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<document id="6CD1580C0DA65E01440BD38FACEF2B62" ID-DOI="10.1186/s13358-024-00307-8" ID-ISSN="1664-2384" ID-PMC="PMC10991048" ID-PubMed="38584612" ID-Zenodo-Dep="12005092" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1718654698973" checkinUser="felipe" docAuthor="Pohle, Alexander &amp; Klug, Christian" docDate="2024" docId="761887DBFFA12C2B10BBF8B8FEEC32C2" docLanguage="en" docName="SwissJPalaeontol.143.14.pdf" docOrigin="Swiss Journal of Palaeontology (14) 143 (1)" docSource="http://dx.doi.org/10.1186/s13358-024-00307-8" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Ticinoteuthis Pohle &amp; Klug 2024, gen. nov." docType="treatment" docUuid="F7E76450-CFD4-4BCA-8D4E-7FF4D6D83094" docUuidSource="ZooBank" docVersion="3" lastPageNumber="21" masterDocId="8A21FFA3FFB22C3F102CFFA9FFEF3307" masterDocTitle="Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea" masterLastPageNumber="32" masterPageNumber="1" pageNumber="20" updateTime="1721670897711" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2964F6AA0D984150622049DA9FD6176E">Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea</mods:title>
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<mods:namePart id="F7E909252FF816BBB8995B5FD52FE8A9">Pohle, Alexander</mods:namePart>
<mods:affiliation id="8D5CD4D15CA0E689CC4231A30CF771A1">Institut fÜr Geologie, Mineralogie und Geophysik, Ruhr-Universität Bochum, 44801 Bochum, Germany &amp; Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:affiliation id="6A6812BC1E9E2802FE96542D3172D379">Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<subSubSection id="B6AB6546FFA12C2C10BBF8B8FE0B342F" box="[151,484,1808,1833]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="FE0E36CDFFA12C2C10BBF8B8FE0B342F" blockId="19.[151,484,1808,1833]" box="[151,484,1808,1833]" pageId="19" pageNumber="20">
<heading id="A54681A1FFA12C2C10BBF8B8FE0B342F" box="[151,484,1808,1833]" fontSize="36" level="2" pageId="19" pageNumber="20" reason="3">
Genus
<taxonomicName id="39B14D4EFFA12C2C10C8F8B9FE92342F" authority="Pohle &amp; Klug, 2024" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[228,381,1808,1832]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="CCC5EADFFFA12C2C10C8F8B9FE92342F" bold="true" box="[228,381,1808,1832]" italics="true" pageId="19" pageNumber="20">Ticinoteuthis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7F657A4FFA12C2C11AFF8B8FE0B342F" box="[387,484,1809,1833]" pageId="19" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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.
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFA12C2C1301FF42FB0D3224" pageId="19" pageNumber="20" type="etymology">
<paragraph id="FE0E36CDFFA12C2C1301FF42FB0D3224" blockId="19.[813,1436,235,292]" pageId="19" pageNumber="20">
<emphasis id="CCC5EADFFFA12C2C1301FF42FC463204" bold="true" box="[813,937,235,259]" pageId="19" pageNumber="20">Etymology</emphasis>
: Named after the
<collectingRegion id="3C75F82FFFA12C2C14A4FF42FAC23204" box="[1160,1325,235,259]" country="Switzerland" name="Ticino" pageId="19" pageNumber="20">Canton Ticino</collectingRegion>
(
<collectingCountry id="86A6765DFFA12C2C156DFF42FCB03224" name="Switzerland" pageId="19" pageNumber="20">Switzerland</collectingCountry>
), where the
<typeStatus id="210A886FFFA12C2C13CEFEA5FBFE3223" box="[994,1041,268,292]" pageId="19" pageNumber="20">type</typeStatus>
locality is situated.
</paragraph>
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<subSubSection id="B6AB6546FFA12C2C1301FEE4FCB53283" pageId="19" pageNumber="20" type="type_taxon">
<paragraph id="FE0E36CDFFA12C2C1301FEE4FCB53283" blockId="19.[813,1437,332,388]" pageId="19" pageNumber="20">
<emphasis id="CCC5EADFFFA12C2C1301FEE4FC2C3263" bold="true" box="[813,963,332,357]" pageId="19" pageNumber="20">
<typeStatus id="210A886FFFA12C2C1301FEE4FC893262" box="[813,870,333,357]" pageId="19" pageNumber="20">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="39B14D4EFFA12C2C13FFFEE4FAF33262" authority="Pohle &amp; Klug, 2024" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[979,1308,333,357]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="chuchichaeschtli" status="gen. et sp. nov.">
<emphasis id="CCC5EADFFFA12C2C13FFFEE4FAF33262" box="[979,1308,333,357]" italics="true" pageId="19" pageNumber="20">Ticinoteuthis chuchichaeschtli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7F657A4FFA12C2C150BFEE4FCB53283" pageId="19" pageNumber="20" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFA12C2C1301FE07FB0632C1" box="[813,1257,430,454]" pageId="19" pageNumber="20" type="description">
<paragraph id="FE0E36CDFFA12C2C1301FE07FB0632C1" blockId="19.[813,1257,430,454]" box="[813,1257,430,454]" pageId="19" pageNumber="20">
<emphasis id="CCC5EADFFFA12C2C1301FE07FC0232C1" bold="true" box="[813,1005,430,454]" pageId="19" pageNumber="20">Included species</emphasis>
: Only the
<typeStatus id="210A886FFFA12C2C1472FE07FB6232C1" box="[1118,1165,430,454]" pageId="19" pageNumber="20">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFA12C2C1301FE46FB6D31A0" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="FE0E36CDFFA12C2C1301FE46FB6D31A0" blockId="19.[813,1436,495,679]" pageId="19" pageNumber="20">
<emphasis id="CCC5EADFFFA12C2C1301FE46FC4F3100" bold="true" box="[813,928,495,519]" pageId="19" pageNumber="20">Diagnosis</emphasis>
: Small orthoconic (longiconic) phragmocone with slender expansion rate of about 59° and depressed cross section. Septa apparently inclined towards the venter in anterior direction. Siphuncle submarginal, probably with retrochoanitic septal necks. Neither primordial nor proper rostrum are known.
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFA12C2B1301FD58FD4F3264" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="FE0E36CDFFA12C2C1301FD58FB06352E" blockId="19.[813,1437,753,1929]" pageId="19" pageNumber="20">
<emphasis id="CCC5EADFFFA12C2C1301FD58FC75300E" bold="true" box="[813,922,753,777]" pageId="19" pageNumber="20">Remarks:</emphasis>
<taxonomicName id="39B14D4EFFA12C2C1386FD58FBD8300E" authority="Pohle &amp; Klug, 2024" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[938,1079,753,777]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="CCC5EADFFFA12C2C1386FD58FBD8300E" box="[938,1079,753,777]" italics="true" pageId="19" pageNumber="20">Ticinoteuthis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7F657A4FFA12C2C1464FD58FB5C300F" box="[1096,1203,753,777]" pageId="19" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="39B14D4EFFA12C2C1544FD58FC2A302E" authorityName="Jeletzky" authorityYear="1965" class="Cephalopoda" family="Xiphoteuthidae" genus="Mojsisovicsteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFA12C2C1544FD58FC2A302E" italics="true" pageId="19" pageNumber="20">Mojsisovicsteuthis</emphasis>
</taxonomicName>
but differs in its depressed rather than compressed cross section, slenderer apical angle and apparently retrochoanitic instead of prochoanitic septal necks. For both
<taxonomicName id="39B14D4EFFA12C2C1422FCD8FB3E308E" authorityName="Jeletzky" authorityYear="1965" box="[1038,1233,881,905]" class="Cephalopoda" family="Xiphoteuthidae" genus="Mojsisovicsteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFA12C2C1422FCD8FB3E308E" box="[1038,1233,881,905]" italics="true" pageId="19" pageNumber="20">Mojsisovicsteuthis</emphasis>
</taxonomicName>
and
<taxonomicName id="39B14D4EFFA12C2C1523FCD8FA73308E" authority="Pohle &amp; Klug, 2024" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[1295,1436,881,905]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="CCC5EADFFFA12C2C1523FCD8FA73308E" box="[1295,1436,881,905]" italics="true" pageId="19" pageNumber="20">Ticinoteuthis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7F657A4FFA12C2C1301FC38FC7E30AF" box="[813,913,913,937]" pageId="19" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
, a rostrum (or telum) is undocumented, which was interpreted by Jeletzky (1966) as primary absence. If this is the case, then they cannot be assigned to the Aulacoceratida, an approach that was already proposed by Mariotti and Pignatti (1992, 1993) and by the Treatise on Invertebrate Palaeontology (Mariotti et al., 2021). The absence of a rostrum may be taken as evidence to classify these taxa within the Phragmoteuthida, but representatives of this order are generally characterised by much larger apical angles and possess a typical three-lobed proostracum (Fuchs &amp; Donovan, 2018). In contrast,
<bibRefCitation id="9A204B3CFFA12C2C1301FB58FC29360E" author="Rieber, H." box="[813,966,1265,1289]" pageId="19" pageNumber="20" refId="ref26613" refString="Rieber, H. (1973). Cephalopoden aus der Grenznitumenzone (Mittlere Trias)" type="book" year="1973">Rieber (1973)</bibRefCitation>
described body chambers of
<taxonomicName id="39B14D4EFFA12C2C150FFB58FC6F362E" authorityName="Jeletzky" authorityYear="1965" class="Cephalopoda" family="Xiphoteuthidae" genus="Mojsisovicsteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFA12C2C150FFB58FC6F362E" italics="true" pageId="19" pageNumber="20">Mojsisovicsteuthis</emphasis>
</taxonomicName>
, which is mutually exclusive with any proostracum. For
<taxonomicName id="39B14D4EFFA12C2C138CFA98FB9E364E" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[928,1137,1329,1353]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="chuchichaeschtli">
<emphasis id="CCC5EADFFFA12C2C138CFA98FB9E364E" box="[928,1137,1329,1353]" italics="true" pageId="19" pageNumber="20">T. chuchichaeschtli</emphasis>
</taxonomicName>
, it is unclear if a tubular body chamber or a proostracum was present, although the similarity to
<taxonomicName id="39B14D4EFFA12C2C13D8FAD8FB58368E" authorityName="Jeletzky" authorityYear="1965" box="[1012,1207,1393,1417]" class="Cephalopoda" family="Xiphoteuthidae" genus="Mojsisovicsteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFA12C2C13D8FAD8FB58368E" box="[1012,1207,1393,1417]" italics="true" pageId="19" pageNumber="20">Mojsisovicsteuthis</emphasis>
</taxonomicName>
perhaps suggests a tubular body chamber as well. Thus, they do not fit any of the known Mesozoic coleoid orders or even families, implying that besides desperately needed new data, current family- or order-level definitions either need to be adjusted or new names to be established.
</paragraph>
<paragraph id="FE0E36CDFFA12C2C136EF998FBDD342E" blockId="19.[813,1437,753,1929]" pageId="19" pageNumber="20">
It is possible that “
<taxonomicName id="39B14D4EFFA12C2C1439F998FC8E356E" authority="Mojsisovics, 1873" authorityName="Mojsisovics" authorityYear="1873" class="Cephalopoda" family="Orthoceratidae" genus="Orthoceras" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="styriacum">
<emphasis id="CCC5EADFFFA12C2C1439F998FB62354E" box="[1045,1165,1585,1609]" italics="true" pageId="19" pageNumber="20">Orthoceras</emphasis>
<emphasis id="CCC5EADFFFA12C2C148CF998FAE2354F" box="[1184,1293,1585,1608]" italics="true" pageId="19" pageNumber="20">styriacum</emphasis>
<bibRefCitation id="9A204B3CFFA12C2C153AF998FC8E356E" author="Mojsisovics, E. von" pageId="19" pageNumber="20" pagination="1 - 82" refId="ref25409" refString="Mojsisovics, E. von (1873). Die Cephalopoden der Hallstatter Kalke [The cephalopods from the Hallstatt limestones]. Abhandlungen Der Geologischen Reichsanstalt Wien, 6, 1 - 82. (in German)." type="journal article" year="1873">Mojsisovics, 1873</bibRefCitation>
</taxonomicName>
from the Carnian of
<collectingCountry id="86A6765DFFA12C2C1471F9F8FB42356E" box="[1117,1197,1617,1641]" name="Austria" pageId="19" pageNumber="20">Austria</collectingCountry>
belongs to the same group of species, perhaps even to
<taxonomicName id="39B14D4EFFA12C2C14E4F9D8FABA358E" authorityName="Pohle &amp; Klug" authorityYear="2024" box="[1224,1365,1649,1673]" class="Cephalopoda" family="Orthoceratidae" genus="Ticinoteuthis" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="CCC5EADFFFA12C2C14E4F9D8FABA358E" box="[1224,1365,1649,1673]" italics="true" pageId="19" pageNumber="20">Ticinoteuthis</emphasis>
</taxonomicName>
, as it has a similar expansion rate and cross section and further appears to agree with in the inclination of the septa. However, as the position of the siphuncle is unknown in “
<taxonomicName id="39B14D4EFFA12C2C131AF958FC25340F" authorityName="Mojsisovics" authorityYear="1873" box="[822,970,1777,1801]" class="Cephalopoda" family="Orthoceratidae" genus="Orthoceras" kingdom="Animalia" order="Orthocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="styriacum">
<emphasis id="CCC5EADFFFA12C2C131AF958FCA6340F" box="[822,841,1777,1800]" italics="true" pageId="19" pageNumber="20">O</emphasis>
.”
<emphasis id="CCC5EADFFFA12C2C1371F958FC25340F" box="[861,970,1777,1800]" italics="true" pageId="19" pageNumber="20">styriacum</emphasis>
</taxonomicName>
, it cannot be excluded that it represents a crushed trematoceratid.
</paragraph>
<paragraph id="FE0E36CDFFA12C2B136EF898FD4F3264" blockId="19.[813,1437,753,1929]" lastBlockId="20.[151,775,235,355]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Aulacoceratids differ in having a thick aragonitic rostrum, but their phragmocones generally expand more rapidly and are (sub-) circular in cross section. The only aulacoceratid with a similarly depressed cross section of the phragmocone is
<emphasis id="CCC5EADFFFA62C2B1151FEA2FDFD3224" box="[381,530,267,291]" italics="true" pageId="20" pageNumber="21">Miyagiteuthis</emphasis>
, which further differs in having a strictly marginal siphuncle and less inclined septa (Mariotti et al., 2021;
<bibRefCitation id="9A204B3CFFA62C2B11EDFEE2FD7D3264" author="Niko, S. &amp; Ehiro, M." box="[449,658,331,355]" pageId="20" pageNumber="21" pagination="1 - 8" refId="ref25821" refString="Niko, S., &amp; Ehiro, M. (2018). Aulacocerid coleoids from the Triassic of the South Kitakami Belt, Northeast Japan. Bulletin of the Tohoku University Museum, 17, 1 - 8." type="journal article" year="2018">Niko &amp; Ehiro, 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B6AB6546FFA62C2B10BBFE25FEEC32C2" pageId="20" pageNumber="21" type="distribution">
<paragraph id="FE0E36CDFFA62C2B10BBFE25FEEC32C2" blockId="20.[151,774,396,453]" pageId="20" pageNumber="21">
<emphasis id="CCC5EADFFFA62C2B10BBFE25FEC932A3" bold="true" box="[151,294,396,420]" pageId="20" pageNumber="21">Occurrence:</emphasis>
<collectingRegion id="3C75F82FFFA62C2B116DFE24FE6632A2" box="[321,393,397,421]" country="Switzerland" name="Ticino" pageId="20" pageNumber="21">Ticino</collectingRegion>
,
<collectingCountry id="86A6765DFFA62C2B1184FE24FDC532A2" box="[424,554,397,421]" name="Switzerland" pageId="20" pageNumber="21">Switzerland</collectingCountry>
; Middle Triassic (Anisian).
</paragraph>
</subSubSection>
</treatment>
</document>