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<document id="7732376F31498C2375DFDE23BFD7B903" ID-DOI="10.11646/zootaxa.5395.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10438327" ID-ZooBank="CCA38E8E-26E5-47D4-A27D-305554F6097F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703753817594" checkinUser="plazi" docAuthor="Feng, Zegang, Zhang, Feng &amp; Chen, Jun" docDate="2023" docId="0392327B7E6F6316FF37FE05FC274ABC" docLanguage="en" docName="zootaxa.5395.1.1.pdf" docOrigin="Zootaxa 5395 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5395.1.1/52550" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Bisetocreagris changbing Feng &amp; Zhang &amp; Chen 2023, sp. nov." docType="treatment" docUuid="CAC0EFFE-9041-47FD-BF51-5E4D4B3B8248" docUuidSource="ZooBank" docVersion="2" lastPageNumber="28" masterDocId="FFAB4A037E7A630DFFA0FA66FFBA4815" masterDocTitle="Cave-dwelling Neobisiidae in South China Karst, with descriptions of twelve new species of Bisetocreagris (Pseudoscorpiones, Neobisiidae) from Guizhou Province" masterLastPageNumber="77" masterPageNumber="1" pageNumber="22" updateTime="1703755787862" updateUser="ExternalLinkService">
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<mods:title id="A2F8D4CA9614F5F469A48BFE85B8C137">Cave-dwelling Neobisiidae in South China Karst, with descriptions of twelve new species of Bisetocreagris (Pseudoscorpiones, Neobisiidae) from Guizhou Province</mods:title>
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<mods:namePart id="AC9167D1BCDE8B2C98DA0152DB04BBE3">Feng, Zegang</mods:namePart>
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<mods:affiliation id="B3B30DAB3F31B76AB992836526C9BF81">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 &amp; College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532 &amp; Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author</mods:affiliation>
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<mods:namePart id="DF5E1D140753D1854FEB324FD4AF9C5B">Zhang, Feng</mods:namePart>
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<mods:affiliation id="1D4E3D88804594BF2062714D76F5033C">Key Laboratory of Zoological Systematics and Application, Hebei Basic Science Center for Biotic Interaction, College of Life sciences, Hebei University, Baoding, Hebei 071002, P. R. China dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031 * Corresponding author</mods:affiliation>
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<mods:namePart id="1740A6BA8BF968F070439CFF162342FC">Chen, Jun</mods:namePart>
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<mods:affiliation id="5A39174D5A73C47F0D93EE8B88CBF7AB">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, P. R. China https: // orcid. org / 0000 - 0002 - 5809 - 5984 &amp; College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, P. R. China chenj @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0325 - 2532</mods:affiliation>
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<taxonomicName id="4C3BF8EE7E6F6318FF37FE05FE764C68" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[151,460,1122,1149]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E6F6318FF37FE05FE764C68" bold="true" box="[151,460,1122,1149]" italics="true" pageId="21" pageNumber="22">Bisetocreagris changbing</emphasis>
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(Rae双ŁÞ伪AE)
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<figureCitation id="13009FE87E6F6318FF37FECDFEAE4CD0" box="[151,276,1195,1221]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStartId-0="22.[151,250,1788,1812]" captionStartId-1="23.[151,250,808,832]" captionStartId-2="23.[151,250,1633,1657]" captionStartId-3="24.[151,250,1943,1967]" captionStartId-4="25.[151,250,1874,1898]" captionTargetBox-0="[151,1435,309,1765]" captionTargetBox-1="[151,1435,181,784]" captionTargetBox-2="[151,1435,856,1609]" captionTargetBox-3="[164,1423,181,1919]" captionTargetBox-4="[217,1372,186,1846]" captionTargetId-0="figure-73@22.[151,1435,309,1765]" captionTargetId-1="figure-17@23.[151,1435,181,784]" captionTargetId-2="figure-36@23.[151,1435,856,1609]" captionTargetId-3="figure-21@24.[164,1423,181,1919]" captionTargetId-4="figure-17@25.[210,1377,181,1850]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionText-0="FIGURE 17. Guanyin and Sanyuan caves, type locality of Bisetocreagris changbing sp. nov., A. Surrounding surface environment with the entrance of Guanyin Cave, the temple is built in front of the entrance; B. Entrance of Guanyin Cave; C. Surrounding surface environment with the entrance of Sanyuan Cave; D. Entrance of Sanyuan Cave, locked by an iron door E. The artificial trail inside the Sanyuan Cave; E. Area where B. changbing sp. nov. specimens were collected." captionText-1="FIGURE 18. Live individuals of Bisetocreagris changbing sp. nov. (A: male; B: female) on the ground." captionText-2="FIGURE 19. Bisetocreagris changbing sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." captionText-3="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." captionText-4="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi-0="http://doi.org/10.5281/zenodo.10438363" figureDoi-1="http://doi.org/10.5281/zenodo.10438365" figureDoi-2="http://doi.org/10.5281/zenodo.10438367" figureDoi-3="http://doi.org/10.5281/zenodo.10438369" figureDoi-4="http://doi.org/10.5281/zenodo.10438371" httpUri-0="https://zenodo.org/record/10438363/files/figure.png" httpUri-1="https://zenodo.org/record/10438365/files/figure.png" httpUri-2="https://zenodo.org/record/10438367/files/figure.png" httpUri-3="https://zenodo.org/record/10438369/files/figure.png" httpUri-4="https://zenodo.org/record/10438371/files/figure.png" pageId="21" pageNumber="22">Figs 1721</figureCitation>
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<emphasis id="B94F5F7F7E6F6318FF37FE95FEF94D19" bold="true" box="[151,323,1266,1293]" pageId="21" pageNumber="22">Type material.</emphasis>
<materialsCitation id="3B5389307E6F6318FEE7FE95FB244D40" collectingDate="2020-08-20" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Fenggang County" elevation="755" location="Longquan Town" municipality="Zunyi City" pageId="21" pageNumber="22" specimenCode="MHBU-GZ20082001" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
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male (Ps.-
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):
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,
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,
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,
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,
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,
<location id="8EE4D5B67E6F6318FEFEFF71FD9D4D24" LSID="urn:lsid:plazi:treatment:0392327B7E6F6316FF37FE05FC274ABC:8EE4D5B67E6F6318FEFEFF71FD9D4D24" box="[350,551,1303,1329]" country="China" county="Fenggang County" municipality="Zunyi City" name="Yingchun Village" pageId="21" pageNumber="22" stateProvince="Guizhou">Yingchun Village</location>
,
<location id="8EE4D5B67E6F6318FD93FF71FD6C4D24" LSID="urn:lsid:plazi:treatment:0392327B7E6F6316FF37FE05FC274ABC:8EE4D5B67E6F6318FD93FF71FD6C4D24" box="[563,726,1303,1329]" country="China" county="Fenggang County" municipality="Zunyi City" name="Guanyin Cave" pageId="21" pageNumber="22" stateProvince="Guizhou">Guanyin Cave</location>
(
<emphasis id="B94F5F7F7E6F6318FD47FF71FB654D24" bold="true" box="[743,1247,1303,1329]" pageId="21" pageNumber="22">âṅåǎ义Ṅ凤冈县kȃDzȗǝťūḆñ</emphasis>
), [27°5625.72″, 107°4418.19″],
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a.s.l.
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,
<date id="FF85A5AD7E6F6318FE40FF5DFD2F4D40" box="[480,661,1339,1365]" pageId="21" pageNumber="22" value="2020-08-20">
<collectingDate id="EFC15C457E6F6318FE40FF5DFD2F4D40" box="[480,661,1339,1365]" pageId="21" pageNumber="22" value="2020-08-20">20 August 2020</collectingDate>
</date>
,
<collectorName id="26CEE6BB7E6F6318FD00FF5DFC8C4D40" box="[672,822,1339,1365]" pageId="21" pageNumber="22">Zegang Feng</collectorName>
,
<collectorName id="26CEE6BB7E6F6318FCE1FF5DFC784D40" box="[833,962,1339,1365]" pageId="21" pageNumber="22">Hongru Xu</collectorName>
,
<collectorName id="26CEE6BB7E6F6318FC6CFF5DFBD74D40" box="[972,1133,1339,1365]" pageId="21" pageNumber="22">Yanmeng Hou</collectorName>
leg.
</materialsCitation>
</paragraph>
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<materialsCitation id="3B5389307E6F6318FF67FF39FD734D6C" box="[199,713,1375,1401]" pageId="21" pageNumber="22" specimenCode="MHBU-GZ20082002" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54803DCF7E6F6318FF67FF39FE824D6C" box="[199,312,1375,1401]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
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(Ps.-
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)
</materialsCitation>
,
<materialsCitation id="3B5389307E6F6318FD74FF39FEC24D88" collectingDate="2020-08-20" collectionCode="MHBU-GZ" collectorName="Zegang Feng &amp; Hongru Xu &amp; Yanmeng Hou" country="China" county="Fenggang County" elevation="755" location="Longquan Town" municipality="Zunyi City" pageId="21" pageNumber="22" specimenCode="GZ20082003-05" specimenCount="3" specimenCount-female="3" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D3D48E47E6F6318FD74FF39FCFB4D6C" box="[724,833,1375,1401]" count="3" pageId="21" pageNumber="22" type="female">3 females</specimenCount>
(Ps.-
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<specimenCode id="DB9D2B167E6F6318FBA6FF38FB704D6C" box="[1030,1226,1374,1401]" collectionCode="GZ" pageId="21" pageNumber="22">GZ2008200305</specimenCode>
), all with the same data as the holotype
</materialsCitation>
.
<materialsCitation id="3B5389307E6F6318FE21FFE5FCDC4DF0" collectingDate="2016-10-15" collectorName="Mingyi Tian" country="China" county="Fenggang County" elevation="702" location="Longquan Town" municipality="Zunyi City" pageId="21" pageNumber="22" specimenCode="MHBU-GZ16101501" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="9D3D48E47E6F6318FE21FFE5FE5B4D88" box="[385,481,1411,1437]" count="1" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
(Ps.-
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):
<collectingCountry id="F32CC3FD7E6F6318FC85FFE5FCD14D88" box="[805,875,1411,1437]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
,
<collectingRegion id="49FF4D8F7E6F6318FCD5FFE5FBFB4D88" box="[885,1089,1411,1437]" country="China" name="Guizhou" pageId="21" pageNumber="22">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="6BE019177E6F6318FBECFFE5FB7C4D88" box="[1100,1222,1411,1437]" pageId="21" pageNumber="22">Zunyi City</collectingMunicipality>
,
<collectingCounty id="62E5FBE17E6F6318FB70FFE5FA234D88" box="[1232,1433,1411,1437]" pageId="21" pageNumber="22">Fenggang County</collectingCounty>
,
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,
<location id="8EE4D5B67E6F6318FEFCFFC1FD9E4DD4" LSID="urn:lsid:plazi:treatment:0392327B7E6F6316FF37FE05FC274ABC:8EE4D5B67E6F6318FEFCFFC1FD9E4DD4" box="[348,548,1447,1473]" country="China" county="Fenggang County" municipality="Zunyi City" name="Yingchun Village" pageId="21" pageNumber="22" stateProvince="Guizhou">Yingchun Village</location>
,
<location id="8EE4D5B67E6F6318FD8FFFC1FD6B4DD4" LSID="urn:lsid:plazi:treatment:0392327B7E6F6316FF37FE05FC274ABC:8EE4D5B67E6F6318FD8FFFC1FD6B4DD4" box="[559,721,1447,1473]" country="China" county="Fenggang County" municipality="Zunyi City" name="Sanyuan Cave" pageId="21" pageNumber="22" stateProvince="Guizhou">Sanyuan Cave</location>
(
<emphasis id="B94F5F7F7E6F6318FD41FFC1FB634DD4" bold="true" box="[737,1241,1447,1473]" pageId="21" pageNumber="22">âṅåǎ义Ṅ凤冈县kȃDzȗǝť三元ñ</emphasis>
), [27°5629.85″, 107°4421.55″],
<elevation id="0016645E7E6F6318FEF1FFADFE6E4DF0" box="[337,468,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="7.02" pageId="21" pageNumber="22" unit="m" value="702.0">
<quantity id="4CC32E887E6F6318FEF1FFADFE224DF0" box="[337,408,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="7.02" pageId="21" pageNumber="22" unit="m" value="702.0">702 m</quantity>
a.s.l.
</elevation>
,
<date id="FF85A5AD7E6F6318FE40FFACFD1A4DF0" box="[480,672,1482,1509]" pageId="21" pageNumber="22" value="2016-10-15">
<collectingDate id="EFC15C457E6F6318FE40FFACFD1A4DF0" box="[480,672,1482,1509]" pageId="21" pageNumber="22" value="2016-10-15">15 October 2016</collectingDate>
</date>
,
<collectorName id="26CEE6BB7E6F6318FD0BFFADFC8F4DF0" box="[683,821,1483,1509]" pageId="21" pageNumber="22">Mingyi Tian</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6F6318FF67FF89FBFF4E38" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8B84836D7E6F6318FF67FF89FBFF4E38" blockId="21.[151,1437,1266,2013]" pageId="21" pageNumber="22">
<emphasis id="B94F5F7F7E6F6318FF67FF89FEF44E1C" bold="true" box="[199,334,1519,1545]" pageId="21" pageNumber="22">Etymology.</emphasis>
The species name,
<taxonomicName id="4C3BF8EE7E6F6318FD9EFF89FD0E4E1C" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[574,692,1519,1545]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="changbing">
<emphasis id="B94F5F7F7E6F6318FD9EFF89FD0E4E1C" box="[574,692,1519,1545]" italics="true" pageId="21" pageNumber="22">changbing</emphasis>
</taxonomicName>
, was derived from the Latinized Mandarin phrase for “slender pedicel” cháng bǐng (
<emphasis id="B94F5F7F7E6F6318FE27FC75FE054E38" bold="true" box="[391,447,1555,1581]" pageId="21" pageNumber="22">ẗḣ</emphasis>
), which refers to the shape of the pedicel of the pedipalp.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6F6318FF67FC50FBF34F3C" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8B84836D7E6F6318FF67FC50FBF34F3C" blockId="21.[151,1437,1266,2013]" pageId="21" pageNumber="22">
<emphasis id="B94F5F7F7E6F6318FF67FC50FE494E45" bold="true" box="[199,499,1590,1617]" pageId="21" pageNumber="22">Distribution and habitat.</emphasis>
This new species is known from the
<typeStatus id="54803DCF7E6F6318FC3DFC51FC774E44" box="[925,973,1591,1617]" pageId="21" pageNumber="22">type</typeStatus>
locality, Guanyin (
<figureCitation id="13009FE87E6F6318FB06FC51FABE4E44" box="[1190,1284,1591,1617]" captionStart="FIGURE 1" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[156,1436,186,1039]" captionTargetId="figure-21@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Bisetocreagris species in China. A. B. maomaotou Gao, Wynne et Zhang, 2018 (Maomaotou Cave); B. B. que sp. nov. (Yanggong Cave); C. B. longidigitata sp. nov. (Maoer Cave); D. B. brevis sp. nov. (Yehu Cave); E. B. aspera sp. nov. (Tu Cave); F. B. jiangi sp. nov. (Anjialin Cave); G. B. tenuipalma sp. nov. (Sanjie Cave); H. B. changbing sp. nov. (Sanyuan and Guanyin caves); I. B. ampla sp. nov. (Hei Cave); J. B. gracilenta Gao et Zhang, 2017 (Shanmu Cave); K. B. guangshanensis Gao et Zhang, 2017 (Guangshan Cave); L. B. chumi sp. nov. (Tufei Cave); M. B. sanmen sp. nov. (Sanmen Cave); N. B. xishui sp. nov. (Dongkoushang Cave); O. B. baozinensis Mahnert et Li, 2016 (Baozi Cave); P. B. chuanensis Mahnert et Li, 2016 (Chuan Cave); Q. B. grandis sp. nov. (Wulibeidadong Cave); R. B. scaura (Mahnert, 2003) (Dahei Cave); S. B. martii (Mahnert, 2003) and B. titania (Mahnert, 2003) (Guoquan Cave); T. B. xiaoensis Li et Liu, 2017 (Xiao Cave); U. B. juanxuae Mahnert et Li, 2016 (Dao Cave); V. B. cavernarum Mahnert et Li, 2016 (Ert Long Cave); W. B. chinacavernicola (Schawaller, 1995) (Xian Nin and Chao-Tian caves)." figureDoi="http://doi.org/10.5281/zenodo.10438331" httpUri="https://zenodo.org/record/10438331/files/figure.png" pageId="21" pageNumber="22">Figs 1H</figureCitation>
,
<figureCitation id="13009FE87E6F6318FAB2FC51FADE4E44" box="[1298,1380,1591,1617]" captionStart="FIGURE 17" captionStartId="22.[151,250,1788,1812]" captionTargetBox="[151,1435,309,1765]" captionTargetId="figure-73@22.[151,1435,309,1765]" captionTargetPageId="22" captionText="FIGURE 17. Guanyin and Sanyuan caves, type locality of Bisetocreagris changbing sp. nov., A. Surrounding surface environment with the entrance of Guanyin Cave, the temple is built in front of the entrance; B. Entrance of Guanyin Cave; C. Surrounding surface environment with the entrance of Sanyuan Cave; D. Entrance of Sanyuan Cave, locked by an iron door E. The artificial trail inside the Sanyuan Cave; E. Area where B. changbing sp. nov. specimens were collected." figureDoi="http://doi.org/10.5281/zenodo.10438363" httpUri="https://zenodo.org/record/10438363/files/figure.png" pageId="21" pageNumber="22">17AB</figureCitation>
) and Sanyuan caves (
<figureCitation id="13009FE87E6F6318FEF1FC3DFE0A4E60" box="[337,432,1627,1653]" captionStart="FIGURE 1" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[156,1436,186,1039]" captionTargetId="figure-21@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Bisetocreagris species in China. A. B. maomaotou Gao, Wynne et Zhang, 2018 (Maomaotou Cave); B. B. que sp. nov. (Yanggong Cave); C. B. longidigitata sp. nov. (Maoer Cave); D. B. brevis sp. nov. (Yehu Cave); E. B. aspera sp. nov. (Tu Cave); F. B. jiangi sp. nov. (Anjialin Cave); G. B. tenuipalma sp. nov. (Sanjie Cave); H. B. changbing sp. nov. (Sanyuan and Guanyin caves); I. B. ampla sp. nov. (Hei Cave); J. B. gracilenta Gao et Zhang, 2017 (Shanmu Cave); K. B. guangshanensis Gao et Zhang, 2017 (Guangshan Cave); L. B. chumi sp. nov. (Tufei Cave); M. B. sanmen sp. nov. (Sanmen Cave); N. B. xishui sp. nov. (Dongkoushang Cave); O. B. baozinensis Mahnert et Li, 2016 (Baozi Cave); P. B. chuanensis Mahnert et Li, 2016 (Chuan Cave); Q. B. grandis sp. nov. (Wulibeidadong Cave); R. B. scaura (Mahnert, 2003) (Dahei Cave); S. B. martii (Mahnert, 2003) and B. titania (Mahnert, 2003) (Guoquan Cave); T. B. xiaoensis Li et Liu, 2017 (Xiao Cave); U. B. juanxuae Mahnert et Li, 2016 (Dao Cave); V. B. cavernarum Mahnert et Li, 2016 (Ert Long Cave); W. B. chinacavernicola (Schawaller, 1995) (Xian Nin and Chao-Tian caves)." figureDoi="http://doi.org/10.5281/zenodo.10438331" httpUri="https://zenodo.org/record/10438331/files/figure.png" pageId="21" pageNumber="22">Figs 1H</figureCitation>
,
<figureCitation id="13009FE87E6F6318FE1FFC3DFDAE4E60" box="[447,532,1627,1653]" captionStart="FIGURE 17" captionStartId="22.[151,250,1788,1812]" captionTargetBox="[151,1435,309,1765]" captionTargetId="figure-73@22.[151,1435,309,1765]" captionTargetPageId="22" captionText="FIGURE 17. Guanyin and Sanyuan caves, type locality of Bisetocreagris changbing sp. nov., A. Surrounding surface environment with the entrance of Guanyin Cave, the temple is built in front of the entrance; B. Entrance of Guanyin Cave; C. Surrounding surface environment with the entrance of Sanyuan Cave; D. Entrance of Sanyuan Cave, locked by an iron door E. The artificial trail inside the Sanyuan Cave; E. Area where B. changbing sp. nov. specimens were collected." figureDoi="http://doi.org/10.5281/zenodo.10438363" httpUri="https://zenodo.org/record/10438363/files/figure.png" pageId="21" pageNumber="22">17CD</figureCitation>
), located ~
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east of Longquan Town, Fenggang County. Guanyin Cave is located behind a temple on the hillside and has a a triangular entrance (about
<quantity id="4CC32E887E6F6318FBA1FC19FB964E8C" box="[1025,1068,1663,1689]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="m" value="4.0">4 m</quantity>
high and
<quantity id="4CC32E887E6F6318FB3DFC19FB724E8C" box="[1181,1224,1663,1689]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="m" value="2.0">2 m</quantity>
wide). There is no agricultural activity around the entrance. Sanyuan Cave is located next to a countryside road and has a rectangular entrance (
<quantity id="4CC32E887E6F6318FEA6FCA0FEF34EF4" box="[262,329,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="22" unit="m" value="2.5">2.5 m</quantity>
high and
<quantity id="4CC32E887E6F6318FE1BFCA0FE474EF4" box="[443,509,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="22" unit="m" value="1.5">1.5 m</quantity>
wide), surrounded by a wall, that is blocked by iron gates. It had been developed as a tourist area with cement pathways built from the entrance to an underground river, but has since then been abandoned. All specimens were collected under the rock (
<figureCitation id="13009FE87E6F6318FCBEFD69FCC44F3C" box="[798,894,1807,1833]" captionStart="FIGURE 17" captionStartId="22.[151,250,1788,1812]" captionTargetBox="[151,1435,309,1765]" captionTargetId="figure-73@22.[151,1435,309,1765]" captionTargetPageId="22" captionText="FIGURE 17. Guanyin and Sanyuan caves, type locality of Bisetocreagris changbing sp. nov., A. Surrounding surface environment with the entrance of Guanyin Cave, the temple is built in front of the entrance; B. Entrance of Guanyin Cave; C. Surrounding surface environment with the entrance of Sanyuan Cave; D. Entrance of Sanyuan Cave, locked by an iron door E. The artificial trail inside the Sanyuan Cave; E. Area where B. changbing sp. nov. specimens were collected." figureDoi="http://doi.org/10.5281/zenodo.10438363" httpUri="https://zenodo.org/record/10438363/files/figure.png" pageId="21" pageNumber="22">Fig. 17F</figureCitation>
) in the dark zone.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6F631BFF67FD54FB4A48C0" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B84836D7E6F631BFF67FD54FB4A48C0" blockId="21.[151,1437,1266,2013]" lastBlockId="22.[151,1437,151,285]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B94F5F7F7E6F6318FF67FD54FEF84F59" bold="true" box="[199,322,1842,1868]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
(
<specimenCount id="9D3D48E47E6F6318FEF3FD55FED64F58" box="[339,364,1843,1869]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
<specimenCount id="9D3D48E47E6F6318FECCFD55FEC74F58" box="[364,381,1843,1869]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
). Troglomorphic habitus; carapace with 2 faint eyespots on a flat surface; epistome small and triangular; carapace with 46 setae on posterior margin; pedipalp smooth and slender, both chelal fingers with 155175 teeth; femur 7.808.34 (
<specimenCount id="9D3D48E47E6F6318FDAFFD1DFD9C4F80" box="[527,550,1915,1941]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
), 7.677.97 (
<specimenCount id="9D3D48E47E6F6318FD1CFD1DFD744F80" box="[700,718,1915,1941]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
) times (length 2.732.92 (
<specimenCount id="9D3D48E47E6F6318FC59FD1DFBAA4F80" box="[1017,1040,1915,1941]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
), 2.852.99 (
<specimenCount id="9D3D48E47E6F6318FB06FD1DFB0D4F80" box="[1190,1207,1915,1941]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
)), patella 6.707.22 times (length 2.883.12) longer than wide, pedicel more than half the patella length. Hand with pedicel 3.784.04 (
<specimenCount id="9D3D48E47E6F6318FF01FDA5FF024FC8" box="[161,184,1987,2013]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
), 3.293.62 (
<specimenCount id="9D3D48E47E6F6318FEF1FDA5FED94FC8" box="[337,355,1987,2013]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
) times (length 1.932.02 (
<specimenCount id="9D3D48E47E6F6318FD33FDA5FD104FC8" box="[659,682,1987,2013]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
), 1.942.32 (
<specimenCount id="9D3D48E47E6F6318FCE2FDA5FCE94FC8" box="[834,851,1987,2013]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
)), chela with pedicel 8.518.56 (
<specimenCount id="9D3D48E47E6F6318FB6FFDA5FB5C4FC8" box="[1231,1254,1987,2013]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
), 7.327.68 (
<specimenCount id="9D3D48E47E6F6318FADFFDA5FA2B4FC8" box="[1407,1425,1987,2013]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
) times (length 4.284.34 (
<specimenCount id="9D3D48E47E6C631BFE18FAF1FE7548A4" box="[440,463,151,177]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
), 4.444.76 (
<specimenCount id="9D3D48E47E6C631BFDC9FAF1FDC048A4" box="[617,634,151,177]" count="1" pageId="22" pageNumber="23" type="female"></specimenCount>
)) longer than wide, finger 1.211.32 (
<specimenCount id="9D3D48E47E6C631BFB97FAF1FBF448A4" box="[1079,1102,151,177]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
), 1.141.22 (
<specimenCount id="9D3D48E47E6C631BFB49FAF1FB4148A4" box="[1257,1275,151,177]" count="1" pageId="22" pageNumber="23" type="female"></specimenCount>
) times longer than hand with pedicel. Chelicera: rallum with 89 pinnate setae, distal one with an expanded base.
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E6C6316FF67FAB8FD714954" lastPageId="27" lastPageNumber="28" pageId="22" pageNumber="23" type="description">
<paragraph id="8B84836D7E6C631BFF67FAB8FD374908" blockId="22.[151,1437,151,285]" pageId="22" pageNumber="23">
<emphasis id="B94F5F7F7E6C631BFF67FAB8FEE348ED" bold="true" box="[199,345,222,248]" pageId="22" pageNumber="23">Description.</emphasis>
Male (
<typeStatus id="54803DCF7E6C631BFE03FAB9FDB048EC" box="[419,522,223,249]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
;
<typeStatus id="54803DCF7E6C631BFDB3FAB9FDC948EC" box="[531,627,223,249]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
male in parentheses;
<figureCitation id="13009FE87E6C631BFCFDFAB9FC7E48EC" box="[861,964,223,249]" captionStart="FIGURE 18" captionStartId="23.[151,250,808,832]" captionTargetBox="[151,1435,181,784]" captionTargetId="figure-17@23.[151,1435,181,784]" captionTargetPageId="23" captionText="FIGURE 18. Live individuals of Bisetocreagris changbing sp. nov. (A: male; B: female) on the ground." figureDoi="http://doi.org/10.5281/zenodo.10438365" httpUri="https://zenodo.org/record/10438365/files/figure.png" pageId="22" pageNumber="23">Figs 18A</figureCitation>
,
<figureCitation id="13009FE87E6C631BFC6EFAB9FBBB48EC" box="[974,1025,223,249]" captionStart="FIGURE 19" captionStartId="23.[151,250,1633,1657]" captionTargetBox="[151,1435,856,1609]" captionTargetId="figure-36@23.[151,1435,856,1609]" captionTargetPageId="23" captionText="FIGURE 19. Bisetocreagris changbing sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10438367" httpUri="https://zenodo.org/record/10438367/files/figure.png" pageId="22" pageNumber="23">19A</figureCitation>
). Carapace, chelicerae, and pedipalps reddish brown; abdomen and legs yellowish.
</paragraph>
<caption id="DF44D3E57E6C631BFF37FC9AFB674F95" ID-DOI="http://doi.org/10.5281/zenodo.10438363" ID-Zenodo-Dep="10438363" httpUri="https://zenodo.org/record/10438363/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1788,1812]" targetBox="[151,1435,309,1765]" targetPageId="22" targetType="figure">
<paragraph id="8B84836D7E6C631BFF37FC9AFB674F95" blockId="22.[151,1437,1788,1920]" pageId="22" pageNumber="23">
<emphasis id="B94F5F7F7E6C631BFF37FC9AFE9C4F01" bold="true" box="[151,294,1788,1812]" pageId="22" pageNumber="23">FIGURE 17.</emphasis>
Guanyin and Sanyuan caves, type locality of
<taxonomicName id="4C3BF8EE7E6C631BFC85FC9BFB954F01" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[805,1071,1788,1812]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E6C631BFC85FC9BFB954F01" box="[805,1071,1788,1812]" italics="true" pageId="22" pageNumber="23">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6C631BFB9BFC9AFB284F01" bold="true" box="[1083,1170,1788,1812]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A27CE2047E6C631BFB9BFC9AFB284F01" box="[1083,1170,1788,1812]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Surrounding surface environment with the entrance of Guanyin Cave, the temple is built in front of the entrance; B. Entrance of Guanyin Cave; C. Surrounding surface environment with the entrance of Sanyuan Cave; D. Entrance of Sanyuan Cave, locked by an iron door E. The artificial trail inside the Sanyuan Cave; E. Area where
<taxonomicName id="4C3BF8EE7E6C631BFD4EFD0FFCCF4F95" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[750,885,1896,1920]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E6C631BFD4EFD0FFCCF4F95" box="[750,885,1896,1920]" italics="true" pageId="22" pageNumber="23">B. changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6C631BFCDBFD0EFC764F95" bold="true" box="[891,972,1896,1920]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A27CE2047E6C631BFCDBFD0EFC764F95" box="[891,972,1896,1920]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected.
</paragraph>
</caption>
<caption id="DF44D3E57E6D631AFF37F94EFB064B55" ID-DOI="http://doi.org/10.5281/zenodo.10438365" ID-Zenodo-Dep="10438365" box="[151,1212,808,832]" httpUri="https://zenodo.org/record/10438365/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,808,832]" targetBox="[151,1435,181,784]" targetPageId="23" targetType="figure">
<paragraph id="8B84836D7E6D631AFF37F94EFB064B55" blockId="23.[151,1212,808,832]" box="[151,1212,808,832]" pageId="23" pageNumber="24">
<emphasis id="B94F5F7F7E6D631AFF37F94EFE9A4B55" bold="true" box="[151,288,808,832]" pageId="23" pageNumber="24">FIGURE 18.</emphasis>
Live individuals of
<taxonomicName id="4C3BF8EE7E6D631AFE4DF94FFD4B4B55" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[493,753,808,832]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E6D631AFE4DF94FFD4B4B55" box="[493,753,808,832]" italics="true" pageId="23" pageNumber="24">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6D631AFD58F94EFCF24B55" bold="true" box="[760,840,808,832]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="A27CE2047E6D631AFD58F94EFCF24B55" box="[760,840,808,832]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A: male; B: female) on the ground.
</paragraph>
</caption>
<caption id="DF44D3E57E6D631AFF37FC07FAA04E6C" ID-DOI="http://doi.org/10.5281/zenodo.10438367" ID-Zenodo-Dep="10438367" box="[151,1306,1633,1658]" httpUri="https://zenodo.org/record/10438367/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1633,1657]" targetBox="[151,1435,856,1609]" targetPageId="23" targetType="figure">
<paragraph id="8B84836D7E6D631AFF37FC07FAA04E6C" blockId="23.[151,1306,1633,1658]" box="[151,1306,1633,1658]" pageId="23" pageNumber="24">
<emphasis id="B94F5F7F7E6D631AFF37FC07FE9A4E6C" bold="true" box="[151,288,1633,1657]" pageId="23" pageNumber="24">FIGURE 19.</emphasis>
<taxonomicName id="4C3BF8EE7E6D631AFE86FC04FD904E6C" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[294,554,1633,1657]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E6D631AFE86FC04FD904E6C" box="[294,554,1633,1657]" italics="true" pageId="23" pageNumber="24">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E6D631AFD90FC07FD3B4E6C" bold="true" box="[560,641,1633,1657]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="A27CE2047E6D631AFD90FC07FD3B4E6C" box="[560,641,1633,1657]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, dorsal view; B. Paratype female, dorsal view.
</paragraph>
</caption>
<paragraph id="8B84836D7E6D631AFF67FCAAFC354F38" blockId="23.[151,1437,1739,2018]" pageId="23" pageNumber="24">
Carapace (
<figureCitation id="13009FE87E6D631AFE9CFCADFE1C4EF3" box="[316,422,1739,1766]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="23" pageNumber="24">Figs 20A</figureCitation>
,
<figureCitation id="13009FE87E6D631AFE10FCAAFDBC4EF3" box="[432,518,1740,1766]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="23" pageNumber="24">21AB</figureCitation>
): smooth, 1.28 (1.31) times longer than wide, with a total of 24 (25) setae, including 4 (4) on anterior margin, 5 (6) on posterior margin, and 1 (1) on each side of anteroantero-lateral margin; with 2 faint eyespots on a flat surface (
<figureCitation id="13009FE87E6D631AFE62FD75FD924F3B" box="[450,552,1811,1838]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="23" pageNumber="24">Fig. 20B</figureCitation>
); epistome small and triangular.
</paragraph>
<paragraph id="8B84836D7E6D631AFF67FD51FC924FF4" blockId="23.[151,1437,1739,2018]" pageId="23" pageNumber="24">
Chelicera (
<figureCitation id="13009FE87E6D631AFEE4FD51FE084F47" box="[324,434,1847,1874]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="23" pageNumber="24">Figs 20B</figureCitation>
,
<figureCitation id="13009FE87E6D631AFE60FD5EFD8C4F47" box="[448,566,1848,1874]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="23" pageNumber="24">21BC, E</figureCitation>
): hand with 7 (67) setae, movable finger with 1 (1) submedial seta; fixed finger with 1516 (1315) teeth; movable finger with 1213 (1214) teeth; serrula exterior with 4246 lamellae; serrula interior with 2730 lamellae. Galea (
<figureCitation id="13009FE87E6D631AFD22FD19FD5E4F8F" box="[642,740,1919,1946]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="23" pageNumber="24">Fig. 21E</figureCitation>
) divided into 23 main branches, and each branch divided into multiple sub-branches. Rallum (
<figureCitation id="13009FE87E6D631AFDA3FDC5FDD34FAB" box="[515,617,1955,1982]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="23" pageNumber="24">Fig. 21C</figureCitation>
) with 8 pinnate blades, distal-most blade with an expanded base, distal-most 2 blades separated from others, proximal blade short.
</paragraph>
<caption id="DF44D3E57E626315FF37FDF1FC394FED" ID-DOI="http://doi.org/10.5281/zenodo.10438369" ID-Zenodo-Dep="10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1943,1967]" targetBox="[164,1423,181,1919]" targetPageId="24" targetType="figure">
<paragraph id="8B84836D7E626315FF37FDF1FC394FED" blockId="24.[151,1437,1943,2040]" pageId="24" pageNumber="25">
<emphasis id="B94F5F7F7E626315FF37FDF1FE9A4FBA" bold="true" box="[151,288,1943,1967]" pageId="24" pageNumber="25">FIGURE 20.</emphasis>
<taxonomicName id="4C3BF8EE7E626315FE85FDFEFD934FA5" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[293,553,1944,1968]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E626315FE85FDFEFD934FA5" box="[293,553,1944,1968]" italics="true" pageId="24" pageNumber="25">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E626315FD8FFDFEFDC54FA5" bold="true" box="[559,639,1944,1968]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A27CE2047E626315FD8FFDFEFDC54FA5" box="[559,639,1944,1968]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view.
</paragraph>
</caption>
<caption id="DF44D3E57E636314FF37FD34FDD64FEF" ID-DOI="http://doi.org/10.5281/zenodo.10438371" ID-Zenodo-Dep="10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1874,1898]" targetBox="[217,1372,186,1846]" targetPageId="25" targetType="figure">
<paragraph id="8B84836D7E636314FF37FD34FDD64FEF" blockId="25.[151,1436,1874,2042]" pageId="25" pageNumber="26">
<emphasis id="B94F5F7F7E636314FF37FD34FEA14F7F" bold="true" box="[151,283,1874,1898]" pageId="25" pageNumber="26">FIGURE 21</emphasis>
<taxonomicName id="4C3BF8EE7E636314FE83FD35FD934F7F" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[291,553,1874,1898]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E636314FE83FD35FD934F7F" box="[291,553,1874,1898]" italics="true" pageId="25" pageNumber="26">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E636314FD90FD34FD384F7F" bold="true" box="[560,642,1874,1898]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A27CE2047E636314FD90FD34FD384F7F" box="[560,642,1874,1898]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM).
</paragraph>
</caption>
<paragraph id="8B84836D7E606317FF67FAF1FEC04A28" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Pedipalps (
<figureCitation id="13009FE87E606317FEE4FAF1FE6A48A4" box="[324,464,151,177]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20DE</figureCitation>
,
<figureCitation id="13009FE87E606317FE7DFAF1FD9348A4" box="[477,553,151,177]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21IK</figureCitation>
): apex of coxae rounded, with 5 (5) setae on each side, pedipalpal coxae with 78 (7) setae. Pedipalp smooth and slender. Trochanter 3.39 (3.49) times longer than wide, femur 8.34 (7.80), patella 7.22 (7.02) times longer than wide, pedicel obviously more than half the entire length of patella, chela (with pedicel) 8.56 (8.51), chela (without pedicel) 7.58 (7.55) times longer than wide, movable finger 1.58 (1.75) times longer than hand (without pedicel). Fixed chelal finger with 8 trichobothria, movable finger with 4,
<emphasis id="B94F5F7F7E606317FBC5FB41FBC54954" box="[1125,1151,295,321]" italics="true" pageId="26" pageNumber="27">eb</emphasis>
and
<emphasis id="B94F5F7F7E606317FB14FB41FB634954" box="[1204,1241,295,321]" italics="true" pageId="26" pageNumber="27">esb</emphasis>
on lateral margin of hand;
<emphasis id="B94F5F7F7E606317FF5BFB2DFEAB4970" box="[251,273,331,357]" italics="true" pageId="26" pageNumber="27">ib</emphasis>
,
<emphasis id="B94F5F7F7E606317FEBFFB2DFE834971" box="[287,313,331,356]" italics="true" pageId="26" pageNumber="27">ist</emphasis>
and
<emphasis id="B94F5F7F7E606317FECFFB2DFE2A4970" box="[367,400,331,357]" italics="true" pageId="26" pageNumber="27">isb</emphasis>
closely grouped at the base of the fixed finger;
<emphasis id="B94F5F7F7E606317FC07FB2DFC7C4971" box="[935,966,331,356]" italics="true" pageId="26" pageNumber="27">est</emphasis>
slightly base of finger middle;
<emphasis id="B94F5F7F7E606317FA86FB2DFA8C4971" box="[1318,1334,331,356]" italics="true" pageId="26" pageNumber="27">it</emphasis>
closer to fingertip than
<emphasis id="B94F5F7F7E606317FE94FB09FEF2499D" box="[308,328,367,392]" italics="true" pageId="26" pageNumber="27">et</emphasis>
; on movable finger
<emphasis id="B94F5F7F7E606317FD87FB09FD80499D" box="[551,570,367,392]" italics="true" pageId="26" pageNumber="27">st</emphasis>
nearer to
<emphasis id="B94F5F7F7E606317FD08FB09FD0A499D" box="[680,688,367,392]" italics="true" pageId="26" pageNumber="27">t</emphasis>
than to
<emphasis id="B94F5F7F7E606317FCA8FB09FC9B499C" box="[776,801,367,393]" italics="true" pageId="26" pageNumber="27">sb</emphasis>
, the latter nearer
<emphasis id="B94F5F7F7E606317FC42FB09FC4A499C" box="[994,1008,367,393]" italics="true" pageId="26" pageNumber="27">b</emphasis>
than to
<emphasis id="B94F5F7F7E606317FBE8FB09FBE1499D" box="[1096,1115,367,392]" italics="true" pageId="26" pageNumber="27">st</emphasis>
, the distance between
<emphasis id="B94F5F7F7E606317FAF5FB09FAD4499C" box="[1365,1390,367,393]" italics="true" pageId="26" pageNumber="27">sb</emphasis>
and
<emphasis id="B94F5F7F7E606317FF37FBF5FF1F49B8" box="[151,165,403,429]" italics="true" pageId="26" pageNumber="27">b</emphasis>
is somewhat equal to that of
<emphasis id="B94F5F7F7E606317FE4EFBF5FDBD49B8" box="[494,519,403,429]" italics="true" pageId="26" pageNumber="27">sb</emphasis>
and
<emphasis id="B94F5F7F7E606317FD9DFBF5FDEA49B9" box="[573,592,403,428]" italics="true" pageId="26" pageNumber="27">st</emphasis>
(
<figureCitation id="13009FE87E606317FDFEFBF5FD7049B8" box="[606,714,403,429]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20D</figureCitation>
,
<figureCitation id="13009FE87E606317FD75FBF5FCA249B8" box="[725,792,403,429]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21IJ</figureCitation>
). Venom apparatus present only in the fixed chelal finger, venom duct short, not extending past half of the distance to
<emphasis id="B94F5F7F7E606317FC9AFBD1FCF449C5" box="[826,846,439,464]" italics="true" pageId="26" pageNumber="27">et</emphasis>
. Fixed chelal finger with 169 (160) teeth of unequal length, which are blunt and round in distal and middle part, flat and retrorse in basal part; movable finger with 171 (168) teeth, which are flat and retrorse in distal and middle part, flat, retrorse and the length is unequal in basal part; all teeth contiguous.
</paragraph>
<paragraph id="8B84836D7E606317FF67F821FEFB4AE4" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Abdomen:pleural membrane granulated.Tergal chaetotaxy(IXI):Tergal chaetotaxy(IXI):6:6:6:6:6:6:7:8:9:9:8 (6:6:6:6:7:7:7:8:8:8:7); sternal chaetotaxy (VXI):10:15:15:15:15:14:11:4 (10:14:14:14:15:12:11:4); stigmata with 46 (56) setae; anal cone with 2 dorsal and 2 ventral setae. Male genital area (
<figureCitation id="13009FE87E606317FBB8F8E9FBC74ABC" box="[1048,1149,655,681]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20F</figureCitation>
,
<figureCitation id="13009FE87E606317FB29F8E9FB044ABC" box="[1161,1214,655,681]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21G</figureCitation>
): sternite II with 38 (34) scattered setae; sternite III with anteromedian groove flanked by one small seta on each side, with 22 (17) posterior setae.
</paragraph>
<paragraph id="8B84836D7E606317FF67F89DFC544B48" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Legs: coxae chaetotaxy (IIV): 8:67:45:1012 (9:5:3:9). Leg I (
<figureCitation id="13009FE87E606317FC19F89DFB9F4B00" box="[953,1061,763,789]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20H</figureCitation>
,
<figureCitation id="13009FE87E606317FB93F89DFBDE4B00" box="[1075,1124,763,789]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21L</figureCitation>
): femur 7.23 (7.23), patella 5.10 (4.76), tibia 8.69 (9.79), basitarsus 5.31 (4.92), telotarsus 7.50 (7.67) times longer than wide, femur 1.56 (1.59) times longer than patella, telotarsus 1.30 (1.44) times longer than basitarsus.
</paragraph>
<paragraph id="8B84836D7E606317FF67F901FEDA4C04" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Leg IV(
<figureCitation id="13009FE87E606317FE80F901FE3A4B94" box="[288,384,871,897]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20I</figureCitation>
,
<figureCitation id="13009FE87E606317FE2BF901FE5C4B94" box="[395,486,871,897]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21D, M</figureCitation>
): femur + patella 8.26 (8.53) times longer than wide, femur shorter than patella, tibia 12.10 (12.05), basitarsus 5.56 (5.80), telotarsus 8.21 (8.00) times longer than wide, telotarsus 1.29 (1.29) times longer than basitarsus; basitarsus with one tactile setae (TS=0.11), telotarsus with one tactile setae (TS=0.420.50); subterminal tarsal seta (
<figureCitation id="13009FE87E606317FE05F9B5FDB64BF8" box="[421,524,979,1005]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">Fig. 21D</figureCitation>
) bifurcate, both branches dentated; arolium not divided, shorter than the slender and simple claws.
</paragraph>
<paragraph id="8B84836D7E606317FF67FE7DFBDE4C20" blockId="26.[151,1437,150,2013]" box="[199,1124,1051,1077]" pageId="26" pageNumber="27">
Females (
<typeStatus id="54803DCF7E606317FE93FE7DFE1E4C20" box="[307,420,1051,1077]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
;
<figureCitation id="13009FE87E606317FE0FFE7DFDAD4C20" box="[431,535,1051,1077]" captionStart="FIGURE 18" captionStartId="23.[151,250,808,832]" captionTargetBox="[151,1435,181,784]" captionTargetId="figure-17@23.[151,1435,181,784]" captionTargetPageId="23" captionText="FIGURE 18. Live individuals of Bisetocreagris changbing sp. nov. (A: male; B: female) on the ground." figureDoi="http://doi.org/10.5281/zenodo.10438365" httpUri="https://zenodo.org/record/10438365/files/figure.png" pageId="26" pageNumber="27">Figs 18B</figureCitation>
,
<figureCitation id="13009FE87E606317FD83FE7DFDEC4C20" box="[547,598,1051,1077]" captionStart="FIGURE 19" captionStartId="23.[151,250,1633,1657]" captionTargetBox="[151,1435,856,1609]" captionTargetId="figure-36@23.[151,1435,856,1609]" captionTargetPageId="23" captionText="FIGURE 19. Bisetocreagris changbing sp. nov., A. Holotype male, dorsal view; B. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10438367" httpUri="https://zenodo.org/record/10438367/files/figure.png" pageId="26" pageNumber="27">19B</figureCitation>
): mostly same as
<typeStatus id="54803DCF7E606317FCBCFE7DFCC44C20" box="[796,894,1051,1077]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
and
<typeStatus id="54803DCF7E606317FC14FE7DFBAE4C20" box="[948,1044,1051,1077]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
males.
</paragraph>
<paragraph id="8B84836D7E606317FF67FE59FEBB4CD0" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Chelicera. Hand with 67 setae, movable finger with 1 submedial seta; fixed finger with 1216 teeth; movable finger with 1317 teeth; serrula exterior with 4345 lamellae; serrula interior with 3132 lamellae. Galea divided into 2 main branches, and each one branch divided into multiple sub-branches; rallum of 89 blades, and similar to
<typeStatus id="54803DCF7E606317FF37FECDFF474CD0" box="[151,253,1195,1221]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B84836D7E606317FF67FEA9FCDA4D40" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">Pedipalps. Apex of coxae rounded, with 56 setae on each side. Pedipalpal coxae with 68 setae. Trochanter 3.123.36, femur 7.677.97, patella 6.707.19, chela (with pedicel) 7.327.68, chela (without pedicel) 6.426.83 times longer than wide, movable finger 1.501.60 times longer than hand (without pedicel). Fixed chelal finger with about 155170 teeth, movable finger with about 160175 teeth.</paragraph>
<paragraph id="8B84836D7E606317FF67FF39FC3C4DD4" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Abdomen. Tergal chaetotaxy (IXI): 6:6:56:6:6:67:6:69:8:79:67; sternal chaetotaxy (IVXI): 811:13 15:1215:1314:1214:1114:1114:34. Female genital area (
<figureCitation id="13009FE87E606317FCC3FFE5FC754D88" box="[867,975,1411,1437]" captionStart="FIGURE 20" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1919]" captionTargetId="figure-21@24.[164,1423,181,1919]" captionTargetPageId="24" captionText="FIGURE 20. Bisetocreagris changbing sp. nov., holotype male (AF, HI), female (G). A. Carapace, dorsal view; B. Eye area, lateral view; C. Right chelicera, dorsal view; D. Right chela, lateral view; E. Right pedipalp, dorsal view; F. Male genitalia; G. Female genitalia; H. Right leg I, lateral view; I. Right leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10438369" httpUri="https://zenodo.org/record/10438369/files/figure.png" pageId="26" pageNumber="27">Figs 20G</figureCitation>
,
<figureCitation id="13009FE87E606317FC7CFFE5FBAA4D88" box="[988,1040,1411,1437]" captionStart="FIGURE 21" captionStartId="25.[151,250,1874,1898]" captionTargetBox="[217,1372,186,1846]" captionTargetId="figure-17@25.[210,1377,181,1850]" captionTargetPageId="25" captionText="FIGURE 21 Bisetocreagris changbing sp. nov., holotype male (AG, IM), female (H). A. Carapace, dorsal view; B. Right chelicera, dorsal view; C. Rallum; D. Subterminal tarsal seta; E. Galea; F. Movable finger of chelicera, showing galea; G. Male genitalia, ventral view; H. Female genitalia, ventral view; I. Right chela, dorsal view; J. Right chelal fingers, retrolateral view; K. Right pedipalp, minus chela, dorsal view; L. Right leg I, lateral view; M. Right leg IV, lateral view. Scale bars: 0.1 mm (CF), 0.25 mm (B, GH), 0.5 mm (A), 1 mm (IM)." figureDoi="http://doi.org/10.5281/zenodo.10438371" httpUri="https://zenodo.org/record/10438371/files/figure.png" pageId="26" pageNumber="27">21H</figureCitation>
): sternite II with 49 setae on each side; sternite III with a row of 1215 setae on the posterior margin.
</paragraph>
<paragraph id="8B84836D7E606317FF67FFADFF514F58" blockId="26.[151,1437,150,2013]" pageId="26" pageNumber="27">
Measurements: (length/breadth or depth in mm; ratios for most characters in parentheses).
<typeStatus id="54803DCF7E606317FB52FFADFAE04DF0" box="[1266,1370,1483,1509]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
male (
<typeStatus id="54803DCF7E606317FF3FFF89FEBB4E1C" box="[159,257,1519,1545]" pageId="26" pageNumber="27" type="paratype">paratype</typeStatus>
male in square brackets). Body length 3.96 [3.76]. Carapace 1.36/1.06 (1.28) [1.35/1.03(1.31)]. Pedipalpal trochanter 1.29/0.38 (3.39) [1.29/0.37 (3.49)], femur 2.92/0.35 (8.34) [2.73/0.35 (7.80)], patella 2.96/0.41 (7.22) [2.88/0.41 (7.02)], chela (with pedicel) 4.28/0.50 (8.56) [4.34/0.51 (8.51)], chela (without pedicel) 3.79/0.50 (7.58) [3.85/0.51 (7.55)], hand length (without pedicel) 1.55 [1.46], movable finger length 2.45 (1.58 times longer than hand without pedicel) [2.55 (1.75 times longer than hand without pedicel)]. Leg I: trochanter 0.44/0.28 (1.57) [0.47/0.28 (1.68)], femur 1.59/0.22 (7.23) [1.59/0.22 (7.23)], patella 1.02/0.20 (5.10) [1.00/0.21 (4.76)], tibia 1.39/0.16 (8.69) [1.37/0.14 (9.79)], basitarsus 0.69/0.13 (5.31) [0.64/0.13 (4.92)], telotarsus 0.90/0.12 (7.50) [0.92/0.12 (7.67)]. Leg IV: trochanter 0.73/0.30 (2.43) [0.78/0.31 (2.52)], femur + patella 2.56/0.31 (8.26) [2.56/0.30 (8.53)], tibia 2.42/0.20 (12.10) [2.41/0.20 (12.05)], basitarsus 0.89/0.16 (5.56) [0.87/0.15 (5.80)], telotarsus 1.15/0.14 (8.21) [1.12/0.14 (8.00)].
</paragraph>
<paragraph id="8B84836D7E606316FF67FD31FD714954" blockId="26.[151,1437,150,2013]" lastBlockId="27.[151,1437,150,681]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Females (
<typeStatus id="54803DCF7E606317FE93FD31FE1F4F64" box="[307,421,1879,1905]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
). Body length 3.735.41. Carapace 1.371.52/1.111.19 (1.211.28). Pedipalpal trochanter 1.271.34/0.380.43 (3.123.36), femur 2.852.99/0.360.39 (7.677.97), patella 2.883.12/0.420.46 (6.707.19), chela (with pedicel) 4.444.76/0.590.65 (7.327.68), chela (without pedicel) 3.904.17/0.590.65 (6.426.83), hand length (without pedicel) 1.501.78, movable finger length 2.482.67 (1.501.60 times longer than hand without pedicel). Leg I: trochanter 0.430.45/0.270.30 (1.431.67), femur 1.581.64/0.220.25 (6.567.41), patella 0.96 1.01/0.200.22 (4.575.05), tibia 1.361.44/0.150.16 (8.509.4), basitarsus 0.640.66/0.120.14 (4.575.33), telotarsus 0.900.95/0.120.13 (7.157.92). Leg IV: trochanter 0.770.82/0.310.32 (2.412.65), femur + patella 2.562.73/0.300.31 (8.538.81), tibia 2.432.56/0.180.21 (12.1514.17), basitarsus 0.870.93/0.150.16 (5.75 6.20), telotarsus 1.131.20/0.140.16 (7.138.29).
</paragraph>
</subSubSection>
<subSubSection id="C321D0E67E616316FF67FB2DFC274ABC" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8B84836D7E616316FF67FB2DFBC449E0" blockId="27.[151,1437,150,681]" pageId="27" pageNumber="28">
<emphasis id="B94F5F7F7E616316FF67FB2DFE814970" bold="true" box="[199,315,331,357]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName id="4C3BF8EE7E616316FEE0FB2DFDE44970" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[320,606,331,357]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="changbing" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FEE0FB2DFDE44970" box="[320,606,331,357]" italics="true" pageId="27" pageNumber="28">Bisetocreagris changbing</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FDC4FB2DFD014970" bold="true" box="[612,699,331,357]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FDC4FB2DFD014970" box="[612,699,331,357]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C3BF8EE7E616316FCE7FB2DFB724970" authority="Mahnert et Li, 2016" authorityName="Mahnert et Li" authorityYear="2016" box="[839,1224,331,357]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="baozinensis">
<emphasis id="B94F5F7F7E616316FCE7FB2DFC524970" box="[839,1000,331,357]" italics="true" pageId="27" pageNumber="28">B. baozinensis</emphasis>
<bibRefCitation id="EFAAFE9C7E616316FC4DFB2DFB724970" author="Mahnert, V. &amp; Li, Y. C." box="[1005,1224,331,357]" pageId="27" pageNumber="28" pagination="259 - 268" refId="ref48128" refString="Mahnert, V. &amp; Li, Y. C. (2016) Cave-inhabiting Neobisiidae (Arachnida: Pseudoscorpiones) from China, with description of four new species of Bisetocreagris Curcic. Revue suisse de Zoologie, 123, 259 - 268. https: // doi. org / 10.5281 / zenodo. 155299" type="journal article" year="2016">
Mahnert et Li,
<quantity id="4CC32E887E616316FB30FB2DFB724970" box="[1168,1224,331,357]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="27" pageNumber="28" unit="in" value="2016.0">2016</quantity>
</bibRefCitation>
</taxonomicName>
in having a slender pedipalpal and a pedipalpal pedicel about half the entire length of pedipalpal patella, but it can be distinguished by the presence of 2 faint eyespots (
<emphasis id="B94F5F7F7E616316FDAFFBF5FD7049B8" box="[527,714,403,429]" italics="true" pageId="27" pageNumber="28">
vs.
<taxonomicName id="4C3BF8EE7E616316FD95FBF5FD7049B8" authorityName="Mahnert &amp; Li" authorityYear="2016" box="[565,714,403,429]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="baozinesis">B. baozinesis</taxonomicName>
</emphasis>
without eyes or eyespots); the pedicel distinctly more than half the patella length (
<emphasis id="B94F5F7F7E616316FECAFBD1FE3349C5" box="[362,393,439,464]" italics="true" pageId="27" pageNumber="28">vs.</emphasis>
pedicel about half the patella length in
<taxonomicName id="4C3BF8EE7E616316FCE9FBD1FC5149C4" authorityName="Mahnert et Li" authorityYear="2016" box="[841,1003,439,465]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="baozinensis">
<emphasis id="B94F5F7F7E616316FCE9FBD1FC5149C4" box="[841,1003,439,465]" italics="true" pageId="27" pageNumber="28">B. baozinensis</emphasis>
</taxonomicName>
); and the pedipalpal patella 7.027.22 (
<specimenCount id="9D3D48E47E616316FF01FBBDFF0249E0" box="[161,184,475,501]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
), 6.707.22 (
<specimenCount id="9D3D48E47E616316FEEFFBBDFEDB49E0" box="[335,353,475,501]" count="1" pageId="27" pageNumber="28" type="female"></specimenCount>
)
<emphasis id="B94F5F7F7E616316FED2FBBDFE2B49E1" box="[370,401,475,500]" italics="true" pageId="27" pageNumber="28">vs.</emphasis>
6.47.0 (
<specimenCount id="9D3D48E47E616316FE5CFBBDFDA949E0" box="[508,531,475,501]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
), 6.06.1 (
<specimenCount id="9D3D48E47E616316FD2EFBBDFD1A49E0" box="[654,672,475,501]" count="1" pageId="27" pageNumber="28" type="female"></specimenCount>
) times longer than wide in
<taxonomicName id="4C3BF8EE7E616316FC76FBBDFBC249E0" authorityName="Mahnert et Li" authorityYear="2016" box="[982,1144,475,501]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="baozinensis">
<emphasis id="B94F5F7F7E616316FC76FBBDFBC249E0" box="[982,1144,475,501]" italics="true" pageId="27" pageNumber="28">B. baozinensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84836D7E616316FF67FB99FC274ABC" blockId="27.[151,1437,150,681]" pageId="27" pageNumber="28">
This new species is similar, in pedipalpal proportions, to
<taxonomicName id="4C3BF8EE7E616316FC9DFB99FC634A0C" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[829,985,511,537]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tenuipalma" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FC9DFB99FC634A0C" box="[829,985,511,537]" italics="true" pageId="27" pageNumber="28">B. tenuipalma</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FC7FFB99FB824A0C" bold="true" box="[991,1080,511,537]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FC7FFB99FB824A0C" box="[991,1080,511,537]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3BF8EE7E616316FBCCFB99FB5A4A0D" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[1132,1248,511,536]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sanmen" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FBCCFB99FB5A4A0D" box="[1132,1248,511,536]" italics="true" pageId="27" pageNumber="28">B. sanmen</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FB46FB99FA854A0C" bold="true" box="[1254,1343,511,537]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FB46FB99FA854A0C" box="[1254,1343,511,537]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished by the presence of 2 faint eyespots on a flat surface of carapace (
<emphasis id="B94F5F7F7E616316FB84F845FBF84A29" box="[1060,1090,547,572]" italics="true" pageId="27" pageNumber="28">vs.</emphasis>
with 4 eyespots on the slightly raised surface in
<taxonomicName id="4C3BF8EE7E616316FEF7F821FE4F4A74" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[343,501,583,609]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tenuipalma" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FEF7F821FE4F4A74" box="[343,501,583,609]" italics="true" pageId="27" pageNumber="28">B. tenuipalma</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FE5CF821FDED4A74" bold="true" box="[508,599,583,609]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FE5CF821FDED4A74" box="[508,599,583,609]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); the pedicel distinctly more than half the patella length (
<emphasis id="B94F5F7F7E616316FB40F821FB444A75" box="[1248,1278,583,608]" italics="true" pageId="27" pageNumber="28">vs.</emphasis>
pedicel about half the patella length in
<taxonomicName id="4C3BF8EE7E616316FE11F80DFDF54A90" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[433,591,619,645]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tenuipalma" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FE11F80DFDF54A90" box="[433,591,619,645]" italics="true" pageId="27" pageNumber="28">B. tenuipalma</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FDF7F80DFD0B4A90" bold="true" box="[599,689,619,645]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FDF7F80DFD0B4A90" box="[599,689,619,645]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3BF8EE7E616316FD48F80DFCE44A91" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[744,862,619,644]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sanmen" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FD48F80DFCE44A91" box="[744,862,619,644]" italics="true" pageId="27" pageNumber="28">B. sanmen</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FCC5F80DFC054A90" bold="true" box="[869,959,619,645]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FCC5F80DFC054A90" box="[869,959,619,645]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
); and the smooth pedipalp (
<emphasis id="B94F5F7F7E616316FB5DF80DFAA64A91" box="[1277,1308,619,644]" italics="true" pageId="27" pageNumber="28">vs.</emphasis>
chela hand with fine granular in
<taxonomicName id="4C3BF8EE7E616316FE24F8E9FD9B4ABC" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[388,545,655,681]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tenuipalma" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FE24F8E9FD9B4ABC" box="[388,545,655,681]" italics="true" pageId="27" pageNumber="28">B. tenuipalma</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FD88F8E9FD3B4ABC" bold="true" box="[552,641,655,681]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FD88F8E9FD3B4ABC" box="[552,641,655,681]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3BF8EE7E616316FD18F8E9FC974ABD" authority="Feng &amp; Zhang &amp; Chen, 2023" authorityName="Feng &amp; Zhang &amp; Chen" authorityYear="2023" box="[696,813,655,680]" class="Arachnida" family="Neobisiidae" genus="Bisetocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sanmen" status="sp. nov.">
<emphasis id="B94F5F7F7E616316FD18F8E9FC974ABD" box="[696,813,655,680]" italics="true" pageId="27" pageNumber="28">B. sanmen</emphasis>
</taxonomicName>
<emphasis id="B94F5F7F7E616316FC93F8E9FC374ABC" bold="true" box="[819,909,655,681]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27CE2047E616316FC93F8E9FC374ABC" box="[819,909,655,681]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>