treatments-xml/data/BE/47/E5/BE47E558FFDDFFE95ED5CCE1FB0BF3F4.xml
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<document id="2C45ED9304DF70D509EC10628B85347B" ID-DOI="10.5281/zenodo.201624" ID-GBIF-Dataset="cb51feb9-68d1-481e-bf51-1d846277cf15" ID-ISSN="1175-5326" ID-Zenodo-Dep="201624" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460266258637" checkinUser="plazi" docAuthor="Salcedo, Diana L., Martín, Guillermo San &amp; Solís-Weiss, Vivianne" docDate="2011" docId="BE47E558FFDDFFE95ED5CCE1FB0BF3F4" docLanguage="en" docName="zt02800p052.pdf" docOrigin="Zootaxa 2800" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Trypanosyllis microdenticulata Salcedo, Martín &amp; Solís-Weiss, 2011, n. sp." docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="427E9D20FFD8FFE35E42CD72FFB2F303" masterDocTitle="Two new species of Syllidae (Annelida: Polychaeta) from the Southern Mexican Pacific" masterLastPageNumber="52" masterPageNumber="41" pageNumber="46" updateTime="1698245785610" updateUser="plazi">
<mods:mods id="7DD5AF375E1F79D3E5706C8CDA74C926" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FE44200688AE497CF7BF97EF7CB173BD">
<mods:title id="A51154411AC52F6E2524BCB335FDCB78">Two new species of Syllidae (Annelida: Polychaeta) from the Southern Mexican Pacific</mods:title>
</mods:titleInfo>
<mods:name id="2417EF93F3F5D1C8E5B45272985557C2" type="personal">
<mods:role id="B5902FED75F2A8D31C09D348DAC66312">
<mods:roleTerm id="59682B942AAFE162866B40EC41C50998">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4567E705160FD9B1ED9C9572CFEBD3C8">Salcedo, Diana L.</mods:namePart>
</mods:name>
<mods:name id="65222777CC864704A1C83FB1450A6AF6" type="personal">
<mods:role id="C7B9EF126B53F449AB50177D68E9BC92">
<mods:roleTerm id="C2C5602ABA4AF4765262CA2A3E3F2EBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63155D0F124B157845A99C0201A85F39">Martín, Guillermo San</mods:namePart>
</mods:name>
<mods:name id="7D059A4B1ED45AE2B5807FA8C79D440C" type="personal">
<mods:role id="4F9E015A345F49A72E65F09723CB6B28">
<mods:roleTerm id="36C13227C065D89EDAAFB22E83398511">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A0FA9F44B47EB9C18B590FECA235C5E">Solís-Weiss, Vivianne</mods:namePart>
</mods:name>
<mods:typeOfResource id="9F51966C9F71594EFFE888475579DA61">text</mods:typeOfResource>
<mods:relatedItem id="688FA3F6EC729E36849539DDB761BF85" type="host">
<mods:titleInfo id="CF2126A927E6896B674D7ADE653C4408">
<mods:title id="1E5AD55681666200022DD3F514F8EC33">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="49A8EE9879E23E3153783816B6BB7938">
<mods:date id="FB3D07E9121187B95A4DE0E3CF320647">2011</mods:date>
<mods:detail id="A1A9F7198EDFF2F3F760343076100F98" type="volume">
<mods:number id="93E73EAD860C80AF08FDD25C637B576A">2800</mods:number>
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<mods:start id="C00E2E5501D6CE0EB719F4F915DE51BF">41</mods:start>
<mods:end id="F3B0E5BC74BB55BDD46E47DC2B7E7842">52</mods:end>
</mods:extent>
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<mods:classification id="DB69321B45029003A00CD77DC2BCCCA5">journal article</mods:classification>
<mods:identifier id="8AE20D71F8322021AA683684E03A2F07" type="DOI">10.5281/zenodo.201624</mods:identifier>
<mods:identifier id="767B55D32E5875D2109957A3C1AC4CB2" type="GBIF-Dataset">cb51feb9-68d1-481e-bf51-1d846277cf15</mods:identifier>
<mods:identifier id="42C0C34AC560439A4A9C0611B322A415" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="14344E42CAE70FC46BB6BD6E8E9967BD" type="Zenodo-Dep">201624</mods:identifier>
</mods:mods>
<treatment id="BE47E558FFDDFFE95ED5CCE1FB0BF3F4" ID-DOI="http://doi.org/10.5281/zenodo.6183566" ID-GBIF-Taxon="119394172" ID-Zenodo-Dep="6183566" LSID="urn:lsid:plazi:treatment:BE47E558FFDDFFE95ED5CCE1FB0BF3F4" httpUri="http://treatment.plazi.org/id/BE47E558FFDDFFE95ED5CCE1FB0BF3F4" lastPageId="10" lastPageNumber="51" pageId="5" pageNumber="46">
<subSubSection id="7EF407C5FFDDFFE65ED5CCE1FDAAF2CC" pageId="5" pageNumber="46" type="nomenclature">
<paragraph id="3651544EFFDDFFE65ED5CCE1FDF0F2AF" blockId="5.[151,578,403,463]" box="[151,578,403,428]" pageId="5" pageNumber="46">
<heading id="6D19E322FFDDFFE65ED5CCE1FDF0F2AF" bold="true" box="[151,578,403,428]" fontSize="10" level="2" pageId="5" pageNumber="46" reason="2">
<emphasis id="049A885CFFDDFFE65ED5CCE1FDF0F2AF" bold="true" box="[151,578,403,428]" pageId="5" pageNumber="46">
<taxonomicName id="F1EE2FCDFFDDFFE65ED5CCE1FE4EF2AF" ID-CoL="59FJ7" box="[151,508,403,428]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="46" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFDDFFE65ED5CCE1FE4EF2AF" bold="true" box="[151,508,403,428]" italics="true" pageId="5" pageNumber="46">Trypanosyllis microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFDDFFE65C41CCE6FDF0F2AF" box="[515,578,404,428]" pageId="5" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3651544EFFDDFFE65ED5CCC5FDAAF2CC" blockId="5.[151,578,403,463]" box="[151,536,439,463]" pageId="5" pageNumber="46">
(
<figureCitation id="AED548CBFFDDFFE65EDDCCC5FF58F2CC" box="[159,234,439,463]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Figs. 3</figureCitation>
AB, 4AG, 5AF, 6AD)
</paragraph>
</subSubSection>
<subSubSection id="7EF407C5FFDDFFE65ED5CC8DFD49F0EF" pageId="5" pageNumber="46" type="materials_examined">
<paragraph id="3651544EFFDDFFE65ED5CC8DFE92F07C" blockId="5.[151,1437,511,2013]" pageId="5" pageNumber="46">
<emphasis id="049A885CFFDDFFE65ED5CC8DFE31F11B" bold="true" box="[151,387,511,536]" pageId="5" pageNumber="46">Material examined.</emphasis>
<typeStatus id="E955EAECFFDDFFE65FCECC8DFE4BF114" box="[396,505,511,535]" pageId="5" pageNumber="46" type="holotype">Holotype</typeStatus>
: complete specimen (CPICML POH37003), El Jardín E3B1,
<geoCoordinate id="53DA3289FFDDFFE65A9ACC8DFAD8F114" box="[1240,1386,511,535]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823933">16°49.436N</geoCoordinate>
and
<geoCoordinate id="53DA3289FFDDFFE65ED5CF56FE9DF13F" box="[151,303,548,572]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91635">99°54.981W</geoCoordinate>
,
<date id="4250728EFFDDFFE65F7ECF56FE64F13F" box="[316,470,548,573]" pageId="5" pageNumber="46" value="2006-05-26">26 May 2006</date>
, 12 m, coarse sand.
<typeStatus id="E955EAECFFDDFFE65CFDCF57FC82F13E" box="[703,816,549,573]" pageId="5" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="20E89FC7FFDDFFE65D7ECF56FC75F13F" box="[828,967,548,572]" pageId="5" pageNumber="46" type="generic">9 specimens</specimenCount>
(CPICML POP-37-004) El Jardín E3B1,
<geoCoordinate id="53DA3289FFDDFFE65ED5CF35FE9BF15C" box="[151,297,583,607]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823933">16°49.436N</geoCoordinate>
and
<geoCoordinate id="53DA3289FFDDFFE65F26CF35FE4EF15C" box="[356,508,583,607]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91635">99°54.981W</geoCoordinate>
,
<date id="4250728EFFDDFFE65C49CF35FD1AF15C" box="[523,680,583,608]" pageId="5" pageNumber="46" value="2006-05-26">26 May 2006</date>
, 12 m, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65DD5CF35FB96F15C" box="[919,1060,583,607]" pageId="5" pageNumber="46" type="generic">6 specimens</specimenCount>
(
<collectionCode id="50FFCC8BFFDDFFE65A77CF35FB39F15C" LSID="urn:lsid:biocol.org:col:35275" box="[1077,1163,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="46">MNCN</collectionCode>
16.01/13263), Palmitas E2B2
<geoCoordinate id="53DA3289FFDDFFE65E9CCF1EFEC0F187" box="[222,370,620,644]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823668">16°49.420N</geoCoordinate>
,
<geoCoordinate id="53DA3289FFDDFFE65F3CCF1EFDA4F187" box="[382,534,620,644]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91222">99°54.733W</geoCoordinate>
,
<date id="4250728EFFDDFFE65C63CF1EFD08F187" box="[545,698,620,645]" pageId="5" pageNumber="46" value="2006-05-25">25 May 2006</date>
,
<quantity id="F116F9ABFFDDFFE65C87CF1EFCA7F187" box="[709,789,620,644]" metricMagnitude="1" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="46" unit="m" value="10.5">10.5 m</quantity>
, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65DF0CF1EFB8EF187" box="[946,1084,620,644]" pageId="5" pageNumber="46" type="generic">6 specimens</specimenCount>
(LACM-AHF
<collectionCode id="50FFCC8BFFDDFFE65AA4CF1EFA9EF187" box="[1254,1324,620,644]" pageId="5" pageNumber="46">POLY</collectionCode>
2640), El Jardín E3B2,
<geoCoordinate id="53DA3289FFDDFFE65F7ACFFDFE78F1A4" box="[312,458,655,679]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823933">16°49.436N</geoCoordinate>
and
<geoCoordinate id="53DA3289FFDDFFE65C4ACFFDFD12F1A4" box="[520,672,655,679]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91635">99°54.981W</geoCoordinate>
,
<date id="4250728EFFDDFFE65CEDCFFDFCFDF1A4" box="[687,847,655,680]" pageId="5" pageNumber="46" value="2006-05-26">26 May 2006</date>
, 12 m, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65A07CFFDFB61F1A4" box="[1093,1235,655,679]" pageId="5" pageNumber="46" type="generic">6 specimens</specimenCount>
(AM: W.37087), Palmitas E2B1,
<geoCoordinate id="53DA3289FFDDFFE65F0ECFC6FE6DF1CF" box="[332,479,692,716]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823668">16°49.420N</geoCoordinate>
and
<geoCoordinate id="53DA3289FFDDFFE65C57CFC6FD1FF1CF" box="[533,685,692,716]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91222">99°54.733W</geoCoordinate>
,
<date id="4250728EFFDDFFE65CFBCFC6FCE3F1CF" box="[697,849,692,717]" pageId="5" pageNumber="46" value="2006-05-25">25 May 2006</date>
,
<quantity id="F116F9ABFFDDFFE65D1ECFC6FC1FF1CF" box="[860,941,692,716]" metricMagnitude="1" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="46" unit="m" value="10.5">10.5 m</quantity>
, coarse sand.
<emphasis id="049A885CFFDDFFE65A08CFC1FA8FF1CF" bold="true" box="[1098,1341,691,716]" pageId="5" pageNumber="46">Additional material.</emphasis>
<specimenCount id="20E89FC7FFDDFFE65B06CFC6FF60F1F3" pageId="5" pageNumber="46" type="generic">2 specimens</specimenCount>
(CPICML PO37080), Palmitas E2B1,
<geoCoordinate id="53DA3289FFDDFFE65CE7CFA5FC8AF1EC" box="[677,824,727,751]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823668">16°49.420N</geoCoordinate>
and
<geoCoordinate id="53DA3289FFDDFFE65D2CCFA5FBB5F1EC" box="[878,1031,727,751]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91222">99°54.733W</geoCoordinate>
,
<date id="4250728EFFDDFFE65A50CFA5FB19F1EC" box="[1042,1195,727,753]" pageId="5" pageNumber="46" value="2006-05-25">25 May 2006</date>
,
<quantity id="F116F9ABFFDDFFE65AF4CFA5FAB5F1F3" box="[1206,1287,727,752]" metricMagnitude="1" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="46" unit="m" value="10.5">10.5 m</quantity>
, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65ED5CF8FFE96F017" box="[151,292,764,788]" pageId="5" pageNumber="46" type="generic">7 specimens</specimenCount>
(CPICML PO37080), Palmitas E2B2
<geoCoordinate id="53DA3289FFDDFFE65CBFCF8EFC20F017" box="[765,914,764,788]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823668">16°49.420N</geoCoordinate>
,
<geoCoordinate id="53DA3289FFDDFFE65DE2CF8EFB8BF017" box="[928,1081,764,788]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91222">99°54.733W</geoCoordinate>
,
<date id="4250728EFFDDFFE65A04CF8EFB56F017" box="[1094,1252,764,789]" pageId="5" pageNumber="46" value="2006-05-25">25 May 2006</date>
,
<quantity id="F116F9ABFFDDFFE65AB0CF8EFAF7F017" box="[1266,1349,764,788]" metricMagnitude="1" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="46" unit="m" value="10.5">10.5 m</quantity>
, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65E9ECE6DFED8F034" box="[220,362,799,823]" pageId="5" pageNumber="46" type="generic">4 specimens</specimenCount>
(CPICML PO37 080), El Jardín E3B2,
<geoCoordinate id="53DA3289FFDDFFE65D22CE6DFC46F034" box="[864,1012,799,823]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.823933">16°49.436N</geoCoordinate>
,
<geoCoordinate id="53DA3289FFDDFFE65A41CE6DFB29F034" box="[1027,1179,799,823]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.91635">99°54.981W</geoCoordinate>
,
<date id="4250728EFFDDFFE65AE8CE6DFAF8F034" box="[1194,1354,799,825]" pageId="5" pageNumber="46" value="2006-05-26">26 May 2006</date>
, 12 m, coarse sand.
<specimenCount id="20E89FC7FFDDFFE65F68CE36FE19F05F" box="[298,427,836,860]" pageId="5" pageNumber="46" type="generic">1 specimen</specimenCount>
(CPICML PO37080), Caleta E1B2,
<geoCoordinate id="53DA3289FFDDFFE65D2ECE36FBB2F05F" box="[876,1024,836,860]" direction="north" orientation="latitude" pageId="5" pageNumber="46" precision="1" value="16.82995">16°49.797N</geoCoordinate>
,
<geoCoordinate id="53DA3289FFDDFFE65A4FCE36FB17F05F" box="[1037,1189,836,860]" direction="west" orientation="longitude" pageId="5" pageNumber="46" precision="1" value="-99.90103">99°54.062W</geoCoordinate>
,
<date id="4250728EFFDDFFE65AF0CE36FAFFF05F" box="[1202,1357,836,861]" pageId="5" pageNumber="46" value="2006-05-25">25 May 2006</date>
, 12 m, coarse sand.
</paragraph>
<paragraph id="3651544EFFDDFFE65E87CEF9FD49F0EF" blockId="5.[151,1437,511,2013]" pageId="5" pageNumber="46">
<emphasis id="049A885CFFDDFFE65E87CEF9FDFDF0A7" bold="true" box="[197,591,907,932]" pageId="5" pageNumber="46">Comparative material examined.</emphasis>
<taxonomicName id="F1EE2FCDFFDDFFE65C14CEFEFCE1F0A7" box="[598,851,908,932]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="46" phylum="Annelida" rank="species" species="coeliaca">
<emphasis id="049A885CFFDDFFE65C14CEFEFCE1F0A7" box="[598,851,908,932]" italics="true" pageId="5" pageNumber="46">Trypanosyllis coeliaca</emphasis>
</taxonomicName>
:
<specimenCount id="20E89FC7FFDDFFE65D23CEFEFC59F0A7" box="[865,1003,908,932]" pageId="5" pageNumber="46" type="generic">4 specimens</specimenCount>
(
<collectionCode id="50FFCC8BFFDDFFE65DBBCEFEFBFDF0A7" LSID="urn:lsid:biocol.org:col:35275" box="[1017,1103,908,932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="46">MNCN</collectionCode>
16.01/6860).
<collectingCountry id="4EF914DEFFDDFFE65AAECEFEFA82F0A7" box="[1260,1328,908,932]" name="Spain" pageId="5" pageNumber="46">Spain</collectingCountry>
: Mediterranean Sea, Columbrete Is., N of Columbrete Grande,
<specimenCount id="20E89FC7FFDDFFE65D46CEDDFCF7F0CB" box="[772,837,943,968]" pageId="5" pageNumber="46" type="generic">
<quantity id="F116F9ABFFDDFFE65D46CEDDFCF3F0CB" box="[772,833,943,968]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="46" unit="m" value="47.0">47 m</quantity>
.
</specimenCount>
<taxonomicName id="F1EE2FCDFFDDFFE65D0FCEC2FBFEF0CB" box="[845,1100,944,968]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="46" phylum="Annelida" rank="species" species="coeliaca">
<emphasis id="049A885CFFDDFFE65D0FCEC2FBFEF0CB" box="[845,1100,944,968]" italics="true" pageId="5" pageNumber="46">Trypanosyllis coeliaca</emphasis>
</taxonomicName>
:
<specimenCount id="20E89FC7FFDDFFE65A1ECEDDFB55F0C4" box="[1116,1255,943,967]" pageId="5" pageNumber="46" type="generic">3 specimens</specimenCount>
(
<collectionCode id="50FFCC8BFFDDFFE65AB5CEDDFAFFF0C4" LSID="urn:lsid:biocol.org:col:35275" box="[1271,1357,943,967]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="46">MNCN</collectionCode>
16.01/ 6863).
<collectingCountry id="4EF914DEFFDDFFE65EA4CEA6FE98F0EF" box="[230,298,980,1004]" name="Spain" pageId="5" pageNumber="46">Spain</collectingCountry>
: Cantabric Sea, “El Cachucho”, Asturias.
</paragraph>
</subSubSection>
<subSubSection id="7EF407C5FFDDFFE55E87CE85FB51F267" lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46" type="description">
<paragraph id="3651544EFFDDFFE65E87CE85FDA6F633" blockId="5.[151,1437,511,2013]" pageId="5" pageNumber="46">
<emphasis id="049A885CFFDDFFE65E87CE85FEE4F713" bold="true" box="[197,342,1015,1040]" pageId="5" pageNumber="46">Description.</emphasis>
<typeStatus id="E955EAECFFDDFFE65F1DCE8AFE75F713" box="[351,455,1016,1040]" pageId="5" pageNumber="46" type="holotype">Holotype</typeStatus>
complete specimen
<quantity id="F116F9ABFFDDFFE65CF3CE8AFD46F713" box="[689,756,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="46" unit="mm" value="6.0">6 mm</quantity>
long,
<quantity id="F116F9ABFFDDFFE65D7CCE8AFC11F713" box="[830,931,1016,1040]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="46" unit="mm" value="0.62">0.62 mm</quantity>
wide, with 66 chaetigers. Longest complete specimen
<quantity id="F116F9ABFFDDFFE65F48C96EFED1F737" box="[266,355,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="46" unit="mm" value="9.5">9.5 mm</quantity>
long,
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wide, with 88 chaetigers. Body dorsoventrally flattened, colourless when preserved. Two dorsal transverse ciliated bands on each segment, visible under SEM: one with small tufts of cilia at midlength (
<figureCitation id="AED548CBFFDDFFE65F5AC916FEE8F77F" box="[280,346,1124,1148]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
C, indicated by superior arrow), the other continuous, close to anterior margin (
<figureCitation id="AED548CBFFDDFFE65A90C916FAA7F77F" box="[1234,1301,1124,1148]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
C, indicated by inferior arrow). Additional small ciliated bands on parapodial lobes (
<figureCitation id="AED548CBFFDDFFE65D83C9FAFBB6F7A3" box="[961,1028,1160,1184]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
D, indicated by arrow) as well as on dorsal cirri and bases of lateral antennae (
<figureCitation id="AED548CBFFDDFFE65C3AC9DEFD0FF7C7" box="[632,701,1196,1220]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
A, indicated by inferior arrow). Prostomium oval to trapezoidal, wider than long, dorsally bilobed, with two pairs of eyes in trapezoidal arrangement, close to each other, anterior pair ventral and posterior pair dorsal; one pair of ventral eyespots (lacking in a few
<typeStatus id="E955EAECFFDDFFE65A1CC986FB7CF60F" box="[1118,1230,1268,1292]" pageId="5" pageNumber="46" type="paratype">paratypes</typeStatus>
), close to anterior prostomial margin (
<figureCitation id="AED548CBFFDDFFE65F36C86AFE72F633" box="[372,448,1304,1328]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Figs. 3</figureCitation>
A, 3B).
</paragraph>
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Palps short, ovate, about half as long as prostomium, completely separated (
<figureCitation id="AED548CBFFDDFFE65A7BC84EFB3BF657" box="[1081,1161,1340,1364]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Figs. 3</figureCitation>
B, 5B). Antennae short, inserted on anterior prostomial margin; median antenna slightly longer than combined length of prostomium and palps, with 12 articles (
<quantity id="F116F9ABFFDDFFE65FE5C8F6FE51F69F" box="[423,483,1412,1436]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="46" unit="in" value="13.0">13 in</quantity>
some
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); lateral ones shorter, with 10 articles (
<figureCitation id="AED548CBFFDDFFE65A27C8F6FB06F69F" box="[1125,1204,1412,1436]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Figs. 3</figureCitation>
A, 5A). Two nuchal organs as postero-lateral rows of cilia, one around each prostomial lobe (
<figureCitation id="AED548CBFFDDFFE65D82C8DAFBB0F6C3" box="[960,1026,1448,1472]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
A, indicated by superior arrow). Peristomium well defined, shorter than following segments, with two pairs of tentacular cirri; dorsal one similar to median antennae, with 1415 articles; ventral one shorter, with 8 articles. Dorsal cirri short, thick, shorter than body width, slightly tapering distally, containing fibrillar material in each article, originating on short ceratophores (
<figureCitation id="AED548CBFFDDFFE65EDDCB4AFF59F553" box="[159,235,1592,1616]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Figs. 3</figureCitation>
A, 5A); longer on anterior segments, alternating long (1418 articles) and short (1011 articles) in midbody and posterior chaetigers, alternating long (1314 articles) and short (910 articles). Ventral cirri short, digitiform to conical, similar in size or slightly shorter than parapodial lobes. Parapodial lobes conical (
<figureCitation id="AED548CBFFDDFFE65AD5CBF2FB69F59B" box="[1175,1243,1664,1688]" captionStart="FIGURE 5" captionStartId="8.[151,250,1743,1766]" captionTargetBox="[160,1428,187,1727]" captionTargetId="figure@8.[151,1436,161,1758]" captionTargetPageId="8" captionText="FIGURE 5. SEM of Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (A B) 50 µm. (C) 100 µm. (D E) 10 µm. (F) 5 µm." httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="5" pageNumber="46">Fig. 5</figureCitation>
D). About 1315 heterogomph compound chaetae per parapodium, with thick shafts and bidentate blades, having short and coarse spines on margin; both teeth similar in size on anterior falcigers, subdistal tooth smaller than distal one on median and posterior chaetae (
<figureCitation id="AED548CBFFDDFFE65FDCCB9EFE42F407" box="[414,496,1772,1796]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
A, 4C, 4E). Marked dorso-ventral and antero-posterior gradation in blade length; anterior chaetae elongated, with slender blades (10.716.8 μm) (
<figureCitation id="AED548CBFFDDFFE65D27CA7DFC01F424" box="[869,947,1807,1831]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
A, 5E), becoming shorter and thicker posteriorly (1013.6 μm length) (
<figureCitation id="AED548CBFFDDFFE65F90CA46FD92F44F" box="[466,544,1844,1868]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
C, 4E, 5F). Dorsal and ventral simple chaetae only in most posterior chaetigers; dorsal one slender, smooth, bidentate, with subdistal tooth smaller (
<figureCitation id="AED548CBFFDDFFE65DD3CA25FC6DF46C" box="[913,991,1879,1903]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
G, 6A); ventral simple chaeta similar to dorsal one, but with short spines on subdistal margin (Fig. 6B). Two, sometimes three aciculae in anterior parapodia (one larger and thicker) (
<figureCitation id="AED548CBFFDDFFE65F94CAD2FD97F4BB" box="[470,549,1952,1976]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
B, 5D), and a single acicula in midbody and posterior parapodia (
<figureCitation id="AED548CBFFDDFFE65ABCCAD2FAFFF4BB" box="[1278,1357,1952,1976]" captionStart="FIGURE 4" captionStartId="7.[151,250,1717,1740]" captionTargetBox="[227,1355,201,1681]" captionTargetId="figure@7.[220,1368,193,1696]" captionTargetPageId="7" captionText="FIGURE 4. Trypanosyllis microdenticulata n. sp. (A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm." httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="5" pageNumber="46">Figs. 4</figureCitation>
D, 4F), all stout, straight, acute, protruding from parapodial lobes. Pharynx long, through nine segments (
<figureCitation id="AED548CBFFDDFFE65AABCAB6FA99F4DF" box="[1257,1323,1988,2012]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="5" pageNumber="46">Fig. 3</figureCitation>
A),(variation in
<typeStatus id="E955EAECFFDEFFE55EA4CDEAFEA4F3B3" box="[230,278,152,176]" pageId="6" pageNumber="47">type</typeStatus>
material is from 7 to 9), with a crown of ten soft papillae, surrounded by sinuous glands (
<figureCitation id="AED548CBFFDEFFE55B40CDE5FAF1F3AC" box="[1282,1347,151,175]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="6" pageNumber="47">Fig. 3</figureCitation>
A, indicated by arrow) and dark ovate glands inserted between the folds of latter (
<figureCitation id="AED548CBFFDEFFE55D96CDCEFBA6F3D7" box="[980,1044,188,212]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="6" pageNumber="47">Fig. 3</figureCitation>
A, indicated by arrow). Pharyngeal tooth conical, small and thin, trepan with 10 small, short and transparent teeth, very difficult to distinguish in some specimens (Fig. 6C, indicated by arrow). Proventricle short, extending through 4 segments (variation among
<typeStatus id="E955EAECFFDEFFE55B20CC77FF60F243" pageId="6" pageNumber="47" type="paratype">paratypes</typeStatus>
from 3 to 6), with 19 muscle cell rows (
<figureCitation id="AED548CBFFDEFFE55CDFCC55FD6DF23C" box="[669,735,295,319]" captionStart="FIGURE 3" captionStartId="6.[151,250,1943,1966]" captionTargetBox="[394,1171,424,1905]" captionTargetId="figure@6.[388,1198,409,1922]" captionTargetPageId="6" captionText="FIGURE 3. Trypanosyllis microdenticulata n. sp. (A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm." httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="6" pageNumber="47">Fig. 3</figureCitation>
A) (variation among
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from 17 to 21). Pygidium with two anal cirri, similar in length to dorsal ones at midbody, with 7 articles (or
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some
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).
</paragraph>
</subSubSection>
<caption id="629104C6FFDEFFE55ED5CAE5FCFFF4CD" httpUri="https://zenodo.org/record/201627/files/figure.png" pageId="6" pageNumber="47" targetBox="[394,1171,424,1905]" targetPageId="6">
<paragraph id="3651544EFFDEFFE55ED5CAE5FCFFF4CD" blockId="6.[151,1436,1943,2001]" pageId="6" pageNumber="47">
<emphasis id="049A885CFFDEFFE55ED5CAE5FEBFF4AE" bold="true" box="[151,269,1943,1966]" pageId="6" pageNumber="47">FIGURE 3</emphasis>
.
<taxonomicName id="F1EE2FCDFFDEFFE55F5ECAE5FDE5F4AE" box="[284,599,1943,1965]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="47" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFDEFFE55F5ECAE5FDE5F4AE" box="[284,599,1943,1965]" italics="true" pageId="6" pageNumber="47">Trypanosyllis microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFDEFFE55C1DCAEAFD28F4AE" box="[607,666,1944,1965]" pageId="6" pageNumber="47" rank="species">
<emphasis id="049A885CFFDEFFE55C1DCAEAFD28F4AE" bold="true" box="[607,666,1944,1965]" pageId="6" pageNumber="47">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Anterior end, dorsal view (Superior arrow: ovate dark glands. Inferior arrow: sinuous glands). (B) Anterior end, ventral view. Scale 40 µm.
</paragraph>
</caption>
<caption id="629104C6FFDFFFE45ED5CBC7FBC2F5EA" httpUri="https://zenodo.org/record/201628/files/figure.png" pageId="7" pageNumber="48" targetBox="[227,1355,201,1681]" targetPageId="7">
<paragraph id="3651544EFFDFFFE45ED5CBC7FBC2F5EA" blockId="7.[151,1436,1717,1771]" pageId="7" pageNumber="48">
<emphasis id="049A885CFFDFFFE45ED5CBC7FEA1F5C8" bold="true" box="[151,275,1717,1740]" pageId="7" pageNumber="48">FIGURE 4.</emphasis>
<taxonomicName id="F1EE2FCDFFDFFFE45F5BCBC4FDE6F5CF" box="[281,596,1718,1740]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="48" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFDFFFE45F5BCBC4FDE6F5CF" box="[281,596,1718,1740]" italics="true" pageId="7" pageNumber="48">Trypanosyllis microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFDFFFE45C18CBC4FD21F5C8" box="[602,659,1718,1739]" pageId="7" pageNumber="48" rank="species">
<emphasis id="049A885CFFDFFFE45C18CBC4FD21F5C8" bold="true" box="[602,659,1718,1739]" pageId="7" pageNumber="48">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Anterior chaetae. (B) Anterior aciculae. (C) Midbody chaetae. (D) Midbody acicula. (E) Posterior chaetae. (F) Posterior acicula. (G) Dorsal simple chaeta. Scale 40 µm.
</paragraph>
</caption>
<subSubSection id="7EF407C5FFDFFFE95E87CA65FD4FF3AC" lastPageId="10" lastPageNumber="51" pageId="7" pageNumber="48" type="discussion">
<paragraph id="3651544EFFDFFFE45E87CA65FF54F4E0" blockId="7.[151,1436,1815,2019]" pageId="7" pageNumber="48">
<emphasis id="049A885CFFDFFFE45E87CA65FE8BF42C" bold="true" box="[197,313,1815,1839]" pageId="7" pageNumber="48">Remarks.</emphasis>
This species is characterized by having the anterior pair of eyes in ventral position and the posterior one dorsally, as well as by having bidentate compound chaetae and a small trepan with transparent teeth. The genus
<taxonomicName id="F1EE2FCDFFDFFFE45ED5CA2DFE9CF474" box="[151,302,1887,1911]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="049A885CFFDFFFE45ED5CA2DFE9CF474" box="[151,302,1887,1911]" italics="true" pageId="7" pageNumber="48">Trypanosyllis</emphasis>
</taxonomicName>
currently has more than 20 valid species (
<bibRefCitation id="527F29BFFFDFFFE45D50CA2DFC24F474" author="Cinar" box="[786,918,1887,1911]" pageId="7" pageNumber="48" refString="Cinar, M. E. (2007) A new species of Trypanosyllis (Polychaeta: Syllidae) from the Levantine coast of Turkey (eastern Mediterranean). Journal of the Marine Biological Association of the United Kingdom, 87, 451 - 457." type="journal article" year="2007">Çinar 2007</bibRefCitation>
;
<bibRefCitation id="527F29BFFFDFFFE45DE0CA2DFB78F474" author="Nogueira" box="[930,1226,1887,1911]" pageId="7" pageNumber="48" refString="Nogueira, J. M. &amp; Fukuda, M. (2008) A new species of Trypanosyllis (Polychaeta: Syllidae) from Brazil, with a redescription of Brazilian material of Trypanosyllis zebra. Journal of the Marine Biological Association of the United Kingdom, 88, 913 - 924." type="journal article" year="2008">Nogueira &amp; Fukuda 2008</bibRefCitation>
). Of these, only
<taxonomicName id="F1EE2FCDFFDFFFE45BC5CA12FE7FF498" authority="Perkins, 1981" authorityName="Perkins" authorityYear="1981" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="48" phylum="Annelida" rank="species" species="parvidentata">
<emphasis id="049A885CFFDFFFE45BC5CA12FE94F498" italics="true" pageId="7" pageNumber="48">T. parvidentata</emphasis>
<bibRefCitation id="527F29BFFFDFFFE45F72CAF1FE7FF498" author="Perkins" box="[304,461,1923,1947]" pageId="7" pageNumber="48" refString="Perkins, T. H. (1981) Syllidae (Polychaeta), principally from Florida, with descriptions of a new genus and twenty-one new species. Proceedings of the Biological Society of Washington, 93 (4), 1080 - 1172." type="journal article" year="1981">Perkins, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F1EE2FCDFFDFFFE45C4BCAF6FCF9F498" authority="Claparede, 1868" authorityName="Claparede" authorityYear="1868" box="[521,843,1923,1947]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="48" phylum="Annelida" rank="species" species="coeliaca">
<emphasis id="049A885CFFDFFFE45C4BCAF6FD35F498" box="[521,647,1923,1947]" italics="true" pageId="7" pageNumber="48">T. coeliaca</emphasis>
<bibRefCitation id="527F29BFFFDFFFE45CD3CAF1FCF9F498" author="Claparede" box="[657,843,1923,1947]" pageId="7" pageNumber="48" refString="Claparede, E. (1868) Les Annelides Chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d´Histoire Naturelle de Geneve, 19 (2), 1 - 500." type="journal article" year="1868">Claparède, 1868</bibRefCitation>
</taxonomicName>
resemble
<taxonomicName id="F1EE2FCDFFDFFFE45D86CAF6FB13F498" box="[964,1185,1923,1947]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="48" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFDFFFE45D86CAF6FB13F498" box="[964,1185,1923,1947]" italics="true" pageId="7" pageNumber="48">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFDFFFE45AE9CAF6FB5FF49F" box="[1195,1261,1924,1948]" pageId="7" pageNumber="48" rank="species">
<emphasis id="049A885CFFDFFFE45AE9CAF6FB5FF49F" bold="true" box="[1195,1261,1924,1948]" pageId="7" pageNumber="48">n. sp.</emphasis>
</taxonomicNameLabel>
All three have small trepans, sinuous glands surrounding the pharynx, and antennae, dorsal cirri, pharynx and proventricle similar in size.
</paragraph>
<caption id="629104C6FFD0FFEB5ED5CBBDFCDBF443" httpUri="https://zenodo.org/record/201629/files/figure.png" pageId="8" pageNumber="49" targetBox="[160,1428,187,1727]" targetPageId="8">
<paragraph id="3651544EFFD0FFEB5ED5CBBDFCDBF443" blockId="8.[151,1436,1743,1858]" pageId="8" pageNumber="49">
<emphasis id="049A885CFFD0FFEB5ED5CBBDFEA0F5E6" bold="true" box="[151,274,1743,1766]" pageId="8" pageNumber="49">FIGURE 5.</emphasis>
SEM of
<taxonomicName id="F1EE2FCDFFD0FFEB5F2DCBA2FD18F5E5" box="[367,682,1744,1766]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD0FFEB5F2DCBA2FD18F5E5" box="[367,682,1744,1766]" italics="true" pageId="8" pageNumber="49">Trypanosyllis microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD0FFEB5CF2CBA2FD5BF5E6" box="[688,745,1744,1765]" pageId="8" pageNumber="49" rank="species">
<emphasis id="049A885CFFD0FFEB5CF2CBA2FD5BF5E6" bold="true" box="[688,745,1744,1765]" pageId="8" pageNumber="49">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Anterior end, dorsal view (Superior arrow: nuchal organs. Inferior arrow: ciliated band) (B) Anterior end, ventral view. (C) Detail of dorsum most anterior segments (Superior arrow: central ciliated band. Inferior arrow: lateral ciliated band). (D) Parapodial lobe (Arrow: ciliated band). (E) Anterior chaetae. (F) Posterior chaetae. Scale: (AB) 50 µm. (C) 100 µm. (DE) 10 µm. (F) 5 µm.
</paragraph>
</caption>
<paragraph id="3651544EFFD0FFEA5E87CA18FB8FF21F" blockId="8.[151,1437,1897,2030]" lastBlockId="9.[151,1436,151,824]" lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
However, unlike
<taxonomicName id="F1EE2FCDFFD0FFEB5FC8CA18FDD4F482" box="[394,614,1897,1921]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD0FFEB5FC8CA18FDD4F482" box="[394,614,1897,1921]" italics="true" pageId="8" pageNumber="49">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD0FFEB5C32CA18FD03F481" box="[624,689,1898,1922]" pageId="8" pageNumber="49" rank="species">
<emphasis id="049A885CFFD0FFEB5C32CA18FD03F481" bold="true" box="[624,689,1898,1922]" pageId="8" pageNumber="49">n. sp.</emphasis>
</taxonomicNameLabel>
, the other two species have two pairs of eyes in dorsal position. Additionally,
<taxonomicName id="F1EE2FCDFFD0FFEB5F72CAFCFE68F4A6" box="[304,474,1933,1957]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="parvidentata">
<emphasis id="049A885CFFD0FFEB5F72CAFCFE68F4A6" box="[304,474,1933,1957]" italics="true" pageId="8" pageNumber="49">T. parvidentata</emphasis>
</taxonomicName>
has a solitary acicula throughout the body. Unlike
<taxonomicName id="F1EE2FCDFFD0FFEB5A52CAFCFB5BF4A6" box="[1040,1257,1933,1957]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD0FFEB5A52CAFCFB5BF4A6" box="[1040,1257,1933,1957]" italics="true" pageId="8" pageNumber="49">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD0FFEB5AADCAFFFA9FF4A6" box="[1263,1325,1933,1957]" pageId="8" pageNumber="49" rank="species">
<emphasis id="049A885CFFD0FFEB5AADCAFFFA9FF4A6" bold="true" box="[1263,1325,1933,1957]" pageId="8" pageNumber="49">n. sp.</emphasis>
</taxonomicNameLabel>
, the anterior falcigers of
<taxonomicName id="F1EE2FCDFFD0FFEB5F0DCAC0FE79F4CA" box="[335,459,1969,1993]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="coeliaca">
<emphasis id="049A885CFFD0FFEB5F0DCAC0FE79F4CA" box="[335,459,1969,1993]" italics="true" pageId="8" pageNumber="49">T. coeliaca</emphasis>
</taxonomicName>
have slender and longer blades and longer spines on distal margin, while
<taxonomicName id="F1EE2FCDFFD0FFEB5B51CAC0FF71F4ED" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="parvidentata">
<emphasis id="049A885CFFD0FFEB5B51CAC0FF71F4ED" italics="true" pageId="8" pageNumber="49">T. parvidentata</emphasis>
</taxonomicName>
has longer falcigers with both teeth similar in size (in
<taxonomicName id="F1EE2FCDFFD0FFEB5D68CAA5FBB6F4ED" box="[810,1028,2006,2030]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="49" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD0FFEB5D68CAA5FBB6F4ED" box="[810,1028,2006,2030]" italics="true" pageId="8" pageNumber="49">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD0FFEB5A4ECAA4FBF9F4ED" box="[1036,1099,2006,2030]" pageId="8" pageNumber="49" rank="species">
<emphasis id="049A885CFFD0FFEB5A4ECAA4FBF9F4ED" bold="true" box="[1036,1099,2006,2030]" pageId="8" pageNumber="49">n. sp.</emphasis>
</taxonomicNameLabel>
, the subdistal tooth is smaller in most falcigers).
<taxonomicName id="F1EE2FCDFFD1FFEA5F2BCDEAFD5EF3AC" authority="Grube, 1860" authorityName="Grube" authorityYear="1860" box="[361,748,151,176]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="zebra">
<emphasis id="049A885CFFD1FFEA5F2BCDEAFDF7F3B3" box="[361,581,152,176]" italics="true" pageId="9" pageNumber="50">Trypanosyllis zebra</emphasis>
(
<bibRefCitation id="527F29BFFFD1FFEA5C17CDE5FD51F3AC" author="Grube" box="[597,739,151,175]" pageId="9" pageNumber="50" refString="Grube, A. E. (1860) Beschreibung neuer oder wenig bekannter Anneliden. Funter Beitrag. Archiv fur Naturgeschichte, 26, 71 - 118." type="journal article" year="1860">Grube, 1860</bibRefCitation>
)
</taxonomicName>
is one of the most common littoral syllids, even in the Mexican Pacific, and is indeed similar to
<taxonomicName id="F1EE2FCDFFD1FFEA5C75CDCFFCA0F3D7" box="[567,786,188,212]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD1FFEA5C75CDCFFCA0F3D7" box="[567,786,188,212]" italics="true" pageId="9" pageNumber="50">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD1FFEA5D59CDCEFCE9F3D7" box="[795,859,188,212]" pageId="9" pageNumber="50" rank="species">
<emphasis id="049A885CFFD1FFEA5D59CDCEFCE9F3D7" bold="true" box="[795,859,188,212]" pageId="9" pageNumber="50">n. sp.</emphasis>
</taxonomicNameLabel>
, but the former has a distinctive colour pattern, as well as much longer antennae and dorsal cirri. Furthermore, the pharynx in
<taxonomicName id="F1EE2FCDFFD1FFEA5DB4CD93FBFDF3FB" box="[1014,1103,224,248]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="zebra">
<emphasis id="049A885CFFD1FFEA5DB4CD93FBFDF3FB" box="[1014,1103,224,248]" italics="true" pageId="9" pageNumber="50">T. zebra</emphasis>
</taxonomicName>
is proportionally shorter and the proventricle is longer, and the species lacks the dorsal transverse ciliated bands.
</paragraph>
<paragraph id="3651544EFFD1FFEA5E87CC5AFD0EF187" blockId="9.[151,1436,151,824]" pageId="9" pageNumber="50">Most specimens were dissected and their pharynx examined. All had a small and thin pharyngeal tooth, very difficult to distinguish, and a trepan with about 10 very small and transparent teeth. Even after dissection of the pharynx, the trepan is very difficult to observe in most of the specimens, since sometimes the teeth are shorter than the length of the papillae and also because of its colour and texture, it can be easily confused with pharyngeal tissues. This could be due to the fact that the teeth of the trepan are made up of soft and fibrous tissues when they begin to develop in young organisms (Fig. 6C, indicated by arrow), becoming larger and rigid structures in adults (Fig. 6D). Therefore, such a structure can be overlooked and assumed to be absent in some specimens depending on their stage of development and it could also be lost accidentally (San Martín 2003). However, after careful examination of a large number of specimens, it was concluded that all have a trepan, even small individuals although they may be minute and difficult to see.</paragraph>
<paragraph id="3651544EFFD1FFEA5E87CFE2FCFAF034" blockId="9.[151,1436,151,824]" pageId="9" pageNumber="50">
In some species such as
<taxonomicName id="F1EE2FCDFFD1FFEA5F9BCFE3FD83F1AB" box="[473,561,656,680]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="zebra">
<emphasis id="049A885CFFD1FFEA5F9BCFE3FD83F1AB" box="[473,561,656,680]" italics="true" pageId="9" pageNumber="50">T. zebra</emphasis>
</taxonomicName>
, the presence of a pharyngeal tooth seems to be size-dependent, as mid-sized and large specimens lack one (
<bibRefCitation id="527F29BFFFD1FFEA5FACCFC6FD1DF1CF" author="San" box="[494,687,692,716]" pageId="9" pageNumber="50" refString="San Martin, G. (1984) Estudio biogeografico, faunistico y sistematico de los Poliquetos de la familia Syllidae (Annelida: Polychaeta) en Baleares. Ediciones de la Universidad Complutense de Madrid, 187, 529 pp." type="book" year="1984">San Martín 1984</bibRefCitation>
, 2003;
<bibRefCitation id="527F29BFFFD1FFEA5D43CFC6FC49F1CF" author="San" box="[769,1019,692,716]" pageId="9" pageNumber="50" refString="San Martin, G., Hutchings, P. &amp; Aguado, M. A. (2008) Syllinae (Polychaeta, Syllidae) from Australia. Part. 2. Genera Inermosyllis, Megasyllis n. gen., Opisthosyllis and Trypanosyllis. Zootaxa, 1840, 1 - 53." type="journal article" year="2008">
San Martín
<emphasis id="049A885CFFD1FFEA5DC4CFC7FC05F1CF" box="[902,951,692,716]" italics="true" pageId="9" pageNumber="50">et al</emphasis>
2008
</bibRefCitation>
). Nonetheless, in
<taxonomicName id="F1EE2FCDFFD1FFEA5A80CFC7FA2EF1CF" box="[1218,1436,692,716]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD1FFEA5A80CFC7FA2EF1CF" box="[1218,1436,692,716]" italics="true" pageId="9" pageNumber="50">T. microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD1FFEA5ED5CFAAFF64F1F3" box="[151,214,728,752]" pageId="9" pageNumber="50" rank="species">
<emphasis id="049A885CFFD1FFEA5ED5CFAAFF64F1F3" bold="true" box="[151,214,728,752]" pageId="9" pageNumber="50">n. sp.</emphasis>
</taxonomicNameLabel>
the pharyngeal tooth is always present, while the only size-related differences is trepan teeth. The trepan is a diagnostic character of the genus (San Martín 2003), and it is considered homoplasic, so that it should be used with care to differentiate lower taxa (
<bibRefCitation id="527F29BFFFD1FFEA5FBFCE6DFC8EF034" author="Aguado" box="[509,828,799,823]" pageId="9" pageNumber="50" refString="Aguado, M. T. &amp; San Martin, G. (2009) Phylogeny of Syllidae (Polychaeta) based on morphological data. Zoologica Scripta, 38 (4), 379 - 402." type="journal article" year="2009">Aguado &amp; San Martín 2009</bibRefCitation>
).
</paragraph>
<paragraph id="3651544EFFD1FFEA5F43CAF4FF0DF4D4" blockId="9.[151,1436,1925,2007]" pageId="9" pageNumber="50">
<emphasis id="049A885CFFD1FFEA5F43CAF4FEA6F498" bold="true" box="[257,276,1926,1947]" pageId="9" pageNumber="50">6.</emphasis>
SEM of
<taxonomicName id="F1EE2FCDFFD1FFEA5F37CAF4FD00F49F" box="[373,690,1926,1948]" class="Polychaeta" family="Syllidae" genus="Trypanosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="50" phylum="Annelida" rank="species" species="microdenticulata" status="sp. nov.">
<emphasis id="049A885CFFD1FFEA5F37CAF4FD00F49F" box="[373,690,1926,1948]" italics="true" pageId="9" pageNumber="50">Trypanosyllis microdenticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FA93527FFD1FFEA5CF8CAF4FD47F498" box="[698,757,1926,1947]" pageId="9" pageNumber="50" rank="species">
<emphasis id="049A885CFFD1FFEA5CF8CAF4FD47F498" bold="true" box="[698,757,1926,1947]" pageId="9" pageNumber="50">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Dorsal simple chaeta. (B) Ventral simple chaeta. (C) Everted pharynx, small specimen (Arrow: soft teeth). (D) Hard teeth, larger specimen. Scale: (A) 2 µm. (B) 1 µm. (C) 10 µm. (D) 20 µm.
</paragraph>
<paragraph id="3651544EFFD2FFE95E87CDE5FD4FF3AC" blockId="10.[197,1209,151,248]" box="[197,765,151,176]" pageId="10" pageNumber="51">
<emphasis id="049A885CFFD2FFE95E87CDE5FE9AF3B3" bold="true" box="[197,296,151,176]" pageId="10" pageNumber="51">Habitat.</emphasis>
Subtidal coarse sand,
<quantity id="F116F9ABFFD2FFE95C61CDE5FD02F3AC" box="[547,688,151,175]" metricMagnitude="1" metricUnit="m" metricValue="1.125" metricValueMax="1.2" metricValueMin="1.05" pageId="10" pageNumber="51" unit="m" value="11.25" valueMax="12.0" valueMin="10.5">10.5 to 12 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="7EF407C5FFD2FFE95E87CDC9FB0BF3F4" pageId="10" pageNumber="51" type="distribution">
<paragraph id="3651544EFFD2FFE95E87CDC9FB0BF3F4" blockId="10.[197,1209,151,248]" pageId="10" pageNumber="51">
<emphasis id="049A885CFFD2FFE95E87CDC9FEECF3D7" bold="true" box="[197,350,187,212]" pageId="10" pageNumber="51">Distribution.</emphasis>
Western coast of Acapulco Bay, Southern Mexican Pacific.
<emphasis id="049A885CFFD2FFE95E87CD92FEF9F3FB" bold="true" box="[197,331,224,248]" pageId="10" pageNumber="51">Etymology.</emphasis>
The specific name refers to the small size of the teeth belonging to the trepan.
</paragraph>
</subSubSection>
</treatment>
</document>