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<document id="12CDAFECB97109079CC40DFCDFA39378" ID-DOI="10.5281/zenodo.13609085" ID-ISSN="2345-7600" ID-Zenodo-Dep="13609085" ID-ZooBank="http://zoobank.org/urn:lsid:zoobank.org:pub:4B0A7B82-1507-4576-988A-7A0B3710B307" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1723049599405" checkinUser="felipe" docAuthor="Stasolla, Gianluca &amp; Innocenti, Gianna" docDate="2014" docId="03CA87F0FFC6FFB5FF50D79BF005F72A" docLanguage="en" docName="RafflesBZool.62.591-599.pdf" docOrigin="Raffles Bulletin Of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Orcovita angulata Ng, Guinot &amp; Iliffe 1996" docType="treatment" docVersion="4" lastPageNumber="595" masterDocId="FFF3FF88FFC7FFB1FFBED01EF376FF94" masterDocTitle="A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae)" masterLastPageNumber="599" masterPageNumber="591" pageNumber="592" updateTime="1725019373397" updateUser="ExternalLinkService">
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<mods:title id="EC694E9287212CDBF834225A90AD7483">A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae)</mods:title>
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<mods:namePart id="9D8D2AB726D15B9E9A7436518FE761EC">Stasolla, Gianluca</mods:namePart>
<mods:affiliation id="504817D603ACC98325FB9070152DC0ED">Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Università di Firenze, via Romana 17, I- 50125 Firenze, Italy</mods:affiliation>
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<mods:namePart id="E21444A8CE99730402DC71CB66C2088B">Innocenti, Gianna</mods:namePart>
<mods:affiliation id="D0E82B28FD2B01705EA017C62BA7E49A">Museo di Storia Naturale, Sezione di Zoologia “ La Specola ”, Università di Firenze, via Romana 17, I- 50125 Firenze, Italy</mods:affiliation>
<mods:nameIdentifier id="B5BC45A7FA1086CE8D4323F1480D728B" type="email">gianna.innocenti@unifi.it</mods:nameIdentifier>
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<mods:date id="8D272735831C21BA300BAED80E15B0CC">2014</mods:date>
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<treatment id="03CA87F0FFC6FFB5FF50D79BF005F72A" ID-DOI="http://doi.org/10.5281/zenodo.13285390" ID-Zenodo-Dep="13285390" LSID="urn:lsid:plazi:treatment:03CA87F0FFC6FFB5FF50D79BF005F72A" httpUri="http://treatment.plazi.org/id/03CA87F0FFC6FFB5FF50D79BF005F72A" lastPageId="4" lastPageNumber="595" pageId="1" pageNumber="592">
<subSubSection id="C379656DFFC6FFB0FF50D79BF195F809" box="[238,739,1925,1949]" pageId="1" pageNumber="592" type="nomenclature">
<paragraph id="8BDC36E6FFC6FFB0FF50D79BF195F809" blockId="1.[169,789,1925,2042]" box="[238,739,1925,1949]" pageId="1" pageNumber="592">
<heading id="D094818AFFC6FFB0FF50D79BF195F809" bold="true" box="[238,739,1925,1949]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="592" reason="2">
<taxonomicName id="4C634D65FFC6FFB0FF50D79BF195F809" ID-CoL="754ZC" authority="Ng, Guinot &amp; Iliffe, 1996" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[238,739,1925,1949]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC6FFB0FF50D79BF195F809" bold="true" box="[238,739,1925,1949]" pageId="1" pageNumber="592">
<emphasis id="B917EAF4FFC6FFB0FF50D79BF2CEF809" bold="true" box="[238,440,1925,1949]" italics="true" pageId="1" pageNumber="592">Orcovita angulata</emphasis>
<bibRefCitation id="EFF24B17FFC6FFB0FE7ED79BF195F809" author="Ng PKL &amp; Guinot D &amp; Iliffe TM" box="[448,739,1925,1949]" pageId="1" pageNumber="592" pagination="109 - 134" refId="ref5431" refString="Ng PKL, Guinot D &amp; Iliffe TM (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1): 109 - 134." type="journal article" year="1996">Ng, Guinot &amp; Iliffe, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C379656DFFC6FFB0FE12D7BBF153F829" box="[428,549,1957,1981]" pageId="1" pageNumber="592" type="description">
<paragraph id="8BDC36E6FFC6FFB0FE12D7BBF153F829" blockId="1.[169,789,1925,2042]" box="[428,549,1957,1981]" pageId="1" pageNumber="592">
(
<figureCitation id="13582A63FFC6FFB0FE0AD7BBF177F829" box="[436,513,1957,1981]" captionStart="Fig" captionStartId="3.[868,903,2118,2140]" captionTargetBox="[869,1508,865,2093]" captionTargetId="figure-312@3.[868,1508,865,2093]" captionTargetPageId="3" captionText="Fig. 3. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268)." figureDoi="http://doi.org/10.5281/zenodo.13609100" httpUri="https://zenodo.org/record/13609100/files/figure.png" pageId="1" pageNumber="592">Figs. 3</figureCitation>
,
<figureCitation id="13582A63FFC6FFB0FDB1D7BBF16BF829" box="[527,541,1957,1981]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="1" pageNumber="592">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C379656DFFC6FFB0FF17D7FAF063F86E" box="[169,789,2020,2042]" pageId="1" pageNumber="592" type="reference_group">
<paragraph id="8BDC36E6FFC6FFB0FF17D7FAF063F86E" blockId="1.[169,789,1925,2042]" box="[169,789,2020,2042]" pageId="1" pageNumber="592">
<treatmentCitationGroup id="AB7311C8FFC6FFB0FF17D7FAF063F86E" box="[169,789,2020,2042]" pageId="1" pageNumber="592">
<treatmentCitation id="0AC210F7FFC6FFB0FF17D7FAF1E3F86D" author="Ng PKL &amp; Guinot D &amp; Iliffe TM" box="[169,661,2020,2042]" page="129" pageId="1" pageNumber="592" year="1996">
<taxonomicName id="4C634D65FFC6FFB0FF17D7FAF1E3F86D" ID-CoL="754ZC" authority="Ng, Guinot &amp; Iliffe, 1996: 129" authorityName="Ng, Guinot &amp; Iliffe" authorityPageNumber="129" authorityYear="1996" box="[169,661,2020,2042]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC6FFB0FF17D7FAF22DF86D" box="[169,347,2020,2041]" italics="true" pageId="1" pageNumber="592">Orcovita angulata</emphasis>
<bibRefCitation id="EFF24B17FFC6FFB0FEDDD7FAF1E3F86D" author="Ng PKL &amp; Guinot D &amp; Iliffe TM" box="[355,661,2020,2042]" pageId="1" pageNumber="592" pagination="109 - 134" refId="ref5431" refString="Ng PKL, Guinot D &amp; Iliffe TM (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1): 109 - 134." type="journal article" year="1996">Ng, Guinot &amp; Iliffe, 1996: 129</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 2b, 16.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C379656DFFC6FFB5FF17D803F005F72A" lastPageId="4" lastPageNumber="595" pageId="1" pageNumber="592" type="materials_examined">
<paragraph id="8BDC36E6FFC6FFB0FF17D803F05EF7DB" blockId="1.[169,808,2077,2128]" pageId="1" pageNumber="592">
<emphasis id="B917EAF4FFC6FFB0FF17D803F20CF7A7" bold="true" box="[169,378,2077,2099]" pageId="1" pageNumber="592">Material examined.</emphasis>
<materialsCitation id="3B0B3CBBFFC6FFB0FE3AD800F052F7DB" collectionCode="R" collectorName="Pukaway" country="Philippines" county="Berti" location="Coron Island" municipality="Pukaway Cave" pageId="1" pageNumber="592" specimenCode="MZUF 4268" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D65FD6FFFC6FFB0FE3AD800F2BFF7A7" box="[388,457,2077,2099]" count="1" pageId="1" pageNumber="592" type="male">1 male</specimenCount>
(16.2 ×
<quantity id="4C9B9B03FFC6FFB0FD99D800F1F4F7A7" box="[551,642,2078,2099]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="592" unit="mm" value="11.7">11.7 mm</quantity>
) (
<specimenCode id="DBC59E9DFFC6FFB0FD27D800F06BF7A7" box="[665,797,2077,2099]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="1" pageNumber="592">MZUF 4268</specimenCode>
),
<collectingCountry id="F3747676FFC6FFB0FF17D824F26AF7C4" box="[169,284,2106,2128]" name="Philippines" pageId="1" pageNumber="592">Philippines</collectingCountry>
,
<location id="8EBC603DFFC6FFB0FE94D824F2C7F7C4" LSID="urn:lsid:plazi:treatment:03CA87F0FFC6FFB5FF50D79BF005F72A:8EBC603DFFC6FFB0FE94D824F2C7F7C4" box="[298,433,2106,2128]" country="Philippines" county="Berti" municipality="Pukaway Cave" name="Coron Island" pageId="1" pageNumber="592">Coron Island</location>
,
<collectingMunicipality id="6BB8AC9CFFC6FFB0FE01D824F12EF7DB" box="[447,600,2106,2128]" pageId="1" pageNumber="592">
<collectorName id="26965330FFC6FFB0FE01D824F16CF7C4" box="[447,538,2106,2128]" pageId="1" pageNumber="592">Pukaway</collectorName>
Cave
</collectingMunicipality>
, coll.
<collectionCode id="ED72AE23FFC6FFB0FD22D825F1D9F7C4" box="[668,687,2107,2128]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="592">R</collectionCode>
.
<collectingCounty id="62BD4E6AFFC6FFB0FD02D824F198F7C4" box="[700,750,2106,2128]" pageId="1" pageNumber="592">Berti</collectingCounty>
&amp; S
</materialsCitation>
.
</paragraph>
<caption id="DF1C666EFFC6FFB0FCDDD85BF692F7CE" ID-DOI="http://doi.org/10.5281/zenodo.13609090" ID-Zenodo-Dep="13609090" box="[867,1508,2117,2139]" httpUri="https://zenodo.org/record/13609090/files/figure.png" pageId="1" pageNumber="592" startId="1.[867,902,2117,2139]" targetBox="[868,1508,888,2093]" targetPageId="1" targetType="figure">
<paragraph id="8BDC36E6FFC6FFB0FCDDD85BF692F7CE" blockId="1.[867,1508,2117,2139]" box="[867,1508,2117,2139]" pageId="1" pageNumber="592">
Fig. 1.
<taxonomicName id="4C634D65FFC6FFB0FC19D85BF723F7CE" authorityName="Ng &amp; Ng, 2009:" authorityYear="1098" box="[935,1109,2117,2138]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B917EAF4FFC6FFB0FC19D85BF723F7CE" box="[935,1109,2117,2138]" italics="true" pageId="1" pageNumber="592">Orcovita holthuisi</emphasis>
</taxonomicName>
, female (12.2 × 9.99 mm) (MZUF 3922).
</paragraph>
</caption>
<paragraph id="8BDC36E6FFC6FFB0FF17D849F2B6F7F8" blockId="1.[169,448,2135,2157]" box="[169,448,2135,2157]" pageId="1" pageNumber="592">
Meggiorini,
<date id="FFDD1026FFC6FFB0FE9DD849F2CAF7F8" box="[291,444,2135,2157]" pageId="1" pageNumber="592" value="1991-03-15">15 March 1991</date>
.
</paragraph>
<paragraph id="8BDC36E6FFC6FFB0FC8CD8B8F02CF72A" blockId="1.[818,858,2214,2238]" box="[818,858,2214,2238]" pageId="1" pageNumber="592">592</paragraph>
<paragraph id="8BDC36E6FFC5FFB3FDD0D0B2F73AFF50" blockId="2.[622,1100,172,196]" box="[622,1100,172,196]" pageId="2" pageNumber="593">
<emphasis id="B917EAF4FFC5FFB3FDD0D0B2F77BFF57" box="[622,1037,172,196]" italics="true" pageId="2" pageNumber="593">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2014
</paragraph>
<paragraph id="8BDC36E6FFC5FFB3FF7ED0E8F1E6FEF9" blockId="2.[192,831,245,366]" pageId="2" pageNumber="593">row of rounded granules present just below infraorbital margin on suborbital region, extending to sub-branchial region. Antennules broad. Antennal articles entering orbital hiatus. Eyes developed, cornea pigmented.</paragraph>
<paragraph id="8BDC36E6FFC5FFB3FF7ED18BF23BFDD9" blockId="2.[192,831,405,589]" pageId="2" pageNumber="593">
Epistome (
<figureCitation id="13582A63FFC5FFB3FE8CD18BF2F4FE39" box="[306,386,405,429]" captionStart="Fig" captionStartId="3.[868,903,2118,2140]" captionTargetBox="[869,1508,865,2093]" captionTargetId="figure-312@3.[868,1508,865,2093]" captionTargetPageId="3" captionText="Fig. 3. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268)." figureDoi="http://doi.org/10.5281/zenodo.13609100" httpUri="https://zenodo.org/record/13609100/files/figure.png" pageId="2" pageNumber="593">Fig. 3C</figureCitation>
) narrow, posterior margin with one lateral cleft, median part broadly triangular, margin smooth. Third maxilliped (
<figureCitation id="13582A63FFC5FFB3FEFFD1CBF2E7FE79" box="[321,401,469,493]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="2" pageNumber="593">Fig. 4F</figureCitation>
) with broad, stout exopod, as narrow as ischium of endopod, flagellum well developed; ischium stout; merus broad, antero-external angle edge gently granulate, auriculiform.
</paragraph>
<paragraph id="8BDC36E6FFC5FFB3FF7ED26BF6B5FED9" blockId="2.[192,831,629,749]" lastBlockId="2.[891,1530,245,333]" pageId="2" pageNumber="593">
Male chelae (
<figureCitation id="13582A63FFC5FFB3FEECD26BF2B2FD19" box="[338,452,629,653]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="2" pageNumber="593">Fig. 4D, E</figureCitation>
) subequal, fingers as long as palm; proximal region of cutting edges of fingers with very small pulvinus. Margins of merus serrated. Carpus rounded, serrated, inner distal angle with low, obtuse tooth. Chelipeds relatively long, surface of articles smooth, without any traces of a longitudinal sulcus; tips of fingers corneous, spoon-like, outer surface of tips of finger with dense, short setae.
</paragraph>
<paragraph id="8BDC36E6FFC5FFB3FCC5D16BF68FFD79" blockId="2.[891,1530,373,749]" pageId="2" pageNumber="593">
Ambulatory legs (
<figureCitation id="13582A63FFC5FFB3FB81D16BF7C0FE19" box="[1087,1206,373,397]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="2" pageNumber="593">Fig. 4G, H</figureCitation>
) flattened in cross section, all articles very long, slender; third ambulatory leg longest; surface smooth. Anterior dorsal margin of merus with acute, inconspicuous subdistal tooth; anterior, posterior margins of propodus with short and long setae, respectively; dactylus slender, with acute tip. Dactylus slightly rectangular in cross section, flexor margin comb-like in the first three pereiopods; 3M long, slender (length to width ratio 5.4), 3P long, slender (length to width ratio 5.1), 3D very long, slender (length to width ratio 12.4); 4M long, slender (length to width ratio 4.9), 4P long (length to width ratio 3.2), 4D long, slender (length to width ratio 7.3) (see Table 1). Male abdomen
</paragraph>
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<paragraph id="8BDC36E6FFC5FFB3FF7ED859F68CF7C9" blockId="2.[192,1530,2119,2141]" box="[192,1530,2119,2141]" pageId="2" pageNumber="593">
Fig. 2.
<taxonomicName id="4C634D65FFC5FFB3FEB6D856F2CFF7C8" authorityName="Ng &amp; Ng, 2009:" authorityYear="1098" box="[264,441,2119,2141]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="holthuisi">
<emphasis id="B917EAF4FFC5FFB3FEB6D856F2CFF7C8" box="[264,441,2119,2141]" italics="true" pageId="2" pageNumber="593">Orcovita holthuisi</emphasis>
</taxonomicName>
, female (10.9 × 9.38 mm) (MZUF 3922). A, carapace; B, dorsal view of the right cheliped; C, dorsal view of
</paragraph>
</caption>
<paragraph id="8BDC36E6FFC5FFB3FCF5D8B8F005F72A" blockId="2.[843,883,2214,2238]" box="[843,883,2214,2238]" pageId="2" pageNumber="593">593</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D105F3B7FE60" blockId="3.[169,222,283,2165]" box="[170,193,283,500]" pageId="3" pageNumber="594">width / length) dactylus</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D1EBF3B7FD96" blockId="3.[169,222,283,2165]" box="[170,193,501,514]" pageId="3" pageNumber="594">=</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D21CF3B7FD87" blockId="3.[169,222,283,2165]" box="[170,193,514,531]" pageId="3" pageNumber="594">D</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D204F3B7FD8A" blockId="3.[169,222,283,2165]" box="[170,193,538,542]" pageId="3" pageNumber="594">,</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D203F3B7FDED" blockId="3.[169,222,283,2165]" box="[170,193,541,633]" pageId="3" pageNumber="594">propodus</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D264F3B7FD13" blockId="3.[169,222,283,2165]" box="[170,193,634,647]" pageId="3" pageNumber="594">=</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D299F3B7FD00" blockId="3.[169,222,283,2165]" box="[170,193,647,660]" pageId="3" pageNumber="594">P</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D285F3B7FD0B" blockId="3.[169,222,283,2165]" box="[170,193,667,671]" pageId="3" pageNumber="594">,</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D281F3B7FD49" blockId="3.[169,222,283,2165]" box="[170,193,671,733]" pageId="3" pageNumber="594">merus</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D2C3F3B7FC94" blockId="3.[169,222,283,2165]" box="[170,193,733,768]" pageId="3" pageNumber="594">= M</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D31EF3B7FC93" blockId="3.[169,222,283,2165]" box="[170,193,768,775]" pageId="3" pageNumber="594">(</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D310F3B7FCA1" blockId="3.[169,222,283,2165]" box="[170,193,782,821]" pageId="3" pageNumber="594">legs</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF17D322F3C1FCD3" blockId="3.[169,222,283,2165]" box="[169,183,828,839]" pageId="3" pageNumber="594">th</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D359F3B7FCC7" blockId="3.[169,222,283,2165]" box="[170,193,839,851]" pageId="3" pageNumber="594">4</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D344F3B7FCE9" blockId="3.[169,222,283,2165]" box="[170,193,858,893]" pageId="3" pageNumber="594">and</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF17D39AF3C1FC04" blockId="3.[169,222,283,2165]" box="[169,183,900,912]" pageId="3" pageNumber="594">rd</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D38EF3B7FC7C" blockId="3.[169,222,283,2165]" box="[170,193,912,1000]" pageId="3" pageNumber="594">3 the For</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D3EEF3B7FC67" blockId="3.[169,222,283,2165]" box="[170,193,1008,1011]" pageId="3" pageNumber="594">.</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D3EDF3B7FAAC" blockId="3.[169,222,283,2165]" box="[170,193,1011,1336]" pageId="3" pageNumber="594">characters morphological various</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D521F3B7FAC7" blockId="3.[169,222,283,2165]" box="[170,193,1343,1363]" pageId="3" pageNumber="594">of</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D544F3B7FA52" blockId="3.[169,222,283,2165]" box="[170,193,1370,1478]" pageId="3" pageNumber="594">
<collectingCountry id="F3747676FFC4FFB2FF14D544F3B7FA52" box="[170,193,1370,1478]" name="Philippines" pageId="3" pageNumber="594">Philippines</collectingCountry>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D5D0F3B7FA7F" blockId="3.[169,222,283,2165]" box="[170,193,1486,1515]" pageId="3" pageNumber="594">the</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D5ECF3B7F9E8" blockId="3.[169,222,283,2165]" box="[170,193,1522,1660]" pageId="3" pageNumber="594">
from
<taxonomicName id="4C634D65FFC4FFB2FF14D636F3B7F9E8" authorityName="Ng &amp; Tomascik" authorityYear="1994" box="[170,193,1576,1660]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAF4FFC4FFB2FF14D636F3B7F9E8" box="[170,193,1576,1660]" italics="true" pageId="3" pageNumber="594">Orcovita</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D69AF3B7F90C" blockId="3.[169,222,283,2165]" box="[170,193,1668,1688]" pageId="3" pageNumber="594">of</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D681F3B7F8B8" blockId="3.[169,222,283,2165]" box="[170,193,1695,1836]" pageId="3" pageNumber="594">species known</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D72AF3B7F8C5" blockId="3.[169,222,283,2165]" box="[170,193,1844,1873]" pageId="3" pageNumber="594">the</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D746F3A8F789" blockId="3.[169,222,283,2165]" box="[170,222,1880,2080]" pageId="3" pageNumber="594">among Comparisons. indicated</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D839F3A8F7D6" blockId="3.[169,222,283,2165]" box="[170,222,2085,2114]" pageId="3" pageNumber="594">. 1 are</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FF14D85EF3A8F7E1" blockId="3.[169,222,283,2165]" box="[170,222,2112,2165]" pageId="3" pageNumber="594">Table ratio</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE75D0B2F7B4FF50" blockId="3.[459,1218,172,196]" box="[459,1218,172,196]" pageId="3" pageNumber="594">
Stasolla &amp; Innocenti: A new species of
<taxonomicName id="4C634D65FFC4FFB2FCC3D0B2F0ADFF57" authorityName="Ng &amp; Tomascik" authorityYear="1994" box="[893,987,172,195]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B917EAF4FFC4FFB2FCC3D0B2F0ADFF57" box="[893,987,172,195]" italics="true" pageId="3" pageNumber="594">Orcovita</emphasis>
</taxonomicName>
from the
<collectingCountry id="F3747676FFC4FFB2FBF4D0B2F7B4FF50" box="[1098,1218,172,196]" name="Philippines" pageId="3" pageNumber="594">Philippines</collectingCountry>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD0E7F26EFF6A" blockId="3.[257,792,249,2155]" box="[257,280,249,254]" pageId="3" pageNumber="594">.</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD0E0F26EFEA5" blockId="3.[257,792,249,2155]" box="[257,280,254,305]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD0E0F26EFEA5" bold="true" box="[257,280,254,305]" pageId="3" pageNumber="594">sp. n</emphasis>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD126F06EFE2C" blockId="3.[257,792,249,2155]" box="[257,792,304,440]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD126F26EFE30" bold="true" box="[257,280,312,440]" italics="true" pageId="3" pageNumber="594">
<taxonomicName id="4C634D65FFC4FFB2FEBFD126F26EFE0D" authorityName="Stasolla &amp; Innocenti" authorityYear="2014" box="[257,280,312,409]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="tabiacoud">tabiacoud</taxonomicName>
O.
</emphasis>
2 broad broad very no setae but long stout absent smooth 1.38 4.33 4.15 6.69 4.07 3.04 5.30
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE2BD1EAF2DAFD83" blockId="3.[257,792,249,2155]" box="[405,428,500,535]" pageId="3" pageNumber="594">tips</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE2BD200F2DAFDFB" blockId="3.[257,792,249,2155]" box="[405,428,542,623]" pageId="3" pageNumber="594">cheliped</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD264F26EFD21" blockId="3.[257,792,249,2155]" box="[257,280,634,693]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD264F26EFD21" bold="true" box="[257,280,634,693]" pageId="3" pageNumber="594">) male</emphasis>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE2BD268F2A6FD3F" blockId="3.[257,792,249,2155]" box="[405,464,614,683]" pageId="3" pageNumber="594">on slender</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD2ABF2A6FD42" blockId="3.[257,792,249,2155]" box="[257,464,660,727]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD2ABF26EFD28" bold="true" box="[257,280,693,700]" pageId="3" pageNumber="594">(</emphasis>
broad setae and
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD2DDF2A6FC9C" blockId="3.[257,792,249,2155]" box="[257,464,707,795]" pageId="3" pageNumber="594">
<taxonomicName id="4C634D65FFC4FFB2FEBFD2DDF26EFC8F" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[257,280,707,795]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC4FFB2FEBFD2DDF26EFC8F" bold="true" box="[257,280,707,795]" italics="true" pageId="3" pageNumber="594">angulata</emphasis>
</taxonomicName>
and short long
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD33CF26EFC52" blockId="3.[257,792,249,2155]" pageId="3" pageNumber="594">
.
<emphasis id="B917EAF4FFC4FFB2FEBFD338F26EFCAD" bold="true" box="[257,280,806,825]" italics="true" pageId="3" pageNumber="594">O</emphasis>
2 broad acute dense very absent serrated 1.30 5.44 5.14 12.48 4.94 3.22 7.33
<emphasis id="B917EAF4FFC4FFB2FEBFD367F26EFC52" bold="true" box="[257,280,889,966]" pageId="3" pageNumber="594">) female</emphasis>
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD3D8F283FC71" blockId="3.[257,792,249,2155]" box="[257,501,936,1005]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD3D8F26EFC58" bold="true" box="[257,280,966,972]" pageId="3" pageNumber="594">(</emphasis>
setae slender low
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD3CAF283FB84" blockId="3.[257,792,249,2155]" box="[257,501,980,1068]" pageId="3" pageNumber="594">
<taxonomicName id="4C634D65FFC4FFB2FEBFD3CAF26EFBB8" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[257,280,980,1068]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC4FFB2FEBFD3CAF26EFBB8" bold="true" box="[257,280,980,1068]" italics="true" pageId="3" pageNumber="594">angulata</emphasis>
</taxonomicName>
broad soft, and and
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD429F2DAFB4C" blockId="3.[257,792,249,2155]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD429F26EFBA3" bold="true" box="[257,280,1075,1098]" italics="true" pageId="3" pageNumber="594">O.</emphasis>
2 broad very short long weak serrated 1.3 5.4 5.4 16.1 4.6 3.7 6.6 pulvinus
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE2BD4C1F2A6FA94" blockId="3.[257,792,249,2155]" box="[405,464,1233,1280]" pageId="3" pageNumber="594">the stout</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE2BD51DF2A6FABE" blockId="3.[257,792,249,2155]" box="[405,464,1283,1322]" pageId="3" pageNumber="594">on and</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FECFD53DF2A6FAF4" blockId="3.[257,792,249,2155]" box="[369,464,1313,1376]" pageId="3" pageNumber="594">strong setae short</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD552F2A6F9D6" blockId="3.[257,792,249,2155]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD552F26EFA56" bold="true" box="[257,280,1356,1474]" italics="true" pageId="3" pageNumber="594">
<taxonomicName id="4C634D65FFC4FFB2FEBFD552F26EFA30" authorityName="Ng &amp; Ng, 2009:" authorityYear="1098" box="[257,280,1356,1444]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="holthuisi">holthuisi</taxonomicName>
. O
</emphasis>
3 narrow and acute dense long relatively present smooth 1.2 4.3 3.9 8.1 3.9 2.7 2.2 setae slender
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD65CF2A6F9F9" blockId="3.[257,792,249,2155]" box="[257,464,1597,1665]" pageId="3" pageNumber="594">
<taxonomicName id="4C634D65FFC4FFB2FEBFD65CF26EF915" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[257,280,1602,1665]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="fictilia">
<emphasis id="B917EAF4FFC4FFB2FEBFD65CF26EF915" bold="true" box="[257,280,1602,1665]" italics="true" pageId="3" pageNumber="594">fictilia</emphasis>
</taxonomicName>
soft, and
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FEBFD693F16FF88C" blockId="3.[257,792,249,2155]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FEBFD693F26EF918" bold="true" box="[257,280,1672,1696]" italics="true" pageId="3" pageNumber="594">O.</emphasis>
2 broad broad short long absent granular 1.5 1.3 5.3 4.2 9.1 4.2 4.4 2.6 3.0 6.9 6.2 carpus
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FDBCD701F14BF8E4" blockId="3.[257,792,249,2155]" box="[514,573,1823,1904]" pageId="3" pageNumber="594">cheliped ratio</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FDBCD7B3F14BF79A" blockId="3.[257,792,249,2155]" pageId="3" pageNumber="594">the male carapace teeth shape teeth chelipeds legs cristae of length</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FE96D7F2F06EF7CA" blockId="3.[257,792,249,2155]" box="[296,792,2028,2155]" pageId="3" pageNumber="594">Anterolateral Carapace Anterolateral Male Ambulatory Epigastric Inner margin to Width M P D M P D</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FCDBD0EBF7B9FD39" blockId="3.[869,1508,245,685]" pageId="3" pageNumber="594">
(
<figureCitation id="13582A63FFC4FFB2FCD3D0EBF0BDFE99" box="[877,971,245,269]" captionStart="Fig" captionStartId="3.[868,903,2118,2140]" captionTargetBox="[869,1508,865,2093]" captionTargetId="figure-312@3.[868,1508,865,2093]" captionTargetPageId="3" captionText="Fig. 3. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268)." figureDoi="http://doi.org/10.5281/zenodo.13609100" httpUri="https://zenodo.org/record/13609100/files/figure.png" pageId="3" pageNumber="594">Figs. 3B</figureCitation>
,
<figureCitation id="13582A63FFC4FFB2FC66D0EBF08EFE99" box="[984,1016,245,269]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="3" pageNumber="594">4B</figureCitation>
) narrowly triangular; lateral margins covered with short, rare setae; first abdominal somite, distal margin concave, swollen laterally lateral margin; second abdominal somite narrow, short, proximal margin concave; third abdominal somite broad; fourth abdominal somite broader; fifth abdominal somite with proximal and distal margins gently sinuously; sixth abdominal somite with proximal and distal margins slightly convex and concave medially respectively, lateral margins weakly convex; telson with lateral margins weakly concave, distal margin rounded. G1 (
<figureCitation id="13582A63FFC4FFB2FCD3D22BF0CCFDD9" box="[877,954,565,589]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="3" pageNumber="594">Fig. 4I</figureCitation>
) slender, gently curving outwards, reaching to or slightly beyond anterior margin of sternite 5, proximal part swollen; terminal lobe elongate; subterminal lobe densely setose. G2 (
<figureCitation id="13582A63FFC4FFB2FC58D28BF74FFD39" box="[998,1081,661,685]" captionStart="Fig" captionStartId="4.[192,227,2090,2112]" captionTargetBox="[251,1474,233,2053]" captionTargetId="figure-7@4.[242,1482,228,2064]" captionTargetPageId="4" captionText="Fig. 4. Orcovita angulata, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I," figureDoi="http://doi.org/10.5281/zenodo.13609104" httpUri="https://zenodo.org/record/13609104/files/figure.png" pageId="3" pageNumber="594">Fig. 4L</figureCitation>
) short, small.
</paragraph>
<paragraph id="8BDC36E6FFC4FFB2FCDBD2CBF76EFC99" blockId="3.[869,1508,725,781]" pageId="3" pageNumber="594">
<emphasis id="B917EAF4FFC4FFB2FCDBD2CBF0CFFD79" bold="true" box="[869,953,725,749]" pageId="3" pageNumber="594">Colour.</emphasis>
Unknown. The preserved specimens are uniformly yellowish white.
</paragraph>
<caption id="DF1C666EFFC4FFB2FCDAD858F692F7C8" ID-DOI="http://doi.org/10.5281/zenodo.13609100" ID-Zenodo-Dep="13609100" box="[868,1508,2118,2140]" httpUri="https://zenodo.org/record/13609100/files/figure.png" pageId="3" pageNumber="594" startId="3.[868,903,2118,2140]" targetBox="[869,1508,865,2093]" targetPageId="3" targetType="figure">
<paragraph id="8BDC36E6FFC4FFB2FCDAD858F692F7C8" blockId="3.[868,1508,2118,2140]" box="[868,1508,2118,2140]" pageId="3" pageNumber="594">
Fig. 3.
<taxonomicName id="4C634D65FFC4FFB2FC12D859F72AF7CF" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[940,1116,2118,2140]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC4FFB2FC12D859F72AF7CF" box="[940,1116,2118,2140]" italics="true" pageId="3" pageNumber="594">Orcovita angulata</emphasis>
</taxonomicName>
, male (16.2 × 11.7 mm) (MZUF 4268).
</paragraph>
</caption>
<paragraph id="8BDC36E6FFC4FFB2FC8CD8B8F02CF72A" blockId="3.[818,858,2214,2238]" box="[818,858,2214,2238]" pageId="3" pageNumber="594">594</paragraph>
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<emphasis id="B917EAF4FFC3FFB5FDD0D0B2F77BFF57" box="[622,1037,172,196]" italics="true" pageId="4" pageNumber="595">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2014
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Fig. 4.
<taxonomicName id="4C634D65FFC3FFB5FEB4D835F2CAF7AB" authorityName="Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[266,444,2090,2112]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="B917EAF4FFC3FFB5FEB4D835F2CAF7AB" box="[266,444,2090,2112]" italics="true" pageId="4" pageNumber="595">Orcovita angulata</emphasis>
</taxonomicName>
, male (16.2 × 11.7 mm) (MZUF 4268); A, carapace; B, abdomen; C, epibranchial region; D, dorsal view of right cheliped; E, dorsal view of carpus of cheliped; F, third maxilliped; G, right third ambulatory leg; H, right fourth ambulatory leg; I,
</paragraph>
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