164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document id="C549B0670340F2FCF52D1C8765575021" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. & Liang, Hongbin" docDate="2016" docId="03D6879DFFC6FFDD3D1E6568FB587563" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Epaphiotrechus Deuve & Kavanaugh & Liang 2016, gen. nov." docType="treatment" docVersion="2" lastPageNumber="425" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="424" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="76B8FB2C7A1ABA47CBE09428C83FE0CB">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
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<mods:name id="F8396336ACA249EBDF1B41F0D2C1DFD4" type="personal">
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<mods:roleTerm id="0277CAD10D58EBA028026631B7883B95">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2F56285F18106C2C7FCF2266242B541C">Deuve, Thierry</mods:namePart>
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<mods:affiliation id="E8494E3F4CBDCE03ADC55402E3A80799">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National d’Histoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
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</mods:name>
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<mods:name id="116717C1380EBFDC59130722877AA9DF" type="personal">
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<mods:role id="FEBA491E520DE26FF2D4D5549B2C1A4E">
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<mods:roleTerm id="06261ECE706DE2BF8A85E6E3644C1F9B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="01C92B58D389867381ED1E690166DEC1">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="0DB60B06B24B58C70644C2AEAC4C493A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="94043828CB513BC440EBB2276A2207D8" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:roleTerm id="2743BCBF77F32F2A5A76022EF189D0C3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A98B3100D2E0F31A2B41AE97301EC8AA">Liang, Hongbin</mods:namePart>
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<mods:affiliation id="D8DCBE82E97C62EE2F75FCD0B92ED6A2">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:title id="3206A61405CD5B6E3613CE1CAC5C1213">Proceedings of the California Academy of Sciences</mods:title>
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</mods:titleInfo>
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<mods:part id="D63DBFB8FCA199A90B52DB5CFE1ED6C7">
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<mods:date id="2BE853C691D5D40E5F59E79349D27CFF">2016</mods:date>
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<mods:number id="460E74D0FC8669DBCCA8B7F5B19CFC38">2016-10-14</mods:number>
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<mods:detail id="8EBC3FAE7FC7033E549313560950DEF7" type="volume">
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<mods:number id="437FB6B9B9C6881997DF23A36487AC89">63</mods:number>
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<mods:detail id="17C6C579691F3A32F7F8FA0E57997BF5" type="issue">
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<mods:number id="6E2D2A56E20096917AD6D19AC8986901">12</mods:number>
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<mods:start id="E6792BC1E855C7234A771993BAFB8FA8">341</mods:start>
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<mods:classification id="187876C3762CA6E6AFE1F1D2B993AF1B">journal article</mods:classification>
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<mods:identifier id="D8587FCC3A66633B6811E9338DBD977E" type="DOI">10.5281/zenodo.13155283</mods:identifier>
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<mods:identifier id="4FFE6CB78D257FE5A88931ADD530F9E6" type="ISSN">0068-547X</mods:identifier>
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<treatment id="03D6879DFFC6FFDD3D1E6568FB587563" LSID="urn:lsid:plazi:treatment:03D6879DFFC6FFDD3D1E6568FB587563" httpUri="http://treatment.plazi.org/id/03D6879DFFC6FFDD3D1E6568FB587563" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424">
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<subSubSection id="C3656500FFC6FFDA3D1E6568FC097261" box="[329,967,1305,1329]" pageId="83" pageNumber="424" type="nomenclature">
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<paragraph id="8BC0368BFFC6FFDA3D1E6568FC097261" blockId="83.[329,967,1305,1329]" box="[329,967,1305,1329]" pageId="83" pageNumber="424">
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<heading id="D08881E7FFC6FFDA3D1E6568FC097261" box="[329,967,1305,1329]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="424" reason="2">
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<emphasis id="B90BEA99FFC6FFDA3D1E6568FC097261" bold="true" box="[329,967,1305,1329]" pageId="83" pageNumber="424">
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Genus
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<taxonomicName id="4C7F4D08FFC6FFDA3DCE6568FD587261" ID-CoL="8M7XY" authority="Deuve" authorityName="Deuve & Kavanaugh & Liang" authorityYear="2016" box="[409,662,1305,1329]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEA99FFC6FFDA3DCE6568FD867261" bold="true" box="[409,584,1305,1329]" italics="true" pageId="83" pageNumber="424">Epaphiotrechus</emphasis>
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Deuve
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</taxonomicName>
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and Kavanaugh,
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<taxonomicNameLabel id="A23857E2FFC6FFDA3F356568FC097261" box="[866,967,1305,1329]" pageId="83" pageNumber="424" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3656500FFC6FFDA3CFC6535FD2C72CC" pageId="83" pageNumber="424" type="type_taxon">
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||
<paragraph id="8BC0368BFFC6FFDA3CFC6535FD32720C" blockId="83.[123,1174,1348,1725]" box="[171,764,1348,1372]" pageId="83" pageNumber="424">
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||
<emphasis id="B90BEA99FFC6FFDA3CFC6535FEF1720B" bold="true" box="[171,319,1348,1372]" pageId="83" pageNumber="424">
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||
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<typeStatus id="54C48829FFC6FFDA3CFC6535FF2A720B" box="[171,228,1348,1372]" pageId="83" pageNumber="424">TYPE</typeStatus>
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</smallCapsWord>
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<smallCapsWord id="8D26A057FFC6FFDA3CBB6539FEF1720B" baselines="1366" box="[236,319,1352,1371]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="83" pageNumber="424">SPECIES</smallCapsWord>
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</emphasis>
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.—
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<taxonomicName id="4C7F4D08FFC6FFDA3D3F6535FD6F720C" ID-CoL="8M9KW" authorityName="Deuve & Kavanaugh & Liang" authorityYear="2016" box="[360,673,1348,1372]" class="Insecta" family="Carabidae" genus="Epaphiotrechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="species" species="fortipesoides" status="sp. nov.">
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<emphasis id="B90BEA99FFC6FFDA3D3F6535FD6F720C" box="[360,673,1348,1372]" italics="true" pageId="83" pageNumber="424">Epaphiotrechus fortipesoides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857E2FFC6FFDA3EFF6534FD32720C" box="[680,764,1349,1372]" pageId="83" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BC0368BFFC6FFDA3CFC6515FD2C72CC" blockId="83.[123,1174,1348,1725]" pageId="83" pageNumber="424">
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<emphasis id="B90BEA99FFC6FFDA3CFC6515FDD5722C" bold="true" box="[171,539,1380,1404]" pageId="83" pageNumber="424">Derivation of genus group name</emphasis>
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.— The genus group name (masculine) is a combination of two other trechine generic names,
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<taxonomicName id="4C7F4D08FFC6FFDA3DBA65F5FD9C72CC" ID-CoL="8KWTX" authorityName="Samouelle" authorityYear="1819" box="[493,594,1412,1436]" class="Insecta" family="Carabidae" genus="Epaphius" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEA99FFC6FFDA3DBA65F5FD9C72CC" box="[493,594,1412,1436]" italics="true" pageId="83" pageNumber="424">Epaphius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7F4D08FFC6FFDA3ED065F5FD1472CC" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[647,730,1412,1436]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEA99FFC6FFDA3ED065F5FD1472CC" box="[647,730,1412,1436]" italics="true" pageId="83" pageNumber="424">Trechus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656500FFC6FFDD3CFC65D4FDA87603" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424" type="diagnosis">
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<paragraph id="8BC0368BFFC6FFDD3CFC65D4FDA87603" blockId="83.[123,1174,1348,1725]" lastBlockId="84.[123,1174,219,563]" lastPageId="84" lastPageNumber="425" pageId="83" pageNumber="424">
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<smallCapsWord id="8D26A057FFC6FFDA3CFC65D4FEEE72EB" baselines="1463,1462" box="[171,288,1445,1469]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="83" pageNumber="424">
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<emphasis id="B90BEA99FFC6FFDA3CFC65D4FEEE72EB" bold="true" box="[171,288,1445,1469]" pageId="83" pageNumber="424">DIAGNOSIS</emphasis>
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||
</smallCapsWord>
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||
.— Adults of this genus (
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<figureCitation id="13442A0EFFC6FFDA3E7C65D5FD4D72ED" box="[555,643,1444,1469]" captionStart="FIGURE" captionStartId="87.[171,183,1591,1610]" captionTargetBox="[120,1176,216,1574]" captionTargetId="figure-15@87.[120,1176,216,1575]" captionTargetPageId="87" captionText="FIGURE 33. Epaphiotrechus fortipesoides sp. nov.; a. Dorsal habitus (CASENT1039089). b. Map of locality record (red circle) for E. fortipesoides in the Gaoligong Shan region. Scale lines a = 0.5 mm, b = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155346" httpUri="https://zenodo.org/record/13155346/files/figure.png" pageId="83" pageNumber="424">Fig. 33a</figureCitation>
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) can be recognized by the following combination of character states: size large (BL =
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<quantity id="4C879B6EFFC6FFDA3DA965B5FD59728D" box="[510,663,1476,1501]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="83" pageNumber="424" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5 to 4.7 mm</quantity>
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), apterous; body color dark piceous with elytral interval 1 and lateral areas of pronotum and elytra more or less reddish; dorsum shiny, distinctly iridescent, dorsal surface glabrous except for isolated fixed setae typical for trechines, eyes also glabrous; right mandible (
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<figureCitation id="13442A0EFFC6FFDA3DC76654FE25716D" box="[400,491,1573,1597]" captionStart="FIGURE" captionStartId="43.[171,183,1611,1630]" captionTargetBox="[153,1126,230,1591]" captionTargetId="figure-15@43.[134,1152,216,1601]" captionTargetPageId="43" captionText="FIGURE 16. Trechine right mandibles, ventral aspect. a. Agonotrechus xiaoheishan sp. nov. b. Trechus indicus Putzeys with dentition enlarged. c. Trechus gongshanensis sp. nov. d. Trechus shibalicus sp. nov. e. Pseudepaphius gonggaicus Deuve with dentition enlarged (China, Sichuan Province, Moxi Township, NE slope of Gongga Shan). f. Trechepaphiopsis uniporosa sp. nov. with dentition enlarged. g. Epaphiotrechus fortipesoides sp. nov. with dentition enlarged. h. Trechepaphiama gaoligong sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13155313" httpUri="https://zenodo.org/record/13155313/files/figure.png" pageId="83" pageNumber="424">
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Fig.
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<quantity id="4C879B6EFFC6FFDA3D956654FE25716D" box="[450,491,1573,1597]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="83" pageNumber="424" unit="g" value="16.0">16g</quantity>
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</figureCitation>
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) with premolar not fused with retinaculum but closely associat- ed with the latter [possibly representative of an intermediate state in the evolution of the “bidentate” mandibular
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<typeStatus id="54C48829FFC6FFDA3D176617FEA0712E" box="[320,366,1638,1662]" pageId="83" pageNumber="424">type</typeStatus>
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(see discussion above for genus
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<taxonomicName id="4C7F4D08FFC6FFDA3EA56614FC8B712D" authorityName="Clairville" authorityYear="1806" box="[754,837,1637,1661]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="424" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEA99FFC6FFDA3EA56614FC8B712D" box="[754,837,1637,1661]" italics="true" pageId="83" pageNumber="424">Trechus</emphasis>
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</taxonomicName>
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)], anterior tip of retinaculum enlarged as a distinct tooth and displaced anteriorly (but not quite as far as in members of
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<taxonomicName id="4C7F4D08FFC6FFDA386066F4FEF071ED" pageId="83" pageNumber="424">
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<emphasis id="B90BEA99FFC6FFDA386066F4FF2F71ED" italics="true" pageId="83" pageNumber="424">Trechepaphiopsis</emphasis>
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species
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</taxonomicName>
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); pronotum with basal angles small and rectangular; elytra elongate, oblong, slightly flattened along the median suture area, striae finely impressed, crenulate or finely punctate, lateral striae partly effaced but striae 6 and 7 still evident; two discal setae present on interval 3 next to stria 3, inserted near the anterior one-sixth and near mid-elytral length, respectively; preapical seta present, inserted next to stria 2.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656500FFC1FFDD3CFC612AFB587563" pageId="84" pageNumber="425" type="discussion">
|
||
<paragraph id="8BC0368BFFC1FFDD3CFC612AFD9176A3" blockId="84.[123,1174,219,563]" pageId="84" pageNumber="425">
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||
<emphasis id="B90BEA99FFC1FFDD3CFC612AFEE67622" bold="true" box="[171,296,347,371]" pageId="84" pageNumber="425">COMMENTS</emphasis>
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</smallCapsWord>
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.— When Uéno (1999) described
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<taxonomicName id="4C7F4D08FFC1FFDD3ECD612AFC547623" box="[666,922,347,371]" class="Insecta" family="Carabidae" genus="Trechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="species" species="fortipes">
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||
<emphasis id="B90BEA99FFC1FFDD3ECD612AFD237623" box="[666,749,347,371]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
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(
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<emphasis id="B90BEA99FFC1FFDD3EA8612DFCF77623" box="[767,825,348,371]" italics="true" pageId="84" pageNumber="425">s. str.</emphasis>
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||
)
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||
<emphasis id="B90BEA99FFC1FFDD3F1E612AFC547623" box="[841,922,347,371]" italics="true" pageId="84" pageNumber="425">fortipes</emphasis>
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</taxonomicName>
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, he noted that it was a “strange species similar to certain
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<taxonomicName id="4C7F4D08FFC1FFDD3DBA610AFDBF76C3" authorityName="Ueno" authorityYear="1953" box="[493,625,379,403]" class="Insecta" family="Carabidae" genus="Epaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90BEA99FFC1FFDD3DBA610AFDBF76C3" box="[493,625,379,403]" italics="true" pageId="84" pageNumber="425">Epaphiopsis</emphasis>
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</taxonomicName>
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”, with the distinct premolar on the right mandible. However, characters of the male aedeagus, particularly the presence of a distinct endophallic sclerite, led him to consider this species as a basal member of genus
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<taxonomicName id="4C7F4D08FFC1FFDD3F6E61CAFC427683" authorityName="Clairville" authorityYear="1806" box="[825,908,443,467]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="425" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90BEA99FFC1FFDD3F6E61CAFC427683" box="[825,908,443,467]" italics="true" pageId="84" pageNumber="425">Trechus</emphasis>
|
||
</taxonomicName>
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in which the plesiomorphic right mandibular dentition was retained.
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</paragraph>
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<paragraph id="8BC0368BFFC1FFDD3CFC618AFB587563" blockId="84.[123,1174,219,563]" pageId="84" pageNumber="425">
|
||
<emphasis id="B90BEA99FFC1FFDD3CFC618AFE347543" bold="true" box="[171,506,507,531]" pageId="84" pageNumber="425">
|
||
<smallCapsWord id="8D26A057FFC1FFDD3CFC618AFE987542" baselines="525,525" box="[171,342,507,531]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="84" pageNumber="425">GEOGRAPHICAL</smallCapsWord>
|
||
<smallCapsWord id="8D26A057FFC1FFDD3D37618EFE3A7542" baselines="525" box="[352,500,511,530]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="84" pageNumber="425">DISTRIBUTION</smallCapsWord>
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.
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</emphasis>
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This genus currently is known from only two species, both found only in the southern part of the Gaoligong Shan region of western
|
||
<collectingRegion id="49BBF869FFC1FFDD3FDD626DFB8A7563" box="[906,1092,539,563]" country="China" name="Yunnan" pageId="84" pageNumber="425">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F368761BFFC1FFDD3818626AFB5C7563" box="[1103,1170,539,563]" name="China" pageId="84" pageNumber="425">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |