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<document id="A4F74B9364441DD95FEBA279089B7E53" ID="10.11646/zootaxa.4170.2.12" ID-DOI="10.11646/zootaxa.4170.2.12" ID-GBIF-Dataset="89fc4d5e-607d-4cc4-84fe-58c99b9de56f" ID-ISSN="1175-5326" ID-Zenodo-Dep="257464" ID-ZooBank="A3692741-1169-4FBF-94CA-6AF0E94C0288" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474615847330" checkinUser="plazi" docAuthor="Krikken, J." docDate="2016" docId="9272877D5324105CD59EFF4302931184" docLanguage="en" docName="zootaxa.4170.2.12.pdf" docOrigin="Zootaxa 4170 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clinterocera heinrichi Krikken, new species" docType="treatment" docVersion="7" lastPageNumber="400" masterDocId="6E4BFF055326105BD509FFD40111100C" masterDocTitle="First records of the cremastocheiline genus Clinterocera Motschulsky, 1858 from the Indonesian island of Sulawesi, with description of a new species (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="400" masterPageNumber="393" pageNumber="395" updateTime="1698387158409" updateUser="plazi">
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<mods:title id="FEEEE0F066248CA2737F9F83510A72E3">First records of the cremastocheiline genus Clinterocera Motschulsky, 1858 from the Indonesian island of Sulawesi, with description of a new species (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
</mods:titleInfo>
<mods:name id="4BE2597D23DE10F68C2DCF50C8AF652E" type="personal">
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<mods:roleTerm id="001C5E2451A44B227CF8441886332CBF">Author</mods:roleTerm>
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<mods:namePart id="E9134F96C9629E5C3B42B4E34830A7C0">Krikken, J.</mods:namePart>
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<mods:title id="3DC4AC150D0EA110E0832724B394F13D">Zootaxa</mods:title>
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<mods:date id="6E3E03BF08C4EA9E267D89B246FEDE32">2016</mods:date>
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<mods:number id="8CB2DB149CFDFE9D7546322F3D02CA69">4170</mods:number>
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<mods:classification id="A0A7983DAD8F7314AE1ECDF5C3FBBDEA">journal article</mods:classification>
<mods:identifier id="C117B7C4A74997BC2DD6ABCB9EA52D39" type="DOI">10.11646/zootaxa.4170.2.12</mods:identifier>
<mods:identifier id="53710F2268370505F3578B7D9CA8D78B" type="GBIF-Dataset">89fc4d5e-607d-4cc4-84fe-58c99b9de56f</mods:identifier>
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<treatment id="9272877D5324105CD59EFF4302931184" ID-DOI="http://doi.org/10.5281/zenodo.3507760" ID-GBIF-Taxon="124552822" ID-Zenodo-Dep="3507760" LSID="urn:lsid:plazi:treatment:9272877D5324105CD59EFF4302931184" httpUri="http://treatment.plazi.org/id/9272877D5324105CD59EFF4302931184" lastPageId="7" lastPageNumber="400" pageId="2" pageNumber="395">
<subSubSection id="52C165E053241059D59EFF43030E10DF" pageId="2" pageNumber="395" type="nomenclature">
<paragraph id="1A64366B53241059D59EFF4303A510BE" blockId="2.[151,692,151,211]" box="[151,692,151,178]" pageId="2" pageNumber="395">
<heading id="412C810753241059D59EFF4303A510BE" bold="true" box="[151,692,151,178]" fontSize="11" level="1" pageId="2" pageNumber="395" reason="1">
<emphasis id="28AFEA7953241059D59EFF4303A510BE" bold="true" box="[151,692,151,178]" pageId="2" pageNumber="395">
<taxonomicName id="DDDB4DE853241059D59EFF43030910BE" authority="Krikken" authorityName="Krikken" box="[151,536,151,178]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="species" species="heinrichi" status="sp. nov.">
<emphasis id="28AFEA7953241059D59EFF4300B510BD" bold="true" box="[151,420,151,177]" italics="true" pageId="2" pageNumber="395">Clinterocera heinrichi</emphasis>
Krikken
</taxonomicName>
,
<taxonomicNameLabel id="339C570253241059D72DFF4C03A510BE" box="[548,692,152,178]" pageId="2" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1A64366B53241059D59EFF6E030E10DF" blockId="2.[151,692,151,211]" box="[151,543,186,211]" pageId="2" pageNumber="395">
<figureCitation id="82E02AEE53241059D59EFF6E001010DF" box="[151,257,186,211]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1911,1934]" captionTargetBox="[161,1419,301,1879]" captionTargetId="figure@3.[151,1435,301,1890]" captionTargetPageId="3" captionText="FIGURES 1 6. Clinterocera Sulawesi. 1 3, habitus oblique; 4 6, forebody dorsofrontal. 1, 4. C. heinrichi, holotype; 2, 5, male, He-He; 3, 6, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257465/files/figure.png" pageId="2" pageNumber="395">Figs. 1, 4</figureCitation>
,
<figureCitation id="82E02AEE53241059D404FF6F005610DF" box="[269,327,187,211]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[170,1429,187,1641]" captionTargetId="figure@4.[151,1435,173,1649]" captionTargetPageId="4" captionText="FIGURES 7 12. Clinterocera Sulawesi. 7 9, habitus dorsal; 10 12, pygidium and adjacent parts, dorsocaudal. 7, 10. C. heinrichi, holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="2" pageNumber="395">7, 10</figureCitation>
,
<figureCitation id="82E02AEE53241059D45AFF6F030A10DF" box="[339,539,186,211]" captionStart-0="FIGURES 13 17" captionStart-1="FIGURES 18 24" captionStartId-0="5.[151,264,1853,1876]" captionStartId-1="6.[151,264,1812,1835]" captionTargetBox-0="[162,1414,208,1818]" captionTargetBox-1="[164,1423,209,1779]" captionTargetId-0="figure@5.[150,1436,193,1833]" captionTargetId-1="figure@6.[150,1436,193,1792]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." captionText-1="FIGURES 18 24. Clinterocera Sulawesi. 18 20, habitus ventral; 21 24, parameres full-face and lateral. 18, 21 22, C. heinrichi, holotype; 23 24, male, He-He. Scale lines 2 mm." httpUri-0="https://zenodo.org/record/257467/files/figure.png" httpUri-1="https://zenodo.org/record/257468/files/figure.png" pageId="2" pageNumber="395">13, 1618, 2122</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52C165E053241059D59EFED6045A11C3" pageId="2" pageNumber="395" type="materials_examined">
<paragraph id="1A64366B53241059D59EFED605A91117" blockId="2.[151,1437,258,2011]" box="[151,1208,258,283]" pageId="2" pageNumber="395">
<emphasis id="28AFEA7953241059D59EFED600911117" bold="true" box="[151,384,258,283]" pageId="2" pageNumber="395">Material examined.</emphasis>
Holotype male from Central Sulawesi, with two original labels (printed):
</paragraph>
<paragraph id="1A64366B53241059D5CEFEF3045A11C3" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
<materialsCitation id="AAB33C3653241059D5CEFEF305B9116F" ID-GBIF-Occurrence="1314772816" collectingDate="1980-04-03" collectingDateMax="1980-04-13" collectingDateMin="1980-04-03" collectorName="M. J. D. Brendell" location="Mt. Tambusisi" pageId="2" pageNumber="395" specimenCount="2" stateProvince="SULAWESI TENGAH" typeStatus="holotype">
<collectingRegion id="D81FF88953241059D5D9FEF300CF1133" box="[208,478,295,319]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="395">SULAWESI TENGAH</collectingRegion>
: \
<location id="1F0460B053241059D4F0FEF203871133" LSID="urn:lsid:plazi:treatment:9272877D5324105CD59EFF4302931184:1F0460B053241059D4F0FEF203871133" box="[505,662,294,319]" name="Mt. Tambusisi" pageId="2" pageNumber="395" stateProvince="SULAWESI TENGAH">Mt.Tambusisi</location>
, 4000, \ 1˚39S-121˚21E. \
<date id="6E6510AB53241059D6E9FEF205601133" box="[992,1137,294,319]" pageId="2" pageNumber="395" value="1980-04-03" valueMax="1980-04-13" valueMin="1980-04-03">
<collectingDate id="7E21E94353241059D6E9FEF205601133" box="[992,1137,294,319]" pageId="2" pageNumber="395" value="1980-04-03" valueMax="1980-04-13" valueMin="1980-04-03">3-13.iv.1980</collectingDate>
</date>
”, “
<collectorName id="B72E53BD53241059D19CFEF3045B1133" box="[1173,1354,294,319]" pageId="2" pageNumber="395">M.J.D. Brendell</collectorName>
\ B.M. 1980-280”; relevant
<typeStatus id="C56088C953241059D488FE9E00F2116F" box="[385,483,330,355]" pageId="2" pageNumber="395" type="holotype">holotype</typeStatus>
label added. Additional specimens (not designated
<typeStatus id="C56088C953241059D126FE9F05B1116F" box="[1071,1184,331,355]" pageId="2" pageNumber="395" type="paratype">paratypes</typeStatus>
)
</materialsCitation>
:
<materialsCitation id="AAB33C3653241059D1BDFE9E0394118B" ID-GBIF-Occurrence="1314772814" collectingDate="1930-12" collectorName="G. Heinrich" elevation="500" location="He" pageId="2" pageNumber="395" specimenCount="1" specimenCount-male="1" stateProvince="North Sulawesi" typeStatus="holotype">
<collectingRegion id="D81FF88953241059D1BDFE9E0477116F" box="[1204,1382,330,355]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="395">North Sulawesi</collectingRegion>
: He-
<location id="1F0460B053241059D59EFEBB01AB118A" LSID="urn:lsid:plazi:treatment:9272877D5324105CD59EFF4302931184:1F0460B053241059D59EFEBB01AB118A" box="[151,186,367,390]" name="He" pageId="2" pageNumber="395" stateProvince="North Sulawesi">He</location>
,
<quantity id="DD239B8E53241059D5C5FEBA001E118B" box="[204,271,366,391]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="395" unit="m" value="500.0">
<elevation id="91F6D15853241059D5C5FEBA001E118B" box="[204,271,366,391]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="395" unit="m" value="500.0">500m</elevation>
</quantity>
,
<date id="6E6510AB53241059D428FEBA0090118B" box="[289,385,366,391]" pageId="2" pageNumber="395" value="1930-12">
<collectingDate id="7E21E94353241059D428FEBA0090118B" box="[289,385,366,391]" pageId="2" pageNumber="395" value="1930-12">xii.1930</collectingDate>
</date>
,
<collectorName id="B72E53BD53241059D49BFEBB030C118B" box="[402,541,366,391]" pageId="2" pageNumber="395">G. Heinrich</collectorName>
,
<specimenCount id="0CDDFDE253241059D726FEBB0394118B" box="[559,645,366,391]" pageId="2" pageNumber="395" type="male">1 male</specimenCount>
</materialsCitation>
;
<materialsCitation id="AAB33C3653241059D79EFEBA045A11C3" ID-GBIF-Occurrence="1314772815" collectingDate="1932-01-01" collectingDateMax="1932-01-15" collectingDateMin="1932-01-01" collectorName="G. Heinrich &amp; The Natural History Museum &amp; London &amp; The &amp; Heinrich" country="United Kingdom" elevation="1500" location="Mt Tangke Salokko" pageId="2" pageNumber="395" specimenCount="1" specimenCount-female="1" stateProvince="Southeast Sulawesi" typeStatus="holotype">
<collectingRegion id="D81FF88953241059D79EFEBA026A118B" box="[663,891,366,391]" country="Indonesia" name="Sulawesi Tenggara" pageId="2" pageNumber="395">Southeast Sulawesi</collectingRegion>
:
<location id="1F0460B053241059D684FEBB0568118B" LSID="urn:lsid:plazi:treatment:9272877D5324105CD59EFF4302931184:1F0460B053241059D684FEBB0568118B" box="[909,1145,366,391]" country="United Kingdom" name="Mt Tangke Salokko" pageId="2" pageNumber="395" stateProvince="Southeast Sulawesi">Mt Tangke Salokko</location>
,
<quantity id="DD239B8E53241059D185FEBA05CD118B" box="[1164,1244,366,391]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="395" unit="m" value="1500.0">
<elevation id="91F6D15853241059D185FEBA05CD118B" box="[1164,1244,366,391]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="395" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="6E6510AB53241059D1E7FEBA0460118B" box="[1262,1393,366,391]" pageId="2" pageNumber="395" value="1932-01-01" valueMax="1932-01-15" valueMin="1932-01-01">
<collectingDate id="7E21E94353241059D1E7FEBA0460118B" box="[1262,1393,366,391]" pageId="2" pageNumber="395" value="1932-01-01" valueMax="1932-01-15" valueMin="1932-01-01">1-15.i.1932</collectingDate>
</date>
,
<collectorName id="B72E53BD53241059D08CFEBB01ED11A7" pageId="2" pageNumber="395">G. Heinrich</collectorName>
,
<specimenCount id="0CDDFDE253241059D40EFE47007911A7" box="[263,360,402,427]" pageId="2" pageNumber="395" type="female">1 female</specimenCount>
(all three in
<collectorName id="B72E53BD53241059D4F1FE46025711A6" box="[504,838,402,427]" pageId="2" pageNumber="395">The Natural History Museum</collectorName>
,
<collectorName id="B72E53BD53241059D658FE4602BA11A7" box="[849,939,402,427]" pageId="2" pageNumber="395">London</collectorName>
,
<collectingCountry id="62CC76FB53241059D6BEFE46056811A7" box="[951,1145,402,427]" name="United Kingdom" pageId="2" pageNumber="395">United Kingdom</collectingCountry>
).
<collectorName id="B72E53BD53241059D182FE4605A611A7" box="[1163,1207,402,427]" pageId="2" pageNumber="395">The</collectorName>
localities of the two
<collectorName id="B72E53BD53241059D59EFE6201E811C3" box="[151,249,438,463]" pageId="2" pageNumber="395">Heinrich</collectorName>
specimens are indicated on maps in books on the collectors
<collectingRegion id="D81FF88953241059D6AEFE62051D11C3" box="[935,1036,438,463]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="395">Sulawesi</collectingRegion>
expedition (
<bibRefCitation id="7E4A4B9A53241059D191FE62042E11C3" author="Heinrich" box="[1176,1343,438,463]" pageId="2" pageNumber="395" refString="Heinrich, G. (1943) Celebes. Seltsame Jagd durch seltsames Land. Reimer, Berlin, 198 pp." type="book" year="1943">
<collectorName id="B72E53BD53241059D191FE6205ED11C3" box="[1176,1276,438,463]" pageId="2" pageNumber="395">Heinrich</collectorName>
1943
</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52C165E053241059D5CEFE0E04741746" pageId="2" pageNumber="395" type="description">
<paragraph id="1A64366B53241059D5CEFE0E0032128F" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
<emphasis id="28AFEA7953241059D5CEFE0E033411FF" bold="true" box="[199,549,474,499]" pageId="2" pageNumber="395">
Description (
<typeStatus id="C56088C953241059D46BFE0E00DD11FF" box="[354,460,474,499]" pageId="2" pageNumber="395" type="holotype">holotype</typeStatus>
, male).
</emphasis>
Habitus (Fig, 1) robust, length
<quantity id="DD239B8E53241059D69DFE0F02F711FE" box="[916,998,475,499]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="395" unit="mm" value="22.0">22 mm</quantity>
(with head bent down). Black, elytral base with small red marking; some parts shiny, others more matt; also with symmetric markings of distinct pale khaki tomentum, mainly on sides of pronotum and elytra, and on scutellum; also limited tomentum on ventral side and legs (pectoral parts, sides of abdomen, and femora). Pilosity abundant but inconspicuous, minute, brown, stubble-like.
</paragraph>
<paragraph id="1A64366B53241059D5CEFD5A025D131F" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Head surface (
<figureCitation id="82E02AEE53241059D465FD5A00AD12AB" box="[364,444,654,679]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1911,1934]" captionTargetBox="[161,1419,301,1879]" captionTargetId="figure@3.[151,1435,301,1890]" captionTargetPageId="3" captionText="FIGURES 1 6. Clinterocera Sulawesi. 1 3, habitus oblique; 4 6, forebody dorsofrontal. 1, 4. C. heinrichi, holotype; 2, 5, male, He-He; 3, 6, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257465/files/figure.png" pageId="2" pageNumber="395">Figs. 4</figureCitation>
,
<figureCitation id="82E02AEE53241059D4C0FD5B00F712AB" box="[457,486,655,679]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1853,1876]" captionTargetBox="[162,1414,208,1818]" captionTargetId="figure@5.[150,1436,193,1833]" captionTargetPageId="5" captionText="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="2" pageNumber="395">13</figureCitation>
) largely shiny, distinctly evenly convex, apical margin not reflexed (not concave in profile), and sides steep. Punctation dense, minute, superficially pustulate behind clypeal apex, rapidly becoming more distinct and annulate-striolate toward other parts of head. Eye-canthi with light tomentum, frons with indistinct dark band between eyes (looks like thin tomentum).
</paragraph>
<paragraph id="1A64366B53241059D5CEFCCB033A1373" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Pronotum (
<figureCitation id="82E02AEE53241059D44EFCCA009A133B" box="[327,395,798,823]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[170,1429,187,1641]" captionTargetId="figure@4.[151,1435,173,1649]" captionTargetPageId="4" captionText="FIGURES 7 12. Clinterocera Sulawesi. 7 9, habitus dorsal; 10 12, pygidium and adjacent parts, dorsocaudal. 7, 10. C. heinrichi, holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="2" pageNumber="395">Fig. 7</figureCitation>
) with dorsal outline virtually fully rounded, surface black, somewhat matt, lateral margins with light tomentum. Pronotal surface abundantly to densely arcuate-annulate-striolate. Scutellum tomentous, somewhat vaguely arcuate-striolate.
</paragraph>
<paragraph id="1A64366B53241059D5CEFC5E02C7145B" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Elytral surface (
<figureCitation id="82E02AEE53241059D474FC5E00D313AF" box="[381,450,906,931]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[170,1429,187,1641]" captionTargetId="figure@4.[151,1435,173,1649]" captionTargetPageId="4" captionText="FIGURES 7 12. Clinterocera Sulawesi. 7 9, habitus dorsal; 10 12, pygidium and adjacent parts, dorsocaudal. 7, 10. C. heinrichi, holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="2" pageNumber="395">Fig. 7</figureCitation>
) largely black, disc generally evenly deplanate, with distinct indication of longitudinal juxtasutural striola at short distance along entire suture; humeral umbone more shiny than most of elytral surface, between umbone and scutellum a small red marking is present; elytral sides with pale tomentous band over most of lateral and caudal declivities. Elytral disc (
<figureCitation id="82E02AEE53241059D771FC2203DB1403" box="[632,714,1014,1039]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1853,1876]" captionTargetBox="[162,1414,208,1818]" captionTargetId="figure@5.[150,1436,193,1833]" captionTargetPageId="5" captionText="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="2" pageNumber="395">Fig. 17</figureCitation>
) with abundant to dense, mainly elliptic arcuate striolae, denser and coarser near lateral tomentous band, apparently finer and more annular in that band. Posthumeral elytral emargination distinct; apicosutural elytral angle (
<figureCitation id="82E02AEE53241059D7B4FBEA021F145B" box="[701,782,1086,1111]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[170,1429,187,1641]" captionTargetId="figure@4.[151,1435,173,1649]" captionTargetPageId="4" captionText="FIGURES 7 12. Clinterocera Sulawesi. 7 9, habitus dorsal; 10 12, pygidium and adjacent parts, dorsocaudal. 7, 10. C. heinrichi, holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="2" pageNumber="395">Fig. 10</figureCitation>
) shortly rounded.
</paragraph>
<paragraph id="1A64366B53241059D5CEFBB6009A155E" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Expanded mentum (
<figureCitation id="82E02AEE53241059D4BBFBB603191477" box="[434,520,1122,1147]" captionStart="FIGURES 18 24" captionStartId="6.[151,264,1812,1835]" captionTargetBox="[164,1423,209,1779]" captionTargetId="figure@6.[150,1436,193,1792]" captionTargetPageId="6" captionText="FIGURES 18 24. Clinterocera Sulawesi. 18 20, habitus ventral; 21 24, parameres full-face and lateral. 18, 21 22, C. heinrichi, holotype; 23 24, male, He-He. Scale lines 2 mm." httpUri="https://zenodo.org/record/257468/files/figure.png" pageId="2" pageNumber="395">Fig. 18</figureCitation>
) with exposed side minutely, evenly, densely, superficially punctate. Concave upper side of expanded antennal scapus finely rugulate-striolate. Preprosternal apophysis robust, obliquely directed forward, tapering, apex blunt (in profile). Tomentous markings laterally, almost entire underside of body black, largely shiny. Prosternal sides densely striolate-hemipunctate, each puncture with minute bristle. Mesoventral and metaventral surface densely set with distinct arcuate-striolate (hemi)punctures; metaventral disc deplanate, with distinctly impressed, narrow midline, striolate punctures less dense there, smaller; lateral metaventral elements with some tomentum.
</paragraph>
<paragraph id="1A64366B53241059D5CEFA8A05B6160B" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Abdominal underside (
<figureCitation id="82E02AEE53241059D4C3FA8A030A157B" box="[458,539,1374,1399]" captionStart="FIGURES 18 24" captionStartId="6.[151,264,1812,1835]" captionTargetBox="[164,1423,209,1779]" captionTargetId="figure@6.[150,1436,193,1792]" captionTargetPageId="6" captionText="FIGURES 18 24. Clinterocera Sulawesi. 18 20, habitus ventral; 21 24, parameres full-face and lateral. 18, 21 22, C. heinrichi, holotype; 23 24, male, He-He. Scale lines 2 mm." httpUri="https://zenodo.org/record/257468/files/figure.png" pageId="2" pageNumber="395">Fig. 18</figureCitation>
) largely shiny black (medially somewhat brown), medially concave (distinct in profile); ventrites with (medially) finely arcuate-striolate and (laterally) denser annulate-striolate punctation. Propygidium and abdominal ventrites laterally (near elytral edge) tomentous; pygidium (
<figureCitation id="82E02AEE53241059D1A4FA72041E15B3" box="[1197,1295,1446,1471]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[170,1429,187,1641]" captionTargetId="figure@4.[151,1435,173,1649]" captionTargetPageId="4" captionText="FIGURES 7 12. Clinterocera Sulawesi. 7 9, habitus dorsal; 10 12, pygidium and adjacent parts, dorsocaudal. 7, 10. C. heinrichi, holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm." httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="2" pageNumber="395">Figs. 10</figureCitation>
,
<figureCitation id="82E02AEE53241059D029FA73042C15B3" box="[1312,1341,1447,1471]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1853,1876]" captionTargetBox="[162,1414,208,1818]" captionTargetId="figure@5.[150,1436,193,1833]" captionTargetPageId="5" captionText="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="2" pageNumber="395">13</figureCitation>
) evenly convex, lightly tomentous, generally with strong, crowded annulate-striolate punctation; propygidial spiracles conically produced. Parameres robustly bilobate, interparameral split widened (
<figureCitation id="82E02AEE53241059D118FA3A058A160B" box="[1041,1179,1518,1543]" captionStart="FIGURES 18 24" captionStartId="6.[151,264,1812,1835]" captionTargetBox="[164,1423,209,1779]" captionTargetId="figure@6.[150,1436,193,1792]" captionTargetPageId="6" captionText="FIGURES 18 24. Clinterocera Sulawesi. 18 20, habitus ventral; 21 24, parameres full-face and lateral. 18, 21 22, C. heinrichi, holotype; 23 24, male, He-He. Scale lines 2 mm." httpUri="https://zenodo.org/record/257468/files/figure.png" pageId="2" pageNumber="395">Figs. 2122</figureCitation>
).
</paragraph>
<paragraph id="1A64366B53241059D5CEF9C60534170F" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Protibia (
<figureCitation id="82E02AEE53241059D43BF9C600961627" box="[306,391,1554,1579]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1853,1876]" captionTargetBox="[162,1414,208,1818]" captionTargetId="figure@5.[150,1436,193,1833]" captionTargetPageId="5" captionText="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="2" pageNumber="395">Fig. 16</figureCitation>
) with short, obtuse apico-external denticle and 1 obtuse, more proximal external denticle; underside with longitudinal ridge ending in robust, tapering, downward projection; terminal spur short; most of upper side densely punctate, more or less striolate. Mesotibiae and metatibiae with distinct external protrusion, their apex with 3 distinct protrusions,
<emphasis id="28AFEA7953241059D754F9AB0397169A" box="[605,646,1663,1686]" italics="true" pageId="2" pageNumber="395">i.e.,</emphasis>
2 simply acute and 1 internal dilated extension supporting pair of tapering spurs; tibial surface densely to crowdedly striolate-punctate(-rugulate), external side tomentous. All femora equally striolate-punctate; posterior side of mesofemora and metafemora with tomentum. Tarsi all with 4 short densely, finely punctate tarsomeres, protarsi distinctly thickset; claws short.
</paragraph>
<paragraph id="1A64366B53241059D5CEF8DA04741746" blockId="2.[151,1437,258,2011]" pageId="2" pageNumber="395">
Head maximum width (tips of eye-canthi) 4.4 mm. Pronotum median length 5.1 mm, maximum width 6.3 mm. Elytra maximum (longitudinal) length approximately
<quantity id="DD239B8E53241059D7FBF8E702521746" box="[754,835,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="395" unit="mm" value="13.0">13 mm</quantity>
, combined maximum width (at humeri) 9.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="52C165E05324105CD5CEF882036D114C" lastPageId="7" lastPageNumber="400" pageId="2" pageNumber="395" type="diagnosis">
<paragraph id="1A64366B53241058D5CEF88202A010F4" blockId="2.[151,1437,258,2011]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="396" pageId="2" pageNumber="395">
<emphasis id="28AFEA7953241059D5CEF88203AC1763" bold="true" box="[199,701,1878,1903]" pageId="2" pageNumber="395">Diagnosis, variation, sexual dimorphism</emphasis>
(see all comparative figures). The distribution of tomentous markings on the
<typeStatus id="C56088C953241059D45FF8AE00A9179F" box="[342,440,1914,1939]" pageId="2" pageNumber="395" type="holotype">holotype</typeStatus>
along the sides of the pronotum and elytra serves to tentatively distinguish this species from other known
<taxonomicName id="DDDB4DE853241059D461F84A00E717BA" box="[360,502,1950,1974]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA7953241059D461F84A00E717BA" box="[360,502,1950,1974]" italics="true" pageId="2" pageNumber="395">Clinterocera</emphasis>
</taxonomicName>
,
<emphasis id="28AFEA7953241059D70DF84B033C17BA" box="[516,557,1951,1974]" italics="true" pageId="2" pageNumber="395">i.e.,</emphasis>
the taxa reported from the western mainland and nearby islands. Males have a long projection on the protibial underside, near the insertion of the compacted tarsus. The red markings, if present at all, remain small and limited to the elytra, contrary to the more conspicuously black/red symmetric pattern of several continental species in the
<taxonomicName id="DDDB4DE853251058D722FF6803BC10D8" box="[555,685,188,212]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="jucunda">
<emphasis id="28AFEA7953251058D722FF68032F10DF" box="[555,574,188,211]" italics="true" pageId="3" pageNumber="396">C</emphasis>
.
<emphasis id="28AFEA7953251058D75BFF6803BC10D8" box="[594,685,188,212]" italics="true" pageId="3" pageNumber="396">jucunda</emphasis>
</taxonomicName>
group. Elytral disc flat, not particularly modified (
<emphasis id="28AFEA7953251058D01DFF69042C10D8" box="[1300,1341,189,212]" italics="true" pageId="3" pageNumber="396">i.e.,</emphasis>
lacking longitudinal costae, heavy punctation, and white velutinous markings).
</paragraph>
<caption id="4EA466E353251058D59EF8A303AF17A0" httpUri="https://zenodo.org/record/257465/files/figure.png" pageId="3" pageNumber="396" targetBox="[161,1419,301,1879]" targetPageId="3">
<paragraph id="1A64366B53251058D59EF8A303AF17A0" blockId="3.[151,1436,1911,1964]" pageId="3" pageNumber="396">
<emphasis id="28AFEA7953251058D59EF8A3002D1781" bold="true" box="[151,316,1911,1934]" pageId="3" pageNumber="396">FIGURES 16.</emphasis>
<taxonomicName id="DDDB4DE853251058D44CF8AC00D71782" box="[325,454,1912,1934]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA7953251058D44CF8AC00D71782" box="[325,454,1912,1934]" italics="true" pageId="3" pageNumber="396">Clinterocera</emphasis>
</taxonomicName>
Sulawesi. 13, habitus oblique; 46, forebody dorsofrontal. 1, 4.
<emphasis id="28AFEA7953251058D170F8AD05EA1782" box="[1145,1275,1912,1934]" italics="true" pageId="3" pageNumber="396">
<taxonomicName id="DDDB4DE853251058D170F8AD05E61782" box="[1145,1271,1912,1934]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="396" phylum="Arthropoda" rank="species" species="heinrichi">C. heinrichi</taxonomicName>
,
</emphasis>
holotype; 2, 5, male, He-He; 3, 6, female, Salokko. Scale lines 5 mm.
</paragraph>
</caption>
<caption id="4EA466E35322105FD59EF95402C616B9" httpUri="https://zenodo.org/record/257466/files/figure.png" pageId="4" pageNumber="397" targetBox="[170,1429,187,1641]" targetPageId="4">
<paragraph id="1A64366B5322105FD59EF95402C616B9" blockId="4.[151,1436,1664,1717]" pageId="4" pageNumber="397">
<emphasis id="28AFEA795322105FD59EF954005A169A" bold="true" box="[151,331,1664,1687]" pageId="4" pageNumber="397">FIGURES 712.</emphasis>
<taxonomicName id="DDDB4DE85322105FD45CF95500C7169B" box="[341,470,1665,1687]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA795322105FD45CF95500C7169B" box="[341,470,1665,1687]" italics="true" pageId="4" pageNumber="397">Clinterocera</emphasis>
</taxonomicName>
Sulawesi. 79, habitus dorsal; 1012, pygidium and adjacent parts, dorsocaudal. 7, 10.
<emphasis id="28AFEA795322105FD08DF95601E816B9" italics="true" pageId="4" pageNumber="397">
<taxonomicName id="DDDB4DE85322105FD08DF95601E416B9" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="heinrichi">C. heinrichi</taxonomicName>
,
</emphasis>
holotype; 8, 11, male, He-He; 9, 12, female, Salokko. Scale lines 5 mm.
</paragraph>
</caption>
<paragraph id="1A64366B5322105FD5CEF90800AA17FD" blockId="4.[151,1437,1756,2034]" pageId="4" pageNumber="397">
Head largely shiny. Pygidium evenly strongly rounded (not impressed over midline), crowdedly punctate. Propygidial spiracles conically raised. Characteristic of the
<typeStatus id="C56088C95322105FD64FF8D502B91716" box="[838,936,1793,1818]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
is the surface of the clypeus, which slopes evenly down to the apical border, not being distinctly reflexed (
<emphasis id="28AFEA795322105FD671F8F202B31731" box="[888,930,1830,1853]" italics="true" pageId="4" pageNumber="397">i.e.,</emphasis>
margin hardly concave in lateral view); the male from the Minahassa (
<emphasis id="28AFEA795322105FD4CFF89E00FE176D" box="[454,495,1866,1889]" italics="true" pageId="4" pageNumber="397">i.e.,</emphasis>
<collectingRegion id="D81FF8895322105FD4FEF89D03B6176E" box="[503,679,1865,1890]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="397">North Sulawesi</collectingRegion>
) has a different, more concave clypeal margin. This male lacks the abundant tomentum, and its microsculpture is different from the
<typeStatus id="C56088C95322105FD6A3F8B9051E178A" box="[938,1039,1901,1926]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
, being coarser on the head. The parameres are slightly different, as in the shape of the interparameral split, which is more widened in the
<typeStatus id="C56088C95322105FD03AF845048917A6" box="[1331,1432,1937,1962]" pageId="4" pageNumber="397" type="holotype">holotype</typeStatus>
, narrower in the Minahassa male. The body length of both males reaches approximately
<quantity id="DD239B8E5322105FD187F86105F117C1" box="[1166,1248,1973,1997]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="397" unit="mm" value="22.0">22 mm</quantity>
(with head bent down, in protective state).
</paragraph>
<caption id="4EA466E35323105ED59EF8E90524177F" httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="5" pageNumber="398" targetBox="[162,1414,208,1818]" targetPageId="5">
<paragraph id="1A64366B5323105ED59EF8E90524177F" blockId="5.[151,1437,1853,1907]" pageId="5" pageNumber="398">
<emphasis id="28AFEA795323105ED59EF8E90045175F" bold="true" box="[151,340,1853,1876]" pageId="5" pageNumber="398">FIGURES 1317.</emphasis>
<taxonomicName id="DDDB4DE85323105ED452F8EA00CD1758" box="[347,476,1854,1876]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA795323105ED452F8EA00CD1758" box="[347,476,1854,1876]" italics="true" pageId="5" pageNumber="398">Clinterocera</emphasis>
</taxonomicName>
Sulawesi. 1315, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 1617.
<emphasis id="28AFEA795323105ED400F88A0099177F" box="[265,392,1885,1907]" italics="true" pageId="5" pageNumber="398">
<taxonomicName id="DDDB4DE85323105ED400F88A0095177F" box="[265,388,1885,1907]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="heinrichi">C. heinrichi</taxonomicName>
,
</emphasis>
holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm.
</paragraph>
</caption>
<caption id="4EA466E35320105DD59EF8C003E21746" httpUri="https://zenodo.org/record/257468/files/figure.png" pageId="6" pageNumber="399" targetBox="[164,1423,209,1779]" targetPageId="6">
<paragraph id="1A64366B5320105DD59EF8C003E21746" blockId="6.[151,1436,1812,1866]" pageId="6" pageNumber="399">
<emphasis id="28AFEA795320105DD59EF8C000481726" bold="true" box="[151,345,1812,1835]" pageId="6" pageNumber="399">FIGURES 1824.</emphasis>
<taxonomicName id="DDDB4DE85320105DD46CF8C100F71727" box="[357,486,1813,1835]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA795320105DD46CF8C100F71727" box="[357,486,1813,1835]" italics="true" pageId="6" pageNumber="399">Clinterocera</emphasis>
</taxonomicName>
Sulawesi. 1820, habitus ventral; 2124, parameres full-face and lateral. 18, 2122,
<emphasis id="28AFEA795320105DD08DF8C201E81746" italics="true" pageId="6" pageNumber="399">
<taxonomicName id="DDDB4DE85320105DD08DF8C201E41746" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="heinrichi">C. heinrichi</taxonomicName>
,
</emphasis>
holotype; 2324, male, He-He. Scale lines 2 mm.
</paragraph>
</caption>
<paragraph id="1A64366B5320105DD5CEF8A2048C17DB" blockId="6.[151,1437,1910,2007]" pageId="6" pageNumber="399">
The female from Southeast Sulawesi (total body length reaches approximately
<quantity id="DD239B8E5320105DD152F8A305BE1783" box="[1115,1199,1911,1935]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="399" unit="mm" value="24.0">24 mm</quantity>
) is virtually entirely black, but agrees with the holotype in its clypeal shape and several details. The ridge on the underside of the protibia is well developed, terminally slightly protuberant, but lacking the long, downward projection of the males.
</paragraph>
<paragraph id="1A64366B5321105CD59EFF43000110FB" blockId="7.[151,1436,151,392]" pageId="7" pageNumber="400">
The abdominal venter is not concave—as usual in most female
<taxonomicName id="DDDB4DE85321105CD660FF4302F710BC" box="[873,998,151,176]" class="Insecta" family="Cetoniinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="400" phylum="Arthropoda" rank="family">Cetoniinae</taxonomicName>
. Pygidium moderately, evenly convex, surface plane more upward, crowdedly punctate (compare
<figureCitation id="82E02AEE5321105CD645FF6802C910D9" box="[844,984,188,213]" captionStart="FIGURES 13 17" captionStartId="5.[151,264,1853,1876]" captionTargetBox="[162,1414,208,1818]" captionTargetId="figure@5.[150,1436,193,1833]" captionTargetPageId="5" captionText="FIGURES 13 17. Clinterocera Sulawesi. 13 15, habitus lateral; 16, protibia lateral; 17, section from suture over left elytron. 13, 16 17. C. heinrichi, holotype; 14, male, He-He; 15, female, Salokko. Scale lines 2 mm." httpUri="https://zenodo.org/record/257467/files/figure.png" pageId="7" pageNumber="400">Figs. 1315</figureCitation>
). Some parts show thin indications of tomentum.
</paragraph>
<paragraph id="1A64366B5321105CD5CEFED0036D114C" blockId="7.[151,1436,151,392]" pageId="7" pageNumber="400">
The large
<taxonomicName id="DDDB4DE85321105CD43EFED000D51110" box="[311,452,260,284]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="400" phylum="Arthropoda" rank="species" species="cyprilrusi">
<emphasis id="28AFEA795321105CD43EFED000D51110" box="[311,452,260,284]" italics="true" pageId="7" pageNumber="400">C. cyprilrusi</emphasis>
</taxonomicName>
from Borneo mentioned in the Introduction belongs to a different group of
<taxonomicName id="DDDB4DE85321105CD007FED0048D1110" box="[1294,1436,260,284]" class="Insecta" family="Cetoniidae" genus="Clinterocera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="400" phylum="Arthropoda" rank="genus">
<emphasis id="28AFEA795321105CD007FED0048D1110" box="[1294,1436,260,284]" italics="true" pageId="7" pageNumber="400">Clinterocera</emphasis>
</taxonomicName>
species (Legrand &amp; Chew Keas
<bibRefCitation id="7E4A4B9A5321105CD709FEFC0361114C" author="Foo" box="[512,624,296,320]" pageId="7" pageNumber="400" refString="Legrand, J. - P. &amp; Chew Kea Foo, S. (2010) Les Cetoniinae du Sabah. Collection ex Natura (Magellanes), 1, 1 - 127." type="journal article" year="2010">Foo 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="52C165E05321105CD5CEFE9802931184" pageId="7" pageNumber="400" type="etymology">
<paragraph id="1A64366B5321105CD5CEFE9802931184" blockId="7.[151,1436,151,392]" pageId="7" pageNumber="400">
<emphasis id="28AFEA795321105CD5CEFE9800B41169" bold="true" box="[199,421,332,357]" pageId="7" pageNumber="400">Derivatio nominis.</emphasis>
Named after the classical natural history explorer and specialist in
<taxonomicName id="DDDB4DE85321105CD1A1FE9801E81184" authority="Gerd Heinrich" authorityName="Gerd Heinrich" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="400" phylum="Arthropoda" rank="family">Ichneumonidae, Gerd Heinrich</taxonomicName>
(
<date id="6E6510AB5321105CD401FEBB00971184" box="[264,390,367,392]" pageId="7" pageNumber="400" value="1896" valueMax="1984">1896-1984</date>
), collector of two of the
<collectingRegion id="D81FF8895321105CD793FEBB03EE1184" box="[666,767,367,392]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="400">Sulawesi</collectingRegion>
specimens.
</paragraph>
</subSubSection>
</treatment>
</document>