286 lines
35 KiB
XML
286 lines
35 KiB
XML
<document id="A30EC588A67FA02E627BFEBD6956A053" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel & Solís, Angel" docDate="2001" docId="03A887FE5D4815382D03D0A1FC9DF9D6" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) guapiloides Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="28" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="26" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:mods id="95FB120637ED8D17FA283F3562FB8A60" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B933A77CE125AE3536C5119A2AE69192">
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<mods:title id="305F567F01DCB2B56B67827DCF99C845">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2D59EB587554A12C8FA6E81AD1F41822" type="personal">
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<mods:role id="D7C05A901127A432771F55C9F8B414F4">
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<mods:roleTerm id="4E834789EB5582318E56AE169658977E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D3067EEE2B33650A5B26876864B23BFE">Morón, Miguel Angel</mods:namePart>
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<mods:affiliation id="D494B4FA4217D655DA1DF1588691E455">Departamento de Entomología Instituto de Ecología, A. C. (sector SEPCONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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</mods:name>
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<mods:name id="E23315D869D7DD59D57F0B1A1550A9FD" type="personal">
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<mods:role id="802AFC9C307AFC087DC94596DD2E404D">
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<mods:roleTerm id="7DB57396A9B1608EDEE71E1B90BDA3B3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AEB7F70E97F3014F488F18CBC8D116D2">Solís, Angel</mods:namePart>
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<mods:affiliation id="DC7B51928057815D6353C4159BBDDAC4">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="57C24F732584ABB8776E98FC73BFC3E3">text</mods:typeOfResource>
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<mods:relatedItem id="D5B0751D57560BF75D679DEC11EEC8DC" type="host">
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<mods:titleInfo id="2861DA605E06AB14ABF8CC5B580CEDE6">
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<mods:title id="F32ECAD589053F8AF9E15B7F89127A6C">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="B1ECD1FA92C72FF975983CD22D523B81">
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<mods:date id="49AD67AE9C329FFFA7F54939AC7EB1E2">2001</mods:date>
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<mods:detail id="E0D9882B7B7D5F00F24BD7EDFD066E74" type="pubDate">
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<mods:number id="EB9720A1FB9BF8FF3B9CBA977D8A8C65">2001-03-01</mods:number>
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</mods:detail>
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<mods:detail id="30D9600B729059CF4BC7E42E2A562C79" type="volume">
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<mods:number id="0B9EA9FCC6426B3B86DF5289E6261117">55</mods:number>
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</mods:detail>
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<mods:detail id="531E1F5C90AFD6E4161D2CBF7AA8FF28" type="issue">
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<mods:number id="24006F5D958D9CFB0F096AB84A7A53DC">1</mods:number>
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</mods:detail>
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<mods:extent id="3BD8E93524C4811ED4CDC4906B9F53DF" unit="page">
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<mods:start id="B77B9546564566EBC2DC1AB3069A14A1">11</mods:start>
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<mods:end id="CBED72D0453498182B4D01CD63BE84AC">29</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="80FFD55E915F9632D6AB9FFA0551042E">journal article</mods:classification>
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<mods:identifier id="8BDA437877788A397E0292C78A9C333E" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
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<mods:identifier id="270A2655C19B9CCA2B7527F9DD001772" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="2CCF109CAB04819108B1CD3255AB7ED7" type="Zenodo-Dep">4900311</mods:identifier>
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</mods:mods>
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<treatment id="03A887FE5D4815382D03D0A1FC9DF9D6" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6" httpUri="http://treatment.plazi.org/id/03A887FE5D4815382D03D0A1FC9DF9D6" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26">
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<subSubSection id="C31B65635D4815262D03D0A1FBADFADA" box="[316,1075,1261,1283]" pageId="15" pageNumber="26" type="nomenclature">
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<paragraph id="8BBE36E85D4815262D03D0A1FBADFADA" blockId="15.[316,1075,1261,1309]" box="[316,1075,1261,1283]" pageId="15" pageNumber="26">
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<taxonomicName id="4C014D6B5D4815262D03D0A1FC3DFADB" authority="Moron and Solis" authorityName="Moron and Solis" box="[316,931,1262,1283]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="26" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov." subGenus="Phyllophaga">
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<emphasis id="B975EAFA5D4815262D03D0A1FE5CFADB" box="[316,450,1262,1283]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
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(
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<emphasis id="B975EAFA5D4815262DEBD0A1FDC7FADB" box="[468,601,1262,1283]" italics="true" pageId="15" pageNumber="26">Phyllophaga</emphasis>
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)
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<emphasis id="B975EAFA5D4815262E54D0A1FD7BFADB" box="[619,741,1262,1283]" italics="true" pageId="15" pageNumber="26">guapiloides</emphasis>
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Morón and Solís
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</taxonomicName>
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,
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<emphasis id="B975EAFA5D4815262F8FD0A2FBADFADA" bold="true" box="[944,1075,1261,1282]" pageId="15" pageNumber="26">
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<taxonomicNameLabel id="A24657815D4815262F8FD0A2FBADFADA" box="[944,1075,1261,1282]" pageId="15" pageNumber="26" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B65635D4815382E46D147FCDAFCA0" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26" type="description">
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<paragraph id="8BBE36E85D4815262E46D147FD69FAC5" blockId="15.[316,1075,1261,1309]" box="[633,759,1288,1309]" pageId="15" pageNumber="26">
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<figureCitation id="133A2A6D5D4815262E46D147FD69FAC5" box="[633,759,1288,1309]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="15" pageNumber="26">Figs. 28–31</figureCitation>
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</paragraph>
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<paragraph id="8BBE36E85D4815382D10D160FC2FFDE8" blockId="15.[279,1113,1327,1658]" lastBlockId="17.[279,1113,277,1550]" lastPageId="17" lastPageNumber="28" pageId="15" pageNumber="26">
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<emphasis id="B975EAFA5D4815262D10D160FE11FA9A" bold="true" box="[303,399,1327,1346]" pageId="15" pageNumber="26">
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<typeStatus id="54BA884A5D4815262D10D160FE15FA9A" box="[303,395,1327,1346]" pageId="15" pageNumber="26" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male. Clypeus, frons and pronotum shiny dark brown; elytra shiny dark brown, head and pronotum with erect, mixed long and short setae on entire surface; elytra with abundant decumbent short setae on entire surface and some scattered erect setae along the suture and toward the sides of scutellum; mouth parts, sterna, pygidium and legs shiny reddish brown. Clypeus 3.8× wider than long, anterior border slightly sinuate, with clearly elevated margin, surface slightly concave, densely rugopunctate, with many short erect setae. Frontoclypeal suture sinuate and partially hidden by punctures. Frons 1.7× wider than long, convex,densely rugopunctate, with mixed short and long setae on entire surface. Antenna 10segmented, with 3segmented club, lamellae of 8th to 10th segments 1.2× longer than length of preceeding 6 segments combined. Frons 5.0× wider than dorsal diameter of eye. Eye canthus long and wide, with 12–15 setae. Labrum bilobed, deeply notched, with abundant slender setae. Mentum slightly convex, with slender setae on each side, moderately sinuate anteriorly. Pronotum 1.8× wider than long and 2.2× wider than frons. Pronotal disk shiny, with deep, round punctures irregularly separated from one another by less than 1 diameter; lateral borders strongly angled, lateral marginal bead crenulate and progressively dentate toward posterior corner, with regularly located, long, slender setae; basal bead indicated by punctures, mainly toward the sides; anterior angles right, clearly prominent; posterior angles obtuse, clearly prominent. Scutellum 1.7× wider than long, with numerous shallow setiferous punctures. Elytron 2.4× longer than wider, shiny, densely setiferous punctate; epipleural border very narrow, extended along the complete margin, provided with dense fringe of short setae for entire length; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, with fine punctures densely placed and setiferous punctures. Pygidium moderately convex, shiny, with fine punctures densely placed and setiferous punctures, with scattered long erect setae on distal half of disk; apical margin with 16 long, slender setae; basal margin effaced medially. Pterosterna with long, dense, yellowish setae. Visible abdominal sternites II–IV convex, with scattered fine setiferous punctures; sternite V convex, finely rugose with setiferous punctures on the middle, with a shallow transverse sulcus before the posterior border; anal plate moderately narrowed, slightly concave, with scattered setiferous punctures, anterior and posterior border clearly elevated nearly all along the complete width, with 14 long setae near the posterior border. Protibia slightly shorter than protarsus (0.8:1), with external border tridentate, basal tooth short, preapical spur acute, straight, as long as 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and as long as lower spur. Metatibia 0.8× shorter than metatarsus, with one oblique, feeble, setiferous carina on external side; upper apical spur articulated, slightly curved, with apex acute, 0.9× shorter than basal metatarsomere, and 1.6× longer than lower spur; lower apical spur articulated with tibial border, slightly curved, with rounded apex. Tarsomeres semicylindrical, not much elongate, with enlarged apex and some setae apically; meso and metatarsomeres 2–4 ventrally with a serrulate longitudinal carina and one line of short setae at each side; protarsomeres 1–3 each with subapical small toothlike blade. Tarsal claws symmetrical, similar on all legs, each with a large acute tooth near the middle of ventral border with some minute incisions before the base (
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<figureCitation id="133A2A6D5D5615382DC5D53AFDDAFE50" box="[506,580,373,392]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Fig. 28</figureCitation>
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). Genital capsule with short and narrowed parameres, dorsally and ventrally fused, ringshaped, apex straight, progresively narrowed, with toothlike apex finely notched. Aedeagus long with preapical patches of strongly sclerotized short spines and one tuft of 7–8 very long setae on the dorsoapical area of sclerotized support, this support is a canaliculated wide structure with the base articulated with membranous parts of trema; ventral border of the apex slightly bisinuate (
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<figureCitation id="133A2A6D5D5615382FC0D5A2FEAFFDC0" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Figs. 29– 30</figureCitation>
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). Tectum convex. Length of genital capsule from apex of parameres to border of basal piece:
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<quantity id="4CF99B0D5D5615382D68D652FE3DFDE8" box="[343,419,541,560]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="17" pageNumber="28" unit="mm" value="4.7">4.7 mm</quantity>
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. Total body length:
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<quantity id="4CF99B0D5D5615382E5CD652FD25FDE8" box="[611,699,541,560]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="17" pageNumber="28" unit="mm" value="19.2">19.2 mm</quantity>
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. Humeral width:
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<quantity id="4CF99B0D5D5615382F5ED652FC33FDE8" box="[865,941,541,560]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="17" pageNumber="28" unit="mm" value="8.9">8.9 mm</quantity>
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.
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</paragraph>
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||
<caption id="DF7E66605D5715392D75D7C8FDB3FC12" ID-DOI="http://doi.org/10.5281/zenodo.4900325" ID-Zenodo-Dep="4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="16" pageNumber="27" startId="16.[330,373,903,922]" targetBox="[331,1116,273,880]" targetPageId="16">
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<paragraph id="8BBE36E85D5715392D75D7C8FDB3FC12" blockId="16.[307,1140,903,970]" pageId="16" pageNumber="27">
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<emphasis id="B975EAFA5D5715392D75D7C8FE21FC42" bold="true" box="[330,447,903,922]" pageId="16" pageNumber="27">Figs. 28–31.</emphasis>
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<emphasis id="B975EAFA5D5715392DE8D7C8FD59FC42" box="[471,711,903,922]" italics="true" pageId="16" pageNumber="27">
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<taxonomicName id="4C014D6B5D5715392DE8D7C8FD5DFC42" box="[471,707,903,922]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="27" phylum="Arthropoda" rank="species" species="guapiloides">Phyllophaga guapiloides</taxonomicName>
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.
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</emphasis>
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<emphasis id="B975EAFA5D5715392EF2D7C8FD7AFC42" bold="true" box="[717,740,903,922]" pageId="16" pageNumber="27">28</emphasis>
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) Protarsal claw, male;
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<emphasis id="B975EAFA5D5715392F82D7C8FC4AFC42" bold="true" box="[957,980,903,922]" pageId="16" pageNumber="27">29</emphasis>
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) genital capsule, lateral view;
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<emphasis id="B975EAFA5D5715392D8FD7D0FE59FC6A" bold="true" box="[432,455,927,946]" pageId="16" pageNumber="27">30</emphasis>
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) paramera, distal view;
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<emphasis id="B975EAFA5D5715392E8ED7D0FD56FC6A" bold="true" box="[689,712,927,946]" pageId="16" pageNumber="27">31</emphasis>
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) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="8BBE36E85D5615382D0DD672FBACFCD5" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
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<emphasis id="B975EAFA5D5615382D0DD672FE09FD8A" bold="true" box="[306,407,573,594]" pageId="17" pageNumber="28">
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<typeStatus id="54BA884A5D5615382D0DD672FE0CFD8A" box="[306,402,573,594]" pageId="17" pageNumber="28" type="allotype">Allotype</typeStatus>
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||
.
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</emphasis>
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Female. Similar to the male except as follows: antennal club 0.8× shorter than the preceeding 6 segments; visible abdominal sternites II to V convex with more setae; anal plate convex. Metafemur 1.4× wider than femur of male. Metatibia at apex 1.4× wider than male metatibia. Both apical spurs of metatibia wider than in male. Pygidium less convex. Ventral genital plates strongly sclerotized, nearly symmetrical, round with wide, slightly elongated, setiferous apex; dorsal genital plates sclerotized, shortened, with wide setiferous apex (
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<figureCitation id="133A2A6D5D5615382DB8D6B7FE49FCD5" box="[391,471,760,781]" captionStart="Figs" captionStartId="16.[330,373,903,922]" captionTargetBox="[331,1116,273,880]" captionTargetId="figure-12@16.[325,1120,272,883]" captionTargetPageId="16" captionText="Figs. 28–31. Phyllophaga guapiloides. 28) Protarsal claw, male; 29) genital capsule, lateral view; 30) paramera, distal view; 31) genital plates, female. Scale lines 5 1 mm, except Fig. 28 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900325" httpUri="https://zenodo.org/record/4900325/files/figure.png" pageId="17" pageNumber="28">Fig. 31</figureCitation>
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). Total body length:
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<quantity id="4CF99B0D5D5615382E86D6B7FC82FCD5" box="[697,796,760,781]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="17" pageNumber="28" unit="mm" value="21.9">21.9 mm</quantity>
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. Humeral width:
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<quantity id="4CF99B0D5D5615382FE7D6B7FBB0FCD5" box="[984,1070,760,781]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="17" pageNumber="28" unit="mm" value="9.2">9.2 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BBE36E85D5615382D0DD75DFCDAFCA0" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
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<emphasis id="B975EAFA5D5615382D0DD75DFD8FFCFF" bold="true" box="[306,529,786,807]" pageId="17" pageNumber="28">
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||
<typeStatus id="54BA884A5D5615382D0DD75DFE06FCFF" box="[306,408,786,807]" pageId="17" pageNumber="28" type="paratype">Paratype</typeStatus>
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Variation.
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</emphasis>
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Males. Similar to
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<typeStatus id="54BA884A5D5615382EEAD75CFCAEFCF0" box="[725,816,787,808]" pageId="17" pageNumber="28" type="holotype">holotype</typeStatus>
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||
except in total body length: 18.0–
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||
<quantity id="4CF99B0D5D5615382D6DD762FE28FC9A" box="[338,438,813,834]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="17" pageNumber="28" unit="mm" value="21.2">21.2 mm</quantity>
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, humeral width: 8.0–
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<quantity id="4CF99B0D5D5615382E9FD762FD69FC9A" box="[672,759,813,834]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="28" unit="mm" value="9.5">9.5 mm</quantity>
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; some specimens slightly darker or lighter than
|
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<typeStatus id="54BA884A5D5615382DFBD707FDBDFC85" box="[452,547,840,861]" pageId="17" pageNumber="28" type="holotype">holotype</typeStatus>
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. Females. Similar to
|
||
<typeStatus id="54BA884A5D5615382F28D707FCF5FC85" box="[791,875,840,861]" pageId="17" pageNumber="28" type="allotype">allotype</typeStatus>
|
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except in total body length: 18.2–21.0 mm, humeral width:
|
||
<quantity id="4CF99B0D5D5615382E82D72CFCDEFCA0" box="[701,832,867,888]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.1" metricValueMin="8.3" pageId="17" pageNumber="28" unit="mm" value="8.7" valueMax="9.1" valueMin="8.3">8.3–9.1 mm</quantity>
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.
|
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C31B65635D5615382D0DD732FC97FB95" pageId="17" pageNumber="28" type="materials_examined">
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<paragraph id="8BBE36E85D5615382D0DD732FC35FBC0" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
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||
<emphasis id="B975EAFA5D5615382D0DD732FE22FC4A" bold="true" box="[306,444,893,914]" pageId="17" pageNumber="28">Type Series.</emphasis>
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Described from
|
||
<specimenCount id="9D07FD615D5615382E4FD732FD4EFC4A" box="[624,720,893,914]" pageId="17" pageNumber="28" type="male">19 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D07FD615D5615382F3AD732FCE2FC4A" box="[773,892,893,914]" pageId="17" pageNumber="28" type="female">13 females</specimenCount>
|
||
.
|
||
<materialsCitation id="3B693CB55D5615382FB8D732FEE2FC3A" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="IN- BIO" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="Estacion Biologica Las Alturas" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR 1000 909226" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
|
||
<typeStatus id="54BA884A5D5615382FB8D732FC76FC4A" box="[903,1000,893,914]" pageId="17" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D07FD615D5615382FCFD732FBBDFC4A" box="[1008,1059,893,914]" pageId="17" pageNumber="28" type="male">male</specimenCount>
|
||
(
|
||
<collectionCode id="ED10AE2D5D561538280DD732FED7FC75" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||
): ‘‘
|
||
<collectingCountry id="F31676785D5615382D55D7D7FE77FC75" box="[362,489,920,941]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C5F80A5D5615382DC7D7D7FDF3FC75" box="[504,621,920,941]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="28">Puntarenas</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDAAC925D5615382E42D7D7FD71FC75" box="[637,751,920,941]" pageId="17" pageNumber="28">Coto Brus</collectingMunicipality>
|
||
,
|
||
<location id="8EDE60335D5615382EC1D7D7FBCAFC75" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6:8EDE60335D5615382EC1D7D7FBCAFC75" box="[766,1108,920,941]" country="Costa Rica" municipality="Coto Brus" name="Estacion Biologica Las Alturas" pageId="17" pageNumber="28" stateProvince="Puntarenas">Estación Biológica Las Alturas</location>
|
||
,
|
||
<date id="FFBF10285D5615382D28D7FCFE50FC10" box="[279,462,947,968]" pageId="17" pageNumber="28" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">
|
||
<collectingDate id="EFFBE9C05D5615382D28D7FCFE50FC10" box="[279,462,947,968]" pageId="17" pageNumber="28" value="1992-03-23" valueMax="1992-05-02" valueMin="1992-03-23">23III to 2V92</collectingDate>
|
||
</date>
|
||
,
|
||
<quantity id="4CF99B0D5D5615382DDFD7FCFDA1FC10" box="[480,575,947,968]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">
|
||
<elevation id="002CD1DB5D5615382DDFD7FCFDA1FC10" box="[480,575,947,968]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">1,500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26F4533E5D5615382E6FD7FCFD30FC10" box="[592,686,947,968]" pageId="17" pageNumber="28">F. Araya</collectorName>
|
||
LS 322500, 591300/
|
||
<specimenCode id="DBA79E935D5615382FA0D7FCFEF0FC3A" pageId="17" pageNumber="28">INBIO CR 1000 909226</specimenCode>
|
||
.’’
|
||
</materialsCitation>
|
||
<materialsCitation id="3B693CB55D5615382DBAD782FE3CFC25" accessionNumber="LS 322500" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="INBIO" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="El Rosario de Naranjo" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR I000 909233" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="allotype">
|
||
<typeStatus id="54BA884A5D5615382DBAD782FE7EFC3A" box="[389,480,973,994]" pageId="17" pageNumber="28" type="allotype">Allotype</typeStatus>
|
||
<specimenCount id="9D07FD615D5615382DD6D782FDAEFC3A" box="[489,560,973,994]" pageId="17" pageNumber="28" type="female">female</specimenCount>
|
||
(
|
||
<collectionCode id="ED10AE2D5D5615382E7FD782FD11FC3A" box="[576,655,973,994]" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||
): same data as holotype except
|
||
<specimenCode id="DBA79E935D5615382FDBD782FE3CFC25" pageId="17" pageNumber="28">INBIO CR I000 909233</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B693CB55D5615382D90D7A7FB87FC25" accessionNumber="LS 322500" box="[431,1049,1000,1021]" collectingDate="1992-03-23" collectingDateMax="1992-05-02" collectingDateMin="1992-03-23" collectionCode="CASC, CNC, INBIO, MXAL, ZMHU" collectorName="F. Araya" country="Costa Rica" elevation="1500" latitude="8.966666" location="El Rosario de Naranjo" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" pageId="17" pageNumber="28" specimenCode="INBIO CR 1000 909226" specimenCount="30" specimenCount-female="12" specimenCount-male="18" stateProvince="Alajuela" typeStatus="paratype">
|
||
<typeStatus id="54BA884A5D5615382D90D7A7FD87FC25" box="[431,537,1000,1021]" pageId="17" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||
: same data as holotype (
|
||
<specimenCount id="9D07FD615D5615382F1CD7A7FC19FC25" box="[803,903,1000,1021]" count="18" pageId="17" pageNumber="28" type="male">18 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D07FD615D5615382FABD7A7FB93FC25" box="[916,1037,1000,1021]" count="12" pageId="17" pageNumber="28" type="female">12 females</specimenCount>
|
||
).
|
||
</materialsCitation>
|
||
<typeStatus id="54BA884A5D561538281ED7A7FED0FBC0" pageId="17" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode id="ED10AE2D5D5615382DDDD04CFDB7FBC0" box="[482,553,1027,1048]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="17" pageNumber="28">CASC</collectionCode>
|
||
,
|
||
<collectionCode id="ED10AE2D5D5615382E09D04CFDF0FBC0" box="[566,622,1027,1048]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="17" pageNumber="28">CNC</collectionCode>
|
||
,
|
||
<collectionCode id="ED10AE2D5D5615382E44D04CFD58FBC0" box="[635,710,1027,1048]" pageId="17" pageNumber="28">INBIO</collectionCode>
|
||
,
|
||
<collectionCode id="ED10AE2D5D5615382EECD04CFCBEFBC0" box="[723,800,1027,1048]" pageId="17" pageNumber="28">MXAL</collectionCode>
|
||
and
|
||
<collectionCode id="ED10AE2D5D5615382F68D04CFC39FBC0" box="[855,935,1027,1048]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="17" pageNumber="28">ZMHU</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE36E85D5615382D0DD052FC97FB95" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||
<emphasis id="B975EAFA5D5615382D0DD052FE4AFBEA" bold="true" box="[306,468,1053,1074]" pageId="17" pageNumber="28">Type Locality.</emphasis>
|
||
<location id="8EDE60335D5615382DE5D052FCBBFBEA" LSID="urn:lsid:plazi:treatment:03A887FE5D4815382D03D0A1FC9DF9D6:8EDE60335D5615382DE5D052FCBBFBEA" box="[474,805,1053,1074]" country="Costa Rica" latitude="8.966666" longLatPrecision="1300" longitude="-83.1" municipality="Coto Brus" name="Estacion Biologica Las Alturas" pageId="17" pageNumber="28" stateProvince="Puntarenas">Estación Biológica Las Alturas</location>
|
||
,
|
||
<collectingMunicipality id="6BDAAC925D5615382F10D052FC03FBEA" box="[815,925,1053,1074]" pageId="17" pageNumber="28">Coto Brus</collectingMunicipality>
|
||
, province of
|
||
<collectingRegion id="49C5F80A5D5615382819D052FEFAFB95" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="28">Puntarenas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31676785D5615382D4ED077FE7BFB95" box="[369,485,1080,1101]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||
(approx.
|
||
<geoCoordinate id="EE35502F5D5615382E70D077FD02FB95" box="[591,668,1080,1101]" degrees="8" direction="north" minutes="58" orientation="latitude" pageId="17" pageNumber="28" precision="925" value="8.966666">8°58'N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE35502F5D5615382E96D077FD63FB95" box="[681,765,1080,1101]" degrees="83" direction="west" minutes="6" orientation="longitude" pageId="17" pageNumber="28" precision="925" value="-83.1">83°6'W</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D5615382D0DD01DFD0FFB44" pageId="17" pageNumber="28" type="biology_ecology">
|
||
<paragraph id="8BBE36E85D5615382D0DD01DFD0FFB44" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||
<emphasis id="B975EAFA5D5615382D0DD01DFE78FBBF" bold="true" box="[306,486,1106,1127]" pageId="17" pageNumber="28">Biological Data.</emphasis>
|
||
All males and females of
|
||
<taxonomicName id="4C014D6B5D5615382F35D01CFC3AFBB0" box="[778,932,1107,1128]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov.">
|
||
<emphasis id="B975EAFA5D5615382F35D01CFC3AFBB0" box="[778,932,1107,1128]" italics="true" pageId="17" pageNumber="28">P. guapiloides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657815D5615382F91D01CFC77FBB0" box="[942,1001,1107,1128]" pageId="17" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
||
were collected at lights on the border of coffee plantations located at
|
||
<quantity id="4CF99B0D5D5615382F92D022FB9BFB5A" box="[941,1029,1133,1154]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="28" unit="m" value="1500.0">1,500 m</quantity>
|
||
of altitude. Phenology: March–May (22).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D5615382D0DD0EEFD1DFA67" pageId="17" pageNumber="28" type="discussion">
|
||
<paragraph id="8BBE36E85D5615382D0DD0EEFD1DFA67" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||
<emphasis id="B975EAFA5D5615382D0DD0EEFE00FB6E" bold="true" box="[306,414,1185,1206]" pageId="17" pageNumber="28">Remarks.</emphasis>
|
||
<taxonomicName id="4C014D6B5D5615382D96D0EDFCC8FB6F" box="[425,854,1186,1207]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiloides" status="sp. nov." subGenus="Phyllophaga">
|
||
<emphasis id="B975EAFA5D5615382D96D0EDFDB1FB6F" box="[425,559,1186,1207]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||
(
|
||
<emphasis id="B975EAFA5D5615382E7DD0EDFD59FB6F" box="[578,711,1186,1207]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||
)
|
||
<emphasis id="B975EAFA5D5615382EE3D0EDFCC8FB6F" box="[732,854,1186,1207]" italics="true" pageId="17" pageNumber="28">guapiloides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657815D5615382F5ED0EDFC00FB6F" box="[865,926,1186,1207]" pageId="17" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
|
||
probably belong to ‘‘
|
||
<emphasis id="B975EAFA5D5615382D78D0F3FE35FB09" box="[327,427,1212,1233]" italics="true" pageId="17" pageNumber="28">setidorsis</emphasis>
|
||
’’ species group (
|
||
<emphasis id="B975EAFA5D5615382E50D0F3FD37FB09" box="[623,681,1212,1233]" italics="true" pageId="17" pageNumber="28">sensu</emphasis>
|
||
<bibRefCitation id="EF904B195D5615382E8AD0F3FCA1FB09" author="Moron" box="[693,831,1212,1233]" firstAuthor="Moron" pageId="17" pageNumber="28" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
|
||
), but the structure of the aedeagus is much different from that species. It is closely related to
|
||
<taxonomicName id="4C014D6B5D5615382834D098FD2FFADE" authority="Saylor" authorityName="Saylor" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles" subGenus="Phyllophaga">
|
||
<emphasis id="B975EAFA5D5615382834D098FEC6FADE" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||
(
|
||
<emphasis id="B975EAFA5D5615382D51D0BEFE6AFADE" box="[366,500,1265,1286]" italics="true" pageId="17" pageNumber="28">Phyllophaga</emphasis>
|
||
)
|
||
<emphasis id="B975EAFA5D5615382E37D0BEFDFFFADE" box="[520,609,1265,1286]" italics="true" pageId="17" pageNumber="28">guapiles</emphasis>
|
||
Saylor
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49C5F80A5D5615382EC3D0BEFCEAFADE" box="[764,884,1265,1286]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="28">Guanacaste</collectingRegion>
|
||
area and is without external differences, but the structure of parameres and the sclerotized support of aedeagus are clearly different (
|
||
<figureCitation id="133A2A6D5D5615382EBCD169FC98FAE3" box="[643,774,1318,1339]" captionStart="Figs" captionStartId="18.[330,373,687,706]" captionTargetBox="[341,1102,275,666]" captionTargetId="figure-12@18.[340,1105,272,667]" captionTargetPageId="18" captionText="Figs. 32–34. Phyllophaga guapiles. 32) apex of genital capsule, lateral view; 33) apex of paramera and sclerotized support of aedeagus, ventrodistal view; 34) genital plates, female. Scale lines 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900327" httpUri="https://zenodo.org/record/4900327/files/figure.png" pageId="17" pageNumber="28">Figs. 32–33</figureCitation>
|
||
). Female genital plates are also slightly different from
|
||
<taxonomicName id="4C014D6B5D5615382E2BD10FFD10FA8D" box="[532,654,1344,1365]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles">
|
||
<emphasis id="B975EAFA5D5615382E2BD10FFD10FA8D" box="[532,654,1344,1365]" italics="true" pageId="17" pageNumber="28">P. guapiles</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="133A2A6D5D5615382E9FD10FFD6AFA8D" box="[672,756,1344,1365]" captionStart="Figs" captionStartId="18.[330,373,687,706]" captionTargetBox="[341,1102,275,666]" captionTargetId="figure-12@18.[340,1105,272,667]" captionTargetPageId="18" captionText="Figs. 32–34. Phyllophaga guapiles. 32) apex of genital capsule, lateral view; 33) apex of paramera and sclerotized support of aedeagus, ventrodistal view; 34) genital plates, female. Scale lines 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4900327" httpUri="https://zenodo.org/record/4900327/files/figure.png" pageId="17" pageNumber="28">Fig. 34</figureCitation>
|
||
). Density, size and shape of the punctures on the head, pronotum, elytra and pygidium; body vestiture; shape of hind tibial spurs, and tarsal claws; structure of tarsomeres and general pattern of aedeagus, will aid to separate this pair of sister species from other species in the ‘‘
|
||
<emphasis id="B975EAFA5D5615382DFDD1E5FDB8FA67" box="[450,550,1450,1471]" italics="true" pageId="17" pageNumber="28">setidorsis</emphasis>
|
||
’’ group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B65635D5615382D0DD18BFC9DF9D6" pageId="17" pageNumber="28" type="etymology">
|
||
<paragraph id="8BBE36E85D5615382D0DD18BFC9DF9D6" blockId="17.[279,1113,277,1550]" pageId="17" pageNumber="28">
|
||
<emphasis id="B975EAFA5D5615382D0DD18BFE2FFA01" bold="true" box="[306,433,1476,1497]" pageId="17" pageNumber="28">Etymology.</emphasis>
|
||
Derived from latin sufix
|
||
<emphasis id="B975EAFA5D5615382EE2D18BFC85FA01" box="[733,795,1476,1497]" italics="true" pageId="17" pageNumber="28">oides,</emphasis>
|
||
similar, false (Jaeger 1955), meaning ‘‘similar to
|
||
<taxonomicName id="4C014D6B5D5615382DCCD190FDD2FA2C" box="[499,588,1503,1524]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="28" phylum="Arthropoda" rank="species" species="guapiles">
|
||
<emphasis id="B975EAFA5D5615382DCCD190FDD2FA2C" box="[499,588,1503,1524]" italics="true" pageId="17" pageNumber="28">guapiles</emphasis>
|
||
</taxonomicName>
|
||
.’’ In the popular Spanish language of
|
||
<collectingCountry id="F31676785D5615382FE0D190FBCAFA2C" box="[991,1108,1503,1524]" name="Costa Rica" pageId="17" pageNumber="28">Costa Rica</collectingCountry>
|
||
, the term ‘‘
|
||
<emphasis id="B975EAFA5D5615382DB6D1B6FE52F9D6" box="[393,460,1529,1550]" italics="true" pageId="17" pageNumber="28">guapil</emphasis>
|
||
’’ is applied to paired things.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |