emphasis>
<materialsCitation id="3B2645405155FFE9FEBC1568FD347210" box="[334,663,808,835]" country="Iraq" location="Duhok" pageId="14" pageNumber="274" specimenCount="1" stateProvince="Dihok">
<emphasis id="B93A930F5155FFE9FEBC1568FE297211" bold="true" box="[334,394,809,835]" pageId="14" pageNumber="274">
<collectingCountry id="F3590F8D5155FFE9FEBC1568FE267211" box="[334,389,809,835]" name="Iraq" pageId="14" pageNumber="274">Iraq</collectingCountry>
.
</emphasis>
<collectingRegion id="498A81FF5155FFE9FE631569FE427210" box="[401,481,808,834]" country="Iraq" name="Dihok" pageId="14" pageNumber="274">Duhok</collectingRegion>
. New record for
</materialsCitation>
<collectingRegion id="498A81FF5155FFE9FD6C1569FCE17210" box="[670,834,808,834]" country="Iraq" name="Slemani" pageId="14" pageNumber="274">Sulaymaniyah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C965155FFE9FF35150DFDAC7235" box="[199,527,844,871]" pageId="14" pageNumber="274" type="distribution">
<paragraph id="8BF14F1D5155FFE9FF35150DFDAC7235" blockId="14.[151,1436,700,871]" box="[199,527,844,871]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF35150DFEC27234" bold="true" box="[199,353,844,870]" pageId="14" pageNumber="274">Distribution.</emphasis>
<collectingCountry id="F3590F8D5155FFE9FE95150CFE157235" box="[359,438,845,871]" name="Greece" pageId="14" pageNumber="274">Greece</collectingCountry>
to
<collectingCountry id="F3590F8D5155FFE9FE28150CFDA87235" box="[474,523,845,871]" name="Iran" pageId="14" pageNumber="274">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,124 @@
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<heading id="D0B9F8715155FFE9FF6515EAFDB57297" bold="true" box="[151,534,939,966]" fontSize="11" level="1" pageId="14" pageNumber="274" reason="1">
<taxonomicName id="4C4E349E5155FFE9FF6515EAFDB57297" ID-CoL="4G4SW" authority="(Poda, 1761)" baseAuthorityName="Poda" baseAuthorityYear="1761" box="[151,534,939,966]" class="Arachnida" family="Salticidae" genus="Philaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="274" phylum="Arthropoda" rank="species" species="chrysops">
<emphasis id="B93A930F5155FFE9FF6515EAFDB57297" bold="true" box="[151,534,939,966]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF6515EAFED17294" bold="true" box="[151,370,939,966]" italics="true" pageId="14" pageNumber="274">Philaeus chrysops</emphasis>
(Poda, 1761)
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<paragraph id="8BF14F1D5155FFE9FF6515B2FE637560" blockId="14.[151,1436,1011,1145]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF6515B2FEA5755F" bold="true" box="[151,262,1011,1037]" pageId="14" pageNumber="274">Material.</emphasis>
<materialsCitation id="3B2645405155FFE9FEE315B2FE1F7560" collectingDate="2021-04-05" collectionCode="ZMUT" country="Iraq" latitude="35.76461" location="Piramagroon Mt." longLatPrecision="1" longitude="45.24" municipality="Dukan" pageId="14" pageNumber="274" specimenCount="2" specimenCount-male="2" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D5155FFE9FEE315B2FEF6755F" box="[273,341,1011,1037]" name="Iraq" pageId="14" pageNumber="274">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5155FFE9FE9515B2FD3C755C" box="[359,671,1011,1038]" country="Iraq" name="As Sulaymaniyah" pageId="14" pageNumber="274">Sulaymaniyah Governorate</collectingRegion>
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:
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(
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),
<collectingMunicipality id="6B95D5675155FFE9FCA315B2FC3C755F" box="[849,927,1011,1037]" pageId="14" pageNumber="274">Dukan</collectingMunicipality>
,
<location id="8E9119C65155FFE9FC5F15B2FBD1755C" LSID="urn:lsid:plazi:treatment:03E7FE0B5155FFE9FF6515EAFDE4752B:8E9119C65155FFE9FC5F15B2FBD1755C" box="[941,1138,1011,1038]" country="Iraq" latitude="35.76461" longLatPrecision="1" longitude="45.24" municipality="Dukan" name="Piramagroon Mt." pageId="14" pageNumber="274" stateProvince="As Sulaymaniyah">Piramagroon Mt.</location>
, Zewe Vil.,
<geoCoordinate id="EE7A29DA5155FFE9FAF115B2FA3F755C" box="[1283,1436,1011,1038]" degrees="35" direction="north" minutes="45" orientation="latitude" pageId="14" pageNumber="274" precision="1" seconds="52.6" value="35.76461">35°45'52.6&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA5155FFE9FF651256FE8C7560" box="[151,303,1047,1074]" degrees="45" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="274" precision="1" seconds="24.0" value="45.24">45°14'24.0&quot;E</geoCoordinate>
,
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<collectingDate id="EFB490355155FFE9FEC91256FE1F7560" box="[315,444,1047,1074]" pageId="14" pageNumber="274" value="2021-04-05">04.05.2021</collectingDate>
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.
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<paragraph id="8BF14F1D5155FFE9FF35127AFC927507" blockId="14.[151,1436,1011,1145]" box="[199,817,1083,1110]" pageId="14" pageNumber="274">
<emphasis id="B93A930F5155FFE9FF35127AFEE47507" bold="true" box="[199,327,1083,1109]" pageId="14" pageNumber="274">Records in</emphasis>
<materialsCitation id="3B2645405155FFE9FEBC127AFD207507" box="[334,643,1083,1110]" country="Iraq" location="Erbil" pageId="14" pageNumber="274" specimenCount="1" stateProvince="Hewler">
<emphasis id="B93A930F5155FFE9FEBC127AFE297507" bold="true" box="[334,394,1083,1109]" pageId="14" pageNumber="274">
<collectingCountry id="F3590F8D5155FFE9FEBC127AFE267507" box="[334,389,1083,1109]" name="Iraq" pageId="14" pageNumber="274">Iraq</collectingCountry>
.
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<collectingRegion id="498A81FF5155FFE9FE63127AFE6E7507" box="[401,461,1083,1109]" country="Iraq" name="Hewler" pageId="14" pageNumber="274">Erbil</collectingRegion>
. New record for
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<collectingRegion id="498A81FF5155FFE9FD7B127AFC8E7507" box="[649,813,1083,1109]" country="Iraq" name="Slemani" pageId="14" pageNumber="274">Sulaymaniyah</collectingRegion>
.
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<emphasis id="B93A930F5155FFE9FF35121EFEC2752B" bold="true" box="[199,353,1119,1145]" pageId="14" pageNumber="274">Distribution.</emphasis>
Circum-Palaearctic.
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View file

@ -0,0 +1,178 @@
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<emphasis id="B93A930F5155FFE9FF65169EFECA71A8" bold="true" box="[151,361,223,250]" italics="true" pageId="14" pageNumber="274">Pisaura mirabilis</emphasis>
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<emphasis id="B93A930F5155FFE9FF651766FEA57013" bold="true" box="[151,262,295,321]" pageId="14" pageNumber="274">Material.</emphasis>
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,
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.
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Records in
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<collectingRegion id="498A81FF5155FFE9FE6317F6FE427083" box="[401,481,439,465]" country="Iraq" name="Dihok" pageId="14" pageNumber="274">Duhok</collectingRegion>
. New records for
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and
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.
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<emphasis id="B93A930F5155FFE9FF35179AFEC270A7" bold="true" box="[199,353,475,501]" pageId="14" pageNumber="274">Distribution.</emphasis>
Trans-Palaearctic.
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</document>

View file

@ -0,0 +1,111 @@
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<taxonomicName id="4C4E349E5156FFEAFF6517F7FD5B7082" ID-CoL="4SVNQ" authority="(Logunov &amp; Huseynov, 2008)" baseAuthorityName="Logunov &amp; Huseynov" baseAuthorityYear="2008" box="[151,760,438,465]" class="Arachnida" family="Philodromidae" genus="Rhysodromus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="273" phylum="Arthropoda" rank="species" species="rikhteri">
<emphasis id="B93A930F5156FFEAFF6517F7FD5B7082" bold="true" box="[151,760,438,465]" pageId="13" pageNumber="273">
<emphasis id="B93A930F5156FFEAFF6517F7FE377083" bold="true" box="[151,404,438,465]" italics="true" pageId="13" pageNumber="273">Rhysodromus rikhteri</emphasis>
(Logunov &amp; Huseynov, 2008)
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<figureCitation id="137553985156FFEAFF65179AFF5370A7" box="[151,240,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="13" pageNumber="273">Figs 1H</figureCitation>
,
<figureCitation id="137553985156FFEAFF0D179AFE8270A7" box="[255,289,475,501]" captionStart="FIGURE 9" captionStartId="13.[152,255,1975,1999]" captionTargetBox="[166,1425,633,1952]" captionTargetId="figure-142@13.[151,1436,633,1952]" captionTargetPageId="13" captionText="FIGURE 9. Male palp of Enoplognatha gemina (AC) and Xysticus acerbus (EF), and intact epigyne of Rhysodromus rikhteri (D). A, D, E ventral; B, F retrolateral; C prolateral. Scale bars: 0.2 mm." pageId="13" pageNumber="273">9D</figureCitation>
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<emphasis id="B93A930F5156FFEAFF651463FEA5736E" bold="true" box="[151,262,546,572]" pageId="13" pageNumber="273">Material.</emphasis>
<materialsCitation id="3B2645405156FFEAFEFF1462FA287333" collectingDate="2022-05-17" collectionCode="ZMUT" country="Iraq" latitude="36.58275" location="Erbil Governorate" longLatPrecision="1" longitude="44.99125" pageId="13" pageNumber="273" specimenCount="1" specimenCount-female="1" stateProvince="Arbil">
<collectingCountry id="F3590F8D5156FFEAFEFF1462FEF2736F" box="[269,337,547,573]" name="Iraq" pageId="13" pageNumber="273">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF5156FFEAFE931462FD93736F" box="[353,560,547,573]" country="Iraq" name="Arbil" pageId="13" pageNumber="273">Erbil Governorate</collectingRegion>
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:
<specimenCount id="9D4884945156FFEAFDCD1462FDC1736F" box="[575,610,547,573]" count="1" pageId="13" pageNumber="273" type="female">1♀</specimenCount>
(
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), Choman, Weze Vil.,
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<geoCoordinate id="EE7A29DA5156FFEAFB921463FB5A736F" box="[1120,1273,546,573]" degrees="44" direction="east" minutes="59" orientation="longitude" pageId="13" pageNumber="273" precision="1" seconds="28.5" value="44.99125">44°59'28.5&quot;E</geoCoordinate>
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<collectingDate id="EFB490355156FFEAFAF61463FA25736F" box="[1284,1414,546,573]" pageId="13" pageNumber="273" value="2022-05-17">17.05.2022</collectingDate>
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<emphasis id="B93A930F5156FFEAFF351407FEC27332" bold="true" box="[199,353,582,608]" pageId="13" pageNumber="273">Distribution.</emphasis>
Previously known only from Armenia and Azerbaijan. New genus and species records for Iraq
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,225 @@
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<mods:title id="95A4740420DC809B7A07040FA2A22DAE">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:namePart id="A63B56F3E87D0B593422EB83F0304599">Zamani, Alireza</mods:namePart>
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<mods:namePart id="DD6DE0F5BEB9633538914AEED5FF2F71">Khudhur, Farhad A.</mods:namePart>
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<mods:affiliation id="31881AB26742A562F08D0B6B81EC9EA5">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<taxonomicName id="4C4E349E515AFFE6FF651592FDC172BF" authority="Zamani &amp; Marusik" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[151,610,979,1005]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="261" phylum="Arthropoda" rank="species" species="sorani" status="sp. nov.">
<emphasis id="B93A930F515AFFE6FF651592FEC672BF" bold="true" box="[151,357,979,1005]" italics="true" pageId="1" pageNumber="261">Tegenaria sorani</emphasis>
Zamani &amp; Marusik
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<emphasis id="B93A930F515AFFE6FF65127EFEE5750B" bold="true" box="[151,326,1087,1113]" pageId="1" pageNumber="261">Type material.</emphasis>
<materialsCitation id="3B264540515AFFE6FEBC127EFDD2752F" collectingDate="2023-08-05" collectionCode="ZMUT" collectorName="F. A. Khudhur" country="Iraq" county="Soran" latitude="36.70828" location="Beston Cave" longLatPrecision="1" longitude="44.392365" municipality="Bradost Mt." pageId="1" pageNumber="261" specimenCount="1" specimenCount-male="1" stateProvince="Arbil" typeStatus="holotype">
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(
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),
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:
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<collectingRegion id="498A81FF515AFFE6FD6D127EFCD37508" box="[671,880,1087,1114]" country="Iraq" name="Arbil" pageId="1" pageNumber="261">Erbil Governorate</collectingRegion>
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</materialsCitation>
.
<materialsCitation id="3B264540515AFFE6FD891225FB05752F" box="[635,1190,1123,1150]" collectionCode="ZMUT" country="Iraq" pageId="1" pageNumber="261" specimenCount="2" specimenCount-male="2" stateProvince="Arbil" typeStatus="paratype">
<typeStatus id="54F5F1BF515AFFE6FD891225FD4F752C" box="[635,748,1124,1150]" pageId="1" pageNumber="261" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D488494515AFFE6FD041222FCBA752F" box="[758,793,1123,1149]" count="2" pageId="1" pageNumber="261" type="male">2♂</specimenCount>
(
<collectionCode id="ED5FD7D8515AFFE6FCDA1222FCDE752F" box="[808,893,1123,1149]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="1" pageNumber="261">ZMUT</collectionCode>
), same as for the
<typeStatus id="54F5F1BF515AFFE6FBB21222FB05752F" box="[1088,1190,1123,1149]" pageId="1" pageNumber="261" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515AFFE6FF3512C6FD9675B8" pageId="1" pageNumber="261" type="etymology">
<paragraph id="8BF14F1D515AFFE6FF3512C6FD9675B8" blockId="1.[151,1437,1087,2014]" pageId="1" pageNumber="261">
<emphasis id="B93A930F515AFFE6FF3512C6FEED75F3" bold="true" box="[199,334,1159,1185]" pageId="1" pageNumber="261">Etymology.</emphasis>
The specific epithet is a noun in apposition, referring to a dialect of the Kurdish language that is spoken in the most parts of Iraqi
<collectingRegion id="498A81FF515AFFE6FDEB12EAFD297597" box="[537,650,1195,1221]" country="Iran" name="Kordestan" pageId="1" pageNumber="261">Kurdistan</collectingRegion>
, as well as the
<collectingRegion id="498A81FF515AFFE6FCCC12EAFBB87597" box="[830,1051,1195,1221]" country="Iran" name="Kordestan" pageId="1" pageNumber="261">Kurdistan Province</collectingRegion>
,
<collectingRegion id="498A81FF515AFFE6FBDB12EAFA857597" box="[1065,1318,1195,1221]" country="Iran" name="Kermanshah" pageId="1" pageNumber="261">Kermanshah Province</collectingRegion>
, and
<collectingRegion id="498A81FF515AFFE6FA9712EDFEDE75BB" country="Iran" name="Azarbayjan-e Gharbi" pageId="1" pageNumber="261">West Azerbaijan Province</collectingRegion>
of western
<collectingCountry id="F3590F8D515AFFE6FDF21291FD9275B8" box="[512,561,1232,1258]" name="Iran" pageId="1" pageNumber="261">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515AFFE6FF3512B2FBBC7493" pageId="1" pageNumber="261" type="diagnosis">
<paragraph id="8BF14F1D515AFFE6FF3512B2FBBC7493" blockId="1.[151,1437,1087,2014]" pageId="1" pageNumber="261">
<emphasis id="B93A930F515AFFE6FF3512B2FEE1745F" bold="true" box="[199,322,1267,1293]" pageId="1" pageNumber="261">Diagnosis.</emphasis>
The new species is most similar to
<taxonomicName id="4C4E349E515AFFE6FD2512B5FBB2745C" authority="(Clerck, 1757)" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[727,1041,1267,1294]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="261" phylum="Arthropoda" rank="species" species="domestica">
<emphasis id="B93A930F515AFFE6FD2512B5FCC5745F" box="[727,870,1267,1293]" italics="true" pageId="1" pageNumber="261">T. domestica</emphasis>
(Clerck, 1757)
</taxonomicName>
and
<taxonomicName id="4C4E349E515AFFE6FBBB12B5FE327463" authority="Zamani, Marusik &amp; Malek-Hosseini, 2018" authorityName="Zamani, Marusik &amp; Malek-Hosseini" authorityYear="2018" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="261" phylum="Arthropoda" rank="species" species="alamto">
<emphasis id="B93A930F515AFFE6FBBB12B5FBFA745F" box="[1097,1113,1268,1293]" italics="true" pageId="1" pageNumber="261">T</emphasis>
.
<emphasis id="B93A930F515AFFE6FB9512B2FB16745F" box="[1127,1205,1267,1293]" italics="true" pageId="1" pageNumber="261">alamto</emphasis>
<bibRefCitation id="EFDF32EC515AFFE6FB4F12B2FE327463" author="Zamani, A. &amp; Marusik, Y. M. &amp; Malek-Hosseini, M. J." pageId="1" pageNumber="261" pagination="95 - 97" refId="ref8577" refString="Zamani, A., Marusik, Y. M. &amp; Malek-Hosseini, M. J. (2018) A new species of Tegenaria Latreille, 1804 (Araneae: Agelenidae) from western Iran. Zootaxa, 4444 (1), 95 - 97. https: // doi. org / 10.11646 / zootaxa. 4444.1.7" type="journal article" year="2018">
Zamani, Marusik &amp; Malek-Hosseini,
<quantity id="4CB6E2F8515AFFE6FEAB1356FE327463" box="[345,401,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="1" pageNumber="261" unit="in" value="2018.0">2018</quantity>
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</taxonomicName>
in having two retrolateral apophyses, a relatively short conductor with only the retrolateral arm developed, and a short embolus. It can be distinguished by the total absence of the prolateral arm of the conductor (vs. a small extension present), a thin embolus originating at ca. 10 oclock position (vs. the embolus in
<taxonomicName id="4C4E349E515AFFE6FF6513C5FE8674CF" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[151,293,1411,1437]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="261" phylum="Arthropoda" rank="species" species="domestica">
<emphasis id="B93A930F515AFFE6FF6513C5FE8674CF" box="[151,293,1411,1437]" italics="true" pageId="1" pageNumber="261">T. domestica</emphasis>
</taxonomicName>
thick, and in both species originating at 9 oclock position), and the aspinose palpal tibia (vs. spinose) (cf.
<figureCitation id="13755398515AFFE6FF3613E6FEBB7493" box="[196,280,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[152,255,1818,1842]" captionTargetBox="[157,1430,181,1794]" captionTargetId="figure-35@3.[151,1436,181,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male palp of Tegenaria sorani sp. n. (AD) and Cheiracanthium montanum (E, F). A, E ventral; B prolateral; C, F retrolateral; D dorsal. Abbreviations: Co—conductor, Cr—retrolateral arm of conductor, Ta—tegular apophysis, Ts—heavily sclerotized retrolateral tibial apophysis, Tw—weakly sclerotized retrolateral tibial apophysis. Scale bars: 0.2 mm." pageId="1" pageNumber="261">Fig. 2A</figureCitation>
and
<bibRefCitation id="EFDF32EC515AFFE6FEBD13E6FD9C7493" author="Cazanove, G. &amp; Cressent, P. - H. &amp; Tamon, J. - M. &amp; Derepas, B." box="[335,575,1447,1474]" pageId="1" pageNumber="261" pagination="30 - 35" refId="ref8193" refString="Cazanove, G., Cressent, P. - H., Tamon, J. - M. &amp; Derepas, B. (2022) Le cas de Agelena borbonica Vinson, 1863 a La Reunion: un nouveau synonyme de Tegenaria domestica (Clerck, 1758). Revue Arachnologique, Series 2, 9, 30 - 35." type="journal article" year="2022">
Cazanove
<emphasis id="B93A930F515AFFE6FE3613E9FE567493" box="[452,501,1447,1473]" italics="true" pageId="1" pageNumber="261">et al</emphasis>
. 2022
</bibRefCitation>
: fig. 2A, D and
<bibRefCitation id="EFDF32EC515AFFE6FD0013E6FC6A7493" author="Zamani, A. &amp; Marusik, Y. M. &amp; Malek-Hosseini, M. J." box="[754,969,1447,1474]" pageId="1" pageNumber="261" pagination="95 - 97" refId="ref8577" refString="Zamani, A., Marusik, Y. M. &amp; Malek-Hosseini, M. J. (2018) A new species of Tegenaria Latreille, 1804 (Araneae: Agelenidae) from western Iran. Zootaxa, 4444 (1), 95 - 97. https: // doi. org / 10.11646 / zootaxa. 4444.1.7" type="journal article" year="2018">
Zamani
<emphasis id="B93A930F515AFFE6FCBC13E9FC237493" box="[846,896,1447,1473]" italics="true" pageId="1" pageNumber="261">et al</emphasis>
. 2018
</bibRefCitation>
: fig. 1).
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515AFFE6FF35138AFE3D76EB" pageId="1" pageNumber="261" type="description">
<paragraph id="8BF14F1D515AFFE6FF35138AFF5C77EF" blockId="1.[151,1437,1087,2014]" pageId="1" pageNumber="261">
<emphasis id="B93A930F515AFFE6FF35138AFEFA74B7" bold="true" box="[199,345,1483,1509]" pageId="1" pageNumber="261">Description.</emphasis>
Male (
<typeStatus id="54F5F1BF515AFFE6FE5F138AFDB974B7" box="[429,538,1483,1509]" pageId="1" pageNumber="261" type="holotype">Holotype</typeStatus>
). Habitus as in
<figureCitation id="13755398515AFFE6FD3F138AFC8574B7" box="[717,806,1483,1509]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="1" pageNumber="261">Fig. 1A</figureCitation>
. Total length 6.55. Carapace 3.75 long, 2.70 wide. Eye sizes: AME: 0.15, ALE: 0.20, PME: 0.18, PLE: 0.19. Carapace and sternum yellowish brown, carapace darker in pars cephalica, without any pattern. Labium, chelicerae, and maxillae brownish. Legs of same color as carapace and with faint annulations. Abdomen light beige, with scattered dark gray patches on dorsum and without any pattern ventrally. Spinnerets dark beige. Measurements of legs: I: 25.55 (6.90, 1.60, 7.07, 7.18, 2.80), Fe II: 6.74 (the rest of the segments missing), III: 20.62 (5.80, 1.35, 5.12, 6.40, 1.95), IV: 23.02+missing Ta (6.95, 1.50, 6.27, 8.30, missing).
</paragraph>
<paragraph id="8BF14F1D515AFFE6FF351086FE3176C7" blockId="1.[151,1437,1087,2014]" pageId="1" pageNumber="261">
Palp as in
<figureCitation id="13755398515AFFE6FEB11086FE6277B3" box="[323,449,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[152,255,1818,1842]" captionTargetBox="[157,1430,181,1794]" captionTargetId="figure-35@3.[151,1436,181,1794]" captionTargetPageId="3" captionText="FIGURE 2. Male palp of Tegenaria sorani sp. n. (AD) and Cheiracanthium montanum (E, F). A, E ventral; B prolateral; C, F retrolateral; D dorsal. Abbreviations: Co—conductor, Cr—retrolateral arm of conductor, Ta—tegular apophysis, Ts—heavily sclerotized retrolateral tibial apophysis, Tw—weakly sclerotized retrolateral tibial apophysis. Scale bars: 0.2 mm." pageId="1" pageNumber="261">Fig. 2AD</figureCitation>
; tibia 2 times longer than wide with 2 retrolateral apophyses, one rectangular heavily sclerotized (
<emphasis id="B93A930F515AFFE6FED210ADFE9B7657" box="[288,312,1772,1797]" italics="true" pageId="1" pageNumber="261">Ts</emphasis>
) and another weakly sclerotized (
<emphasis id="B93A930F515AFFE6FD5F10ADFD6E7657" box="[685,717,1772,1797]" italics="true" pageId="1" pageNumber="261">Tw</emphasis>
) as long as tibia wide; cymbium 1.9 times longer than wide, with tip equal to length of bulb; bulb almost as wide as long; tegular apophysis (
<emphasis id="B93A930F515AFFE6FC281151FC56767B" box="[986,1013,1808,1833]" italics="true" pageId="1" pageNumber="261">Ta</emphasis>
) located posteriorly, originating at ca. 6 oclock position, bent at right angle, terminal part as long as basal; conductor (
<emphasis id="B93A930F515AFFE6FBE11175FB97761C" box="[1043,1076,1844,1870]" italics="true" pageId="1" pageNumber="261">Co</emphasis>
) wider than long, prolateral arm lacking, retrolateral arm (
<emphasis id="B93A930F515AFFE6FE4B1119FE747620" box="[441,471,1880,1906]" italics="true" pageId="1" pageNumber="261">Cr</emphasis>
) wide and not pointed; embolus short and thin, originated at ca. 10 oclock position, forming ca. 180º loop.
</paragraph>
<paragraph id="8BF14F1D515AFFE6FF3511DEFE3D76EB" blockId="1.[151,1437,1087,2014]" box="[199,414,1951,1977]" pageId="1" pageNumber="261">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541C96515AFFE6FF351182FAA0768F" box="[199,1283,1987,2014]" pageId="1" pageNumber="261" type="distribution">
<paragraph id="8BF14F1D515AFFE6FF351182FAA0768F" blockId="1.[151,1437,1087,2014]" box="[199,1283,1987,2014]" pageId="1" pageNumber="261">
<emphasis id="B93A930F515AFFE6FF351182FEC2768F" bold="true" box="[199,353,1987,2013]" pageId="1" pageNumber="261">Distribution.</emphasis>
Known only from the
<typeStatus id="54F5F1BF515AFFE6FD921185FD33768C" box="[608,656,1988,2014]" pageId="1" pageNumber="261">type</typeStatus>
locality in
<collectingRegion id="498A81FF515AFFE6FCFD1182FC7E768F" box="[783,989,1987,2013]" country="Iraq" name="Arbil" pageId="1" pageNumber="261">Erbil Governorate</collectingRegion>
, Iraqi
<collectingRegion id="498A81FF515AFFE6FBD61182FB31768F" box="[1060,1170,1987,2013]" country="Iran" name="Kordestan" pageId="1" pageNumber="261">Kurdistan</collectingRegion>
(
<figureCitation id="13755398515AFFE6FB501182FB56768F" box="[1186,1269,1987,2013]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="1" pageNumber="261">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,205 @@
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<emphasis id="B93A930F515DFFE1FF651052FD45777C" bold="true" box="[151,742,1555,1582]" pageId="6" pageNumber="266">
<taxonomicName id="4C4E349E515DFFE1FF651052FD34777F" authority="Zamani &amp; Marusik" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[151,663,1555,1582]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="266" phylum="Arthropoda" rank="species" species="kurdistanica" status="sp. nov.">
<emphasis id="B93A930F515DFFE1FF651052FE39777C" bold="true" box="[151,410,1555,1582]" italics="true" pageId="6" pageNumber="266">Dysdera kurdistanica</emphasis>
Zamani &amp; Marusik
</taxonomicName>
,
<taxonomicNameLabel id="A2092E74515DFFE1FD561055FD45777C" box="[676,742,1556,1582]" pageId="6" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE1FF651079FE9D7700" box="[151,318,1591,1618]" pageId="6" pageNumber="266" type="description">
<paragraph id="8BF14F1D515DFFE1FF651079FE9D7700" blockId="6.[151,742,1555,1618]" box="[151,318,1591,1618]" pageId="6" pageNumber="266">
<figureCitation id="13755398515DFFE1FF651079FF517700" box="[151,242,1592,1618]" captionStart="FIGURE 4" captionStartId="6.[152,255,1116,1140]" captionTargetBox="[151,1402,186,1087]" captionTargetId="figure-18@6.[151,1436,181,1092]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Dysdera goyzha sp. n. (A, B) and Dysdera kurdistanica sp. n. (C), dorsal. A female; B, C males." pageId="6" pageNumber="266">Figs 4C</figureCitation>
,
<figureCitation id="13755398515DFFE1FF0F1076FE9D7700" box="[253,318,1591,1618]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="6" pageNumber="266">5EH</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE1FF651038FCBC77E5" pageId="6" pageNumber="266" type="materials_examined">
<paragraph id="8BF14F1D515DFFE1FF651038FCBC77E5" blockId="6.[151,1437,1657,2044]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF651038FEE877C1" bold="true" box="[151,331,1657,1683]" pageId="6" pageNumber="266">Type material.</emphasis>
<materialsCitation id="3B264540515DFFE1FEA51038FCBC77E5" collectingDate="2023-04-15" collectionCode="ZMUT" collectorName="F. A. Khudhur" country="Iraq" latitude="35.50089" location="Hazar Merd" longLatPrecision="1" longitude="45.315277" municipality="Bakrajo" pageId="6" pageNumber="266" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah" typeStatus="holotype">
<typeStatus id="54F5F1BF515DFFE1FEA51038FE1C77C1" box="[343,447,1657,1683]" pageId="6" pageNumber="266" type="holotype">Holotype</typeStatus>
<specimenCount id="9D488494515DFFE1FE3E103BFE4277C6" box="[460,481,1658,1684]" count="1" pageId="6" pageNumber="266" type="male"></specimenCount>
(
<collectionCode id="ED5FD7D8515DFFE1FE071038FDE977C1" box="[501,586,1657,1683]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="6" pageNumber="266">ZMUT</collectionCode>
),
<collectingCountry id="F3590F8D515DFFE1FD911038FD0F77C1" box="[611,684,1657,1683]" name="Iraq" pageId="6" pageNumber="266">IRAQ</collectingCountry>
:
<emphasis id="B93A930F515DFFE1FD491038FC5677C1" box="[699,1013,1657,1683]" italics="true" pageId="6" pageNumber="266">
<collectingRegion id="498A81FF515DFFE1FD491038FC5677C1" box="[699,1013,1657,1683]" country="Iraq" name="As Sulaymaniyah" pageId="6" pageNumber="266">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B95D567515DFFE1FBF81038FBC577C1" box="[1034,1126,1657,1683]" pageId="6" pageNumber="266">Bakrajo</collectingMunicipality>
,
<location id="8E9119C6515DFFE1FB84103BFAA677C1" LSID="urn:lsid:plazi:treatment:03E7FE0B515DFFE0FF651052FAC87607:8E9119C6515DFFE1FB84103BFAA677C1" box="[1142,1285,1657,1684]" country="Iraq" latitude="35.50089" longLatPrecision="1" longitude="45.315277" municipality="Bakrajo" name="Hazar Merd" pageId="6" pageNumber="266" stateProvince="As Sulaymaniyah">Hazar Merd</location>
, Dekon Vil.,
<geoCoordinate id="EE7A29DA515DFFE1FF6510DCFE9677EA" box="[151,309,1693,1720]" degrees="35" direction="north" minutes="30" orientation="latitude" pageId="6" pageNumber="266" precision="1" seconds="03.2" value="35.50089">35°3003.2“N</geoCoordinate>
<geoCoordinate id="EE7A29DA515DFFE1FECE10DCFE7977EA" box="[316,474,1693,1720]" degrees="45" direction="east" minutes="18" orientation="longitude" pageId="6" pageNumber="266" precision="1" seconds="55.0" value="45.315277">45°1855.0“E</geoCoordinate>
,
<date id="FFF069DD515DFFE1FE1710DCFDC077EA" box="[485,611,1693,1720]" pageId="6" pageNumber="266" value="2023-04-15">
<collectingDate id="EFB49035515DFFE1FE1710DCFDC077EA" box="[485,611,1693,1720]" pageId="6" pageNumber="266" value="2023-04-15">15.04.2023</collectingDate>
</date>
(
<collectorName id="26BB2ACB515DFFE1FD8110DCFCB077E5" box="[627,787,1693,1719]" pageId="6" pageNumber="266">F.A. Khudhur</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE1FF351080FAA27789" box="[199,1281,1729,1756]" pageId="6" pageNumber="266" type="etymology">
<paragraph id="8BF14F1D515DFFE1FF351080FAA27789" blockId="6.[151,1437,1657,2044]" box="[199,1281,1729,1756]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF351080FEED7789" bold="true" box="[199,334,1729,1755]" pageId="6" pageNumber="266">Etymology.</emphasis>
The specific epithet refers to the distribution of the species in the
<collectingRegion id="498A81FF515DFFE1FBC11080FB5E7789" box="[1075,1277,1729,1755]" country="Iran" name="Kordestan" pageId="6" pageNumber="266">Kurdistan Region</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE1FF3510A4FDEE761A" pageId="6" pageNumber="266" type="diagnosis">
<paragraph id="8BF14F1D515DFFE1FF3510A4FDEE761A" blockId="6.[151,1437,1657,2044]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF3510A4FEE177AD" bold="true" box="[199,322,1765,1791]" pageId="6" pageNumber="266">Diagnosis.</emphasis>
The new species differs from all congeners by having a strongly bent posterior apophysis (
<emphasis id="B93A930F515DFFE1FAC310A7FAE177AD" box="[1329,1346,1766,1791]" italics="true" pageId="6" pageNumber="266">P</emphasis>
), a long stylus longer than the crest (
<emphasis id="B93A930F515DFFE1FE281148FE4E7671" box="[474,493,1801,1827]" italics="true" pageId="6" pageNumber="266">C</emphasis>
), and a unique shape of the spermophor (
<emphasis id="B93A930F515DFFE1FC3B1148FC467671" box="[969,997,1801,1827]" italics="true" pageId="6" pageNumber="266">Sp</emphasis>
), forming some loops, but not spirally coiled like in other species of
<taxonomicName id="4C4E349E515DFFE1FE15116CFDE57615" authorityName="Latreille" authorityYear="1804" box="[487,582,1837,1863]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis id="B93A930F515DFFE1FE15116CFDE57615" box="[487,582,1837,1863]" italics="true" pageId="6" pageNumber="266">Dysdera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE0FF351110FE3D7663" lastPageId="7" lastPageNumber="267" pageId="6" pageNumber="266" type="description">
<paragraph id="8BF14F1D515DFFE1FF351110FB4576A9" blockId="6.[151,1437,1657,2044]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF351110FEFA7639" bold="true" box="[199,345,1873,1899]" pageId="6" pageNumber="266">Description.</emphasis>
Male. Habitus as in
<figureCitation id="13755398515DFFE1FDBE1110FD067639" box="[588,677,1873,1899]" captionStart="FIGURE 4" captionStartId="6.[152,255,1116,1140]" captionTargetBox="[151,1402,186,1087]" captionTargetId="figure-18@6.[151,1436,181,1092]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Dysdera goyzha sp. n. (A, B) and Dysdera kurdistanica sp. n. (C), dorsal. A female; B, C males." pageId="6" pageNumber="266">Fig. 4C</figureCitation>
. Total length 4.87. Carapace 2.03 long, 1.54 wide. Eye diameters: AME 0.12, PME 0.10, PLE 0.09. Carapace, sternum, chelicerae, labium, and maxillae light reddish brown. Legs yellowish. Abdomen light beige, without any pattern. Spinnerets uniformly light beige. Measurements of legs: I: 5.61 (1.48, 1.01, 1.26, 1.32, 0.54), II: 5.03 (1.34, 0.86, 1.09, 1.25, 0.49), III: 4.04 (1.15, 0.73, 0.76, 0.96, 0.44), IV: 4.86+missing Ta (1.56, 0.88, 1.14, 1.28, missing). Spination: III: Ti: 1v; Mt: 2v. IV: Ti: 1v; Mt: 2v.
</paragraph>
<caption id="DF311F95515CFFE0FF6A13CCFE0B74BE" pageId="7" pageNumber="267" startId="7.[152,255,1421,1445]" targetBox="[164,1436,181,1397]" targetPageId="7" targetType="figure">
<paragraph id="8BF14F1D515CFFE0FF6A13CCFE0B74BE" blockId="7.[151,1437,1421,1517]" pageId="7" pageNumber="267">
<emphasis id="B93A930F515CFFE0FF6A13CCFEB474F7" bold="true" box="[152,279,1421,1445]" pageId="7" pageNumber="267">FIGURE 5.</emphasis>
Male palp of
<taxonomicName id="4C4E349E515CFFE0FE5213CCFDE174F7" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[416,578,1421,1445]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="goyzha" status="sp. nov.">
<emphasis id="B93A930F515CFFE0FE5213CCFDE174F7" box="[416,578,1421,1445]" italics="true" pageId="7" pageNumber="267">Dysdera goyzha</emphasis>
</taxonomicName>
<emphasis id="B93A930F515CFFE0FDB513CCFDDC74F7" bold="true" box="[583,639,1421,1445]" pageId="7" pageNumber="267">
<taxonomicNameLabel id="A2092E74515CFFE0FDB513CCFDDC74F7" box="[583,639,1421,1445]" pageId="7" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(AD) and
<taxonomicName id="4C4E349E515CFFE0FD0613CCFC6E74F7" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[756,973,1421,1445]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="267" phylum="Arthropoda" rank="species" species="kurdistanica" status="sp. nov.">
<emphasis id="B93A930F515CFFE0FD0613CCFC6E74F7" box="[756,973,1421,1445]" italics="true" pageId="7" pageNumber="267">Dysdera kurdistanica</emphasis>
</taxonomicName>
<emphasis id="B93A930F515CFFE0FC2013CCFBA974F7" bold="true" box="[978,1034,1421,1445]" pageId="7" pageNumber="267">
<taxonomicNameLabel id="A2092E74515CFFE0FC2013CCFBA974F7" box="[978,1034,1421,1445]" pageId="7" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(EH). A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations:
<emphasis id="B93A930F515CFFE0FDAF13F0FD27749A" box="[605,644,1457,1480]" italics="true" pageId="7" pageNumber="267">C—</emphasis>
crest,
<emphasis id="B93A930F515CFFE0FD3113F3FD71749B" box="[707,722,1458,1481]" italics="true" pageId="7" pageNumber="267">P</emphasis>
—posterior apophysis,
<emphasis id="B93A930F515CFFE0FC3113F0FC53749B" box="[963,1008,1457,1481]" italics="true" pageId="7" pageNumber="267">Rl—</emphasis>
retrolateral lamina,
<emphasis id="B93A930F515CFFE0FB4913F0FB48749A" box="[1211,1259,1457,1480]" italics="true" pageId="7" pageNumber="267">Sp—</emphasis>
spermophor,
<emphasis id="B93A930F515CFFE0FA8013F0FA3F749A" box="[1394,1436,1457,1480]" italics="true" pageId="7" pageNumber="267">St—</emphasis>
stylus. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF14F1D515CFFE0FF35107EFE8D765F" blockId="7.[151,1436,1599,1878]" pageId="7" pageNumber="267">
Palp as in
<figureCitation id="13755398515CFFE0FECD107EFE1A7708" box="[319,441,1599,1626]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="7" pageNumber="267">Fig. 5EH</figureCitation>
; cymbium relatively short, 2.7 times longer than wide, gradually tapering in distal half; bulb attached to cymbium in middle part of cymbium; bulb ca. 1.4 times longer than cymbium; tegular part very short, ca. 1/3 of bulb length, in anterior view as long as wide; spermophor (
<emphasis id="B93A930F515CFFE0FC2C10C9FC5977F0" box="[990,1018,1672,1698]" italics="true" pageId="7" pageNumber="267">Sp</emphasis>
) not spirally coiled, but bent forming several loops, in anterior view forming S-shaped figure; posterior apophysis (
<emphasis id="B93A930F515CFFE0FC0C10EDFBAC7797" box="[1022,1039,1708,1733]" italics="true" pageId="7" pageNumber="267">P</emphasis>
) strongly bent, hook-like; crest (
<emphasis id="B93A930F515CFFE0FA7210EDFA307794" box="[1408,1427,1708,1734]" italics="true" pageId="7" pageNumber="267">C</emphasis>
) well sclerotized, bill-shaped, 4 times longer than wide; stylus very long, as long as tegulum wide, bent anteriorly at ca. 75º angle.
</paragraph>
<paragraph id="8BF14F1D515CFFE0FF351156FE3D7663" blockId="7.[151,1436,1599,1878]" box="[199,414,1815,1841]" pageId="7" pageNumber="267">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541C96515CFFE0FF35117AFAC87607" box="[199,1387,1851,1878]" pageId="7" pageNumber="267" type="distribution">
<paragraph id="8BF14F1D515CFFE0FF35117AFAC87607" blockId="7.[151,1436,1599,1878]" box="[199,1387,1851,1878]" pageId="7" pageNumber="267">
<emphasis id="B93A930F515CFFE0FF35117AFEC27607" bold="true" box="[199,353,1851,1877]" pageId="7" pageNumber="267">Distribution.</emphasis>
Known only from the
<typeStatus id="54F5F1BF515CFFE0FD92117DFD337604" box="[608,656,1852,1878]" pageId="7" pageNumber="267">type</typeStatus>
locality in
<collectingRegion id="498A81FF515CFFE0FCFD117AFBE57607" box="[783,1094,1851,1877]" country="Iraq" name="As Sulaymaniyah" pageId="7" pageNumber="267">Sulaymaniyah Governorate</collectingRegion>
, Iraqi
<collectingRegion id="498A81FF515CFFE0FB7E117AFB597607" box="[1164,1274,1851,1877]" country="Iran" name="Kordestan" pageId="7" pageNumber="267">Kurdistan</collectingRegion>
(
<figureCitation id="13755398515CFFE0FAF8117AFAFE7607" box="[1290,1373,1851,1877]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="7" pageNumber="267">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,242 @@
<document id="8CD67A84EDEF2FA752322FF79234EC3C" ID-DOI="10.11646/zootaxa.5492.2.6" ID-ISSN="1175-5326" ID-ZooBank="4C877712-B5C4-451A-87BE-B0C08B2BEC24" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722841366327" checkinUser="plazi" docAuthor="Zamani, Alireza, Khudhur, Farhad A. &amp; Marusik, Yuri M." docDate="2024" docId="03E7FE0B515EFFE1FF6512BAFAC8749D" docLanguage="en" docName="zootaxa.5492.1.6.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dysdera goyzha Zamani &amp; Khudhur &amp; Marusik 2024, sp. n." docType="treatment" docVersion="1" lastPageNumber="266" masterDocId="FFDE8673515BFFE7FFF21641FFA37152" masterDocTitle="New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records" masterLastPageNumber="278" masterPageNumber="260" pageNumber="265" updateTime="1722841495305" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
<mods:mods id="C20D5E7A337E6DACB793DB272D3C9FCE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DC6BBA5539279C4482632774A637C29D">
<mods:title id="FA263D7D59882F6C85437B424D2332D2">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
</mods:titleInfo>
<mods:name id="087DA1AEE15E92777CD7D1EB1095A34A" type="personal">
<mods:role id="B22C02132D09D142720DAFD1EB324881">
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</mods:role>
<mods:namePart id="32E6A9E62F9D936E2974D53393F3F097">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="DBE7429506D47612753EF3335FBC051A" type="ORCID">0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation id="85152DB686D4E15806BC85001E7D8847">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku 20500, Finland</mods:affiliation>
<mods:nameIdentifier id="46CDD529CF97D070ADE4291E50000F2A" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8335D1BCC9CFECF778264CF49EB1942C" type="personal">
<mods:role id="9F0E6B4C97D5105C76E95B3AC649A887">
<mods:roleTerm id="1C8487F4ABBF2F49E3AE23049E993C91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8866893AC843E128855BBAFF085A1696">Khudhur, Farhad A.</mods:namePart>
<mods:nameIdentifier id="A248C96B84463AE1764B4BA5AFEAA908" type="ORCID">0000-0001-5267-6334</mods:nameIdentifier>
<mods:affiliation id="E865C6884340A15EEF26FA6D58447A23">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="385A8B72F6C48D8B86B2A74CA6B2B861" type="email">qi.ude.lusvinu@ruhduhk.dahraf</mods:nameIdentifier>
</mods:name>
<mods:name id="8600933B67EEF8C56AFC4D0735FDE85A" type="personal">
<mods:role id="C1FD35CC39784DFF74A89C220E1CD1E5">
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</mods:role>
<mods:namePart id="E3EBBED3BD9B22AC2DC510E9A869CCEE">Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier id="388DE89F1907C5D79EF1A31B455340F6" type="ORCID">0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation id="F4B1528839488E97C0CE2E150AFB2BD8">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
<mods:nameIdentifier id="FF1EB7848ADE41D39BA969BED5C107E8" type="email">yurmar@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F44F29A2545C814AA887DA1DA7952737">text</mods:typeOfResource>
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<mods:title id="AC51B4CDB00A00A3263CE05021F644F6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="17B9863B8A21450FCAAB1EE09F06829D">
<mods:date id="66EDC5E48213DA0A6BF0E1CD918A4DE1">2024</mods:date>
<mods:detail id="58E78F527D47326FB555BA298A2430CA" type="pubDate">
<mods:number id="8E78DE007F4FB95C1160D6867101A9B3">2024-08-05</mods:number>
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<mods:detail id="F5C4B0B71EA7B4A8FF6ED0716E443082" type="volume">
<mods:number id="82FCDFAB5D5D4969EBD73F609811C8F2">5492</mods:number>
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<mods:url id="982DCCDEA6E9F8049F02F977DB7737E1">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
<mods:classification id="8F7BC52AC9C3685A28DA04F007654E96">journal article</mods:classification>
<mods:identifier id="835C4F8504BFF31D2F49D9D6F8013E43" type="DOI">10.11646/zootaxa.5492.2.6</mods:identifier>
<mods:identifier id="15137BFE8B52E71F95E6D4A3CA0ABDCD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E7FE0B515EFFE1FF6512BAFAC8749D" LSID="urn:lsid:plazi:treatment:03E7FE0B515EFFE1FF6512BAFAC8749D" httpUri="http://treatment.plazi.org/id/03E7FE0B515EFFE1FF6512BAFAC8749D" lastPageId="6" lastPageNumber="266" pageId="5" pageNumber="265">
<subSubSection id="C3541C96515EFFE2FF6512BAFD007447" box="[151,675,1275,1302]" pageId="5" pageNumber="265" type="nomenclature">
<paragraph id="8BF14F1D515EFFE2FF6512BAFD007447" blockId="5.[151,675,1275,1338]" box="[151,675,1275,1302]" pageId="5" pageNumber="265">
<heading id="D0B9F871515EFFE2FF6512BAFD007447" bold="true" box="[151,675,1275,1302]" fontSize="11" level="1" pageId="5" pageNumber="265" reason="1">
<emphasis id="B93A930F515EFFE2FF6512BAFD007447" bold="true" box="[151,675,1275,1302]" pageId="5" pageNumber="265">
<taxonomicName id="4C4E349E515EFFE2FF6512BAFDF77447" authority="Zamani &amp; Marusik" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[151,596,1275,1302]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="goyzha" status="sp. nov.">
<emphasis id="B93A930F515EFFE2FF6512BAFEF47444" bold="true" box="[151,343,1275,1302]" italics="true" pageId="5" pageNumber="265">Dysdera goyzha</emphasis>
Zamani &amp; Marusik
</taxonomicName>
,
<taxonomicNameLabel id="A2092E74515EFFE2FD9312BAFD007447" box="[609,675,1275,1301]" pageId="5" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515EFFE2FF65135EFE377468" box="[151,404,1311,1338]" pageId="5" pageNumber="265" type="description">
<paragraph id="8BF14F1D515EFFE2FF65135EFE377468" blockId="5.[151,675,1275,1338]" box="[151,404,1311,1338]" pageId="5" pageNumber="265">
<figureCitation id="13755398515EFFE2FF65135EFEB7746B" box="[151,276,1311,1337]" captionStart="FIGURE 4" captionStartId="6.[152,255,1116,1140]" captionTargetBox="[151,1402,186,1087]" captionTargetId="figure-18@6.[151,1436,181,1092]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Dysdera goyzha sp. n. (A, B) and Dysdera kurdistanica sp. n. (C), dorsal. A female; B, C males." pageId="5" pageNumber="265">Figs 4AB</figureCitation>
,
<figureCitation id="13755398515EFFE2FEED135EFEC77468" box="[287,356,1311,1338]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="5" pageNumber="265">5AD</figureCitation>
,
<figureCitation id="13755398515EFFE2FE80135EFE377468" box="[370,404,1311,1338]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="5" pageNumber="265">6D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515EFFE2FF651326FADD74F7" pageId="5" pageNumber="265" type="materials_examined">
<paragraph id="8BF14F1D515EFFE2FF651326FADD74F7" blockId="5.[151,1437,1383,2022]" pageId="5" pageNumber="265">
<emphasis id="B93A930F515EFFE2FF651326FEEF74D3" bold="true" box="[151,332,1383,1409]" pageId="5" pageNumber="265">Type material.</emphasis>
<materialsCitation id="3B264540515EFFE2FEA81326FCB874F7" collectingDate="2022-08-03" collectionCode="ZMUT" collectorName="F. A. Khudhur" country="Iraq" latitude="35.62419" location="Hawary Shar Park" longLatPrecision="1" longitude="45.458" municipality="Goyzha Mt." pageId="5" pageNumber="265" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah" typeStatus="holotype">
<typeStatus id="54F5F1BF515EFFE2FEA81326FE6174D3" box="[346,450,1383,1409]" pageId="5" pageNumber="265" type="holotype">Holotype</typeStatus>
<specimenCount id="9D488494515EFFE2FE231329FE4574D0" box="[465,486,1384,1410]" count="1" pageId="5" pageNumber="265" type="male"></specimenCount>
(
<collectionCode id="ED5FD7D8515EFFE2FE0E1326FDF274D3" box="[508,593,1383,1409]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="5" pageNumber="265">ZMUT</collectionCode>
),
<collectingCountry id="F3590F8D515EFFE2FD991326FD1774D3" box="[619,692,1383,1409]" name="Iraq" pageId="5" pageNumber="265">IRAQ</collectingCountry>
:
<emphasis id="B93A930F515EFFE2FD371326FBA274D0" box="[709,1025,1383,1410]" italics="true" pageId="5" pageNumber="265">
<collectingRegion id="498A81FF515EFFE2FD371326FBA274D0" box="[709,1025,1383,1410]" country="Iraq" name="As Sulaymaniyah" pageId="5" pageNumber="265">Sulaymaniyah Governorate</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B95D567515EFFE2FBE51326FB0474D0" box="[1047,1191,1383,1410]" pageId="5" pageNumber="265">Goyzha Mt.</collectingMunicipality>
,
<location id="8E9119C6515EFFE2FB481329FA3B74D3" LSID="urn:lsid:plazi:treatment:03E7FE0B515EFFE1FF6512BAFAC8749D:8E9119C6515EFFE2FB481329FA3B74D3" box="[1210,1432,1383,1410]" country="Iraq" latitude="35.62419" longLatPrecision="1" longitude="45.458" municipality="Goyzha Mt." name="Hawary Shar Park" pageId="5" pageNumber="265" stateProvince="As Sulaymaniyah">Hawary Shar Park</location>
,
<geoCoordinate id="EE7A29DA515EFFE2FF6513CAFE9574F4" box="[151,310,1419,1446]" degrees="35" direction="north" minutes="37" orientation="latitude" pageId="5" pageNumber="265" precision="1" seconds="27.1" value="35.62419">35°3727.1“N</geoCoordinate>
<geoCoordinate id="EE7A29DA515EFFE2FECE13CAFE7974F4" box="[316,474,1419,1446]" degrees="45" direction="east" minutes="27" orientation="longitude" pageId="5" pageNumber="265" precision="1" seconds="28.8" value="45.458">45°2728.8“E</geoCoordinate>
,
<date id="FFF069DD515EFFE2FE1713CAFDC074F7" box="[485,611,1419,1445]" pageId="5" pageNumber="265" value="2022-08-03">
<collectingDate id="EFB49035515EFFE2FE1713CAFDC074F7" box="[485,611,1419,1445]" pageId="5" pageNumber="265" value="2022-08-03">08.03.2022</collectingDate>
</date>
(
<collectorName id="26BB2ACB515EFFE2FD8113CAFCB074F7" box="[627,787,1419,1445]" pageId="5" pageNumber="265">F.A. Khudhur</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B264540515EFFE2FCD713CDFAD974F7" box="[805,1402,1419,1446]" collectionCode="ZMUT" country="Iraq" pageId="5" pageNumber="265" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="As Sulaymaniyah" typeStatus="paratype">
<typeStatus id="54F5F1BF515EFFE2FCD713CDFC3574F4" box="[805,918,1420,1446]" pageId="5" pageNumber="265" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D488494515EFFE2FC5313CAFC6774F7" box="[929,964,1419,1445]" count="1" pageId="5" pageNumber="265" type="male">1♂</specimenCount>
<specimenCount id="9D488494515EFFE2FC3913CAFC4D74F7" box="[971,1006,1419,1445]" count="2" pageId="5" pageNumber="265" type="female">2♀</specimenCount>
(
<collectionCode id="ED5FD7D8515EFFE2FC0F13CAFBF174F7" box="[1021,1106,1419,1445]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="5" pageNumber="265">ZMUT</collectionCode>
), same as for the
<typeStatus id="54F5F1BF515EFFE2FAE613CAFAD974F7" box="[1300,1402,1419,1445]" pageId="5" pageNumber="265" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515EFFE2FF3513EEFAF8749B" box="[199,1371,1455,1482]" pageId="5" pageNumber="265" type="etymology">
<paragraph id="8BF14F1D515EFFE2FF3513EEFAF8749B" blockId="5.[151,1437,1383,2022]" box="[199,1371,1455,1482]" pageId="5" pageNumber="265">
<emphasis id="B93A930F515EFFE2FF3513EEFEED749B" bold="true" box="[199,334,1455,1481]" pageId="5" pageNumber="265">Etymology.</emphasis>
The specific epithet is a noun in apposition, referring to the
<typeStatus id="54F5F1BF515EFFE2FC0113F1FB807498" box="[1011,1059,1456,1482]" pageId="5" pageNumber="265">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515EFFE2FF351392FBFC772F" pageId="5" pageNumber="265" type="diagnosis">
<paragraph id="8BF14F1D515EFFE2FF351392FBFC772F" blockId="5.[151,1437,1383,2022]" pageId="5" pageNumber="265">
<emphasis id="B93A930F515EFFE2FF351392FEE174BF" bold="true" box="[199,322,1491,1517]" pageId="5" pageNumber="265">Diagnosis.</emphasis>
The male of the new species is somewhat similar to that of
<taxonomicName id="4C4E349E515EFFE2FC001395FF6C7743" authority="Zamani, Marusik &amp; Szuts, 2023" authorityName="Zamani, Marusik &amp; Szuts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="tapuria">
<emphasis id="B93A930F515EFFE2FC001395FBC474BF" box="[1010,1127,1492,1517]" italics="true" pageId="5" pageNumber="265">D. tapuria</emphasis>
<bibRefCitation id="EFDF32EC515EFFE2FB821392FF6C7743" author="Zamani, A. &amp; Marusik, Y. M. &amp; Szuts, T." pageId="5" pageNumber="265" pagination="43 - 86" refId="ref8638" refString="Zamani, A., Marusik, Y. M. &amp; Szuts, T. (2023) A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. ZooKeys, 1146, 43 - 86. https: // doi. org / 10.3897 / zookeys. 1146.97517" type="journal article" year="2023">Zamani, Marusik &amp; Szűts, 2023</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3590F8D515EFFE2FEE413B9FEE57740" box="[278,326,1528,1554]" name="Iran" pageId="5" pageNumber="265">Iran</collectingCountry>
, in having a low, poorly sclerotized crest (
<emphasis id="B93A930F515EFFE2FCDA13B9FC987740" box="[808,827,1528,1554]" italics="true" pageId="5" pageNumber="265">C</emphasis>
), but it differs by the lack of a stylus (cf.
<figureCitation id="13755398515EFFE2FAE513B6FACF7740" box="[1303,1388,1527,1554]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="5" pageNumber="265">Fig. 5A</figureCitation>
and Zamani
<emphasis id="B93A930F515EFFE2FF01105DFE877767" box="[243,292,1563,1589]" italics="true" pageId="5" pageNumber="265">et al</emphasis>
. 2023: fig. 30A).
<taxonomicName id="4C4E349E515EFFE2FE18105AFD3D7767" authorityName="Zamani &amp; Khudhur &amp; Marusik" authorityYear="2024" box="[490,670,1563,1589]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="265" phylum="Arthropoda" rank="species" species="goyzha" status="sp. nov.">
<emphasis id="B93A930F515EFFE2FE18105AFD3D7767" box="[490,670,1563,1589]" italics="true" pageId="5" pageNumber="265">Dysdera goyzha</emphasis>
</taxonomicName>
<emphasis id="B93A930F515EFFE2FD57105AFD477767" bold="true" box="[677,740,1563,1589]" pageId="5" pageNumber="265">
<taxonomicNameLabel id="A2092E74515EFFE2FD57105AFD477767" box="[677,740,1563,1589]" pageId="5" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from all congeners in having a large retrolateral lamina (
<emphasis id="B93A930F515EFFE2FF53107EFF19770B" box="[161,186,1599,1625]" italics="true" pageId="5" pageNumber="265">Rl</emphasis>
) forming a claw- or bill-like shape in dorsal and ventral views (
<figureCitation id="13755398515EFFE2FC84107EFC487708" box="[886,1003,1599,1626]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="5" pageNumber="265">Fig. 5C, D</figureCitation>
). The female of the new species differs from all congeners by having a ragged anterior surface of the receptacle (
<emphasis id="B93A930F515EFFE2FC3E1025FC49772F" box="[972,1002,1636,1661]" italics="true" pageId="5" pageNumber="265">Re</emphasis>
;
<figureCitation id="13755398515EFFE2FC0A1022FBF1772F" box="[1016,1106,1635,1661]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="5" pageNumber="265">Fig. 6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515EFFE1FF3510C6FAFC74F9" lastPageId="6" lastPageNumber="266" pageId="5" pageNumber="265" type="description">
<paragraph id="8BF14F1D515EFFE2FF3510C6FC427604" blockId="5.[151,1437,1383,2022]" pageId="5" pageNumber="265">
<emphasis id="B93A930F515EFFE2FF3510C6FEFA77F3" bold="true" box="[199,345,1671,1697]" pageId="5" pageNumber="265">Description.</emphasis>
Male (
<typeStatus id="54F5F1BF515EFFE2FE5910C6FDBB77F3" box="[427,536,1671,1697]" pageId="5" pageNumber="265" type="holotype">Holotype</typeStatus>
). Habitus as in
<figureCitation id="13755398515EFFE2FD3A10C6FC8377F3" box="[712,800,1671,1697]" captionStart="FIGURE 4" captionStartId="6.[152,255,1116,1140]" captionTargetBox="[151,1402,186,1087]" captionTargetId="figure-18@6.[151,1436,181,1092]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Dysdera goyzha sp. n. (A, B) and Dysdera kurdistanica sp. n. (C), dorsal. A female; B, C males." pageId="5" pageNumber="265">Fig. 4B</figureCitation>
. Total length 10.83. Carapace 4.00 long, 3.20 wide. Eye diameters: AME 0.22, PME 0.16, PLE 0.18. Carapace, sternum, chelicerae, labium, and maxillae light reddish brown. Legs light orange. Abdomen beige, without any pattern. Spinnerets uniformly beige. Measurements of legs: I: 14.47 (4.00, 2.44, 3.50, 3.62, 0.91), II: 13.44 (3.78, 2.17, 3.21, 3.40, 0.88), III: 10.31 (2.95, 1.63, 2.10, 2.83, 0.80), IV: 12.90+missing Ta (4.11, 2.06, 2.95, 3.78, missing). Spination: I: Fe: 4pl. II: Fe: 3pl. III: Fe: 2(3)d, 2pl; Ti: 1d, 2pl, 1rl, 6v; Mt: 3pl, 2rl, 4v. IV: Fe: 9d; Ti: 1d, 3pl, 3rl, 9v; Mt: 3pl, 5rl, 5v.
</paragraph>
<paragraph id="8BF14F1D515EFFE2FF35111EFD0676B7" blockId="5.[151,1437,1383,2022]" pageId="5" pageNumber="265">
Palp as in
<figureCitation id="13755398515EFFE2FEB5111EFE657628" box="[327,454,1887,1914]" captionStart="FIGURE 5" captionStartId="7.[152,255,1421,1445]" captionTargetBox="[164,1436,181,1397]" captionTargetId="figure-35@7.[151,1436,181,1397]" captionTargetPageId="7" captionText="FIGURE 5. Male palp of Dysdera goyzha sp. n. (AD) and Dysdera kurdistanica sp. n. (EH).A, E retrolateral; B, F prolateral; C, G distal; D, H proximal. Abbreviations: C—crest, P—posterior apophysis, Rl—retrolateral lamina, Sp—spermophor, St— stylus. Scale bars: 0.2 mm." pageId="5" pageNumber="265">Fig. 5AD</figureCitation>
; cymbium long, 3.5 times longer than wide; bulb attached to cymbium in proximal half; bulb ca. 1.6 times longer than cymbium, and 3.3 times longer than wide; tegular part 1.2 times longer than wide; psembolus longer than tegulum, with weakly sclerotized and low crest (
<emphasis id="B93A930F515EFFE2FBF111E9FBB57690" box="[1027,1046,1960,1986]" italics="true" pageId="5" pageNumber="265">C</emphasis>
) and large retrolateral lamina (
<emphasis id="B93A930F515EFFE2FA8111E6FA2F7693" box="[1395,1420,1959,1985]" italics="true" pageId="5" pageNumber="265">Rl</emphasis>
), proximal part of lamina bent toward cymbium.
</paragraph>
<caption id="DF311F95515DFFE1FF6A121DFAC07526" box="[152,1379,1116,1140]" pageId="6" pageNumber="266" startId="6.[152,255,1116,1140]" targetBox="[151,1402,186,1087]" targetPageId="6" targetType="figure">
<paragraph id="8BF14F1D515DFFE1FF6A121DFAC07526" blockId="6.[152,1379,1116,1140]" box="[152,1379,1116,1140]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF6A121DFEBB7526" bold="true" box="[152,280,1116,1140]" pageId="6" pageNumber="266">FIGURE 4.</emphasis>
Habitus of
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<emphasis id="B93A930F515DFFE1FE7D121DFD907526" box="[399,563,1116,1140]" italics="true" pageId="6" pageNumber="266">Dysdera goyzha</emphasis>
</taxonomicName>
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</emphasis>
(A, B) and
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<emphasis id="B93A930F515DFFE1FD19121DFC667526" box="[747,965,1116,1140]" italics="true" pageId="6" pageNumber="266">Dysdera kurdistanica</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A2092E74515DFFE1FC39121DFBA77526" box="[971,1028,1116,1140]" pageId="6" pageNumber="266" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(C), dorsal. A female; B, C males.
</paragraph>
</caption>
<paragraph id="8BF14F1D515DFFE1FF3512F8FE727436" blockId="6.[151,1437,1209,1488]" pageId="6" pageNumber="266">
Female. Habitus as in
<figureCitation id="13755398515DFFE1FE3712F8FDBC7581" box="[453,543,1209,1235]" captionStart="FIGURE 4" captionStartId="6.[152,255,1116,1140]" captionTargetBox="[151,1402,186,1087]" captionTargetId="figure-18@6.[151,1436,181,1092]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Dysdera goyzha sp. n. (A, B) and Dysdera kurdistanica sp. n. (C), dorsal. A female; B, C males." pageId="6" pageNumber="266">Fig. 4A</figureCitation>
. Total length 12.63. Carapace 4.65 long, 3.32 wide. Eye diameters: AME 0.22, PME 0.17, PLE 0.19. Coloration as in male. Measurements of legs: I: 15.15 (4.24, 2.82, 3.55, 3.49, 1.05), II: 14.75 (4.20, 2.45, 3.43, 3.62, 1.05), III: 11.31 (3.32, 1.80, 2.26, 2.98, 0.95), IV: 15.45 (4.54, 2.27, 3.30, 4.27, 1.07). Spination: I: Fe: 4pl. II: Fe: 3(4)pl. III: Fe: 2(5)d, 1(0)pl; Ti: 1d, 2pl, 2rl, 6v; Mt: 3pl, 2rl, 4v. IV: Fe: 7(9)d; Ti: 1d, 3pl, 3rl, 9v; Mt: 3pl, 5rl, 5v.
</paragraph>
<paragraph id="8BF14F1D515DFFE1FF35132CFAFC74F9" blockId="6.[151,1437,1209,1488]" pageId="6" pageNumber="266">
Endogyne as in
<figureCitation id="13755398515DFFE1FE86132CFE6874D5" box="[372,459,1389,1415]" captionStart="FIGURE 6" captionStartId="8.[152,255,1147,1171]" captionTargetBox="[151,1436,181,1123]" captionTargetId="figure-18@8.[151,1436,181,1123]" captionTargetPageId="8" captionText="FIGURE 6. Female copulatory organs of Haplodrassus qashqai (A), Zelotes hazarmerdensis sp. n. (B, C) and Dysdera goyzha sp. n. (D). A intact, ventral; B macerated, ventral; C, D macerated, dorsal. Abbreviations: Ah—anterior hood, Ag—accessorial gland, Da—dorsal arch, Fo—fovea, Re—receptacle, Tb—transversal bar. Scale bars: 0.2 mm." pageId="6" pageNumber="266">Fig. 6D</figureCitation>
; receptacle (
<emphasis id="B93A930F515DFFE1FDA6132FFDD174D5" box="[596,626,1390,1415]" italics="true" pageId="6" pageNumber="266">Re</emphasis>
) straight with ragged anterior margin; dorsal arch of anterior diverticulum (
<emphasis id="B93A930F515DFFE1FF5313D3FF6074F9" box="[161,195,1426,1451]" italics="true" pageId="6" pageNumber="266">Da</emphasis>
) semiround, 2 times wider than long; transversal bar (
<emphasis id="B93A930F515DFFE1FCD113D0FCE274F9" box="[803,833,1425,1451]" italics="true" pageId="6" pageNumber="266">Tb</emphasis>
) long and thin, 1.5 times longer than receptacle.
</paragraph>
</subSubSection>
<subSubSection id="C3541C96515DFFE1FF3513F4FAC8749D" box="[199,1387,1461,1488]" pageId="6" pageNumber="266" type="distribution">
<paragraph id="8BF14F1D515DFFE1FF3513F4FAC8749D" blockId="6.[151,1437,1209,1488]" box="[199,1387,1461,1488]" pageId="6" pageNumber="266">
<emphasis id="B93A930F515DFFE1FF3513F4FEC2749D" bold="true" box="[199,353,1461,1487]" pageId="6" pageNumber="266">Distribution.</emphasis>
Known only from the
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locality in
<collectingRegion id="498A81FF515DFFE1FCFD13F4FBE5749D" box="[783,1094,1461,1487]" country="Iraq" name="As Sulaymaniyah" pageId="6" pageNumber="266">Sulaymaniyah Governorate</collectingRegion>
, Iraqi
<collectingRegion id="498A81FF515DFFE1FB7E13F4FB59749D" box="[1164,1274,1461,1487]" country="Iran" name="Kordestan" pageId="6" pageNumber="266">Kurdistan</collectingRegion>
(
<figureCitation id="13755398515DFFE1FAF813F4FAFE749D" box="[1290,1373,1461,1487]" captionStart="FIGURE 10" captionStartId="16.[152,255,1294,1318]" captionTargetBox="[151,1436,381,1271]" captionTargetId="figure-58@16.[151,1436,381,1271]" captionTargetPageId="16" captionText="FIGURE 10. Collection localities of the new species described in this paper in Iraqi Kurdistan: Tegenaria sorani sp. n. (circle), Dysdera goyzha sp. n. (triangle), Dysdera kurdistanica sp. n. and Zelotes hazarmerdensis sp. n. (square), and Palpimanus garmiyanus sp. n. (hexagon)." pageId="6" pageNumber="266">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,108 @@
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Genus
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Latreille, 1804
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<emphasis id="B93A930F515EFFE2FF6515BEFEB7754B" bold="true" box="[151,276,1023,1049]" pageId="5" pageNumber="265">Comment.</emphasis>
<taxonomicName id="4C4E349E515EFFE2FEE915BEFED9754B" authorityName="Latreille" authorityYear="1804" box="[283,378,1023,1049]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B93A930F515EFFE2FEE915BEFED9754B" box="[283,378,1023,1049]" italics="true" pageId="5" pageNumber="265">Dysdera</emphasis>
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is one of the largest spider genera, with 326 named species (
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). Only one species,
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<emphasis id="B93A930F515EFFE2FA731241FA36754B" box="[1409,1429,1024,1049]" italics="true" pageId="5" pageNumber="265">D</emphasis>
.
<emphasis id="B93A930F515EFFE2FF651265FF4F756F" box="[151,236,1060,1085]" italics="true" pageId="5" pageNumber="265">crocata</emphasis>
C.L. Koch, 1838
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, has a global distribution due to anthropogenic dispersal. The vast majority of the species have very limited ranges.Almost all species known from neighboring
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and
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are endemics or subendemics (Zamani
<emphasis id="B93A930F515EFFE2FEF3122DFE9575D7" box="[257,310,1131,1157]" italics="true" pageId="5" pageNumber="265">et al</emphasis>
. 2023;
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Danışman
<emphasis id="B93A930F515EFFE2FDFC122DFDEA75D7" box="[526,585,1131,1157]" italics="true" pageId="5" pageNumber="265">et al.</emphasis>
2024
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). Therefore, for the two new species described here, we have only compared them with species from nearby regions.
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<emphasis id="B93A930F515FFFE3FF6510E2FE0F77EF" bold="true" box="[151,428,1699,1726]" italics="true" pageId="4" pageNumber="264">Porrhoclubiona vegeta</emphasis>
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(
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<emphasis id="B93A930F515FFFE3FF351172FEC2761F" bold="true" box="[199,353,1843,1869]" pageId="4" pageNumber="264">Distribution.</emphasis>
Mediterranean to
<collectingCountry id="F3590F8D515FFFE3FDDC1175FDFC761C" box="[558,607,1844,1870]" name="Iran" pageId="4" pageNumber="264">Iran</collectingCountry>
. New genus and species records for
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.
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View file

@ -0,0 +1,121 @@
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L. Koch, 1877
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,
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<emphasis id="B93A930F515FFFE3FF6513E6FEA57493" bold="true" box="[151,262,1447,1473]" pageId="4" pageNumber="264">Material.</emphasis>
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<geoCoordinate id="EE7A29DA515FFFE3FB0D13E6FA3B7490" box="[1279,1432,1447,1474]" degrees="44" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="264" precision="1" seconds="43.7" value="44.978806">44°58'43.7&quot;E</geoCoordinate>
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<collectingDate id="EFB49035515FFFE3FF65138AFEBA74B7" box="[151,281,1483,1509]" pageId="4" pageNumber="264" value="2022-06-04">06.04.2022</collectingDate>
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.
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<emphasis id="B93A930F515FFFE3FF3513AEFEC2775B" bold="true" box="[199,353,1519,1545]" pageId="4" pageNumber="264">Distribution.</emphasis>
<collectingCountry id="F3590F8D515FFFE3FE9513AEFE04775B" box="[359,423,1519,1545]" name="Spain" pageId="4" pageNumber="264">Spain</collectingCountry>
to
<collectingCountry id="F3590F8D515FFFE3FE3913B1FE5F7758" box="[459,508,1520,1546]" name="Iran" pageId="4" pageNumber="264">Iran</collectingCountry>
. New family, genus, and species records for
<collectingCountry id="F3590F8D515FFFE3FC1D13B1FB837758" box="[1007,1056,1520,1546]" name="Iraq" pageId="4" pageNumber="264">Iraq</collectingCountry>
.
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View file

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<mods:namePart id="8480B8C512C74FDF65F3682DBAA8988C">Khudhur, Farhad A.</mods:namePart>
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<mods:affiliation id="4B460D5624E7C221C712F8F50ACE2947">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<mods:affiliation id="36AC99BB3371FC2C76213821BF60B791">Institute for Biological Problems of the North FEB RAS, Magadan, Russia &amp; Altai State University, Barnaul, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<paragraph id="8BF14F1D515FFFE3FF651496FD6473A3" blockId="4.[151,711,727,789]" box="[151,711,727,754]" pageId="4" pageNumber="264">
<heading id="D0B9F871515FFFE3FF651496FD6473A3" bold="true" box="[151,711,727,754]" fontSize="11" level="1" pageId="4" pageNumber="264" reason="1">
<taxonomicName id="4C4E349E515FFFE3FF651496FD6473A3" ID-CoL="3FYVJ" authority="(Walckenaer, 1802)" baseAuthorityName="Walckenaer" baseAuthorityYear="1802" box="[151,711,727,754]" class="Arachnida" family="Araneidae" genus="Gibbaranea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="264" phylum="Arthropoda" rank="species" species="bituberculata">
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<emphasis id="B93A930F515FFFE3FF651496FE6C73A0" bold="true" box="[151,463,727,754]" italics="true" pageId="4" pageNumber="264">Gibbaranea bituberculata</emphasis>
(Walckenaer, 1802)
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<figureCitation id="13755398515FFFE3FF6514BAFF527247" box="[151,241,763,789]" captionStart="FIGURE 1" captionStartId="2.[152,255,1886,1910]" captionTargetBox="[157,1433,181,1858]" captionTargetId="figure-18@2.[151,1436,181,1863]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (AF, H, I) and cephalothorax (G) of Tegenaria sorani sp. n. (A), Gibbaranea bituberculata (B), Enoplognatha gemina (C), Zelotes hazarmerdensis sp. n. (D), Haplodrassus qashqai (E), Cheiracanthium montanum (F, G), Rhysodromus rikhteri (H) and Xysticus acerbus (I), dorsal (AF, H, I) and frontal (G). AC, FG, I males; D, E, H females." pageId="4" pageNumber="264">Figs 1B</figureCitation>
,
<figureCitation id="13755398515FFFE3FF0F14BAFEE37247" box="[253,320,763,789]" captionStart="FIGURE 3" captionStartId="5.[152,255,773,797]" captionTargetBox="[169,1419,181,749]" captionTargetId="figure-35@5.[151,1436,181,749]" captionTargetPageId="5" captionText="FIGURE 3. Male palp of Gibbaranea bituberculata. A ventral; B prolateral; C retrolateral. Scale bars: 0.2 mm." pageId="4" pageNumber="264">3AC</figureCitation>
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<emphasis id="B93A930F515FFFE3FF651502FEA5720F" bold="true" box="[151,262,835,861]" pageId="4" pageNumber="264">Material.</emphasis>
<materialsCitation id="3B264540515FFFE3FEE01502FEBA72D3" collectingDate="2022-06-04" collectionCode="ZMUT" country="Iraq" latitude="35.78" location="Chamy Razan valley" longLatPrecision="1" longitude="44.978806" pageId="4" pageNumber="264" specimenCount="1" specimenCount-male="1" stateProvince="As Sulaymaniyah">
<collectingCountry id="F3590F8D515FFFE3FEE01502FEF5720F" box="[274,342,835,861]" name="Iraq" pageId="4" pageNumber="264">IRAQ</collectingCountry>
:
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<collectingRegion id="498A81FF515FFFE3FE9B1502FD01720C" box="[361,674,835,862]" country="Iraq" name="As Sulaymaniyah" pageId="4" pageNumber="264">Sulaymaniyah Governorate</collectingRegion>
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:
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(
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,
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;
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<specimenCount id="9D488494515FFFE3FED61526FEE472D3" box="[292,327,871,897]" count="1" pageId="4" pageNumber="264" type="male">1♂</specimenCount>
(
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),
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,
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,
<geoCoordinate id="EE7A29DA515FFFE3FCDF1526FC6572D0" box="[813,966,871,898]" degrees="35" direction="north" minutes="36" orientation="latitude" pageId="4" pageNumber="264" precision="1" seconds="57.2" value="35.615887">35°36'57.2&quot;N</geoCoordinate>
<geoCoordinate id="EE7A29DA515FFFE3FC3F1526FBC572D0" box="[973,1126,871,898]" degrees="45" direction="east" minutes="26" orientation="longitude" pageId="4" pageNumber="264" precision="1" seconds="47.0" value="45.44639">45°26'47.0&quot;E</geoCoordinate>
,
<date id="FFF069DD515FFFE3FB831526FB5172D3" box="[1137,1266,871,897]" pageId="4" pageNumber="264" value="2022-03-30">
<collectingDate id="EFB49035515FFFE3FB831526FB5172D3" box="[1137,1266,871,897]" pageId="4" pageNumber="264" value="2022-03-30">30.03.2022</collectingDate>
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.
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<emphasis id="B93A930F515FFFE3FF3515CAFEC272F7" bold="true" box="[199,353,907,933]" pageId="4" pageNumber="264">Distribution.</emphasis>
West Palaearctic. New genus and species records for
<collectingCountry id="F3590F8D515FFFE3FC4915CDFC4F72F4" box="[955,1004,908,934]" name="Iraq" pageId="4" pageNumber="264">Iraq</collectingCountry>
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View file

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<emphasis id="B93A930F515FFFE3FF6515B6FEDF7540" bold="true" box="[151,380,1015,1042]" italics="true" pageId="4" pageNumber="264">Mangora acalypha</emphasis>
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<emphasis id="B93A930F515FFFE3FF65127EFEA5750B" bold="true" box="[151,262,1087,1113]" pageId="4" pageNumber="264">Material.</emphasis>
<materialsCitation id="3B264540515FFFE3FEFF127EFC31752F" collectingDate="2021-06-13" collectionCode="ZMUT" country="Iraq" county="Records in Iraq." latitude="35.161083" location="Iraqi Kurdistan" longLatPrecision="1" longitude="46.11647" municipality="New" pageId="4" pageNumber="264" specimenCount="1" specimenCount-female="1" stateProvince="Salah ad Din">
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:
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(
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), Hawar Vil.,
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<geoCoordinate id="EE7A29DA515FFFE3FBD1127EFB1F7508" box="[1059,1212,1087,1114]" degrees="46" direction="east" minutes="06" orientation="longitude" pageId="4" pageNumber="264" precision="1" seconds="59.3" value="46.11647">46°06'59.3&quot;E</geoCoordinate>
,
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<collectingDate id="EFB49035515FFFE3FB35127EFAEB750B" box="[1223,1352,1087,1113]" pageId="4" pageNumber="264" value="2021-06-13">13.06.2021</collectingDate>
</date>
.
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<collectingCounty id="62903791515FFFE3FF351222FE29752F" box="[199,394,1123,1149]" pageId="4" pageNumber="264">Records in Iraq.</collectingCounty>
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<collectingRegion id="498A81FF515FFFE3FE631222FD8B752F" box="[401,552,1123,1149]" country="Iraq" name="Salah ad Din" pageId="4" pageNumber="264">Salah Al-Din</collectingRegion>
.
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record for
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.
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<emphasis id="B93A930F515FFFE3FF3512C6FEC275F3" bold="true" box="[199,353,1159,1185]" pageId="4" pageNumber="264">Distribution.</emphasis>
West Palaearctic, east to
<collectingRegion id="498A81FF515FFFE3FD8C12C6FD4775F3" box="[638,740,1159,1185]" country="China" name="Xinjiang Uygur" pageId="4" pageNumber="264">Xinjiang</collectingRegion>
.
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</document>

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@ -0,0 +1,109 @@
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<mods:title id="D73F1EE484F6DE8C388DA13E26115770">New data on spiders (Arachnida: Araneae) of Iraqi Kurdistan, with new species and records</mods:title>
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<mods:affiliation id="D4186E5D60E90607C2D78505624F959A">University of Sulaimani, Sulaymaniyah, Kurdistan Region, Iraq &amp; University of Mendel, Brno, Czech Republic</mods:affiliation>
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<emphasis id="B93A930F515FFFE3FF65169EFE3B71AB" bold="true" box="[151,408,223,250]" italics="true" pageId="4" pageNumber="264">Aculepeira ceropegia</emphasis>
(Walckenaer, 1802)
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<emphasis id="B93A930F515FFFE3FF651766FEA57013" bold="true" box="[151,262,295,321]" pageId="4" pageNumber="264">Material.</emphasis>
<materialsCitation id="3B264540515FFFE3FEE11766FE1E7034" collectingDate="2023-01-05" collectionCode="ZMUT" country="Iraq" latitude="34.77211" location="Sulaymaniyah Governorate" longLatPrecision="1" longitude="45.652195" pageId="4" pageNumber="264" specimenCount="1" specimenCount-female="1" stateProvince="As Sulaymaniyah">
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<collectingRegion id="498A81FF515FFFE3FE9E1766FD047010" box="[364,679,295,322]" country="Iraq" name="As Sulaymaniyah" pageId="4" pageNumber="264">Sulaymaniyah Governorate</collectingRegion>
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(
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), Garmyan, Maydan, Gorgnosh Vil.,
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,
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<collectingDate id="EFB49035515FFFE3FEC9170AFE1E7034" box="[315,445,331,358]" pageId="4" pageNumber="264" value="2023-01-05">01.05.2023</collectingDate>
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.
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<emphasis id="B93A930F515FFFE3FF35172EFEC270DB" bold="true" box="[199,353,367,393]" pageId="4" pageNumber="264">Distribution.</emphasis>
West Palaearctic. New species record for
<collectingCountry id="F3590F8D515FFFE3FCCA1731FCCA70D8" box="[824,873,368,394]" name="Iraq" pageId="4" pageNumber="264">Iraq</collectingCountry>
.
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West Palaearctic. New genus and species records for
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(
<figureCitation id="3B8A2A063E50FFDDDCDAF92BEB6FF8E2" box="[159,288,1699,1725]" captionStart="FIGURES 3748" captionStartId="15.[151,264,1812,1836]" captionTargetBox="[163,1414,207,1776]" captionTargetId="figure-19@15.[160,1427,199,1789]" captionTargetPageId="15" captionText="FIGURES 3748. Adults (all in coll. ANHRT and collected in MSR). 37. Streblote polydora, ♀. 38. Cleopatrina phocea, ♂. 39. Eutricha morosa, ♂. 40. E. fulvida, ♂. 41. Id., ♀. 42. Id., ♂. 43. E. giustii sp. n., holotype ♂ (gen. slide No. LG 6228). 44. Id., paratype ♂. 45. Pallastica cf. pallens, ♂. 46. Id., ♂. 47. Stoermeriana aculeata, ♂. 48. Id., ♂." pageId="4" pageNumber="235">Figs 4344</figureCitation>
,
<figureCitation id="3B8A2A063E50FFDDDD69F92BEB3FF8E2" box="[300,368,1699,1725]" captionStart="FIGURES 9091" captionStartId="21.[151,264,1810,1834]" captionTargetBox="[160,1419,186,1773]" captionTargetId="figure-19@21.[155,1431,181,1786]" captionTargetPageId="21" captionText="FIGURES 9091. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c) (all in coll. ANHRT). 90. Eutricha giustii sp. n., holotype, LG 6228. 91. E. fulvida, LG 6239." pageId="4" pageNumber="235">90ac</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E50FFDDDCD2F963EBD3F95A" blockId="4.[151,1437,1771,1906]" box="[151,412,1771,1798]" pageId="4" pageNumber="235">
<materialsCitation id="13D93CDE3E50FFDDDCD2F963EBD3F95A" box="[151,412,1771,1798]" collectionCode="ANHRT" pageId="4" pageNumber="235" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7C0A88213E50FFDDDCD2F963EAB0F95A" box="[151,255,1771,1797]" pageId="4" pageNumber="235" type="holotype">Holotype</typeStatus>
<specimenCount id="B5B7FD0A3E50FFDDDD43F964EB54F959" box="[262,283,1772,1798]" count="1" pageId="4" pageNumber="235" type="male"></specimenCount>
(
<collectionCode id="C5A0AE463E50FFDDDD6FF963EBDFF95A" box="[298,400,1771,1797]" pageId="4" pageNumber="235">ANHRT</collectionCode>
):
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E50FFDDDCD2F887E9C1F982" pageId="4" pageNumber="235" type="materials_examined">
<paragraph id="A30E36833E50FFDDDCD2F887EF54F92E" blockId="4.[151,1437,1771,1906]" pageId="4" pageNumber="235">
<materialsCitation id="13D93CDE3E50FFDDDCE4F887EF54F92E" collectingDate="2018-02-17" collectingDateMax="2018-02-21" collectingDateMin="2018-02-17" collectionCode="ANHRT" collectorName="Smith, L. &amp; Gy. M. Laszlo" country="Mozambique" county="Special Reserve" elevation="15" latitude="-26.44961" location="Milibangalala" longLatPrecision="1" longitude="32.924946" municipality="Ponta" pageId="4" pageNumber="235" specimenCount="1" specimenCount-male="1" stateProvince="Maputo">
<collectingCountry id="DBA676133E50FFDDDCE4F887EB2AF976" box="[161,357,1807,1833]" name="Mozambique" pageId="4" pageNumber="235">MOZAMBIQUE</collectingCountry>
<quantity id="64499B663E50FFDDDD36F887EBEAF975" box="[371,421,1807,1834]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="235" unit="m" value="15.0">
<elevation id="289CD1B03E50FFDDDD36F887EBEAF975" box="[371,421,1807,1834]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="235" unit="m" value="15.0">15m</elevation>
</quantity>
/
<collectingRegion id="6175F8613E50FFDDDD8DF898E850F975" box="[456,543,1808,1834]" country="Mozambique" name="Maputo" pageId="4" pageNumber="235">Maputo</collectingRegion>
<collectingCounty id="4A6F4E0F3E50FFDDDE68F887E8A4F975" box="[557,747,1807,1834]" pageId="4" pageNumber="235">Special Reserve</collectingCounty>
,
<collectingMunicipality id="436AACF93E50FFDDDEB8F898E972F975" box="[765,829,1808,1834]" pageId="4" pageNumber="235">Ponta</collectingMunicipality>
/
<location id="A66E60583E50FFDDDF24F887E9B2F976" LSID="urn:lsid:plazi:treatment:2B1887953E50FFDCDCD2F9F7EABEF8C6:A66E60583E50FFDDDF24F887E9B2F976" box="[865,1021,1807,1833]" country="Mozambique" county="Special Reserve" latitude="-26.44961" longLatPrecision="1" longitude="32.924946" municipality="Ponta" name="Milibangalala" pageId="4" pageNumber="235" stateProvince="Maputo">Milibangalala</location>
(
<location id="A66E60583E50FFDDD857F887EE9FF976" LSID="urn:lsid:plazi:treatment:2B1887953E50FFDCDCD2F9F7EABEF8C6:A66E60583E50FFDDD857F887EE9FF976" box="[1042,1232,1807,1833]" country="Mozambique" county="Special Reserve" latitude="-26.44961" longLatPrecision="1" longitude="32.924946" municipality="Ponta" name="Dune Grassland" pageId="4" pageNumber="235" stateProvince="Maputo">Dune Grassland</location>
) /
<geoCoordinate id="C68550443E50FFDDD8B9F887EFD7F975" box="[1276,1432,1807,1834]" degrees="26" direction="south" minutes="26" orientation="latitude" pageId="4" pageNumber="235" precision="1" seconds="58.6" value="-26.44961">26°2658.6”S</geoCoordinate>
,
<geoCoordinate id="C68550443E50FFDDDCD2F8BBEB7CF911" box="[151,307,1843,1870]" degrees="32" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="235" precision="1" seconds="29.8" value="32.924946">32°5529.8”E</geoCoordinate>
/
<date id="D70F10433E50FFDDDD08F8BBEBABF912" box="[333,484,1843,1869]" pageId="4" pageNumber="235" value="2018-02-17" valueMax="2018-02-21" valueMin="2018-02-17">
<collectingDate id="C74BE9AB3E50FFDDDD08F8BBEBABF912" box="[333,484,1843,1869]" pageId="4" pageNumber="235" value="2018-02-17" valueMax="2018-02-21" valueMin="2018-02-17">17-21.ii.2018</collectingDate>
</date>
<collectingMethod id="7AF04E943E50FFDDDDAEF8BBE88FF911" box="[491,704,1843,1870]" pageId="4" pageNumber="235">Actinic Light Trap</collectingMethod>
/ Laszlo, G., Mulvaney, J., /
<collectorName id="0E4453553E50FFDDD851F8BBEECEF911" box="[1044,1153,1843,1870]" pageId="4" pageNumber="235">Smith, L.</collectorName>
, leg. /
<collectionCode id="C5A0AE463E50FFDDD894F8BBEF7CF912" box="[1233,1331,1843,1869]" pageId="4" pageNumber="235">ANHRT</collectionCode>
:2018.2 // ANHRTUK / 00035169 // Gen. slide No. / LG 6228
<specimenCount id="B5B7FD0A3E50FFDDDEA0F8D0E8B5F92D" box="[741,762,1880,1906]" count="1" pageId="4" pageNumber="235" type="male"></specimenCount>
/ prep. by
<collectorName id="0E4453553E50FFDDDF37F8DFEE44F92E" box="[882,1035,1879,1905]" pageId="4" pageNumber="235">Gy.M. Laszlo</collectorName>
[partially handwritten]”
</materialsCitation>
</paragraph>
<paragraph id="A30E36833E50FFDDDCD2F828E82BF9E6" blockId="4.[151,910,1951,2013]" box="[151,612,1951,1978]" pageId="4" pageNumber="235">
<materialsCitation id="13D93CDE3E50FFDDDCD2F828E82FF9E6" box="[151,608,1951,1978]" collectionCode="ANHRT, MHN" pageId="4" pageNumber="235" specimenCount="26" specimenCount-male="26" typeStatus="paratype">
<typeStatus id="7C0A88213E50FFDDDCD2F828EB4BF9E5" box="[151,260,1952,1978]" pageId="4" pageNumber="235" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="B5B7FD0A3E50FFDDDD56F817EB15F9E6" box="[275,346,1951,1977]" count="26" pageId="4" pageNumber="235" type="male">26♂♂</specimenCount>
in
<collectionCode id="C5A0AE463E50FFDDDD38F817EB94F9E6" box="[381,475,1951,1977]" pageId="4" pageNumber="235">ANHRT</collectionCode>
and
<collectionCode id="C5A0AE463E50FFDDDE54F817E817F9E6" box="[529,600,1951,1977]" pageId="4" pageNumber="235">MHN</collectionCode>
)
</materialsCitation>
:
</paragraph>
<paragraph id="A30E36833E50FFDDDCD2F84BE9C1F982" blockId="4.[151,910,1951,2013]" box="[151,910,1987,2013]" pageId="4" pageNumber="235">
[3A]—
<specimenCount id="B5B7FD0A3E50FFDDDCAFF84BEB6CF982" box="[234,291,1987,2013]" count="2" pageId="4" pageNumber="235" type="male">2♂♂</specimenCount>
; [3B]—
<specimenCount id="B5B7FD0A3E50FFDDDD3AF84BEBF5F982" box="[383,442,1987,2013]" count="2" pageId="4" pageNumber="235" type="male">2♂♂</specimenCount>
; [3C]—
<specimenCount id="B5B7FD0A3E50FFDDDE53F84BE800F982" box="[534,591,1987,2013]" count="3" pageId="4" pageNumber="235" type="male">3♂♂</specimenCount>
; [3D]—
<specimenCount id="B5B7FD0A3E50FFDDDEE8F84BE8BAF982" box="[685,757,1987,2013]" count="16" pageId="4" pageNumber="235" type="male">16♂♂</specimenCount>
; [3E]—
<specimenCount id="B5B7FD0A3E50FFDDDF14F84BE9C5F982" box="[849,906,1987,2013]" count="3" pageId="4" pageNumber="235" type="male">3♂♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E51FFDCDCD2FF1EEF21FC62" pageId="5" pageNumber="236" type="diagnosis">
<paragraph id="A30E36833E51FFDCDCD2FF1EEF21FC62" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDCD2FF1EEB45FEEF" bold="true" box="[151,266,150,176]" pageId="5" pageNumber="236">Diagnosis</emphasis>
. With its uniform fawn-coloured wings,
<taxonomicName id="64B14D003E51FFDCDE94FF1FE930FEEF" authorityName="Takano &amp; László" authorityYear="2024" box="[721,895,151,177]" class="Insecta" family="Lasiocampidae" genus="Eutricha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="giustii" status="sp. nov.">
<emphasis id="91C5EA913E51FFDCDE94FF1FE930FEEF" box="[721,895,151,177]" italics="true" pageId="5" pageNumber="236">Eutricha giustii</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E51FFDCDFC3FF1FE98AFEEE" bold="true" box="[902,965,151,177]" pageId="5" pageNumber="236">
<taxonomicNameLabel id="8AF657EA3E51FFDCDFC3FF1FE98AFEEE" box="[902,965,151,177]" pageId="5" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is a highly distinctive moth that cannot be readily confused with any other species in this or any other allied genera and it is a great surprise that such an insect had evaded discovery until now. All other members of the
<taxonomicName id="64B14D003E51FFDCDF7AFF57EE15FEA6" authority="Walker, 1855" authorityName="Walker" authorityYear="1855" box="[831,1114,222,249]" class="Insecta" family="Lasiocampidae" genus="Pachypasa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E51FFDCDF7AFF57E9F5FEA6" box="[831,954,223,249]" italics="true" pageId="5" pageNumber="236">Pachypasa</emphasis>
<bibRefCitation id="C7204B723E51FFDCDF87FF57EE15FEA6" author="Walker, F." box="[962,1114,222,249]" pageId="5" pageNumber="236" pagination="1259 - 1507" refId="ref14828" refString="Walker, F. (1855) Lepidoptera Heterocera. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VI. Trustees of the British Museum (Natural History), London, pp. i - iv + 1259 - 1507." type="book chapter" year="1855">Walker, 1855</bibRefCitation>
</taxonomicName>
group revised by
<bibRefCitation id="C7204B723E51FFDCD96BFF57EBCAFF42" author="Zolotuhin, V. V. &amp; Gurkovich, A. V." pageId="5" pageNumber="236" pagination="1 - 75" refId="ref15268" refString="Zolotuhin, V. V. &amp; Gurkovich, A. V. (2009) A review of the genus Pachypasa Walker, 1855 sensu lato in Africa (Lepidoptera, Lasiocampidae). Neue Entomologische Nachrichten, 63, 1 - 75." type="book chapter" year="2009">Zolotuhin &amp; Gurkovich (2009)</bibRefCitation>
display well-defined fasciae upon darker ground-colours (ranging from brown through to slate-grey) on the forewings. The male genital configuration of the sympatric
<taxonomicName id="64B14D003E51FFDCD845FEAFEE22FF1E" baseAuthorityName="Distant" baseAuthorityYear="1897" box="[1024,1133,295,321]" class="Insecta" family="Lasiocampidae" genus="Eutricha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="fulvida">
<emphasis id="91C5EA913E51FFDCD845FEAFEE22FF1E" box="[1024,1133,295,321]" italics="true" pageId="5" pageNumber="236">E. fulvida</emphasis>
</taxonomicName>
(
<figureCitation id="3B8A2A063E51FFDCD839FEAFEE81FF1E" box="[1148,1230,295,321]" captionStart="FIGURES 9091" captionStartId="21.[151,264,1810,1834]" captionTargetBox="[160,1419,186,1773]" captionTargetId="figure-19@21.[155,1431,181,1786]" captionTargetPageId="21" captionText="FIGURES 9091. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c) (all in coll. ANHRT). 90. Eutricha giustii sp. n., holotype, LG 6228. 91. E. fulvida, LG 6239." pageId="5" pageNumber="236">Fig. 91</figureCitation>
) differs markedly, expressed by the considerably broader tegumen, shorter membranous-setose socii, and markedly larger bilobate valves where the dorsal lobe has an additional triangular medioventral process; the ventral valval lobe has a much larger, semi-circular subbasal protrusion followed by a short, finely dentate ventromedial prominence continued in a gradually tapered, apically pointed distal section. Moreover, the base of the vinculum is markedly wider, the dorsal plate of saccus has sinuous margins and the cubile is noticeably longer with narrower, more heavily sclerotized caudolateral and much wider, sharply dentate, apically pointed caudal projections. Additionally, the phallus has a much longer and more pointed dilation of the coecum penis and a markedly longer and stouter carinal process.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E51FFDCDC82FDCEEEF1FBBA" pageId="5" pageNumber="236" type="description">
<paragraph id="A30E36833E51FFDCDC82FDCEEE25FC3E" blockId="5.[151,1437,150,1689]" box="[199,1130,582,609]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDC82FDCEEB1EFC3F" bold="true" box="[199,337,582,608]" pageId="5" pageNumber="236">Description</emphasis>
. Forewing length:
<typeStatus id="7C0A88213E51FFDCDE62FDCFE8C6FC3E" box="[551,649,583,609]" pageId="5" pageNumber="236" type="holotype">holotype</typeStatus>
male:
<quantity id="64499B663E51FFDCDE90FDCFE968FC3E" box="[725,807,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="236" unit="mm" value="21.0">21 mm</quantity>
;
<typeStatus id="7C0A88213E51FFDCDF77FDCFE9DDFC3E" box="[818,914,583,609]" pageId="5" pageNumber="236" type="paratype">paratype</typeStatus>
males:
<quantity id="64499B663E51FFDCDFAFFDCFEE29FC3E" box="[1002,1126,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="5" pageNumber="236" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">2022 mm</quantity>
.
</paragraph>
<paragraph id="A30E36833E51FFDCDC82FDE3EED4FD96" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">
Male. Upperside. Ground colour of head, thorax, forewing and abdomen fawn; hindwing pale fawn. Antenna bipectinate, brown. Veins densely packed with golden scales giving definition against ground colour. Forewing elongate, rounded at apex, outer margin very gently arcuate; paler distally.
<collectingRegion id="6175F8613E51FFDCDFAAFD3BEE60FC92" box="[1007,1071,691,717]" country="Tanzania" name="Pwani" pageId="5" pageNumber="236">Costa</collectingRegion>
and fringe bronze. Anal margin with a patch of dark brown scales medially. Postmedial fascia brown, narrow, indistinct, running broadly parallel to outer margin. Hindwing rounded, fringe bronze. Ground colour darker basally and towards tornus. Underside. Ground colour as on upperside but head and legs with a ferruginous tint. Forewing darker and more ferruginous in costal half, paler distally. Postmedial fascia absent, but indicated by boundary of pale distal area. Hindwing with dark ferruginous scaling basally and along costal margin. Abdomen. Eighth segment unmodified, sternite very short, somewhat trapezoidal, anterior margin with shallow medial depression, posterior margin straight; tergite elongate quadrangular with straight anterior and lateral margins and evenly convex posterior margin.
</paragraph>
<paragraph id="A30E36833E51FFDCDC82FC5BEEF1FB26" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">Male genitalia. Uncus heavily-reduced; socius strongly reduced, represented by short, rounded, membranous, sparsely setose bulge situated near dorsal base of valve; tegumen shorter than width of valve base, with very narrow lateral plates. Valve heavily sclerotized, broad at base, very short (as long as phallus), bilobate; dorsal lobe with concave costal and straight ventral margin, subapically gently curved, apically rounded; ventral lobe with straight dorsal margin and slightly concave ventral margin following a broadly dilated, evenly rounded subbasal section; apex pointed. Juxta heavily sclerotized, semi-circular, tightly fused with phallus. Vinculum strongly modified, dorsal plate inverted-triangular with slightly concave lateral margins; ventral plate (cubile) consisting of medially deeply split, posteriorly gradually dilated elongate plates with bifurcate distal end possessing a moderately sclerotized, subapically curved apically pointed lateral projection and a shorter, heavily sclerotized, posteriorly gradually tapered, apically truncate caudal projection. Phallus strongly sclerotized, coecum penis rounded triangular, carina well-developed, robust thorn-shaped. Vesica membranous, as wide as the distal end of the phallus and almost as long as phallus, without cornuti, abruptly constricted at base of rather thin vesica ejaculatorius.</paragraph>
<paragraph id="A30E36833E51FFDCDC82FA0BEBDEFBC2" blockId="5.[151,1437,150,1689]" box="[199,401,1411,1437]" pageId="5" pageNumber="236">Female unknown.</paragraph>
<paragraph id="A30E36833E51FFDCDC82FA2EEEF1FBBA" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDC82FA2EEBDBFB9F" bold="true" box="[199,404,1446,1472]" pageId="5" pageNumber="236">DNA divergences</emphasis>
. The new species has been assigned the BIN BOLD:AFG3313. Intraspecific PWD was 0.0% (n=2) and diverged from
<taxonomicName id="64B14D003E51FFDCDDF7FA43E850FBBA" baseAuthorityName="Distant" baseAuthorityYear="1897" box="[434,543,1483,1509]" class="Insecta" family="Lasiocampidae" genus="Eutricha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="fulvida">
<emphasis id="91C5EA913E51FFDCDDF7FA43E850FBBA" box="[434,543,1483,1509]" italics="true" pageId="5" pageNumber="236">E. fulvida</emphasis>
</taxonomicName>
by 4.54.6% (n=3) and the MSR
<taxonomicName id="64B14D003E51FFDCDFDEFA43EE42FBBB" authorityName="Walker" authorityYear="1865" box="[923,1037,1483,1508]" class="Insecta" family="Lasiocampidae" genus="Lebeda" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="morosa">
<emphasis id="91C5EA913E51FFDCDFDEFA43EE42FBBB" box="[923,1037,1483,1508]" italics="true" pageId="5" pageNumber="236">E. morosa</emphasis>
</taxonomicName>
by 4.3% (n=1).
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E51FFDCDC82FA67E829F80E" pageId="5" pageNumber="236" type="etymology">
<paragraph id="A30E36833E51FFDCDC82FA67E829F80E" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDC82FA67EBD7F856" bold="true" box="[199,408,1519,1545]" italics="true" pageId="5" pageNumber="236">Derivatio nominis</emphasis>
. The new species is dedicated with great pleasure to Alessandro Giusti, Senior Curator of
<taxonomicName id="64B14D003E51FFDCDCD2F99BEBD7F872" authority=", NHMUK" authorityName="NHMUK" box="[151,408,1555,1581]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Tracheophyta" rank="order">Lepidoptera, NHMUK</taxonomicName>
, who by facilitating access to the
<taxonomicName id="64B14D003E51FFDCDF41F99BE9E1F872" authorityName="Harris" authorityYear="1841" box="[772,942,1555,1581]" class="Insecta" family="Lasiocampidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="family">Lasiocampidae</taxonomicName>
under his care, has ensured the completeness of this paper. A noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E51FFDCDC82F9D2EABEF8C6" pageId="5" pageNumber="236" type="distribution">
<paragraph id="A30E36833E51FFDCDC82F9D2EABEF8C6" blockId="5.[151,1437,150,1689]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDC82F9D2EB16F82B" bold="true" box="[199,345,1626,1652]" pageId="5" pageNumber="236">Distribution</emphasis>
. The new species is known only from its
<typeStatus id="7C0A88213E51FFDCDF62F9D3E918F82A" box="[807,855,1627,1653]" pageId="5" pageNumber="236">type</typeStatus>
locality and is almost certainly endemic to the MPA hotspot.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="91C5EA913E50FFDDDCD2FA9FEB10FB6D" bold="true" box="[151,351,1303,1330]" italics="true" pageId="4" pageNumber="235">Eutricha fulvida</emphasis>
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<bibRefCitation id="C7204B723E50FFDDDD2BFA9FE857FB6E" author="Distant, W. L." box="[366,536,1303,1329]" pageId="4" pageNumber="235" pagination="197 - 211" refId="ref12783" refString="Distant, W. L. (1897) On a collection of Heterocera made in the Transvaal. Annals and Magazine of Natural History, Series 6, 20 (116), 197 - 211. https: // doi. org / 10.1080 / 00222939708680612" type="journal article" year="1897">Distant, 1897</bibRefCitation>
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*
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(
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)
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[3D]—
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; [3E]—
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; [4E]—
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.
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<emphasis id="91C5EA913E50FFDDDCD2FA2FEB4BFB9E" bold="true" box="[151,260,1447,1473]" pageId="4" pageNumber="235">Remarks</emphasis>
. Described from a single female specimen from Pretoria,
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,
<bibRefCitation id="C7204B723E50FFDDD879FA2FEFD3FB9E" author="Zolotuhin, V. V. &amp; Gurkovich, A. V." box="[1084,1436,1447,1473]" pageId="4" pageNumber="235" pagination="1 - 75" refId="ref15268" refString="Zolotuhin, V. V. &amp; Gurkovich, A. V. (2009) A review of the genus Pachypasa Walker, 1855 sensu lato in Africa (Lepidoptera, Lasiocampidae). Neue Entomologische Nachrichten, 63, 1 - 75." type="book chapter" year="2009">Zolotuhin &amp; Gurkovich (2009)</bibRefCitation>
were only aware of
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from Witbank [=Emalahleni],
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, stating that it was a very rare species. It is reasonably common in MSR suggesting that its distribution may be centred more around the low-lying areas of southern Africa. Barcode-confirmed specimens of both sexes are figured herein.
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[4C]—
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<emphasis id="91C5EA913E50FFDDDCD2FD73EBEAFD49" bold="true" box="[151,421,763,790]" italics="true" pageId="4" pageNumber="235">Catalebeda cuneilinea</emphasis>
(
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(
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)
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[3D]—
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; [3E]—
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; [4E]—
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; [4F]—
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; [4G]—
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; [4H]—
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; [4J]—
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.
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<emphasis id="91C5EA913E50FFDDDCD2FC03EB4BFDFA" bold="true" box="[151,260,907,933]" pageId="4" pageNumber="235">Remarks</emphasis>
. Based on
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with identical barcodes from MSR, this species appears to be highly variable both in size and colour. A single Tanzanian specimen from near the
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locality of
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<emphasis id="91C5EA913E50FFDDD825FC38EE9DFD96" box="[1120,1234,943,970]" italics="true" pageId="4" pageNumber="235">C. strandi</emphasis>
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was recovered in the same BIN (BOLD:AAL9879) and it is possible that it is merely a form; however without examination of the primary
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(
<figureCitation id="3B8A2A063E50FFDDDCDAFF33EB50FE89" box="[159,287,187,214]" captionStart="FIGURES 4959" captionStartId="16.[151,264,1847,1871]" captionTargetBox="[155,1432,187,1810]" captionTargetId="figure-23@16.[151,1435,181,1822]" captionTargetPageId="16" captionText="FIGURES 4959. Adults (all in coll. ANHRT and collected in MSR). 49. Bombycopsis nigrovittata, ♂. 50. Id., ♀. 51. Id., ♂. 52. Id., ♂. 53. Id., ♂. 54. B. bipars, ♂. 55. Hypotrabala argenteoguttata, ♂. 56. Grammodora nigrolineata, ♂. 57. Id., ♀. 58. Catalebeda cuneilinea, ♂. 59. Id., ♂." pageId="4" pageNumber="235">Figs 4953</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E50FFDDDCD2FF57EB53FF41" blockId="4.[151,1437,151,286]" pageId="4" pageNumber="235">
[3B]—
<specimenCount id="B5B7FD0A3E50FFDDDCA9FF57EB43FEA6" box="[236,268,223,249]" count="1" pageId="4" pageNumber="235" type="male">1♂</specimenCount>
; [3C]—
<specimenCount id="B5B7FD0A3E50FFDDDD34FF57EBDEFEA6" box="[369,401,223,249]" count="1" pageId="4" pageNumber="235" type="male">1♂</specimenCount>
; [3D]—
<specimenCount id="B5B7FD0A3E50FFDDDDB2FF57E859FEA6" box="[503,534,223,249]" count="1" pageId="4" pageNumber="235" type="male">1♂</specimenCount>
; [4A]—
<specimenCount id="B5B7FD0A3E50FFDDDE3FFF57E8FEFEA6" box="[634,689,223,249]" count="3" pageId="4" pageNumber="235" type="male">3♂♂</specimenCount>
; [4B]—
<specimenCount id="B5B7FD0A3E50FFDDDF5EFF57E97AFEA6" box="[795,821,223,249]" count="1" pageId="4" pageNumber="235" type="female">1♀</specimenCount>
; [4C]—
<specimenCount id="B5B7FD0A3E50FFDDDFD3FF57E980FEA6" box="[918,975,223,249]" count="6" pageId="4" pageNumber="235" type="male">6♂♂</specimenCount>
; [4D]—
<specimenCount id="B5B7FD0A3E50FFDDD874FF57EE38FEA6" box="[1073,1143,223,249]" count="17" pageId="4" pageNumber="235" type="male">17♂♂</specimenCount>
; [4G]—
<specimenCount id="B5B7FD0A3E50FFDDD89BFF57EEB3FEA6" box="[1246,1276,223,249]" count="1" pageId="4" pageNumber="235" type="male">1♂</specimenCount>
; [4H]—
<specimenCount id="B5B7FD0A3E50FFDDD91AFF57EFD8FEA6" box="[1375,1431,223,249]" count="4" pageId="4" pageNumber="235" type="male">4♂♂</specimenCount>
; [4I]—
<specimenCount id="B5B7FD0A3E50FFDDDCA5FE8BEB57FF41" box="[224,280,259,286]" count="5" pageId="4" pageNumber="235" type="male">5♂♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E50FFDDDCD2FEC3E849FCF6" pageId="4" pageNumber="235" type="discussion">
<paragraph id="A30E36833E50FFDDDCD2FEC3E849FCF6" blockId="4.[151,1437,331,681]" pageId="4" pageNumber="235">
<emphasis id="91C5EA913E50FFDDDCD2FEC3EB4BFF3A" bold="true" box="[151,260,331,357]" pageId="4" pageNumber="235">Remarks</emphasis>
. In
<bibRefCitation id="C7204B723E50FFDDDD7DFEC4E836FF3A" author="Joannou, J. G. &amp; Kruger, M." box="[312,633,331,358]" pageId="4" pageNumber="235" pagination="1 - 192" refId="ref13635" refString="Joannou, J. G. &amp; Kruger, M. (2009) Revision of the genus Bombycopsis C. &amp; R. Felder, 1874 (Lepidoptera: Lasiocampoidea: Lasiocampidae: Lasiocampinae: Lasiocampini). Transvaal Museum Monograph, 14, 1 - 192." type="journal article" year="2009">Joannou &amp; Krügers (2009)</bibRefCitation>
revision of
<taxonomicName id="64B14D003E50FFDDDF4FFEC3EEB7FF3A" authority="C. Felder &amp; R. Felder, 1874" authorityName="C. Felder &amp; R. Felder" authorityYear="1874" box="[778,1272,331,358]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E50FFDDDF4FFEC3E9D1FF3A" box="[778,926,331,357]" italics="true" pageId="4" pageNumber="235">Bombycopsis</emphasis>
C. Felder &amp; R.
<bibRefCitation id="C7204B723E50FFDDD821FEC3EEB7FF3A" author="Felder, C. &amp; Felder, R." box="[1124,1272,331,357]" pageId="4" pageNumber="235" refId="ref13172" refString="Felder, C. &amp; Felder, R. (1874) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Zweiter Band. Abtheilung 2, Heft 4, Lepidoptera. Atlas der Heterocera. aus der kaiserlich-koniglichen Hof- und Staatsdruckerei, Wien, 20 pp., 140 pls." type="book" year="1874">Felder, 1874</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="64B14D003E50FFDDD943FEC4E85FFFD6" authority="Joannou &amp; Kruger, 2009" authorityName="Joannou &amp; Kruger" authorityYear="2009" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="pallida" subGenus="Bufoidia">
<emphasis id="91C5EA913E50FFDDD943FEC4EAA9FFD6" italics="true" pageId="4" pageNumber="235">B. (Bufoidia) pallida</emphasis>
<bibRefCitation id="C7204B723E50FFDDDCB5FEF8E85FFFD6" author="Joannou, J. G. &amp; Kruger, M." box="[240,528,367,394]" pageId="4" pageNumber="235" pagination="1 - 192" refId="ref13635" refString="Joannou, J. G. &amp; Kruger, M. (2009) Revision of the genus Bombycopsis C. &amp; R. Felder, 1874 (Lepidoptera: Lasiocampoidea: Lasiocampidae: Lasiocampinae: Lasiocampini). Transvaal Museum Monograph, 14, 1 - 192." type="journal article" year="2009">Joannou &amp; Krüger, 2009</bibRefCitation>
</taxonomicName>
, a sympatric new species allied to
<taxonomicName id="64B14D003E50FFDDDFE3FEF8EE05FFD6" authorityName="Aurivillius" authorityYear="1927" box="[934,1098,368,393]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="nigrovittata">
<emphasis id="91C5EA913E50FFDDDFE3FEF8EE05FFD6" box="[934,1098,368,393]" italics="true" pageId="4" pageNumber="235">B. nigrovittata</emphasis>
</taxonomicName>
, was described based mainly on phenotypic differences despite noting the latter was very variable in colouration.
<bibRefCitation id="C7204B723E50FFDDD812FE1CEF1CFFF2" author="Prozorov, A. M. &amp; Prozorova, T. A. &amp; Spitsyn, V. M. &amp; Spitsyna, E. A. &amp; Kondakov, A. V. &amp; Soboleva, A. A. &amp; Volkova, J. S. &amp; Yakovlev, R. V. &amp; Saldaitis, A. &amp; Sulak, H. E. &amp; Revay, E. E. &amp; Muller, G. C." box="[1111,1363,403,430]" pageId="4" pageNumber="235" pagination="417 - 445" refId="ref14145" refString="Prozorov, A. M., Prozorova, T. A., Spitsyn, V. M., Spitsyna, E. A., Kondakov, A. V., Soboleva, A. A., Volkova, J. S., Yakovlev, R. V., Saldaitis, A., Sulak, H. E., Revay, E. E. &amp; Muller, G. C. (2023) New records of Lasiocampidae (Lepidoptera) from Zanzibar Island with taxonomic notes and description of one new species of Odontopacha Aurivillius, 1909. Zootaxa, 5311 (3), 417 - 445. https: // doi. org / 10.11646 / zootaxa. 5311.3.6" type="journal article" year="2023">
Prozorov
<emphasis id="91C5EA913E50FFDDD883FE1CEEB6FFF2" box="[1222,1273,403,429]" italics="true" pageId="4" pageNumber="235">et al</emphasis>
. (2023)
</bibRefCitation>
found barcodes of both phenotypes to be almost identical (maximum PWD of 0.3%) and yet they maintained that “stable morphological differences” were not supported by the genetics in spite of the aforementioned variability of
<taxonomicName id="64B14D003E50FFDDD9C1FE54EB55FC46" authorityName="Aurivillius" authorityYear="1927" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="nigrovittata">
<emphasis id="91C5EA913E50FFDDD9C1FE54EB55FC46" italics="true" pageId="4" pageNumber="235">B. nigrovittata</emphasis>
</taxonomicName>
and the identical genitalia. Numerous specimens in ANHRT of
<taxonomicName id="64B14D003E50FFDDDFB5FD88EEDCFC46" authorityName="Aurivillius" authorityYear="1927" box="[1008,1171,512,537]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="nigrovittata">
<emphasis id="91C5EA913E50FFDDDFB5FD88EEDCFC46" box="[1008,1171,512,537]" italics="true" pageId="4" pageNumber="235">B. nigrovittata</emphasis>
</taxonomicName>
(including insects with the
<taxonomicName id="64B14D003E50FFDDDC85FDABEB40FC62" authorityName="Joannou &amp; Kruger" authorityYear="2009" box="[192,271,547,573]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="91C5EA913E50FFDDDC85FDABEB40FC62" box="[192,271,547,573]" italics="true" pageId="4" pageNumber="235">pallida</emphasis>
</taxonomicName>
phenotype) from throughout its range (
<collectingCountry id="DBA676133E50FFDDDE8CFDABE913FC62" box="[713,860,547,573]" name="South Africa" pageId="4" pageNumber="235">South Africa</collectingCountry>
,
<collectingCountry id="DBA676133E50FFDDDF22FDABE983FC62" box="[871,972,547,573]" name="Tanzania" pageId="4" pageNumber="235">Tanzania</collectingCountry>
and
<collectingCountry id="DBA676133E50FFDDD847FDABEE16FC62" box="[1026,1113,547,573]" name="Zambia" pageId="4" pageNumber="235">Zambia</collectingCountry>
) were recently barcoded and it is apparent that
<taxonomicName id="64B14D003E50FFDDDD24FDC0EB9FFC3E" authorityName="Joannou &amp; Kruger" authorityYear="2009" box="[353,464,583,609]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="91C5EA913E50FFDDDD24FDC0EB9FFC3E" box="[353,464,583,609]" italics="true" pageId="4" pageNumber="235">B. pallida</emphasis>
</taxonomicName>
is merely a pale form of
<taxonomicName id="64B14D003E50FFDDDEABFDC0E9DFFC3E" authorityName="Aurivillius" authorityYear="1927" box="[750,912,584,609]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="nigrovittata">
<emphasis id="91C5EA913E50FFDDDEABFDC0E9DFFC3E" box="[750,912,584,609]" italics="true" pageId="4" pageNumber="235">B. nigrovittata</emphasis>
</taxonomicName>
and the former is synonymised with the latter, thus:
<taxonomicName id="64B14D003E50FFDDDC93FDE3E977FCDA" authority="Aurivillius, 1927" authorityName="Aurivillius" authorityYear="1927" box="[214,824,619,645]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="nigrovittata" subGenus="Bufoidia">
<emphasis id="91C5EA913E50FFDDDC93FDE3E83FFCDA" box="[214,624,619,645]" italics="true" pageId="4" pageNumber="235">Bombycopsis (Bufoidia) nigrovittata</emphasis>
<bibRefCitation id="C7204B723E50FFDDDE32FDE3E977FCDA" author="Aurivillius, C." box="[631,824,619,645]" pageId="4" pageNumber="235" pagination="205 - 281" refId="ref12495" refString="Aurivillius, C. (1927) Lasiocampidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Eine Systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. Die Afrikanischen Spinner und Schwarmer. Verlag des Seitzschen Werkes, Alfred Kernen, Stuttgart, pp. 205 - 281." type="book chapter" year="1927">Aurivillius, 1927</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="64B14D003E50FFDDDF1DFDE4EF2BFCDA" authority="Joannou &amp; Kruger, 2009" authorityName="Joannou &amp; Kruger" authorityYear="2009" box="[856,1380,619,646]" class="Insecta" family="Lasiocampidae" genus="Bombycopsis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="pallida" status="syn. n." subGenus="Bufoidia">
<emphasis id="91C5EA913E50FFDDDF1DFDE4EE0CFCDA" box="[856,1091,619,645]" italics="true" pageId="4" pageNumber="235">B. (Bufoidia) pallida</emphasis>
<bibRefCitation id="C7204B723E50FFDDD80EFDE4EF2BFCDA" author="Joannou, J. G. &amp; Kruger, M." box="[1099,1380,619,646]" pageId="4" pageNumber="235" pagination="1 - 192" refId="ref13635" refString="Joannou, J. G. &amp; Kruger, M. (2009) Revision of the genus Bombycopsis C. &amp; R. Felder, 1874 (Lepidoptera: Lasiocampoidea: Lasiocampidae: Lasiocampinae: Lasiocampini). Transvaal Museum Monograph, 14, 1 - 192." type="journal article" year="2009">Joannou &amp; Krüger, 2009</bibRefCitation>
</taxonomicName>
<emphasis id="91C5EA913E50FFDDD928FDE3EAE1FCF6" bold="true" pageId="4" pageNumber="235">
<taxonomicNameLabel id="8AF657EA3E50FFDDD928FDE3EAE1FCF6" pageId="4" pageNumber="235">syn. n.</taxonomicNameLabel>
</emphasis>
(junior subjective synonymy).
</paragraph>
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</treatment>
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@ -0,0 +1,149 @@
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<paragraph id="A30E36833E51FFDCDCD2F962E85FF95B" blockId="5.[151,528,1770,1869]" box="[151,528,1770,1797]" pageId="5" pageNumber="236">
<heading id="F84681EF3E51FFDCDCD2F962E85FF95B" bold="true" box="[151,528,1770,1797]" fontSize="11" level="1" pageId="5" pageNumber="236" reason="1">
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<emphasis id="91C5EA913E51FFDCDCD2F962E85FF95B" bold="true" box="[151,528,1770,1797]" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDCD2F962EB2BF95A" bold="true" box="[151,356,1770,1797]" italics="true" pageId="5" pageNumber="236">Eutricha morosa</emphasis>
(
<bibRefCitation id="C7204B723E51FFDCDD37F962E847F95B" author="Walker, F." box="[370,520,1770,1796]" pageId="5" pageNumber="236" pagination="323 - 706" refId="ref14997" refString="Walker, F. (1865) n. k. In: List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII - Supplement Part 2. Trustees of the British Museum (Natural History), London, pp. i - iv + 323 - 706." type="book chapter" year="1865">Walker, 1865</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</subSubSection>
<subSubSection id="EBAB65083E51FFDCDCD2F887EB40F912" pageId="5" pageNumber="236" type="description">
<paragraph id="A30E36833E51FFDCDCD2F887EAB6F976" blockId="5.[151,528,1770,1869]" box="[151,249,1807,1833]" pageId="5" pageNumber="236">
(
<figureCitation id="3B8A2A063E51FFDCDCDAF887EABEF976" box="[159,241,1807,1833]" captionStart="FIGURES 3748" captionStartId="15.[151,264,1812,1836]" captionTargetBox="[163,1414,207,1776]" captionTargetId="figure-19@15.[160,1427,199,1789]" captionTargetPageId="15" captionText="FIGURES 3748. Adults (all in coll. ANHRT and collected in MSR). 37. Streblote polydora, ♀. 38. Cleopatrina phocea, ♂. 39. Eutricha morosa, ♂. 40. E. fulvida, ♂. 41. Id., ♀. 42. Id., ♂. 43. E. giustii sp. n., holotype ♂ (gen. slide No. LG 6228). 44. Id., paratype ♂. 45. Pallastica cf. pallens, ♂. 46. Id., ♂. 47. Stoermeriana aculeata, ♂. 48. Id., ♂." pageId="5" pageNumber="236">Fig. 39</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E51FFDCDCD2F8BBEB40F912" blockId="5.[151,528,1770,1869]" box="[151,271,1843,1869]" pageId="5" pageNumber="236">
[4L]—
<specimenCount id="B5B7FD0A3E51FFDCDCA9F8BBEB44F912" box="[236,267,1843,1869]" count="1" pageId="5" pageNumber="236" type="male">1♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E51FFDFDCD2F8F3E961FD66" lastPageId="6" lastPageNumber="237" pageId="5" pageNumber="236" type="discussion">
<paragraph id="A30E36833E51FFDFDCD2F8F3E961FD66" blockId="5.[151,1437,1914,2013]" lastBlockId="6.[151,1437,151,825]" lastPageId="6" lastPageNumber="237" pageId="5" pageNumber="236">
<emphasis id="91C5EA913E51FFDCDCD2F8F3EB4BF9CA" bold="true" box="[151,260,1915,1941]" pageId="5" pageNumber="236">Remarks</emphasis>
.
<taxonomicName id="64B14D003E51FFDCDD56F8F3E813F9CA" authority="Walker, 1865" authorityName="Walker" authorityYear="1865" box="[275,604,1914,1941]" class="Insecta" family="Lasiocampidae" genus="Lebeda" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="morosa">
<emphasis id="91C5EA913E51FFDCDD56F8F3EB8FF9CB" box="[275,448,1915,1941]" italics="true" pageId="5" pageNumber="236">Lebeda morosa</emphasis>
<bibRefCitation id="C7204B723E51FFDCDD82F8F3E813F9CA" author="Walker, F." box="[455,604,1914,1941]" pageId="5" pageNumber="236" pagination="323 - 706" refId="ref14997" refString="Walker, F. (1865) n. k. In: List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII - Supplement Part 2. Trustees of the British Museum (Natural History), London, pp. i - iv + 323 - 706." type="book chapter" year="1865">Walker, 1865</bibRefCitation>
</taxonomicName>
was one of several species of African moths described by Francis Walker from the collection of William DUrban (18361934). Walker stated the type locality simply as “
<collectingCountry id="DBA676133E51FFDCD891F817EF28F9E6" box="[1236,1383,1951,1977]" name="South Africa" pageId="5" pageNumber="236">South Africa</collectingCountry>
” but it can be presumed the specimens originated from the East London/King Williams Town area where DUrban was based in the early 1860s (
<bibRefCitation id="C7204B723E52FFDFDDF8FF1FE81BFEEE" author="Trimen, R." box="[445,596,151,177]" pageId="6" pageNumber="237" refId="ref14708" refString="Trimen, R. (1887) South-African Butterflies: a monograph of the extra-tropical species. Vol. I. Trubner &amp; Co., London, xiv + 355 pp, 6 pls. https: // doi. org / 10.5962 / bhl. title. 8997" type="book" year="1887">Trimen 1887</bibRefCitation>
). Upon his return to
<collectingCountry id="DBA676133E52FFDFDF7EFF1FE9D4FEEE" box="[827,923,151,177]" name="United Kingdom" pageId="6" pageNumber="237">England</collectingCountry>
, he became the first curator of RAMM where his
<taxonomicName id="64B14D003E52FFDFDC85FF33EB08FE8A" box="[192,327,187,213]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
collections were on public display as recently as the 1930s and the cabinets could be browsed by visitors at their leisure (H. Morgenroth, pers. comm.).
<bibRefCitation id="C7204B723E52FFDFDEB3FF57EE1EFEA6" author="Zolotuhin, V. V. &amp; Gurkovich, A. V." box="[758,1105,223,249]" pageId="6" pageNumber="237" pagination="1 - 75" refId="ref15268" refString="Zolotuhin, V. V. &amp; Gurkovich, A. V. (2009) A review of the genus Pachypasa Walker, 1855 sensu lato in Africa (Lepidoptera, Lasiocampidae). Neue Entomologische Nachrichten, 63, 1 - 75." type="book chapter" year="2009">Zolotuhin &amp; Gurkovich (2009)</bibRefCitation>
could not locate the
<typeStatus id="7C0A88213E52FFDFD97FFF57EFD3FEA6" box="[1338,1436,223,249]" pageId="6" pageNumber="237" type="holotype">holotype</typeStatus>
in NHMUK and designated a
<typeStatus id="7C0A88213E52FFDFDDB3FE8BE800FF42" box="[502,591,259,285]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
to “clear up the nomenclatural situation” (
<bibRefCitation id="C7204B723E52FFDFD805FE8BEAF8FF1E" author="Zolotuhin, V. V. &amp; Gurkovich, A. V." pageId="6" pageNumber="237" pagination="1 - 75" refId="ref15268" refString="Zolotuhin, V. V. &amp; Gurkovich, A. V. (2009) A review of the genus Pachypasa Walker, 1855 sensu lato in Africa (Lepidoptera, Lasiocampidae). Neue Entomologische Nachrichten, 63, 1 - 75." type="book chapter" year="2009">Zolotuhin &amp; Gurkovich 2009: 37</bibRefCitation>
). Perhaps an assumption was made that all of Walkers types are found in NHMUK (which is certainly not the case—there are many in OUMNH), or they were aware of the reference in
<bibRefCitation id="C7204B723E52FFDFDF9FFEC3EEECFF3A" author="Horn, W. &amp; Kahle, I. &amp; Friese, G. &amp; Gaedike, R." box="[986,1187,331,357]" pageId="6" pageNumber="237" refId="ref13563" refString="Horn, W., Kahle, I., Friese, G. &amp; Gaedike, R. (1990) Collectiones entomologicae. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 573 pp." type="book" year="1990">
Horn
<emphasis id="91C5EA913E52FFDFD85FFEC3EE04FF3A" box="[1050,1099,331,357]" italics="true" pageId="6" pageNumber="237">et al</emphasis>
. (1990)
</bibRefCitation>
to some of DUrbans South African
<taxonomicName id="64B14D003E52FFDFDD79FEE7EB8CFFD6" box="[316,451,367,393]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
being in NHMUK via the Joicey Bequest. The
<typeStatus id="7C0A88213E52FFDFDFA7FEE7EE74FFD6" box="[994,1083,367,393]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
designation is however invalid as it was not sufficiently in accordance with the qualifying conditions listed under Article 75.3 of the
<bibRefCitation id="C7204B723E52FFDFD947FE1BEFD8FFF2" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[1282,1431,403,429]" pageId="6" pageNumber="237" refId="ref13601" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International code of zoological nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, xx + 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
, namely Articles 75.3.4 (“reasons for believing the name-bearing type specimen to be lost or destroyed, and the steps that had been taken to trace it”) and 75.3.6 (“evidence that the
<typeStatus id="7C0A88213E52FFDFDFD4FE53E9A5FFAA" box="[913,1002,475,501]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
came as nearly as practicable from the original type locality”). The specimen designated as the
<typeStatus id="7C0A88213E52FFDFDF4FFE77E92CFC46" box="[778,867,511,537]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
in NHMUK from Mvuma in central
<collectingCountry id="DBA676133E52FFDFD945FE77EF37FC46" box="[1280,1400,511,537]" name="Zimbabwe" pageId="6" pageNumber="237">Zimbabwe</collectingCountry>
(at an elevation of
<quantity id="64499B663E52FFDFDD00FDABEBD5FC62" box="[325,410,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="237" unit="m" value="1400.0">1400 m</quantity>
a.s.l.) is over
<quantity id="64499B663E52FFDFDE7DFDAAE8D4FC62" box="[568,667,546,573]" metricMagnitude="6" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="237" unit="km" value="1500.0">1500 km</quantity>
as the crow flies from where DUrbans specimen was likely caught within the MPA in coastal
<collectingCountry id="DBA676133E52FFDFDDFFFDCFE804FC3E" box="[442,587,583,609]" name="South Africa" pageId="6" pageNumber="237">South Africa</collectingCountry>
. Based on the close proximity and similar habitat of MSR to the type locality of Walkers taxon, it is believed that the figured specimen (
<figureCitation id="3B8A2A063E52FFDFDF77FDE3E9CBFCDA" box="[818,900,619,645]" captionStart="FIGURES 3748" captionStartId="15.[151,264,1812,1836]" captionTargetBox="[163,1414,207,1776]" captionTargetId="figure-19@15.[160,1427,199,1789]" captionTargetPageId="15" captionText="FIGURES 3748. Adults (all in coll. ANHRT and collected in MSR). 37. Streblote polydora, ♀. 38. Cleopatrina phocea, ♂. 39. Eutricha morosa, ♂. 40. E. fulvida, ♂. 41. Id., ♀. 42. Id., ♂. 43. E. giustii sp. n., holotype ♂ (gen. slide No. LG 6228). 44. Id., paratype ♂. 45. Pallastica cf. pallens, ♂. 46. Id., ♂. 47. Stoermeriana aculeata, ♂. 48. Id., ♂." pageId="6" pageNumber="237">Fig. 39</figureCitation>
) refers to the true
<taxonomicName id="64B14D003E52FFDFD810FDE3EE87FCDB" authorityName="Walker" authorityYear="1865" box="[1109,1224,619,644]" class="Insecta" family="Lasiocampidae" genus="Lebeda" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="morosa">
<emphasis id="91C5EA913E52FFDFD810FDE3EE87FCDB" box="[1109,1224,619,644]" italics="true" pageId="6" pageNumber="237">E. morosa</emphasis>
</taxonomicName>
. It is interesting to note that a publicly available barcode (GWOSG024-10) of an
<taxonomicName id="64B14D003E52FFDFDF11FD07E98AFCF7" authorityName="Walker" authorityYear="1865" box="[852,965,655,680]" class="Insecta" family="Lasiocampidae" genus="Lebeda" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="morosa">
<emphasis id="91C5EA913E52FFDFDF11FD07E98AFCF7" box="[852,965,655,680]" italics="true" pageId="6" pageNumber="237">E. morosa</emphasis>
</taxonomicName>
specimen from eastern
<collectingCountry id="DBA676133E52FFDFD893FD07EF01FCF6" box="[1238,1358,655,681]" name="Zimbabwe" pageId="6" pageNumber="237">Zimbabwe</collectingCountry>
differs from MSR, Zambian and Tanzanian specimens with PWDs of 2.22.7% (n=5) suggesting that the
<typeStatus id="7C0A88213E52FFDFD891FD3BEF62FC92" box="[1236,1325,691,717]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
specimen may refer to a different taxon entirely. The natural history stores at RAMM could not be accessed during the period of this present research but it is possible that Walkers
<typeStatus id="7C0A88213E52FFDFDEB9FD73E911FD4A" box="[764,862,763,789]" pageId="6" pageNumber="237" type="holotype">holotype</typeStatus>
specimen is extant, and a new
<typeStatus id="7C0A88213E52FFDFD8FEFD73EF5BFD4A" box="[1211,1300,763,789]" pageId="6" pageNumber="237" type="neotype">neotype</typeStatus>
designation will not be made until further investigations are conducted.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="EBAB65083E52FFDFDCD2F8F6E87EF9C7" box="[151,561,1918,1945]" pageId="6" pageNumber="237" type="nomenclature">
<paragraph id="A30E36833E52FFDFDCD2F8F6E87EF9C7" blockId="6.[151,561,1918,2017]" box="[151,561,1918,1945]" pageId="6" pageNumber="237">
<heading id="F84681EF3E52FFDFDCD2F8F6E87EF9C7" bold="true" box="[151,561,1918,1945]" fontSize="11" level="1" pageId="6" pageNumber="237" reason="1">
<taxonomicName id="64B14D003E52FFDFDCD2F8F6E87EF9C7" ID-CoL="4P36S" authority="(Walker, 1865)" baseAuthorityName="Walker" baseAuthorityYear="1865" box="[151,561,1918,1945]" class="Insecta" family="Lasiocampidae" genus="Pseudometa" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="91C5EA913E52FFDFDCD2F8F6E87EF9C7" bold="true" box="[151,561,1918,1945]" pageId="6" pageNumber="237">
<emphasis id="91C5EA913E52FFDFDCD2F8F6EBCBF9C6" bold="true" box="[151,388,1918,1945]" italics="true" pageId="6" pageNumber="237">Pseudometa basalis</emphasis>
(
<bibRefCitation id="C7204B723E52FFDFDDD7F8F6E867F9C7" author="Walker, F." box="[402,552,1918,1944]" pageId="6" pageNumber="237" pagination="323 - 706" refId="ref14997" refString="Walker, F. (1865) n. k. In: List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII - Supplement Part 2. Trustees of the British Museum (Natural History), London, pp. i - iv + 323 - 706." type="book chapter" year="1865">Walker, 1865</bibRefCitation>
)
</emphasis>
</taxonomicName>
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<subSubSection id="EBAB65083E52FFDFDCD2F82BEB5DF9BE" pageId="6" pageNumber="237" type="description">
<paragraph id="A30E36833E52FFDFDCD2F82BEAB6F9E2" blockId="6.[151,561,1918,2017]" box="[151,249,1955,1981]" pageId="6" pageNumber="237">
(
<figureCitation id="3B8A2A063E52FFDFDCDAF82BEABEF9E2" box="[159,241,1955,1981]" captionStart="FIGURES 2536" captionStartId="10.[151,264,1754,1778]" captionTargetBox="[172,1415,189,1717]" captionTargetId="figure-23@10.[159,1427,181,1730]" captionTargetPageId="10" captionText="FIGURES 2536. Adults (all in coll. ANHRT and collected in MSR). 25. Typhonoya longipennis, ♂. 26. Id., ♀. 27. Id., ♂. 28. Pseudometa basalis, ♀. 29. Nirbiana micha, ♀. 30. Morongea missdebeerae, ♂. 31. Euwallengrenia reducta, ♂. 32. Id., ♀. 33. Soligena juna juna, ♂. 34. Id., ♂. 35. Id., ♀. 36. Odontopacha fenestrata, ♂." pageId="6" pageNumber="237">Fig. 28</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E52FFDFDCD2F84EEB5DF9BE" blockId="6.[151,561,1918,2017]" box="[151,274,1990,2017]" pageId="6" pageNumber="237">
[5A]—
<specimenCount id="B5B7FD0A3E52FFDFDCB1F84EEB41F9BE" box="[244,270,1990,2017]" count="1" pageId="6" pageNumber="237" type="female">1♀</specimenCount>
.
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<emphasis id="91C5EA913E52FFDFDCD2FB55EB41FAA7" bold="true" box="[151,270,1245,1272]" italics="true" pageId="6" pageNumber="237">Pallastica</emphasis>
cf.
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(
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)
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(
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)
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[2B]—
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; [3B]—
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; [4B]—
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; [4C] -
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; [4D]—
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.
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<emphasis id="91C5EA913E52FFDFDCD2FAE5EB4BFBD8" bold="true" box="[151,260,1389,1415]" pageId="6" pageNumber="237">Remarks</emphasis>
. The genus
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can be diagnosed from allied genera based on the curved processes of the cubile lacking any serrations or projections. In the puzzling original description of the genus,
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stated that the forewing pattern was “specific”, yet contradicted themselves by implicitly providing variation of forewing markings as an autapomorphy, whilst also noting that the male genitalia was not suitable for specific diagnoses; they concluded that the genus was “very difficult to determine” and that when more specimens were examined, a reliable identification was “impossible” (
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). This genus together with several other poorly-defined allied genera described by
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are in need of a full reassessment integrating genetic markers which is beyond the scope of this present paper. However, based on barcoded specimens from
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and
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, it is very apparent that both pale and dark specimens with differing phenotypes are conspecific and the genus contains a surplus of names to species.
<bibRefCitation id="C7204B723E52FFDFDCD2F95EEBC4F8B0" author="Prozorov, A. M. &amp; Prozorova, T. A. &amp; Spitsyn, V. M. &amp; Spitsyna, E. A. &amp; Kondakov, A. V. &amp; Soboleva, A. A. &amp; Volkova, J. S. &amp; Yakovlev, R. V. &amp; Saldaitis, A. &amp; Sulak, H. E. &amp; Revay, E. E. &amp; Muller, G. C." box="[151,395,1749,1776]" pageId="6" pageNumber="237" pagination="417 - 445" refId="ref14145" refString="Prozorov, A. M., Prozorova, T. A., Spitsyn, V. M., Spitsyna, E. A., Kondakov, A. V., Soboleva, A. A., Volkova, J. S., Yakovlev, R. V., Saldaitis, A., Sulak, H. E., Revay, E. E. &amp; Muller, G. C. (2023) New records of Lasiocampidae (Lepidoptera) from Zanzibar Island with taxonomic notes and description of one new species of Odontopacha Aurivillius, 1909. Zootaxa, 5311 (3), 417 - 445. https: // doi. org / 10.11646 / zootaxa. 5311.3.6" type="journal article" year="2023">
Prozorov
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suggested that there were two taxa in southern Africa, with
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, described from Chipata in
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, possibly being a species distinct from a more widely distributed eastern African taxon to which the MSR insects belong.
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<emphasis id="91C5EA913E53FFDEDCD2F9A4EB2AF818" bold="true" box="[151,357,1580,1607]" italics="true" pageId="7" pageNumber="238">Diapalpus smithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF657EA3E53FFDEDD28F9A5EBE0F818" box="[365,431,1581,1607]" pageId="7" pageNumber="238" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E53FFDEDCD2F9D8EB05F834" box="[151,330,1616,1643]" pageId="7" pageNumber="238" type="description">
<paragraph id="A30E36833E53FFDEDCD2F9D8EB05F834" blockId="7.[151,431,1580,1643]" box="[151,330,1616,1643]" pageId="7" pageNumber="238">
(
<figureCitation id="3B8A2A063E53FFDEDCDAF9D8EABCF834" box="[159,243,1616,1643]" captionStart="FIGURES 1324" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[157,1435,187,1820]" captionTargetId="figure-19@9.[151,1435,181,1833]" captionTargetPageId="9" captionText="FIGURES 1324. Adults (all in coll. ANHRT and collected in MSR unless otherwise stated). 13. Sena donaldsoni marshalli, ♂. 14. Id., ♂. 15. S. prompta, ♂. 16. Id., ♂. 17. Id., ♀. 18. Id., ♀. 19. Ocinaropsis obscura, ♂. 20. Odontocheilopteryx myxa, ♂. 21. Id., ♀. 22. Diapalpus smithi sp. n., holotype ♂ (gen. slide No. LG 6267). 23. D. griseus, Zambia, Mwinilunga, ♂ (gen. slide No. LG 6356). 24. D. congregarius, Tanzania, Iringa, ♂ (NHMUK 010606818, gen. prep. by László) (NHMUK)." pageId="7" pageNumber="238">Fig. 22</figureCitation>
,
<figureCitation id="3B8A2A063E53FFDEDCBBF9D9EB0DF834" box="[254,322,1617,1643]" captionStart="FIGURES 8789" captionStartId="20.[151,264,1304,1328]" captionTargetBox="[179,1409,188,1267]" captionTargetId="figure-23@20.[166,1422,181,1280]" captionTargetPageId="20" captionText="FIGURES 8789. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c). 87. Diapalpus smithi sp. n., holotype, LG 6267 (ANHRT). 88. D. congregarius, NHMUK 010606818 (prepared by G. László) (NHMUK). 89. D. griseus, LG 6356 (ANHRT)." pageId="7" pageNumber="238">87ac</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E53FFDEDCD2F911EB5DF9D4" pageId="7" pageNumber="238" type="materials_examined">
<paragraph id="A30E36833E53FFDEDCD2F911EBEEF8EC" blockId="7.[151,1437,1689,1823]" box="[151,417,1689,1715]" pageId="7" pageNumber="238">
<materialsCitation id="13D93CDE3E53FFDEDCD2F911EBEEF8EC" box="[151,417,1689,1715]" collectionCode="ANHRT" pageId="7" pageNumber="238" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="91C5EA913E53FFDEDCD2F911EB4BF8EC" bold="true" box="[151,260,1689,1715]" pageId="7" pageNumber="238">
<typeStatus id="7C0A88213E53FFDEDCD2F911EB4BF8EC" box="[151,260,1689,1715]" pageId="7" pageNumber="238" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="B5B7FD0A3E53FFDEDD4EF911EB6FF8EC" box="[267,288,1689,1715]" count="1" pageId="7" pageNumber="238" type="male"></specimenCount>
(
<collectionCode id="C5A0AE463E53FFDEDD6AF911EBDAF8EC" box="[303,405,1689,1715]" pageId="7" pageNumber="238">ANHRT</collectionCode>
):
</materialsCitation>
</paragraph>
<paragraph id="A30E36833E53FFDEDCD2F935EE48F941" blockId="7.[151,1437,1689,1823]" pageId="7" pageNumber="238">
<materialsCitation id="13D93CDE3E53FFDEDCE4F935EE48F941" collectingDate="2018-02-09" collectingDateMax="2018-02-17" collectingDateMin="2018-02-09" collectionCode="ANHRT" collectorName="Smith, L. &amp; Gy. M. Laszlo" country="Mozambique" county="Special Reserve" elevation="22" latitude="-26.503944" location="Sand Thicket" longLatPrecision="1" longitude="32.716557" municipality="West Gate" pageId="7" pageNumber="238" specimenCount="1" specimenCount-male="1" stateProvince="Maputo">
<collectingCountry id="DBA676133E53FFDEDCE4F935EB2AF888" box="[161,357,1725,1751]" name="Mozambique" pageId="7" pageNumber="238">MOZAMBIQUE</collectingCountry>
<quantity id="64499B663E53FFDEDD28F935EBD1F888" box="[365,414,1725,1751]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="238" unit="m" value="22.0">
<elevation id="289CD1B03E53FFDEDD28F935EBD1F888" box="[365,414,1725,1751]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="238" unit="m" value="22.0">22m</elevation>
</quantity>
/
<collectingRegion id="6175F8613E53FFDEDDF2F935E841F888" box="[439,526,1725,1751]" country="Mozambique" name="Maputo" pageId="7" pageNumber="238">Maputo</collectingRegion>
<collectingCounty id="4A6F4E0F3E53FFDEDE52F934E880F888" box="[535,719,1724,1751]" pageId="7" pageNumber="238">Special Reserve</collectingCounty>
, /
<collectingMunicipality id="436AACF93E53FFDEDEAEF935E92FF888" box="[747,864,1725,1751]" pageId="7" pageNumber="238">West Gate</collectingMunicipality>
(
<location id="A66E60583E53FFDEDF2AF934EE46F889" LSID="urn:lsid:plazi:treatment:2B1887953E53FFD1DCD2F9A4EABEF9CA:A66E60583E53FFDEDF2AF934EE46F889" box="[879,1033,1724,1750]" country="Mozambique" county="Special Reserve" latitude="-26.503944" longLatPrecision="1" longitude="32.716557" municipality="West Gate" name="Sand Thicket" pageId="7" pageNumber="238" stateProvince="Maputo">Sand Thicket</location>
) /
<geoCoordinate id="C68550443E53FFDED86DF934EE8BF889" box="[1064,1220,1724,1750]" degrees="26" direction="south" minutes="30" orientation="latitude" pageId="7" pageNumber="238" precision="1" seconds="14.2" value="-26.503944">26°3014.2”S</geoCoordinate>
,
<geoCoordinate id="C68550443E53FFDED894F934EF23F888" box="[1233,1388,1724,1751]" degrees="32" direction="east" minutes="42" orientation="longitude" pageId="7" pageNumber="238" precision="1" seconds="59.6" value="32.716557">32°4259.6”E</geoCoordinate>
/
<date id="D70F10433E53FFDED9C0F935EB47F8A5" pageId="7" pageNumber="238" value="2018-02-09" valueMax="2018-02-17" valueMin="2018-02-09">
<collectingDate id="C74BE9AB3E53FFDED9C0F935EB47F8A5" pageId="7" pageNumber="238" value="2018-02-09" valueMax="2018-02-17" valueMin="2018-02-09">9- 17.ii.2018</collectingDate>
</date>
<collectingMethod id="7AF04E943E53FFDEDD48F968EB94F8A4" box="[269,475,1760,1787]" pageId="7" pageNumber="238">Actinic Light Trap</collectingMethod>
/ Laszlo, G., Mulvaney, J., /
<collectorName id="0E4453553E53FFDEDF53F968E9CFF8A4" box="[790,896,1760,1787]" pageId="7" pageNumber="238">Smith, L.</collectorName>
, leg. /
<collectionCode id="C5A0AE463E53FFDEDF83F969EE67F8A4" box="[966,1064,1761,1787]" pageId="7" pageNumber="238">ANHRT</collectionCode>
:2018.2 // ANHRTUK / 00039664 // Gen. slide No. / LG 6267
<specimenCount id="B5B7FD0A3E53FFDEDD94F88DEBA9F940" box="[465,486,1797,1823]" count="1" pageId="7" pageNumber="238" type="male"></specimenCount>
/ prep. by
<collectorName id="0E4453553E53FFDEDE1BF88DE8B8F941" box="[606,759,1796,1823]" pageId="7" pageNumber="238">Gy.M. Laszlo</collectorName>
[partially handwritten]”
</materialsCitation>
</paragraph>
<paragraph id="A30E36833E53FFDEDCD2F8C5EBEEF938" blockId="7.[151,417,1869,1931]" box="[151,417,1869,1895]" pageId="7" pageNumber="238">
<materialsCitation id="13D93CDE3E53FFDEDCD2F8C5EBD2F938" box="[151,413,1869,1895]" collectionCode="ANHRT" pageId="7" pageNumber="238" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="91C5EA913E53FFDEDCD2F8C5EB4BF938" bold="true" box="[151,260,1869,1895]" pageId="7" pageNumber="238">
<typeStatus id="7C0A88213E53FFDEDCD2F8C5EB4BF938" box="[151,260,1869,1895]" pageId="7" pageNumber="238" type="paratype">Paratype</typeStatus>
</emphasis>
<specimenCount id="B5B7FD0A3E53FFDEDD4EF8C5EB6FF938" box="[267,288,1869,1895]" count="1" pageId="7" pageNumber="238" type="male"></specimenCount>
(
<collectionCode id="C5A0AE463E53FFDEDD6AF8C5EBDAF938" box="[303,405,1869,1895]" pageId="7" pageNumber="238">ANHRT</collectionCode>
)
</materialsCitation>
:
</paragraph>
<paragraph id="A30E36833E53FFDEDCD2F8F9EB5DF9D4" blockId="7.[151,417,1869,1931]" box="[151,274,1905,1931]" pageId="7" pageNumber="238">
[4H]—
<specimenCount id="B5B7FD0A3E53FFDEDCAAF8F9EB41F9D4" box="[239,270,1905,1931]" count="1" pageId="7" pageNumber="238" type="male">1♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E53FFD1DCD2F830EEFCFC92" lastPageId="8" lastPageNumber="239" pageId="7" pageNumber="238" type="diagnosis">
<paragraph id="A30E36833E53FFD1DCD2F830EEFCFC92" blockId="7.[151,1436,1976,2039]" lastBlockId="8.[151,1437,151,1941]" lastPageId="8" lastPageNumber="239" pageId="7" pageNumber="238">
<emphasis id="91C5EA913E53FFDEDCD2F830EB45F98D" bold="true" box="[151,266,1976,2002]" pageId="7" pageNumber="238">Diagnosis</emphasis>
.
<taxonomicName id="64B14D003E53FFDEDD5DF830EB97F98D" authorityName="Takano &amp; László" authorityYear="2024" box="[280,472,1976,2002]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="91C5EA913E53FFDEDD5DF830EB97F98D" box="[280,472,1976,2002]" italics="true" pageId="7" pageNumber="238">Diapalpus smithi</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E53FFDEDD9AF831E850F98C" bold="true" box="[479,543,1977,2003]" pageId="7" pageNumber="238">
<taxonomicNameLabel id="8AF657EA3E53FFDEDD9AF831E850F98C" box="[479,543,1977,2003]" pageId="7" pageNumber="238" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is the fourth species to be described in the genus and is most similar to the
<typeStatus id="7C0A88213E53FFDED929F831EFD3F98C" box="[1388,1436,1977,2003]" pageId="7" pageNumber="238">type</typeStatus>
species,
<taxonomicName id="64B14D003E53FFDEDCB0F855E871F9A8" authority="Strand, 1913" authorityName="Strand" authorityYear="1913" box="[245,574,2012,2039]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="congregarius">
<emphasis id="91C5EA913E53FFDEDCB0F855EBE5F9A9" box="[245,426,2013,2038]" italics="true" pageId="7" pageNumber="238">D. congregarius</emphasis>
<bibRefCitation id="C7204B723E53FFDEDDF5F854E871F9A8" author="Strand, E." box="[432,574,2012,2039]" pageId="7" pageNumber="238" pagination="67 - 69" refId="ref14561" refString="Strand, E. (1913) Eine neue afrikanische Lasiocampiden-Gattung und - Art. Archiv fur Naturgeschichte. Abteilung A, 79 (1), 67 - 69. https: // doi. org / 10.5962 / bhl. part. 11220" type="journal article" year="1913">Strand, 1913</bibRefCitation>
</taxonomicName>
(
<figureCitation id="3B8A2A063E53FFDEDE09F854E8D3F9A8" box="[588,668,2012,2039]" captionStart="FIGURES 1324" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[157,1435,187,1820]" captionTargetId="figure-19@9.[151,1435,181,1833]" captionTargetPageId="9" captionText="FIGURES 1324. Adults (all in coll. ANHRT and collected in MSR unless otherwise stated). 13. Sena donaldsoni marshalli, ♂. 14. Id., ♂. 15. S. prompta, ♂. 16. Id., ♂. 17. Id., ♀. 18. Id., ♀. 19. Ocinaropsis obscura, ♂. 20. Odontocheilopteryx myxa, ♂. 21. Id., ♀. 22. Diapalpus smithi sp. n., holotype ♂ (gen. slide No. LG 6267). 23. D. griseus, Zambia, Mwinilunga, ♂ (gen. slide No. LG 6356). 24. D. congregarius, Tanzania, Iringa, ♂ (NHMUK 010606818, gen. prep. by László) (NHMUK)." pageId="7" pageNumber="238">Fig. 24</figureCitation>
) described from
<collectingCountry id="DBA676133E53FFDEDF12F854E9F3F9A9" box="[855,956,2012,2038]" name="Tanzania" pageId="7" pageNumber="238">Tanzania</collectingCountry>
and
<taxonomicName id="64B14D003E53FFDEDFAAF855EEB5F9A8" authority="Hering, 1941" authorityName="Hering" authorityYear="1941" box="[1007,1274,2012,2039]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="91C5EA913E53FFDEDFAAF855EE2EF9A9" box="[1007,1121,2013,2038]" italics="true" pageId="7" pageNumber="238">D. griseus</emphasis>
<bibRefCitation id="C7204B723E53FFDED822F854EEB5F9A8" author="Hering, E. M." box="[1127,1274,2012,2039]" pageId="7" pageNumber="238" pagination="72 - 84" refId="ref13396" refString="Hering, E. M. (1941) Neue Heteroceren aus dem Congo-Gebiet. Revue de Zoologie et Botanique Africaines, 35 (1), 72 - 84." type="journal article" year="1941">Hering, 1941</bibRefCitation>
</taxonomicName>
(
<figureCitation id="3B8A2A063E53FFDED94DF854EF17F9A8" box="[1288,1368,2012,2039]" captionStart="FIGURES 1324" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[157,1435,187,1820]" captionTargetId="figure-19@9.[151,1435,181,1833]" captionTargetPageId="9" captionText="FIGURES 1324. Adults (all in coll. ANHRT and collected in MSR unless otherwise stated). 13. Sena donaldsoni marshalli, ♂. 14. Id., ♂. 15. S. prompta, ♂. 16. Id., ♂. 17. Id., ♀. 18. Id., ♀. 19. Ocinaropsis obscura, ♂. 20. Odontocheilopteryx myxa, ♂. 21. Id., ♀. 22. Diapalpus smithi sp. n., holotype ♂ (gen. slide No. LG 6267). 23. D. griseus, Zambia, Mwinilunga, ♂ (gen. slide No. LG 6356). 24. D. congregarius, Tanzania, Iringa, ♂ (NHMUK 010606818, gen. prep. by László) (NHMUK)." pageId="7" pageNumber="238">Fig. 23</figureCitation>
) from D.R.
<collectingCountry id="DBA676133E5CFFD1DC91FF1FEB6DFEEE" box="[212,290,151,177]" name="Democratic Republic of the Congo" pageId="8" pageNumber="239">Congo</collectingCountry>
. However, the forewing of
<taxonomicName id="64B14D003E5CFFD1DE1EFF1FE95DFEEF" authorityName="Strand" authorityYear="1913" box="[603,786,151,176]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="congregarius">
<emphasis id="91C5EA913E5CFFD1DE1EFF1FE95DFEEF" box="[603,786,151,176]" italics="true" pageId="8" pageNumber="239">D. congregarius</emphasis>
</taxonomicName>
is more elongate and its ground colour brown, the discal spot is more finely defined, and the dark submarginal spots are well-defined contrasting against a paler terminal fascia. In
<taxonomicName id="64B14D003E5CFFD1DD46FF57EB39FEA7" authorityName="Hering" authorityYear="1941" box="[259,374,223,248]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="91C5EA913E5CFFD1DD46FF57EB39FEA7" box="[259,374,223,248]" italics="true" pageId="8" pageNumber="239">D. griseus</emphasis>
</taxonomicName>
, the forewing shape is more similar to
<taxonomicName id="64B14D003E5CFFD1DF60FF57E9C4FEA6" authorityName="Takano &amp; László" authorityYear="2024" box="[805,907,223,249]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="smithi" status="sp. n.">
<emphasis id="91C5EA913E5CFFD1DF60FF57E9C4FEA6" box="[805,907,223,249]" italics="true" pageId="8" pageNumber="239">D. smithi</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E5CFFD1DFD7FF57E99FFEA6" bold="true" box="[914,976,223,249]" pageId="8" pageNumber="239">
<taxonomicNameLabel id="8AF657EA3E5CFFD1DFD7FF57E99FFEA6" box="[914,976,223,249]" pageId="8" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
but in the former, the forewing is heavily irrorated with dark greyish-brown scales, the postmedial band is indistinct and the hindwing is a paler cream. In the male genitalia,
<taxonomicName id="64B14D003E5CFFD1DD2BFEAFE86AFF1F" authorityName="Strand" authorityYear="1913" box="[366,549,295,320]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="congregarius">
<emphasis id="91C5EA913E5CFFD1DD2BFEAFE86AFF1F" box="[366,549,295,320]" italics="true" pageId="8" pageNumber="239">D. congregarius</emphasis>
</taxonomicName>
(
<figureCitation id="3B8A2A063E5CFFD1DE73FEAFE8C7FF1E" box="[566,648,295,321]" captionStart="FIGURES 8789" captionStartId="20.[151,264,1304,1328]" captionTargetBox="[179,1409,188,1267]" captionTargetId="figure-23@20.[166,1422,181,1280]" captionTargetPageId="20" captionText="FIGURES 8789. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c). 87. Diapalpus smithi sp. n., holotype, LG 6267 (ANHRT). 88. D. congregarius, NHMUK 010606818 (prepared by G. László) (NHMUK). 89. D. griseus, LG 6356 (ANHRT)." pageId="8" pageNumber="239">Fig. 88</figureCitation>
) and
<taxonomicName id="64B14D003E5CFFD1DE8DFEAFE973FF1F" authorityName="Hering" authorityYear="1941" box="[712,828,295,320]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="91C5EA913E5CFFD1DE8DFEAFE973FF1F" box="[712,828,295,320]" italics="true" pageId="8" pageNumber="239">D. griseus</emphasis>
</taxonomicName>
(
<figureCitation id="3B8A2A063E5CFFD1DF09FEAFE9D1FF1E" box="[844,926,295,321]" captionStart="FIGURES 8789" captionStartId="20.[151,264,1304,1328]" captionTargetBox="[179,1409,188,1267]" captionTargetId="figure-23@20.[166,1422,181,1280]" captionTargetPageId="20" captionText="FIGURES 8789. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c). 87. Diapalpus smithi sp. n., holotype, LG 6267 (ANHRT). 88. D. congregarius, NHMUK 010606818 (prepared by G. László) (NHMUK). 89. D. griseus, LG 6356 (ANHRT)." pageId="8" pageNumber="239">Fig. 89</figureCitation>
) are both distinguished from
<taxonomicName id="64B14D003E5CFFD1D8A8FEAFEF1BFF1E" authorityName="Takano &amp; László" authorityYear="2024" box="[1261,1364,295,321]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="smithi" status="sp. n.">
<emphasis id="91C5EA913E5CFFD1D8A8FEAFEF1BFF1E" box="[1261,1364,295,321]" italics="true" pageId="8" pageNumber="239">D. smithi</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E5CFFD1D919FEAFEFD2FF1E" bold="true" box="[1372,1437,295,321]" pageId="8" pageNumber="239">
<taxonomicNameLabel id="8AF657EA3E5CFFD1D919FEAFEFD2FF1E" box="[1372,1437,295,321]" pageId="8" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by their noticeably shorter and apically less sharply pointed posterior, and the shorter, more rounded anterior lobe of the valve. The vinculum is considerably shorter, caudally broader and more tapered in the former two species. The phalli are similarly curved medially and tapered distally in all three taxa, however the vesicae of both
<taxonomicName id="64B14D003E5CFFD1D9C4FE1BEB64FF8F" authorityName="Strand" authorityYear="1913" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="congregarius">
<emphasis id="91C5EA913E5CFFD1D9C4FE1BEB64FF8F" italics="true" pageId="8" pageNumber="239">D. congregarius</emphasis>
</taxonomicName>
and
<taxonomicName id="64B14D003E5CFFD1DD21FE3FEB97FF8F" authorityName="Hering" authorityYear="1941" box="[356,472,439,464]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="91C5EA913E5CFFD1DD21FE3FEB97FF8F" box="[356,472,439,464]" italics="true" pageId="8" pageNumber="239">D. griseus</emphasis>
</taxonomicName>
lack sclerotized dentate crests. Despite the similar ground plan of the male genitalia, the single, short triangular or thorn-like medial process of the eighth sternite readily distinguishes these two taxa from
<taxonomicName id="64B14D003E5CFFD1DC93FE77EB71FC46" authorityName="Takano &amp; László" authorityYear="2024" box="[214,318,511,537]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="smithi" status="sp. n.">
<emphasis id="91C5EA913E5CFFD1DC93FE77EB71FC46" box="[214,318,511,537]" italics="true" pageId="8" pageNumber="239">D. smithi</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E5CFFD1DD02FE77EBC7FC46" bold="true" box="[327,392,511,537]" pageId="8" pageNumber="239">
<taxonomicNameLabel id="8AF657EA3E5CFFD1DD02FE77EBC7FC46" box="[327,392,511,537]" pageId="8" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
which lacks this process and possesses a U-shaped medial notch instead. It is worth noting that the habitus of
<taxonomicName id="64B14D003E5CFFD1DD36FDABEB93FC62" authorityName="Takano &amp; László" authorityYear="2024" box="[371,476,547,573]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="smithi" status="sp. n.">
<emphasis id="91C5EA913E5CFFD1DD36FDABEB93FC62" box="[371,476,547,573]" italics="true" pageId="8" pageNumber="239">D. smithi</emphasis>
</taxonomicName>
<emphasis id="91C5EA913E5CFFD1DDA3FDABE866FC62" bold="true" box="[486,553,547,573]" pageId="8" pageNumber="239">
<taxonomicNameLabel id="8AF657EA3E5CFFD1DDA3FDABE866FC62" box="[486,553,547,573]" pageId="8" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is particularly reminiscent of members of the Madagascan genus
<taxonomicName id="64B14D003E5CFFD1D963FDABEB6BFC3E" authority="Butler, 1880" authorityName="Butler" authorityYear="1880" class="Insecta" family="Lasiocampidae" genus="Raphipeza" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E5CFFD1D963FDABEFD3FC62" box="[1318,1436,547,573]" italics="true" pageId="8" pageNumber="239">Raphipeza</emphasis>
<bibRefCitation id="C7204B723E5CFFD1DCD2FDCFEB6BFC3E" author="Butler, A. G." box="[151,292,583,609]" pageId="8" pageNumber="239" pagination="333 - 344" refId="ref12724" refString="Butler, A. G. (1880) On a collection of Lepidoptera from Madagascar with descriptions of new genera and species. Annals and Magazine of Natural History, Series 5, 5 (28 &amp; 29), 333 - 344 &amp; 384 - 395. https: // doi. org / 10.1080 / 00222938009459430" type="journal article" year="1880">Butler, 1880</bibRefCitation>
</taxonomicName>
and with the knowledge that this region of coastal southern African shares sibling or even the same taxa as
<collectingCountry id="DBA676133E5CFFD1DCACFDE3EB3FFCDA" box="[233,368,619,645]" name="Madagascar" pageId="8" pageNumber="239">Madagascar</collectingCountry>
(e.g.,
<bibRefCitation id="C7204B723E5CFFD1DDF0FDE3E8E4FCDA" author="Laszlo, G. M. &amp; Vetina, A. A." box="[437,683,619,645]" pageId="8" pageNumber="239" pagination="225 - 246" refId="ref13963" refString="Laszlo, G. M. &amp; Vetina, A. A. (2019) Contribution to the knowledge of the Nolini of the Maputo Special Reserve in South Mozambique with description of two new species and several taxonomic updates (Lepidoptera, Nolidae, Nolinae). Zootaxa, 4571 (2), 225 - 246. https: // doi. org / 10.11646 / zootaxa. 4571.2.3" type="journal article" year="2019">László &amp; Vetina 2019</bibRefCitation>
), the
<typeStatus id="7C0A88213E5CFFD1DEA0FDE3E95AFCDA" box="[741,789,619,645]" pageId="8" pageNumber="239">type</typeStatus>
species
<taxonomicName id="64B14D003E5CFFD1DF37FDE3EEC9FCDA" authority="(Butler, 1879)" baseAuthorityName="Butler" baseAuthorityYear="1879" box="[882,1158,619,645]" class="Insecta" family="Lasiocampidae" genus="Raphipeza" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="turbata">
<emphasis id="91C5EA913E5CFFD1DF37FDE3E9ACFCDA" box="[882,995,619,645]" italics="true" pageId="8" pageNumber="239">R. turbata</emphasis>
(
<bibRefCitation id="C7204B723E5CFFD1DFB4FDE3EE32FCDA" author="Butler, A. G." box="[1009,1149,619,645]" pageId="8" pageNumber="239" pagination="227 - 246" refId="ref12667" refString="Butler, A. G. (1879) Descriptions of new species of Lepidoptera from Madagascar, with notes on some of the forms already described. Annals and Magazine of Natural History, Series 5, 4 (21), 227 - 246. https: // doi. org / 10.1080 / 00222937908679820" type="journal article" year="1879">Butler, 1879</bibRefCitation>
)
</taxonomicName>
was investigated further. Although the configuration of the male genital capsule is not dissimilar, the phallus-vesica complex of
<taxonomicName id="64B14D003E5CFFD1D957FD07EFCDFCF6" baseAuthorityName="Butler" baseAuthorityYear="1879" box="[1298,1410,655,681]" class="Insecta" family="Lasiocampidae" genus="Raphipeza" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="turbata">
<emphasis id="91C5EA913E5CFFD1D957FD07EFCDFCF6" box="[1298,1410,655,681]" italics="true" pageId="8" pageNumber="239">R. turbata</emphasis>
</taxonomicName>
is fundamentally different and
<taxonomicName id="64B14D003E5CFFD1DD93FD3BE803FC92" authorityName="Butler" authorityYear="1880" box="[470,588,691,717]" class="Insecta" family="Lasiocampidae" genus="Raphipeza" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E5CFFD1DD93FD3BE803FC92" box="[470,588,691,717]" italics="true" pageId="8" pageNumber="239">Raphipeza</emphasis>
</taxonomicName>
is considered to be an allied but clearly distinct genus.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E5CFFD1DC82FD5EE96AF8BE" pageId="8" pageNumber="239" type="description">
<paragraph id="A30E36833E5CFFD1DC82FD5EE9CAFCAE" blockId="8.[151,1437,151,1941]" box="[199,901,726,753]" pageId="8" pageNumber="239">
<emphasis id="91C5EA913E5CFFD1DC82FD5EEB1EFCAF" bold="true" box="[199,337,726,752]" pageId="8" pageNumber="239">Description</emphasis>
. Forewing length:
<typeStatus id="7C0A88213E5CFFD1DE62FD5FE8C6FCAE" box="[551,649,727,753]" pageId="8" pageNumber="239" type="holotype">holotype</typeStatus>
and
<typeStatus id="7C0A88213E5CFFD1DEFAFD5FE96BFCAE" box="[703,804,727,753]" pageId="8" pageNumber="239" type="paratype">paratype</typeStatus>
:
<quantity id="64499B663E5CFFD1DF6BFD5EE9CFFCAE" box="[814,896,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="239" unit="mm" value="15.0">15 mm</quantity>
.
</paragraph>
<paragraph id="A30E36833E5CFFD1DC82FD73E9CCFB0A" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="239">Male. Upperside. Ground colour of head, thorax, forewing and abdomen pale beige; hindwing cream. Antenna bipectinate, beige. Forewing ovoid, rounded at apex, outer margin arcuate. Anal margin with long dark brown acicular scales. Basal area with light cream scaling. Antemedial fascia dark brown, bilineate, weakly sinuate, inner of two more pronounced, arising perpendicularly from costa, and angled in cell to terminate perpendicularly along anal margin. Discal spot dark brown, half width of cell. Area between antemedial and postmedial fascia with heavy irroration of dark beige scales except around discal spot; veins R5 to 1A+2A highlighted with black scales from antemedial fascia to beyond postmedial fascia. Postmedial fascia dark brown, bilineate, crenulate (interneural concavities distad), outer of two indistinct, arising perpendicularly from costa, and angled in space M1 running broadly parallel to outer margin, terminating perpendicularly along anal margin. Submarginal fasciae dark beige, poorly-defined, running parallel to outer margin. Terminal area with two dark beige spots in spaces R5 and M1. Fringe dark beige. Hindwing rounded, outer margin arcuate; paler anally. Costal margin with dark brown irroration. Medial fascia pale beige, arcuate, indistinct. Fringe pale beige. Underside. Ground colour as on upperside except for a band of dark brown scales along anterior margin of thorax. Forewing with dark brown irroration between cell and costal margin to just beyond postmedial fascia. Postmedial fascia as on upperside but lacking crenulations and poorly-defined, becoming weaker anally. Hindwing medial fascia as on upperside but dark brown and slightly better defined. Abdomen. Eighth sternite heavily sclerotized, square, possessing two short, broad, rounded, apically finely setose distal lobes encompassing a narrow U-shaped medial notch.</paragraph>
<paragraph id="A30E36833E5CFFD1DC82FAD7E9DDF80E" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="239">Male genitalia. Uncus fully reduced; socius reduced, represented by very short bulge covered in sparse, long setae. Tegumen very short, ca. twice width of lateral plates. Juxta inverted-triangular, heavily sclerotized. Valve bilobate, anterior lobe nearly half length of posterior lobe, triangular with strongly setose ventral margin; posterior lobe narrow, gradually tapered into pointed apex, slightly curved dorsad, finely setose. Vinculum as long as valve, broad, apically rounded. Phallus short and relatively thin (ca. twice as long as vinculum), tubular, medially slightly curved dorsad; coecum penis membranous, sack-like, carina short (ca. a third of length of phallus), pointed triangular, heavily sclerotized. Vesica with two parallel, elongate serrate crests.</paragraph>
<paragraph id="A30E36833E5CFFD1DC82F9D3EBDEF82A" blockId="8.[151,1437,151,1941]" box="[199,401,1627,1653]" pageId="8" pageNumber="239">Female unknown.</paragraph>
<paragraph id="A30E36833E5CFFD1DC82F9F6E96AF8BE" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="239">
<emphasis id="91C5EA913E5CFFD1DC82F9F6EBDBF8C7" bold="true" box="[199,404,1662,1688]" pageId="8" pageNumber="239">DNA divergences</emphasis>
. The new species has been assigned the BIN BOLD:AFG3475. When rooted with
<taxonomicName id="64B14D003E5CFFD1D96FF9F7EB97F8E2" authority="Prozorov &amp; Zolotuhin, 2012" authorityName="Prozorov &amp; Zolotuhin" authorityYear="2012" class="Insecta" family="Lasiocampidae" genus="Weirdonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E5CFFD1D96FF9F7EFD2F8C6" box="[1322,1437,1663,1689]" italics="true" pageId="8" pageNumber="239">Weirdonia</emphasis>
<bibRefCitation id="C7204B723E5CFFD1DCD2F92BEB97F8E2" author="Prozorov, A. M. &amp; Zolotuhin, V. V." box="[151,472,1699,1725]" pageId="8" pageNumber="239" pagination="548 - 558" refId="ref14277" refString="Prozorov, A. M. &amp; Zolotuhin, V. V. (2012) A new genus of African Lasiocampidae (Lepidoptera). Entomological Review, 92, 548 - 558. https: // doi. org / 10.1134 / S 0013873812050077" type="book chapter" year="2012">Prozorov &amp; Zolotuhin, 2012</bibRefCitation>
</taxonomicName>
, the new species is recovered as sister to the rest of
<taxonomicName id="64B14D003E5CFFD1D852F92BEEC3F8E2" authorityName="Strand" authorityYear="1913" box="[1047,1164,1699,1725]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E5CFFD1D852F92BEEC3F8E2" box="[1047,1164,1699,1725]" italics="true" pageId="8" pageNumber="239">Diapalpus</emphasis>
</taxonomicName>
. Intraspecific PWD was 0.0% (n=2) and diverged from
<taxonomicName id="64B14D003E5CFFD1DDB6F94FE829F8BF" authorityName="Hering" authorityYear="1941" box="[499,614,1735,1760]" class="Insecta" family="Lasiocampidae" genus="Diapalpus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="239" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="91C5EA913E5CFFD1DDB6F94FE829F8BF" box="[499,614,1735,1760]" italics="true" pageId="8" pageNumber="239">D. griseus</emphasis>
</taxonomicName>
by 10.0% (n=3).
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E5CFFD1DC82F963E861F912" pageId="8" pageNumber="239" type="etymology">
<paragraph id="A30E36833E5CFFD1DC82F963E861F912" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="239">
<emphasis id="91C5EA913E5CFFD1DC82F963EBD8F95A" bold="true" box="[199,407,1771,1797]" italics="true" pageId="8" pageNumber="239">Derivatio nominis</emphasis>
. The new species is dedicated with great pleasure to Richard Smith, Chairman of the Board of Trustees, ANHRT, whose dedication to African entomology has enabled the discovery of so many new species to science. A noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E5CFFD1DC82F8DEEABEF9CA" pageId="8" pageNumber="239" type="distribution">
<paragraph id="A30E36833E5CFFD1DC82F8DEEABEF9CA" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="239">
<emphasis id="91C5EA913E5CFFD1DC82F8DEEB16F92F" bold="true" box="[199,345,1878,1904]" pageId="8" pageNumber="239">Distribution</emphasis>
. The new species is known only from its
<typeStatus id="7C0A88213E5CFFD1DF62F8DFE918F92E" box="[807,855,1879,1905]" pageId="8" pageNumber="239">type</typeStatus>
locality and is almost certainly endemic to the MPA hotspot.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,80 @@
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<emphasis id="91C5EA913E53FFDEDCD2FAB8E86AFB15" bold="true" box="[151,549,1328,1355]" pageId="7" pageNumber="238">
<emphasis id="91C5EA913E53FFDEDCD2FAB8EB3BFB14" bold="true" box="[151,372,1328,1355]" italics="true" pageId="7" pageNumber="238">Streblote polydora</emphasis>
(
<bibRefCitation id="C7204B723E53FFDEDDC6FAB8E853FB15" author="Druce, H." box="[387,540,1328,1354]" pageId="7" pageNumber="238" pagination="668 - 686" refId="ref12877" refString="Druce, H. (1887) Descriptions of some new species of Lepidoptera Heterocera, mostly from tropical Africa. Proceedings of the Zoological Society of London, 1887, 668 - 686. https: // doi. org / 10.1111 / j. 1469 - 7998.1887. tb 08160. x" type="book chapter" year="1887">Druce, 1887</bibRefCitation>
)
</emphasis>
</taxonomicName>
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; [3D]—
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; [3E]—
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; [4A]—
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; [4B]—
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; [4E]—
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; [4F]—
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; [4G]—
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; [4H]—
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; [4I]—
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; [4K]—
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; [4L]—
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.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E53FFDEDCD2FC9DE8DEFA74" pageId="7" pageNumber="238" type="discussion">
<paragraph id="A30E36833E53FFDEDCD2FC9DE8DEFA74" blockId="7.[151,1437,788,1067]" pageId="7" pageNumber="238">
<emphasis id="91C5EA913E53FFDEDCD2FC9DEB44FD70" bold="true" box="[151,267,789,815]" pageId="7" pageNumber="238">Remarks.</emphasis>
This widespread southern African species is phenotypically variable, with the ground colour ranging from roseate-brown to greyish-brown. Both extremities of the colour-range have been described, the testaceous form as
<taxonomicName id="64B14D003E53FFDEDCB2FCD5E8C7FD28" authority="Walker, 1855" authorityName="Walker" authorityYear="1855" box="[247,648,860,887]" class="Insecta" family="Lasiocampidae" genus="Lasiocampa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="prompta">
<emphasis id="91C5EA913E53FFDEDCB2FCD5EBA7FD29" box="[247,488,861,886]" italics="true" pageId="7" pageNumber="238">Lasiocampa prompta</emphasis>
<bibRefCitation id="C7204B723E53FFDEDDB5FCD4E8C7FD28" author="Walker, F." box="[496,648,860,887]" pageId="7" pageNumber="238" pagination="1259 - 1507" refId="ref14828" refString="Walker, F. (1855) Lepidoptera Heterocera. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VI. Trustees of the British Museum (Natural History), London, pp. i - iv + 1259 - 1507." type="book chapter" year="1855">Walker, 1855</bibRefCitation>
</taxonomicName>
(
<typeStatus id="7C0A88213E53FFDEDEDDFCD4E8B3FD29" box="[664,764,860,886]" pageId="7" pageNumber="238" type="holotype">holotype</typeStatus>
male from “Caffraria” examined in NHMUK) and a short while later, the greyish form as
<taxonomicName id="64B14D003E53FFDEDE43FC09E9E5FDC5" authority="Walker, 1862" authorityName="Walker" authorityYear="1862" box="[518,938,896,922]" class="Insecta" family="Lasiocampidae" genus="Lasiocampa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="plusioides">
<emphasis id="91C5EA913E53FFDEDE43FC09E944FDC5" box="[518,779,896,922]" italics="true" pageId="7" pageNumber="238">Lasiocampa plusioides</emphasis>
<bibRefCitation id="C7204B723E53FFDEDF56FC08E9E5FDC5" author="Walker, F." box="[787,938,896,922]" pageId="7" pageNumber="238" pagination="263 - 279" refId="ref14938" refString="Walker, F. (1862) Characters of undescribed Lepidoptera in the collection of W. W. Saunders, Esq. Transactions of the Entomological Society of London, Series 3, 1, 263 - 279. https: // doi. org / 10.1111 / j. 1365 - 2311.1862. tb 00605. x" type="journal article" year="1862">Walker, 1862</bibRefCitation>
</taxonomicName>
(
<typeStatus id="7C0A88213E53FFDEDFFEFC09EE5BFDC4" box="[955,1044,897,923]" pageId="7" pageNumber="238" type="syntype">syntype</typeStatus>
female from “Natal” examined in OUMNH). These two taxa were treated as synonymous by
<bibRefCitation id="C7204B723E53FFDEDF76FC2CE9B0FDE0" author="Aurivillius, C." box="[819,1023,932,959]" pageId="7" pageNumber="238" pagination="205 - 281" refId="ref12495" refString="Aurivillius, C. (1927) Lasiocampidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Eine Systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. Die Afrikanischen Spinner und Schwarmer. Verlag des Seitzschen Werkes, Alfred Kernen, Stuttgart, pp. 205 - 281." type="book chapter" year="1927">Aurivillius (1927)</bibRefCitation>
and
<bibRefCitation id="C7204B723E53FFDED872FC2CEE98FDE0" author="Pinhey, E. C. G." box="[1079,1239,932,959]" pageId="7" pageNumber="238" refId="ref14118" refString="Pinhey, E. C. G. (1975) Moths of Southern Africa. Tafelberg Publishers, Cape Town, iv + 273 pp." type="book" year="1975">Pinhey (1975)</bibRefCitation>
whilst
<bibRefCitation id="C7204B723E53FFDED96FFC2CEAADFDBC" author="Vari, L. &amp; Kroon, D. M. &amp; Kruger, M." pageId="7" pageNumber="238" refId="ref14758" refString="Vari, L., Kroon, D. M. &amp; Kruger, M. (2002) Classification and Checklist of the Species of Lepidoptera Recorded in Southern Africa. Simple Solutions Australia, Chastwood, 384 pp." type="book" year="2002">
Vári
<emphasis id="91C5EA913E53FFDED921FC2DEFD2FDE1" box="[1380,1437,932,958]" italics="true" pageId="7" pageNumber="238">et al.</emphasis>
(2002)
</bibRefCitation>
and
<bibRefCitation id="C7204B723E53FFDEDD5DFC41EBF7FDBC" author="Kruger, M." box="[280,440,969,995]" pageId="7" pageNumber="238" pagination="1 - 201" refId="ref13730" refString="Kruger, M. (2020) Checklist of the Lepidoptera of southern Africa. Metamorphosis, 31 (2), 1 - 201. https: // doi. org / 10.4314 / met. v 31 i 2.2" type="journal article" year="2020">Krüger (2020)</bibRefCitation>
considered the latter as a subspecies of the former; it is however clear that these two taxa are conspecific and the synonymy is revived thus:
<taxonomicName id="64B14D003E53FFDEDE97FC65EE51FA58" authority="(Walker, 1855)" baseAuthorityName="Walker" baseAuthorityYear="1855" box="[722,1054,1004,1031]" class="Insecta" family="Lasiocampidae" genus="Sena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="prompta">
<emphasis id="91C5EA913E53FFDEDE97FC65E921FA59" box="[722,878,1005,1031]" italics="true" pageId="7" pageNumber="238">Sena prompta</emphasis>
(
<bibRefCitation id="C7204B723E53FFDEDF38FC64EE59FA58" author="Walker, F." box="[893,1046,1004,1031]" pageId="7" pageNumber="238" pagination="1259 - 1507" refId="ref14828" refString="Walker, F. (1855) Lepidoptera Heterocera. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VI. Trustees of the British Museum (Natural History), London, pp. i - iv + 1259 - 1507." type="book chapter" year="1855">Walker, 1855</bibRefCitation>
)
</taxonomicName>
=
<taxonomicName id="64B14D003E53FFDED878FC65EA80FA75" authority="Walker, 1862" authorityName="Walker" authorityYear="1862" class="Insecta" family="Lasiocampidae" genus="Lasiocampa" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="plusioides">
<emphasis id="91C5EA913E53FFDED878FC65EF0FFA59" box="[1085,1344,1004,1030]" italics="true" pageId="7" pageNumber="238">Lasiocampa plusioides</emphasis>
<bibRefCitation id="C7204B723E53FFDED903FC64EA80FA75" author="Walker, F." pageId="7" pageNumber="238" pagination="263 - 279" refId="ref14938" refString="Walker, F. (1862) Characters of undescribed Lepidoptera in the collection of W. W. Saunders, Esq. Transactions of the Entomological Society of London, Series 3, 1, 263 - 279. https: // doi. org / 10.1111 / j. 1365 - 2311.1862. tb 00605. x" type="journal article" year="1862">Walker, 1862</bibRefCitation>
</taxonomicName>
<emphasis id="91C5EA913E53FFDEDC93FB99EB76FA74" bold="true" box="[214,313,1041,1067]" pageId="7" pageNumber="238">syn. rev.</emphasis>
(junior subjective synonymy).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,132 @@
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<paragraph id="A30E36833E53FFDEDCD2FF1FE88AFEEE" blockId="7.[151,709,151,249]" box="[151,709,151,178]" pageId="7" pageNumber="238">
<heading id="F84681EF3E53FFDEDCD2FF1FE88AFEEE" bold="true" box="[151,709,151,178]" fontSize="11" level="1" pageId="7" pageNumber="238" reason="1">
<taxonomicName id="64B14D003E53FFDEDCD2FF1FE88AFEEE" ID-CoL="5L5CK" authority="(Aurivillius, 1901)" baseAuthorityName="Aurivillius" baseAuthorityYear="1901" box="[151,709,151,178]" class="Insecta" family="Lasiocampidae" genus="Sena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="subSpecies" species="donaldsoni" subSpecies="marshalli">
<emphasis id="91C5EA913E53FFDEDCD2FF1FE88AFEEE" bold="true" box="[151,709,151,178]" pageId="7" pageNumber="238">
<emphasis id="91C5EA913E53FFDEDCD2FF1FEB93FEED" bold="true" box="[151,476,151,178]" italics="true" pageId="7" pageNumber="238">Sena donaldsoni marshalli</emphasis>
(
<bibRefCitation id="C7204B723E53FFDEDDAEFF1FE8F2FEEE" author="Aurivillius, C." box="[491,701,151,177]" pageId="7" pageNumber="238" pagination="113 - 128" refId="ref12288" refString="Aurivillius, C. (1901) Diagnosen neuer Lepidopteren aus Afrika 6. Entomologisk Tidskrift, 22 (2), 113 - 128." type="journal article" year="1901">Aurivillius, 1901</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="EBAB65083E53FFDEDCD2FF33E844FEA6" pageId="7" pageNumber="238" type="description">
<paragraph id="A30E36833E53FFDEDCD2FF33EB68FE8A" blockId="7.[151,709,151,249]" box="[151,295,187,213]" pageId="7" pageNumber="238">
(
<figureCitation id="3B8A2A063E53FFDEDCDAFF33EB50FE8A" box="[159,287,187,213]" captionStart="FIGURES 1324" captionStartId="9.[151,264,1856,1880]" captionTargetBox="[157,1435,187,1820]" captionTargetId="figure-19@9.[151,1435,181,1833]" captionTargetPageId="9" captionText="FIGURES 1324. Adults (all in coll. ANHRT and collected in MSR unless otherwise stated). 13. Sena donaldsoni marshalli, ♂. 14. Id., ♂. 15. S. prompta, ♂. 16. Id., ♂. 17. Id., ♀. 18. Id., ♀. 19. Ocinaropsis obscura, ♂. 20. Odontocheilopteryx myxa, ♂. 21. Id., ♀. 22. Diapalpus smithi sp. n., holotype ♂ (gen. slide No. LG 6267). 23. D. griseus, Zambia, Mwinilunga, ♂ (gen. slide No. LG 6356). 24. D. congregarius, Tanzania, Iringa, ♂ (NHMUK 010606818, gen. prep. by László) (NHMUK)." pageId="7" pageNumber="238">Figs 1314</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E53FFDEDCD2FF57E844FEA6" blockId="7.[151,709,151,249]" box="[151,523,223,249]" pageId="7" pageNumber="238">
[4A]—
<specimenCount id="B5B7FD0A3E53FFDEDCAAFF57EB41FEA6" box="[239,270,223,249]" count="1" pageId="7" pageNumber="238" type="male">1♂</specimenCount>
; [4B]—
<specimenCount id="B5B7FD0A3E53FFDEDD35FF57EBC0FEA6" box="[368,399,223,249]" count="1" pageId="7" pageNumber="238" type="male">1♂</specimenCount>
; [4I]—
<specimenCount id="B5B7FD0A3E53FFDEDDA2FF57E848FEA6" box="[487,519,223,249]" count="1" pageId="7" pageNumber="238" type="male">1♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E53FFDEDCD2FEAFEB45FFA9" pageId="7" pageNumber="238" type="discussion">
<paragraph id="A30E36833E53FFDEDCD2FEAFEB45FFA9" blockId="7.[151,1437,295,502]" pageId="7" pageNumber="238">
<emphasis id="91C5EA913E53FFDEDCD2FEAFEB4BFF1E" bold="true" box="[151,260,295,321]" pageId="7" pageNumber="238">Remarks</emphasis>
. Dissections of specimens referable to
<taxonomicName id="64B14D003E53FFDEDEEAFEAFE929FF1E" box="[687,870,295,321]" class="Insecta" family="Lasiocampidae" genus="Sena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="donaldsoni">
<emphasis id="91C5EA913E53FFDEDEEAFEAFE929FF1E" box="[687,870,295,321]" italics="true" pageId="7" pageNumber="238">Sena donaldsoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="C7204B723E53FFDEDF36FEAFEE5BFF1E" author="Holland, W. J." box="[883,1044,295,321]" pageId="7" pageNumber="238" pagination="408 - 421" refId="ref13510" refString="Holland, W. J. (1897) List of the Lepidoptera Heterocera collected by A. Donaldson Smith. In: Donaldson Smith, A. Through unknown African countries. The first expedition from Somaliland to Lake Rudolf. Edward Arnold, London, pp. 408 - 421." type="book chapter" year="1897">
<collectingCountry id="DBA676133E53FFDEDF36FEAFE99DFF1E" box="[883,978,295,321]" name="Netherlands" pageId="7" pageNumber="238">Holland</collectingCountry>
, 1897
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) with minimal barcode divergences from northern
<collectingCountry id="DBA676133E53FFDEDD01FEC3EBE6FF3A" box="[324,425,331,357]" name="Tanzania" pageId="7" pageNumber="238">Tanzania</collectingCountry>
and MSR revealed near identical male genital configurations and the southern taxon
<taxonomicName id="64B14D003E53FFDEDCD2FEE7EB4DFFD6" authorityName="Aurivillius" authorityYear="1909" box="[151,258,367,393]" class="Insecta" family="Lasiocampidae" genus="Metajana" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="91C5EA913E53FFDEDCD2FEE7EB4DFFD6" box="[151,258,367,393]" italics="true" pageId="7" pageNumber="238">marshalli</emphasis>
</taxonomicName>
, previously treated as a form (
<bibRefCitation id="C7204B723E53FFDEDE20FEE7E96AFFD6" author="Aurivillius, C." box="[613,805,367,393]" pageId="7" pageNumber="238" pagination="205 - 281" refId="ref12495" refString="Aurivillius, C. (1927) Lasiocampidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Eine Systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. Die Afrikanischen Spinner und Schwarmer. Verlag des Seitzschen Werkes, Alfred Kernen, Stuttgart, pp. 205 - 281." type="book chapter" year="1927">Aurivillius 1927</bibRefCitation>
), synonym (
<bibRefCitation id="C7204B723E53FFDEDFF7FEE7EE0BFFD5" author="Pinhey, E. C. G." box="[946,1092,367,394]" pageId="7" pageNumber="238" refId="ref14118" refString="Pinhey, E. C. G. (1975) Moths of Southern Africa. Tafelberg Publishers, Cape Town, iv + 273 pp." type="book" year="1975">Pinhey 1975</bibRefCitation>
) and a subspecies (
<bibRefCitation id="C7204B723E53FFDED960FEE7EA9EFFF2" author="Vari, L. &amp; Kroon, D. M. &amp; Kruger, M." pageId="7" pageNumber="238" refId="ref14758" refString="Vari, L., Kroon, D. M. &amp; Kruger, M. (2002) Classification and Checklist of the Species of Lepidoptera Recorded in Southern Africa. Simple Solutions Australia, Chastwood, 384 pp." type="book" year="2002">
Vári
<emphasis id="91C5EA913E53FFDED927FEF8EFDAFFD6" box="[1378,1429,367,393]" italics="true" pageId="7" pageNumber="238">et al</emphasis>
. 2002
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) of
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<emphasis id="91C5EA913E53FFDEDD47FE1BEBD4FFF2" box="[258,411,403,429]" italics="true" pageId="7" pageNumber="238">S. donaldsoni</emphasis>
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, is at most subspecific. The barcoded MSR specimens fell into a BIN (BOLD:ACF5827) containing specimens from
<collectingCountry id="DBA676133E53FFDEDD89FE3FE815FF8E" box="[460,602,439,465]" name="South Africa" pageId="7" pageNumber="238">South Africa</collectingCountry>
and it is possible that
<taxonomicName id="64B14D003E53FFDEDF17FE3FEEF8FF8E" authority="(Aurivillius, 1922)" baseAuthorityName="Aurivillius" baseAuthorityYear="1922" box="[850,1207,439,465]" class="Insecta" family="Lasiocampidae" genus="Sena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="breyeri">
<emphasis id="91C5EA913E53FFDEDF17FE3FE990FF8E" box="[850,991,439,465]" italics="true" pageId="7" pageNumber="238">Sena breyeri</emphasis>
(
<bibRefCitation id="C7204B723E53FFDEDFA8FE3FEEE1FF8E" author="Aurivillius, C." box="[1005,1198,439,465]" pageId="7" pageNumber="238" pagination="139 - 141" refId="ref12464" refString="Aurivillius, C. (1922) New forms of Lasiocampidae from the Transvaal Museum. Annals of the Transvaal Museum, 9 (2), 139 - 141." type="journal article" year="1922">Aurivillius, 1922</bibRefCitation>
)
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is synonymous with
<taxonomicName id="64B14D003E53FFDEDCD2FE53EB4DFFAA" authorityName="Aurivillius" authorityYear="1909" box="[151,258,475,501]" class="Insecta" family="Lasiocampidae" genus="Metajana" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="238" phylum="Arthropoda" rank="species" species="marshalli">
<emphasis id="91C5EA913E53FFDEDCD2FE53EB4DFFAA" box="[151,258,475,501]" italics="true" pageId="7" pageNumber="238">marshalli</emphasis>
</taxonomicName>
.
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</subSubSection>
</treatment>
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(
<figureCitation id="3B8A2A063E56FFDBDCDAF817EABEF9E6" box="[159,241,1950,1977]" captionStart="FIGURES 4959" captionStartId="16.[151,264,1847,1871]" captionTargetBox="[155,1432,187,1810]" captionTargetId="figure-23@16.[151,1435,181,1822]" captionTargetPageId="16" captionText="FIGURES 4959. Adults (all in coll. ANHRT and collected in MSR). 49. Bombycopsis nigrovittata, ♂. 50. Id., ♀. 51. Id., ♂. 52. Id., ♂. 53. Id., ♂. 54. B. bipars, ♂. 55. Hypotrabala argenteoguttata, ♂. 56. Grammodora nigrolineata, ♂. 57. Id., ♀. 58. Catalebeda cuneilinea, ♂. 59. Id., ♂." pageId="2" pageNumber="233">Fig. 54</figureCitation>
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[4A]—
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)
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[4B]—
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<emphasis id="91C5EA913E56FFDBDCD2FAD6EBEAFB27" bold="true" box="[151,421,1374,1401]" italics="true" pageId="2" pageNumber="233">Anadiasa punctifascia</emphasis>
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)
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<emphasis id="91C5EA913E56FFDBDCD2F99BEB4BF872" bold="true" box="[151,260,1555,1581]" pageId="2" pageNumber="233">Remarks</emphasis>
. Although the three barcoded specimens from MSR fall into two BINs (BOLD:AFG5578 and BOLD: AAL9749), male genital dissections seemed to confirm that this species is phenotypically variable with some specimens displaying uniform forewing colouration (
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<emphasis id="91C5EA913E58FFD5DCD2F963EB43F95A" bold="true" box="[151,268,1771,1797]" italics="true" pageId="12" pageNumber="243">Leipoxais</emphasis>
cf.
<emphasis id="91C5EA913E58FFD5DD7CF963EBDCF959" bold="true" box="[313,403,1771,1798]" italics="true" pageId="12" pageNumber="243">acharis</emphasis>
<bibRefCitation id="C7204B723E58FFD5DDDFF963E873F95A" author="Hering, E. M." box="[410,572,1771,1797]" pageId="12" pageNumber="243" pagination="487 - 499" refId="ref13357" refString="Hering, E. M. (1928) Alte und neue Lasiocampiden von Afrika im Zoologischen Museum Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 14 (3 - 4), 487 - 499." type="journal article" year="1928">Hering, 1928</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E58FFD5DCD2F887EBC5F92D" pageId="12" pageNumber="243" type="description">
<paragraph id="A30E36833E58FFD5DCD2F887EB68F976" blockId="12.[151,1436,1771,1906]" box="[151,295,1807,1833]" pageId="12" pageNumber="243">
(
<figureCitation id="3B8A2A063E58FFD5DCDAF887EB50F976" box="[159,287,1807,1833]" captionStart="FIGURES 7986" captionStartId="19.[151,264,1327,1351]" captionTargetBox="[190,1398,190,1290]" captionTargetId="figure-19@19.[177,1410,181,1302]" captionTargetPageId="19" captionText="FIGURES 7986. Adults (all in coll.ANHRT and collected in MSR unless otherwise stated). 79. Rhinobombyx maputoensis sp. n., holotype ♂ (gen. slide No. LG 6261). 80. Id., paratype ♂. 81. Id., paratype ♂. 82. Leipoxais cf. acharis, ♂. 83. Id., ♀. 84. Id., ♂. 85. L. peraffinis, Republic of Congo, Nouabalé-Ndoki NP, ♂ (gen. slide No. LG 6276). 86. Rhinobombyx cuneata, Zambia, Kasanka NP, ♂ (gen. slide No. LG 6350)." pageId="12" pageNumber="243">Figs 8284</figureCitation>
)
</paragraph>
<paragraph id="A30E36833E58FFD5DCD2F8BBEBC5F92D" blockId="12.[151,1436,1771,1906]" pageId="12" pageNumber="243">
[2B]—
<specimenCount id="B5B7FD0A3E58FFD5DCACF8BBEB6DF912" box="[233,290,1843,1869]" count="3" pageId="12" pageNumber="243" type="male">3♂♂</specimenCount>
; [3B]—
<specimenCount id="B5B7FD0A3E58FFD5DDC7F8BBEBEDF912" box="[386,418,1843,1869]" count="1" pageId="12" pageNumber="243" type="male">1♂</specimenCount>
; [4A]—
<specimenCount id="B5B7FD0A3E58FFD5DE44F8BBE876F912" box="[513,569,1843,1869]" count="4" pageId="12" pageNumber="243" type="male">4♂♂</specimenCount>
; [4B]—
<specimenCount id="B5B7FD0A3E58FFD5DED2F8BBE89FF912" box="[663,720,1843,1869]" count="3" pageId="12" pageNumber="243" type="male">3♂♂</specimenCount>
; [4C]—
<specimenCount id="B5B7FD0A3E58FFD5DF68F8BBE92AF912" box="[813,869,1843,1869]" count="3" pageId="12" pageNumber="243" type="male">3♂♂</specimenCount>
; [4D]—
<specimenCount id="B5B7FD0A3E58FFD5DF86F8BBEE45F912" box="[963,1034,1843,1869]" count="14" pageId="12" pageNumber="243" type="male">14♂♂</specimenCount>
; [4E]—
<specimenCount id="B5B7FD0A3E58FFD5D828F8BBEED1F912" box="[1133,1182,1843,1869]" count="2" pageId="12" pageNumber="243" type="female">2♀♀</specimenCount>
; [4H]—
<specimenCount id="B5B7FD0A3E58FFD5D944F8BBEF6FF912" box="[1281,1312,1843,1869]" count="1" pageId="12" pageNumber="243" type="male">1♂</specimenCount>
; [4I]—
<specimenCount id="B5B7FD0A3E58FFD5D93DF8BBEFD7F912" box="[1400,1432,1843,1869]" count="1" pageId="12" pageNumber="243" type="male">1♂</specimenCount>
; [4J]—
<specimenCount id="B5B7FD0A3E58FFD5DCA0F8DFEB4AF92E" box="[229,261,1879,1905]" count="1" pageId="12" pageNumber="243" type="male">1♂</specimenCount>
; [5B]—
<specimenCount id="B5B7FD0A3E58FFD5DD23F8DFEBC9F92D" box="[358,390,1879,1906]" count="1" pageId="12" pageNumber="243" type="male">1♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E58FFD4DCD2F817E903FDDE" lastPageId="13" lastPageNumber="244" pageId="12" pageNumber="243" type="discussion">
<paragraph id="A30E36833E58FFD4DCD2F817E9C7FFF2" blockId="12.[151,1436,1951,2013]" lastBlockId="13.[151,1437,150,897]" lastPageId="13" lastPageNumber="244" pageId="12" pageNumber="243">
<emphasis id="91C5EA913E58FFD5DCD2F817EB4BF9E6" bold="true" box="[151,260,1951,1977]" pageId="12" pageNumber="243">Remarks</emphasis>
. Similar to many other Afrotropical
<taxonomicName id="64B14D003E58FFD5DEE1F817E901F9E6" authorityName="Harris" authorityYear="1841" box="[676,846,1951,1977]" class="Insecta" family="Lasiocampidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="family">Lasiocampidae</taxonomicName>
genera,
<taxonomicName id="64B14D003E58FFD5DFEAF828EE53F9E6" authorityName="Holland" authorityYear="1893" box="[943,1052,1952,1977]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E58FFD5DFEAF828EE53F9E6" box="[943,1052,1952,1977]" italics="true" pageId="12" pageNumber="243">Leipoxais</emphasis>
</taxonomicName>
<bibRefCitation id="C7204B723E58FFD5D866F817EE8AF9E6" author="Holland, W. J." box="[1059,1221,1951,1977]" pageId="12" pageNumber="243" pagination="373 - 376" refId="ref13427" refString="Holland, W. J. (1893) Descriptions of new species and genera of West African Lepidoptera. Psyche, 6 (201), 373 - 376 + 393 - 400 + 411 - 418 + 431 - 434 + 451 - 454 + 469 - 476 + 487 - 490 + 513 - 520 + 531 - 538 + 549 - 552 + 565 - 568. https: // doi. org / 10.1155 / 1893 / 67239" type="journal article" year="1893">
<collectingCountry id="DBA676133E58FFD5D866F817EECFF9E6" box="[1059,1152,1951,1977]" name="Netherlands" pageId="12" pageNumber="243">Holland</collectingCountry>
, 1893
</bibRefCitation>
is in need of a full holistic revision. Dissections of the male genitalia of several species revealed structurally different configurations especially in the phallus-vesica complex, suggesting the genus likely comprises several genera.
<bibRefCitation id="C7204B723E59FFD4D8FAFF1FEF2FFEEE" author="Pinhey, E. C. G." box="[1215,1376,150,177]" pageId="13" pageNumber="244" refId="ref14118" refString="Pinhey, E. C. G. (1975) Moths of Southern Africa. Tafelberg Publishers, Cape Town, iv + 273 pp." type="book" year="1975">Pinhey (1975)</bibRefCitation>
,
<bibRefCitation id="C7204B723E59FFD4D92FFF1FEB69FE8A" author="Vari, L. &amp; Kroon, D. M. &amp; Kruger, M." pageId="13" pageNumber="244" refId="ref14758" refString="Vari, L., Kroon, D. M. &amp; Kruger, M. (2002) Classification and Checklist of the Species of Lepidoptera Recorded in Southern Africa. Simple Solutions Australia, Chastwood, 384 pp." type="book" year="2002">
Vári
<emphasis id="91C5EA913E59FFD4DCD2FF33EA9DFE8A" box="[151,210,187,213]" italics="true" pageId="13" pageNumber="244">et al.</emphasis>
(2002)
</bibRefCitation>
and
<bibRefCitation id="C7204B723E59FFD4DD25FF33E84BFE8A" author="Kruger, M." box="[352,516,187,213]" pageId="13" pageNumber="244" pagination="1 - 201" refId="ref13730" refString="Kruger, M. (2020) Checklist of the Lepidoptera of southern Africa. Metamorphosis, 31 (2), 1 - 201. https: // doi. org / 10.4314 / met. v 31 i 2.2" type="journal article" year="2020">Krüger (2020)</bibRefCitation>
list only two species of
<taxonomicName id="64B14D003E59FFD4DF64FF33E9C1FE8B" authorityName="Holland" authorityYear="1893" box="[801,910,187,212]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="91C5EA913E59FFD4DF64FF33E9C1FE8B" box="[801,910,187,212]" italics="true" pageId="13" pageNumber="244">Leipoxais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF657EA3E59FFD4DFD2FF33E9F7FE8A" box="[919,952,187,213]" pageId="13" pageNumber="244" sensu="lato">s.l.</taxonomicNameLabel>
for southern Africa of which only one,
<taxonomicName id="64B14D003E59FFD4D9C3FF33EB93FEA6" authority="Aurivillius, 1911" authorityName="Aurivillius" authorityYear="1911" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="91C5EA913E59FFD4D9C3FF33EB58FEA7" italics="true" pageId="13" pageNumber="244">L. emarginata</emphasis>
<bibRefCitation id="C7204B723E59FFD4DD59FF57EB93FEA6" author="Aurivillius, C." box="[284,476,223,249]" pageId="13" pageNumber="244" pagination="161 - 167" refId="ref12407" refString="Aurivillius, C. (1911) New genera and species of African Striphnopterygidae and Lasiocampidae in the British Museum. Transactions of the Entomological Society of London, 59 (1), 161 - 167. https: // doi. org / 10.1111 / j. 1365 - 2311.1911. tb 03081. x" type="journal article" year="1911">Aurivillius, 1911</bibRefCitation>
</taxonomicName>
, is described from
<collectingCountry id="DBA676133E59FFD4DEF5FF57E90EFEA6" box="[688,833,223,249]" name="South Africa" pageId="13" pageNumber="244">South Africa</collectingCountry>
. The other is
<taxonomicName id="64B14D003E59FFD4DF93FF57EE11FEA6" box="[982,1118,223,249]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="peraffinis">
<emphasis id="91C5EA913E59FFD4DF93FF57EE11FEA6" box="[982,1118,223,249]" italics="true" pageId="13" pageNumber="244">L. peraffinis</emphasis>
</taxonomicName>
<bibRefCitation id="C7204B723E59FFD4D820FF57EF4BFEA6" author="Holland, W. J." box="[1125,1284,223,249]" pageId="13" pageNumber="244" pagination="373 - 376" refId="ref13427" refString="Holland, W. J. (1893) Descriptions of new species and genera of West African Lepidoptera. Psyche, 6 (201), 373 - 376 + 393 - 400 + 411 - 418 + 431 - 434 + 451 - 454 + 469 - 476 + 487 - 490 + 513 - 520 + 531 - 538 + 549 - 552 + 565 - 568. https: // doi. org / 10.1155 / 1893 / 67239" type="journal article" year="1893">
<collectingCountry id="DBA676133E59FFD4D820FF57EE8DFEA6" box="[1125,1218,223,249]" name="Netherlands" pageId="13" pageNumber="244">Holland</collectingCountry>
, 1893
</bibRefCitation>
(
<figureCitation id="3B8A2A063E59FFD4D956FF57EF29FEA6" box="[1299,1382,222,249]" captionStart="FIGURES 7986" captionStartId="19.[151,264,1327,1351]" captionTargetBox="[190,1398,190,1290]" captionTargetId="figure-19@19.[177,1410,181,1302]" captionTargetPageId="19" captionText="FIGURES 7986. Adults (all in coll.ANHRT and collected in MSR unless otherwise stated). 79. Rhinobombyx maputoensis sp. n., holotype ♂ (gen. slide No. LG 6261). 80. Id., paratype ♂. 81. Id., paratype ♂. 82. Leipoxais cf. acharis, ♂. 83. Id., ♀. 84. Id., ♂. 85. L. peraffinis, Republic of Congo, Nouabalé-Ndoki NP, ♂ (gen. slide No. LG 6276). 86. Rhinobombyx cuneata, Zambia, Kasanka NP, ♂ (gen. slide No. LG 6350)." pageId="13" pageNumber="244">Fig. 85</figureCitation>
), the type species of the genus described from
<collectingCountry id="DBA676133E59FFD4DE1CFE8BE8EBFF42" box="[601,676,259,285]" name="Gabon" pageId="13" pageNumber="244">Gabon</collectingCountry>
which is absent from southern Africa (current observation). Previous references (e.g.,
<bibRefCitation id="C7204B723E59FFD4DD11FEAFEBAAFF1E" author="Pinhey, E. C. G." box="[340,485,294,321]" pageId="13" pageNumber="244" refId="ref14118" refString="Pinhey, E. C. G. (1975) Moths of Southern Africa. Tafelberg Publishers, Cape Town, iv + 273 pp." type="book" year="1975">Pinhey 1975</bibRefCitation>
;
<bibRefCitation id="C7204B723E59FFD4DDB4FEAFE8E8FF1E" author="Vari, L. &amp; Kroon, D. M. &amp; Kruger, M." box="[497,679,295,321]" pageId="13" pageNumber="244" refId="ref14758" refString="Vari, L., Kroon, D. M. &amp; Kruger, M. (2002) Classification and Checklist of the Species of Lepidoptera Recorded in Southern Africa. Simple Solutions Australia, Chastwood, 384 pp." type="book" year="2002">
Vári
<emphasis id="91C5EA913E59FFD4DE6EFEAFE82AFF1E" box="[555,613,295,321]" italics="true" pageId="13" pageNumber="244">et al.</emphasis>
2002
</bibRefCitation>
) to
<taxonomicName id="64B14D003E59FFD4DE90FEAFE910FF1E" box="[725,863,295,321]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="peraffinis">
<emphasis id="91C5EA913E59FFD4DE90FEAFE910FF1E" box="[725,863,295,321]" italics="true" pageId="13" pageNumber="244">L. peraffinis</emphasis>
</taxonomicName>
likely refer to
<taxonomicName id="64B14D003E59FFD4D84EFEAFEE32FF1E" authorityName="Hering" authorityYear="1928" box="[1035,1149,295,321]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="acharis">
<emphasis id="91C5EA913E59FFD4D84EFEAFEE32FF1E" box="[1035,1149,295,321]" italics="true" pageId="13" pageNumber="244">L. acharis</emphasis>
</taxonomicName>
described from
<collectingCountry id="DBA676133E59FFD4D97DFEAFEFD2FF1E" box="[1336,1437,295,321]" name="Tanzania" pageId="13" pageNumber="244">Tanzania</collectingCountry>
which appears to be a widespread species throughout much of Africa. However, there is a second phenotypically indistinguishable taxon found sympatrically in East Africa which can only be separated based on genital characters and the
<typeStatus id="7C0A88213E59FFD4DCB5FE1BEB1DFFF2" box="[240,338,403,429]" pageId="13" pageNumber="244" type="holotype">holotype</typeStatus>
of
<taxonomicName id="64B14D003E59FFD4DD32FE1BEBA6FFF2" authorityName="Hering" authorityYear="1928" box="[375,489,403,429]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="acharis">
<emphasis id="91C5EA913E59FFD4DD32FE1BEBA6FFF2" box="[375,489,403,429]" italics="true" pageId="13" pageNumber="244">L. acharis</emphasis>
</taxonomicName>
requires dissection for confirmation.
</paragraph>
<paragraph id="A30E36833E59FFD4DC82FE3FE97EFC3E" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="244">
As part of investigations into this genus, a female
<typeStatus id="7C0A88213E59FFD4DEBBFE3FE91AFF8E" box="[766,853,439,465]" pageId="13" pageNumber="244" type="syntype">syntype</typeStatus>
of the little-known
<taxonomicName id="64B14D003E59FFD4D873FE3FEFD3FF8E" authority="Distant, 1899" authorityName="Distant" authorityYear="1899" box="[1078,1436,439,465]" class="Insecta" family="Lasiocampidae" genus="Lebeda" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="mustelina">
<emphasis id="91C5EA913E59FFD4D873FE3FEEB2FF8E" box="[1078,1277,439,465]" italics="true" pageId="13" pageNumber="244">Lebeda mustelina</emphasis>
<bibRefCitation id="C7204B723E59FFD4D940FE3FEFD3FF8E" author="Distant, W. L." box="[1285,1436,439,465]" pageId="13" pageNumber="244" pagination="461 - 465" refId="ref12831" refString="Distant, W. L. (1899) Some apparently undescribed insects from the Transvaal. Annals and Magazine of Natural History, Series 7, 3 (18), 461 - 465. https: // doi. org / 10.1080 / 00222939908678151" type="journal article" year="1899">Distant, 1899</bibRefCitation>
</taxonomicName>
from Lydenburg [=Mashishing],
<collectingCountry id="DBA676133E59FFD4DE56FE53E8E9FFAA" box="[531,678,475,501]" name="South Africa" pageId="13" pageNumber="244">South Africa</collectingCountry>
was examined in NHMUK and together with a photograph of a male with similar markings from Thabazimbi,
<collectingCountry id="DBA676133E59FFD4DEE3FE77E979FC46" box="[678,822,511,537]" name="South Africa" pageId="13" pageNumber="244">South Africa</collectingCountry>
in the research collection of H. Staude, Magaliesburg,
<collectingCountry id="DBA676133E59FFD4DCD2FDABEB67FC62" box="[151,296,547,573]" name="South Africa" pageId="13" pageNumber="244">South Africa</collectingCountry>
, it has become apparent that this taxon is congeneric with
<taxonomicName id="64B14D003E59FFD4DFF4FDABEE51FC63" authorityName="Holland" authorityYear="1893" box="[945,1054,547,572]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="91C5EA913E59FFD4DFF4FDABEE51FC63" box="[945,1054,547,572]" italics="true" pageId="13" pageNumber="244">Leipoxais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF657EA3E59FFD4D860FDABEE09FC62" box="[1061,1094,547,573]" pageId="13" pageNumber="244" sensu="lato">s.l.</taxonomicNameLabel>
resulting in the following new combination:
<taxonomicName id="64B14D003E59FFD4DD77FDCFE88CFC3E" authority="(Distant, 1899)" authorityName="Takano &amp; László" authorityYear="2024" baseAuthorityName="Distant" baseAuthorityYear="1899" box="[306,707,583,609]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="mustelina" status="comb. nov.">
<emphasis id="91C5EA913E59FFD4DD77FDCFE85CFC3E" box="[306,531,583,609]" italics="true" pageId="13" pageNumber="244">Leipoxais mustelina</emphasis>
(
<bibRefCitation id="C7204B723E59FFD4DE64FDCFE8F4FC3E" author="Distant, W. L." box="[545,699,583,609]" pageId="13" pageNumber="244" pagination="461 - 465" refId="ref12831" refString="Distant, W. L. (1899) Some apparently undescribed insects from the Transvaal. Annals and Magazine of Natural History, Series 7, 3 (18), 461 - 465. https: // doi. org / 10.1080 / 00222939908678151" type="journal article" year="1899">Distant, 1899</bibRefCitation>
)
</taxonomicName>
<emphasis id="91C5EA913E59FFD4DE8FFDCFE97EFC3E" bold="true" box="[714,817,583,609]" pageId="13" pageNumber="244">
<taxonomicNameLabel id="8AF657EA3E59FFD4DE8FFDCFE97EFC3E" box="[714,817,583,609]" pageId="13" pageNumber="244" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="A30E36833E59FFD4DC82FDE3E903FDDE" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="244">
A second species of
<taxonomicName id="64B14D003E59FFD4DDEDFDE3E85AFCDB" authorityName="Holland" authorityYear="1893" box="[424,533,619,644]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="91C5EA913E59FFD4DDEDFDE3E85AFCDB" box="[424,533,619,644]" italics="true" pageId="13" pageNumber="244">Leipoxais</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AF657EA3E59FFD4DE5EFDE3E873FCDA" box="[539,572,619,645]" pageId="13" pageNumber="244" sensu="lato">s.l.</taxonomicNameLabel>
recorded from MSR has proved to be undescribed and was recovered as sister to
<taxonomicName id="64B14D003E59FFD4DCF3FD07E855FCF6" authority="Aurivillius, 1879" authorityName="Aurivillius" authorityYear="1879" box="[182,538,655,681]" class="Insecta" family="Lasiocampidae" genus="Rhinobombyx" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="91C5EA913E59FFD4DCF3FD07EB1FFCF6" box="[182,336,655,681]" italics="true" pageId="13" pageNumber="244">Rhinobombyx</emphasis>
<bibRefCitation id="C7204B723E59FFD4DD1DFD07E855FCF6" author="Aurivillius, C." box="[344,538,655,681]" pageId="13" pageNumber="244" pagination="39 - 69" refId="ref12214" refString="Aurivillius, C. (1879) Lepidoptera Damarensia. Forteckning pa fjarilar insamlade i Damaralandet af G. de Vylder aren 1873 och 1874 jemte beskrifning ofver forut okanda arter. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 36 (7), 39 - 69." type="journal article" year="1879">
Aurivillius,
<quantity id="64499B663E59FFD4DDA7FD07E855FCF6" box="[482,538,655,681]" metricMagnitude="1" metricUnit="m" metricValue="4.77266" pageId="13" pageNumber="244" unit="in" value="1879.0">1879</quantity>
</bibRefCitation>
</taxonomicName>
in the barcode analysis. This result was perhaps not all that surprising—aside from its distinctive grey colouration, the habitus, wing shape, resting position and larva (see
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Staude
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. 2023
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) of the
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species,
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(
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) are very similar to
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. Moreover, there are close affinities in the male genitalia of the two (
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), sharing the small overall size of the capsule, the short valval lobes, the robust triangular carinal process of the phallus and the unilobate configuration of the vesica. Based on the different ground plan of the genitalia in
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, as well as the results of the barcode analysis, the new species from MSR is attributed to
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)
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[4A]—
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<emphasis id="91C5EA913E58FFD5DCD2FF1FEB88FEEE" bold="true" box="[151,455,151,178]" italics="true" pageId="12" pageNumber="243">Trichopisthia igneotincta</emphasis>
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<bibRefCitation id="C7204B723E58FFD5DD92FF1FE8E6FEEE" author="Aurivillius, C." box="[471,681,151,177]" pageId="12" pageNumber="243" pagination="1 - 29" refId="ref12365" refString="Aurivillius, C. (1909) Diagnosen neuer Lepidopteren aus Afrika. 9. Arkiv for Zoologi, 5 (5), 1 - 29. https: // doi. org / 10.5962 / bhl. part. 3496" type="journal article" year="1909">Aurivillius, 1909</bibRefCitation>
)
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(
<figureCitation id="3B8A2A063E58FFD5DCDAFF33EB50FE89" box="[159,287,187,214]" captionStart="FIGURES 6069" captionStartId="17.[151,264,1775,1799]" captionTargetBox="[166,1432,188,1738]" captionTargetId="figure-19@17.[153,1434,181,1750]" captionTargetPageId="17" captionText="FIGURES 6069. Adults (all in coll. ANHRT and collected in MSR). 60. Mallocampa leighi, ♂. 61. Id., ♂. 62. Id., ♂. 63. Id., ♀. 64. Trichopisthia monteiroi, ♂. 65. T. igneotincta, ♂. 66. Id., ♂. 67. Id., ♂. 68. Id., ♀. 69. Id., ♀." pageId="12" pageNumber="243">Figs 6569</figureCitation>
)
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[4A]—
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; [4B]—
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.
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<emphasis id="91C5EA913E58FFD5DCD2FEAFEB4BFF1E" bold="true" box="[151,260,295,321]" pageId="12" pageNumber="243">Remarks</emphasis>
. Three barcoded specimens from MSR with varying colouration and pattern were recovered in the same BIN (BOLD:AAJ8392) supporting the viewpoint of both
<bibRefCitation id="C7204B723E58FFD5DF66FEC3E9A0FF3A" author="Aurivillius, C." box="[803,1007,331,357]" pageId="12" pageNumber="243" pagination="205 - 281" refId="ref12495" refString="Aurivillius, C. (1927) Lasiocampidae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Eine Systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. Die Afrikanischen Spinner und Schwarmer. Verlag des Seitzschen Werkes, Alfred Kernen, Stuttgart, pp. 205 - 281." type="book chapter" year="1927">Aurivillius (1927)</bibRefCitation>
and
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that
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is a single variable species. It is worth noting that specimens from southern Africa (southern
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and
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<emphasis id="91C5EA913E59FFD4DCD2FC5AE8C0FDB3" bold="true" box="[151,655,978,1005]" pageId="13" pageNumber="244">
<emphasis id="91C5EA913E59FFD4DCD2FC5AEBF5FDB3" bold="true" box="[151,442,978,1005]" italics="true" pageId="13" pageNumber="244">Odontopacha fenestrata</emphasis>
<bibRefCitation id="C7204B723E59FFD4DD85FC5AE8C0FDB3" author="Aurivillius, C." box="[448,655,978,1004]" pageId="13" pageNumber="244" pagination="1 - 29" refId="ref12365" refString="Aurivillius, C. (1909) Diagnosen neuer Lepidopteren aus Afrika. 9. Arkiv for Zoologi, 5 (5), 1 - 29. https: // doi. org / 10.5962 / bhl. part. 3496" type="journal article" year="1909">Aurivillius, 1909</bibRefCitation>
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(
<figureCitation id="3B8A2A063E59FFD4DCDAFC7FEABEFA4E" box="[159,241,1015,1041]" captionStart="FIGURES 2536" captionStartId="10.[151,264,1754,1778]" captionTargetBox="[172,1415,189,1717]" captionTargetId="figure-23@10.[159,1427,181,1730]" captionTargetPageId="10" captionText="FIGURES 2536. Adults (all in coll. ANHRT and collected in MSR). 25. Typhonoya longipennis, ♂. 26. Id., ♀. 27. Id., ♂. 28. Pseudometa basalis, ♀. 29. Nirbiana micha, ♀. 30. Morongea missdebeerae, ♂. 31. Euwallengrenia reducta, ♂. 32. Id., ♀. 33. Soligena juna juna, ♂. 34. Id., ♂. 35. Id., ♀. 36. Odontopacha fenestrata, ♂." pageId="13" pageNumber="244">Fig. 36</figureCitation>
)
</paragraph>
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[4I]—
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</treatment>
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View file

@ -0,0 +1,217 @@
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(
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,
<figureCitation id="3B8A2A063E59FFD4DD69FB23EB06FA9A" box="[300,329,1195,1221]" captionStart="FIGURES 9294" captionStartId="22.[151,264,1412,1436]" captionTargetBox="[158,1435,187,1375]" captionTargetId="figure-23@22.[151,1435,181,1387]" captionTargetPageId="22" captionText="FIGURES 9294. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c) (all in coll.ANHRT).92. Rhinobombyx maputoensis sp. n., holotype, LG 6261 (cubile attached to abdomen). 93. R. cuneata, LG 6350. 94. Leipoxais peraffinis, LG 6276 (cubile attached to abdomen)." pageId="13" pageNumber="244">92</figureCitation>
)
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Actinic LightTrap / Laszlo, G., Mulvaney, J., /
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, leg. /
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/ prep. by
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[partially handwritten]”
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<typeStatus id="7C0A88213E59FFD4DCD2FA2FEB40FB9E" box="[151,271,1447,1473]" pageId="13" pageNumber="244" type="paratype">Paratypes</typeStatus>
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(
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in
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and
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; [4G]—
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; [4I]—
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; [4J]—
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.
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<emphasis id="91C5EA913E59FFD4DCD2F99AEB45F873" bold="true" box="[151,266,1554,1580]" pageId="13" pageNumber="244">Diagnosis</emphasis>
.
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<emphasis id="91C5EA913E59FFD4DD65F99BE81AF873" box="[288,597,1555,1581]" italics="true" pageId="13" pageNumber="244">Rhinobombyx maputoensis</emphasis>
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<taxonomicNameLabel id="8AF657EA3E59FFD4DE26F99BE8E5F872" box="[611,682,1555,1581]" pageId="13" pageNumber="244" rank="species">sp. n.</taxonomicNameLabel>
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is phenotypically reminiscent of
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<emphasis id="91C5EA913E59FFD4D803F99BEF62F872" box="[1094,1325,1555,1581]" italics="true" pageId="13" pageNumber="244">Leipoxais peraffinis</emphasis>
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(
<typeStatus id="7C0A88213E59FFD4D906F99BEFD3F872" box="[1347,1436,1555,1581]" pageId="13" pageNumber="244" type="syntype">syntype</typeStatus>
examined in NHMUK), but in the latter, the medial fascia of the forewing upperside is irregularly but strongly crenulate, a diffuse brown postmedial fascia is present and the terminal area is heavily irrorated with silvery-grey scales. In the male genitalia,
<taxonomicName id="64B14D003E59FFD4DD93F9F7E811F8C6" box="[470,606,1663,1689]" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="peraffinis">
<emphasis id="91C5EA913E59FFD4DD93F9F7E811F8C6" box="[470,606,1663,1689]" italics="true" pageId="13" pageNumber="244">L. peraffinis</emphasis>
</taxonomicName>
(
<figureCitation id="3B8A2A063E59FFD4DE29F9F7E8F2F8C6" box="[620,701,1663,1689]" captionStart="FIGURES 9294" captionStartId="22.[151,264,1412,1436]" captionTargetBox="[158,1435,187,1375]" captionTargetId="figure-23@22.[151,1435,181,1387]" captionTargetPageId="22" captionText="FIGURES 9294. Male genitalia (a: clasping apparatus, b: phallus) and eighth sternite (c) (all in coll.ANHRT).92. Rhinobombyx maputoensis sp. n., holotype, LG 6261 (cubile attached to abdomen). 93. R. cuneata, LG 6350. 94. Leipoxais peraffinis, LG 6276 (cubile attached to abdomen)." pageId="13" pageNumber="244">Fig. 94</figureCitation>
) has ca. three times longer valval lobes where the proximal lobe is very narrow digitiform and the distal one is strongly curved caudad, and a markedly longer, sub-quadrangular cubile with a medially slightly depressed anterior margin without well-developed anterolateral processes. Additionally,
<taxonomicName id="64B14D003E59FFD4D9C3F94FEB4DF95A" class="Insecta" family="Lasiocampidae" genus="Leipoxais" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="peraffinis">
<emphasis id="91C5EA913E59FFD4D9C3F94FEB4DF95A" italics="true" pageId="13" pageNumber="244">L. peraffinis</emphasis>
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possesses a characteristically bilobate vesica as well as a very wide, dentate carinal plate of the phallus.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E59FFD7DC82F886EE96FA4E" lastPageId="14" lastPageNumber="245" pageId="13" pageNumber="244" type="description">
<paragraph id="A30E36833E59FFD4DC82F886E9A0F976" blockId="13.[151,1437,1554,2013]" box="[199,1007,1806,1833]" pageId="13" pageNumber="244">
<emphasis id="91C5EA913E59FFD4DC82F886EB1EF977" bold="true" box="[199,337,1806,1832]" pageId="13" pageNumber="244">Description</emphasis>
. Forewing length:
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:
<quantity id="64499B663E59FFD4DEDDF887E8A5F976" box="[664,746,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="244" unit="mm" value="17.0">17 mm</quantity>
;
<typeStatus id="7C0A88213E59FFD4DEB0F887E92AF976" box="[757,869,1807,1833]" pageId="13" pageNumber="244" type="paratype">paratypes</typeStatus>
:
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.
</paragraph>
<paragraph id="A30E36833E59FFD7DC82F8BBEB0DFFD6" blockId="13.[151,1437,1554,2013]" lastBlockId="14.[151,1437,151,1185]" lastPageId="14" lastPageNumber="245" pageId="13" pageNumber="244">Male. Upperside. Ground colour of head, thorax, wings and abdomen varying from russet to pale brown; body and base of wings irrorated with pale greyish-silver scales. Labial palp, long, porrect, three times diameter of eye. Antenna bipectinate, brown. Forewing elongate triangular, rounded at apex, outer margin gently arcuate with weak interneural crenulation. Anal margin with dark scales medially. Antemedial fascia dark brown, unevenly arcuate. Discal spot white, crescent- or reniform-shaped, ringed in black. Medial fascia dark brown, gently and unevenly arcuate, with paler scaling on its outer edge. Submarginal fascia dark brown, indistinct in some specimens, irregularly sinuate, indented distad in space CuA1, strongly defined on its outer edge leaving a weaker shadow proximally. Terminal area irrorated with dark scales. Termen with pale greyish-silver scales. Fringe greyish-brown. Hindwing ovoid, costal margin with emargination, outer margin arcuate and crenulate; paler basally and anally. Costal margin with darker brown irroration. Medial fascia dark brown, faint, gently arcuate, with a paler diffuse band on its outer edge. Submarginal fascia dark brown, indistinct, irregularly sinuate. Termen with pale greyish-silver scales. Fringe greyish-brown.</paragraph>
<paragraph id="A30E36833E5AFFD7DC82FE1BEFDBFC62" blockId="14.[151,1437,151,1185]" pageId="14" pageNumber="245">Underside. Ground colour as on upperside but forewings paler; both wings irrorated with pale straw or greyish-silver scales, and generally paler distally and basally. Medial fascia as on upperside but petering out towards anal margin. Postmedial fascia dark brown, very faint, straight. Submarginal fascia as on upperside but weakly-expressed. Terminal fascia dark brown, very narrow. Hindwing medial fascia as on upperside but well-defined. Postmedial fascia dark brown, very faint, sinuate. Submarginal fascia as on upperside, but diffuse, broader and better defined.</paragraph>
<paragraph id="A30E36833E5AFFD7DC82FDCFEBE1FDFA" blockId="14.[151,1437,151,1185]" pageId="14" pageNumber="245">Male genitalia. Uncus fully reduced. Socius reduced, represented by small membranous bulge bearing very long setae. Tegumen moderately long (ca. twice length of valve) with very narrow lateral plates. Juxta quadrangular, heavily sclerotized, fused with phallus with short densely setose brush-like processes basolaterally. Valve very short (ca. half length of phallus), bilobate, densely setose, proximal lobe rounded in basal two-thirds produced into a short but stout, apically rounded process; distal lobe considerably narrower basally and ca. 10% longer than proximal lobe, apex produced into a short, pointed tip. Vinculum short and broad (as long as valve base), cubile elongatequadrangular with thick, apically rounded craniolateral processes of same length as basal part of cubile and gently sinuous caudal margin. Phallus moderately thick, posteriorly dilated; carina short (a third of length of phallus), robust triangular, situated dorsally; ventral side of phallus with scobinate plate ventrally. Vesica as long as carina, inflated, without cornuti.</paragraph>
<paragraph id="A30E36833E5AFFD7DC82FC27EBDEFD96" blockId="14.[151,1437,151,1185]" box="[199,401,943,969]" pageId="14" pageNumber="245">Female unknown.</paragraph>
<paragraph id="A30E36833E5AFFD7DC82FC5AEE96FA4E" blockId="14.[151,1437,151,1185]" pageId="14" pageNumber="245">
<emphasis id="91C5EA913E5AFFD7DC82FC5AEBD7FDB3" bold="true" box="[199,408,978,1004]" pageId="14" pageNumber="245">DNA divergences</emphasis>
. The new species is recovered in two BINs BOLD:AFG8787 and BOLD:AFG8788 with intraspecific PWDs of 0.02.0% (n=7). It differs from
<taxonomicName id="64B14D003E5AFFD7DEBEFC7FE93CFA4F" authorityName="Aurivillius" authorityYear="1879" box="[763,883,1015,1040]" class="Insecta" family="Lasiocampidae" genus="Rhinobombyx" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="245" phylum="Arthropoda" rank="species" species="cuneata">
<emphasis id="91C5EA913E5AFFD7DEBEFC7FE93CFA4F" box="[763,883,1015,1040]" italics="true" pageId="14" pageNumber="245">R. cuneata</emphasis>
</taxonomicName>
with PWDs of 5.66.8% (n=4).
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E5AFFD7DC82FB93EAB7FA06" pageId="14" pageNumber="245" type="etymology">
<paragraph id="A30E36833E5AFFD7DC82FB93EAB7FA06" blockId="14.[151,1437,151,1185]" pageId="14" pageNumber="245">
<emphasis id="91C5EA913E5AFFD7DC82FB93EBD6FA6A" bold="true" box="[199,409,1051,1077]" italics="true" pageId="14" pageNumber="245">Derivatio nominis</emphasis>
. The new species derives its name from the
<typeStatus id="7C0A88213E5AFFD7DFDAFB93E980FA6A" box="[927,975,1051,1077]" pageId="14" pageNumber="245">type</typeStatus>
locality. An adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="EBAB65083E5AFFD7DC82FBEAEABEFAFE" pageId="14" pageNumber="245" type="distribution">
<paragraph id="A30E36833E5AFFD7DC82FBEAEABEFAFE" blockId="14.[151,1437,151,1185]" pageId="14" pageNumber="245">
<emphasis id="91C5EA913E5AFFD7DC82FBEAEB16FA23" bold="true" box="[199,345,1122,1148]" pageId="14" pageNumber="245">Distribution</emphasis>
. The new species is known only from its
<typeStatus id="7C0A88213E5AFFD7DF62FBEBE918FA22" box="[807,855,1123,1149]" pageId="14" pageNumber="245">type</typeStatus>
locality and is almost certainly endemic to the MPA hotspot.
</paragraph>
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<mods:title id="ABE0411887449602EF9AF30051166B8F">The Lasiocampidae (Lepidoptera: Lasiocampoidea) of Maputo Special Reserve, Mozambique: a provisional checklist with descriptions of three new species and taxonomic notes</mods:title>
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<emphasis id="91C5EA913E5AFFD7DCD2FB7AEB31FB52" bold="true" box="[151,382,1266,1293]" italics="true" pageId="14" pageNumber="245">Soligena juna juna</emphasis>
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[4A]—
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; [4B]—
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; [4C]—
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; [4D]—
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; [4F]—
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; [4G]—
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; [4H]—
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; [5B]—
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<emphasis id="91C5EA913E5FFFD2DCD2F99BEAB9F871" bold="true" box="[151,246,1555,1582]" italics="true" pageId="11" pageNumber="242">Trabala</emphasis>
cf.
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*
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(
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[4A]—
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<emphasis id="91C5EA913E5FFFD2DCD2F92BEB4BF8E2" bold="true" box="[151,260,1699,1725]" pageId="11" pageNumber="242">Remarks</emphasis>
. Preliminary investigations have revealed a complicated picture within
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and the genus is in need of revision. All of the Afrotropical species currently standing under the genus were described from West and Central Africa, and it is possible that this single male specimen from MSR, which constitutes a new country record and range extension for the genus, is an undescribed species. However, before any attempt is made to describe this taxon, the specific boundaries of the known taxa need to be established which is beyond the scope of this present work.
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<emphasis id="91C5EA913E5FFFD2DCD2FAD7EB25FB26" bold="true" box="[151,362,1375,1402]" italics="true" pageId="11" pageNumber="242">Philotherma rosa</emphasis>
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<mods:title id="22F54E87DA5167B39AEB5987515D85F8">A new species of the genus Mongolotettix Rehn, 1928 from Zhejiang, China (Acrididae, Acridoidea, Orthoptera)</mods:title>
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<emphasis id="8590EA88A914981ADEE49DF1C019FC31" bold="true" box="[151,533,984,1011]" italics="true" pageId="0" pageNumber="297">Mongolotettix tianmushanensis</emphasis>
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<figureCitation id="2FDF2A1FA914981ADCF19DF1C0F3FC30" box="[642,767,984,1010]" captionStart="FIGURES 115" captionStartId="1.[151,264,1640,1664]" captionTargetBox="[233,1348,195,1562]" captionTargetId="figure-19@1.[201,1386,173,1611]" captionTargetPageId="1" captionText="FIGURES 115. Mongolotettix tianmushanensis sp. nov. 1. Body in lateral view ♂; 2. Mesosternum in ventral view♂; 3. Epiphallus, ♂; 4. End of abdomen in dorsal view♂; 5. Stridulatory pegs ♂; 6. Tegmen ♀; 7. Body in lateral view ♀; 8. Mesosternum in ventral view♀; 9. End of abdomen in ventral view; 10. End of abdomen in lateral view. 1115. Mongolotettix jiangxiensis Zhang, et al, 2023 11. Body in lateral view ♂; 12. Stridulatory pegs ♂; 13. Mesosternum in ventral view♂; 14. Epiphallus, ♂; 15. Head and pronotum ♀." pageId="0" pageNumber="297">Figs. 110</figureCitation>
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<materialsCitation id="078C3CC7A914981ADEE49DD7C2E0FBF8" collectingDate="2011-07-29" collectorName="Wang Pen-Qiang &amp; Zhang De-Zhi." latitude="30.166666" location="Tianmushan" longLatPrecision="1223" longitude="118.86667" pageId="0" pageNumber="297" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
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, collected by
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<paragraph id="B75B369AA914981ADEB49A6FC685FA04" blockId="0.[151,1437,984,1947]" pageId="0" pageNumber="297">Male: Body medium in size. Head shorter than pronotum. Face oblique in profile. Frontal ridge widened downward. Antennae ensiform. Eyes oval, vertical diameter 1.8 times horizontal diameter and 1.3 times length of subocular furrow. Pronotum median keel visible, cut by hind transverse sulcus distinctly, lateral keels almost parallel, prozona 1.4 times metazona in length. Tegmina shorter, almost reaching the base of epiproct, distal margin concave in the middle, width of cubital area wider than width of medial area. Interspace of mesosternum narrow in the middle, length 2.5 times minimum width. Lateral lobes of metasternum separated. Upper keel of hind femur smooth, length of hind femur 5.6 times its maximum width, end of lower knee lobes angular, inner side with 97 stridulatory pegs, apex acuted. Hind tibia with 12 spines on inner side and 13 spines on outer side, external apical spine absent. Second metatarsomere shorter than the first. Tympanum large and round. Epiproct scutiform. Furculae larger, curver. Cercus long conical, extending over the tip of epiproct. Subgenital plate conical, apex acute. Hind margin of tergum of periproct with circular furculae. Width of epiphallus larger than high, both inner sides of epiphallus with projection in the middle (fig.3).</paragraph>
<paragraph id="B75B369AA914981ADEB49BFBC168F9B8" blockId="0.[151,1437,984,1947]" pageId="0" pageNumber="297">Female: Body more robust. Vertical diameter of eyes 1.5 times horizontal diameter and distinctly shorter than length of subocular furrow. Prozona 1.6 times metazoan in length. Tegmina extending over the hind margin of first abdominal tergum, length 2.6 times its maximum width. Length of hind femur 5.4 times its maximum width. Cercus short-conical, not reaching epiproct. Subgenital plate oblong, hind margin prominently acute in the middle. Upper ovipositor valve slender, length 3.8 times its maximum width, outer margin smooth.</paragraph>
<paragraph id="B75B369AA914981ADEB498AFC2EFF924" blockId="0.[151,1437,984,1947]" pageId="0" pageNumber="297">Coloration: Body yellowish brown. Antennae yellow. Pronotum yellowish brown. Tegmina of male without white stripe on the fore margin at base. Tegmina of female with a black longitudinal stripe before the middle.Abdomen yellowish brown.</paragraph>
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Measurement (in mm): Length of body:
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21.7,
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32.1. Length of tegmina:
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11.2,
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5.1. Length of hind femur:
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10.9,
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17.3.
</paragraph>
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Diagnosis. The new species is similar to
<taxonomicName id="70E44D19A914981ADC0B9912C164F890" authority="Zhang" authorityName="Zhang" box="[632,872,1850,1874]" class="Insecta" family="Acrididae" genus="Mongolotettix" kingdom="Animalia" order="Orthoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="species" species="jiangxiensis">
<emphasis id="8590EA88A914981ADC0B9912C116F890" box="[632,794,1851,1874]" italics="true" pageId="0" pageNumber="297">M. jiangxiensis</emphasis>
Zhang
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,
<emphasis id="8590EA88A914981ADD019912C1ADF890" box="[882,929,1850,1874]" italics="true" pageId="0" pageNumber="297">et al</emphasis>
, 2023. The major differences are listed in
<tableCitation id="FA660321A914981ADB129913C2A9F8B5" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Comparison of Mongolotettix tianmushanensis sp. nov. with M. jiangxiensis Zhang, et al, 2023" pageId="0" pageNumber="297">Table 1</tableCitation>
.
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<emphasis id="8590EA88A914981ADEB499AAC34EF859" bold="true" box="[199,322,1923,1947]" pageId="0" pageNumber="297">Etymology</emphasis>
The specific epithet is named for the
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locality Tianmushan.
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<mods:url id="2F4187605120766D71421B0F7ABB39DB">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
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<mods:identifier id="E044D713E40D2E9DDC2D24B21898D5C5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9C5E09220C590357FF58F9FD76E7C044" LSID="urn:lsid:plazi:treatment:9C5E09220C590357FF58F9FD76E7C044" httpUri="http://treatment.plazi.org/id/9C5E09220C590357FF58F9FD76E7C044" lastPageId="5" lastPageNumber="294" pageId="1" pageNumber="290">
<subSubSection id="5CEDEBBF0C590353FF58F9FD7107C230" box="[151,781,1554,1581]" pageId="1" pageNumber="290" type="nomenclature">
<paragraph id="1448B8340C590353FF58F9FD7107C230" blockId="1.[151,781,1554,1617]" box="[151,781,1554,1581]" pageId="1" pageNumber="290">
<heading id="4F000F580C590353FF58F9FD7107C230" bold="true" box="[151,781,1554,1581]" fontSize="11" level="1" pageId="1" pageNumber="290" reason="1">
<emphasis id="268364260C590353FF58F9FD7107C230" bold="true" box="[151,781,1554,1581]" pageId="1" pageNumber="290">
<taxonomicName id="D3F7C3B70C590353FF58F9FD70A9C231" authority="Acosta, Mendes &amp; Zefa" authorityName="Acosta, Mendes &amp; Zefa" authorityYear="2024" box="[151,675,1554,1581]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C590353FF58F9FD737CC230" bold="true" box="[151,374,1554,1581]" italics="true" pageId="1" pageNumber="290">Oecanthus buxixu</emphasis>
Acosta, Mendes &amp; Zefa
</taxonomicName>
,
<taxonomicNameLabel id="3DB0D95D0C590353FD60F9FC7107C230" box="[687,781,1555,1581]" pageId="1" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CEDEBBF0C590353FF58F9D8701DC24C" box="[151,535,1590,1617]" pageId="1" pageNumber="290" type="description">
<paragraph id="1448B8340C590353FF58F9D8701DC24C" blockId="1.[151,781,1554,1617]" box="[151,535,1590,1617]" pageId="1" pageNumber="290">
<figureCitation id="8CCCA4B10C590353FF58F9D873B0C24C" box="[151,442,1590,1617]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="1" pageNumber="290">Figures 16, 810, 1215</figureCitation>
;
<tableCitation id="59758D8F0C590353FE0BF9D8701DC24C" box="[452,535,1591,1617]" captionStart="TABLE 1" captionStartId="3.[152,244,1626,1650]" captionText="TABLE 1. Oecanthus buxixu sp. nov. measurements (mm). BL—body length; ID—interocular distance; PL—pronoto length; PWA—pronoto width anterior region; PWP—pronoto width posterior region; FL—femur III length; TL—tibia III length; FT—file teeth number; CL—cerci length; OL—ovipositor length." pageId="1" pageNumber="290">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="5CEDEBBF0C590353FF58F9907751C2A0" pageId="1" pageNumber="290" type="materials_examined">
<paragraph id="1448B8340C590353FF58F9907751C2A0" blockId="1.[151,1436,1662,1869]" pageId="1" pageNumber="290">
<emphasis id="268364260C590353FF58F9907349C285" bold="true" box="[151,323,1662,1689]" pageId="1" pageNumber="290">Type material.</emphasis>
<materialsCitation id="A49FB2690C590353FE87F9907081C2A0" collectingDate="2024-02-02" collectionCode="INPA" collectorName="Mendes, D. M. M." country="Brazil" county="Base Casa do Bare" latitude="-2.4906666" location="Amazonas" longLatPrecision="1" longitude="-64.70894" municipality="Maraa" pageId="1" pageNumber="290" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="268364260C590353FE87F99073BFC284" bold="true" box="[328,437,1663,1689]" pageId="1" pageNumber="290">
<typeStatus id="CB4C06960C590353FE87F99073BFC284" box="[328,437,1663,1689]" pageId="1" pageNumber="290" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="02F173BD0C590353FE74F99073D9C284" box="[443,467,1663,1689]" count="1" pageId="1" pageNumber="290" type="male"></specimenCount>
:
<collectingCountry id="6CE0F8A40C590353FE12F990702EC284" box="[477,548,1663,1689]" name="Brazil" pageId="1" pageNumber="290">Brazil</collectingCountry>
,
<location id="1128EEEF0C590353FDE3F99070AFC284" LSID="urn:lsid:plazi:treatment:9C5E09220C590357FF58F9FD76E7C044:1128EEEF0C590353FDE3F99070AFC284" box="[556,677,1663,1689]" country="Brazil" county="Base Casa do Bare" latitude="-2.4906666" longLatPrecision="1" longitude="-64.70894" municipality="Maraa" name="Amazonas" pageId="1" pageNumber="290">Amazonas</location>
,
<collectingMunicipality id="F42C224E0C590353FD61F99070F2C284" box="[686,760,1663,1689]" pageId="1" pageNumber="290">Maraã</collectingMunicipality>
,
<collectingCounty id="FD29C0B80C590353FCCDF99071D9C284" box="[770,979,1663,1689]" pageId="1" pageNumber="290">Base Casa do Baré</collectingCounty>
,
<geoCoordinate id="71C3DEF30C590353FC13F9907669C284" box="[988,1123,1663,1689]" degrees="2" direction="south" minutes="29" orientation="latitude" pageId="1" pageNumber="290" precision="1" seconds="26.4" value="-2.4906666">2°2926.4”S</geoCoordinate>
<geoCoordinate id="71C3DEF30C590353FBBCF9907710C284" box="[1139,1306,1663,1689]" degrees="64" direction="west" minutes="42" orientation="longitude" pageId="1" pageNumber="290" precision="1" seconds="32.2" value="-64.70894">64°4232.2”W</geoCoordinate>
,
<date id="60499EF40C590353FAEBF9907792C284" box="[1316,1432,1663,1689]" pageId="1" pageNumber="290" value="2024-02-02">
<collectingDate id="700D671C0C590353FAEBF9907792C284" box="[1316,1432,1663,1689]" pageId="1" pageNumber="290" value="2024-02-02">02.ii.2024</collectingDate>
</date>
, coleta manual,
<collectorName id="B902DDE20C590353FE8EF94C7009C2A0" box="[321,515,1699,1725]" pageId="1" pageNumber="290">Mendes, D.M.M.</collectorName>
<emphasis id="268364260C590353FDC5F94C7026C2A0" box="[522,556,1699,1725]" italics="true" pageId="1" pageNumber="290">leg</emphasis>
. (
<collectionCode id="72E620F10C590353FD8DF94C708FC2A0" box="[578,645,1699,1725]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="290">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A49FB2690C590353FD55F94C7627C2A0" box="[666,1069,1699,1725]" collectionCode="INPA" country="Brazil" pageId="1" pageNumber="290" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="268364260C590353FD55F94C7111C2A0" bold="true" box="[666,795,1699,1725]" pageId="1" pageNumber="290">
<typeStatus id="CB4C06960C590353FD55F94C711CC2A0" box="[666,790,1699,1725]" pageId="1" pageNumber="290" type="paratype">Paratypes</typeStatus>
:
</emphasis>
01 (
<collectionCode id="72E620F10C590353FC83F94C719BC2A0" box="[844,913,1699,1725]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="290">INPA</collectionCode>
), 01
<specimenCount id="02F173BD0C590353FC0EF94C71DFC2A0" box="[961,981,1699,1725]" count="1" pageId="1" pageNumber="290" type="male"></specimenCount>
and 01
</materialsCitation>
<materialsCitation id="A49FB2690C590353FBE3F94C775DC2A0" box="[1068,1367,1699,1725]" collectingDate="2024-05-08" collectionCode="IDSM" country="Brazil" pageId="1" pageNumber="290" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="02F173BD0C590353FBE3F94C7636C2A0" box="[1068,1084,1699,1725]" count="1" pageId="1" pageNumber="290" type="female"></specimenCount>
(
<collectionCode id="72E620F10C590353FB83F94C769DC2A0" box="[1100,1175,1699,1725]" pageId="1" pageNumber="290">IDSM</collectionCode>
), Ibid.
<date id="60499EF40C590353FB2AF94C775DC2A0" box="[1253,1367,1699,1725]" pageId="1" pageNumber="290" value="2024-05-08">
<collectingDate id="700D671C0C590353FB2AF94C775DC2A0" box="[1253,1367,1699,1725]" pageId="1" pageNumber="290" value="2024-05-08">08.v.2024</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CEDEBBF0C590353FF08F92872F7C350" pageId="1" pageNumber="290" type="etymology">
<paragraph id="1448B8340C590353FF08F92872F7C350" blockId="1.[151,1436,1662,1869]" pageId="1" pageNumber="290">
<emphasis id="268364260C590353FF08F9287344C2FC" bold="true" box="[199,334,1735,1761]" pageId="1" pageNumber="290">Etymology.</emphasis>
The specific epithet
<taxonomicName id="D3F7C3B70C590353FDF3F9287092C2FC" authorityName="Acosta, Mendes &amp; Zefa" authorityYear="2024" box="[572,664,1735,1761]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="buxixu">“buxixu</taxonomicName>
” refers to the popular name of the plant
<taxonomicName id="D3F7C3B70C590353FBA9F9287743C2FD" box="[1126,1353,1735,1761]" class="Magnoliopsida" family="Melastomataceae" genus="Clidemia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="290" phylum="Magnoliopsida" rank="species" species="japurensis">
<emphasis id="268364260C590353FBA9F9287743C2FD" box="[1126,1353,1735,1761]" italics="true" pageId="1" pageNumber="290">Clidemia japurensis</emphasis>
</taxonomicName>
, where the tree cricket is commonly found. It is a shrub native to South America, especially found in
<collectingCountry id="6CE0F8A40C590353FB1DF9047713C318" box="[1234,1305,1771,1797]" name="Brazil" pageId="1" pageNumber="290">Brazil</collectingCountry>
,
<collectingCountry id="6CE0F8A40C590353FAE8F9047792C318" box="[1319,1432,1771,1797]" name="Colombia" pageId="1" pageNumber="290">Colombia</collectingCountry>
,
<collectingCountry id="6CE0F8A40C590353FF58F8E07307C334" box="[151,269,1807,1833]" name="Venezuela" pageId="1" pageNumber="290">Venezuela</collectingCountry>
, and
<collectingCountry id="6CE0F8A40C590353FE88F8E073A8C334" box="[327,418,1807,1833]" name="Guyana" pageId="1" pageNumber="290">Guyana</collectingCountry>
. This plant is known for its small edible fruits and attractive flowers, which the tree cricket feeds on.
</paragraph>
</subSubSection>
<caption id="4088E8BC0C5A0350FF58FACF7642C245" pageId="2" pageNumber="291" startId="2.[151,264,1312,1336]" targetBox="[151,1432,179,1285]" targetPageId="2" targetType="figure">
<paragraph id="1448B8340C5A0350FF58FACF7642C245" blockId="2.[151,1437,1312,1624]" pageId="2" pageNumber="291">
<emphasis id="268364260C5A0350FF58FACF734CC125" bold="true" box="[151,326,1312,1336]" pageId="2" pageNumber="291">FIGURES 115.</emphasis>
Male and Female morphological characteristics of
<taxonomicName id="D3F7C3B70C5A0350FC87FACF71F4C125" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[840,1022,1312,1336]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5A0350FC87FACF71F4C125" box="[840,1022,1312,1336]" italics="true" pageId="2" pageNumber="291">Oecanthus buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5A0350FBCCFACF7659C125" bold="true" box="[1027,1107,1312,1336]" pageId="2" pageNumber="291">
<taxonomicNameLabel id="3DB0D95D0C5A0350FBCCFACF7659C125" box="[1027,1107,1312,1336]" pageId="2" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus.
</paragraph>
</caption>
<subSubSection id="5CEDEBBF0C5A0350FF08F92D7706C31D" pageId="2" pageNumber="291" type="diagnosis">
<paragraph id="1448B8340C5A0350FF08F92D7706C31D" blockId="2.[151,1436,1730,1793]" pageId="2" pageNumber="291">
<emphasis id="268364260C5A0350FF08F92D7348C2C1" bold="true" box="[199,322,1730,1756]" pageId="2" pageNumber="291">Diagnosis.</emphasis>
Combination of the following characteristics: marks on scape and pedicel absent (
<figureCitation id="8CCCA4B10C5A0350FB3DF92D7733C2C0" box="[1266,1337,1730,1757]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="2" pageNumber="291">Fig. 4</figureCitation>
); calling song with 3 to 5 chirps/s, 6 to 20 pulses per chirp, and peak frequency 3.3 kHz, at 26ºC (
<figureCitation id="8CCCA4B10C5A0350FBB8F90976F5C31D" box="[1143,1279,1766,1792]" captionStart="FIGURES 1619" captionStartId="3.[151,264,905,929]" captionTargetBox="[161,1425,192,871]" captionTargetId="figure-18@3.[151,1436,181,882]" captionTargetPageId="3" captionText="FIGURES 1619. Spectrogram of the calling song of Oecanthus buxixu sp. nov. 16, Sonogram of the calling song registered in the field; 17, Sonogram of the calling song registered in the Lab.; 18, Oscillogram of the calling song registered in the Lab; 19, peak frequency." pageId="2" pageNumber="291">Figs. 1619</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="4088E8BC0C5B0351FF58FC667354C7F7" pageId="3" pageNumber="292" startId="3.[151,264,905,929]" targetBox="[161,1425,192,871]" targetPageId="3" targetType="figure">
<paragraph id="1448B8340C5B0351FF58FC667354C7F7" blockId="3.[151,1436,905,1002]" pageId="3" pageNumber="292">
<emphasis id="268364260C5B0351FF58FC66735FC7BF" bold="true" box="[151,341,905,930]" pageId="3" pageNumber="292">FIGURES 1619.</emphasis>
Spectrogram of the calling song of
<taxonomicName id="D3F7C3B70C5B0351FD09FC657177C7BF" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[710,893,906,930]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5B0351FD09FC657177C7BF" box="[710,893,906,930]" italics="true" pageId="3" pageNumber="292">Oecanthus buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5B0351FC4BFC6571DFC7BF" bold="true" box="[900,981,906,930]" pageId="3" pageNumber="292">
<taxonomicNameLabel id="3DB0D95D0C5B0351FC4BFC6571DFC7BF" box="[900,981,906,930]" pageId="3" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
16, Sonogram of the calling song registered in the field; 17, Sonogram of the calling song registered in the Lab.; 18, Oscillogram of the calling song registered in the Lab; 19, peak frequency.
</paragraph>
</caption>
<subSubSection id="5CEDEBBF0C5B0357FF08FBD376E7C044" lastPageId="5" lastPageNumber="294" pageId="3" pageNumber="292" type="description">
<paragraph id="1448B8340C5B0351FF08FBD3769FC237" blockId="3.[151,1437,1084,1579]" pageId="3" pageNumber="292">
<emphasis id="268364260C5B0351FF08FBD3735BC04B" bold="true" box="[199,337,1084,1110]" pageId="3" pageNumber="292">Description</emphasis>
,
<typeStatus id="CB4C06960C5B0351FEADFBD373CEC04B" box="[354,452,1084,1110]" pageId="3" pageNumber="292" type="holotype">holotype</typeStatus>
<specimenCount id="02F173BD0C5B0351FE03FBD273EBC04A" box="[460,481,1085,1111]" count="1" pageId="3" pageNumber="292" type="male"></specimenCount>
(
<figureCitation id="8CCCA4B10C5B0351FE3DFBD370F3C04A" box="[498,761,1084,1111]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="3" pageNumber="292">Figs 16, 810, 1215</figureCitation>
). Coloration described
<emphasis id="268364260C5B0351FBCCFBD2765BC04B" box="[1027,1105,1085,1110]" italics="true" pageId="3" pageNumber="292">in vivo</emphasis>
. Body and other appendages light green.
<emphasis id="268364260C5B0351FEEBFB8F736EC067" bold="true" box="[292,356,1120,1146]" pageId="3" pageNumber="292">Head</emphasis>
: prognathous, uniformly light green, including vertex, gena, clypeus, and labrum; dark eyes; ocelli absent; scape and pedicel without spots (
<figureCitation id="8CCCA4B10C5B0351FD77FB6B70F5C083" box="[696,767,1156,1182]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="3" pageNumber="292">Fig. 4</figureCitation>
), same color as the flagellum; maxillary palps translucent green, moderately elongated, first and second palpomeres smaller than the other three, third to fifth subequal in size; last palpomere claviform, slightly curved at apex; labial palps light green, gradually increasing in size towards the apex.
<emphasis id="268364260C5B0351FEC8FB1F736AC117" bold="true" box="[263,352,1264,1290]" pageId="3" pageNumber="292">Thorax</emphasis>
: pronotum longer than wide, posterior margin larger than anterior, dorsal disk pale green, with slightly whitish dorsolateral stripes; pronotum lateral lobe posterior more projected ventrally then the anterior region; tegmina translucent, longer than the abdomen; stridulatory file with 42 teeth; four cross veins in the harp, the smaller one poorly marked and curved; one cross veins in the mirror; well-developed hind wings, extending beyond the tegmina. Legs uniformly pale green; auditory tympana present in the tibiae I, the inner twice the size of the outer; tibia III slightly longer than femur III, serrulate, armed with four inner and three outer subapical spurs, and three outer and four inner apical spurs; spurs and spines of tibiae tipped with black; basitarsus longer than the other two tarsomeres combined.
<emphasis id="268364260C5B0351FE0DFA03703DC21B" bold="true" box="[450,567,1516,1542]" pageId="3" pageNumber="292">Abdomen</emphasis>
: tergites whitish translucent, cerci light green, tipped with black, and reaching the length of the hind wings; subgenital plate rounded posteriorly; measurements (
<tableCitation id="59758D8F0C5B0351FBFFF9FF768EC237" box="[1072,1156,1552,1578]" captionStart="TABLE 1" captionStartId="3.[152,244,1626,1650]" captionText="TABLE 1. Oecanthus buxixu sp. nov. measurements (mm). BL—body length; ID—interocular distance; PL—pronoto length; PWA—pronoto width anterior region; PWP—pronoto width posterior region; FL—femur III length; TL—tibia III length; FT—file teeth number; CL—cerci length; OL—ovipositor length." pageId="3" pageNumber="292">Table 1</tableCitation>
).
</paragraph>
<caption id="4088E8BC0C5B0351FF57F9B571A3C2A7" pageId="3" pageNumber="292" startId="3.[152,244,1626,1650]" targetType="table">
<paragraph id="1448B8340C5B0351FF57F9B571A3C2A7" blockId="3.[151,1437,1626,1722]" pageId="3" pageNumber="292">
<emphasis id="268364260C5B0351FF57F9B5731AC26F" bold="true" box="[152,272,1626,1650]" pageId="3" pageNumber="292">TABLE 1.</emphasis>
<taxonomicName id="D3F7C3B70C5B0351FED7F9B573D3C26F" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[280,473,1626,1650]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5B0351FED7F9B573D3C26F" box="[280,473,1626,1650]" italics="true" pageId="3" pageNumber="292">Oecanthus buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5B0351FE2EF9B57032C26F" bold="true" box="[481,568,1626,1650]" pageId="3" pageNumber="292">
<taxonomicNameLabel id="3DB0D95D0C5B0351FE2EF9B57032C26F" box="[481,568,1626,1650]" pageId="3" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
measurements (mm). BL—body length; ID—interocular distance; PL—pronoto length; PWA—pronoto width anterior region; PWP—pronoto width posterior region; FL—femur III length; TL—tibia III length; FT—file teeth number; CL—cerci length; OL—ovipositor length.
</paragraph>
</caption>
<paragraph id="1448B8340C5B0351FE9EF924777DC3BB" pageId="3" pageNumber="292">
<table id="66F74A940C5BFCADFF50F924777DC3BB" box="[159,1399,1739,1958]" gridcols="11" gridrows="6" pageId="3" pageNumber="292">
<tr id="AAC7BA760C5BFCADFF50F924777DC2FD" box="[159,1399,1739,1760]" gridrow="0" pageId="3" pageNumber="292" rowspan-0="1">
<th id="E916D30A0C5BFCADFE9EF9247371C2FD" box="[337,379,1739,1760]" gridcol="1" gridrow="0" pageId="3" pageNumber="292">BL</th>
<th id="E916D30A0C5BFCADFE09F92473FAC2FD" box="[454,496,1739,1760]" gridcol="2" gridrow="0" pageId="3" pageNumber="292">ID</th>
<th id="E916D30A0C5BFCADFDF3F924706CC2FD" box="[572,614,1739,1760]" gridcol="3" gridrow="0" pageId="3" pageNumber="292">PL</th>
<th id="E916D30A0C5BFCADFD7FF92470E9C2FD" box="[688,739,1739,1760]" gridcol="4" gridrow="0" pageId="3" pageNumber="292">PWA</th>
<th id="E916D30A0C5BFCADFCD0F924715AC2FD" box="[799,848,1739,1760]" gridcol="5" gridrow="0" pageId="3" pageNumber="292">PWP</th>
<th id="E916D30A0C5BFCADFC44F92471BFC2FD" box="[907,949,1739,1760]" gridcol="6" gridrow="0" pageId="3" pageNumber="292">FL</th>
<th id="E916D30A0C5BFCADFC31F9247622C2FD" box="[1022,1064,1739,1760]" gridcol="7" gridrow="0" pageId="3" pageNumber="292">TL</th>
<th id="E916D30A0C5BFCADFBA2F9247683C2FD" box="[1133,1161,1739,1760]" gridcol="8" gridrow="0" pageId="3" pageNumber="292">FT</th>
<th id="E916D30A0C5BFCADFB19F924770AC2FD" box="[1238,1280,1739,1760]" gridcol="9" gridrow="0" pageId="3" pageNumber="292">CL</th>
<th id="E916D30A0C5BFCADFA82F924777DC2FD" box="[1357,1399,1739,1760]" gridcol="10" gridrow="0" pageId="3" pageNumber="292">OL</th>
</tr>
<tr id="AAC7BA760C5BFCADFF50F91D777DC315" box="[159,1399,1778,1800]" gridrow="1" pageId="3" pageNumber="292">
<th id="E916D30A0C5BFCADFF50F91D7300C315" box="[159,266,1778,1800]" gridcol="0" gridrow="1" pageId="3" pageNumber="292">Holotype♂</th>
<td id="E916D30A0C5BFCADFE9EF91D7371C315" box="[337,379,1778,1800]" gridcol="1" gridrow="1" pageId="3" pageNumber="292">9.45</td>
<td id="E916D30A0C5BFCADFE09F91D73FAC315" box="[454,496,1778,1800]" gridcol="2" gridrow="1" pageId="3" pageNumber="292">0.66</td>
<td id="E916D30A0C5BFCADFDF3F91D706CC315" box="[572,614,1778,1800]" gridcol="3" gridrow="1" pageId="3" pageNumber="292">1.62</td>
<td id="E916D30A0C5BFCADFD7FF91D70E9C315" box="[688,739,1778,1800]" gridcol="4" gridrow="1" pageId="3" pageNumber="292">0.82</td>
<td id="E916D30A0C5BFCADFCD0F91D715AC315" box="[799,848,1778,1800]" gridcol="5" gridrow="1" pageId="3" pageNumber="292">1.33</td>
<td id="E916D30A0C5BFCADFC44F91D71BFC315" box="[907,949,1778,1800]" gridcol="6" gridrow="1" pageId="3" pageNumber="292">6.72</td>
<td id="E916D30A0C5BFCADFC31F91D7622C315" box="[1022,1064,1778,1800]" gridcol="7" gridrow="1" pageId="3" pageNumber="292">7.71</td>
<td id="E916D30A0C5BFCADFBA2F91D7683C315" box="[1133,1161,1778,1800]" gridcol="8" gridrow="1" pageId="3" pageNumber="292">42</td>
<td id="E916D30A0C5BFCADFB19F91D770AC315" box="[1238,1280,1778,1800]" gridcol="9" gridrow="1" pageId="3" pageNumber="292">3.65</td>
<td id="E916D30A0C5BFCADFA82F91D777DC315" box="[1357,1399,1778,1800]" gridcol="10" gridrow="1" pageId="3" pageNumber="292">-</td>
</tr>
<tr id="AAC7BA760C5BFCADFF50F8F6777DC332" box="[159,1399,1817,1839]" gridrow="2" pageId="3" pageNumber="292">
<th id="E916D30A0C5BFCADFF50F8F67300C332" box="[159,266,1817,1839]" gridcol="0" gridrow="2" pageId="3" pageNumber="292">Paratype♂</th>
<td id="E916D30A0C5BFCADFE9EF8F67371C332" box="[337,379,1817,1839]" gridcol="1" gridrow="2" pageId="3" pageNumber="292">9.68</td>
<td id="E916D30A0C5BFCADFE09F8F673FAC332" box="[454,496,1817,1839]" gridcol="2" gridrow="2" pageId="3" pageNumber="292">0.69</td>
<td id="E916D30A0C5BFCADFDF3F8F6706CC332" box="[572,614,1817,1839]" gridcol="3" gridrow="2" pageId="3" pageNumber="292">1.64</td>
<td id="E916D30A0C5BFCADFD7FF8F670E9C332" box="[688,739,1817,1839]" gridcol="4" gridrow="2" pageId="3" pageNumber="292">0.89</td>
<td id="E916D30A0C5BFCADFCD0F8F6715AC332" box="[799,848,1817,1839]" gridcol="5" gridrow="2" pageId="3" pageNumber="292">1.39</td>
<td id="E916D30A0C5BFCADFC44F8F671BFC332" box="[907,949,1817,1839]" gridcol="6" gridrow="2" pageId="3" pageNumber="292">6.83</td>
<td id="E916D30A0C5BFCADFC31F8F67622C332" box="[1022,1064,1817,1839]" gridcol="7" gridrow="2" pageId="3" pageNumber="292">7.9</td>
<td id="E916D30A0C5BFCADFBA2F8F67683C332" box="[1133,1161,1817,1839]" gridcol="8" gridrow="2" pageId="3" pageNumber="292">37</td>
<td id="E916D30A0C5BFCADFB19F8F6770AC332" box="[1238,1280,1817,1839]" gridcol="9" gridrow="2" pageId="3" pageNumber="292">3.72</td>
<td id="E916D30A0C5BFCADFA82F8F6777DC332" box="[1357,1399,1817,1839]" gridcol="10" gridrow="2" pageId="3" pageNumber="292">-</td>
</tr>
<tr id="AAC7BA760C5BFCADFF50F8AE777DC34A" box="[159,1399,1857,1879]" gridrow="3" pageId="3" pageNumber="292">
<th id="E916D30A0C5BFCADFF50F8AE7300C34A" box="[159,266,1857,1879]" gridcol="0" gridrow="3" pageId="3" pageNumber="292">Paratype♂</th>
<td id="E916D30A0C5BFCADFE9EF8AE7371C34A" box="[337,379,1857,1879]" gridcol="1" gridrow="3" pageId="3" pageNumber="292">10.2</td>
<td id="E916D30A0C5BFCADFE09F8AE73FAC34A" box="[454,496,1857,1879]" gridcol="2" gridrow="3" pageId="3" pageNumber="292">0.7</td>
<td id="E916D30A0C5BFCADFDF3F8AE706CC34A" box="[572,614,1857,1879]" gridcol="3" gridrow="3" pageId="3" pageNumber="292">1.7</td>
<td id="E916D30A0C5BFCADFD7FF8AE70E9C34A" box="[688,739,1857,1879]" gridcol="4" gridrow="3" pageId="3" pageNumber="292">1</td>
<td id="E916D30A0C5BFCADFCD0F8AE715AC34A" box="[799,848,1857,1879]" gridcol="5" gridrow="3" pageId="3" pageNumber="292">1.4</td>
<td id="E916D30A0C5BFCADFC44F8AE71BFC34A" box="[907,949,1857,1879]" gridcol="6" gridrow="3" pageId="3" pageNumber="292">7.1</td>
<td id="E916D30A0C5BFCADFC31F8AE7622C34A" box="[1022,1064,1857,1879]" gridcol="7" gridrow="3" pageId="3" pageNumber="292">8.1</td>
<td id="E916D30A0C5BFCADFBA2F8AE7683C34A" box="[1133,1161,1857,1879]" gridcol="8" gridrow="3" pageId="3" pageNumber="292">-</td>
<td id="E916D30A0C5BFCADFB19F8AE770AC34A" box="[1238,1280,1857,1879]" gridcol="9" gridrow="3" pageId="3" pageNumber="292">3.82</td>
<td id="E916D30A0C5BFCADFA82F8AE777DC34A" box="[1357,1399,1857,1879]" gridcol="10" gridrow="3" pageId="3" pageNumber="292">-</td>
</tr>
<tr id="AAC7BA760C5BFCADFF50F886777DC362" box="[159,1399,1897,1919]" gridrow="4" pageId="3" pageNumber="292">
<th id="E916D30A0C5BFCADFF50F8867300C362" box="[159,266,1897,1919]" gridcol="0" gridrow="4" pageId="3" pageNumber="292">Paratype♀</th>
<td id="E916D30A0C5BFCADFE9EF8867371C362" box="[337,379,1897,1919]" gridcol="1" gridrow="4" pageId="3" pageNumber="292">11.8</td>
<td id="E916D30A0C5BFCADFE09F88673FAC362" box="[454,496,1897,1919]" gridcol="2" gridrow="4" pageId="3" pageNumber="292">0.72</td>
<td id="E916D30A0C5BFCADFDF3F886706CC362" box="[572,614,1897,1919]" gridcol="3" gridrow="4" pageId="3" pageNumber="292">1.6</td>
<td id="E916D30A0C5BFCADFD7FF88670E9C362" box="[688,739,1897,1919]" gridcol="4" gridrow="4" pageId="3" pageNumber="292">1.2</td>
<td id="E916D30A0C5BFCADFCD0F886715AC362" box="[799,848,1897,1919]" gridcol="5" gridrow="4" pageId="3" pageNumber="292">1.3</td>
<td id="E916D30A0C5BFCADFC44F88671BFC362" box="[907,949,1897,1919]" gridcol="6" gridrow="4" pageId="3" pageNumber="292">7.5</td>
<td id="E916D30A0C5BFCADFC31F8867622C362" box="[1022,1064,1897,1919]" gridcol="7" gridrow="4" pageId="3" pageNumber="292">9.1</td>
<td id="E916D30A0C5BFCADFBA2F8867683C362" box="[1133,1161,1897,1919]" gridcol="8" gridrow="4" pageId="3" pageNumber="292">-</td>
<td id="E916D30A0C5BFCADFB19F886770AC362" box="[1238,1280,1897,1919]" gridcol="9" gridrow="4" pageId="3" pageNumber="292">4.36</td>
<td id="E916D30A0C5BFCADFA82F886777DC362" box="[1357,1399,1897,1919]" gridcol="10" gridrow="4" pageId="3" pageNumber="292">4.41</td>
</tr>
<tr id="AAC7BA760C5BFCADFF50F87F777DC3BB" box="[159,1399,1936,1958]" gridrow="5" pageId="3" pageNumber="292">
<th id="E916D30A0C5BFCADFF50F87F7300C3BB" box="[159,266,1936,1958]" gridcol="0" gridrow="5" pageId="3" pageNumber="292">Paratype♀</th>
<td id="E916D30A0C5BFCADFE9EF87F7371C3BB" box="[337,379,1936,1958]" gridcol="1" gridrow="5" pageId="3" pageNumber="292">12.4</td>
<td id="E916D30A0C5BFCADFE09F87F73FAC3BB" box="[454,496,1936,1958]" gridcol="2" gridrow="5" pageId="3" pageNumber="292">0.75</td>
<td id="E916D30A0C5BFCADFDF3F87F706CC3BB" box="[572,614,1936,1958]" gridcol="3" gridrow="5" pageId="3" pageNumber="292">1.65</td>
<td id="E916D30A0C5BFCADFD7FF87F70E9C3BB" box="[688,739,1936,1958]" gridcol="4" gridrow="5" pageId="3" pageNumber="292">1.19</td>
<td id="E916D30A0C5BFCADFCD0F87F715AC3BB" box="[799,848,1936,1958]" gridcol="5" gridrow="5" pageId="3" pageNumber="292">1.6</td>
<td id="E916D30A0C5BFCADFC44F87F71BFC3BB" box="[907,949,1936,1958]" gridcol="6" gridrow="5" pageId="3" pageNumber="292">7</td>
<td id="E916D30A0C5BFCADFC31F87F7622C3BB" box="[1022,1064,1936,1958]" gridcol="7" gridrow="5" pageId="3" pageNumber="292">8.54</td>
<td id="E916D30A0C5BFCADFBA2F87F7683C3BB" box="[1133,1161,1936,1958]" gridcol="8" gridrow="5" pageId="3" pageNumber="292">-</td>
<td id="E916D30A0C5BFCADFB19F87F770AC3BB" box="[1238,1280,1936,1958]" gridcol="9" gridrow="5" pageId="3" pageNumber="292">4.33</td>
<td id="E916D30A0C5BFCADFA82F87F777DC3BB" box="[1357,1399,1936,1958]" gridcol="10" gridrow="5" pageId="3" pageNumber="292">4.76</td>
</tr>
</table>
</paragraph>
<caption id="4088E8BC0C5C0356FF58F878763BC3CE" pageId="4" pageNumber="293" startId="4.[151,264,1943,1967]" targetBox="[178,1409,181,1918]" targetPageId="4" targetType="figure">
<paragraph id="1448B8340C5C0356FF58F878763BC3CE" blockId="4.[151,1436,1943,2003]" pageId="4" pageNumber="293">
<emphasis id="268364260C5C0356FF58F878735FC3B2" bold="true" box="[151,341,1943,1967]" pageId="4" pageNumber="293">FIGURES 2024.</emphasis>
Photographs of
<taxonomicName id="D3F7C3B70C5C0356FE32F87870BEC3B2" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[509,692,1943,1967]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5C0356FE32F87870BEC3B2" box="[509,692,1943,1967]" italics="true" pageId="4" pageNumber="293">Oecanthus buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5C0356FD74F8787106C3B2" bold="true" box="[699,780,1943,1967]" pageId="4" pageNumber="293">
<taxonomicNameLabel id="3DB0D95D0C5C0356FD74F8787106C3B2" box="[699,780,1943,1967]" pageId="4" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the field. 20, Adult male; 21, Adult female; 22, Nymph f; 23, Adult female feeding; 24, shrub
<taxonomicName id="D3F7C3B70C5C0356FE2FF85470A7C3CE" box="[480,685,1979,2003]" class="Magnoliopsida" family="Melastomataceae" genus="Clidemia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="293" phylum="Magnoliopsida" rank="species" species="japurensis">
<emphasis id="268364260C5C0356FE2FF85470A7C3CE" box="[480,685,1979,2003]" italics="true" pageId="4" pageNumber="293">Clidemia japurensis</emphasis>
</taxonomicName>
(buxixu). Photos by Mendes, D.M.M.
</paragraph>
</caption>
<paragraph id="1448B8340C5D0357FF08FF78709EC4E4" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="294">
<emphasis id="268364260C5D0357FF08FF787025C4A9" bold="true" box="[199,559,150,180]" pageId="5" pageNumber="294">
Metanotal gland (
<typeStatus id="CB4C06960C5D0357FE54FF797000C4AD" box="[411,522,150,176]" pageId="5" pageNumber="294" type="paratype">Paratype</typeStatus>
<specimenCount id="02F173BD0C5D0357FDDEFF77702CC4A9" box="[529,550,152,180]" count="1" pageId="5" pageNumber="294" type="male"></specimenCount>
)
</emphasis>
(
<figureCitation id="8CCCA4B10C5D0357FDF0FF787098C4AC" box="[575,658,151,177]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="5" pageNumber="294">Fig. 12</figureCitation>
). Scutum with a pair of protuberances, each one with two translucent tufts of bristles; scutellum with a pair of protuberances curved inward; posterior median lobe triangular, sclerotized, with apex translucent curved up and forward.
</paragraph>
<paragraph id="1448B8340C5D0357FF08FEEC71BEC5CC" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="294">
<emphasis id="268364260C5D0357FF08FEEC73FEC501" bold="true" box="[199,500,258,285]" pageId="5" pageNumber="294">
Male genitalia (
<typeStatus id="CB4C06960C5D0357FEB3FEED73E6C501" box="[380,492,258,284]" pageId="5" pageNumber="294" type="holotype">Holotype</typeStatus>
)
</emphasis>
(
<figureCitation id="8CCCA4B10C5D0357FDCEFEEC7088C500" box="[513,642,258,285]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="5" pageNumber="294">Figs 1415</figureCitation>
). Pseudepiphallic sclerite: median lophi (MLophi) projected posteriorly, forming an V-shaped invagination; MLophi apex slightly curved outward; pseudepiphallic apodema (PsAp) anterior margin curved posteriorly in dorsal view; rami (r) long, slightly curved inward, apex single, straight and not connected each other; ectophallic invagination well developed, ectophallic arc (arc) curved anteriorly, distal prolongation present; ectophallic fold (EctF) poorly sclerotized; ectophallic invagination ventral projection (EctVp) longer than ectophallic apodema (EctAp); endophalus (End) U-shaped.
</paragraph>
<paragraph id="1448B8340C5D0357FF08FE3571F5C6D0" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="294">
<emphasis id="268364260C5D0357FF08FE357351C5E8" bold="true" box="[199,347,474,501]" pageId="5" pageNumber="294">Calling song</emphasis>
(
<emphasis id="268364260C5D0357FEA3FE3473D3C5E8" bold="true" box="[364,473,475,501]" pageId="5" pageNumber="294">
<typeStatus id="CB4C06960C5D0357FEA3FE3473D3C5E8" box="[364,473,475,501]" pageId="5" pageNumber="294" type="holotype">Holotype</typeStatus>
</emphasis>
,
<figureCitation id="8CCCA4B10C5D0357FE28FE34706DC5E8" box="[487,615,475,501]" captionStart="FIGURES 1619" captionStartId="3.[151,264,905,929]" captionTargetBox="[161,1425,192,871]" captionTargetId="figure-18@3.[151,1436,181,882]" captionTargetPageId="3" captionText="FIGURES 1619. Spectrogram of the calling song of Oecanthus buxixu sp. nov. 16, Sonogram of the calling song registered in the field; 17, Sonogram of the calling song registered in the Lab.; 18, Oscillogram of the calling song registered in the Lab; 19, peak frequency." pageId="5" pageNumber="294">Figs 1619</figureCitation>
).
<emphasis id="268364260C5D0357FDB4FE347173C5E9" bold="true" box="[635,889,474,501]" pageId="5" pageNumber="294">Under field condition</emphasis>
: 3 ± 0.6 (24) chirps/s; chirp duration 0.28 ± 0.1 (0.050.81) s; interval between chirps 0.06 ± 0.03 (0.020.39) s; 9.1 ± 3.5 (226) pulses per chirp; peak frequency 2.9 ± 0.02 (2.922.96) kHz, at 24ºC. The first chirp has a higher number of pulses and exhibits a lower initial frequency and intensity (dB) (
<figureCitation id="8CCCA4B10C5D0357FE22FDA87048C67C" box="[493,578,583,609]" captionStart="FIGURES 1619" captionStartId="3.[151,264,905,929]" captionTargetBox="[161,1425,192,871]" captionTargetId="figure-18@3.[151,1436,181,882]" captionTargetPageId="3" captionText="FIGURES 1619. Spectrogram of the calling song of Oecanthus buxixu sp. nov. 16, Sonogram of the calling song registered in the field; 17, Sonogram of the calling song registered in the Lab.; 18, Oscillogram of the calling song registered in the Lab; 19, peak frequency." pageId="5" pageNumber="294">Fig. 16</figureCitation>
).
<emphasis id="268364260C5D0357FD97FDA8715FC67D" bold="true" box="[600,853,582,609]" pageId="5" pageNumber="294">Under lab conditions</emphasis>
: 3.5 ± 0.7 (35) chirps/s; chirp duration 0.3 ± 0.12 (0.150.53) s; interval between chirps 0.07 ± 0.02 (0.040.11) s; 11.2 ± 4.5 (620) pulses per chirp; peak frequency 3.1 ± 0.06 (33.3) kHz, at 26ºC. Similar to field conditions, the first chirp has a higher number of pulses and exhibits a lower initial frequency and intensity (dB) compared to the rest of the signal.
</paragraph>
<paragraph id="1448B8340C5D0357FF08FD3876F0C724" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="294">
<emphasis id="268364260C5D0357FF08FD387314C6EC" bold="true" box="[199,286,727,753]" pageId="5" pageNumber="294">Female</emphasis>
(
<figureCitation id="8CCCA4B10C5D0357FEE0FD3873ACC6EC" box="[303,422,727,753]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="5" pageNumber="294">Figs 7, 11</figureCitation>
,
<figureCitation id="8CCCA4B10C5D0357FE7BFD38700AC6EC" box="[436,512,727,753]" captionStart="FIGURES 2024" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1409,181,1918]" captionTargetId="figure-29@4.[178,1409,181,1918]" captionTargetPageId="4" captionText="FIGURES 2024. Photographs of Oecanthus buxixu sp. nov. in the field. 20, Adult male; 21, Adult female; 22, Nymph f; 23, Adult female feeding; 24, shrub Clidemia japurensis (buxixu). Photos by Mendes, D.M.M." pageId="5" pageNumber="294">21, 23</figureCitation>
). Body color similar to male, slightly longer than the male; tegmina slender and translucent; hind wings surpass the tegmina and cover the ovipositor and cerci; ovipositor as long as the cerci, apex denticulated; cerci pale green, tipped with black; supra-anal plate as
<figureCitation id="8CCCA4B10C5D0357FC58FCF071D0C724" box="[919,986,799,825]" captionStart="FIGURES 115" captionStartId="2.[151,264,1312,1336]" captionTargetBox="[151,1432,179,1285]" captionTargetId="figure-29@2.[151,1436,179,1286]" captionTargetPageId="2" captionText="FIGURES 115. Male and Female morphological characteristics of Oecanthus buxixu sp. nov. after alcohol fixation. 13, Male in dorsal, ventral and lateral view; 4, Part of head in dorsal view, with scape and pedicel in ventral view; 5, Male terminalia in ventral view, showing subgenital plate and cerci; 6, Male supranal plate, dorsal view; 7, Female terminalia in dorsal view, showing supranal plate; 8, Tibia III serrulate, with four inner and three outer subapical spurs (*); 9, Tibia III apex, with three outer apical spurs (a, b, c); 10, Tibia III apex, with four inner apical spurs (d, e, f, g); 11, Ovipositor apex, ventral view; 12, Male metanotal gland, dorsal view; 13, Male right tegmina, ventral view; 14, Phallic sclerites, ventral view; 15, Phallic sclerites, dorsal view. Conventions: sp—scape, pd—pedicel, Sc—scutum, St—scutellum, pml—posterior median lobe, MLophi—median lophi of pseudepiphallus; PsAp—pseudepiphallic apodema; r—rami; Arc—ectophallic arc; EctF—ectophallic fold; EctVp— ectophallic invagination ventral projection; EctAp—ectophallic apodema; End—endophalus." pageId="5" pageNumber="294">Fig. 7</figureCitation>
; measurements (
<tableCitation id="59758D8F0C5D0357FB5AFCF076E3C724" box="[1173,1257,799,825]" captionStart="TABLE 1" captionStartId="3.[152,244,1626,1650]" captionText="TABLE 1. Oecanthus buxixu sp. nov. measurements (mm). BL—body length; ID—interocular distance; PL—pronoto length; PWA—pronoto width anterior region; PWP—pronoto width posterior region; FL—femur III length; TL—tibia III length; FT—file teeth number; CL—cerci length; OL—ovipositor length." pageId="5" pageNumber="294">Table 1</tableCitation>
).
</paragraph>
<paragraph id="1448B8340C5D0357FF08FCAD76E7C044" blockId="5.[151,1437,150,1113]" pageId="5" pageNumber="294">
<emphasis id="268364260C5D0357FF08FCAD73D9C741" bold="true" box="[199,467,834,861]" pageId="5" pageNumber="294">Habitat and behavior.</emphasis>
<taxonomicName id="D3F7C3B70C5D0357FE12FCAC7047C740" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[477,589,835,861]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5D0357FE12FCAC7047C740" box="[477,589,835,861]" italics="true" pageId="5" pageNumber="294">O. buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5D0357FD98FCAC70BEC740" bold="true" box="[599,692,835,861]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="3DB0D95D0C5D0357FD98FCAC70BEC740" box="[599,692,835,861]" pageId="5" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found on the bush
<taxonomicName id="D3F7C3B70C5D0357FC1EFCAC76BCC741" box="[977,1206,835,861]" class="Magnoliopsida" family="Melastomataceae" genus="Clidemia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="294" phylum="Magnoliopsida" rank="species" species="japurensis">
<emphasis id="268364260C5D0357FC1EFCAC76BCC741" box="[977,1206,835,861]" italics="true" pageId="5" pageNumber="294">Clidemia japurensis</emphasis>
</taxonomicName>
(
<taxonomicName id="D3F7C3B70C5D0357FB07FCAC779BC740" box="[1224,1425,835,861]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
), present in “Terra Firme” forests near the edge (
<figureCitation id="8CCCA4B10C5D0357FD7BFC887100C79C" box="[692,778,871,897]" captionStart="FIGURES 2024" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1409,181,1918]" captionTargetId="figure-29@4.[178,1409,181,1918]" captionTargetPageId="4" captionText="FIGURES 2024. Photographs of Oecanthus buxixu sp. nov. in the field. 20, Adult male; 21, Adult female; 22, Nymph f; 23, Adult female feeding; 24, shrub Clidemia japurensis (buxixu). Photos by Mendes, D.M.M." pageId="5" pageNumber="294">Fig. 24</figureCitation>
). In the locality, the bushes were around
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tall, and the tree crickets were found between
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and
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in height, in the abaxial part of the leaves. We found only one individual per leaf. Nymphs were seen both on the leaves, walking among the branches. Males stridulate on the lower part of the leaves, generally close to the edge, and they were very skittish when approaching them during the song, stopping the song and closing their wings quickly. Adults and nymphs were observed feeding on fruits (
<figureCitation id="8CCCA4B10C5D0357FAA0FC1872BEC028" captionStart="FIGURES 2024" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[178,1409,181,1918]" captionTargetId="figure-29@4.[178,1409,181,1918]" captionTargetPageId="4" captionText="FIGURES 2024. Photographs of Oecanthus buxixu sp. nov. in the field. 20, Adult male; 21, Adult female; 22, Nymph f; 23, Adult female feeding; 24, shrub Clidemia japurensis (buxixu). Photos by Mendes, D.M.M." pageId="5" pageNumber="294">Fig. 23</figureCitation>
) and leaves, indicating a close relationship between the tree-crickets and this plant. Despite extensive search in the locality, specimens of
<taxonomicName id="D3F7C3B70C5D0357FE74FBD07022C044" authorityName="Acosta &amp; Mendes &amp; Ramalho &amp; Zefa" authorityYear="2024" box="[443,552,1087,1113]" class="Insecta" family="Gryllidae" genus="Oecanthus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="buxixu" status="sp. nov.">
<emphasis id="268364260C5D0357FE74FBD07022C044" box="[443,552,1087,1113]" italics="true" pageId="5" pageNumber="294">O. buxixu</emphasis>
</taxonomicName>
<emphasis id="268364260C5D0357FDE1FBD07082C044" bold="true" box="[558,648,1087,1113]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="3DB0D95D0C5D0357FDE1FBD07082C044" box="[558,648,1087,1113]" pageId="5" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were only found on on bushes of
<taxonomicName id="D3F7C3B70C5D0357FBCAFBD076EDC045" box="[1029,1255,1087,1113]" class="Magnoliopsida" family="Melastomataceae" genus="Clidemia" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="294" phylum="Magnoliopsida" rank="species" species="japurensis">
<emphasis id="268364260C5D0357FBCAFBD076EDC045" box="[1029,1255,1087,1113]" italics="true" pageId="5" pageNumber="294">Clidemia japurensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,180 @@
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<mods:title id="FD0841FA9FEB16A260EC3B12F54F2DA2">Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae)</mods:title>
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<mods:namePart id="9F49B98725340C4376140D155F0E34B6">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="C3C950CF8711DF24C31EDA2F2BB71A9E">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024 - 5192, USA &amp; Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Lima 15081, Perú &amp; Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima 15072, Perú</mods:affiliation>
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<mods:classification id="6ACBF01AF77778139EA6651BB0A6793E">journal article</mods:classification>
<mods:identifier id="4B492F6EDBCBC7975C3D2CDA3A4C7296" type="DOI">10.11646/zootaxa.5492.2.4</mods:identifier>
<mods:identifier id="D2518CBD38DF148ACB5AC7655F2AA684" type="ISSN">1175-5326</mods:identifier>
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<treatment id="D37C87A62734FFFB2DEBC69FFB400C9F" ID-DOI="http://doi.org/10.5281/zenodo.13219252" ID-Zenodo-Dep="13219252" LSID="urn:lsid:plazi:treatment:D37C87A62734FFFB2DEBC69FFB400C9F" httpUri="http://treatment.plazi.org/id/D37C87A62734FFFB2DEBC69FFB400C9F" lastPageId="10" lastPageNumber="224" pageId="9" pageNumber="223">
<subSubSection id="13CF653B2734FFF82DEBC69FFDC50F4C" box="[151,612,1255,1282]" pageId="9" pageNumber="223" type="nomenclature">
<paragraph id="5B6A36B02734FFF82DEBC69FFDC50F4C" blockId="9.[151,612,1255,1318]" box="[151,612,1255,1282]" pageId="9" pageNumber="223">
<heading id="002281DC2734FFF82DEBC69FFDC50F4C" bold="true" box="[151,612,1255,1282]" fontSize="11" level="1" pageId="9" pageNumber="223" reason="1">
<emphasis id="69A1EAA22734FFF82DEBC69FFDC50F4C" bold="true" box="[151,612,1255,1282]" pageId="9" pageNumber="223">
<taxonomicName id="9CD54D332734FFF82DEBC69FFE5E0F4F" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[151,511,1255,1282]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="zhenhuai" status="sp. nov.">
<emphasis id="69A1EAA22734FFF82DEBC69FFE280F4C" bold="true" box="[151,393,1255,1282]" italics="true" pageId="9" pageNumber="223">Eucinetus zhenhuai</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="729257D92734FFF82F7BC69FFDC50F4C" box="[519,612,1256,1282]" pageId="9" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2734FFF82DEBC77BFEB80F68" box="[151,281,1292,1318]" pageId="9" pageNumber="223" type="description">
<paragraph id="5B6A36B02734FFF82DEBC77BFEB80F68" blockId="9.[151,612,1255,1318]" box="[151,281,1292,1318]" pageId="9" pageNumber="223">
(
<figureCitation id="C3EE2A352734FFF82DE3C77BFEB00F68" box="[159,273,1292,1318]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStartId-0="10.[151,250,1252,1276]" captionStartId-1="11.[151,250,1600,1624]" captionStartId-2="12.[151,250,1503,1527]" captionStartId-3="13.[151,250,928,952]" captionTargetBox-0="[151,1435,181,1229]" captionTargetBox-1="[151,1435,181,1576]" captionTargetBox-2="[151,1435,181,1479]" captionTargetBox-3="[151,1435,181,904]" captionTargetId-0="figure-18@10.[151,1435,181,1229]" captionTargetId-1="figure-26@11.[151,1435,181,1576]" captionTargetId-2="figure-18@12.[151,1435,181,1479]" captionTargetId-3="figure-26@13.[151,1435,181,904]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 9. General habitus of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-1="FIGURE 10. General habitus of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-2="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." captionText-3="FIGURE 12. X-ray microtomographic reconstruction of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bar: 500 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13219258" figureDoi-1="http://doi.org/10.5281/zenodo.13219260" figureDoi-2="http://doi.org/10.5281/zenodo.13219262" figureDoi-3="http://doi.org/10.5281/zenodo.13219264" httpUri-0="https://zenodo.org/record/13219258/files/figure.png" httpUri-1="https://zenodo.org/record/13219260/files/figure.png" httpUri-2="https://zenodo.org/record/13219262/files/figure.png" httpUri-3="https://zenodo.org/record/13219264/files/figure.png" pageId="9" pageNumber="223">Figs 912</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2734FFF82DEBC724FDDA0F20" box="[151,635,1363,1390]" pageId="9" pageNumber="223" type="materials_examined">
<paragraph id="5B6A36B02734FFF82DEBC724FDDA0F20" blockId="9.[151,1437,1363,1966]" box="[151,635,1363,1390]" pageId="9" pageNumber="223">
<emphasis id="69A1EAA22734FFF82DEBC724FEA70F23" bold="true" box="[151,262,1363,1389]" pageId="9" pageNumber="223">Material.</emphasis>
<materialsCitation id="EBBD3CED2734FFF82C71C723FDDA0F20" box="[269,635,1364,1390]" collectionCode="NIGP" pageId="9" pageNumber="223" specimenCode="NIGP203941" specimenCount="1" typeStatus="holotype">
<typeStatus id="846E88122734FFF82C71C723FED90F20" box="[269,376,1364,1390]" pageId="9" pageNumber="223" type="holotype">Holotype</typeStatus>
,
<specimenCode id="0B739ECB2734FFF82CF8C723FDBA0F20" box="[388,539,1364,1390]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="9" pageNumber="223">NIGP203941</specimenCode>
, female.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2734FFF82DBBC70FFB430F94" pageId="9" pageNumber="223" type="etymology">
<paragraph id="5B6A36B02734FFF82DBBC70FFC590FDC" blockId="9.[151,1437,1363,1966]" box="[199,1016,1400,1426]" pageId="9" pageNumber="223">
<emphasis id="69A1EAA22734FFF82DBBC70FFEEF0FDC" bold="true" box="[199,334,1400,1426]" pageId="9" pageNumber="223">Etymology.</emphasis>
The species is named after the coleopterist Dr. Zhenhua Liu.
</paragraph>
<paragraph id="5B6A36B02734FFF82DBBC7ECFB430F94" blockId="9.[151,1437,1363,1966]" pageId="9" pageNumber="223">
<emphasis id="69A1EAA22734FFF82DBBC7ECFE6E0FFB" bold="true" box="[199,463,1435,1462]" pageId="9" pageNumber="223">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="9911F8522734FFF82DEBC7B7FE8B0F94" box="[151,298,1472,1498]" country="Myanmar" name="Kachin" pageId="9" pageNumber="223">Kachin State</collectingRegion>
,
<collectingCountry id="23C276202734FFF82C49C7B7FE070F94" box="[309,422,1472,1498]" name="Myanmar" pageId="9" pageNumber="223">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2734FFF82DBBC794FAC50CFC" pageId="9" pageNumber="223" type="diagnosis">
<paragraph id="5B6A36B02734FFF82DBBC794FAC50CFC" blockId="9.[151,1437,1363,1966]" pageId="9" pageNumber="223">
<emphasis id="69A1EAA22734FFF82DBBC794FEE30FB3" bold="true" box="[199,322,1507,1533]" pageId="9" pageNumber="223">Diagnosis.</emphasis>
Labrum weakly biemarginate (
<figureCitation id="C3EE2A352734FFF82FD3C793FCBB0FB0" box="[687,794,1508,1534]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11A</figureCitation>
). Apical maxillary palpomere asymmetrical (
<figureCitation id="C3EE2A352734FFF82859C793FA310FB0" box="[1317,1424,1508,1534]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11A</figureCitation>
). Basal antennomeres relatively elongate (especially antennomeres 35); antennomere 3 about 0.85 times as long as 4 (
<figureCitation id="C3EE2A352734FFF82DE3C45BFEA60C08" box="[159,263,1580,1606]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11A</figureCitation>
); antennomere 11 about 2.1 times as long as wide (
<figureCitation id="C3EE2A352734FFF82E38C45BFC0B0C08" box="[836,938,1580,1606]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11B</figureCitation>
). Scutellar shield acute at apex (
<figureCitation id="C3EE2A352734FFF82868C45BFAC20C08" box="[1300,1379,1580,1606]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11</figureCitation>
GA). Elytra about 2.1 times as long as combined width. Hind wings present. Metanepisternum at least twice as long as wide. Meso- and metatibiae with relatively dense spines along outer edge (
<figureCitation id="C3EE2A352734FFF82EA8C403FBF20CC0" box="[980,1107,1652,1678]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11D,E</figureCitation>
). Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia (
<figureCitation id="C3EE2A352734FFF82E2DC4EFFC190CFC" box="[849,952,1688,1714]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11D</figureCitation>
). Metatibial spurs unequal (
<figureCitation id="C3EE2A352734FFF8298CC4EFFAF60CFC" box="[1264,1367,1688,1714]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="9" pageNumber="223">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2734FFFB2DBBC4CCFB400C9F" lastPageId="10" lastPageNumber="224" pageId="9" pageNumber="223" type="description">
<paragraph id="5B6A36B02734FFF82DBBC4CCFAFC0C98" blockId="9.[151,1437,1363,1966]" box="[199,1373,1723,1750]" pageId="9" pageNumber="223">
<emphasis id="69A1EAA22734FFF82DBBC4CCFEF80C9B" bold="true" box="[199,345,1723,1749]" pageId="9" pageNumber="223">Description.</emphasis>
Body fusiform and slender, about
<quantity id="9C2D9B552734FFF82FA1C4CCFC920C98" box="[733,819,1723,1750]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="223" unit="mm" value="2.5">2.5 mm</quantity>
long, 1.0 mm wide; surface with fine short setae.
</paragraph>
<paragraph id="5B6A36B02734FFF82DBBC497FDD60DE0" blockId="9.[151,1437,1363,1966]" pageId="9" pageNumber="223">Head deflexed against ventral side of prothorax; entirely concealed from above by pronotum. Compound eyes weakly emarginate anteriorly, with interfacetal setae. Antennae 11-segmented, filiform; basal antennomeres distinctly elongate (especially antennomeres 25); distal antennomeres somewhat shortened; antennomere 3 about 0.85 times as long as 4; antennomere 11 about 2.1 times as long as wide. Frontoclypeal suture straight anteriorly and with distinct anterolateral angles. Labrum transverse, apically very weakly biemarginate. Mouthparts non-suctorial. Apical maxillary palpomere asymmetrical.</paragraph>
<caption id="0FAA66382737FFFB2DEBC693FDE70F6F" ID-DOI="http://doi.org/10.5281/zenodo.13219258" ID-Zenodo-Dep="13219258" httpUri="https://zenodo.org/record/13219258/files/figure.png" pageId="10" pageNumber="224" startId="10.[151,250,1252,1276]" targetBox="[151,1435,181,1229]" targetPageId="10" targetType="figure">
<paragraph id="5B6A36B02737FFFB2DEBC693FDE70F6F" blockId="10.[151,1437,1252,1313]" pageId="10" pageNumber="224">
<emphasis id="69A1EAA22737FFFB2DEBC693FEB20EB3" bold="true" box="[151,275,1252,1277]" pageId="10" pageNumber="224">FIGURE 9.</emphasis>
General habitus of
<taxonomicName id="9CD54D332737FFFB2CA1C692FCA40EB3" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[477,773,1253,1277]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="224" phylum="Arthropoda" rank="species" species="zhenhuai" status="sp. nov.">
<emphasis id="69A1EAA22737FFFB2CA1C692FD020EB3" box="[477,675,1253,1277]" italics="true" pageId="10" pageNumber="224">Eucinetus zhenhuai</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA22737FFFB2E70C692FCFC0EB3" bold="true" box="[780,861,1253,1277]" pageId="10" pageNumber="224">
<taxonomicNameLabel id="729257D92737FFFB2E70C692FCFC0EB3" box="[780,861,1253,1277]" pageId="10" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203941, under incident light.
<emphasis id="69A1EAA22737FFFB2845C693FAEA0EB2" bold="true" box="[1337,1355,1252,1276]" pageId="10" pageNumber="224">A</emphasis>
, Dorsal view.
<emphasis id="69A1EAA22737FFFB2DAFC77EFF450F6F" bold="true" box="[211,228,1289,1313]" pageId="10" pageNumber="224">B</emphasis>
, Ventral view. Scale bars: 500 μm.
</paragraph>
</caption>
<paragraph id="5B6A36B02737FFFB2DBBC704FB010C53" blockId="10.[151,1436,1395,1745]" pageId="10" pageNumber="224">Prothorax widest at base. Lateral pronotal carinae complete, simple. Pronotal disc simple. Procoxae obliquely oriented, well-projecting, narrowly separated. Scutellar shield triangular, acute at apex. Elytra about 2.1 times as long as combined width; transversely strigulate; sutural stria present; epipleuron narrow, incomplete. Hind wings present. Metaventral posterior process without externally discernable discrimen. Metanepisternum at least twice as long as wide. Metacoxae contiguous; metacoxal plates well-developed and greatly enlarged.</paragraph>
<paragraph id="5B6A36B02737FFFB2DBBC450FD430CE3" blockId="10.[151,1436,1395,1745]" pageId="10" pageNumber="224">
Mid and hind legs with fringe of spines at apices of tibiae and tarsomeres; fore legs without apical fringes of spines. Meso- and metatibiae with relatively dense spines along outer edge. Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia. Metatibial spurs unequal. Tarsi 5-5-5, simple. Meso- and metapretarsal claws with accessory lobes (
<figureCitation id="C3EE2A352737FFFB2F0FC4E4FD770CE3" box="[627,726,1683,1709]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="10" pageNumber="224">Fig. 11F</figureCitation>
).
</paragraph>
<paragraph id="5B6A36B02737FFFB2DBBC4C0FB400C9F" blockId="10.[151,1436,1395,1745]" box="[199,1249,1719,1745]" pageId="10" pageNumber="224">
Abdomen with five ventrites. Ovipositor simple, with gonocoxites and gonostyli (
<figureCitation id="C3EE2A352737FFFB2920C4C0FB700C9F" box="[1116,1233,1719,1745]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="10" pageNumber="224">Fig. 11H,I</figureCitation>
).
</paragraph>
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@ -0,0 +1,222 @@
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<mods:title id="4E1C046DB4C76131B138D685A8BC72B6">Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae)</mods:title>
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<mods:namePart id="FE87EE24633BB9535B72802139CF6EE4">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="9DD52BB6812061A3B5C175A3C254E75A">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China &amp; Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Bristol BS 8 1 TQ, UK</mods:affiliation>
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<mods:namePart id="2B38FD1AC728A581E1ED90F87D1F38C3">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="F03F766B8A33536166BF8AD7586219F1">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024 - 5192, USA &amp; Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Lima 15081, Perú &amp; Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima 15072, Perú</mods:affiliation>
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<mods:namePart id="F4818E64244EC4A62064673DA14CB1F9">Cai, Chen-Yang</mods:namePart>
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<subSubSection id="13CF653B2739FFF52DEBC2E0FD240AFF" box="[151,645,151,178]" pageId="4" pageNumber="218" type="nomenclature">
<paragraph id="5B6A36B02739FFF52DEBC2E0FD240AFF" blockId="4.[151,645,151,213]" box="[151,645,151,178]" pageId="4" pageNumber="218">
<heading id="002281DC2739FFF52DEBC2E0FD240AFF" bold="true" box="[151,645,151,178]" fontSize="11" level="1" pageId="4" pageNumber="218" reason="1">
<emphasis id="69A1EAA22739FFF52DEBC2E0FD240AFF" bold="true" box="[151,645,151,178]" pageId="4" pageNumber="218">
<taxonomicName id="9CD54D332739FFF52DEBC2E0FD810AFF" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[151,544,151,178]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="debilispinus" status="sp. nov.">
<emphasis id="69A1EAA22739FFF52DEBC2E0FE0B0AFC" bold="true" box="[151,426,151,178]" italics="true" pageId="4" pageNumber="218">Eucinetus debilispinus</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="729257D92739FFF52F5BC2E0FD240AFF" box="[551,645,151,177]" pageId="4" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="13CF653B2739FFF52DEBC2CCFEAA0A9B" box="[151,267,187,213]" pageId="4" pageNumber="218" type="description">
<paragraph id="5B6A36B02739FFF52DEBC2CCFEAA0A9B" blockId="4.[151,645,151,213]" box="[151,267,187,213]" pageId="4" pageNumber="218">
(
<figureCitation id="C3EE2A352739FFF52DE3C2CCFEA20A9B" box="[159,259,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1912,1936]" captionStartId-1="3.[151,250,1943,1967]" captionStartId-2="4.[151,250,1739,1763]" captionStartId-3="5.[151,250,1104,1128]" captionTargetBox-0="[151,1435,933,1889]" captionTargetBox-1="[169,1419,181,1919]" captionTargetBox-2="[151,1435,417,1715]" captionTargetBox-3="[151,1436,181,1081]" captionTargetId-0="figure-654@2.[151,1435,933,1889]" captionTargetId-1="figure-26@3.[169,1419,181,1919]" captionTargetId-2="figure-71@4.[151,1435,417,1715]" captionTargetId-3="figure-26@5.[151,1436,181,1081]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. General habitus of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under incident light. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bars: 500 μm." captionText-1="FIGURE 2. General habitus of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bars: 500 μm." captionText-2="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." captionText-3="FIGURE 4. X-ray microtomographic reconstruction of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bar: 500 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13219236" figureDoi-1="http://doi.org/10.5281/zenodo.13219238" figureDoi-2="http://doi.org/10.5281/zenodo.13219240" figureDoi-3="http://doi.org/10.5281/zenodo.13219244" httpUri-0="https://zenodo.org/record/13219236/files/figure.png" httpUri-1="https://zenodo.org/record/13219238/files/figure.png" httpUri-2="https://zenodo.org/record/13219240/files/figure.png" httpUri-3="https://zenodo.org/record/13219244/files/figure.png" pageId="4" pageNumber="218">Figs 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2739FFF52DEBC374FDC40B53" box="[151,613,259,285]" pageId="4" pageNumber="218" type="materials_examined">
<paragraph id="5B6A36B02739FFF52DEBC374FDC40B53" blockId="4.[151,1436,259,394]" box="[151,613,259,285]" pageId="4" pageNumber="218">
<emphasis id="69A1EAA22739FFF52DEBC374FEA70B53" bold="true" box="[151,262,259,285]" pageId="4" pageNumber="218">Material.</emphasis>
<materialsCitation id="EBBD3CED2739FFF52C71C374FDC40B53" box="[269,613,259,285]" collectionCode="NIGP" pageId="4" pageNumber="218" specimenCode="NIGP203939" specimenCount="1" typeStatus="holotype">
<typeStatus id="846E88122739FFF52C71C374FED90B53" box="[269,376,259,285]" pageId="4" pageNumber="218" type="holotype">Holotype</typeStatus>
,
<specimenCode id="0B739ECB2739FFF52CF8C374FDBD0B53" box="[388,540,259,285]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="4" pageNumber="218">NIGP203939</specimenCode>
, male.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2739FFF52DBBC350FB430BC7" pageId="4" pageNumber="218" type="etymology">
<paragraph id="5B6A36B02739FFF52DBBC350FB810B0C" blockId="4.[151,1436,259,394]" box="[199,1056,295,322]" pageId="4" pageNumber="218">
<emphasis id="69A1EAA22739FFF52DBBC350FEEF0B0F" bold="true" box="[199,334,295,321]" pageId="4" pageNumber="218">Etymology.</emphasis>
The specific name refers to its relatively small mesotibial spurs.
</paragraph>
<paragraph id="5B6A36B02739FFF52DBBC33CFB430BC7" blockId="4.[151,1436,259,394]" pageId="4" pageNumber="218">
<emphasis id="69A1EAA22739FFF52DBBC33CFE6E0B2B" bold="true" box="[199,463,331,357]" pageId="4" pageNumber="218">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="9911F8522739FFF52DEBC318FE8B0BC7" box="[151,298,367,393]" country="Myanmar" name="Kachin" pageId="4" pageNumber="218">Kachin State</collectingRegion>
,
<collectingCountry id="23C276202739FFF52C49C307FE070BC4" box="[309,422,368,394]" name="Myanmar" pageId="4" pageNumber="218">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
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<paragraph id="5B6A36B02739FFF52DEBC4BCFAEC0D3D" blockId="4.[151,1437,1739,1907]" pageId="4" pageNumber="218">
<emphasis id="69A1EAA22739FFF52DEBC4BCFEB20CAD" bold="true" box="[151,275,1739,1763]" pageId="4" pageNumber="218">FIGURE 3.</emphasis>
Details of
<taxonomicName id="9CD54D332739FFF52CFEC4BBFD660CAD" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[386,711,1739,1763]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="debilispinus" status="sp. nov.">
<emphasis id="69A1EAA22739FFF52CFEC4BBFDC70CAD" box="[386,614,1739,1763]" italics="true" pageId="4" pageNumber="218">Eucinetus debilispinus</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA22739FFF52FB1C4BCFCBC0CAD" bold="true" box="[717,797,1739,1763]" pageId="4" pageNumber="218">
<taxonomicNameLabel id="729257D92739FFF52FB1C4BCFCBC0CAD" box="[717,797,1739,1763]" pageId="4" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203939, under confocal microscopy.
<emphasis id="69A1EAA22739FFF5283EC4BCFAF50CAD" bold="true" box="[1346,1364,1739,1763]" pageId="4" pageNumber="218">A</emphasis>
, Head, dorsal view.
<emphasis id="69A1EAA22739FFF52C65C498FE8B0D49" bold="true" box="[281,298,1775,1799]" pageId="4" pageNumber="218">B</emphasis>
, Head and prothorax, lateral view.
<emphasis id="69A1EAA22739FFF52FE8C498FD070D49" bold="true" box="[660,678,1775,1799]" pageId="4" pageNumber="218">C</emphasis>
, Antennae.
<emphasis id="69A1EAA22739FFF52E5DC498FC920D49" bold="true" box="[801,819,1775,1799]" pageId="4" pageNumber="218">D</emphasis>
, Meso- and metathorax, ventral view.
<emphasis id="69A1EAA22739FFF529C3C498FB710D49" bold="true" box="[1215,1232,1775,1799]" pageId="4" pageNumber="218">E</emphasis>
, Fore and mid legs.
<emphasis id="69A1EAA22739FFF52DEBC564FF070D65" bold="true" box="[151,166,1811,1835]" pageId="4" pageNumber="218">F</emphasis>
, Hind legs.
<emphasis id="69A1EAA22739FFF52C54C564FE9D0D65" bold="true" box="[296,316,1811,1835]" pageId="4" pageNumber="218">G</emphasis>
, Scutellar shield, dorsal view.
<emphasis id="69A1EAA22739FFF52F02C564FD330D65" bold="true" box="[638,658,1811,1835]" pageId="4" pageNumber="218">H</emphasis>
, Elytral apex, dorsal view.
<emphasis id="69A1EAA22739FFF52ECEC564FC1D0D65" bold="true" box="[946,956,1811,1835]" pageId="4" pageNumber="218">I</emphasis>
, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm.
</paragraph>
</caption>
<caption id="0FAA66382738FFF42DEBC627FC8E0EC3" ID-DOI="http://doi.org/10.5281/zenodo.13219244" ID-Zenodo-Dep="13219244" httpUri="https://zenodo.org/record/13219244/files/figure.png" pageId="5" pageNumber="219" startId="5.[151,250,1104,1128]" targetBox="[151,1436,181,1081]" targetPageId="5" targetType="figure">
<paragraph id="5B6A36B02738FFF42DEBC627FC8E0EC3" blockId="5.[151,1436,1104,1165]" pageId="5" pageNumber="219">
<emphasis id="69A1EAA22738FFF42DEBC627FEB50E26" bold="true" box="[151,276,1104,1128]" pageId="5" pageNumber="219">FIGURE 4.</emphasis>
X-ray microtomographic reconstruction of
<taxonomicName id="9CD54D332738FFF42FAAC626FB810E27" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[726,1056,1105,1129]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="debilispinus" status="sp. nov.">
<emphasis id="69A1EAA22738FFF42FAAC626FC1D0E27" box="[726,956,1105,1129]" italics="true" pageId="5" pageNumber="219">Eucinetus debilispinus</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA22738FFF42954C626FBDB0E27" bold="true" box="[1064,1146,1105,1129]" pageId="5" pageNumber="219">
<taxonomicNameLabel id="729257D92738FFF42954C626FBDB0E27" box="[1064,1146,1105,1129]" pageId="5" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203939.
<emphasis id="69A1EAA22738FFF428F8C627FA370E26" bold="true" box="[1412,1430,1104,1128]" pageId="5" pageNumber="219">A</emphasis>
, Dorsal view.
<emphasis id="69A1EAA22738FFF42C61C602FE8F0EC3" bold="true" box="[285,302,1141,1165]" pageId="5" pageNumber="219">B</emphasis>
, Ventral view.
<emphasis id="69A1EAA22738FFF42CBAC603FE790EC2" bold="true" box="[454,472,1140,1164]" pageId="5" pageNumber="219">C</emphasis>
, Lateral view. Scale bar: 500 μm.
</paragraph>
</caption>
<subSubSection id="13CF653B2738FFF42DBBC6A6FD110FD2" pageId="5" pageNumber="219" type="diagnosis">
<paragraph id="5B6A36B02738FFF42DBBC6A6FD110FD2" blockId="5.[151,1437,1233,2040]" pageId="5" pageNumber="219">
<emphasis id="69A1EAA22738FFF42DBBC6A6FEE30EA5" bold="true" box="[199,322,1233,1259]" pageId="5" pageNumber="219">Diagnosis.</emphasis>
Labrum emarginate (
<figureCitation id="C3EE2A352738FFF42F48C6A6FD2F0EA5" box="[564,654,1233,1259]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3A</figureCitation>
). Apical maxillary palpomere narrow and symmetrical, apically acute (
<figureCitation id="C3EE2A352738FFF42DE3C683FEB30F40" box="[159,274,1268,1294]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3A,B</figureCitation>
). Antennomere 3 about 0.70 times as long as 4 (
<figureCitation id="C3EE2A352738FFF42E5DC683FCDB0F40" box="[801,890,1268,1294]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3A</figureCitation>
); antennomere 11 about 2.7 times as long as wide (
<figureCitation id="C3EE2A352738FFF42DE3C76FFF560F7C" box="[159,247,1304,1330]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3C</figureCitation>
). Elytra about 1.8 times as long as combined width. Hind wings present. Metanepisternum at least twice as long as wide (
<figureCitation id="C3EE2A352738FFF42C4EC74CFE2A0F1B" box="[306,395,1339,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3D</figureCitation>
). Meso- and metatibiae at most with only one or two spines along outer edge (
<figureCitation id="C3EE2A352738FFF42991C74CFAC30F1B" box="[1261,1378,1339,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[169,1419,181,1919]" captionTargetId="figure-26@3.[169,1419,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.13219238" httpUri="https://zenodo.org/record/13219238/files/figure.png" pageId="5" pageNumber="219">Figs 2B,C</figureCitation>
,
<figureCitation id="C3EE2A352738FFF42811C74CFA310F1B" box="[1389,1424,1339,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">3E</figureCitation>
). Mesotibial spurs small; the smaller one not distinctly stouter than spines along apical fringe of mesotibia (
<figureCitation id="C3EE2A352738FFF42845C728FA310F37" box="[1337,1424,1375,1401]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3E</figureCitation>
). Parameres distinctly bowed outwardly (
<figureCitation id="C3EE2A352738FFF42F29C7F5FD020FD2" box="[597,675,1410,1436]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="13CF653B2738FFF72DBBC7D2FEC10A9B" lastPageId="6" lastPageNumber="220" pageId="5" pageNumber="219" type="description">
<paragraph id="5B6A36B02738FFF42DBBC7D2FAFC0F8E" blockId="5.[151,1437,1233,2040]" box="[199,1373,1445,1472]" pageId="5" pageNumber="219">
<emphasis id="69A1EAA22738FFF42DBBC7D2FEF80FF1" bold="true" box="[199,345,1445,1471]" pageId="5" pageNumber="219">Description.</emphasis>
Body fusiform and slender, about
<quantity id="9C2D9B552738FFF42FA1C7D1FC920F8E" box="[733,819,1446,1472]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="219" unit="mm" value="1.7">1.7 mm</quantity>
long,
<quantity id="9C2D9B552738FFF42E05C7D1FC6E0F8E" box="[889,975,1446,1472]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="219" unit="mm" value="0.8">0.8 mm</quantity>
wide; surface with fine short setae.
</paragraph>
<paragraph id="5B6A36B02738FFF42DBBC7BEFE9B0CF6" blockId="5.[151,1437,1233,2040]" pageId="5" pageNumber="219">
Head deflexed against ventral side of prothorax; entirely concealed from above by pronotum. Compound eyes weakly emarginate anteriorly, with interfacetal setae. Antennae 11-segmented, filiform; antennomere 2 elongate; antennomeres 310 slightly longer than wide, somewhat submoniliform; antennomere 3 about 0.70 times as long as 4, distinctly narrower than 4; antennomere 11 about 2.7 times as long as wide, with distinct pattern of sensillae (
<figureCitation id="C3EE2A352738FFF42DE3C420FF580C3F" box="[159,249,1623,1649]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3C</figureCitation>
). Frontoclypeal suture straight anteriorly and with distinct anterolateral angles. Labrum weakly narrowed toward apex, apically emarginate. Mouthparts non-suctorial. Apical maxillary palpomere narrow and symmetrical, apically acute.
</paragraph>
<paragraph id="5B6A36B02738FFF42DBBC4B6FE5C0D27" blockId="5.[151,1437,1233,2040]" pageId="5" pageNumber="219">Prothorax widest at base. Lateral pronotal carinae complete, simple. Pronotal disc simple. Procoxae obliquely oriented, well-projecting, narrowly separated. Scutellar shield triangular, possibly with acute apex. Elytra about 1.8 times as long as combined width; transversely strigulate; sutural stria present; epipleuron narrow, incomplete. Hind wings present. Metanepisternum at least twice as long as wide. Metacoxae contiguous; metacoxal plates well-developed and greatly enlarged.</paragraph>
<paragraph id="5B6A36B02738FFF42DBBC504FD740DB6" blockId="5.[151,1437,1233,2040]" pageId="5" pageNumber="219">
Mid and hind legs with fringe of spines at apices of tibiae and tarsomeres; fore legs without apical fringes of spines. Meso- and metatibiae at most with only one or two spines along outer edge. Mesotibial spurs small; the smaller one not distinctly stouter than spines along apical fringe of mesotibia. Tarsi 5-5-5, simple. Meso- and metapretarsal claws with accessory lobes (
<figureCitation id="C3EE2A352738FFF42F0FC5A9FD690DB6" box="[627,712,2014,2040]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="5" pageNumber="219">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="5B6A36B0273BFFF72DBBC2E0FEC10A9B" blockId="6.[151,1436,151,213]" pageId="6" pageNumber="220">
Abdomen with six ventrites. Aedeagus of trilobate
<typeStatus id="846E8812273BFFF72E64C2E0FCED0AFF" box="[792,844,151,177]" pageId="6" pageNumber="220">type</typeStatus>
; penis shorter than parameres; parameres distinctly bowed outwardly.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,251 @@
<document id="96991FB9DA414C2D49EA1E60CC29D7F7" ID-DOI="10.11646/zootaxa.5492.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13219234" ID-ZooBank="880AEC67-7542-4E96-98CD-86A2C2A31DA3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722841220849" checkinUser="plazi" docAuthor="Li, Yan-Da, Engel, Michael S., Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2024" docId="D37C87A6273BFFF82DEBC350FD6F0A9B" docLanguage="en" docName="zootaxa.5492.1.4.pdf" docOrigin="Zootaxa 5492 (2)" docSource="http://zoobank.org/880aec67-7542-4e96-98cd-86a2c2a31da3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eucinetus panghongae Li &amp; Cai 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="222" masterDocId="2F45FFDE273DFFF12D7CC277FFA10A4E" masterDocTitle="Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae)" masterLastPageNumber="230" masterPageNumber="214" pageNumber="220" updateTime="1722841643882" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F8FF8D2F45F1A11208A26821ED7255C2">Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae)</mods:title>
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<mods:namePart id="01C8850FAB4AE64DD36E204E18B1DBE2">Li, Yan-Da</mods:namePart>
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<mods:affiliation id="AD123634E9FCCDA1C146288E86BC9DA3">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China &amp; Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Bristol BS 8 1 TQ, UK</mods:affiliation>
<mods:nameIdentifier id="4C8D259ED5BAD08436B7259D0DC2830A" type="email">ydli@pku.edu.cn</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="A281658B1797026C1E0D330C41B52DD8">Engel, Michael S.</mods:namePart>
<mods:nameIdentifier id="1182B46AD3C859412A026CF83D066FEC" type="ORCID">0000-0003-3067-077X</mods:nameIdentifier>
<mods:affiliation id="EE34838676D394E7859F0DF6C159B87E">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024 - 5192, USA &amp; Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Lima 15081, Perú &amp; Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima 15072, Perú</mods:affiliation>
<mods:nameIdentifier id="F96CA88E57639A7F28D1138B34891C66" type="email">mengel@amnh.org</mods:nameIdentifier>
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<mods:namePart id="14D869F6ACCF9302CF6D83F943C8FCD3">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="7156CDEFAC00BA08495CD2AB64122AE5">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
<mods:nameIdentifier id="AA9912ADB435503D5F9151EBD2D40530" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:namePart id="1F917B16C734C978B955F4FE5A751925">Cai, Chen-Yang</mods:namePart>
<mods:nameIdentifier id="BD091210451BFD569BDFE6DCB8A45020" type="ORCID">0000-0002-9283-8323</mods:nameIdentifier>
<mods:affiliation id="A09ADE9CBFB6721354BC13686B459090">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
<mods:nameIdentifier id="A695621E5E92C38EFAB672852E7AB321" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource id="CFB7286E1B762CA177840D0BB03B8CBB">text</mods:typeOfResource>
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<mods:title id="6156E068DE4E0CD3D6B9EDFC890FB62C">Zootaxa</mods:title>
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<mods:date id="52EBA4E0202A1A237925EB8B04ECA607">2024</mods:date>
<mods:detail id="6A2427EB5D622760EAB5D34D5E1CD507" type="pubDate">
<mods:number id="6A8F5F116CB5A5EE930D41AF3AE441B4">2024-08-05</mods:number>
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<mods:number id="C8F87A686DCBDB7A0AF5453B49A78885">5492</mods:number>
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<mods:identifier id="735BC4EBC93A8B736EA8F68F0C59D5E0" type="DOI">10.11646/zootaxa.5492.2.4</mods:identifier>
<mods:identifier id="117FA755839B10222F3F9F2507653334" type="ISSN">1175-5326</mods:identifier>
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<treatment id="D37C87A6273BFFF82DEBC350FD6F0A9B" ID-DOI="http://doi.org/10.5281/zenodo.13219246" ID-Zenodo-Dep="13219246" LSID="urn:lsid:plazi:treatment:D37C87A6273BFFF82DEBC350FD6F0A9B" httpUri="http://treatment.plazi.org/id/D37C87A6273BFFF82DEBC350FD6F0A9B" lastPageId="9" lastPageNumber="222" pageId="6" pageNumber="220">
<subSubSection id="13CF653B273BFFF72DEBC350FD2B0B0F" box="[151,650,294,321]" pageId="6" pageNumber="220" type="nomenclature">
<paragraph id="5B6A36B0273BFFF72DEBC350FD2B0B0F" blockId="6.[151,650,294,357]" box="[151,650,294,321]" pageId="6" pageNumber="220">
<heading id="002281DC273BFFF72DEBC350FD2B0B0F" bold="true" box="[151,650,294,321]" fontSize="11" level="1" pageId="6" pageNumber="220" reason="1">
<emphasis id="69A1EAA2273BFFF72DEBC350FD2B0B0F" bold="true" box="[151,650,294,321]" pageId="6" pageNumber="220">
<taxonomicName id="9CD54D33273BFFF72DEBC350FD840B0E" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[151,549,294,321]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="panghongae" status="sp. nov.">
<emphasis id="69A1EAA2273BFFF72DEBC350FE0E0B0F" bold="true" box="[151,431,294,321]" italics="true" pageId="6" pageNumber="220">Eucinetus panghongae</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="729257D9273BFFF72F50C350FD2B0B0F" box="[556,650,295,321]" pageId="6" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B273BFFF72DEBC33CFEAA0B2B" box="[151,267,330,357]" pageId="6" pageNumber="220" type="description">
<paragraph id="5B6A36B0273BFFF72DEBC33CFEAA0B2B" blockId="6.[151,650,294,357]" box="[151,267,330,357]" pageId="6" pageNumber="220">
(
<figureCitation id="C3EE2A35273BFFF72DE3C33CFEA20B2B" box="[159,259,330,357]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,250,1871,1895]" captionStartId-1="7.[151,250,1392,1416]" captionStartId-2="8.[151,250,1503,1527]" captionStartId-3="9.[151,250,1113,1137]" captionTargetBox-0="[151,1435,777,1848]" captionTargetBox-1="[151,1435,181,1368]" captionTargetBox-2="[151,1435,181,1479]" captionTargetBox-3="[151,1436,237,1090]" captionTargetId-0="figure-190@6.[151,1435,777,1848]" captionTargetId-1="figure-26@7.[151,1435,181,1368]" captionTargetId-2="figure-18@8.[151,1435,181,1479]" captionTargetId-3="figure-49@9.[151,1436,237,1090]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. General habitus of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-1="FIGURE 6. General habitus of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." captionText-2="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." captionText-3="FIGURE 8. X-ray microtomographic reconstruction of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bar: 500 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13219248" figureDoi-1="http://doi.org/10.5281/zenodo.13219250" figureDoi-2="http://doi.org/10.5281/zenodo.13219254" figureDoi-3="http://doi.org/10.5281/zenodo.13219256" httpUri-0="https://zenodo.org/record/13219248/files/figure.png" httpUri-1="https://zenodo.org/record/13219250/files/figure.png" httpUri-2="https://zenodo.org/record/13219254/files/figure.png" httpUri-3="https://zenodo.org/record/13219256/files/figure.png" pageId="6" pageNumber="220">Figs 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13CF653B273BFFF72DEBC3E5FDC40BE3" box="[151,613,402,429]" pageId="6" pageNumber="220" type="materials_examined">
<paragraph id="5B6A36B0273BFFF72DEBC3E5FDC40BE3" blockId="6.[151,1437,402,753]" box="[151,613,402,429]" pageId="6" pageNumber="220">
<emphasis id="69A1EAA2273BFFF72DEBC3E5FEA70BE2" bold="true" box="[151,262,402,428]" pageId="6" pageNumber="220">Material.</emphasis>
<materialsCitation id="EBBD3CED273BFFF72C71C3E4FDC40BE3" box="[269,613,403,429]" collectionCode="NIGP" pageId="6" pageNumber="220" specimenCode="NIGP203940" specimenCount="1" typeStatus="holotype">
<typeStatus id="846E8812273BFFF72C71C3E4FED90BE3" box="[269,376,403,429]" pageId="6" pageNumber="220" type="holotype">Holotype</typeStatus>
,
<specimenCode id="0B739ECB273BFFF72CF8C3E4FDBD0BE3" box="[388,540,403,429]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="6" pageNumber="220">NIGP203940</specimenCode>
, male.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B273BFFF72DBBC3C0FB430857" pageId="6" pageNumber="220" type="etymology">
<paragraph id="5B6A36B0273BFFF72DBBC3C0FC470B9F" blockId="6.[151,1437,402,753]" box="[199,998,439,465]" pageId="6" pageNumber="220">
<emphasis id="69A1EAA2273BFFF72DBBC3C0FEEF0B9F" bold="true" box="[199,334,439,465]" pageId="6" pageNumber="220">Etymology.</emphasis>
The species is named after the coleopterist Dr. Hong Pang.
</paragraph>
<paragraph id="5B6A36B0273BFFF72DBBC3ADFB430857" blockId="6.[151,1437,402,753]" pageId="6" pageNumber="220">
<emphasis id="69A1EAA2273BFFF72DBBC3ADFE6E0BBA" bold="true" box="[199,463,474,501]" pageId="6" pageNumber="220">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="9911F852273BFFF72DEBC388FE8B0857" box="[151,298,511,537]" country="Myanmar" name="Kachin" pageId="6" pageNumber="220">Kachin State</collectingRegion>
,
<collectingCountry id="23C27620273BFFF72C49C388FE070857" box="[309,422,511,537]" name="Myanmar" pageId="6" pageNumber="220">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="13CF653B273BFFF72DBBC055FCAB08BF" pageId="6" pageNumber="220" type="diagnosis">
<paragraph id="5B6A36B0273BFFF72DBBC055FCAB08BF" blockId="6.[151,1437,402,753]" pageId="6" pageNumber="220">
<emphasis id="69A1EAA2273BFFF72DBBC055FEE30872" bold="true" box="[199,322,546,572]" pageId="6" pageNumber="220">Diagnosis.</emphasis>
Labrum emarginate (
<figureCitation id="C3EE2A35273BFFF72F46C054FD360873" box="[570,663,547,573]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7A</figureCitation>
). Apical maxillary palpomere asymmetrical, apically with aciculate tip (
<figureCitation id="C3EE2A35273BFFF72DB9C030FE80082F" box="[197,289,583,609]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7A</figureCitation>
). Antennomere 3 about 0.85 times as long as 4 (
<figureCitation id="C3EE2A35273BFFF72E3BC030FC02082F" box="[839,931,583,609]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7A</figureCitation>
); antennomere 11 about 2.5 times as long as wide (
<figureCitation id="C3EE2A35273BFFF72DA1C01CFE9708CB" box="[221,310,619,645]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7B</figureCitation>
). Scutellar shield rounded at apex (
<figureCitation id="C3EE2A35273BFFF72FBEC01CFCB608CB" box="[706,791,619,645]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7F</figureCitation>
). Elytra about 1.8 times as long as combined width. Hind wings present. Metanepisternum at least twice as long as wide. Meso- and metatibiae at most with only one or two spines along outer edge (
<figureCitation id="C3EE2A35273BFFF72CD0C0C4FDA60883" box="[428,519,691,717]" captionStart="FIGURE 6" captionStartId="7.[151,250,1392,1416]" captionTargetBox="[151,1435,181,1368]" captionTargetId="figure-26@7.[151,1435,181,1368]" captionTargetPageId="7" captionText="FIGURE 6. General habitus of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Dorsal view. B, Ventral view. Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.13219250" httpUri="https://zenodo.org/record/13219250/files/figure.png" pageId="6" pageNumber="220">Figs 6B</figureCitation>
,
<figureCitation id="C3EE2A35273BFFF72F6EC0C4FD940883" box="[530,565,691,717]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">7E</figureCitation>
). Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia (
<figureCitation id="C3EE2A35273BFFF72C4EC0A0FE2D08BF" box="[306,396,727,753]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7D</figureCitation>
). Metatibial spurs equal (
<figureCitation id="C3EE2A35273BFFF72FD4C0A0FD5C08BF" box="[680,765,727,753]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="6" pageNumber="220">Fig. 7F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="0FAA6638273BFFF72DEBC538FD310DC2" ID-DOI="http://doi.org/10.5281/zenodo.13219248" ID-Zenodo-Dep="13219248" httpUri="https://zenodo.org/record/13219248/files/figure.png" pageId="6" pageNumber="220" startId="6.[151,250,1871,1895]" targetBox="[151,1435,777,1848]" targetPageId="6" targetType="figure">
<paragraph id="5B6A36B0273BFFF72DEBC538FD310DC2" blockId="6.[151,1436,1871,1932]" pageId="6" pageNumber="220">
<emphasis id="69A1EAA2273BFFF72DEBC538FEB70D29" bold="true" box="[151,278,1871,1895]" pageId="6" pageNumber="220">FIGURE 5.</emphasis>
General habitus of
<taxonomicName id="9CD54D33273BFFF72C95C527FC9D0D29" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[489,828,1871,1895]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="panghongae" status="sp. nov.">
<emphasis id="69A1EAA2273BFFF72C95C527FD720D29" box="[489,723,1871,1895]" italics="true" pageId="6" pageNumber="220">Eucinetus panghongae</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA2273BFFF72E3AC538FC380D29" bold="true" box="[838,921,1871,1895]" pageId="6" pageNumber="220">
<taxonomicNameLabel id="729257D9273BFFF72E3AC538FC380D29" box="[838,921,1871,1895]" pageId="6" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203940, under incident light.
<emphasis id="69A1EAA2273BFFF728F8C538FA370D29" bold="true" box="[1412,1430,1871,1895]" pageId="6" pageNumber="220">A</emphasis>
, Dorsal view.
<emphasis id="69A1EAA2273BFFF72C61C504FE8F0DC5" bold="true" box="[285,302,1907,1931]" pageId="6" pageNumber="220">B</emphasis>
, Ventral view. Scale bars: 500 μm.
</paragraph>
</caption>
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<paragraph id="5B6A36B0273AFFF62DEBC707FD0E0FE2" blockId="7.[151,1437,1392,1452]" pageId="7" pageNumber="221">
<emphasis id="69A1EAA2273AFFF62DEBC707FEB20FC6" bold="true" box="[151,275,1392,1416]" pageId="7" pageNumber="221">FIGURE 6.</emphasis>
General habitus of
<taxonomicName id="9CD54D33273AFFF62CA0C706FC840FC6" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[476,805,1392,1416]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="species" species="panghongae" status="sp. nov.">
<emphasis id="69A1EAA2273AFFF62CA0C706FD650FC6" box="[476,708,1392,1416]" italics="true" pageId="7" pageNumber="221">Eucinetus panghongae</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA2273AFFF62E50C707FCDD0FC6" bold="true" box="[812,892,1392,1416]" pageId="7" pageNumber="221">
<taxonomicNameLabel id="729257D9273AFFF62E50C707FCDD0FC6" box="[812,892,1392,1416]" pageId="7" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203940, under confocal microscopy.
<emphasis id="69A1EAA2273AFFF62DEBC7E3FF080FE2" bold="true" box="[151,169,1428,1452]" pageId="7" pageNumber="221">A</emphasis>
, Dorsal view.
<emphasis id="69A1EAA2273AFFF62C40C7E3FEEC0FE2" bold="true" box="[316,333,1428,1452]" pageId="7" pageNumber="221">B</emphasis>
, Ventral view. Scale bars: 500 μm.
</paragraph>
</caption>
<subSubSection id="13CF653B273AFFF92DBBC789FB3A0DCD" lastPageId="8" lastPageNumber="222" pageId="7" pageNumber="221" type="description">
<paragraph id="5B6A36B0273AFFF62DBBC789FAFC0C57" blockId="7.[151,1437,1534,1957]" box="[199,1373,1534,1561]" pageId="7" pageNumber="221">
<emphasis id="69A1EAA2273AFFF62DBBC789FEF80C56" bold="true" box="[199,345,1534,1560]" pageId="7" pageNumber="221">Description.</emphasis>
Body fusiform and slender, about
<quantity id="9C2D9B55273AFFF62FA1C789FC920C57" box="[733,819,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="221" unit="mm" value="1.6">1.6 mm</quantity>
long,
<quantity id="9C2D9B55273AFFF62E05C789FC6E0C57" box="[889,975,1534,1561]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="221" unit="mm" value="0.7">0.7 mm</quantity>
wide; surface with fine short setae.
</paragraph>
<paragraph id="5B6A36B0273AFFF62DBBC455FDC50CBE" blockId="7.[151,1437,1534,1957]" pageId="7" pageNumber="221">Head deflexed against ventral side of prothorax; entirely concealed from above by pronotum. Compound eyes weakly emarginate anteriorly, with interfacetal setae. Antennae 11-segmented, filiform; antennomere 2 elongate; antennomeres 310 moderately longer than wide; antennomere 3 about 0.85 times as long as 4; antennomere 11 about 2.5 times as long as wide. Frontoclypeal suture straight anteriorly and with distinct anterolateral angles. Labrum weakly narrowed toward apex, apically emarginate. Mouthparts non-suctorial. Apical maxillary palpomere asymmetrical, apically with aciculate tip.</paragraph>
<paragraph id="5B6A36B0273AFFF62DBBC48DFBA50DEA" blockId="7.[151,1437,1534,1957]" pageId="7" pageNumber="221">Prothorax widest at base. Lateral pronotal carinae complete, simple. Pronotal disc simple. Procoxae obliquely oriented, well-projecting, narrowly separated. Scutellar shield rounded at apex. Elytra about 1.8 times as long as combined width; transversely strigulate; sutural stria present; epipleuron narrow, incomplete. Hind wings present. Metaventral posterior process with distinct median discrimen. Metanepisternum at least twice as long as wide. Metacoxae contiguous; metacoxal plates well-developed and greatly enlarged.</paragraph>
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<paragraph id="5B6A36B02735FFF92DEBC7A8FD910CC9" blockId="8.[151,1436,1503,1671]" pageId="8" pageNumber="222">
<emphasis id="69A1EAA22735FFF92DEBC7A8FEB20FB9" bold="true" box="[151,275,1503,1527]" pageId="8" pageNumber="222">FIGURE 7.</emphasis>
Details of
<taxonomicName id="9CD54D332735FFF92CFEC797FD690FB9" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2024" box="[386,712,1503,1527]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="222" phylum="Arthropoda" rank="species" species="panghongae" status="sp. nov.">
<emphasis id="69A1EAA22735FFF92CFEC797FDC80FB9" box="[386,617,1503,1527]" italics="true" pageId="8" pageNumber="222">Eucinetus panghongae</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="69A1EAA22735FFF92FB2C7A8FCBE0FB9" bold="true" box="[718,799,1503,1527]" pageId="8" pageNumber="222">
<taxonomicNameLabel id="729257D92735FFF92FB2C7A8FCBE0FB9" box="[718,799,1503,1527]" pageId="8" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP203940, under confocal microscopy.
<emphasis id="69A1EAA22735FFF9283EC7A8FAF50FB9" bold="true" box="[1346,1364,1503,1527]" pageId="8" pageNumber="222">A</emphasis>
, Head, dorsal view.
<emphasis id="69A1EAA22735FFF92C60C474FE8C0C55" bold="true" box="[284,301,1539,1563]" pageId="8" pageNumber="222">B</emphasis>
, Antennae.
<emphasis id="69A1EAA22735FFF92CD5C474FE1A0C55" bold="true" box="[425,443,1539,1563]" pageId="8" pageNumber="222">C</emphasis>
, Mid leg.
<emphasis id="69A1EAA22735FFF92F56C474FD9D0C55" bold="true" box="[554,572,1539,1563]" pageId="8" pageNumber="222">D</emphasis>
, Fore and mid legs.
<emphasis id="69A1EAA22735FFF92E6AC474FC860C55" bold="true" box="[790,807,1539,1563]" pageId="8" pageNumber="222">E</emphasis>
, Mid leg.
<emphasis id="69A1EAA22735FFF92EE9C474FC050C55" bold="true" box="[917,932,1539,1563]" pageId="8" pageNumber="222">F</emphasis>
, Abdominal apex, ventral view.
<emphasis id="69A1EAA22735FFF9298BC474FAAA0C55" bold="true" box="[1271,1291,1539,1563]" pageId="8" pageNumber="222">G</emphasis>
, Hind leg.
<emphasis id="69A1EAA22735FFF928FEC474FA370C55" bold="true" box="[1410,1430,1539,1563]" pageId="8" pageNumber="222">H</emphasis>
, Scutellar shield, dorsal view.
<emphasis id="69A1EAA22735FFF92CC1C450FE660C71" bold="true" box="[445,455,1575,1599]" pageId="8" pageNumber="222">I</emphasis>
, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="5B6A36B02735FFF92DBBC4AEFD7B0D11" blockId="8.[151,1436,1753,2032]" pageId="8" pageNumber="222">
Mid and hind legs with fringe of spines at apices of tibiae and tarsomeres; fore legs without apical fringes of spines. Meso- and metatibiae at most with only one or two spines along outer edge. Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia. Metatibial spurs equal. Tarsi 5-5-5, simple. Meso- and metapretarsal claws with accessory lobes (
<figureCitation id="C3EE2A352735FFF92F0FC532FD680D11" box="[627,713,1861,1888]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="8" pageNumber="222">Fig. 7G</figureCitation>
).
</paragraph>
<paragraph id="5B6A36B02735FFF92DBBC51EFB3A0DCD" blockId="8.[151,1436,1753,2032]" box="[199,1179,1897,1924]" pageId="8" pageNumber="222">Abdomen with six ventrites. Posterior margin of ventrite 5 subtruncate, not emarginate.</paragraph>
</subSubSection>
<subSubSection id="13CF653B2735FFF82DBBC5FAFD6F0A9B" lastPageId="9" lastPageNumber="223" pageId="8" pageNumber="222" type="discussion">
<paragraph id="5B6A36B02735FFF82DBBC5FAFD6F0A9B" blockId="8.[151,1436,1753,2032]" lastBlockId="9.[151,1436,151,213]" lastPageId="9" lastPageNumber="223" pageId="8" pageNumber="222">
<emphasis id="69A1EAA22735FFF92DBBC5FAFE9A0DE9" bold="true" box="[199,315,1933,1959]" pageId="8" pageNumber="222">Remarks.</emphasis>
The sternite VIII of NIGP203940 (and also NIGP203939) is densely covered with setae like the preceding ventrites (
<figureCitation id="C3EE2A352735FFF92CFCC5C6FE760D85" box="[384,471,1969,1995]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="8" pageNumber="222">Fig. 7F</figureCitation>
). It should be normally exposed when alive, representing the sixth ventrite. In some recent comprehensive coleopteran compendia, the abdomen is described as having only five ventrites for the whole
<taxonomicName id="9CD54D332734FFF82DEBC2E0FEBA0AFF" authorityName="Lacordaire" authorityYear="1857" box="[151,283,151,177]" class="Insecta" family="Eucinetidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="family">Eucinetidae</taxonomicName>
(
<bibRefCitation id="3F444B412734FFF82C55C2E0FDC00AFF" author="Lawrence, J. F. &amp; Slipinski, A." box="[297,609,151,179]" pageId="9" pageNumber="223" refId="ref6438" refString="Lawrence, J. F. &amp; Slipinski, A. (2013) Australian beetles. Volume 1: morphology, classification and keys. CSIRO Publishing, Collingwood, Australia https: // doi. org / 10.1071 / 9780643097292" type="book" year="2013">Lawrence &amp; Ślipiński 2013</bibRefCitation>
;
<bibRefCitation id="3F444B412734FFF82F11C2E0FCAC0AFF" author="Leschen, R. A. B." box="[621,781,151,177]" pageId="9" pageNumber="223" pagination="206 - 210" refId="ref6480" refString="Leschen, R. A. B. (2016) Eucinetidae Lacordaire, 1857. In: Beutel, R. G. &amp; Leschen, R. A. B. (Eds.), Handbook of Zoology, Arthropoda: Insecta, Coleoptera, beetles, Vol. 1: morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). 2 nd Edition, Walter de Gruyter, Berlin, pp. 206 - 210. https: // doi. org / 10.1515 / 9783110373929 - 013" type="book chapter" year="2016">Leschen 2016</bibRefCitation>
). However, the sternite VIII is actually commonly exposed in many extant members of
<taxonomicName id="9CD54D332734FFF82CADC2CCFD680A9B" authority="(Vit 1977 a)" baseAuthorityName="Vit" baseAuthorityYear="1977" box="[465,713,187,213]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA22734FFF82CADC2CCFD9E0A9A" box="[465,575,187,212]" italics="true" pageId="9" pageNumber="223">Eucinetus</emphasis>
(
<bibRefCitation id="3F444B412734FFF82F31C2CCFD600A9B" author="Vit, S." box="[589,705,187,213]" pageId="9" pageNumber="223" pagination="443 - 451" refId="ref7105" refString="Vit, S. (1977 a) Contribution a la connaissance du genre Eucinetus Germar (Coleoptera, Eucinetidae). Revue Suisse de Zoologie, 84, 443 - 451. https: // doi. org / 10.5962 / bhl. part. 91398" type="journal article" year="1977">Vit 1977a</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B3C740090BC5293A7C23A222811D1E33">Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae)</mods:title>
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<mods:identifier id="44C44E056BE343B6BDFF1AF29C41DE32" type="DOI">10.11646/zootaxa.5492.2.4</mods:identifier>
<mods:identifier id="BFC84A56CDF3F862F994200BE7C302A2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="27F320E0B1F9464BC0B3F17747E3BF28" type="Zenodo-Dep">13219234</mods:identifier>
<mods:identifier id="851296D9AA5BEA90C6FD5B89D1B21F69" type="ZooBank">880AEC67-7542-4E96-98CD-86A2C2A31DA3</mods:identifier>
</mods:mods>
<treatment id="D37C87A6273CFFF32DEBC799FF510BBB" LSID="urn:lsid:plazi:treatment:D37C87A6273CFFF32DEBC799FF510BBB" httpUri="http://treatment.plazi.org/id/D37C87A6273CFFF32DEBC799FF510BBB" lastPageId="2" lastPageNumber="216" pageId="1" pageNumber="215">
<subSubSection id="13CF653B273CFFF02DEBC799FDBD0C46" box="[151,540,1518,1545]" pageId="1" pageNumber="215" type="nomenclature">
<paragraph id="5B6A36B0273CFFF02DEBC799FDBD0C46" blockId="1.[151,540,1518,1545]" box="[151,540,1518,1545]" pageId="1" pageNumber="215">
<heading id="002281DC273CFFF02DEBC799FDBD0C46" bold="true" box="[151,540,1518,1545]" fontSize="11" level="1" pageId="1" pageNumber="215" reason="1">
<emphasis id="69A1EAA2273CFFF02DEBC799FDBD0C46" bold="true" box="[151,540,1518,1545]" pageId="1" pageNumber="215">
Genus
<taxonomicName id="9CD54D33273CFFF02D92C798FDBD0C46" ID-CoL="4F7S" authority="Germar, 1818" authorityName="Germar" authorityYear="1818" box="[238,540,1518,1545]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02D92C798FEC90C47" bold="true" box="[238,360,1519,1545]" italics="true" pageId="1" pageNumber="215">Eucinetus</emphasis>
Germar, 1818
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13CF653B273CFFF32DEBC440FF510BBB" lastPageId="2" lastPageNumber="216" pageId="1" pageNumber="215" type="discussion">
<paragraph id="5B6A36B0273CFFF02DEBC440FAD40D03" blockId="1.[151,1437,1591,2013]" pageId="1" pageNumber="215">
<emphasis id="69A1EAA2273CFFF02DEBC440FEAA0C1F" bold="true" box="[151,267,1591,1617]" pageId="1" pageNumber="215">Remarks.</emphasis>
The three new species described here have unspecialized mouthparts, distinguishing them from the genera with suctorial or semisuctorial mouthparts:
<emphasis id="69A1EAA2273CFFF02FB0C42CFC9F0C3B" box="[716,830,1627,1653]" italics="true" pageId="1" pageNumber="215">Bisayodes</emphasis>
Wakimura &amp; Jałoszyński,
<taxonomicName id="9CD54D33273CFFF02914C42CFAA00C3B" authority="Vit" authorityName="Vit" authorityYear="1985" box="[1128,1281,1627,1653]" class="Insecta" family="Eucinetidae" genus="Eucilodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02914C42CFB770C3B" box="[1128,1238,1627,1653]" italics="true" pageId="1" pageNumber="215">Eucilodes</emphasis>
Vit
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02877C42CFEBA0CD7" authority="Jaloszynski" authorityName="Jaloszynski" class="Insecta" family="Eucinetidae" genus="Cretohlezkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02877C42CFA3D0C3B" box="[1291,1436,1627,1653]" italics="true" pageId="1" pageNumber="215">Cretohlezkus</emphasis>
Jałoszyński
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02C59C408FE640CD7" authority="Reitter" authorityName="Reitter" authorityYear="1884" box="[293,453,1663,1689]" class="Insecta" family="Eucinetidae" genus="Bisaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02C59C408FED00CD6" box="[293,369,1663,1688]" italics="true" pageId="1" pageNumber="215">Bisaya</emphasis>
Reitter
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02CB2C408FDC40CD7" authority="Vit" authorityName="Vit" box="[462,613,1663,1689]" class="Insecta" family="Eucinetidae" genus="Jentozkus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02CB2C408FD9A0CD7" box="[462,571,1663,1689]" italics="true" pageId="1" pageNumber="215">Jentozkus</emphasis>
Vit
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02F13C408FCA20CD7" authority="Vit" authorityName="Vit" box="[623,771,1663,1689]" class="Insecta" family="Eucinetidae" genus="Tohlezkus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02F13C408FD7D0CD7" box="[623,732,1663,1689]" italics="true" pageId="1" pageNumber="215">Tohlezkus</emphasis>
Vit
</taxonomicName>
and
<taxonomicName id="9CD54D33273CFFF02E4BC408FC700CD7" authority="Vit" authorityName="Vit" authorityYear="2000" box="[823,977,1663,1689]" class="Insecta" family="Eucinetidae" genus="Proeuzkus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02E4BC408FC0A0CD7" box="[823,939,1663,1689]" italics="true" pageId="1" pageNumber="215">Proeuzkus</emphasis>
Vit
</taxonomicName>
(
<emphasis id="69A1EAA2273CFFF02E9CC408FBA90CD6" box="[992,1032,1663,1688]" italics="true" pageId="1" pageNumber="215">e.g.</emphasis>
,
<bibRefCitation id="3F444B41273CFFF02969C408FB240CD7" author="Vit, S." box="[1045,1157,1663,1689]" pageId="1" pageNumber="215" pagination="917 - 935" refId="ref7152" refString="Vit, S. (1977 b) Contribution a la connaissance des Eucinetidae (Coleoptera). Revue Suisse de Zoologie, 84, 917 - 935. https: // doi. org / 10.5962 / bhl. part. 91368" type="journal article" year="1977">Vit 1977b</bibRefCitation>
,
<bibRefCitation id="3F444B41273CFFF029F3C408FB680CD7" author="Vit, S." box="[1167,1225,1663,1689]" pageId="1" pageNumber="215" pagination="769 - 774" refId="ref7195" refString="Vit, S. (1981) Une nouvelle espece du genre Tohlezkus Vit de Taiwan (Coleoptera, Eucinetidae). Revue Suisse de Zoologie, 88, 769 - 774. https: // doi. org / 10.5962 / bhl. part. 82406" type="journal article" year="1981">1981</bibRefCitation>
,
<bibRefCitation id="3F444B41273CFFF029AFC408FAAE0CD7" author="Vit, S." box="[1235,1295,1663,1689]" pageId="1" pageNumber="215" pagination="123 - 138" refId="ref7433" refString="Vit, S. (2000) Contribution a la connaissance de la famille Eucinetidae (Coleoptera). Revue Suisse de Zoologie, 107, 123 - 138. https: // doi. org / 10.5962 / bhl. part. 80122" type="journal article" year="2000">2000</bibRefCitation>
;
<bibRefCitation id="3F444B41273CFFF02865C408FA360CD7" author="Sakai, M." box="[1305,1431,1663,1689]" pageId="1" pageNumber="215" pagination="83 - 85" refId="ref7014" refString="Sakai, M. (1980) A new genus of Eucinetidae from Japan (Coleoptera). Transactions of the Shikoku Entomological Society, 15, 83 - 85." type="journal article" year="1980">Sakai 1980</bibRefCitation>
;
<bibRefCitation id="3F444B41273CFFF02DEBC4D4FEFD0CF3" author="Jaloszynski, P." box="[151,348,1699,1725]" pageId="1" pageNumber="215" pagination="126 - 133" refId="ref5959" refString="Jaloszynski, P. (2019) † Cretohlezkus gen. nov. from Upper Cretaceous Burmese amber demonstrates ancient origins of suctorial mouthparts in Eucinetidae (Coleoptera: Scirtoidea). Cretaceous Research, 100, 126 - 133. https: // doi. org / 10.1016 / j. cretres. 2019.03.016" type="journal article" year="2019">Jałoszyński 2019</bibRefCitation>
;
<bibRefCitation id="3F444B41273CFFF02C16C4D4FD6C0CF3" author="Jaloszynski, P. &amp; Wakimura, R." box="[362,717,1699,1725]" pageId="1" pageNumber="215" pagination="122 - 138" refId="ref6014" refString="Jaloszynski, P. &amp; Wakimura, R. (2023) A new genus, new species and new records of Eucinetidae in Japan and Taiwan (Coleoptera). Zootaxa, 5351 (1), 122 - 138. https: // doi. org / 10.11646 / zootaxa. 5351.1.5" type="journal article" year="2023">Jałoszyński &amp; Wakimura 2023</bibRefCitation>
). The remaining “non-suctorial” group of
<taxonomicName id="9CD54D33273CFFF029CEC4D4FA970CF3" authorityName="Lacordaire" authorityYear="1857" box="[1202,1334,1699,1725]" class="Insecta" family="Eucinetidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="family">Eucinetidae</taxonomicName>
includes
<taxonomicName id="9CD54D33273CFFF02DEBC4B0FEDF0CAF" authority="Nikitsky" authorityName="Nikitsky" authorityYear="1996" box="[151,382,1735,1761]" class="Insecta" family="Eucinetidae" genus="Eucinetella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02DEBC4B0FEB60CAF" box="[151,279,1735,1761]" italics="true" pageId="1" pageNumber="215">Eucinetella</emphasis>
Nikitsky
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02CFBC4B0FE540CAE" box="[391,501,1735,1760]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02CFBC4B0FE540CAE" box="[391,501,1735,1760]" italics="true" pageId="1" pageNumber="215">Eucinetus</emphasis>
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02F7EC4B0FD630CAF" authority="Latreille" authorityName="Latreille" authorityYear="1829" box="[514,706,1735,1761]" class="Insecta" family="Eucinetidae" genus="Nycteus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02F7EC4B0FDFA0CAE" box="[514,603,1735,1760]" italics="true" pageId="1" pageNumber="215">Nycteus</emphasis>
Latreille
</taxonomicName>
,
<taxonomicName id="9CD54D33273CFFF02FB7C4B0FC060CAF" authority="Casey" authorityName="Casey" authorityYear="1885" box="[715,935,1735,1761]" class="Insecta" family="Eucinetidae" genus="Euscaphurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02FB7C4B0FCFC0CAF" box="[715,861,1735,1761]" italics="true" pageId="1" pageNumber="215">Euscaphurus</emphasis>
Casey
</taxonomicName>
and
<taxonomicName id="9CD54D33273CFFF02EA7C4B0FAF40CAF" authority="Bullians &amp; Leschen." authorityName="Bullians &amp; Leschen." box="[987,1365,1735,1761]" class="Insecta" family="Eucinetidae" genus="Noteucinetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02EA7C4B0FBCC0CAE" box="[987,1133,1735,1760]" italics="true" pageId="1" pageNumber="215">Noteucinetus</emphasis>
Bullians &amp; Leschen.
</taxonomicName>
These new species differ from
<taxonomicName id="9CD54D33273CFFF02CD4C49CFD890D4B" authorityName="Nikitsky" authorityYear="1996" box="[424,552,1771,1797]" class="Insecta" family="Eucinetidae" genus="Eucinetella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02CD4C49CFD890D4B" box="[424,552,1771,1797]" italics="true" pageId="1" pageNumber="215">Eucinetella</emphasis>
</taxonomicName>
and
<taxonomicName id="9CD54D33273CFFF02F23C49CFD500D4B" authorityName="Casey" authorityYear="1885" box="[607,753,1771,1797]" class="Insecta" family="Eucinetidae" genus="Euscaphurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02F23C49CFD500D4B" box="[607,753,1771,1797]" italics="true" pageId="1" pageNumber="215">Euscaphurus</emphasis>
</taxonomicName>
by having slender and filiform antennae, with antennomeres loosely assembled (antennae claviform to subclaviform in
<taxonomicName id="9CD54D33273CFFF02E5AC578FC070D67" authorityName="Nikitsky" authorityYear="1996" box="[806,934,1807,1833]" class="Insecta" family="Eucinetidae" genus="Eucinetella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02E5AC578FC070D67" box="[806,934,1807,1833]" italics="true" pageId="1" pageNumber="215">Eucinetella</emphasis>
</taxonomicName>
and
<taxonomicName id="9CD54D33273CFFF02EA0C578FBCF0D67" authorityName="Casey" authorityYear="1885" box="[988,1134,1807,1833]" class="Insecta" family="Eucinetidae" genus="Euscaphurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02EA0C578FBCF0D67" box="[988,1134,1807,1833]" italics="true" pageId="1" pageNumber="215">Euscaphurus</emphasis>
</taxonomicName>
:
<bibRefCitation id="3F444B41273CFFF02901C578FA810D67" author="Nikitsky, N. B." box="[1149,1312,1807,1833]" pageId="1" pageNumber="215" pagination="1366 - 1373" refId="ref6975" refString="Nikitsky, N. B. (1996) New Coleoptera from China. Zoologicheskii Zhurnal, 75, 1366 - 1373. [in Russian; English translation in Entomological Review, 76, 1231 - 1237]" type="journal article" year="1996">Nikitsky 1996</bibRefCitation>
;
<bibRefCitation id="3F444B41273CFFF02857C578FA310D67" author="Vit, S." box="[1323,1424,1807,1833]" pageId="1" pageNumber="215" pagination="125 - 137" refId="ref7365" refString="Vit, S. (1996) Deux especes nouvelles d'Eucinetidae d'Amerique du Nord particulierement interessantes (Coleoptera: Eucinetidae). Elytron, 9, 125 - 137." type="journal article" year="1996">Vit 1996</bibRefCitation>
), and from
<taxonomicName id="9CD54D33273CFFF02C78C544FE370D02" box="[260,406,1843,1868]" class="Insecta" family="Eucinetidae" genus="Noteucinetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02C78C544FE370D02" box="[260,406,1843,1868]" italics="true" pageId="1" pageNumber="215">Noteucinetus</emphasis>
</taxonomicName>
in the more elongate and less convex body (
<bibRefCitation id="3F444B41273CFFF02EF5C544FB0D0D03" author="Bullians, M. S. &amp; Leschen, R. A. B." box="[905,1196,1843,1869]" pageId="1" pageNumber="215" pagination="29 - 38" refId="ref5254" refString="Bullians, M. S. &amp; Leschen, R. A. B. (2004) Noteucinetus new genus from New Zealand and Chile and notes on Eucinetus stewarti (Broun) (Coleoptera: Eucinetidae). New Zealand Entomologist, 27, 29 - 38. https: // doi. org / 10.1080 / 00779962.2004.9722121" type="journal article" year="2004">Bullians &amp; Leschen 2004</bibRefCitation>
;
<bibRefCitation id="3F444B41273CFFF029CAC544FAC80D03" author="Lawrence, J. F." box="[1206,1385,1843,1869]" pageId="1" pageNumber="215" pagination="151 - 182" refId="ref6299" refString="Lawrence, J. F. (2019) New species of Eucinetus and Noteucinetus from Australia (Coleoptera: Scirtoidea: Eucinetidae). Zootaxa, 4668 (2), 151 - 182. https: // doi. org / 10.11646 / zootaxa. 4668.2.1" type="journal article" year="2019">Lawrence 2019</bibRefCitation>
).
</paragraph>
<paragraph id="5B6A36B0273CFFF32DBBC520FA800B0F" blockId="1.[151,1437,1591,2013]" lastBlockId="2.[151,1437,150,501]" lastPageId="2" lastPageNumber="216" pageId="1" pageNumber="215">
<bibRefCitation id="3F444B41273CFFF02DBBC520FED90D3F" author="Du, X. &amp; Slipinski, A. &amp; Liu, Z. &amp; Pang, H." box="[199,376,1879,1905]" pageId="1" pageNumber="215" pagination="1 - 9" refId="ref5661" refString="Du, X., Slipinski, A., Liu, Z. &amp; Pang, H. (2020) Description of a new species of Eucinetidae (Coleoptera, Scirtoidea) from Cretaceous Burmese amber. ZooKeys, 982, 1 - 9. https: // doi. org / 10.3897 / zookeys. 982.39335" type="journal article" year="2020">
Du
<emphasis id="69A1EAA2273CFFF02D8CC520FE860D3F" box="[240,295,1879,1905]" italics="true" pageId="1" pageNumber="215">et al.</emphasis>
(2020)
</bibRefCitation>
claimed that the antennomere
<quantity id="9C2D9B55273CFFF02FB1C520FD560D3F" box="[717,759,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="215" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="9CD54D33273CFFF02F81C520FCF70D3E" authorityName="Latreille" authorityYear="1829" box="[765,854,1879,1904]" class="Insecta" family="Eucinetidae" genus="Nycteus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02F81C520FCF70D3E" box="[765,854,1879,1904]" italics="true" pageId="1" pageNumber="215">Nycteus</emphasis>
</taxonomicName>
is distinctly shorter than the adjacent antennomeres, while in
<taxonomicName id="9CD54D33273CFFF02D80C50CFECB0DDA" box="[252,362,1915,1940]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02D80C50CFECB0DDA" box="[252,362,1915,1940]" italics="true" pageId="1" pageNumber="215">Eucinetus</emphasis>
</taxonomicName>
it is almost the same length as antennomere 4. However, according to
<bibRefCitation id="3F444B41273CFFF029EDC50CFAF50DDB" author="Lawrence, J. F." box="[1169,1364,1915,1941]" pageId="1" pageNumber="215" pagination="151 - 182" refId="ref6299" refString="Lawrence, J. F. (2019) New species of Eucinetus and Noteucinetus from Australia (Coleoptera: Scirtoidea: Eucinetidae). Zootaxa, 4668 (2), 151 - 182. https: // doi. org / 10.11646 / zootaxa. 4668.2.1" type="journal article" year="2019">Lawrence (2019)</bibRefCitation>
, some
<taxonomicName id="9CD54D33273CFFF02DEBC5E8FEFA0DF7" box="[151,347,1951,1977]" pageId="1" pageNumber="215">
<emphasis id="69A1EAA2273CFFF02DEBC5E8FEA40DF6" box="[151,261,1951,1976]" italics="true" pageId="1" pageNumber="215">Eucinetus</emphasis>
species
</taxonomicName>
also have a short antennomere 3 (
<emphasis id="69A1EAA2273CFFF02FB4C5E8FD510DF6" box="[712,752,1951,1976]" italics="true" pageId="1" pageNumber="215">e.g.</emphasis>
,
<taxonomicName id="9CD54D33273CFFF02F80C5E8FBBB0DF7" authority="Lawrence" authorityName="Lawrence" authorityYear="2019" box="[764,1050,1951,1977]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="species" species="brindabellae">
<emphasis id="69A1EAA2273CFFF02F80C5E8FC060DF7" box="[764,935,1951,1977]" italics="true" pageId="1" pageNumber="215">E. brindabellae</emphasis>
Lawrence
</taxonomicName>
and
<taxonomicName id="9CD54D33273CFFF02937C5E8FA880DF7" authority="Lawrence" authorityName="Lawrence" authorityYear="2019" box="[1099,1321,1951,1977]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="69A1EAA2273CFFF02937C5E8FB100DF7" box="[1099,1201,1951,1977]" italics="true" pageId="1" pageNumber="215">E. similis</emphasis>
Lawrence
</taxonomicName>
).Actually,
<taxonomicName id="9CD54D33273CFFF02DEBC5B4FF510D92" authorityName="Latreille" authorityYear="1829" box="[151,240,1987,2012]" class="Insecta" family="Eucinetidae" genus="Nycteus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02DEBC5B4FF510D92" box="[151,240,1987,2012]" italics="true" pageId="1" pageNumber="215">Nycteus</emphasis>
</taxonomicName>
could be separated from
<taxonomicName id="9CD54D33273CFFF02F74C5B4FDD70D92" box="[520,630,1987,2012]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273CFFF02F74C5B4FDD70D92" box="[520,630,1987,2012]" italics="true" pageId="1" pageNumber="215">Eucinetus</emphasis>
</taxonomicName>
by the completely exposed antennal insertions and antennal grooves, U- shaped frontoclypeal suture and acute inner anterior angle of mesepimeron (
<bibRefCitation id="3F444B41273FFFF32EA2C2E0FBE50AFF" author="Vit, S." box="[990,1092,150,177]" pageId="2" pageNumber="216" pagination="421 - 460" refId="ref7279" refString="Vit, S. (1985) Etude de la morphologie des especes palearctiques du genre Eucinetus Germar et quelques remarques sur son utilisation taxonomique (Coleoptera Eucinetidae). Revue Suisse de Zoologie, 92, 421 - 460. https: // doi. org / 10.5962 / bhl. part. 81624" type="journal article" year="1985">Vit 1985</bibRefCitation>
: figs 2, 61;
<bibRefCitation id="3F444B41273FFFF329BDC2E0FA820AFF" author="Vit, S." box="[1217,1315,151,177]" pageId="2" pageNumber="216" pagination="387 - 394" refId="ref7395" refString="Vit, S. (1999) Sur les genres Nycteus Latreille, 1829, et Eucinetus Germar, 1818 (Coleoptera, Eucinetidae). Mitteilungen der Schwizerischen Entomologischen Gesellschaft, 72, 387 - 394." type="journal article" year="1999">Vit 1999</bibRefCitation>
;
<bibRefCitation id="3F444B41273FFFF32852C2E0FF750A9B" author="Lawrence, J. F." pageId="2" pageNumber="216" pagination="151 - 182" refId="ref6299" refString="Lawrence, J. F. (2019) New species of Eucinetus and Noteucinetus from Australia (Coleoptera: Scirtoidea: Eucinetidae). Zootaxa, 4668 (2), 151 - 182. https: // doi. org / 10.11646 / zootaxa. 4668.2.1" type="journal article" year="2019">Lawrence 2019</bibRefCitation>
). The three new species all have a frontoclypeal suture straight anteriorly with distinct anterolateral angles (
<figureCitation id="C3EE2A35273FFFF32DE3C2A8FF5D0AB7" box="[159,252,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1739,1763]" captionTargetBox="[151,1435,417,1715]" captionTargetId="figure-71@4.[151,1435,417,1715]" captionTargetPageId="4" captionText="FIGURE 3. Details of Eucinetus debilispinus Li &amp; Cai sp. nov., holotype, NIGP203939, under confocal microscopy. A, Head, dorsal view. B, Head and prothorax, lateral view. C, Antennae. D, Meso- and metathorax, ventral view. E, Fore and mid legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Elytral apex, dorsal view. I, Abdominal apex, dorsal view. Abbreviations: a111, antennomeres 111; lb, labrum; msem, mesepimeron; mstb, mesotibia; msts, mesotarsus; mtes, metanepisternum; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pm, paramere; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219240" httpUri="https://zenodo.org/record/13219240/files/figure.png" pageId="2" pageNumber="216">Figs 3A</figureCitation>
,
<figureCitation id="C3EE2A35273FFFF32C74C2A8FE8C0AB7" box="[264,301,223,249]" captionStart="FIGURE 7" captionStartId="8.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@8.[151,1435,181,1479]" captionTargetPageId="8" captionText="FIGURE 7. Details of Eucinetus panghongae Li &amp; Cai sp. nov., holotype, NIGP203940, under confocal microscopy. A, Head, dorsal view. B, Antennae. C, Mid leg. D, Fore and mid legs. E, Mid leg. F, Abdominal apex, ventral view. G, Hind leg. H, Scutellar shield, dorsal view. I, Abdominal apex, dorsal view.Abbreviations: a211, antennomeres 211; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v56, ventrites 56. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219254" httpUri="https://zenodo.org/record/13219254/files/figure.png" pageId="2" pageNumber="216">7A</figureCitation>
,
<figureCitation id="C3EE2A35273FFFF32C44C2A8FEC80AB7" box="[312,361,223,249]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1479]" captionTargetId="figure-18@12.[151,1435,181,1479]" captionTargetPageId="12" captionText="FIGURE 11. Details of Eucinetus zhenhuai Li &amp; Cai sp. nov., holotype, NIGP203941, under confocal microscopy. A, Head, dorsal view. B, Antenna. C, Fore legs. D, Mid leg. E, Hind legs. F, Hind legs. G, Scutellar shield, dorsal view. H, Abdominal apex, dorsal view. I, Abdominal apex, ventral view. Abbreviations: a211, antennomeres 211; gc, gonocoxite; gs, gonostylus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pts, protarsus; sc, scutellar shield. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.13219262" httpUri="https://zenodo.org/record/13219262/files/figure.png" pageId="2" pageNumber="216">11A</figureCitation>
), which is characteristic of
<taxonomicName id="9CD54D33273FFFF32FE1C2A8FCAA0AB6" box="[669,779,223,248]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273FFFF32FE1C2A8FCAA0AB6" box="[669,779,223,248]" italics="true" pageId="2" pageNumber="216">Eucinetus</emphasis>
</taxonomicName>
(
<emphasis id="69A1EAA2273FFFF32E67C2A8FCE50AB6" box="[795,836,223,248]" italics="true" pageId="2" pageNumber="216">e.g.</emphasis>
,
<bibRefCitation id="3F444B41273FFFF32E2DC2A8FC630AB7" author="Vit, S." box="[849,962,223,249]" pageId="2" pageNumber="216" pagination="443 - 451" refId="ref7105" refString="Vit, S. (1977 a) Contribution a la connaissance du genre Eucinetus Germar (Coleoptera, Eucinetidae). Revue Suisse de Zoologie, 84, 443 - 451. https: // doi. org / 10.5962 / bhl. part. 91398" type="journal article" year="1977">Vit 1977a</bibRefCitation>
: figs 2, 4, 5;
<bibRefCitation id="3F444B41273FFFF3292DC2A8FB170AB7" author="Vit, S." box="[1105,1206,222,249]" pageId="2" pageNumber="216" pagination="421 - 460" refId="ref7279" refString="Vit, S. (1985) Etude de la morphologie des especes palearctiques du genre Eucinetus Germar et quelques remarques sur son utilisation taxonomique (Coleoptera Eucinetidae). Revue Suisse de Zoologie, 92, 421 - 460. https: // doi. org / 10.5962 / bhl. part. 81624" type="journal article" year="1985">Vit 1985</bibRefCitation>
: fig. 60;
<bibRefCitation id="3F444B41273FFFF32864C2A8FF720B53" author="Kim, T. - K. &amp; Ahn, K. - J." pageId="2" pageNumber="216" pagination="232 - 235" refId="ref6068" refString="Kim, T. - K. &amp; Ahn, K. - J. (2008) Little known Eucinetidae (Coleoptera: Scirtoidea) in Korea. Entomological Research, 38, 232 - 235. https: // doi. org / 10.1111 / j. 1748 - 5967.2008.00158. x" type="journal article" year="2008">Kim &amp; Ahn 2008</bibRefCitation>
: fig. 2). Therefore, these fossils can all be assigned to the extant
<taxonomicName id="9CD54D33273FFFF32EBFC374FB900B52" box="[963,1073,259,284]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273FFFF32EBFC374FB900B52" box="[963,1073,259,284]" italics="true" pageId="2" pageNumber="216">Eucinetus</emphasis>
</taxonomicName>
, demonstrating its considerable antiquity and remarkable stasis over the course of 100 million years of global biotic and abiotic change.
</paragraph>
<paragraph id="5B6A36B0273FFFF32DBBC33CFF510BBB" blockId="2.[151,1437,150,501]" pageId="2" pageNumber="216">
The four species of
<taxonomicName id="9CD54D33273FFFF32CD6C33CFDB90B2A" box="[426,536,331,356]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273FFFF32CD6C33CFDB90B2A" box="[426,536,331,356]" italics="true" pageId="2" pageNumber="216">Eucinetus</emphasis>
</taxonomicName>
from
<collectingRegion id="9911F852273FFFF32F1CC33CFD100B2B" box="[608,689,331,357]" country="Myanmar" name="Kachin" pageId="2" pageNumber="216">Kachin</collectingRegion>
amber (those described herein and the one described by
<bibRefCitation id="3F444B41273FFFF32845C33CFF700BC7" author="Du, X. &amp; Slipinski, A. &amp; Liu, Z. &amp; Pang, H." pageId="2" pageNumber="216" pagination="1 - 9" refId="ref5661" refString="Du, X., Slipinski, A., Liu, Z. &amp; Pang, H. (2020) Description of a new species of Eucinetidae (Coleoptera, Scirtoidea) from Cretaceous Burmese amber. ZooKeys, 982, 1 - 9. https: // doi. org / 10.3897 / zookeys. 982.39335" type="journal article" year="2020">
Du
<emphasis id="69A1EAA2273FFFF3281FC33CFA3C0B2B" box="[1379,1437,331,357]" italics="true" pageId="2" pageNumber="216">et al.</emphasis>
2020
</bibRefCitation>
) are known only from the
<typeStatus id="846E8812273FFFF32F7BC318FDD90BC7" box="[519,632,367,393]" pageId="2" pageNumber="216" type="holotype">holotypes</typeStatus>
. Two of the specimens are male, and the other two are female. To our knowledge, there is no notable sexual dimorphism in
<taxonomicName id="9CD54D33273FFFF32F9AC3E4FCF50BE2" box="[742,852,403,428]" class="Insecta" family="Eucinetidae" genus="Eucinetus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="69A1EAA2273FFFF32F9AC3E4FCF50BE2" box="[742,852,403,428]" italics="true" pageId="2" pageNumber="216">Eucinetus</emphasis>
</taxonomicName>
, except that in males one of the two mesotibial spurs is modified with a cleft apex (
<bibRefCitation id="3F444B41273FFFF32C9AC3C0FD3B0B9F" author="Lawrence, J. F." box="[486,666,439,465]" pageId="2" pageNumber="216" pagination="151 - 182" refId="ref6299" refString="Lawrence, J. F. (2019) New species of Eucinetus and Noteucinetus from Australia (Coleoptera: Scirtoidea: Eucinetidae). Zootaxa, 4668 (2), 151 - 182. https: // doi. org / 10.11646 / zootaxa. 4668.2.1" type="journal article" year="2019">Lawrence 2019</bibRefCitation>
). Thus, it would be justified to treat these
<specimenCount id="4DD3FD39273FFFF3290DC3C0FABA0B9F" box="[1137,1307,439,465]" count="4" pageId="2" pageNumber="216" type="generic">four specimens</specimenCount>
as separate species.
</paragraph>
</subSubSection>
</treatment>
</document>