216 lines
21 KiB
XML
216 lines
21 KiB
XML
<document id="C8EFD29A346B15FF2BF18489A3F7E756" ID-DOI="10.1186/s13358-022-00259-x" ID-ISSN="1664-2384" ID-PMC="PMC9681704" ID-PubMed="36439694" ID-Zenodo-Dep="12000082" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718653166208" checkinUser="felipe" docAuthor="Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas" docDate="2022" docId="038B1D1D647FFFA7FF1BE91BD543FDA7" docLanguage="en" docName="SwissJPalaeontol.141.19.pdf" docOrigin="Swiss Journal of Palaeontology (19) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00259-x" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Neospathodus cristagalli Mosher 1968" docType="treatment" docVersion="7" lastPageNumber="23" masterDocId="FFB265656469FFB1FF8CED10D031FFE4" masterDocTitle="A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China" masterLastPageNumber="61" masterPageNumber="1" pageNumber="23" updateTime="1722282637272" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="853DF4EF47174D449C638D7202C72687">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="5EC09536C890D7F4BEA73543E9AFEFC8">Leu, Marc</mods:namePart>
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<mods:affiliation id="F8E8F554FC8C509A1DAED8D5E9D2D7AE">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="69CA010D9C9AC03DB53C8787E182AFC8">Bucher, Hugo</mods:namePart>
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<mods:affiliation id="9A81AA6EBF9872FC79D68463F5CB2063">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="26F2A3776E51E2EB42ECCDB94A2F42FF">Ji, Cheng</mods:namePart>
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<mods:namePart id="FE537F7820D2B167889F18CCA9EF08D3">Brosse, Morgane</mods:namePart>
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<mods:namePart id="0BDD54081DB24759DF1F14B4E0A87DD5">Goudemand, Nicolas</mods:namePart>
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<mods:title id="B62E2B94E962235FC9CCFBBD8B036E1A">Swiss Journal of Palaeontology</mods:title>
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<treatment id="038B1D1D647FFFA7FF1BE91BD543FDA7" LSID="urn:lsid:plazi:treatment:038B1D1D647FFFA7FF1BE91BD543FDA7" httpUri="http://treatment.plazi.org/id/038B1D1D647FFFA7FF1BE91BD543FDA7" lastPageNumber="23" pageId="22" pageNumber="23">
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<subSubSection id="C338FF80647FFFA7FF1BE91BD2ABFBC7" box="[151,666,1035,1060]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8B9DAC0B647FFFA7FF1BE91BD2ABFBC7" blockId="22.[151,666,1035,1091]" box="[151,666,1035,1060]" pageId="22" pageNumber="23">
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<heading id="D0D51B67647FFFA7FF1BE91BD2ABFBC7" box="[151,666,1035,1060]" fontSize="9" level="2" pageId="22" pageNumber="23" reason="8">
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<taxonomicName id="4C22D788647FFFA7FF1BE91BD2ABFBC7" authority="Huckriede, 1958" authorityName="Mosher" authorityYear="1968" box="[151,666,1035,1060]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
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<emphasis id="B9567019647FFFA7FF1BE91BD101FBC7" box="[151,304,1035,1059]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
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ex gr.
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<emphasis id="B9567019647FFFA7FEFAE91BD1D3FBC7" box="[374,482,1035,1059]" italics="true" pageId="22" pageNumber="23">cristagalli</emphasis>
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<bibRefCitation id="EFB3D1FA647FFFA7FE64E91BD2ABFBC7" author="Huckriede, R." box="[488,666,1035,1059]" pageId="22" pageNumber="23" pagination="141 - 175" refId="ref42031" refString="Huckriede, R. (1958). Die Conodonten der Mediterranen Trias und ihr stratigraphischer Wert. Palaontologische Zeitschrift, 32, 141 - 175." type="journal article" year="1958">Huckriede, 1958</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C338FF80647FFFA7FF1BE93BD543FDA7" pageId="22" pageNumber="23" type="description">
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<paragraph id="8B9DAC0B647FFFA7FF1BE93BD0C0FBA7" blockId="22.[151,666,1035,1091]" box="[151,241,1067,1091]" pageId="22" pageNumber="23">
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<figureCitation id="1319B08E647FFFA7FF1BE93BD0C0FBA7" box="[151,241,1067,1091]" captionStart="Fig" captionStartId="21.[169,200,1283,1303]" captionTargetBox="[153,1438,238,1422]" captionTargetId="figure-168@21.[229,1363,254,1271]" captionTargetPageId="21" captionText="Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181" figureDoi="http://doi.org/10.5281/zenodo.12000270" httpUri="https://zenodo.org/record/12000270/files/figure.png" pageId="22" pageNumber="23">Fig. 14E</figureCitation>
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE97BD17BFB47" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
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1970
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<taxonomicName id="4C22D788647FFFA7FF59E97BD26EFB67" authority="Huckriede" authorityName="Huckriede" box="[213,607,1131,1155]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF59E97BD1D2FB67" box="[213,483,1131,1155]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
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Huckriede
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</taxonomicName>
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; Sweet, p. 346, pl. 1, figs. 14, 15.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE9BBD170FB07" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
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1982
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<taxonomicName id="4C22D788647FFFA7FF5FE9BBD26BFB27" authority="Huckriede" authorityName="Huckriede" box="[211,602,1195,1219]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF5FE9BBD1EEFB27" box="[211,479,1195,1219]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
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Huckriede
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</taxonomicName>
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; Matsuda p. 92, pl. 3, figs. 1–12.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE9FBD123FAC7" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
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2005
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<taxonomicName id="4C22D788647FFFA7FF59E9FBD260FAE7" authority="Sweet" authorityName="Sweet" box="[213,593,1259,1283]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF59E9FBD15FFAE7" box="[213,366,1259,1283]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
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cf.
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<emphasis id="B9567019647FFFA7FE15E9FBD234FAE7" box="[409,517,1259,1283]" italics="true" pageId="22" pageNumber="23">cristagalli</emphasis>
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Sweet
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</taxonomicName>
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; Orchard, p. 89, text-fig. 14.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE83BD2A6FA87" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
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2014
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<taxonomicName id="4C22D788647FFFA7FF55E83BD25CFAA7" authority="Huckriede" authorityName="Huckriede" box="[217,621,1323,1347]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF55E83BD1DDFAA7" box="[217,492,1323,1347]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
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Huckriede
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</taxonomicName>
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; Maekawa & Igo in Shigeta et al., p. 223, figs. 161.10–161.12.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE87BD22AFA47" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
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2015
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<taxonomicName id="4C22D788647FFFA7FF55E87BD25DFA67" authority="Huckriede" authorityName="Huckriede" box="[217,620,1387,1411]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF55E87BD1DAFA67" box="[217,491,1387,1411]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
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Huckriede
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</taxonomicName>
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; Maekawa in Maekawa et al., p. 315, figs. 5.4–5.6.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE8DBD1F2FA07" blockId="22.[151,775,1483,1891]" box="[151,451,1483,1507]" pageId="22" pageNumber="23">
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<emphasis id="B9567019647FFFA7FF1BE8DBD0CEFA07" box="[151,255,1483,1507]" italics="true" pageId="22" pageNumber="23">Material.</emphasis>
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ca.
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<specimenCount id="9D246782647FFFA7FEA5E8DBD18EFA07" box="[297,447,1483,1507]" count="30" pageId="22" pageNumber="23" type="generic">30 specimens</specimenCount>
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.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FF1BE8FBD111F887" blockId="22.[151,775,1483,1891]" pageId="22" pageNumber="23">
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<emphasis id="B9567019647FFFA7FF1BE8FBD0CDF9E7" box="[151,252,1515,1539]" italics="true" pageId="22" pageNumber="23">Remarks.</emphasis>
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In comparison with the
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<typeStatus id="549912A9647FFFA7FDA8E8FBD2B5F9E7" box="[548,644,1515,1539]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C22D788647FFFA7FD3EE8FCD0EDF9C7" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FD3EE8FCD0EDF9C7" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFB3D1FA647FFFA7FF7DEB1BD19DF9C7" author="Huckriede, R." box="[241,428,1547,1571]" pageId="22" pageNumber="23" pagination="141 - 175" refId="ref42031" refString="Huckriede, R. (1958). Die Conodonten der Mediterranen Trias und ihr stratigraphischer Wert. Palaontologische Zeitschrift, 32, 141 - 175." type="journal article" year="1958">Huckriede, 1958</bibRefCitation>
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, Pl. 10, fig. 15), this element is much shorter, the basal cavity is oval, posteriorly rounded and only partly inverted, and the posterior triangular cusp is not conspicuously separated from the other denticles. Sweet still included such forms within
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<taxonomicName id="4C22D788647FFFA7FF1BEBBCD112F927" box="[151,291,1707,1731]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FF1BEBBCD112F927" box="[151,291,1707,1731]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
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</taxonomicName>
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, but excluded similar forms with a rounded basal cavity and rounded, pointy denticles and assigned them to the then new species
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<taxonomicName id="4C22D788647FFFA7FE69EBFCD263F8E7" authorityName="Sweet" authorityYear="1970" box="[485,594,1771,1795]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="dieneri">
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<emphasis id="B9567019647FFFA7FE69EBFCD263F8E7" box="[485,594,1771,1795]" italics="true" pageId="22" pageNumber="23">N. dieneri</emphasis>
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</taxonomicName>
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. In our opinion, such short elements where the cusp is not separated from the other denticles would deserve to be differentiated as a new species.
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</paragraph>
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<paragraph id="8B9DAC0B647FFFA7FCA1EDFCD543FDA7" blockId="22.[813,1437,235,579]" pageId="22" pageNumber="23">
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||
<emphasis id="B9567019647FFFA7FCA1EDFCD399FEE7" box="[813,936,236,259]" italics="true" pageId="22" pageNumber="23">Occurrence</emphasis>
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.
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<taxonomicName id="4C22D788647FFFA7FC37EDFCD47FFEE7" box="[955,1102,235,259]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
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<emphasis id="B9567019647FFFA7FC37EDFCD47FFEE7" box="[955,1102,235,259]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
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</taxonomicName>
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is known worldwide in the Dienerian and earliest Smithian. It is not clear yet whether the present form is younger than
|
||
<taxonomicName id="4C22D788647FFFA7FA82EC3CD5ACFEA7" box="[1294,1437,299,323]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
|
||
<emphasis id="B9567019647FFFA7FA82EC3CD5ACFEA7" box="[1294,1437,299,323]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A265CD62647FFFA7FCA1EC5CD38DFE87" box="[813,956,331,355]" pageId="22" pageNumber="23" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
and occurs only close to the Dienerian– Smithian (Induan–Olenekian) boundary: Toad Formation, British Columbia (
|
||
<bibRefCitation id="EFB3D1FA647FFFA7FBC3EC9BD4C9FE47" author="Orchard, M. J." box="[1103,1272,395,419]" pageId="22" pageNumber="23" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
|
||
). Salt Range,
|
||
<collectingCountry id="F335EC9B647FFFA7FCA1ECBBD3B8FE27" box="[813,905,427,451]" name="Pakistan" pageId="22" pageNumber="23">Pakistan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB3D1FA647FFFA7FC10ECBBD415FE27" author="Sweet, W. C." box="[924,1060,427,451]" pageId="22" pageNumber="23" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
|
||
). Guryul Ravine, Kashmir (
|
||
<bibRefCitation id="EFB3D1FA647FFFA7FAEAECBBD395FE07" author="Matsuda, T." pageId="22" pageNumber="23" pagination="87 - 102" refId="ref44946" refString="Matsuda, T. (1982). Early triassic conodonts from Kashmir, India part 2: Neospathodus 1. Journal of Geosciences Osaka City University, 25, 87 - 102." type="journal article" year="1982">Matsuda, 1982</bibRefCitation>
|
||
). Nanpanjiang basin; north-eastern
|
||
<collectingCountry id="F335EC9B647FFFA7FAB3ECDBD5ADFE07" box="[1343,1436,459,483]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
|
||
<taxonomicName id="4C22D788647FFFA7FCA1ECFBD3D4FDE7" authority="beds" authorityName="Beds" box="[813,997,491,515]" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="22" pageNumber="12" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019647FFFA7FCA1ECFBD398FDE7" box="[813,937,491,515]" italics="true" pageId="22" pageNumber="23">Flemingites</emphasis>
|
||
beds
|
||
</taxonomicName>
|
||
within the
|
||
<taxonomicName id="4C22D788647FFFA7FBE0ECFBD53FFDE7" authorityName="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand" authorityYear="2022" box="[1132,1294,491,515]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9567019647FFFA7FBE0ECFBD53FFDE7" box="[1132,1294,491,515]" italics="true" pageId="22" pageNumber="23">Novispathodus</emphasis>
|
||
</taxonomicName>
|
||
ex. gr.
|
||
<taxonomicName id="4C22D788647FFFA7FAE9ECFCD3ACFDC7" authority="Zone" authorityName="Zone" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="waageni">
|
||
<emphasis id="B9567019647FFFA7FAE9ECFCD368FDC7" italics="true" pageId="22" pageNumber="23">waageni</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB3D1FA647FFFA7FC23EF1BD493FDC7" author="Maekawa, T. & Komatsu, T. & Shigeta, Y. & Huyen, D. T. & Tien, D. C." box="[943,1186,523,547]" pageId="22" pageNumber="23" pagination="312 - 320" refId="ref44838" refString="Maekawa, T., Komatsu, T., Shigeta, Y., Huyen, D. T., & Tien, D. C. (2015). First occurrence of Early Triassic conodonts from the Lang Son Formation, northeastern Vietnam. Paleontological Research, 19 (4), 312 - 320. https: // doi. org / 10.2517 / 2015 PR 014" type="journal article" year="2015">Maekawa et al., 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB3D1FA647FFFA7FB38EF1BD5BCFDC7" author="Shigeta, Y. & Komatsu, T. & Maekawa, T. & Huyen, D. T." box="[1204,1421,523,547]" pageId="22" pageNumber="23" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., & Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
|
||
). Luolou Formation,
|
||
<taxonomicName id="4C22D788647FFFA7FC72EF3BD44BFDA7" box="[1022,1146,555,579]" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019647FFFA7FC72EF3BD44BFDA7" box="[1022,1146,555,579]" italics="true" pageId="22" pageNumber="23">Flemingites</emphasis>
|
||
</taxonomicName>
|
||
limestone (this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |