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<taxonomicName id="4C614D2CFFA2FFFF7432A74BFD68FA3C" authority="? (Mantell, 1844)" baseAuthorityName="Mantell" baseAuthorityYear="1844" box="[264,643,1395,1422]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="701" phylum="Chordata" rank="species" species="belli">
<emphasis id="B915EABDFFA2FFFF7432A74BFE2CFA3F" bold="true" box="[264,455,1395,1422]" italics="true" pageId="20" pageNumber="701">Hylaeochelys belli</emphasis>
? (
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)
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(
<figureCitation id="135A2A2AFFA2FFFF74BCA7ABFE39FA1C" box="[390,466,1427,1453]" captionStart="FIG" captionStartId="18.[132,143,1940,1957]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-29@18.[698,1267,768,1324]" captionTargetPageId="18" captionText="FIG. 12. — Helochelydrid and thalassochelydian turtle remains from Angeac-Charente: A, peripheral plate of Helochelydridae indet. in dorsal view;B, pleural plate of Helochelydridae indet. in dorsal view; C, D, right hyoplastron of Hylaeochelys belli? (Mantell, 1844) (ANG18-5915) in ventral (C) and dorsal (D) views; E, left humerus of Hylaeochelys? belli? (ANG18-5937) in ventral view; F, right humerus (ANG18-5938) in dorsal view;; G, H, Peripheral of Hylaeochelys? belli? (ANG 18-5993) in dorsal (G) and ventral (H) views. Scale bar: A, 1 cm; E-H, 2 cm; B, 3 cm; C, D, 4 cm." figureDoi="http://doi.org/10.5281/zenodo.6902061" httpUri="https://zenodo.org/record/6902061/files/figure.png" pageId="20" pageNumber="701">Fig. 12</figureCitation>
C-H)
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<paragraph id="8BDE36AFFFA2FFFF75BEA7EBFEFEFA5C" blockId="20.[131,777,1491,2028]" box="[132,277,1491,1517]" pageId="20" pageNumber="701">DESCRIPTION</paragraph>
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The third turtle taxon is represented by many isolated plates and one incomplete specimen discovered in 2018, consisting of parts of the plastron, some peripherals and both humeri (
<figureCitation id="135A2A2AFFA2FFFF75B7A46AFF31F9DD" box="[141,218,1618,1644]" captionStart="FIG" captionStartId="18.[132,143,1940,1957]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-29@18.[698,1267,768,1324]" captionTargetPageId="18" captionText="FIG. 12. — Helochelydrid and thalassochelydian turtle remains from Angeac-Charente: A, peripheral plate of Helochelydridae indet. in dorsal view;B, pleural plate of Helochelydridae indet. in dorsal view; C, D, right hyoplastron of Hylaeochelys belli? (Mantell, 1844) (ANG18-5915) in ventral (C) and dorsal (D) views; E, left humerus of Hylaeochelys? belli? (ANG18-5937) in ventral view; F, right humerus (ANG18-5938) in dorsal view;; G, H, Peripheral of Hylaeochelys? belli? (ANG 18-5993) in dorsal (G) and ventral (H) views. Scale bar: A, 1 cm; E-H, 2 cm; B, 3 cm; C, D, 4 cm." figureDoi="http://doi.org/10.5281/zenodo.6902061" httpUri="https://zenodo.org/record/6902061/files/figure.png" pageId="20" pageNumber="701">Fig. 12</figureCitation>
C-H). The shell bone surface is nearly smooth with shallow grooves, which clearly distinguishes this taxon from the two others previously described. The shell bones are relatively thin. The vertebral scutes are very wide and cover more than half of the costals laterally. The neurals are narrow and elongated, especially in the mid-posterior part of the carapace. The plastron bears a central plastral fontanelle. The mesoplastron is absent. The humerus shape is unremarkable and similar to that recently described in Late Jurassic thalassochelydians (e.g., 
<bibRefCitation id="EFF04B5EFFA2FFFF745DA54AFDAEF83A" author="PUNTENER C. &amp; BILLON- BRUYAT J. - P. &amp; BOCAT L. &amp; BERGER J. - P. &amp; JOYCE W. G." box="[359,581,1905,1932]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="20" pageNumber="701" pagination="353 - 374" part="34" refId="ref47250" refString="PUNTENER C., BILLON- BRUYAT J. - P., BOCAT L., BERGER J. - P. &amp; JOYCE W. G. 2014. - Taxonomy and phylogeny of the turtle Tropidemys langii Rutimeyer, 1873 based on new specimens from the Kimmeridgian of the Swiss Jura Mountains. Journal of Vertebrate Paleontology 34 (2): 353 - 374. https: // doi. org / 10.1 080 / 02724634.2013.804412" title="Taxonomy and phylogeny of the turtle Tropidemys langii Rutimeyer, 1873 based on new specimens from the Kimmeridgian of the Swiss Jura Mountains" type="journal article" year="2014">
Püntener 
<emphasis id="B915EABDFFA2FFFF74EAA54AFDEEF83A" box="[464,517,1905,1931]" italics="true" pageId="20" pageNumber="701">et al.</emphasis>
2014
</bibRefCitation>
, 
<bibRefCitation id="EFF04B5EFFA2FFFF776EA549FD67F83A" author="ANQUETIN J. &amp; PUNTENER C. &amp; JOYCE W. G." box="[596,652,1905,1931]" journalOrPublisher="Bulletin of the Peabody Museum of Natural History" pageId="20" pageNumber="701" pagination="317 - 369" part="58" refId="ref35760" refString="ANQUETIN J., PUNTENER C. &amp; JOYCE W. G. 2017. - A review of the fossil record of turtles of the clade Thalassochelydia. Bulletin of the Peabody Museum of Natural History 58: 317 - 369. https: // doi. org / 10.3374 / 014.058.0205" title="A review of the fossil record of turtles of the clade Thalassochelydia" type="journal article" year="2017">2017</bibRefCitation>
). Based on the available characters, this form is possibly closely related to the Early Cretaceous (Purbeck and Wealden of the 
<collectingCountry id="F376763FFFA2FFFF7786A589FF02F85D" name="United Kingdom" pageId="20" pageNumber="701">United Kingdom</collectingCountry>
) thalassochelydian 
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<emphasis id="B915EABDFFA2FFFF7496A5E9FDBCF85A" box="[428,599,2001,2027]" italics="true" pageId="20" pageNumber="701">Hylaeochelys belli</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF04B5EFFA2FFFF775CA5E9FCEBF85A" author="Lydekker" box="[614,768,2001,2027]" firstAuthor="Lydekker" pageId="20" pageNumber="701" refId="ref42720" refString="LYDEKKER R. 1889. - Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part III, The Order Chelonia. Trustees of the British Museum, London, 239 p." type="book" year="1889">Lydekker 1889</bibRefCitation>
; 
<bibRefCitation id="EFF04B5EFFA2FFFF7617A6ECFBF1FB5F" author="Hirayama" box="[813,1050,1236,1262]" etAl="et al." firstAuthor="Hirayama" pageId="20" pageNumber="701" pagination="181 - 198" refId="ref41221" refString="HIRAYAMA R., BRINKMAN D. B. &amp; DANILOV I. G. 2000. - Distribution and biogeography of non-marine Cretaceous turtles. Russian Journal of Herpetology 7: 181 - 198." type="journal article" year="2000">
Hirayama 
<emphasis id="B915EABDFFA2FFFF7698A6ECFC33FB5F" box="[930,984,1236,1262]" italics="true" pageId="20" pageNumber="701">et al.</emphasis>
2000
</bibRefCitation>
; Pérez-García 2012; Pérez-García &amp; Ortega 2014; 
<bibRefCitation id="EFF04B5EFFA2FFFF76FFA6CBFB3AFABF" author="Anquetin &amp; Andre" box="[965,1233,1267,1294]" firstAuthor="Anquetin" pageId="20" pageNumber="701" pagination="5" refId="ref35695" refString="ANQUETIN J. &amp; ANDRE C. 2020. - The last surviving Thalassochelydia - A new turtle cranium from the Early Cretaceous of the Purbeck Group (Dorset, UK). PaleorXiv: 7 pa 5 c, version 3, peer-reviewed by PCI Paleo. https: // doi. org / 10.31233 / osf. io / 7 pa 5 c" type="journal article" year="2020">Anquetin &amp; André 2020</bibRefCitation>
).
</paragraph>
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<taxonomicName id="4C614D2CFFA2FFFF7679A72BFC1DFA9C" authorityName="? (Mantell" authorityYear="1844" box="[835,1014,1299,1325]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="20" pageNumber="701" phylum="Chordata" rank="species" species="belli">
<emphasis id="B915EABDFFA2FFFF7679A72BFC1DFA9C" box="[835,1014,1299,1325]" italics="true" pageId="20" pageNumber="701">Hylaeochelys belli</emphasis>
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is generally considered to be a freshwater turtle based on the sedimentary contexts of the localities in which it occurs (
<bibRefCitation id="EFF04B5EFFA2FFFF76D9A76BFB43FADC" author="Milner" box="[995,1192,1363,1389]" etAl="et al." firstAuthor="Milner" pageId="20" pageNumber="701" pagination="28 - 29" refId="ref44377" refString="MILNER A. R., EDMONDS R. P. &amp; GODDEN M. 2012. - The turtle genus Hylaeochelys from the Portland Stone Formation of Dorset. Symposium of Vertebrate Palaeontology and Comparative Anatomy: 28 - 29." type="proceedings paper" year="2012">
Milner 
<emphasis id="B915EABDFFA2FFFF710FA76CFB82FADC" box="[1077,1129,1363,1389]" italics="true" pageId="20" pageNumber="701">et al.</emphasis>
2012
</bibRefCitation>
; Pérez-García &amp; Ortega 2014; 
<bibRefCitation id="EFF04B5EFFA2FFFF7654A74BFB87FA3C" author="Anquetin &amp; Andre" box="[878,1132,1395,1421]" firstAuthor="Anquetin" pageId="20" pageNumber="701" pagination="5" refId="ref35695" refString="ANQUETIN J. &amp; ANDRE C. 2020. - The last surviving Thalassochelydia - A new turtle cranium from the Early Cretaceous of the Purbeck Group (Dorset, UK). PaleorXiv: 7 pa 5 c, version 3, peer-reviewed by PCI Paleo. https: // doi. org / 10.31233 / osf. io / 7 pa 5 c" type="journal article" year="2020">Anquetin &amp; André 2020</bibRefCitation>
). Most of the Angeac-Charente material represents juveniles or sub-adults, which may suggest that the swampy environment was a rookery for this turtle.
</paragraph>
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