496 lines
64 KiB
XML
496 lines
64 KiB
XML
<document id="7174D4CBAC51EDEC786C10B498760E49" ID-DOI="10.1649/0010-065X-77.4.629" ID-ISSN="1938-4394" ID-Zenodo-Dep="13522797" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1722968791930" checkinUser="felipe" docAuthor="Arias-Buriticá, Jorge Armando, Bach, Andressa, Peres, Carlos A., Haugaasen, Torbjørn, Hawes, Joseph E., Azevedo, Renato A. & Vaz-De-Mello, Fernando Z." docDate="2023" docId="03E787CDFFC1FFDEFD59E69CA5FAFC19" docLanguage="en" docName="TheColeopBull.77.4.629-635.pdf" docOrigin="The Coleopterists Bulletin 77 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-77.4.629" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Isocopris rossinii Arias-Buritica, Bach" docType="treatment" docVersion="4" lastPageNumber="633" masterDocId="FFDEFFB5FFC3FFDBFF9CE672A652FF96" masterDocTitle="Figs. 4 - 10 in Descriptions of Two New Species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) in Korea" masterLastPageNumber="635" masterPageNumber="629" pageNumber="630" updateTime="1724955499975" updateUser="ExternalLinkService">
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<mods:title id="64D4ADDEA0BB62F2D5397E887C2B5590">Figs. 4 - 10 in Descriptions of Two New Species of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) in Korea</mods:title>
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<mods:namePart id="71EA42A568D7BE86932C7B7401399EBD">Arias-Buriticá, Jorge Armando</mods:namePart>
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<mods:namePart id="E4268739289A68802DB00EE25B95E014">Bach, Andressa</mods:namePart>
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<mods:namePart id="5BCFAE41C616B3F96B80C51A294E240A">Peres, Carlos A.</mods:namePart>
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<mods:namePart id="67FFE161C5E6A92DF5A6553369DBB6C7">Haugaasen, Torbjørn</mods:namePart>
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<mods:title id="FC3FC43CF69539306BC867F563335C2F">The Coleopterists Bulletin</mods:title>
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<taxonomicName id="4C4E4D58FFC1FFD9FD59E69CA218FE95" ID-CoL="BXTZN" authority="Arias-Buritica, Bach" authorityName="Arias-Buritica, Bach" box="[709,1098,237,259]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="species" species="rossinii">
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<emphasis id="B93AEAC9FFC1FFD9FD59E69CA53CFE95" bold="true" box="[709,878,238,259]" italics="true" pageId="2" pageNumber="630">Isocopris rossinii</emphasis>
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Arias-Buriticá, Bach
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, and Vaz-de-Mello, new species
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(
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<figureCitation id="13752A5EFFC1FFD9FCEBE72AA59AFEF8" box="[887,968,344,366]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[197,229,1704,1723]" captionStartId-1="4.[176,207,1663,1682]" captionStartId-2="5.[197,228,1687,1706]" captionTargetBox-0="[279,1103,205,1672]" captionTargetBox-1="[161,1145,913,1633]" captionTargetBox-2="[192,1173,969,1655]" captionTargetId-0="figure-7@3.[173,1193,193,1686]" captionTargetId-1="figure-391@4.[151,1171,901,1645]" captionTargetId-2="figure-443@5.[173,1193,954,1669]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." captionText-1="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." captionText-2="Fig. 3. Distribution map for Isocopris rossinii, new species. Plot A and Plot B represent the collection sites in Serra do Divisor National Park, Acre, Brazil." figureDoi-0="http://doi.org/10.5281/zenodo.13522805" figureDoi-1="http://doi.org/10.5281/zenodo.13522811" figureDoi-2="http://doi.org/10.5281/zenodo.13522819" httpUri-0="https://zenodo.org/record/13522805/files/figure.png" httpUri-1="https://zenodo.org/record/13522811/files/figure.png" httpUri-2="https://zenodo.org/record/13522819/files/figure.png" pageId="2" pageNumber="630">Fig. 1–3</figureCitation>
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)
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<subSubSection id="C3546550FFC1FFD9FD59E7F4A210FE7D" pageId="2" pageNumber="630" type="etymology">
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<paragraph id="8BF136DBFFC1FFD9FD59E7F4A210FE7D" blockId="2.[685,1172,390,1644]" pageId="2" pageNumber="630">
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<emphasis id="B93AEAC9FFC1FFD9FD59E7F4A56BFE0D" bold="true" box="[709,825,390,411]" pageId="2" pageNumber="630">Etymology.</emphasis>
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This species is named in honor of Dr. Michele Rossini, Italian scarabaeoidologist and good friend of the authors, who carried out the taxonomic revision of
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<taxonomicName id="4C4E4D58FFC1FFD9FCF5E7A4A590FE7D" authorityName="PEREIRA AND MARTINEZ" authorityYear="1960" box="[873,962,470,491]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="630" phylum="Arthropoda" rank="genus">
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<emphasis id="B93AEAC9FFC1FFD9FCF5E7A4A590FE7D" box="[873,962,470,491]" italics="true" pageId="2" pageNumber="630">Isocopris</emphasis>
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with FZVM.
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<subSubSection id="C3546550FFC1FFD9FD59E781A5CDFCB8" pageId="2" pageNumber="630" type="diagnosis">
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<paragraph id="8BF136DBFFC1FFD9FD59E781A5CDFCB8" blockId="2.[685,1172,390,1644]" pageId="2" pageNumber="630">
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<emphasis id="B93AEAC9FFC1FFD9FD59E781A57CFD9F" bold="true" box="[709,814,499,521]" pageId="2" pageNumber="630">Diagnosis.</emphasis>
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This species is recognized by the following combination of characters: head with fronto-clypeal carina and deep depression in the frons (
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<figureCitation id="13752A5EFFC1FFD9FD75E431A54FFDCE" box="[745,797,579,601]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1</figureCitation>
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), both sexes with head margin distinctly notched at the clypeo-genal junction (
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<figureCitation id="13752A5EFFC1FFD9FB82E42CA2D4FDE5" box="[1054,1158,606,628]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Figs. 1a, c</figureCitation>
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), ventral clypeal process in lateral view large and trapezoidal (
|
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<figureCitation id="13752A5EFFC1FFD9FCBAE4E1A5DFFD3F" box="[806,909,659,681]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Figs. 1e, f</figureCitation>
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), metaventral anterior lobe glabrous (
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<figureCitation id="13752A5EFFC1FFD9FC97E4DCA522FD52" box="[779,880,686,708]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Figs. 1b, d</figureCitation>
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), morphology of the male genital organ as in
|
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<figureCitation id="13752A5EFFC1FFD9FCCCE4BAA5DCFD48" box="[848,910,712,734]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="2" pageNumber="630">Fig. 2</figureCitation>
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. Females show secondary sexual dimorphism: head with triangular shape, elytral striae from the first to the fourth widened in the central area (
|
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<figureCitation id="13752A5EFFC1FFD9FCD1E56AA5C0FCB8" box="[845,914,792,814]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1c</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3546550FFC1FFDEFD59E547A7E8FE3B" lastPageId="5" lastPageNumber="633" pageId="2" pageNumber="630" type="description">
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<paragraph id="8BF136DBFFC1FFDFFD59E547A5FDFD7A" blockId="2.[685,1172,390,1644]" lastBlockId="4.[685,1172,194,858]" lastPageId="4" lastPageNumber="632" pageId="2" pageNumber="630">
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<emphasis id="B93AEAC9FFC1FFD9FD59E547A510FCDD" bold="true" box="[709,834,821,843]" pageId="2" pageNumber="630">Description.</emphasis>
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Male. Length
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<quantity id="4CB69B3EFFC1FFD9FC51E544A277FCDD" box="[973,1061,822,844]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="2" pageNumber="630" unit="mm" value="19.6">19.6 mm</quantity>
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, width
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<quantity id="4CB69B3EFFC1FFD9FBF6E544A480FCF0" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="630" unit="mm" value="11.7">11.7 mm</quantity>
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(
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<figureCitation id="13752A5EFFC1FFD9FD43E522A515FCF0" box="[735,839,848,870]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Figs. 1a, b</figureCitation>
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). Body color black and shiny, with brown setae on both sides of head and pronotum (
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<figureCitation id="13752A5EFFC1FFD9FD28E5F4A4AFFC0D" box="[692,765,902,924]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1a</figureCitation>
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).
|
||
<emphasis id="B93AEAC9FFC1FFD9FC88E5F4A500FC0D" bold="true" box="[788,850,902,923]" pageId="2" pageNumber="630">Head:</emphasis>
|
||
Wider than long. Antenna with eight antennomeres. Clypeus emarginate at middle, with two obtuse teeth. Surface of clypeus shiny with smooth wrinkles giving it a wavy appearance. Ventral clypeal process in lateral view large and trapezoidal (
|
||
<figureCitation id="13752A5EFFC1FFD9FCADE279A5D3FBB6" box="[817,897,1035,1057]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1e</figureCitation>
|
||
). Head margin distinctly notched at the clypeo-genal junction, genal margin broader than clypeus. Fronto-clypeal region with a central and transverse carina of about
|
||
<quantity id="4CB69B3EFFC1FFD9FB83E229A23BFBE7" box="[1055,1129,1115,1137]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="630" unit="mm" value="2.5">2.5 mm</quantity>
|
||
and apical ridge flattened (
|
||
<figureCitation id="13752A5EFFC1FFD9FCE3E204A590FB1D" box="[895,962,1142,1164]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1e</figureCitation>
|
||
). Clypeo-genal suture clearly visible. Frons with deep and distinct depression (
|
||
<figureCitation id="13752A5EFFC1FFD9FD7EE2D9A574FB56" box="[738,806,1195,1217]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="2" pageNumber="630">Fig. 1e</figureCitation>
|
||
).
|
||
<emphasis id="B93AEAC9FFC1FFD9FCA5E2D9A5FAFB56" bold="true" box="[825,936,1195,1216]" pageId="2" pageNumber="630">Pronotum:</emphasis>
|
||
Wider than long, simple and convex, anteromedial region without humps. Anterior angles acute. Pronotal punctures dense, very shallow and separated by no more than twice their diameter, near posterior margin with shallow and elliptical ocellate punctures.
|
||
<emphasis id="B93AEAC9FFC1FFD9FB90E343A2C0FAD0" bold="true" box="[1036,1170,1329,1350]" pageId="2" pageNumber="630">Hypomeron:</emphasis>
|
||
Shagreened surface with setose punctures spaced about by their diameters, denser on lateral margins, glabrous and smooth in the central area.
|
||
<emphasis id="B93AEAC9FFC1FFD9FD32E3E9A57DFA26" bold="true" box="[686,815,1435,1456]" pageId="2" pageNumber="630">Prosternum:</emphasis>
|
||
Shagreened surface with smooth central area, glabrous.
|
||
<emphasis id="B93AEAC9FFC1FFD9FCF8E3C7A5A1FA5D" bold="true" box="[868,1011,1461,1483]" pageId="2" pageNumber="630">Mesoventrite:</emphasis>
|
||
Shagreened surface. Smooth in the central area and with large, deep ocellate punctures separated by less than their diameters in the lateral areas.
|
||
<emphasis id="B93AEAC9FFC1FFD9FC4FE077A2C0F98D" bold="true" box="[979,1170,1541,1563]" pageId="2" pageNumber="630">Mesanepisternum:</emphasis>
|
||
Shagreened surface with setose punctures separated by less than one-half their diameter.
|
||
<emphasis id="B93AEAC9FFC1FFD9FB99E049A2C0F9C7" bold="true" box="[1029,1170,1595,1617]" pageId="2" pageNumber="630">Metaventrite:</emphasis>
|
||
Anterior lobe with shagreened surface with wrinkles near suture with mesoventrite. With shallow punctures separated by two and three times their diameters in the lateral areas near the insertion of the mesocoxa. Central area with shiny, glabrous surface (
|
||
<figureCitation id="13752A5EFFC7FFDFFF03E75FA6B6FED5" box="[159,228,301,323]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="4" pageNumber="632">Fig. 1b</figureCitation>
|
||
). Metaventral lateral margins with strongly shagreened surface with large and deep ocellatesetose punctures separated by less than their diameter.
|
||
<emphasis id="B93AEAC9FFC7FFDFFF54E70FA7D5FE05" bold="true" box="[200,391,381,403]" pageId="4" pageNumber="632">Metanepisternum:</emphasis>
|
||
Shagreened surface with large, deep ocellate punctures separated by about their diameters, some points with short setae.
|
||
<emphasis id="B93AEAC9FFC7FFDFFF04E7BFA6B3FE74" bold="true" box="[152,225,461,482]" pageId="4" pageNumber="632">Elytra:</emphasis>
|
||
Striae bicarinate, distinctly impressed and with shagreened surface, with small ocellate punctures throughout, spaced about three to four times their diameters. Interstriae with shagreened surface, with small and shallow punctures, separated by two to three times their diameters.
|
||
<emphasis id="B93AEAC9FFC7FFDFFE35E421A440FDFE" bold="true" box="[425,530,595,616]" pageId="4" pageNumber="632">Abdomen:</emphasis>
|
||
Abdominal ventrites curved and projected at the lateral margin. Shagreened surface with shallow punctures. Anterior area from first to fifth ventrite with a row of large, deep ocellate punctures separated by less than their diameters and denser on the lateral margins. Sixth ventrite compressed in the middle region.
|
||
<emphasis id="B93AEAC9FFC7FFDFFF04E57CA750FCB2" bold="true" box="[152,258,782,804]" pageId="4" pageNumber="632">Pygidium:</emphasis>
|
||
Pygidial margin complete. Surface shiny, with small, shallow punctures separated by three times their diameters throughout.
|
||
<emphasis id="B93AEAC9FFC7FFDFFD98E536A468FCCF" bold="true" box="[516,570,836,857]" pageId="4" pageNumber="632">Legs:</emphasis>
|
||
Apical margin of protibia obliquely truncate, inner and apical angles finely acuminate, with few straight setae at the apex. Apical spur abruptly angled near the apex (
|
||
<figureCitation id="13752A5EFFC7FFDFFD74E760A57EFEB1" box="[744,812,274,296]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="4" pageNumber="632">Fig. 1a</figureCitation>
|
||
). Tarsomere 1 of meso- and metalegs as long as the second and third tarsomeres together. Metatibial spur with apical bifurcation (
|
||
<figureCitation id="13752A5EFFC7FFDFFBA2E735A2D4FECA" box="[1086,1158,327,349]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="4" pageNumber="632">Fig. 1a</figureCitation>
|
||
).
|
||
<emphasis id="B93AEAC9FFC7FFDFFD31E713A54AFEE1" bold="true" box="[685,792,353,375]" pageId="4" pageNumber="632">Aedeagus:</emphasis>
|
||
Parameres subtriangular in lateral view (
|
||
<figureCitation id="13752A5EFFC7FFDFFD28E70EA4ADFE07" box="[692,767,380,402]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2a</figureCitation>
|
||
). In dorsal view parameres symmetrical with curved apex (
|
||
<figureCitation id="13752A5EFFC7FFDFFCFDE7E5A5FBFE3B" box="[865,937,407,429]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2b</figureCitation>
|
||
). In ventral view subgenital plate present, parameres symmetrical with curved apex and base with spine-shaped projection (
|
||
<figureCitation id="13752A5EFFC7FFDFFD28E795A4A5FE6A" box="[692,759,487,509]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2c</figureCitation>
|
||
).
|
||
<emphasis id="B93AEAC9FFC7FFDFFC94E795A5C4FE6B" bold="true" box="[776,918,487,509]" pageId="4" pageNumber="632">Endophallites:</emphasis>
|
||
Lamella copulatrix (LC) is large, asymmetrical with two sclerotized processes and a fleshy process with many bristles (
|
||
<figureCitation id="13752A5EFFC7FFDFFBA2E46EA2DAFDA4" box="[1086,1160,540,562]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2d</figureCitation>
|
||
). Sclerites of axial and subaxial complex (A+SA complex) with indeterminate shape, more sclerotized in the central area surrounded by a semi-sclerotized membrane (
|
||
<figureCitation id="13752A5EFFC7FFDFFC81E4F5A50DFD0B" box="[797,863,647,669]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2f</figureCitation>
|
||
). Superior-right peripheral sclerite (SRP) with a “C” shape, wide and with undefined margins (
|
||
<figureCitation id="13752A5EFFC7FFDFFC9BE4CEA51FFD47" box="[775,845,700,722]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">Fig. 2e</figureCitation>
|
||
). Fronto-lateral peripheral sclerite (FLP) as shown in
|
||
<figureCitation id="13752A5EFFC7FFDFFCF8E4A5A5F8FD7A" box="[868,938,727,749]" captionStart="Fig" captionStartId="4.[176,207,1663,1682]" captionTargetBox="[161,1145,913,1633]" captionTargetId="figure-391@4.[151,1171,901,1645]" captionTargetPageId="4" captionText="Fig. 2. Isocopris rossinii, new species. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632">
|
||
Fig.
|
||
<quantity id="4CB69B3EFFC7FFDFFC0CE4A5A5F8FD7A" box="[912,938,727,748]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="632" unit="g" value="2.0">2g</quantity>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF316653FFC0FFD8FF59E0DAA279F942" ID-DOI="http://doi.org/10.5281/zenodo.13522805" ID-Zenodo-Dep="13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="3" pageNumber="631" startId="3.[197,229,1704,1723]" targetBox="[279,1103,205,1672]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BF136DBFFC0FFD8FF59E0DAA279F942" blockId="3.[173,1193,1704,1748]" pageId="3" pageNumber="631">
|
||
<emphasis id="B93AEAC9FFC0FFD8FF59E0DAA754F92D" bold="true" box="[197,262,1704,1723]" pageId="3" pageNumber="631">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C4E4D58FFC0FFD8FE80E0DBA7EFF92D" authorityName="Kim & Ahn" authorityYear="2023" box="[284,445,1705,1723]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="631" phylum="Arthropoda" rank="species" species="rossinii" status="sp. nov.">
|
||
<emphasis id="B93AEAC9FFC0FFD8FE80E0DBA7EFF92D" box="[284,445,1705,1723]" italics="true" pageId="3" pageNumber="631">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B93AEAC9FFC0FFD8FE53E0DAA410F92D" bold="true" box="[463,578,1704,1723]" pageId="3" pageNumber="631">
|
||
<taxonomicNameLabel id="A20957B2FFC0FFD8FE53E0DAA410F92D" box="[463,578,1704,1723]" pageId="3" pageNumber="631" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF136DBFFC7FFDEFD59E486A7E8FE3B" blockId="4.[685,1172,194,858]" lastBlockId="5.[173,660,193,911]" lastPageId="5" lastPageNumber="633" pageId="4" pageNumber="632">
|
||
<emphasis id="B93AEAC9FFC7FFDFFD59E486A57EFC9C" bold="true" box="[709,812,756,778]" pageId="4" pageNumber="632">Variation.</emphasis>
|
||
In males the length varies from
|
||
<quantity id="4CB69B3EFFC7FFDFFBC1E486A557FCB2" metricMagnitude="-2" metricUnit="m" metricValue="1.86" metricValueMax="1.97" metricValueMin="1.75" pageId="4" pageNumber="632" unit="mm" value="18.6" valueMax="19.7" valueMin="17.5">17.5– 19.7 mm</quantity>
|
||
and the width from
|
||
<quantity id="4CB69B3EFFC7FFDFFC4FE57DA202FCB2" box="[979,1104,783,805]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="4" pageNumber="632" unit="mm" value="10.75" valueMax="11.5" valueMin="10.0">10–11.5 mm</quantity>
|
||
. Some males have a shorter cephalic process (less than
|
||
<quantity id="4CB69B3EFFC7FFDFFBE9E55BA486FCCF" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="632" unit="mm" value="1.5">1.5 mm</quantity>
|
||
) and the apical ridge of the carina is bicuspid.
|
||
<emphasis id="B93AEAC9FFC6FFDEFF31E6B0A752FF41" bold="true" box="[173,256,194,215]" pageId="5" pageNumber="633">Female:</emphasis>
|
||
Length 18.0–
|
||
<quantity id="4CB69B3EFFC6FFDEFE17E6B0A7B7FF41" box="[395,485,194,215]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="5" pageNumber="633" unit="mm" value="21.1">21.1 mm</quantity>
|
||
. Width
|
||
<quantity id="4CB69B3EFFC6FFDEFDA8E6B3A687FF64" metricMagnitude="-2" metricUnit="m" metricValue="1.115" metricValueMax="1.15" metricValueMin="1.08" pageId="5" pageNumber="633" unit="mm" value="11.15" valueMax="11.5" valueMin="10.8">10.8–11.5 mm</quantity>
|
||
. Females differ from males in having a less rounded head, giving the appearance of being triangular, while the cephalic carina changes as mentioned in the variation of males (
|
||
<figureCitation id="13752A5EFFC6FFDEFE40E75EA44FFED4" box="[476,541,300,322]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="5" pageNumber="633">Fig. 1f</figureCitation>
|
||
). Secondary sexual dimorphism present, with the elytral striae from the first to the fourth widened in the central area (
|
||
<figureCitation id="13752A5EFFC6FFDEFF79E70EA77EFE07" box="[229,300,380,402]" captionStart="Fig" captionStartId="3.[197,229,1704,1723]" captionTargetBox="[279,1103,205,1672]" captionTargetId="figure-7@3.[173,1193,193,1686]" captionTargetPageId="3" captionText="Fig. 1. Isocopris rossinii, new species. Male: a) Dorsal view, b) Ventral view, e) Head, lateral view. Female: c) Dorsal view, d) Ventral view, f) Head, lateral view. Scale bars: a, b, d, e = 5 mm; c, f = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13522805" httpUri="https://zenodo.org/record/13522805/files/figure.png" pageId="5" pageNumber="633">Fig. 1c</figureCitation>
|
||
). Sixth abdominal ventrite not compressed towards the middle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF316653FFC7FFDFFF2CE00DA4FEF954" ID-DOI="http://doi.org/10.5281/zenodo.13522811" ID-Zenodo-Dep="13522811" httpUri="https://zenodo.org/record/13522811/files/figure.png" pageId="4" pageNumber="632" startId="4.[176,207,1663,1682]" targetBox="[161,1145,913,1633]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BF136DBFFC7FFDFFF2CE00DA4FEF954" blockId="4.[152,1172,1663,1730]" pageId="4" pageNumber="632">
|
||
<emphasis id="B93AEAC9FFC7FFDFFF2CE00DA6B8F904" bold="true" box="[176,234,1663,1682]" pageId="4" pageNumber="632">Fig. 2.</emphasis>
|
||
<taxonomicName id="4C4E4D58FFC7FFDFFE9CE00DA7C7F907" authorityName="Kim & Ahn" authorityYear="2023" box="[256,405,1663,1681]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="632" phylum="Arthropoda" rank="species" species="rossinii" status="sp. nov.">
|
||
<emphasis id="B93AEAC9FFC7FFDFFE9CE00DA7C7F907" box="[256,405,1663,1681]" italics="true" pageId="4" pageNumber="632">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B93AEAC9FFC7FFDFFE3EE00DA45FF904" bold="true" box="[418,525,1663,1682]" pageId="4" pageNumber="632">
|
||
<taxonomicNameLabel id="A20957B2FFC7FFDFFE3EE00DA45FF904" box="[418,525,1663,1682]" pageId="4" pageNumber="632" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Aedeagus: a) Lateral view, b) Dorsal view, c) Ventral view. d) Lamellae copulatrix, e) Superior-right peripheral sclerite (SRP), f) Sclerites of axial and subaxial complex (A+SA complex), g) Fronto-lateral peripheral sclerite (FLP). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3546550FFC6FFDEFF59E7C6A77EFC19" pageId="5" pageNumber="633" type="materials_examined">
|
||
<paragraph id="8BF136DBFFC6FFDEFF59E7C6A77EFC19" blockId="5.[173,660,193,911]" pageId="5" pageNumber="633">
|
||
<emphasis id="B93AEAC9FFC6FFDEFF59E7C6A704FE5C" bold="true" box="[197,342,436,458]" pageId="5" pageNumber="633">Type Material.</emphasis>
|
||
<materialsCitation id="3B263C86FFC6FFDEFEC6E7C6A45AFDC6" collectingDate="2019-12-11" collectingDateMax="2019-12-15" collectingDateMin="2019-12-11" collectionCode="FIT, R" collectorName="A. Azevedo. Plot" country="Brazil" county="Labels" latitude="-7.446111" location="Par" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" pageId="5" pageNumber="633" specimenCode="FIT 1-3" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="holotype">
|
||
<emphasis id="B93AEAC9FFC6FFDEFEC6E7C6A7EEFE5F" bold="true" box="[346,444,436,457]" pageId="5" pageNumber="633">
|
||
<typeStatus id="54F58879FFC6FFDEFEC6E7C6A7EAFE5F" box="[346,440,436,457]" pageId="5" pageNumber="633" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D48FD52FFC6FFDEFE5CE7C6A7B2FE5F" box="[448,480,436,457]" count="1" pageId="5" pageNumber="633" type="male">1♂</specimenCount>
|
||
.
|
||
<collectingCounty id="62904E57FFC6FFDEFE7BE7C6A47AFE5C" box="[487,552,436,458]" pageId="5" pageNumber="633">Labels</collectingCounty>
|
||
:1: {printed text on white label}
|
||
<collectingCountry id="F359764BFFC6FFDEFEECE7BDA79BFE72" box="[368,457,463,484]" name="Brazil" pageId="5" pageNumber="633">BRASIL</collectingCountry>
|
||
:
|
||
<collectingRegion id="498AF839FFC6FFDEFE4CE7BDA453FE72" box="[464,513,463,484]" country="Brazil" name="Acre" pageId="5" pageNumber="633">Acre</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B95ACA1FFC6FFDEFD97E7BDA4DDFE73" box="[523,655,463,485]" pageId="5" pageNumber="633">Mâncio Lima</collectingMunicipality>
|
||
,
|
||
<location id="8E916000FFC6FFDEFF31E798A682FE69" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFF31E798A682FE69" box="[173,208,490,511]" country="Brazil" county="Labels" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Par" pageId="5" pageNumber="633" stateProvince="Acre">Par</location>
|
||
[que]
|
||
<location id="8E916000FFC6FFDEFE97E798A77BFD96" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFE97E798A77BFD96" box="[267,297,490,512]" country="Brazil" county="Labels" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Na" pageId="5" pageNumber="633" stateProvince="Acre">Na</location>
|
||
[cional]
|
||
<location id="8E916000FFC6FFDEFEE1E798A477FD96" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFEE1E798A477FD96" box="[381,549,490,512]" country="Brazil" county="Labels" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Serra do Divisor" pageId="5" pageNumber="633" stateProvince="Acre">Serra do Divisor</location>
|
||
<geoCoordinate id="EE7A501CFFC6FFDEFDACE79AA4C1FE69" box="[560,659,488,511]" degrees="7" direction="south" minutes="26" orientation="latitude" pageId="5" pageNumber="633" precision="15" seconds="46" value="-7.446111">7°26′46″S</geoCoordinate>
|
||
<geoCoordinate id="EE7A501CFFC6FFDEFF31E471A777FD8C" box="[173,293,515,538]" degrees="73" direction="west" minutes="39" orientation="longitude" pageId="5" pageNumber="633" precision="15" seconds="28" value="-73.657776">73°39′28″W</geoCoordinate>
|
||
,
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFEAAE477A709FD8C" box="[310,347,517,538]" pageId="5" pageNumber="633">FIT</collectionCode>
|
||
.
|
||
<date id="FFF0101BFFC6FFDEFEF7E476A7ABFD8C" box="[363,505,516,538]" pageId="5" pageNumber="633" value="2019-12-11" valueMax="2019-12-15" valueMin="2019-12-11">
|
||
<collectingDate id="EFB4E9F3FFC6FFDEFEF7E476A7ABFD8C" box="[363,505,516,538]" pageId="5" pageNumber="633" value="2019-12-11" valueMax="2019-12-15" valueMin="2019-12-11">11-15.xii.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFD96E476A449FD8C" box="[522,539,516,538]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="633">R</collectionCode>
|
||
.
|
||
<collectorName id="26BB530DFFC6FFDEFD82E476A686FDA3" pageId="5" pageNumber="633">A.Azevedo. Plot</collectorName>
|
||
A
|
||
<specimenCode id="DBE89EA0FFC6FFDEFF6CE46DA768FDA2" box="[240,314,543,564]" collectionCode="FIT" pageId="5" pageNumber="633">FIT 1-3</specimenCode>
|
||
(250) / 2: {printed and handwritten text on red label with black margins
|
||
</materialsCitation>
|
||
}
|
||
<materialsCitation id="3B263C86FFC6FFDEFD88E448A442FD12" collectionCode="CEMT" country="Brazil" pageId="5" pageNumber="633" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="holotype">
|
||
<typeStatus id="54F58879FFC6FFDEFD88E448A4C1FDD9" box="[532,659,570,591]" pageId="5" pageNumber="633" type="holotype">HOLOTYPE</typeStatus>
|
||
<specimenCount id="9D48FD52FFC6FFDEFF31E427A6EDFDFC" box="[173,191,597,618]" count="1" pageId="5" pageNumber="633" type="male">♂</specimenCount>
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFF5BE427A722FDFC" authorityName="Kim & Ahn" authorityYear="2023" box="[199,368,597,618]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii" status="sp. nov.">
|
||
<emphasis id="B93AEAC9FFC6FFDEFF5BE427A722FDFC" box="[199,368,597,618]" italics="true" pageId="5" pageNumber="633">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957B2FFC6FFDEFEE4E427A797FDFC" box="[376,453,597,618]" pageId="5" pageNumber="633" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Arias-BuriticÁ, Bach and Vaz-de-Mello, 2023 [
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFE21E41DA458FD12" box="[445,522,623,644]" pageId="5" pageNumber="633">CEMT</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B263C86FFC6FFDEFD83E41DA7C0FC9F" collectingDate="2019-12-07" collectingDateMax="2019-12-11" collectingDateMin="2019-12-07" collectionCode="CEMT" collectorName="RA Azevedo." country="Brazil" county="Same" elevation="236" latitude="-7.446111" location="Par" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" pageId="5" pageNumber="633" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Acre" typeStatus="paratype">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD83E41DA4C0FD12" bold="true" box="[543,658,623,644]" pageId="5" pageNumber="633">
|
||
<typeStatus id="54F58879FFC6FFDEFD83E41DA4DCFD12" box="[543,654,623,644]" pageId="5" pageNumber="633" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D48FD52FFC6FFDEFF29E4F8A6B5FD09" box="[181,231,650,671]" count="2" pageId="5" pageNumber="633" type="male">2♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D48FD52FFC6FFDEFF6CE4F8A773FD09" box="[240,289,650,671]" count="4" pageId="5" pageNumber="633" type="female">4♀♀</specimenCount>
|
||
):
|
||
<collectingCounty id="62904E57FFC6FFDEFEAFE4F8A73AFD09" box="[307,360,650,671]" pageId="5" pageNumber="633">Same</collectingCounty>
|
||
data as holotype except:
|
||
<date id="FFF0101BFFC6FFDEFDC8E4F8A6ADFD2C" pageId="5" pageNumber="633" value="2019-12-07" valueMax="2019-12-11" valueMin="2019-12-07">
|
||
<collectingDate id="EFB4E9F3FFC6FFDEFDC8E4F8A6ADFD2C" pageId="5" pageNumber="633" value="2019-12-07" valueMax="2019-12-11" valueMin="2019-12-07">07-11- xii.2019</collectingDate>
|
||
</date>
|
||
.
|
||
<specimenCount id="9D48FD52FFC6FFDEFE92E4D6A77CFD2F" box="[270,302,676,697]" count="1" pageId="5" pageNumber="633" type="male">1♂</specimenCount>
|
||
. [
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFED8E4D6A7DCFD2F" box="[324,398,676,697]" pageId="5" pageNumber="633">CEMT</collectionCode>
|
||
].
|
||
<collectingCountry id="F359764BFFC6FFDEFE3FE4D6A7AEFD2F" box="[419,508,676,697]" name="Brazil" pageId="5" pageNumber="633">BRASIL</collectingCountry>
|
||
:
|
||
<collectingRegion id="498AF839FFC6FFDEFD95E4D6A468FD2F" box="[521,570,676,697]" country="Brazil" name="Acre" pageId="5" pageNumber="633">Acre</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B95ACA1FFC6FFDEFDD5E4D6A6B2FD43" pageId="5" pageNumber="633">Mâncio Lima</collectingMunicipality>
|
||
,
|
||
<location id="8E916000FFC6FFDEFF7BE4CDA75AFD42" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFF7BE4CDA75AFD42" box="[231,264,703,724]" country="Brazil" county="Same" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Par" pageId="5" pageNumber="633" stateProvince="Acre">Par</location>
|
||
[que]
|
||
<location id="8E916000FFC6FFDEFEA5E4CDA707FD43" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFEA5E4CDA707FD43" box="[313,341,703,725]" country="Brazil" county="Same" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Na" pageId="5" pageNumber="633" stateProvince="Acre">Na</location>
|
||
[cional]
|
||
<location id="8E916000FFC6FFDEFE02E4CDA463FD43" LSID="urn:lsid:plazi:treatment:03E787CDFFC1FFDEFD59E69CA5FAFC19:8E916000FFC6FFDEFE02E4CDA463FD43" box="[414,561,703,725]" country="Brazil" county="Same" latitude="-7.446111" longLatPrecision="21" longitude="-73.657776" municipality="Mancio Lima" name="Serra do Divisor" pageId="5" pageNumber="633" stateProvince="Acre">Serra do Divisor</location>
|
||
<geoCoordinate id="EE7A501CFFC6FFDEFDA9E4CCA4C6FD43" box="[565,660,702,725]" degrees="7" direction="south" minutes="26" orientation="latitude" pageId="5" pageNumber="633" precision="15" seconds="46" value="-7.446111">7°26′46″S</geoCoordinate>
|
||
<geoCoordinate id="EE7A501CFFC6FFDEFF31E4AAA770FD79" box="[173,290,728,751]" degrees="73" direction="west" minutes="39" orientation="longitude" pageId="5" pageNumber="633" precision="15" seconds="28" value="-73.657776">73°39′28″W</geoCoordinate>
|
||
,
|
||
<quantity id="4CB69B3EFFC6FFDEFEB7E4A8A738FD79" box="[299,362,730,752]" metricMagnitude="2" metricUnit="m" metricValue="2.36" pageId="5" pageNumber="633" unit="m" value="236.0">
|
||
<elevation id="0063D1E8FFC6FFDEFEB7E4A8A738FD79" box="[299,362,730,752]" metricMagnitude="2" metricUnit="m" metricValue="2.36" pageId="5" pageNumber="633" unit="m" value="236.0">236 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFF0101BFFC6FFDEFEE8E4A8A7ADFD66" box="[372,511,730,752]" pageId="5" pageNumber="633" value="2019-12-07" valueMax="2019-12-09" valueMin="2019-12-07">
|
||
<collectingDate id="EFB4E9F3FFC6FFDEFEE8E4A8A7ADFD66" box="[372,511,730,752]" pageId="5" pageNumber="633" value="2019-12-07" valueMax="2019-12-09" valueMin="2019-12-07">07-09.xii.2019</collectingDate>
|
||
</date>
|
||
. hum[an] dung.
|
||
<collectorName id="26BB530DFFC6FFDEFF31E486A763FC9C" box="[173,305,756,778]" pageId="5" pageNumber="633">RA Azevedo.</collectorName>
|
||
<specimenCount id="9D48FD52FFC6FFDEFEA5E486A70AFC9F" box="[313,344,756,777]" count="1" pageId="5" pageNumber="633" type="male">1♂</specimenCount>
|
||
<specimenCount id="9D48FD52FFC6FFDEFEFCE486A7C0FC9F" box="[352,402,756,777]" count="4" pageId="5" pageNumber="633" type="female">4♀♀</specimenCount>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B263C86FFC6FFDEFE03E486A7DAFCB3" collectionCode="CEMT, INPA, CERPE" country="Brazil" pageId="5" pageNumber="633" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Acre" typeStatus="paratype">
|
||
[
|
||
<specimenCount id="9D48FD52FFC6FFDEFE3BE486A785FC9F" box="[423,471,756,777]" count="2" pageId="5" pageNumber="633" type="female">2♀♀</specimenCount>
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFE7CE486A471FC9F" box="[480,547,756,777]" pageId="5" pageNumber="633">CEMT</collectionCode>
|
||
,
|
||
<specimenCount id="9D48FD52FFC6FFDEFDACE486A41CFC9F" box="[560,590,756,777]" count="1" pageId="5" pageNumber="633" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFDCBE486A4DDFC9F" box="[599,655,756,777]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="633">INPA</collectionCode>
|
||
,
|
||
<specimenCount id="9D48FD52FFC6FFDEFF31E57DA69EFCB2" box="[173,204,783,804]" count="1" pageId="5" pageNumber="633" type="male">1♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D48FD52FFC6FFDEFF40E57DA6A8FCB2" box="[220,250,783,804]" count="1" pageId="5" pageNumber="633" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED5FAE1EFFC6FFDEFE99E57DA705FCB2" box="[261,343,783,804]" pageId="5" pageNumber="633">CERPE</collectionCode>
|
||
]. All
|
||
</materialsCitation>
|
||
<typeStatus id="54F58879FFC6FFDEFE08E57DA7A6FCB2" box="[404,500,783,804]" pageId="5" pageNumber="633" type="paratype">Paratypes</typeStatus>
|
||
have a second label: 2: {printed and handwritten text on yellow label with black margins}
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFE22E537A43CFCCC" authorityName="Kim & Ahn" authorityYear="2023" box="[446,622,837,858]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii" status="sp. nov.">
|
||
<emphasis id="B93AEAC9FFC6FFDEFE22E537A43CFCCC" box="[446,622,837,858]" italics="true" pageId="5" pageNumber="633">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957B2FFC6FFDEFDEBE537A685FCE2" pageId="5" pageNumber="633" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Arias-BuriticÁ, Bach and Vaz-de-Mello, 2023
|
||
<typeStatus id="54F58879FFC6FFDEFF32E508A77BFC19" box="[174,297,890,911]" pageId="5" pageNumber="633" type="paratype">PARATYPE</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3546550FFC6FFDEFD47E6B3A5FAFC19" pageId="5" pageNumber="633" type="distribution">
|
||
<paragraph id="8BF136DBFFC6FFDEFD47E6B3A521FE3B" blockId="5.[705,1193,193,912]" pageId="5" pageNumber="633">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD47E6B3A589FF41" bold="true" box="[731,987,193,215]" pageId="5" pageNumber="633">Distribution and Ecology.</emphasis>
|
||
Known only from the southwestern Brazilian Amazon from Serra do Divisor National Park,
|
||
<collectingRegion id="498AF839FFC6FFDEFC37E685A234FE9B" box="[939,1126,247,269]" country="Brazil" name="Rondonia" pageId="5" pageNumber="633">Rondônia province</collectingRegion>
|
||
in the South Brazilian Dominion (
|
||
<bibRefCitation id="EFDF4B2AFFC6FFDEFC49E760A2F0FEB1" author="Morrone, J. J. & Escalante, G. & Rodriguez-Tapia, A. & Carmona, M." box="[981,1186,274,295]" pageId="5" pageNumber="633" refId="ref4088" refString="Morrone, J. J., T. Escalante, G. Rodriguez-Tapia, A. Carmona, M. Arana, and J. D. Mercado-Gomez. 2022. Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias 94 (1), e 20211167. doi. org / 10.1590 / 0001 - 3765202220211167." type="journal volume" year="2022">
|
||
Morrone
|
||
<emphasis id="B93AEAC9FFC6FFDEFBA8E760A233FEB1" box="[1076,1121,274,295]" italics="true" pageId="5" pageNumber="633">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13752A5EFFC6FFDEFD56E75EA550FED7" box="[714,770,300,322]" captionStart="Fig" captionStartId="5.[197,228,1687,1706]" captionTargetBox="[192,1173,969,1655]" captionTargetId="figure-443@5.[173,1193,954,1669]" captionTargetPageId="5" captionText="Fig. 3. Distribution map for Isocopris rossinii, new species. Plot A and Plot B represent the collection sites in Serra do Divisor National Park, Acre, Brazil." figureDoi="http://doi.org/10.5281/zenodo.13522819" httpUri="https://zenodo.org/record/13522819/files/figure.png" pageId="5" pageNumber="633">Fig. 3</figureCitation>
|
||
), a lowland
|
||
<emphasis id="B93AEAC9FFC6FFDEFCE8E75FA588FED7" box="[884,986,300,322]" italics="true" pageId="5" pageNumber="633">terra firme</emphasis>
|
||
site (
|
||
<emphasis id="B93AEAC9FFC6FFDEFB90E75FA27BFED4" box="[1036,1065,301,322]" italics="true" pageId="5" pageNumber="633">i.e.</emphasis>
|
||
, non-flooded area) with primary Amazonian forests up to
|
||
<quantity id="4CB69B3EFFC6FFDEFB18E735A4A1FEEE" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="633" unit="m" value="300.0">300 masl</quantity>
|
||
.
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFC9DE710A5FEFEE1" authorityName="Kim & Ahn" authorityYear="2023" box="[769,940,354,375]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFC9DE710A5FEFEE1" box="[769,940,354,375]" italics="true" pageId="5" pageNumber="633">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
has been sampled using flight interception traps (FIT) and pitfall traps baited with human feces.
|
||
</paragraph>
|
||
<paragraph id="8BF136DBFFC6FFDEFD47E7C6A5FAFC19" blockId="5.[705,1193,193,912]" pageId="5" pageNumber="633">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD47E7C6A5A2FE5C" bold="true" box="[731,1008,436,458]" pageId="5" pageNumber="633">Taxonomic Commentaries.</emphasis>
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFC62E7C7A2FAFE5C" authorityName="Kim & Ahn" authorityYear="2023" box="[1022,1192,437,458]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFC62E7C7A2FAFE5C" box="[1022,1192,437,458]" italics="true" pageId="5" pageNumber="633">Isocopris rossinii</emphasis>
|
||
</taxonomicName>
|
||
belongs to the small- to medium-sized
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFBBAE7A2A4B8FD96" pageId="5" pageNumber="633">
|
||
<emphasis id="B93AEAC9FFC6FFDEFBBAE7A2A22FFE73" box="[1062,1149,464,485]" italics="true" pageId="5" pageNumber="633">Isocopris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, encompassing
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFCE3E798A5A6FE69" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[895,1012,490,511]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFCE3E798A5D5FE69" box="[895,903,490,511]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFC08E798A5A6FE69" box="[916,1012,490,511]" italics="true" pageId="5" pageNumber="633">foveolatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFB9CE798A232FE69" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1024,1120,490,511]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B93AEAC9FFC6FFDEFB9CE798A25AFE69" box="[1024,1032,490,511]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFB89E798A232FE69" box="[1045,1120,490,511]" italics="true" pageId="5" pageNumber="633">imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFBF1E798A4B6FD8F" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFBF1E798A227FE69" box="[1133,1141,490,511]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFB1DE798A4B6FD8F" italics="true" pageId="5" pageNumber="633">nitidus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFD72E477A515FD8F" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[750,839,516,538]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD72E477A4A4FD8C" box="[750,758,517,538]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFC9DE476A515FD8F" box="[769,839,516,537]" italics="true" pageId="5" pageNumber="633">tarsalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFCE6E477A5B2FD8F" box="[890,992,516,538]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="xacriaba">
|
||
<emphasis id="B93AEAC9FFC6FFDEFCE6E477A5D0FD8C" box="[890,898,517,538]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFC11E476A5B2FD8F" box="[909,992,516,537]" italics="true" pageId="5" pageNumber="633">xacriaba</emphasis>
|
||
</taxonomicName>
|
||
. It is easily separated from the last four species by the cephalic process;
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFB06E452A55AFDD9" authorityName="Kim & Ahn" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFB06E452A55AFDD9" italics="true" pageId="5" pageNumber="633">I. rossinii</emphasis>
|
||
</taxonomicName>
|
||
has a transverse fronto-clypeal carina as does
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFD5EE427A560FDFF" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[706,818,596,618]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD5EE427A560FDFF" box="[706,818,596,618]" italics="true" pageId="5" pageNumber="633">I. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
, whereas the other species have a conical horn. By external morphology,
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFC7EE402A26BFD13" authorityName="Kim & Ahn" authorityYear="2023" box="[994,1081,623,645]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFC7EE402A5B8FD13" box="[994,1002,624,645]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFC68E402A26BFD13" box="[1012,1081,624,645]" italics="true" pageId="5" pageNumber="633">rossinii</emphasis>
|
||
</taxonomicName>
|
||
is related to
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFD5DE4F8A568FD09" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[705,826,650,671]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD5DE4F8A49BFD09" box="[705,713,650,671]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFD46E4F8A568FD09" box="[730,826,650,671]" italics="true" pageId="5" pageNumber="633">foveolatus</emphasis>
|
||
</taxonomicName>
|
||
, with which it shares the deeply depressed frons and the structure of the fronto-clypeal process. However, it is easily separated by the shape of the ventral clypeal process, the anterior lobe of the metaventrite, which is glabrous in
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFB9CE487A205FC9C" authorityName="Kim & Ahn" authorityYear="2023" box="[1024,1111,757,778]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFB9CE487A205FC9C" box="[1024,1111,757,778]" italics="true" pageId="5" pageNumber="633">I. rossinii</emphasis>
|
||
</taxonomicName>
|
||
but with long setae present in
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFC1DE562A5A6FCB2" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[897,1012,783,805]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFC1DE562A5DBFCB3" box="[897,905,784,805]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFC08E57DA5A6FCB2" box="[916,1012,783,804]" italics="true" pageId="5" pageNumber="633">foveolatus</emphasis>
|
||
</taxonomicName>
|
||
, as well as the morphology of the aedeagus and endophallites. The female secondary sexual dimorphism present in the elytral striae of
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFCC4E512A5E3FCE3" authorityName="Kim & Ahn" authorityYear="2023" box="[856,945,864,885]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="rossinii">
|
||
<emphasis id="B93AEAC9FFC6FFDEFCC4E512A5E3FCE3" box="[856,945,864,885]" italics="true" pageId="5" pageNumber="633">I. rossinii</emphasis>
|
||
</taxonomicName>
|
||
is shared with
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFBDEE512A2F0FCE3" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1090,1186,863,885]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="imitator">
|
||
<emphasis id="B93AEAC9FFC6FFDEFBDEE512A218FCE3" box="[1090,1098,864,885]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFBCBE512A2F0FCE3" box="[1111,1186,864,885]" italics="true" pageId="5" pageNumber="633">imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFD5FE508A544FC19" baseAuthorityName="Luederwaldt" baseAuthorityYear="1922" box="[707,790,890,911]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="B93AEAC9FFC6FFDEFD5FE508A499FC19" box="[707,715,890,911]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFD4AE508A544FC19" box="[726,790,890,911]" italics="true" pageId="5" pageNumber="633">nitidus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4E4D58FFC6FFDEFCD5E508A5F0FC19" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[841,930,890,911]" class="Insecta" family="Scarabaeidae" genus="Isocopris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="633" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="B93AEAC9FFC6FFDEFCD5E508A503FC19" box="[841,849,890,911]" italics="true" pageId="5" pageNumber="633">I</emphasis>
|
||
.
|
||
<emphasis id="B93AEAC9FFC6FFDEFCC0E508A5F0FC19" box="[860,930,890,911]" italics="true" pageId="5" pageNumber="633">tarsalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |