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<document id="445D2A6B27B2C94A939620DA23FEDC8A" ID-DOI="10.1186/s13358-024-00307-8" ID-ISSN="1664-2384" ID-PMC="PMC10991048" ID-PubMed="38584612" ID-Zenodo-Dep="12005092" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1718654698973" checkinUser="felipe" docAuthor="Pohle, Alexander &amp; Klug, Christian" docDate="2024" docId="761887DBFFAB2C261301FD07FAF0344A" docLanguage="en" docName="SwissJPalaeontol.143.14.pdf" docOrigin="Swiss Journal of Palaeontology (14) 143 (1)" docSource="http://dx.doi.org/10.1186/s13358-024-00307-8" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Paratrematoceras Schastlivceva 1981" docType="treatment" docVersion="3" lastPageNumber="26" masterDocId="8A21FFA3FFB22C3F102CFFA9FFEF3307" masterDocTitle="Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea" masterLastPageNumber="32" masterPageNumber="1" pageNumber="26" updateTime="1721670897711" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FE57613C8E86A16B031D21EF74450DBF">Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea</mods:title>
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<mods:namePart id="F2B7B638AE14E4E7E1D8A228019DD553">Pohle, Alexander</mods:namePart>
<mods:affiliation id="FBD93EA3B2B1AABFC82E57904DE3439A">Institut fÜr Geologie, Mineralogie und Geophysik, Ruhr-Universität Bochum, 44801 Bochum, Germany &amp; Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="E1CE718809A9065089D217C493C408FB">Klug, Christian</mods:namePart>
<mods:affiliation id="74B6FCBFB42CFDB4F01AD68AA98EAA5A">Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<treatment id="761887DBFFAB2C261301FD07FAF0344A" LSID="urn:lsid:plazi:treatment:761887DBFFAB2C261301FD07FAF0344A" httpUri="http://treatment.plazi.org/id/761887DBFFAB2C261301FD07FAF0344A" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection id="B6AB6546FFAB2C261301FD07FACF31C1" box="[813,1312,686,710]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="FE0E36CDFFAB2C261301FD07FACF31C1" blockId="25.[813,1312,686,710]" box="[813,1312,686,710]" pageId="25" pageNumber="26">
<heading id="A54681A1FFAB2C261301FD07FACF31C1" box="[813,1312,686,710]" fontSize="36" level="2" pageId="25" pageNumber="26" reason="3">
Genus
<taxonomicName id="39B14D4EFFAB2C261355FD07FACF31C1" authority="Schastlivceva, 1981" authorityName="Schastlivceva" authorityYear="1981" box="[889,1312,686,710]" class="Cephalopoda" family="Pseudorthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFAB2C261355FD07FBA731C2" bold="true" box="[889,1096,686,709]" italics="true" pageId="25" pageNumber="26">Paratrematoceras</emphasis>
<bibRefCitation id="9A204B3CFFAB2C261462FD07FACF31C1" author="Schastlivceva, N. P." box="[1102,1312,686,710]" pageId="25" pageNumber="26" pagination="76 - 82" refId="ref26976" refString="Schastlivceva, N. P. (1981). O sistematicheskom polozhenii triasovykh ortozeratoidej yuga SSSR [On the systematic position of Triassic orthoceratoids from the southern USSR]. Byulleten' Moskovskogo obshestva ispytatelej prirody. Otdel Geologicheskij, 56, 76 - 82. (in Russian)." type="journal article" year="1981">Schastlivceva, 1981</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="B6AB6546FFAB2C261301FD46FAA73040" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="FE0E36CDFFAB2C261301FD46FAA73040" blockId="25.[813,1437,751,839]" pageId="25" pageNumber="26">
<emphasis id="CCC5EADFFFAB2C261301FD46FC4F3000" bold="true" box="[813,928,751,775]" pageId="25" pageNumber="26">Diagnosis</emphasis>
: Trematoceratid with narrow, tubular siphuncle with short orthochoanitic septal necks that sharply bend in adapically at the transition to the septum.
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFAB2C261301FCD8FAF0308E" box="[813,1311,880,905]" pageId="25" pageNumber="26" type="type_taxon">
<paragraph id="FE0E36CDFFAB2C261301FCD8FAF0308E" blockId="25.[813,1311,880,905]" box="[813,1311,880,905]" pageId="25" pageNumber="26">
<emphasis id="CCC5EADFFFAB2C261301FCD8FC2A308F" bold="true" box="[813,965,880,905]" pageId="25" pageNumber="26">
<typeStatus id="210A886FFFAB2C261301FCD8FC89308E" box="[813,870,881,905]" pageId="25" pageNumber="26">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="39B14D4EFFAB2C2613E0FCD8FAF0308E" authority="Schastlivceva, 1981" authorityName="Schastlivceva" authorityYear="1981" box="[972,1311,881,905]" class="Cephalopoda" family="Orthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="shevyrevi">
<emphasis id="CCC5EADFFFAB2C2613E0FCD8FBA8308E" box="[972,1095,881,905]" italics="true" pageId="25" pageNumber="26">P. shevyrevi</emphasis>
<bibRefCitation id="9A204B3CFFAB2C261461FCD8FAF0308E" author="Schastlivceva, N. P." box="[1101,1311,881,905]" pageId="25" pageNumber="26" pagination="76 - 82" refId="ref26976" refString="Schastlivceva, N. P. (1981). O sistematicheskom polozhenii triasovykh ortozeratoidej yuga SSSR [On the systematic position of Triassic orthoceratoids from the southern USSR]. Byulleten' Moskovskogo obshestva ispytatelej prirody. Otdel Geologicheskij, 56, 76 - 82. (in Russian)." type="journal article" year="1981">Schastlivceva, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFAB2C261301FC1BFB7A30EE" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="FE0E36CDFFAB2C261301FC1BFB7A30EE" blockId="25.[813,1436,946,1002]" pageId="25" pageNumber="26">
<emphasis id="CCC5EADFFFAB2C261301FC1BFC0230CD" bold="true" box="[813,1005,946,970]" pageId="25" pageNumber="26">Included species</emphasis>
:
<taxonomicName id="39B14D4EFFAB2C2613D5FC1AFAA630CD" authority="Schastlivceva, 1981" authorityName="Schastlivceva" authorityYear="1981" box="[1017,1353,946,970]" class="Cephalopoda" family="Orthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="ornatum">
<emphasis id="CCC5EADFFFAB2C2613D5FC1AFB9F30CD" box="[1017,1136,947,970]" italics="true" pageId="25" pageNumber="26">P. ornatum</emphasis>
<bibRefCitation id="9A204B3CFFAB2C26145BFC1BFAA630CD" author="Schastlivceva, N. P." box="[1143,1353,946,970]" pageId="25" pageNumber="26" pagination="76 - 82" refId="ref26976" refString="Schastlivceva, N. P. (1981). O sistematicheskom polozhenii triasovykh ortozeratoidej yuga SSSR [On the systematic position of Triassic orthoceratoids from the southern USSR]. Byulleten' Moskovskogo obshestva ispytatelej prirody. Otdel Geologicheskij, 56, 76 - 82. (in Russian)." type="journal article" year="1981">Schastlivceva, 1981</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="39B14D4EFFAB2C261579FC1AFBF730ED" authority="(Hauer, 1846)" authorityName="Pohle &amp; Klug" authorityYear="2024" baseAuthorityName="Hauer" baseAuthorityYear="1846" class="Cephalopoda" family="Orthoceratidae" genus="Orthoceras" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="salinarium" status="comb. nov.">
<emphasis id="CCC5EADFFFAB2C261579FC1AFC9030ED" italics="true" pageId="25" pageNumber="26">P. salinarium</emphasis>
(Hauer, 1846)
</taxonomicName>
<taxonomicNameLabel id="D7F657A4FFAB2C261432FC7BFB7A30EE" box="[1054,1173,978,1002]" pageId="25" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFAB2C261301FBBAFAFA35EC" pageId="25" pageNumber="26" type="discussion">
<paragraph id="FE0E36CDFFAB2C261301FBBAFA95374C" blockId="25.[813,1437,1043,1771]" pageId="25" pageNumber="26">
<emphasis id="CCC5EADFFFAB2C261301FBBAFC7C372C" bold="true" box="[813,915,1043,1067]" pageId="25" pageNumber="26">Remarks</emphasis>
: Differs from
<taxonomicName id="39B14D4EFFAB2C261402FBBDFB2D372C" authorityName="Eichwald" authorityYear="1851" box="[1070,1218,1044,1067]" class="Cephalopoda" family="Orthoceratidae" genus="Trematoceras" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFAB2C261402FBBDFB2D372C" box="[1070,1218,1044,1067]" italics="true" pageId="25" pageNumber="26">Trematoceras</emphasis>
</taxonomicName>
in its narrower siphuncle with sharply bent, orthochoanitic septal necks.
</paragraph>
<paragraph id="FE0E36CDFFAB2C26136EFBFDFAFA35EC" blockId="25.[813,1437,1043,1771]" pageId="25" pageNumber="26">
In addition to the species listed by
<bibRefCitation id="9A204B3CFFAB2C26152BFBFAFC9A378C" author="Schastlivceva, N. P." pageId="25" pageNumber="26" pagination="1 - 104" refId="ref27077" refString="Schastlivceva, N. P. (1988). Triasovye ortozeratidy i nautilidy SSSR [Triassic orthoceratoids and nautiloids of the USSR]. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta, 229, 1 - 104. (in Russian)." type="journal article" year="1988">Schastlivceva (1988)</bibRefCitation>
, which includes specimens described as
<taxonomicName id="39B14D4EFFAB2C261565FBDDFBFF37AC" baseAuthorityName="Hauer" baseAuthorityYear="1847" class="Cephalopoda" family="Orthoceratidae" genus="Trematoceras" higherTaxonomySource="CoL,GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="dubium">
<emphasis id="CCC5EADFFFAB2C261565FBDDFC9437AC" italics="true" pageId="25" pageNumber="26">Trematoceras</emphasis>
aff.
<emphasis id="CCC5EADFFFAB2C261395FB3AFBFF37AC" box="[953,1040,1171,1195]" italics="true" pageId="25" pageNumber="26">dubium</emphasis>
</taxonomicName>
from the Olenekian of
<collectingCountry id="86A6765DFFAB2C261515FB3AFCB337CC" name="Afghanistan" pageId="25" pageNumber="26">Afghanistan</collectingCountry>
(Kummel &amp; Erben, 1968),
<taxonomicName id="39B14D4EFFAB2C2614B7FB1AFC5C37EC" authority="Hauer, 1846" authorityName="Hauer" authorityYear="1846" class="Cephalopoda" family="Orthoceratidae" genus="Orthoceras" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="salinarium">
<emphasis id="CCC5EADFFFAB2C2614B7FB1AFA7237CC" box="[1179,1437,1203,1227]" italics="true" pageId="25" pageNumber="26">Orthoceras salinarium</emphasis>
Hauer, 1846
</taxonomicName>
from the Norian of
<collectingCountry id="86A6765DFFAB2C2614BFFB7AFB0A37EC" box="[1171,1253,1235,1259]" name="Austria" pageId="25" pageNumber="26">Austria</collectingCountry>
may be referred to the genus, as it has a similarly narrow siphuncle with sharply bent septal necks. However, this is purely based on Hauer’s (1846) drawings and investigation of his
<typeStatus id="210A886FFFAB2C261375FAFDFC66366B" box="[857,905,1364,1388]" pageId="25" pageNumber="26">type</typeStatus>
material is required to test this assignment.
<taxonomicName id="39B14D4EFFAB2C261301FADAFBF3368C" box="[813,1052,1395,1419]" class="Cephalopoda" family="Orthoceratidae" genus="Michelinoceras" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="undefined-A">
<emphasis id="CCC5EADFFFAB2C261301FADAFC3A368C" box="[813,981,1395,1419]" italics="true" pageId="25" pageNumber="26">Michelinoceras</emphasis>
sp. A
</taxonomicName>
in Tongtherm et al. (2016) from the Anisian of
<collectingCountry id="86A6765DFFAB2C2613FFFA3AFBDA36AC" box="[979,1077,1427,1451]" name="Thailand" pageId="25" pageNumber="26">Thailand</collectingCountry>
possibly belongs to
<taxonomicName id="39B14D4EFFAB2C261537FA3DFC9436CC" authorityName="Schastlivceva" authorityYear="1981" class="Cephalopoda" family="Orthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFAB2C261537FA3DFC9436CC" italics="true" pageId="25" pageNumber="26">Paratrematoceras</emphasis>
</taxonomicName>
due to its narrow siphuncle and orthochoanitic septal necks. Several species from Olenekian to Anisian deposits have been assigned to
<taxonomicName id="39B14D4EFFAB2C261482FA5DFA9B350C" authorityName="Schastlivceva" authorityYear="1981" box="[1198,1396,1524,1547]" class="Cephalopoda" family="Orthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFAB2C261482FA5DFA9B350C" box="[1198,1396,1524,1547]" italics="true" pageId="25" pageNumber="26">Paratrematoceras</emphasis>
</taxonomicName>
by Grădinaru et al. (2007), but since they were not accompanied by illustrations, they represent nomina nuda and the presence of the genus in these rocks is unconfirmed. Consequently, the genus appears to be already present and relatively widespread in the early Middle Triassic, although further studies are needed to differentiate it from
<taxonomicName id="39B14D4EFFAB2C2613F8F97DFB8335EC" authorityName="Eichwald" authorityYear="1851" box="[980,1132,1748,1771]" class="Cephalopoda" family="Orthoceratidae" genus="Trematoceras" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthocerida" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFAB2C2613F8F97DFB8335EC" box="[980,1132,1748,1771]" italics="true" pageId="25" pageNumber="26">Trematoceras</emphasis>
</taxonomicName>
with certainty.
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFAB2C261301F8BDFAF0344A" pageId="25" pageNumber="26" type="distribution">
<paragraph id="FE0E36CDFFAB2C261301F8BDFAF0344A" blockId="25.[813,1436,1812,1869]" pageId="25" pageNumber="26">
<emphasis id="CCC5EADFFFAB2C261301F8BDFC54342B" bold="true" box="[813,955,1812,1836]" pageId="25" pageNumber="26">Occurrence:</emphasis>
Northern Caucasus, Alps?,
<collectingCountry id="86A6765DFFAB2C2614C2F8BCFAA1342A" box="[1262,1358,1813,1837]" name="Thailand" pageId="25" pageNumber="26">Thailand</collectingCountry>
?; Middle Triassic (Anisian)?, Late Triassic (Norian).
</paragraph>
</subSubSection>
</treatment>
</document>
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