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<document id="E2C38AC397AA1EE2CFE06259C56BFAB0" ID-CLB-Dataset="9252" ID-DOI="10.11646/zootaxa.4895.3.1" ID-GBIF-Dataset="a5e5d5ad-9d52-48cd-9bf9-ca28a2a7af4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4326650" ID-ZooBank="C6F85190-D79E-429E-9B5B-FD0E54AE96BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608109046279" checkinUser="plazi" docAuthor="Liow, Lee Hsiang &amp; Gordon, Dennis P." docDate="2020" docId="3C7D4825FF80E04B21D5499E51659F0A" docLanguage="en" docName="zootaxa.4895.3.1.pdf" docOrigin="Zootaxa 4895 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adeonellopsis wassi Liow &amp; Gordon 2020, n. sp." docType="treatment" docUuid="B3640502-4E99-4383-9288-AA77A5D70556" docUuidSource="ZooBank" docVersion="7" lastPageNumber="316" masterDocId="C044305DFF8EE0442142490952599A3B" masterDocTitle="New species of Adeonellopsis (Bryozoa: Adeonidae) from southern Zealandia and the western Tasman Sea" masterLastPageNumber="331" masterPageNumber="301" pageNumber="315" updateTime="1698847604300" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AF06EA871ECC1C18ACCFFFDF066004CF">New species of Adeonellopsis (Bryozoa: Adeonidae) from southern Zealandia and the western Tasman Sea</mods:title>
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<mods:namePart id="922C722B7E0C048EB8BF1AE611447286">Liow, Lee Hsiang</mods:namePart>
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<mods:affiliation id="1497AC69D05FF7232FF4C05D664C6AFE">Natural History Museum and Centre for Ecological and Evolutionary Synthesis, University of Oslo, Oslo, Norway. l. h. liow @ ibv. uio. no; https: // orcid. org / 0000 - 0002 - 3732 - 6069</mods:affiliation>
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<mods:namePart id="BDE90D18224B05B23D5996221CD18AE9">Gordon, Dennis P.</mods:namePart>
<mods:affiliation id="8BB688737120ABCE37C865EC31DD3341">National Institute of Water &amp; Atmospheric Research, Private Bag 14901, Kilbirnie, Wellington 6022, New Zealand. dennis. gordon @ niwa. co. nz; http: // orcid. org / 0000 - 0001 - 9398 - 996 X</mods:affiliation>
<mods:nameIdentifier id="81EE944A68C6AD044227E9E51EE8118B" type="email">dennis.gordon@niwa.co.nz</mods:nameIdentifier>
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<mods:date id="AE41470FC58D9F45A793EA7F14A9BC32">2020</mods:date>
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<mods:number id="113355BF9E8947FCC4F92A12761D3FCB">2020-12-16</mods:number>
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<treatment id="3C7D4825FF80E04B21D5499E51659F0A" ID-DOI="http://doi.org/10.5281/zenodo.4326644" ID-GBIF-Taxon="176232380" ID-Zenodo-Dep="4326644" LSID="urn:lsid:zoobank.org:act:B3640502-4E99-4383-9288-AA77A5D70556" httpUri="http://treatment.plazi.org/id/3C7D4825FF80E04B21D5499E51659F0A" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
<subSubSection id="FCCEAAB8FF80E04A21D5499E53889A8A" box="[151,465,151,178]" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="B46BF933FF80E04A21D5499E53889A8A" blockId="14.[151,516,151,213]" box="[151,465,151,178]" pageId="14" pageNumber="315">
<heading id="EF234E5FFF80E04A21D5499E53889A8A" bold="true" box="[151,465,151,178]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<emphasis id="86A02521FF80E04A21D5499E53889A8A" bold="true" box="[151,465,151,178]" pageId="14" pageNumber="315">
<taxonomicName id="73D482B0FF80E04A21D5499E53DE9A8A" authority="Liow &amp; Gordon, 2020" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[151,391,151,178]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="315" phylum="Bryozoa" rank="species" species="wassi" status="sp. nov.">
<emphasis id="86A02521FF80E04A21D5499E53DE9A8A" bold="true" box="[151,391,151,178]" italics="true" pageId="14" pageNumber="315">Adeonellopsis wassi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9D93985AFF80E04A20CD499E53889A8A" box="[399,465,151,177]" pageId="14" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF80E04A21D549B2505D9AEE" box="[151,516,187,213]" pageId="14" pageNumber="315" type="description">
<paragraph id="B46BF933FF80E04A21D549B2505D9AEE" blockId="14.[151,516,151,213]" box="[151,516,187,213]" pageId="14" pageNumber="315">
(
<figureCitation id="2CEFE5B6FF80E04A21DD49B252A09AEE" box="[159,249,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1667,1691]" captionTargetBox="[160,1426,1079,1648]" captionTargetId="figure-349@4.[159,1427,1079,1649]" captionTargetPageId="4" captionText="FIGURE 3. A, B. Adeonellopsis macewindui n. sp.: A. Holotype, NIWA 146070; B. NIWA 146098. C. Adeonellopsis tasmanensis n. sp., composite of three fragments, all NIWA 144892. D. Adeonellopsis periculosa n. sp., holotype, NIWA 146071. E. Adeonellopsis wassi n. sp., composite of three fragments, all NIWA 144898. F, G. Adeonellopsis gemina n. sp.: F. holotype, NIWA 146095; G. Two specimens, NIWA 146073. H. Adeonellopsis minor n. sp., composite of four fragments, NIWA 146152. I. Adeonellopsis gracilis n. sp., composite of two fragments, holotype (larger specimen), NIWA 92745; smaller specimen (separated by thin gap), NIWA 146099. Specimens AI at same scale; scalebar 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4326656" httpUri="https://zenodo.org/record/4326656/files/figure.png" pageId="14" pageNumber="315">Figs 3E</figureCitation>
,
<figureCitation id="2CEFE5B6FF80E04A204749B2534B9AEE" box="[261,274,187,213]" captionStart="FIGURE 9" captionStartId="13.[151,250,1622,1646]" captionTargetBox="[243,1344,420,1595]" captionTargetId="figure-142@13.[239,1347,417,1599]" captionTargetPageId="13" captionText="FIGURE 9. Adeonellopsis tasmanensis n. sp. (NIWA 144892): A, B. Two gonozooids with varied numbers and orientations of adventitious avicularia. Adeonellopsis wassi n. sp. (NIWA 144899): C. Part of stem near axial bifurcation; the black arrow indicates a zooid (gonozooid?) with an unusually large spiramen. D.Autozooids with paired and single suboral avicularia having varied orientations. Scalebars: A, B, 300 μm; C, 2 mm; D, 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4326668" httpUri="https://zenodo.org/record/4326668/files/figure.png" pageId="14" pageNumber="315">9</figureCitation>
CD, 10;
<tableCitation id="F956CC88FF80E04A20C249B253879AEE" box="[384,478,187,213]" captionStart="TABLE 1" captionStartId="9.[151,240,462,488]" captionText="TABLE 1. Tabulation of comparative zooidal metric data (see Methods) for Adeonellopsis species described in the text. The status of the putative gonozooid in A. wassi n. sp., A. gemina n. sp., A. minor n. sp. and A. gracilis n. sp. is uncertain (see text). An em dash signifies that the character is not present. Data for each character are given as the range, plus the number of measurements (N), mean and standard deviation in square brackets. Measurements are based on one or more fragments (probably from the same colony) having the same registration number." pageId="14" pageNumber="315">Tables 1</tableCitation>
,
<tableCitation id="F956CC88FF80E04A20AE49B253A29AEE" box="[492,507,187,213]" captionStart="TABLE 2" captionStartId="10.[151,239,664,690]" captionText="TABLE 2. Tabular comparison of key morphological characters for Adeonellopsis species described in the text. The status of the putative gonozooid in A. wassi n. sp., A. gemina n. sp., A. minor n. sp. and A. gracilis n. sp. is uncertain (see text)." pageId="14" pageNumber="315">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF80E04A21D5480A50FB9B7A" pageId="14" pageNumber="315" type="etymology">
<paragraph id="B46BF933FF80E04A21D5480A50FB9B7A" blockId="14.[151,1437,259,610]" pageId="14" pageNumber="315">
<emphasis id="86A02521FF80E04A21D5480A53479B26" bold="true" box="[151,286,259,285]" pageId="14" pageNumber="315">Etymology.</emphasis>
Honorific for retired bryozoologist Dr Robin E. Wass to acknowledge his contributions to knowledge of Australian Paleozoic and Recent bryozoans.
</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF80E04A2185484253AE9B96" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="B46BF933FF80E04A2185484253AE9B96" blockId="14.[151,1437,259,610]" pageId="14" pageNumber="315">
<emphasis id="86A02521FF80E04A2185484253E89B5E" bold="true" box="[199,433,331,357]" pageId="14" pageNumber="315">Material examined.</emphasis>
<materialsCitation id="04BCF36EFF80E04A20FA484250DD9BB2" ID-GBIF-Occurrence="3001059336" collectionCode="NIWA" collectorName="Australian Museum U. &amp; Stn U" elevation="198" latitude="-34.185" location="Sea" longLatPrecision="6" longitude="151.4333" pageId="14" pageNumber="315" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<emphasis id="86A02521FF80E04A20FA484250719B5E" box="[440,552,331,357]" italics="true" pageId="14" pageNumber="315">
<typeStatus id="6B6F4791FF80E04A20FA4842507B9B5E" box="[440,546,331,357]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectorName id="19219CE5FF80E04A2372484251769B5D" box="[560,815,331,358]" pageId="14" pageNumber="315">Australian Museum U.</collectorName>
5794, from
<collectionCode id="D2C561F6FF80E04A22F7484251A49B5E" box="[949,1021,331,357]" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="14" pageNumber="315" type="Other institutional type">NIWA</collectionCode>
<collectorName id="19219CE5FF80E04A25464842561C9B5E" box="[1028,1093,331,357]" pageId="14" pageNumber="315">Stn U</collectorName>
207,
<geoCoordinate id="D1E09FF4FF80E04A253F484256A59B5E" box="[1149,1276,331,358]" degrees="34.1850" direction="south" orientation="latitude" pageId="14" pageNumber="315" precision="5" value="-34.185">34.1850° S</geoCoordinate>
,
<geoCoordinate id="D1E09FF4FF80E04A244B484257CE9B5D" box="[1289,1431,331,358]" degrees="151.4333" direction="east" orientation="longitude" pageId="14" pageNumber="315" precision="5" value="151.4333">151.4333° E</geoCoordinate>
,
<quantity id="732C54D6FF80E04A21D5486652B99BB1" box="[151,224,367,394]" metricMagnitude="2" metricUnit="m" metricValue="1.98" pageId="14" pageNumber="315" unit="m" value="198.0">
<elevation id="3FF91E00FF80E04A21D5486652B99BB1" box="[151,224,367,394]" metricMagnitude="2" metricUnit="m" metricValue="1.98" pageId="14" pageNumber="315" unit="m" value="198.0">198 m</elevation>
</quantity>
,
<collectingRegion id="761037D1FF80E04A21A9487953E89BB2" box="[235,433,367,394]" country="Australia" name="New South Wales" pageId="14" pageNumber="315">New South Wales</collectingRegion>
shelf,
<collectingRegion id="761037D1FF80E04A20B94879500B9BB1" box="[507,594,368,394]" country="New Zealand" name="Tasman" pageId="14" pageNumber="315">Tasman</collectingRegion>
<location id="B10BAFE8FF80E04A231A486650DD9BB2" LSID="urn:lsid:plazi:treatment:3C7D4825FF80E04B21D5499E51659F0A:B10BAFE8FF80E04A231A486650DD9BB2" box="[600,644,367,393]" latitude="-34.185" longLatPrecision="6" longitude="151.4333" name="Sea" pageId="14" pageNumber="315" stateProvince="New South Wales">Sea</location>
</materialsCitation>
.
<materialsCitation id="04BCF36EFF80E04A23CC487957139BB2" ID-GBIF-Occurrence="3001059337" box="[654,1354,367,394]" collectionCode="NIWA" collectorName="Australian Museum U." elevation="198" latitude="-34.185" location="Sea" longLatPrecision="6" longitude="151.4333" pageId="14" pageNumber="315" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<emphasis id="86A02521FF80E04A23CC487951519BB2" box="[654,776,368,393]" italics="true" pageId="14" pageNumber="315">
<typeStatus id="6B6F4791FF80E04A23CC4879515A9BB2" box="[654,771,368,393]" pageId="14" pageNumber="315" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectorName id="19219CE5FF80E04A224C486656529BB1" box="[782,1035,367,394]" pageId="14" pageNumber="315">Australian Museum U.</collectorName>
5795, same data as holotype
</materialsCitation>
;
<materialsCitation id="04BCF36EFF80E04A2417486653AD9B96" ID-GBIF-Occurrence="3001059322" collectionCode="NIWA" collectorName="Australian Museum U. &amp; Stn U" elevation="198" latitude="-34.185" location="Sea" longLatPrecision="6" longitude="151.4333" pageId="14" pageNumber="315" specimenCode="NIWA 144898" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<specimenCode id="E4725148FF80E04A2417486652B79B96" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="14" pageNumber="315" type="Other institutional type">NIWA 144898</specimenCode>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF80E04A218548BE566D985A" pageId="14" pageNumber="315" type="diagnosis">
<paragraph id="B46BF933FF80E04A218548BE566D985A" blockId="14.[151,1437,259,610]" pageId="14" pageNumber="315">
<emphasis id="86A02521FF80E04A218548BE531B9BEA" bold="true" box="[199,322,439,465]" pageId="14" pageNumber="315">Diagnosis.</emphasis>
Branches flattened, mostly
<quantity id="732C54D6FF80E04A23C048BE509C9BE9" box="[642,709,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="315" unit="mm" value="3.0">3 mm</quantity>
wide. Autozooids averaging 931 μm long, 423 μm wide. Auto-zooidal spiramen mostly with 67 pores. Suboral avicularia 13, moderate-sized; no additional such avicularia in ephebic zooids. Large vicarious avicularia absent; rather, cystids of marginal avicularia subvicarious with functional rostral-opesial component same size as autozooidal suboral avicularia. Putative gonozooid slightly larger than autozooids with larger spiramen of 14 pores. Adventitious avicularia as for autozooids.
</paragraph>
</subSubSection>
<caption id="E0ABA9BBFF80E04A21D54E43512B9DF5" ID-DOI="http://doi.org/10.5281/zenodo.4326670" ID-Zenodo-Dep="4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="14" pageNumber="315" startId="14.[151,250,1866,1890]" targetBox="[249,1339,636,1838]" targetPageId="14">
<paragraph id="B46BF933FF80E04A21D54E43512B9DF5" blockId="14.[151,1436,1866,1998]" pageId="14" pageNumber="315">
<emphasis id="86A02521FF80E04A21D54E4353469D59" bold="true" box="[151,287,1866,1890]" pageId="14" pageNumber="315">FIGURE 10.</emphasis>
<taxonomicName id="73D482B0FF80E04A20644E4353AB9D59" authority="Liow &amp; Gordon, 2020" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[294,498,1866,1890]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="315" phylum="Bryozoa" rank="species" species="wassi" status="sp. nov.">
<emphasis id="86A02521FF80E04A20644E4353AB9D59" box="[294,498,1866,1890]" italics="true" pageId="14" pageNumber="315">Adeonellopsis wassi</emphasis>
</taxonomicName>
<emphasis id="86A02521FF80E04A20BA4E4350689D59" bold="true" box="[504,561,1866,1890]" pageId="14" pageNumber="315">
<taxonomicNameLabel id="9D93985AFF80E04A20BA4E4350689D59" box="[504,561,1866,1890]" pageId="14" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(NIWA 144898): A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm.
</paragraph>
</caption>
<subSubSection id="FCCEAAB8FF81E04B2185499F50DD9E62" pageId="15" pageNumber="316" type="description">
<paragraph id="B46BF933FF81E04B2185499F515F9B7A" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
<emphasis id="86A02521FF81E04B2185499F53009A8B" bold="true" box="[199,345,150,176]" pageId="15" pageNumber="316">Description.</emphasis>
Colony erect, rigid, maximum size not known; longest available branch fragment
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, lateral branching sparse, with distance between bifurcations variable, sometimes quite long; branching in same plane. Colour in life unknown; dried fragments pale creamy-brown. Branch widths varying with age of colony, 2.0
<quantity id="732C54D6FF81E04B240349D657C19AC2" box="[1345,1432,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="316" unit="mm" value="3.8">3.8 mm</quantity>
, mostly
<quantity id="732C54D6FF81E04B21AF480A53689B26" box="[237,305,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="316" unit="mm" value="3.0">3 mm</quantity>
between bifurcations, wider just proximal to bifurcations. Autozooids arranged in quincunx; 610 longitudinal series according to branch width (
<figureCitation id="2CEFE5B6FF81E04B23E3482E50A09B7A" box="[673,761,295,321]" captionStart="FIGURE 9" captionStartId="13.[151,250,1622,1646]" captionTargetBox="[243,1344,420,1595]" captionTargetId="figure-142@13.[239,1347,417,1599]" captionTargetPageId="13" captionText="FIGURE 9. Adeonellopsis tasmanensis n. sp. (NIWA 144892): A, B. Two gonozooids with varied numbers and orientations of adventitious avicularia. Adeonellopsis wassi n. sp. (NIWA 144899): C. Part of stem near axial bifurcation; the black arrow indicates a zooid (gonozooid?) with an unusually large spiramen. D.Autozooids with paired and single suboral avicularia having varied orientations. Scalebars: A, B, 300 μm; C, 2 mm; D, 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4326668" httpUri="https://zenodo.org/record/4326668/files/figure.png" pageId="15" pageNumber="316">Fig. 9C</figureCitation>
).
</paragraph>
<paragraph id="B46BF933FF81E04B2185484251399BEA" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
Neanic autozooids (
<figureCitation id="2CEFE5B6FF81E04B20E94842505F9B5E" box="[427,518,331,357]" captionStart="FIGURE 9" captionStartId="13.[151,250,1622,1646]" captionTargetBox="[243,1344,420,1595]" captionTargetId="figure-142@13.[239,1347,417,1599]" captionTargetPageId="13" captionText="FIGURE 9. Adeonellopsis tasmanensis n. sp. (NIWA 144892): A, B. Two gonozooids with varied numbers and orientations of adventitious avicularia. Adeonellopsis wassi n. sp. (NIWA 144899): C. Part of stem near axial bifurcation; the black arrow indicates a zooid (gonozooid?) with an unusually large spiramen. D.Autozooids with paired and single suboral avicularia having varied orientations. Scalebars: A, B, 300 μm; C, 2 mm; D, 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4326668" httpUri="https://zenodo.org/record/4326668/files/figure.png" pageId="15" pageNumber="316">Fig. 9D</figureCitation>
) more or less rounded-subhexagonal, becoming more elongate-rectangular with ageing and secondary calcification. Interzooidal boundaries indicated by thin lines of calcification in interzooidal furrows, bordered by 1626 areolar pores in single series around entire zooidal margin. Zooids relatively large, mean zooid length and width 931 μm and 423 μm, respectively.
</paragraph>
<paragraph id="B46BF933FF81E04B218548D25001985A" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
Autozooidal peristomial orifice suborbicular to a transversely rounded D-shape. Denticulation (
<figureCitation id="2CEFE5B6FF81E04B25AC48D257089BCE" box="[1262,1361,475,501]" captionStart="FIGURE 10" captionStartId="14.[151,250,1866,1890]" captionTargetBox="[249,1339,636,1838]" captionTargetId="figure-163@14.[246,1342,633,1841]" captionTargetPageId="14" captionText="FIGURE 10. Adeonellopsis wassi n. sp. (NIWA 144898):A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="15" pageNumber="316">Fig. 10B</figureCitation>
) of inner peristomial margin well developed, sparse or absent. Multiporous spiramen in frontal depression (
<figureCitation id="2CEFE5B6FF81E04B244D48F657CD9822" box="[1295,1428,511,537]" captionStart="FIGURE 10" captionStartId="14.[151,250,1866,1890]" captionTargetBox="[249,1339,636,1838]" captionTargetId="figure-163@14.[246,1342,633,1841]" captionTargetPageId="14" captionText="FIGURE 10. Adeonellopsis wassi n. sp. (NIWA 144898):A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="15" pageNumber="316">Fig. 10A, E</figureCitation>
) circularoval, more or less central; averaging 122 μm long, 96 μm wide; spiraminal pores 510, mostly 67, each with 47 short spokes that do not touch.
</paragraph>
<paragraph id="B46BF933FF81E04B21854B62576C9902" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
Adventitious avicularia (
<figureCitation id="2CEFE5B6FF81E04B20984B62506198BE" box="[474,568,619,645]" captionStart="FIGURE 9" captionStartId="13.[151,250,1622,1646]" captionTargetBox="[243,1344,420,1595]" captionTargetId="figure-142@13.[239,1347,417,1599]" captionTargetPageId="13" captionText="FIGURE 9. Adeonellopsis tasmanensis n. sp. (NIWA 144892): A, B. Two gonozooids with varied numbers and orientations of adventitious avicularia. Adeonellopsis wassi n. sp. (NIWA 144899): C. Part of stem near axial bifurcation; the black arrow indicates a zooid (gonozooid?) with an unusually large spiramen. D.Autozooids with paired and single suboral avicularia having varied orientations. Scalebars: A, B, 300 μm; C, 2 mm; D, 500 μm." figureDoi="http://doi.org/10.5281/zenodo.4326668" httpUri="https://zenodo.org/record/4326668/files/figure.png" pageId="15" pageNumber="316">Figs 9D</figureCitation>
;
<figureCitation id="2CEFE5B6FF81E04B23014B6250CD98BE" box="[579,660,619,645]" captionStart="FIGURE 10" captionStartId="14.[151,250,1866,1890]" captionTargetBox="[249,1339,636,1838]" captionTargetId="figure-163@14.[246,1342,633,1841]" captionTargetPageId="14" captionText="FIGURE 10. Adeonellopsis wassi n. sp. (NIWA 144898):A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="15" pageNumber="316">10A, C</figureCitation>
) suboral only, typically 12 but increasing to
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ephebic zooids; one is placed medially, pointing mostly distally, with a second to one side of it pointing transversely or obliquely toward it; an areolar pore on the opposite side of the median one can later develop into a third such avicularium, converging towards its partners so that the tips of all three somewhat converge. All have the typical form, with a narrow openchannelled rostral tip, smooth raised (gymnocystal) margins, common rostral-opesial foramen with slight denticulation and mostly smooth cryptocyst; no pivot bar but short curved pivot ridges at rostral-opesial transition.
</paragraph>
<paragraph id="B46BF933FF81E04B21854A4A517F999E" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
Large vicarious avicularia absent; instead, subvicarious avicularian cystids occur on branch margins (
<figureCitation id="2CEFE5B6FF81E04B246B4A4A57C99966" box="[1321,1424,835,861]" captionStart="FIGURE 10" captionStartId="14.[151,250,1866,1890]" captionTargetBox="[249,1339,636,1838]" captionTargetId="figure-163@14.[246,1342,633,1841]" captionTargetPageId="14" captionText="FIGURE 10. Adeonellopsis wassi n. sp. (NIWA 144898):A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="15" pageNumber="316">Fig. 10D</figureCitation>
). The functional rostral-opesial (gymnocystal) part of the avicularium has the same size and form as the suboral avicularia of autozooids, the rostral tips frontally elevated.
</paragraph>
<paragraph id="B46BF933FF81E04B21854AA653419E0E" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
Only one putative gonozooid seen (
<figureCitation id="2CEFE5B6FF81E04B231E4AA6509F99F2" box="[604,710,943,969]" captionStart="FIGURE 10" captionStartId="14.[151,250,1866,1890]" captionTargetBox="[249,1339,636,1838]" captionTargetId="figure-163@14.[246,1342,633,1841]" captionTargetPageId="14" captionText="FIGURE 10. Adeonellopsis wassi n. sp. (NIWA 144898):A. Autozooids and a possible gonozooid. B. Autozooidal peristomial orifice showing denticulation of inner proximal rim. C. Close-up of suboral autozooidal avicularia; note the pore on the right that may later produce a third avicularium. D. Marginal subvicarious avicularian cystids with small rostral-opesial area. E. Autozooidal spiramen. Scalebars: A, 500 μm; B, C, E, 100 μm; D, 300 μm." figureDoi="http://doi.org/10.5281/zenodo.4326670" httpUri="https://zenodo.org/record/4326670/files/figure.png" pageId="15" pageNumber="316">Fig. 10A</figureCitation>
), 1104 μm long, 511 μm wide, occurring four zooid lengths be-low a bifurcation. Gonozooid peristomial orifice 147 μm wide, the spiramen larger than autozooids, with 14 pores, each with 47 short spokes that do not touch unless pore laterally constricted. Two suboral avicularia like those in autozooids.
</paragraph>
<paragraph id="B46BF933FF81E04B21854D3650DD9E62" blockId="15.[151,1436,150,1329]" box="[199,644,1087,1113]" pageId="15" pageNumber="316">Ancestrula and early astogeny not seen.</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF81E04B21854D6A52A09F36" pageId="15" pageNumber="316" type="discussion">
<paragraph id="B46BF933FF81E04B21854D6A52A09F36" blockId="15.[151,1436,150,1329]" pageId="15" pageNumber="316">
<emphasis id="86A02521FF81E04B21854D6A53629E46" bold="true" box="[199,315,1123,1149]" pageId="15" pageNumber="316">Remarks.</emphasis>
Compared to the two new species described above,
<taxonomicName id="73D482B0FF81E04B22C34D6A563B9E47" authority="Liow &amp; Gordon, 2020" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[897,1122,1123,1149]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="316" phylum="Bryozoa" rank="species" species="wassi" status="sp. nov.">
<emphasis id="86A02521FF81E04B22C34D6A563B9E47" box="[897,1122,1123,1149]" italics="true" pageId="15" pageNumber="316">Adeonellopsis wassi</emphasis>
</taxonomicName>
<emphasis id="86A02521FF81E04B252B4D6A56FE9E46" bold="true" box="[1129,1191,1123,1149]" pageId="15" pageNumber="316">
<taxonomicNameLabel id="9D93985AFF81E04B252B4D6A56FE9E46" box="[1129,1191,1123,1149]" pageId="15" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has relatively smaller suboral avicularia, reminiscent of those in
<taxonomicName id="73D482B0FF81E04B232E4D8E51399E9B" authority="Liow &amp; Gordon, 2020" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[620,864,1159,1185]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="316" phylum="Bryozoa" rank="species" species="gemina" status="sp. nov.">
<emphasis id="86A02521FF81E04B232E4D8E51399E9B" box="[620,864,1159,1185]" italics="true" pageId="15" pageNumber="316">Adeonellopsis gemina</emphasis>
</taxonomicName>
<emphasis id="86A02521FF81E04B22254D8E51FD9E9A" bold="true" box="[871,932,1159,1185]" pageId="15" pageNumber="316">
<taxonomicNameLabel id="9D93985AFF81E04B22254D8E51FD9E9A" box="[871,932,1159,1185]" pageId="15" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="73D482B0FF81E04B229A4D8E56E79E9B" authority="Liow &amp; Gordon, 2020" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[984,1214,1159,1185]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="316" phylum="Bryozoa" rank="species" species="minor" status="sp. nov.">
<emphasis id="86A02521FF81E04B229A4D8E56E79E9B" box="[984,1214,1159,1185]" italics="true" pageId="15" pageNumber="316">Adeonellopsis minor</emphasis>
</taxonomicName>
<emphasis id="86A02521FF81E04B25814D8E57589E9A" bold="true" box="[1219,1281,1159,1185]" pageId="15" pageNumber="316">
<taxonomicNameLabel id="9D93985AFF81E04B25814D8E57589E9A" box="[1219,1281,1159,1185]" pageId="15" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. Colony form in these latter two species differs markedly, however, being smaller with thinner branches. Overall,
<taxonomicName id="73D482B0FF81E04B244C4DA257379EFF" authorityName="Liow &amp; Gordon" authorityYear="2020" box="[1294,1390,1195,1221]" class="Gymnolaemata" family="Adeonidae" genus="Adeonellopsis" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="316" phylum="Bryozoa" rank="species" species="wassi">
<emphasis id="86A02521FF81E04B244C4DA257379EFF" box="[1294,1390,1195,1221]" italics="true" pageId="15" pageNumber="316">A. wassi</emphasis>
</taxonomicName>
has the largest zooids of any of the new species described herein. It also has only subvicarious avicularia on its branch margins.
</paragraph>
</subSubSection>
<subSubSection id="FCCEAAB8FF81E04B21854C1F51659F0A" box="[199,828,1302,1329]" pageId="15" pageNumber="316" type="distribution">
<paragraph id="B46BF933FF81E04B21854C1F51659F0A" blockId="15.[151,1436,150,1329]" box="[199,828,1302,1329]" pageId="15" pageNumber="316">
<emphasis id="86A02521FF81E04B21854C1F53389F0B" bold="true" box="[199,353,1302,1328]" pageId="15" pageNumber="316">Distribution.</emphasis>
<collectingCountry id="CCC3B9A3FF81E04B20244C1E53889F0A" box="[358,465,1303,1329]" name="Australia" pageId="15" pageNumber="316">Australia</collectingCountry>
:
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shelf,
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.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>