287 lines
36 KiB
XML
287 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="D398B4F836BF0DDC787B9DCC0328C1E7" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Lonchodraco microdon Rodrigues & Kellner 2013, comb. n." docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="21" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="20" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044963" LSID="urn:lsid:plazi:treatment:D398B4F836BF0DDC787B9DCC0328C1E7" httpUri="http://treatment.plazi.org/id/D398B4F836BF0DDC787B9DCC0328C1E7" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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<subSubSection pageId="19" pageNumber="20" type="nomenclature">
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName LSID="D398B4F8-36BF-0DDC-787B-9DCC0328C1E7" authority="(Seeley, 1870)" authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Lonchodraco microdon" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">
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<pageBreakToken pageId="19" pageNumber="20" start="start">Lonchodraco</pageBreakToken>
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(?) microdon (Seeley, 1870)
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</taxonomicName>
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<taxonomicNameLabel pageId="19" pageNumber="20">comb. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchodraco (?) microdon comb. n. A-F holotype CAMSM B 54486 (Albian, Cambridge Greensand), anterior fragment of the rostrum A ventral view B respective line drawing C anterior view D respective line drawing E posterior view F respective line drawing. In B dashed lines indicate the portion lost since the original description G-L referred specimen CAMSM B 54439 (Albian, Cambridge Greensand), anterior fragment of the rostrum G ventral view H respective line drawing I anterior view J respective line drawing K posterior view L respective line drawing. Abbreviation: prid - palatal ridge. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22074" pageId="19" pageNumber="20">Fig. 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="reference_group">
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ptenodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">
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<normalizedToken originalValue="“Ptenodactylus”">"Ptenodactylus"</normalizedToken>
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microdon
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</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
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: p. xvi [disclaimed]
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">Ornithocheirus microdon</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 116, pl. XII, fig. 6-7
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">Lonchodectes microdon</taxonomicName>
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(Seeley):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">Ornithocheirus microdon</taxonomicName>
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Seeley:
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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: p. 58
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">Lonchodectes microdon</taxonomicName>
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(Seeley):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: p. 211, table 1
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="oweni" status="comb. n.">Ornithocheirus oweni</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 115
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="oweni" status="comb. n.">Lonchodectes oweni</taxonomicName>
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(Seeley):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 535
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</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon" status="comb. n.">Lonchodectes microdon</taxonomicName>
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(Seeley):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: p. 195, fig. 11C-D, table 1 [synonymy]
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="holotype">
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<paragraph pageId="19" pageNumber="20">Holotype.</paragraph>
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<paragraph pageId="19" pageNumber="20">
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CAMSM B54486, anterior portion of the rostrum (
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchodraco (?) microdon comb. n. A-F holotype CAMSM B 54486 (Albian, Cambridge Greensand), anterior fragment of the rostrum A ventral view B respective line drawing C anterior view D respective line drawing E posterior view F respective line drawing. In B dashed lines indicate the portion lost since the original description G-L referred specimen CAMSM B 54439 (Albian, Cambridge Greensand), anterior fragment of the rostrum G ventral view H respective line drawing I anterior view J respective line drawing K posterior view L respective line drawing. Abbreviation: prid - palatal ridge. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22074" pageId="19" pageNumber="20">Fig. 6A-F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="type locality">
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<paragraph pageId="19" pageNumber="20">Type locality.</paragraph>
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<paragraph pageId="19" pageNumber="20">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="type horizon">
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<paragraph pageId="19" pageNumber="20">Type horizon.</paragraph>
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<paragraph pageId="19" pageNumber="20">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="referred specimen">
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<paragraph pageId="19" pageNumber="20">Referred specimen.</paragraph>
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<paragraph pageId="19" pageNumber="20">
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CAMSM B 54439 (
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchodraco (?) microdon comb. n. A-F holotype CAMSM B 54486 (Albian, Cambridge Greensand), anterior fragment of the rostrum A ventral view B respective line drawing C anterior view D respective line drawing E posterior view F respective line drawing. In B dashed lines indicate the portion lost since the original description G-L referred specimen CAMSM B 54439 (Albian, Cambridge Greensand), anterior fragment of the rostrum G ventral view H respective line drawing I anterior view J respective line drawing K posterior view L respective line drawing. Abbreviation: prid - palatal ridge. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22074" pageId="19" pageNumber="20">Fig. 6G-L</figureCitation>
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) (also from the Cambridge Greensand).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="20" type="diagnosis">
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<paragraph pageId="19" pageNumber="20">Diagnosis.</paragraph>
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<paragraph pageId="19" pageNumber="20">Lonchodraconid pterosaur with the following combination of characters that distinguishes it from other members of the clade (autapomorphies are marked with an asterisk): premaxillary crest absent; dorsal margin of the premaxillae rounded; deep palatal ridge*; palate between the elevation of the alveolar margins and the palatal ridge concave; spacing between alveoli larger than their diameters*; approximately 4.5 alveoli per 3 cm of jaw margin.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
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<paragraph pageId="19" pageNumber="20">Description.</paragraph>
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="microdon">
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<emphasis italics="true" pageId="19" pageNumber="20">Lonchodraco(?) microdon</emphasis>
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</taxonomicName>
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was described by
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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on the basis of CAMSM B54486, a fragmentary upper jaw with a prominent palatal ridge.
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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noted the presence of a concavity on the palate between the elevation of the alveolar margins and the palatal ridge, that the palatal ridge becomes more prominent than the alveolar borders posteriorly, and that the alveolar margins are compressed and rounded.
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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also noted a small tip of jaw
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<normalizedToken originalValue="“associated”">"associated"</normalizedToken>
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with the specimen, but this possibly does not represent the same individual. CAMSM B54486 was recently sampled for histological analysis (M. Riley, pers. comm.). The transverse section of the area that was cut is wider than high, whereas the opposite end is higher than wide, suggesting that the sampled area was located near the anterior end of the snout.
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</paragraph>
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<paragraph pageId="20" pageNumber="21">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oweni">
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<emphasis italics="true" pageId="20" pageNumber="21">
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<pageBreakToken pageId="20" pageNumber="21" start="start">Ornithocheirus</pageBreakToken>
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oweni
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</emphasis>
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</taxonomicName>
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was described in the same work (
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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) on the basis of CAMSM B 54439 (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchodraco (?) microdon comb. n. A-F holotype CAMSM B 54486 (Albian, Cambridge Greensand), anterior fragment of the rostrum A ventral view B respective line drawing C anterior view D respective line drawing E posterior view F respective line drawing. In B dashed lines indicate the portion lost since the original description G-L referred specimen CAMSM B 54439 (Albian, Cambridge Greensand), anterior fragment of the rostrum G ventral view H respective line drawing I anterior view J respective line drawing K posterior view L respective line drawing. Abbreviation: prid - palatal ridge. Arrows indicate alveoli or teeth. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22074" pageId="20" pageNumber="21">Fig. 6G-L</figureCitation>
|
||
), also an upper jaw. This species is quite similar to
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
. Both holotypes share a rostrum with parallel alveolar margins, absence of a premaxillary crest, rounded dorsal margin of the premaxillae and maxillae, deep palatal ridge and the concave palate between the alveolar margins and the palatal ridge.
|
||
<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
|
||
pointed out that
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oweni">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Ornithocheirus oweni</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
in that its jaw margins are not round but flattened, by the presence of rough interspaces between the alveoli, teeth (alveoli?) circular instead of oval, and the dorsal margin of the premaxillae and maxillae rounded instead of having a sharp keel. However, examination of CAMSM B 54439 established that the alveolar margins are poorly preserved so that the number, shape, and spacing of the alveoli cannot be seen. Furthermore,
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
has a rounded dorsal margin of the rostrum, as can be observed in anterior and posterior transverse sections of the holotype. As both species share the same morphological features and come from the same deposit they are here considered conspecific.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="21" type="remarks">
|
||
<paragraph pageId="20" pageNumber="21">Remarks.</paragraph>
|
||
<paragraph pageId="20" pageNumber="21">
|
||
<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="oweni">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Ornithocheirus oweni</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
and, acting as the First Reviser (
|
||
<normalizedToken originalValue="ICZN’s">ICZN's</normalizedToken>
|
||
article 24.2.2), gave
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
priority.
|
||
</paragraph>
|
||
<paragraph pageId="20" pageNumber="21">
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
has small (approximately 3 mm diameter) and evenly spaced alveoli as in
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="machaerorhynchus">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco machaerorhynchus</emphasis>
|
||
</taxonomicName>
|
||
. However, it is distinct from
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="giganteus">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco giganteus</emphasis>
|
||
</taxonomicName>
|
||
, lacking a premaxillary crest, having spacing between the alveoli larger than their diameters, and with parallel alveolar margins. These features constrast with those of the type species of the genus.
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="machaerorhynchus">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco machaerorhynchus</emphasis>
|
||
</taxonomicName>
|
||
the parallel alveolar margins and the same alveolar density (4.5 alveoli per 3 cm of jaw margin), but they differ in the spacing between the alveoli, larger in the first. Due to these differences, it is possible that
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
actually represents a distinct genus. However, in the absence of further evidence, we refrain from naming a new genus and assign the species tentatively to
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/22074" pageId="20" pageNumber="21" start="Figure 6" startId="F6">
|
||
<paragraph pageId="20" pageNumber="21">
|
||
<emphasis bold="true" pageId="20" pageNumber="21">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Seeley" baseAuthorityYear="1870" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="microdon">
|
||
<emphasis italics="true" pageId="20" pageNumber="21">Lonchodraco(?) microdon</emphasis>
|
||
</taxonomicName>
|
||
comb. n.
|
||
<emphasis bold="true" pageId="20" pageNumber="21">A-F</emphasis>
|
||
holotype CAMSM B54486 (Albian, Cambridge Greensand), anterior fragment of the rostrum
|
||
<emphasis bold="true" pageId="20" pageNumber="21">A</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">B</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="20" pageNumber="21">C</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">D</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="20" pageNumber="21">E</emphasis>
|
||
posterior view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">F</emphasis>
|
||
respective line drawing. In
|
||
<emphasis bold="true" pageId="20" pageNumber="21">B</emphasis>
|
||
dashed lines indicate the portion lost since the original description
|
||
<emphasis bold="true" pageId="20" pageNumber="21">G-L</emphasis>
|
||
referred specimen CAMSM B 54439 (Albian, Cambridge Greensand), anterior fragment of the rostrum
|
||
<emphasis bold="true" pageId="20" pageNumber="21">G</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">H</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="20" pageNumber="21">I</emphasis>
|
||
anterior view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">J</emphasis>
|
||
respective line drawing
|
||
<emphasis bold="true" pageId="20" pageNumber="21">K</emphasis>
|
||
posterior view
|
||
<emphasis bold="true" pageId="20" pageNumber="21">L</emphasis>
|
||
respective line drawing. Abbreviation:
|
||
<emphasis bold="true" pageId="20" pageNumber="21">prid</emphasis>
|
||
- palatal ridge. Arrows indicate alveoli or teeth. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |