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<document id="D16FB1C2D4EC3CB0FD5A698BEB2AEBCD" ID-DOI="10.1186/s13358-022-00259-x" ID-ISSN="1664-2384" ID-PMC="PMC9681704" ID-PubMed="36439694" ID-Zenodo-Dep="12000082" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718653166208" checkinUser="felipe" docAuthor="Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane &amp; Goudemand, Nicolas" docDate="2022" docId="038B1D1D6471FFABFCA1EB1BD18EFD87" docLanguage="en" docName="SwissJPalaeontol.141.19.pdf" docOrigin="Swiss Journal of Palaeontology (19) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00259-x" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Novispathodus Orchard 2005" docType="treatment" docVersion="4" lastPageNumber="27" masterDocId="FFB265656469FFB1FF8CED10D031FFE4" masterDocTitle="A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China" masterLastPageNumber="61" masterPageNumber="1" pageNumber="25" updateTime="1722282637272" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9E13B6E4D50E6B5B3D78EFEF2F7B1A33">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="AF03EF8746DC343C34D3602218CFBEFA">Leu, Marc</mods:namePart>
<mods:affiliation id="D629EBD6531376564D01D4F6DC3044F1">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="D9927D5D9D722C70736CF3E881FEFC2F">Bucher, Hugo</mods:namePart>
<mods:affiliation id="1C8988785A7870B537AB7B5922D94585">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="226889E2AE0D237B656DA22C7076175C">Vennemann, Torsten</mods:namePart>
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<mods:namePart id="106935A2D78F8A83010770C6C8AF89D0">Bagherpour, Borhan</mods:namePart>
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<mods:namePart id="4C0CB1C3BA8AE89E7CD1E8CA6FEEFB1E">Ji, Cheng</mods:namePart>
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<mods:namePart id="AF605916F5B87DF74EDDF7FED9726223">Brosse, Morgane</mods:namePart>
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<mods:namePart id="5ECF719FCA1F7D8E11E6EB519E90E17A">Goudemand, Nicolas</mods:namePart>
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<mods:title id="EFF160003B3947B107402EE53EF1EF96">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="D33C6E5DB9DDE9B7D3323F4D15B30E6A">2022</mods:date>
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<treatment id="038B1D1D6471FFABFCA1EB1BD18EFD87" LSID="urn:lsid:plazi:treatment:038B1D1D6471FFABFCA1EB1BD18EFD87" httpUri="http://treatment.plazi.org/id/038B1D1D6471FFABFCA1EB1BD18EFD87" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="C338FF806471FFA9FCA1EB1BD4C8F9C7" box="[813,1273,1547,1571]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8B9DAC0B6471FFA9FCA1EB1BD4C8F9C7" blockId="24.[813,1342,1515,1571]" box="[813,1273,1547,1571]" pageId="24" pageNumber="25">
<heading id="D0D51B676471FFA9FCA1EB1BD4C8F9C7" box="[813,1273,1547,1571]" fontSize="36" level="2" pageId="24" pageNumber="25" reason="3">
Genus
<taxonomicName id="4C22D7886471FFA9FCF5EB1BD4C8F9C7" authority="Orchard, 2005" authorityName="Orchard" authorityYear="2005" box="[889,1273,1547,1571]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
NOVISPATHODUS
<bibRefCitation id="EFB3D1FA6471FFA9FBD1EB1BD4C8F9C7" author="Orchard, M. J." box="[1117,1273,1547,1571]" pageId="24" pageNumber="25" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806471FFABFCA1EB5CD18EFD87" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8B9DAC0B6471FFA9FCA1EB5CD4C8F967" blockId="24.[813,1436,1611,1667]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FCA1EB5CD441F987" box="[813,1136,1611,1635]" italics="true" pageId="24" pageNumber="25">
Type species and
<typeStatus id="549912A96471FFA9FB98EB5BD441F987" box="[1044,1136,1611,1635]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
</emphasis>
.
<taxonomicName id="4C22D7886471FFA9FB07EB5BD46FF967" authority="Orchard, 1995, pp. 118 - 119" authorityName="Orchard" authorityPageNumber="118 - 119" authorityYear="1995" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196471FFA9FB07EB5BD5ADF987" box="[1163,1436,1611,1635]" italics="true" pageId="24" pageNumber="25">Neospathodus abruptus</emphasis>
<bibRefCitation id="EFB3D1FA6471FFA9FCA1EB7BD3FAF967" author="Orchard, M. J." box="[813,971,1643,1667]" pageId="24" pageNumber="25" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
, pp. 118119
</taxonomicName>
, figs. 3.2324.
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF25EBA2D1B2F922" blockId="24.[169,387,1714,1734]" box="[169,387,1714,1734]" pageId="24" pageNumber="25">(See figure on next page.)</paragraph>
<caption id="DF5DFC836471FFA9FF25EBDCD378F864" pageId="24" pageNumber="25" startId="24.[169,200,1740,1760]" targetBox="[153,1439,238,1800]" targetPageId="25">
<paragraph id="8B9DAC0B6471FFA9FF25EBDCD56BF91F" blockId="24.[169,1421,1739,1920]" pageId="24" pageNumber="25">
<figureCitation id="1319B08E6471FFA9FF25EBDCD0D9F904" box="[169,232,1740,1760]" captionStart="Fig" captionStartId="25.[169,200,1776,1796]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000298" httpUri="https://zenodo.org/record/12000298/files/figure.png" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF25EBDCD0D9F904" bold="true" box="[169,232,1740,1760]" pageId="24" pageNumber="25">Fig. 16</emphasis>
</figureCitation>
<taxonomicName id="4C22D7886471FFA9FF7EEBDCD1BDF904" authorityName="Hirsch" authorityYear="1994" box="[242,396,1740,1760]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="21" phylum="Chordata" rank="subFamily" subFamily="Neogondolellinae">Neogondolellinae</taxonomicName>
,
<taxonomicName id="4C22D7886471FFA9FE1FEBDCD215F904" authorityName="Orchard" authorityYear="2005" box="[403,548,1740,1760]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
and
<taxonomicName id="4C22D7886471FFA9FDC2EBDCD299F904" authorityName="Orchard" authorityYear="2005" box="[590,680,1740,1760]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="45" phylum="Chordata" rank="subFamily" subFamily="Mullerinae">Mullerinae</taxonomicName>
from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E6471FFA9FE59EBE1D1EDF91B" attach="left" box="[469,476,1777,1791]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B95670196471FFA9FCF5EBF7D3B6F91F" bold="true" box="[889,903,1767,1787]" pageId="24" pageNumber="25">A</emphasis>
,
<emphasis id="B95670196471FFA9FC03EBF7D3ABF91F" bold="true" box="[911,922,1767,1787]" pageId="24" pageNumber="25">F</emphasis>
,
<emphasis id="B95670196471FFA9FC2EEBF7D385F91F" bold="true" box="[930,948,1767,1787]" pageId="24" pageNumber="25">M</emphasis>
<emphasis id="B95670196471FFA9FC35EBF7D425F91F" box="[953,1044,1767,1787]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C22D7886471FFA9FC35EBF7D43DF91F" authorityName="Orchard" authorityYear="2005" box="[953,1036,1767,1787]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">Discretella</taxonomicName>
?
</emphasis>
n. sp. C;
<emphasis id="B95670196471FFA9FBD1EBF7D45AF91F" bold="true" box="[1117,1131,1767,1787]" pageId="24" pageNumber="25">A</emphasis>
YC16,PIMUZ 39115;
<emphasis id="B95670196471FFA9FA97EBF7D517F91F" bold="true" box="[1307,1318,1767,1787]" pageId="24" pageNumber="25">F</emphasis>
YC16,
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF25EA12D548F8D4" blockId="24.[169,1421,1739,1920]" pageId="24" pageNumber="25">
PIMUZ 39116;
<emphasis id="B95670196471FFA9FEAFEA12D104F8F2" bold="true" box="[291,309,1794,1814]" pageId="24" pageNumber="25">M</emphasis>
SHA342C, PIMUZ 39117.
<emphasis id="B95670196471FFA9FD87EA12D226F8F2" bold="true" box="[523,535,1794,1814]" pageId="24" pageNumber="25">B</emphasis>
<emphasis id="B95670196471FFA9FD90EA12D246F8F2" box="[540,631,1794,1814]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C22D7886471FFA9FD90EA12D25EF8F2" authorityName="Orchard" authorityYear="2005" box="[540,623,1794,1814]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">Discretella</taxonomicName>
?
</emphasis>
n. sp. B;YC16, PIMUZ 39113.
<emphasis id="B95670196471FFA9FCE7EA12D349F8F2" bold="true" box="[875,888,1794,1814]" pageId="24" pageNumber="25">C</emphasis>
<taxonomicName id="4C22D7886471FFA9FCF0EA12D401F8F2" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[892,1072,1794,1814]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196471FFA9FCF0EA12D3E1F8F2" box="[892,976,1794,1814]" italics="true" pageId="24" pageNumber="25">Discretella</emphasis>
aff.
<emphasis id="B95670196471FFA9FC7DEA12D401F8F2" box="[1009,1072,1794,1814]" italics="true" pageId="24" pageNumber="25">discreta</emphasis>
</taxonomicName>
(MÜller);YC22,PIMUZ 39106.
<emphasis id="B95670196471FFA9FAA7EA12D50BF8F2" bold="true" box="[1323,1338,1794,1814]" pageId="24" pageNumber="25">D</emphasis>
,
<emphasis id="B95670196471FFA9FACEEA12D57CF8F2" bold="true" box="[1346,1357,1794,1814]" pageId="24" pageNumber="25">S</emphasis>
<taxonomicName id="4C22D7886471FFA9FF25EA0CD1B9F8D4" authority="(MUller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[169,392,1820,1840]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196471FFA9FF25EA0CD171F8D4" box="[169,320,1820,1840]" italics="true" pageId="24" pageNumber="25">Discretella discreta</emphasis>
(MÜller)
</taxonomicName>
;
<emphasis id="B95670196471FFA9FE02EA0CD1ACF8D4" bold="true" box="[398,413,1820,1840]" pageId="24" pageNumber="25">D</emphasis>
YC16, PIMUZ 39110;
<emphasis id="B95670196471FFA9FDC2EA0CD268F8D4" bold="true" box="[590,601,1820,1840]" pageId="24" pageNumber="25">S</emphasis>
SHA304C, PIMUZ 39111.
<emphasis id="B95670196471FFA9FCA3EA0CD30BF8D4" bold="true" box="[815,826,1820,1840]" pageId="24" pageNumber="25">E</emphasis>
<taxonomicName id="4C22D7886471FFA9FCB3EA0CD38BF8D4" authorityName="Orchard" authorityYear="2005" box="[831,954,1820,1840]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B95670196471FFA9FCB3EA0CD38BF8D4" box="[831,954,1820,1840]" italics="true" pageId="24" pageNumber="25">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886471FFA9FC63EA0CD4BEF8D4" authority="(Orchard)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[1007,1167,1820,1840]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196471FFA9FC63EA0CD408F8D4" box="[1007,1081,1820,1840]" italics="true" pageId="24" pageNumber="25">abruptus</emphasis>
(Orchard)
</taxonomicName>
;YC41, PIMUZ 39225.
<emphasis id="B95670196471FFA9FACEEA0CD550F8D4" bold="true" box="[1346,1377,1820,1840]" pageId="24" pageNumber="25">GI</emphasis>
,
<emphasis id="B95670196471FFA9FAE5EA0CD544F8D4" bold="true" box="[1385,1397,1820,1840]" pageId="24" pageNumber="25">K</emphasis>
,
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF25EA27D378F864" blockId="24.[169,1421,1739,1920]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF25EA27D085F8AF" bold="true" box="[169,180,1847,1867]" pageId="24" pageNumber="25">L</emphasis>
,
<emphasis id="B95670196471FFA9FF30EA27D0FBF8AF" bold="true" box="[188,202,1847,1867]" pageId="24" pageNumber="25">N</emphasis>
,
<emphasis id="B95670196471FFA9FF5FEA27D0D3F8AF" bold="true" box="[211,226,1847,1867]" pageId="24" pageNumber="25">O</emphasis>
<taxonomicName id="4C22D7886471FFA9FF6BEA27D181F8AF" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[231,432,1847,1867]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pseudodieneri" status="sp. nov.">
<emphasis id="B95670196471FFA9FF6BEA27D181F8AF" box="[231,432,1847,1867]" italics="true" pageId="24" pageNumber="25">Discretella pseudodieneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FE39EA27D1D2F8AF" box="[437,483,1847,1867]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196471FFA9FE64EA27D1C7F8AF" bold="true" box="[488,502,1847,1867]" pageId="24" pageNumber="25">G</emphasis>
YC16, PIMUZ 39124;
<emphasis id="B95670196471FFA9FD2BEA27D284F8AF" bold="true" box="[679,693,1847,1867]" pageId="24" pageNumber="25">H</emphasis>
YC25, PIMUZ 39125;
<emphasis id="B95670196471FFA9FCEBEA27D35CF8AF" bold="true" box="[871,877,1847,1867]" pageId="24" pageNumber="25">I</emphasis>
SHA304C,PIMUZ 39126;
<emphasis id="B95670196471FFA9FBCEEA27D47FF8AF" bold="true" box="[1090,1102,1847,1867]" pageId="24" pageNumber="25">K</emphasis>
YC22, PIMUZ 39127,
<emphasis id="B95670196471FFA9FB73EA27D53BF8AF" bold="true" box="[1279,1290,1847,1867]" pageId="24" pageNumber="25">L</emphasis>
YC25, PIMUZ 39128;
<emphasis id="B95670196471FFA9FF6AEA42D0C5F882" bold="true" box="[230,244,1874,1894]" pageId="24" pageNumber="25">N</emphasis>
SHA304C,PIMUZ 39129;
<emphasis id="B95670196471FFA9FE46EA42D1E8F882" bold="true" box="[458,473,1874,1894]" pageId="24" pageNumber="25">O</emphasis>
SHA343C,PIMUZ SQL54979.
<emphasis id="B95670196471FFA9FD5DEA42D2E8F882" bold="true" box="[721,729,1874,1894]" pageId="24" pageNumber="25">J</emphasis>
<taxonomicName id="4C22D7886471FFA9FD52EA42D3ECF882" authority="(MUller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[734,989,1874,1894]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196471FFA9FD52EA42D3A7F882" box="[734,918,1874,1894]" italics="true" pageId="24" pageNumber="25">Guangxidella bransoni</emphasis>
(MÜller)
</taxonomicName>
;YC16, PIMUZ 39140.
<emphasis id="B95670196471FFA9FB1CEA42D483F882" bold="true" box="[1168,1202,1874,1894]" pageId="24" pageNumber="25">PR</emphasis>
<taxonomicName id="4C22D7886471FFA9FB3BEA42D549F882" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1207,1400,1874,1894]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="sp. nov.">
<emphasis id="B95670196471FFA9FB3BEA42D549F882" box="[1207,1400,1874,1894]" italics="true" pageId="24" pageNumber="25">Neospathodus bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FAF1EA42D0F2F864" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B95670196471FFA9FF45EA7CD0E4F864" bold="true" box="[201,213,1900,1920]" pageId="24" pageNumber="25">P</emphasis>
SHA 304C,PIMUZ 39178;
<emphasis id="B95670196471FFA9FE23EA7CD18FF864" bold="true" box="[431,446,1900,1920]" pageId="24" pageNumber="25">Q</emphasis>
SHA333C, PIMUZ 39179;
<emphasis id="B95670196471FFA9FD18EA7CD291F864" bold="true" box="[660,672,1900,1920]" pageId="24" pageNumber="25">R</emphasis>
YC12, PIMUZ 39180
</paragraph>
</caption>
<caption id="DF5DFC836470FFA8FF25EBE0D1CBF8E0" ID-DOI="http://doi.org/10.5281/zenodo.12000298" ID-Zenodo-Dep="12000298" box="[169,506,1776,1796]" httpUri="https://zenodo.org/record/12000298/files/figure.png" pageId="25" pageNumber="26" startId="25.[169,200,1776,1796]" targetBox="[153,1439,238,1800]" targetPageId="25" targetType="figure">
<paragraph id="8B9DAC0B6470FFA8FF25EBE0D1CBF8E0" blockId="25.[169,506,1776,1796]" box="[169,506,1776,1796]" pageId="25" pageNumber="26">
<emphasis id="B95670196470FFA8FF25EBE0D0D9F8E0" bold="true" box="[169,232,1776,1796]" pageId="25" pageNumber="26">Fig. 16</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<paragraph id="8B9DAC0B6473FFABFF21EDFCD292FEE7" blockId="26.[151,775,235,611]" box="[173,675,235,259]" pageId="26" pageNumber="27">
<materialsCitation id="3B4AA6566473FFABFF21EDFCD2AEFEE7" box="[173,671,235,259]" country="Oman" location="Jabral Safra" pageId="26" pageNumber="27" specimenCount="1" typeStatus="holotype">
<emphasis id="B95670196473FFABFF21EDFCD1F2FEE7" box="[173,451,235,259]" italics="true" pageId="26" pageNumber="27">
<typeStatus id="549912A96473FFABFF21EDFCD0D0FEE7" box="[173,225,236,259]" pageId="26" pageNumber="27">Type</typeStatus>
stratum and locality
</emphasis>
.
<location id="8EFDFAD06473FFABFE43EDFBD27CFEE7" LSID="urn:lsid:plazi:treatment:038B1D1D6471FFABFCA1EB1BD18EFD87:8EFDFAD06473FFABFE43EDFBD27CFEE7" box="[463,589,235,259]" country="Oman" name="Jabral Safra" pageId="26" pageNumber="27">Jabral Safra</location>
,
<collectingCountry id="F335EC9B6473FFABFDDBEDFBD2AEFEE7" box="[599,671,235,259]" name="Oman" pageId="26" pageNumber="27">Oman</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DAC0B6473FFABFF21EC1BD18EFD87" blockId="26.[151,775,235,611]" pageId="26" pageNumber="27">
Remarks. Te genus
<taxonomicName id="4C22D7886473FFABFE2FEC1BD278FEC7" authorityName="Orchard" authorityYear="2005" box="[419,585,267,291]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFE2FEC1BD278FEC7" box="[419,585,267,291]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
was introduced as a new genus with a 15-element apparatus by
<bibRefCitation id="EFB3D1FA6473FFABFD24EC3BD0EEFE87" author="Orchard, M. J." pageId="26" pageNumber="27" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard (2005)</bibRefCitation>
and revised by
<bibRefCitation id="EFB3D1FA6473FFABFE19EC5BD28FFE87" author="Goudemand, N. &amp; Orchard, M. J. &amp; Tafforeau, P. &amp; Urdy, S. &amp; Bruehwiler, T. &amp; Brayard, A. &amp; BrUhwiler, T. &amp; Bucher, H." box="[405,702,331,355]" pageId="26" pageNumber="27" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012">Goudemand et al. (2012b)</bibRefCitation>
based on the swapping of the S
<subScript id="17A6AE4E6473FFABFE39EC67D18FFE6C" attach="left" box="[437,446,375,392]" fontSize="7" pageId="26" pageNumber="27">1</subScript>
and S
<subScript id="17A6AE4E6473FFABFD8BEC67D221FE6C" attach="left" box="[519,528,375,392]" fontSize="7" pageId="26" pageNumber="27">2</subScript>
positions. Additional to the
<typeStatus id="549912A96473FFABFF66EC9CD12BFE40" box="[234,282,396,420]" pageId="26" pageNumber="27">type</typeStatus>
species (
<taxonomicName id="4C22D7886473FFABFE01EC9CD212FE47" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[397,547,395,419]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196473FFABFE01EC9CD212FE47" box="[397,547,395,419]" italics="true" pageId="26" pageNumber="27">Nv. abruptus</emphasis>
</taxonomicName>
) and other species whose multi-element apparatus has been reconstructed (e.g.
<taxonomicName id="4C22D7886473FFABFF43ECDCD333FE07" authority="(Goudemand et al., 2012 b)" baseAuthorityName="Goudemand" baseAuthorityYear="2012" box="[207,770,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196473FFABFF43ECDCD1F7FE07" box="[207,454,459,483]" italics="true" pageId="26" pageNumber="27">Nv. pingdingshanensis</emphasis>
(
<bibRefCitation id="EFB3D1FA6473FFABFE5BECDBD2C6FE07" author="Goudemand, N. &amp; Orchard, M. J. &amp; Tafforeau, P. &amp; Urdy, S. &amp; Bruehwiler, T. &amp; Brayard, A. &amp; BrUhwiler, T. &amp; Bucher, H." box="[471,759,459,483]" pageId="26" pageNumber="27" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012">Goudemand et al., 2012b</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C22D7886473FFABFF1BECFCD110FDE7" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[151,289,492,515]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196473FFABFF1BECFCD110FDE7" box="[151,289,492,515]" italics="true" pageId="26" pageNumber="27">Nv. waageni</emphasis>
</taxonomicName>
(unpublished)), we tentatively assign also the following species to
<taxonomicName id="4C22D7886473FFABFE4DEF1BD256FDC7" authorityName="Orchard" authorityYear="2005" box="[449,615,523,547]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFE4DEF1BD256FDC7" box="[449,615,523,547]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
on the basis that they co-occur with S and M elements that are reminiscent of
<taxonomicName id="4C22D7886473FFABFE9FEF5BD188FD87" authorityName="Orchard" authorityYear="2005" box="[275,441,587,611]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95670196473FFABFE9FEF5BD188FD87" box="[275,441,587,611]" italics="true" pageId="26" pageNumber="27">Novispathodus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,252 @@
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<subSubSection id="C338FF806475FFADFF1BEF7BD1E4FD67" box="[151,469,619,643]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B9DAC0B6475FFADFF1BEF7BD1E4FD67" blockId="28.[151,775,235,1667]" box="[151,469,619,643]" pageId="28" pageNumber="29">
<taxonomicName id="4C22D7886475FFADFF1BEF7BD1A4FD67" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,405,619,643]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gryphus" status="sp. nov.">
<emphasis id="B95670196475FFADFF1BEF7BD1A4FD67" box="[151,405,619,643]" italics="true" pageId="28" pageNumber="29">Novispathodus gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFE17EF7CD1E4FD67" box="[411,469,620,643]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFF1BEF9BD17FFD47" box="[151,334,651,675]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B9DAC0B6475FFADFF1BEF9BD17FFD47" blockId="28.[151,775,235,1667]" box="[151,334,651,675]" pageId="28" pageNumber="29">
<figureCitation id="1319B08E6475FFADFF1BEF9BD11AFD47" box="[151,299,651,675]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.12000312" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="28" pageNumber="29">Fig. 17GI, K</figureCitation>
?, L
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFF1BEFDBD1C5FCE7" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B9DAC0B6475FFADFF1BEFDBD336FD07" blockId="28.[151,775,235,1667]" box="[151,775,715,739]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEFDBD139FD07" box="[151,264,715,739]" italics="true" pageId="28" pageNumber="29">Etymology</emphasis>
: from the Greek root gryph referring to the
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEFFBD1C5FCE7" blockId="28.[151,775,235,1667]" box="[151,500,747,776]" pageId="28" pageNumber="29">
hooked shape of the P
<subScript id="17A6AE4E6475FFADFE0BEFE7D1A1FCEC" attach="left" box="[391,400,759,776]" fontSize="7" pageId="28" pageNumber="29">1</subScript>
element.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFF1BEE1BD279FC07" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B9DAC0B6475FFADFF1BEE1BD254FCC7" blockId="28.[151,775,235,1667]" box="[151,613,779,803]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEE1BD0CEFCC7" box="[151,255,779,803]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="549912A96475FFADFF1BEE1BD0C9FCC7" box="[151,248,779,803]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
:
</emphasis>
specimen illustrated in
<figureCitation id="1319B08E6475FFADFD8FEE1BD250FCC7" box="[515,609,779,803]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.12000312" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="28" pageNumber="29">Fig. 17H</figureCitation>
.
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEE3CD250FCA7" blockId="28.[151,775,235,1667]" box="[151,609,811,835]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEE3CD0CEFCA7" box="[151,255,812,835]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="549912A96475FFADFF1BEE3CD0C8FCA7" box="[151,249,812,835]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
:
</emphasis>
specimen illustrated in
<figureCitation id="1319B08E6475FFADFD8FEE3BD26CFCA7" box="[515,605,811,835]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.12000312" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="28" pageNumber="29">
Fig.
<specimenCount id="9D2467826475FFADFDBDEE3BD250FCA7" box="[561,609,811,835]" count="17" pageId="28" pageNumber="29" type="generic">17L.</specimenCount>
</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEE5CD337FC87" blockId="28.[151,775,235,1667]" box="[151,774,843,867]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEE5CD119FC87" box="[151,296,843,867]" italics="true" pageId="28" pageNumber="29">Type locality:</emphasis>
Qiakong, Luolou formation,
<collectingRegion id="49E662E96475FFADFDE9EE5BD0F7FC67" country="China" name="Guizhou" pageId="28" pageNumber="29">
Guizhou Prov-
<paragraph id="8B9DAC0B6475FFADFF1BEE7BD12BFC67" blockId="28.[151,775,235,1667]" box="[151,282,875,899]" pageId="28" pageNumber="29">
ince,
<collectingCountry id="F335EC9B6475FFADFF5CEE7BD124FC67" box="[208,277,875,899]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
.
</paragraph>
</collectingRegion>
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEE9CD336FC47" blockId="28.[151,775,235,1667]" box="[151,775,907,931]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEE9CD13DFC47" box="[151,268,907,931]" italics="true" pageId="28" pageNumber="29">Type level</emphasis>
: Luolou Formation, within latest Smithian
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEEBBD12EFC27" blockId="28.[151,775,235,1667]" box="[151,287,939,963]" pageId="28" pageNumber="29">black shales.</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEEDBD279FC07" blockId="28.[151,775,235,1667]" box="[151,584,971,995]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BEEDBD1B5FC07" box="[151,388,971,995]" italics="true" pageId="28" pageNumber="29">Number of specimens.</emphasis>
ca.
<specimenCount id="9D2467826475FFADFE22EEDBD275FC07" box="[430,580,971,995]" count="10" pageId="28" pageNumber="29" type="generic">10 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFF1BE91CD1ACFA67" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B9DAC0B6475FFADFF1BE91CD1ACFA67" blockId="28.[151,775,235,1667]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BE91CD130FBC7" box="[151,257,1036,1059]" italics="true" pageId="28" pageNumber="29">Diagnosis</emphasis>
. A species with a short segminate-to-segminis- caphateP
<subScript id="17A6AE4E6475FFADFE8EE927D13AFBAC" attach="left" box="[258,267,1079,1096]" fontSize="7" pageId="28" pageNumber="29">1</subScript>
element with 3 to 5 highly recurved denticles and a relatively large, sub-rounded basal cavity.
<emphasis id="B95670196475FFADFF1BE97CD12AFB67" box="[151,283,1132,1155]" italics="true" pageId="28" pageNumber="29">Description.</emphasis>
Te P
<subScript id="17A6AE4E6475FFADFEEFE967D15DFB6C" attach="left" box="[355,364,1143,1160]" fontSize="7" pageId="28" pageNumber="29">1</subScript>
element is segminate to segminisca- phate, the sub-rounded basal cavity extending to most of the length of the element with a tapering at the anterior end. Te denticles are mostly fused and highly recurved posteriorly: although the denticles get larger from the anterior to the posterior, except for the posteriormost one, the height of the carina looks sub-uniform because of the way the denticles are recurved. Te length-to- height ratio is about 1:1.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFF1BE89BD3DDFD07" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B9DAC0B6475FFADFF1BE89BD336FA47" blockId="28.[151,775,235,1667]" box="[151,775,1419,1443]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFF1BE89BD0CDFA47" box="[151,252,1419,1443]" italics="true" pageId="28" pageNumber="29">Remarks.</emphasis>
Tese elements resemble strongly the homolo-
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BE8BCD336FA27" blockId="28.[151,775,235,1667]" box="[151,775,1451,1476]" pageId="28" pageNumber="29">
gous ones of
<taxonomicName id="4C22D7886475FFADFEA7E8BCD22CFA27" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[299,541,1451,1475]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196475FFADFEA7E8BCD17BFA27" box="[299,330,1452,1475]" italics="true" pageId="28" pageNumber="29">Nv</emphasis>
.
<emphasis id="B95670196475FFADFED5E8BBD22CFA27" box="[345,541,1451,1475]" italics="true" pageId="28" pageNumber="29">pingdingshanensis</emphasis>
</taxonomicName>
, except that they are
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BE8DBD336FA07" blockId="28.[151,775,235,1667]" box="[151,775,1483,1507]" pageId="28" pageNumber="29">much shorter and bear less denticles, which are usually</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BE8FCD336F9E7" blockId="28.[151,775,235,1667]" box="[151,775,1515,1540]" pageId="28" pageNumber="29">more recurved. Still, some may be confused with broken</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEB1BD337F9C7" blockId="28.[151,775,235,1667]" box="[151,774,1547,1571]" pageId="28" pageNumber="29">
elements of
<taxonomicName id="4C22D7886475FFADFEABEB1CD22AF9C7" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[295,539,1547,1571]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196475FFADFEABEB1CD177F9C7" box="[295,326,1548,1571]" italics="true" pageId="28" pageNumber="29">Nv</emphasis>
.
<emphasis id="B95670196475FFADFEDBEB1BD22AF9C7" box="[343,539,1547,1571]" italics="true" pageId="28" pageNumber="29">pingdingshanensis</emphasis>
</taxonomicName>
whose anteriormost
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEB3BD336F9A7" blockId="28.[151,775,235,1667]" box="[151,775,1579,1603]" pageId="28" pageNumber="29">
end is missing (
<figureCitation id="1319B08E6475FFADFEB2EB3BD1ACF9A7" box="[318,413,1579,1603]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AC, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. DF, MP, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao &amp; Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao &amp; Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.12000312" httpUri="https://zenodo.org/record/12000312/files/figure.png" pageId="28" pageNumber="29">Fig. 17G</figureCitation>
). Hence the anterior end must be
</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEB5BD336F987" blockId="28.[151,775,235,1667]" box="[151,775,1611,1635]" pageId="28" pageNumber="29">inspected for traces of breakage. Although the specimens</paragraph>
<paragraph id="8B9DAC0B6475FFADFF1BEB7CD3DDFD07" blockId="28.[151,775,235,1667]" lastBlockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
we have seen do not show obvious, surfacial traces of breakage, it is not excluded that some of these elements were broken and were subsequently repaired, thus covering such traces at the surface of the crown. Goudemand and Orchard (in
<bibRefCitation id="EFB3D1FA6475FFADFC6AEC5BD4CBFE87" author="Goudemand, N. &amp; Orchard, M. J. &amp; Tafforeau, P. &amp; Urdy, S. &amp; Bruehwiler, T. &amp; Brayard, A. &amp; BrUhwiler, T. &amp; Bucher, H." box="[998,1274,331,355]" pageId="28" pageNumber="29" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012">Goudemand et al., 2012b</bibRefCitation>
) differentiated similar but longer elements as
<taxonomicName id="4C22D7886475FFADFB0EEC7CD5ADFE67" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[1154,1436,363,387]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196475FFADFB0EEC7CD490FE67" box="[1154,1185,364,387]" italics="true" pageId="28" pageNumber="29">Nv</emphasis>
. aff.
<emphasis id="B95670196475FFADFB54EC7BD5ADFE67" box="[1240,1436,363,387]" italics="true" pageId="28" pageNumber="29">pingdingshanensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA6475FFADFCB8EC9BD47AFE47" author="Goudemand, N. &amp; Orchard, M. J. &amp; Tafforeau, P. &amp; Urdy, S. &amp; Bruehwiler, T. &amp; Brayard, A. &amp; BrUhwiler, T. &amp; Bucher, H." box="[820,1099,395,419]" pageId="28" pageNumber="29" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012">Goudemand et al., 2012b</bibRefCitation>
, figs. 2K, 2L) because they are lower and their denticles more recurved than in elements of
<taxonomicName id="4C22D7886475FFADFCC7ECDCD40DFE07" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[843,1084,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196475FFADFCC7ECDCD35BFE07" box="[843,874,460,483]" italics="true" pageId="28" pageNumber="29">Nv</emphasis>
.
<emphasis id="B95670196475FFADFCF4ECDBD40DFE07" box="[888,1084,459,483]" italics="true" pageId="28" pageNumber="29">pingdingshanensis</emphasis>
</taxonomicName>
: their denticles are so recurved that their upper profile appears straight; as such they are reminiscent of the present species, with whom they may be closely related. In comparison with
<taxonomicName id="4C22D7886475FFADFB74EF3CD5ADFDA7" box="[1272,1436,555,579]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="soleiformis">
<emphasis id="B95670196475FFADFB74EF3CD5ADFDA7" box="[1272,1436,555,579]" italics="true" pageId="28" pageNumber="29">Nv. soleiformis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA6475FFADFCB8EF5BD403FD87" author="Zhao, L. &amp; Tong, J. &amp; Zhang, S. &amp; Sun, Z." box="[820,1074,587,611]" pageId="28" pageNumber="29" pagination="207 - 216" refId="ref49144" refString="Zhao, L., Tong, J., Zhang, S., &amp; Sun, Z. (2008). An update of conodonts in the Induan - Olenekian boundary strata at west Pingdingshan section, Chaohu, Anhui Province. Journal of China University of Geosciences, 19 (3), 207 - 216. https: // doi. org / 10.1016 / S 1002 - 0705 (08) 60040 - 0" type="journal article" year="2008">Zhao &amp; Orchard, 2008</bibRefCitation>
)
<taxonomicName id="4C22D7886475FFADFBC8EF5CD4F7FD87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1092,1222,587,611]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gryphus" status="n. sp.">
<emphasis id="B95670196475FFADFBC8EF5CD4F7FD87" box="[1092,1222,587,611]" italics="true" pageId="28" pageNumber="29">Nv. gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFB43EF5CD53AFD87" box="[1231,1291,588,611]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
has a higher carina with less numerous but more posteriorly recurved denticles. Furthermore, the basal cavity of
<taxonomicName id="4C22D7886475FFADFA8CEF9CD5B1FD47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1280,1408,651,675]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="gryphus" status="n.">
<emphasis id="B95670196475FFADFA8CEF9CD5B1FD47" box="[1280,1408,651,675]" italics="true" pageId="28" pageNumber="29">Nv. gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626475FFADFA04EF9CD5ADFD47" box="[1416,1436,652,675]" pageId="28" pageNumber="29">n.</taxonomicNameLabel>
sp. does not extend as a wide deep groove anteriorly like in
<taxonomicName id="4C22D7886475FFADFCC4EFDCD3D7FD07" box="[840,998,715,739]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="soleiformis">
<emphasis id="B95670196475FFADFCC4EFDCD3D7FD07" box="[840,998,715,739]" italics="true" pageId="28" pageNumber="29">Nv. soleiformis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF806475FFADFCA1EFFCD346FCA7" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8B9DAC0B6475FFADFCA1EFFCD346FCA7" blockId="28.[813,1437,235,1667]" pageId="28" pageNumber="29">
<emphasis id="B95670196475FFADFCA1EFFCD39FFCE7" box="[813,942,748,771]" italics="true" pageId="28" pageNumber="29">Occurrence.</emphasis>
Luolou Formation, within the latest Smith- ian black shales,
<collectingRegion id="49E662E96475FFADFC6DEE1BD40FFCC7" box="[993,1086,779,803]" country="China" name="Guangxi" pageId="28" pageNumber="29">Guangxi</collectingRegion>
and
<collectingRegion id="49E662E96475FFADFBFEEE1BD4E2FCC7" box="[1138,1235,779,803]" country="China" name="Guizhou" pageId="28" pageNumber="29">Guizhou</collectingRegion>
, South
<collectingCountry id="F335EC9B6475FFADFAA8EE1BD557FCC7" box="[1316,1382,779,803]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
(this study).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,558 @@
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<treatment id="038B1D1D6476FF92FCA1E8FBD26EFDE7" LSID="urn:lsid:plazi:treatment:038B1D1D6476FF92FCA1E8FBD26EFDE7" httpUri="http://treatment.plazi.org/id/038B1D1D6476FF92FCA1E8FBD26EFDE7" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32">
<subSubSection id="C338FF806476FFAEFCA1E8FBD536F9E7" box="[813,1287,1515,1539]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B9DAC0B6476FFAEFCA1E8FBD536F9E7" blockId="31.[813,1292,1515,1571]" box="[813,1287,1515,1539]" pageId="31" pageNumber="32">
<heading id="D0D51B676476FFAEFCA1E8FBD536F9E7" box="[813,1287,1515,1539]" fontSize="9" level="2" pageId="31" pageNumber="32" reason="8">
<taxonomicName id="4C22D7886476FFAEFCA1E8FBD536F9E7" authority="(Orchard, 1995)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[813,1287,1515,1539]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196476FFAEFCA1E8FBD462F9E7" box="[813,1107,1515,1539]" italics="true" pageId="31" pageNumber="32">Novispathodus brevissimus</emphasis>
(
<bibRefCitation id="EFB3D1FA6476FFAEFBECE8FBD4CFF9E7" author="Orchard, M. J." box="[1120,1278,1515,1539]" pageId="31" pageNumber="32" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806476FF92FCA1EB1BD26EFDE7" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32" type="description">
<paragraph id="8B9DAC0B6476FFAEFCA1EB1BD53DF9C7" blockId="31.[813,1292,1515,1571]" box="[813,1292,1547,1571]" pageId="31" pageNumber="32">
<figureCitation id="1319B08E6476FFAEFCA1EB1BD3D8F9C7" box="[813,1001,1547,1571]" captionStart="Fig" captionStartId="30.[169,200,1700,1720]" captionTargetBox="[153,1439,238,1918]" captionTargetId="figure-16@30.[229,1363,254,1687]" captionTargetPageId="30" captionText="Fig. 19 Novispathodinae from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A Novispathodus cf.brevissimus (Orchard); LIL515C,PIMUZ 39199. B, F Novispathodus brevissimus (Orchard); B LIL515B, PIMUZ 39200; F BAN1, PIMUZ 39201. C Triassospathodus homeri (Bender); LIL515D,PIMUZ 39317. D, H Novispathodus?brevissimus (Orchard); D LIL515D,PIMUZ 39202; H LIL514A, PIMUZ 39203. E Triassospathodus cf.homeri (Bender); LIL515C, PIMUZ 39318. G, K, OQ Novispathodus n. sp. A; G LIL504,PIMUZ 39251; K LIL504,PIMUZ 39252; O LIL505, PIMUZ 39253; P LIL505, PIMUZ 39254;Q LIL505,PIMUZ 39255. I, L, M, Triassospathodus symmetricus (Orchard); I QIA141, PIMUZ 39336; L LIL512, PIMUZ 39337; M LIL514A, PIMUZ 39338. J Novispathodus ex gr.pingdingshanensis (Zhao &amp; Orchard); QIA138, PIMUZ 39237. N Triassospathodus cf. symmetricus; LIL514B, PIMUZ 39339.R Novispathodus praebrevissimus n.sp; LIL505, PIMUZ 39289" figureDoi="http://doi.org/10.5281/zenodo.12000347" httpUri="https://zenodo.org/record/12000347/files/figure.png" pageId="31" pageNumber="32">Figs. 19A, B, D, F</figureCitation>
;
<figureCitation id="1319B08E6476FFAEFC7FEB1BD44BF9C7" box="[1011,1146,1547,1571]" captionStart="Fig" captionStartId="33.[169,200,1450,1470]" captionTargetBox="[153,1439,238,1588]" captionTargetId="figure-89@33.[229,1363,254,1437]" captionTargetPageId="33" captionText="Fig. 21 Novispathodinae from Qiakong,Laren, Shanggang and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AJ, L, M, Novispathodus brevissimus (Orchard); A LIL515A, PIMUZ 39184; B LIL513A,PIMUZ 39185; C BAN2,PIMUZ 39186; D SHA312, PIMUZ 39187; E SHA313, PIMUZ 39188; F LIL515A, PIMUZ 39189; G LIL509, PIMUZ 39190;H LIL509, PIMUZ 39191; I LIL514B, PIMUZ 39192;J LIL509, PIMUZ 39193;L QIA141, PIMUZ 39195; M QIA144, PIMUZ 39197.K, Novispathodus clinatus (Orchard and Sweet in Orchard); BAN5, PIMUZ 39207" figureDoi="http://doi.org/10.5281/zenodo.12000375" httpUri="https://zenodo.org/record/12000375/files/figure.png" pageId="31" pageNumber="32">21AJ, L, M</figureCitation>
;
<figureCitation id="1319B08E6476FFAEFB08EB1BD532F9C7" box="[1156,1283,1547,1571]" captionStart="Fig" captionStartId="34.[170,201,968,988]" captionTargetBox="[153,1438,238,1081]" captionTargetId="figure-414@34.[229,1363,254,955]" captionTargetPageId="34" captionText="Fig. 22 Novispathodinae from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Icriospathodus cf. crassatus? (Orchard); A BAN1, PIMUZ 39168; D LIL513A,PIMUZ 39169.B, E Novispathodus brevissimus (Orchard); B LIL513B,PIMUZ 39194; E LIL514A,PIMUZ SQL55056. C, F Novispathodus cf, brevissimus (Orchard); C LIL513A, PIMUZ 39196; F, LIL514A, PIMUZ 39198" figureDoi="http://doi.org/10.5281/zenodo.12000396" httpUri="https://zenodo.org/record/12000396/files/figure.png" pageId="31" pageNumber="32">22B, C, E, F</figureCitation>
?
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFF25EB88D1B2F948" blockId="31.[169,387,1688,1708]" box="[169,387,1688,1708]" pageId="31" pageNumber="32">(See figure on next page.)</paragraph>
<caption id="DF5DFC836476FFAEFF25EBA3D4D8F865" pageId="31" pageNumber="32" startId="31.[169,200,1715,1735]" targetBox="[153,1439,238,1800]" targetPageId="32">
<paragraph id="8B9DAC0B6476FFAEFF25EBA3D4D8F865" blockId="31.[169,1420,1713,1921]" pageId="31" pageNumber="32">
<figureCitation id="1319B08E6476FFAEFF25EBA3D0D9F923" box="[169,232,1715,1735]" captionStart="Fig" captionStartId="32.[169,200,1776,1796]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-16@32.[229,1363,254,1763]" captionTargetPageId="32" captionText="Fig. 20 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000362" httpUri="https://zenodo.org/record/12000362/files/figure.png" pageId="31" pageNumber="32">
<emphasis id="B95670196476FFAEFF25EBA3D0D9F923" bold="true" box="[169,232,1715,1735]" pageId="31" pageNumber="32">Fig. 20</emphasis>
</figureCitation>
<taxonomicName id="4C22D7886476FFAEFF7EEBA3D1B2F923" authorityName="Orchard" authorityYear="2005" box="[242,387,1715,1735]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
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elements if not specifically identified otherwise.
<emphasis id="B95670196476FFAEFDCDEBDDD256F905" bold="true" box="[577,615,1741,1761]" pageId="31" pageNumber="32">AG</emphasis>
,
<emphasis id="B95670196476FFAEFDFCEBDDD2A8F905" bold="true" box="[624,665,1741,1761]" pageId="31" pageNumber="32">KM</emphasis>
,
<emphasis id="B95670196476FFAEFD2DEBDDD281F905" bold="true" box="[673,688,1741,1761]" pageId="31" pageNumber="32">Q</emphasis>
,
<emphasis id="B95670196476FFAEFD34EBDDD2F4F905" bold="true" box="[696,709,1741,1761]" pageId="31" pageNumber="32">X</emphasis>
<taxonomicName id="4C22D7886476FFAEFD46EBDDD374F905" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[714,837,1741,1761]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B95670196476FFAEFD46EBDDD374F905" box="[714,837,1741,1761]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886476FFAEFCF6EBDDD42BF905" authority="(Orchard)" baseAuthorityName="Orchard" box="[890,1050,1741,1761]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFCF6EBDDD3F5F905" box="[890,964,1741,1761]" italics="true" pageId="31" pageNumber="32">abruptus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196476FFAEFBADEBDDD41EF905" bold="true" box="[1057,1071,1741,1761]" pageId="31" pageNumber="32">A</emphasis>
LAR212, PIMUZ 39213;
<emphasis id="B95670196476FFAEFB79EBDDD530F905" bold="true" box="[1269,1281,1741,1761]" pageId="31" pageNumber="32">B</emphasis>
LAR207, PIMUZ 39214;
<emphasis id="B95670196476FFAEFF6AEBF8D0C2F918" bold="true" box="[230,243,1768,1788]" pageId="31" pageNumber="32">C</emphasis>
LAR205,PIMUZ 39215;
<emphasis id="B95670196476FFAEFE35EBF8D1F9F918" bold="true" box="[441,456,1768,1788]" pageId="31" pageNumber="32">D</emphasis>
LAR202, PIMUZ 39216;
<emphasis id="B95670196476FFAEFD02EBF8D2A8F918" bold="true" box="[654,665,1768,1788]" pageId="31" pageNumber="32">E</emphasis>
LAR202, PIMUZ 39217;
<emphasis id="B95670196476FFAEFCECEBF8D35AF918" bold="true" box="[864,875,1768,1788]" pageId="31" pageNumber="32">F</emphasis>
LIL506, PIMUZ 39218;
<emphasis id="B95670196476FFAEFBA5EBF8D406F918" bold="true" box="[1065,1079,1768,1788]" pageId="31" pageNumber="32">G</emphasis>
LAR210, PIMUZ 39219;
<emphasis id="B95670196476FFAEFB72EBF8D53BF918" bold="true" box="[1278,1290,1768,1788]" pageId="31" pageNumber="32">K</emphasis>
LIL505,PIMUZ 39220;
<emphasis id="B95670196476FFAEFF6AEA13D0C0F8F3" bold="true" box="[230,241,1795,1815]" pageId="31" pageNumber="32">L</emphasis>
LIL506, PIMUZ 39221;
<emphasis id="B95670196476FFAEFE22EA13D1F1F8F3" bold="true" box="[430,448,1795,1815]" pageId="31" pageNumber="32">M</emphasis>
LIL505,PIMUZ 39222;
<emphasis id="B95670196476FFAEFDF1EA13D2BDF8F3" bold="true" box="[637,652,1795,1815]" pageId="31" pageNumber="32">Q</emphasis>
LAR203,PIMUZ 39223;
<emphasis id="B95670196476FFAEFCDFEA13D351F8F3" bold="true" box="[851,864,1795,1815]" pageId="31" pageNumber="32">X</emphasis>
QIA136,PIMUZ 39224.
<emphasis id="B95670196476FFAEFBA9EA13D402F8F3" bold="true" box="[1061,1075,1795,1815]" pageId="31" pageNumber="32">H</emphasis>
,
<emphasis id="B95670196476FFAEFBB0EA13D478F8F3" bold="true" box="[1084,1097,1795,1815]" pageId="31" pageNumber="32">
<collectionCode id="ED3334CE6476FFAEFBB0EA13D478F8F3" box="[1084,1097,1795,1815]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="32" type="Museum">V</collectionCode>
</emphasis>
<taxonomicName id="4C22D7886476FFAEFBC1EA13D4D4F8F3" box="[1101,1253,1795,1815]" family="Gondolellidae" genus="Novispathodus" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B95670196476FFAEFBC1EA13D4F9F8F3" box="[1101,1224,1795,1815]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
sp.
</taxonomicName>
indet.;
<emphasis id="B95670196476FFAEFAADEA13D51EF8F3" bold="true" box="[1313,1327,1795,1815]" pageId="31" pageNumber="32">H</emphasis>
QIA136, PIMUZ 39291;
<emphasis id="B95670196476FFAEFEAFEA0DD101F8D5" bold="true" box="[291,304,1821,1841]" pageId="31" pageNumber="32">
<collectionCode id="ED3334CE6476FFAEFEAFEA0DD101F8D5" box="[291,304,1821,1841]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="32" type="Museum">V</collectionCode>
</emphasis>
, QIA134, PIMUZ 39292.
<emphasis id="B95670196476FFAEFE75EA0DD1CEF8D5" bold="true" box="[505,511,1821,1841]" pageId="31" pageNumber="32">I</emphasis>
,
<emphasis id="B95670196476FFAEFD8BEA0DD23EF8D5" bold="true" box="[519,527,1821,1841]" pageId="31" pageNumber="32">J</emphasis>
,
<emphasis id="B95670196476FFAEFD94EA0DD215F8D5" bold="true" box="[536,548,1821,1841]" pageId="31" pageNumber="32">P</emphasis>
,
<emphasis id="B95670196476FFAEFDA0EA0DD276F8D5" bold="true" box="[556,583,1821,1841]" pageId="31" pageNumber="32">AA</emphasis>
<taxonomicName id="4C22D7886476FFAEFDC0EA0DD2F6F8D5" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[588,711,1821,1841]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196476FFAEFDC0EA0DD2F6F8D5" box="[588,711,1821,1841]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626476FFAEFD47EA0DD2C9F8D5" box="[715,760,1821,1841]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
Z;
<emphasis id="B95670196476FFAEFC82EA0DD325F8D5" bold="true" box="[782,788,1821,1841]" pageId="31" pageNumber="32">I</emphasis>
QIA138, PIMUZ 39247;
<emphasis id="B95670196476FFAEFC55EA0DD3D0F8D5" bold="true" box="[985,993,1821,1841]" pageId="31" pageNumber="32">J</emphasis>
QIA138,PIMUZ 39248;
<emphasis id="B95670196476FFAEFB2BEA0DD482F8D5" bold="true" box="[1191,1203,1821,1841]" pageId="31" pageNumber="32">P</emphasis>
LIL506, PIMUZ 39249;
<emphasis id="B95670196476FFAEFAFDEA0DD5BDF8D5" bold="true" box="[1393,1420,1821,1841]" pageId="31" pageNumber="32">AA</emphasis>
QIA136, PIMUZ 39250.
<emphasis id="B95670196476FFAEFEE6EA28D149F8A8" bold="true" box="[362,376,1848,1868]" pageId="31" pageNumber="32">N</emphasis>
,
<emphasis id="B95670196476FFAEFE0DEA28D1A1F8A8" bold="true" box="[385,400,1848,1868]" pageId="31" pageNumber="32">O</emphasis>
,
<emphasis id="B95670196476FFAEFE14EA28D192F8A8" bold="true" box="[408,419,1848,1868]" pageId="31" pageNumber="32">S</emphasis>
<taxonomicName id="4C22D7886476FFAEFE24EA28D212F8A8" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[424,547,1848,1868]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="B95670196476FFAEFE24EA28D212F8A8" box="[424,547,1848,1868]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886476FFAEFDD4EA28D3B9F8A8" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[600,904,1848,1868]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFDD4EA28D2C0F8A8" box="[600,753,1848,1868]" italics="true" pageId="31" pageNumber="32">pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
;
<emphasis id="B95670196476FFAEFC1CEA28D3AFF8A8" bold="true" box="[912,926,1848,1868]" pageId="31" pageNumber="32">N</emphasis>
LIL506, PIMUZ 39234;
<emphasis id="B95670196476FFAEFBD0EA28D45AF8A8" bold="true" box="[1116,1131,1848,1868]" pageId="31" pageNumber="32">O</emphasis>
LIL505, PIMUZ 39235;
<emphasis id="B95670196476FFAEFAA5EA28D505F8A8" bold="true" box="[1321,1332,1848,1868]" pageId="31" pageNumber="32">S</emphasis>
QIA136, PIMUZ 39236.
<emphasis id="B95670196476FFAEFEAFEA43D11EF883" bold="true" box="[291,303,1875,1895]" pageId="31" pageNumber="32">R</emphasis>
sp.indet.; LIL511, PIMUZ 39296.
<emphasis id="B95670196476FFAEFDCDEA43D27DF883" bold="true" box="[577,588,1875,1895]" pageId="31" pageNumber="32">T</emphasis>
,
<emphasis id="B95670196476FFAEFDD9EA43D250F883" bold="true" box="[597,609,1875,1895]" pageId="31" pageNumber="32">Y</emphasis>
,
<emphasis id="B95670196476FFAEFDE5EA43D244F883" bold="true" box="[617,629,1875,1895]" pageId="31" pageNumber="32">Z</emphasis>
<taxonomicName id="4C22D7886476FFAEFDF6EA43D2C4F883" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[634,757,1875,1895]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196476FFAEFDF6EA43D2C4F883" box="[634,757,1875,1895]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626476FFAEFD75EA43D317F883" box="[761,806,1875,1895]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
A;
<emphasis id="B95670196476FFAEFCB2EA43D378F883" bold="true" box="[830,841,1875,1895]" pageId="31" pageNumber="32">T</emphasis>
QIA134,PIMUZ 39256,
<emphasis id="B95670196476FFAEFB82EA43D42BF883" bold="true" box="[1038,1050,1875,1895]" pageId="31" pageNumber="32">Y</emphasis>
QIA135, PIMUZ 39257;
<emphasis id="B95670196476FFAEFB6CEA43D4DDF883" bold="true" box="[1248,1260,1875,1895]" pageId="31" pageNumber="32">Z</emphasis>
QIA134, PIMUZ 39258.
<emphasis id="B95670196476FFAEFF6AEA7DD0C5F865" bold="true" box="[230,244,1901,1921]" pageId="31" pageNumber="32">U</emphasis>
<taxonomicName id="4C22D7886476FFAEFF74EA7DD269F865" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" box="[248,600,1901,1921]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="expansus">
<emphasis id="B95670196476FFAEFF74EA7DD1F2F865" box="[248,451,1901,1921]" italics="true" pageId="31" pageNumber="32">Novispathodus expansus</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; QIA134, PIMUZ 39242.
<emphasis id="B95670196476FFAEFCAEEA7DD304F865" bold="true" box="[802,821,1901,1921]" pageId="31" pageNumber="32">W</emphasis>
<taxonomicName id="4C22D7886476FFAEFCB5EA7DD3C8F865" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[825,1017,1901,1921]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="gryphus" status="sp. nov.">
<emphasis id="B95670196476FFAEFCB5EA7DD3C8F865" box="[825,1017,1901,1921]" italics="true" pageId="31" pageNumber="32">Novispathodus gryphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626476FFAEFC72EA7DD41DF865" box="[1022,1068,1901,1921]" pageId="31" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
; QIA135, PIMUZ 39246
</paragraph>
</caption>
<caption id="DF5DFC836449FF91FF25EBE0D1CBF8E0" ID-DOI="http://doi.org/10.5281/zenodo.12000362" ID-Zenodo-Dep="12000362" box="[169,506,1776,1796]" httpUri="https://zenodo.org/record/12000362/files/figure.png" pageId="32" pageNumber="33" startId="32.[169,200,1776,1796]" targetBox="[153,1439,238,1800]" targetPageId="32" targetType="figure">
<paragraph id="8B9DAC0B6449FF91FF25EBE0D1CBF8E0" blockId="32.[169,506,1776,1796]" box="[169,506,1776,1796]" pageId="32" pageNumber="33">
<emphasis id="B95670196449FF91FF25EBE0D0D9F8E0" bold="true" box="[169,232,1776,1796]" pageId="32" pageNumber="33">Fig. 20</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<caption id="DF5DFC836448FF90FF25E8BAD1C2F9CC" ID-DOI="http://doi.org/10.5281/zenodo.12000375" ID-Zenodo-Dep="12000375" httpUri="https://zenodo.org/record/12000375/files/figure.png" pageId="33" pageNumber="34" startId="33.[169,200,1450,1470]" targetBox="[153,1439,238,1588]" targetPageId="33" targetType="figure">
<paragraph id="8B9DAC0B6448FF90FF25E8BAD549FA5A" blockId="33.[169,1413,1449,1576]" box="[169,1400,1449,1470]" pageId="33" pageNumber="34">
<emphasis id="B95670196448FF90FF25E8BAD0D9FA5A" bold="true" box="[169,232,1450,1470]" pageId="33" pageNumber="34">Fig. 21</emphasis>
<taxonomicName id="4C22D7886448FF90FF7EE8BAD1B2FA5A" authorityName="Orchard" authorityYear="2005" box="[242,387,1450,1470]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Qiakong,Laren, Shanggang and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered
</paragraph>
<paragraph id="8B9DAC0B6448FF90FF25E8D4D1C2F9CC" blockId="33.[169,1413,1449,1576]" pageId="33" pageNumber="34">
to be P
<subScript id="17A6AE4E6448FF90FF69E8DED0DDFA38" attach="left" box="[229,236,1486,1500]" fontSize="6" pageId="33" pageNumber="34">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B95670196448FF90FD05E8D4D298FA3C" bold="true" box="[649,681,1476,1496]" pageId="33" pageNumber="34">AJ</emphasis>
,
<emphasis id="B95670196448FF90FD3EE8D4D28CFA3C" bold="true" box="[690,701,1476,1496]" pageId="33" pageNumber="34">L</emphasis>
,
<emphasis id="B95670196448FF90FD49E8D4D2E6FA3C" bold="true" box="[709,727,1476,1496]" pageId="33" pageNumber="34">M</emphasis>
,
<taxonomicName id="4C22D7886448FF90FD53E8D4D420FA3C" authority="(Orchard)" baseAuthorityName="Orchard" box="[735,1041,1476,1496]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196448FF90FD53E8D4D38BFA3C" box="[735,954,1476,1496]" italics="true" pageId="33" pageNumber="34">Novispathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B95670196448FF90FB94E8D4D417FA3C" bold="true" box="[1048,1062,1476,1496]" pageId="33" pageNumber="34">A</emphasis>
LIL515A, PIMUZ 39184;
<emphasis id="B95670196448FF90FB7CE8D4D4CDFA3C" bold="true" box="[1264,1276,1476,1496]" pageId="33" pageNumber="34">B</emphasis>
LIL513A,PIMUZ 39185;
<emphasis id="B95670196448FF90FF6AE8CFD0C2FA17" bold="true" box="[230,243,1503,1523]" pageId="33" pageNumber="34">C</emphasis>
BAN2,PIMUZ 39186;
<emphasis id="B95670196448FF90FE24E8CFD186FA17" bold="true" box="[424,439,1503,1523]" pageId="33" pageNumber="34">D</emphasis>
SHA312, PIMUZ 39187;
<emphasis id="B95670196448FF90FD0DE8CFD2BDFA17" bold="true" box="[641,652,1503,1523]" pageId="33" pageNumber="34">E</emphasis>
SHA313, PIMUZ 39188;
<emphasis id="B95670196448FF90FCDAE8CFD350FA17" bold="true" box="[854,865,1503,1523]" pageId="33" pageNumber="34">F</emphasis>
LIL515A, PIMUZ 39189;
<emphasis id="B95670196448FF90FBA7E8CFD408FA17" bold="true" box="[1067,1081,1503,1523]" pageId="33" pageNumber="34">G</emphasis>
LIL509, PIMUZ 39190;
<emphasis id="B95670196448FF90FB7AE8CFD535FA17" bold="true" box="[1270,1284,1503,1523]" pageId="33" pageNumber="34">H</emphasis>
LIL509, PIMUZ 39191;
<emphasis id="B95670196448FF90FF6AE8EAD0DDF9EA" bold="true" box="[230,236,1530,1550]" pageId="33" pageNumber="34">I</emphasis>
LIL514B, PIMUZ 39192;
<emphasis id="B95670196448FF90FE3FE8EAD18AF9EA" bold="true" box="[435,443,1530,1550]" pageId="33" pageNumber="34">J</emphasis>
LIL509, PIMUZ 39193;
<emphasis id="B95670196448FF90FDF5E8EAD2B5F9EA" bold="true" box="[633,644,1530,1550]" pageId="33" pageNumber="34">L</emphasis>
QIA141, PIMUZ 39195;
<emphasis id="B95670196448FF90FCC5E8EAD36AF9EA" bold="true" box="[841,859,1530,1550]" pageId="33" pageNumber="34">M</emphasis>
QIA144, PIMUZ 39197.
<emphasis id="B95670196448FF90FBACE8EAD41DF9EA" bold="true" box="[1056,1068,1530,1550]" pageId="33" pageNumber="34">K</emphasis>
,
<taxonomicName id="4C22D7886448FF90FBB9E8EAD4C5F9EA" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[1077,1268,1530,1550]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="clinatus">
<emphasis id="B95670196448FF90FBB9E8EAD4C5F9EA" box="[1077,1268,1530,1550]" italics="true" pageId="33" pageNumber="34">Novispathodus clinatus</emphasis>
</taxonomicName>
(Orchard and Sweet in Orchard); BAN5, PIMUZ 39207
</paragraph>
</caption>
<paragraph id="8B9DAC0B6448FF90FF1BEB96D0E3F95A" blockId="33.[151,775,1670,1854]" pageId="33" pageNumber="34">
1981
<taxonomicName id="4C22D7886448FF90FF56EB96D255F97A" authority="Bender" authorityName="Bender" box="[218,612,1670,1694]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B95670196448FF90FF56EB96D235F97A" box="[218,516,1670,1694]" italics="true" pageId="33" pageNumber="34">Neospathodus triangularis</emphasis>
Bender
</taxonomicName>
; Koike, pl. 1, fig. 6.
</paragraph>
<paragraph id="8B9DAC0B6448FF90FF1BEBD6D1AEF91A" blockId="33.[151,775,1670,1854]" pageId="33" pageNumber="34">
*1995
<taxonomicName id="4C22D7886448FF90FF50EBD6D1CBF93A" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[220,506,1734,1758]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="brevissimus" status="sp. nov.">
<emphasis id="B95670196448FF90FF50EBD6D1CBF93A" box="[220,506,1734,1758]" italics="true" pageId="33" pageNumber="34">Neospathodus brevissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626448FF90FD8DEBD7D20DF93A" box="[513,572,1735,1758]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
, Orchard, pp. 117, pl. 3, figs. 1415, 2022.
</paragraph>
<paragraph id="8B9DAC0B6448FF90FF1BEA16D13FF8DA" blockId="33.[151,775,1670,1854]" pageId="33" pageNumber="34">
2011
<taxonomicName id="4C22D7886448FF90FF5FEA16D269F8FA" authority="Orchard" authorityName="Orchard" box="[211,600,1798,1822]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196448FF90FF5FEA16D1C3F8FA" box="[211,498,1798,1822]" italics="true" pageId="33" pageNumber="34">Neospathodus brevissimus</emphasis>
Orchard
</taxonomicName>
; Ji et al., p. 219, fig. 3, nr. 7.
</paragraph>
<paragraph id="8B9DAC0B6448FF90FCA1EB96D3F8F8FA" blockId="33.[813,1437,1670,1886]" pageId="33" pageNumber="34">
2014
<taxonomicName id="4C22D7886448FF90FCE4EB96D4CBF97A" authority="(Bender)" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[872,1274,1670,1694]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B95670196448FF90FCE4EB96D4A2F97A" box="[872,1171,1670,1694]" italics="true" pageId="33" pageNumber="34">Novispathodus triangularis</emphasis>
(Bender)
</taxonomicName>
; Maekawa and Igo in Shigeta et al., p. 241243, figs. 172.19, 172.1327, 173.13, 173.612, 173.1621, 173.2844, 174.1174.24. 2015
<taxonomicName id="4C22D7886448FF90FCE5EBF6D515F91A" authority="(Orchard)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[873,1316,1766,1790]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B95670196448FF90FCE5EBF6D49CF91A" box="[873,1197,1766,1790]" italics="true" pageId="33" pageNumber="34">Triassospathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
; Yan et al., p. 240, fig. 3.4.
</paragraph>
<paragraph id="8B9DAC0B6448FF90FCA1EA36D3CAF8BA" blockId="33.[813,1437,1670,1886]" pageId="33" pageNumber="34">
2016
<taxonomicName id="4C22D7886448FF90FCF9EA36D513F8DA" authority="(Bender)" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[885,1314,1830,1854]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B95670196448FF90FCF9EA36D49CF8DA" box="[885,1197,1830,1854]" italics="true" pageId="33" pageNumber="34">Novispathodus triangularis</emphasis>
(Bender)
</taxonomicName>
; Komatsu et al., p. 69, fig. 5.8.
</paragraph>
<caption id="DF5DFC83644BFF93FF26EED8D157FBC8" ID-DOI="http://doi.org/10.5281/zenodo.12000396" ID-Zenodo-Dep="12000396" httpUri="https://zenodo.org/record/12000396/files/figure.png" pageId="34" pageNumber="35" startId="34.[170,201,968,988]" targetBox="[153,1438,238,1081]" targetPageId="34" targetType="figure">
<paragraph id="8B9DAC0B644BFF93FF26EED8D55FFC38" blockId="34.[169,1418,966,1068]" box="[170,1390,966,991]" pageId="34" pageNumber="35">
<emphasis id="B9567019644BFF93FF26EED8D0D8FC38" bold="true" box="[170,233,968,988]" pageId="34" pageNumber="35">Fig. 22</emphasis>
<taxonomicName id="4C22D788644BFF93FF7BEED8D1B9FC38" authorityName="Orchard" authorityYear="2005" box="[247,392,968,988]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E644BFF93FA88EEC1D53AFC3B" attach="left" box="[1284,1291,977,991]" fontSize="6" pageId="34" pageNumber="35">1</subScript>
elements if
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF25EEF2D157FBC8" blockId="34.[169,1418,966,1068]" pageId="34" pageNumber="35">
not specifically identified otherwise.
<emphasis id="B9567019644BFF93FE53EEF2D1DCFC12" bold="true" box="[479,493,994,1014]" pageId="34" pageNumber="35">A</emphasis>
,
<emphasis id="B9567019644BFF93FE79EEF2D235FC12" bold="true" box="[501,516,994,1014]" pageId="34" pageNumber="35">D</emphasis>
<taxonomicName id="4C22D788644BFF93FD84EEF2D374FC12" authority="? (Orchard)" baseAuthorityName="Orchard" box="[520,837,994,1014]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="crassatus">
<emphasis id="B9567019644BFF93FD84EEF2D2B1FC12" box="[520,640,994,1014]" italics="true" pageId="34" pageNumber="35">Icriospathodus</emphasis>
cf.
<emphasis id="B9567019644BFF93FD10EEF2D2D6FC12" box="[668,743,994,1014]" italics="true" pageId="34" pageNumber="35">crassatus</emphasis>
? (Orchard)
</taxonomicName>
;
<emphasis id="B9567019644BFF93FCC0EEF2D36BFC12" bold="true" box="[844,858,994,1014]" pageId="34" pageNumber="35">A</emphasis>
BAN1, PIMUZ 39168;
<emphasis id="B9567019644BFF93FB9CEEF2D42EFC12" bold="true" box="[1040,1055,994,1014]" pageId="34" pageNumber="35">D</emphasis>
LIL513A,PIMUZ 39169.
<emphasis id="B9567019644BFF93FB64EEF2D4C5FC12" bold="true" box="[1256,1268,994,1014]" pageId="34" pageNumber="35">B</emphasis>
,
<emphasis id="B9567019644BFF93FB71EEF2D539FC12" bold="true" box="[1277,1288,994,1014]" pageId="34" pageNumber="35">E</emphasis>
<taxonomicName id="4C22D788644BFF93FA81EEF2D16CFBF5" authority="(Orchard)" baseAuthorityName="Orchard" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FA81EEF2D137FBF5" italics="true" pageId="34" pageNumber="35">Novispathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B9567019644BFF93FEE8EEEDD141FBF5" bold="true" box="[356,368,1021,1041]" pageId="34" pageNumber="35">B</emphasis>
LIL513B,PIMUZ 39194;
<emphasis id="B9567019644BFF93FDB4EEEDD272FBF5" bold="true" box="[568,579,1021,1041]" pageId="34" pageNumber="35">E</emphasis>
LIL514A,PIMUZ SQL55056.
<emphasis id="B9567019644BFF93FCA3EEEDD30DFBF5" bold="true" box="[815,828,1021,1041]" pageId="34" pageNumber="35">C</emphasis>
,
<emphasis id="B9567019644BFF93FCC8EEEDD37EFBF5" bold="true" box="[836,847,1021,1041]" pageId="34" pageNumber="35">F</emphasis>
<taxonomicName id="4C22D788644BFF93FCDFEEEDD3FFFBF5" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[851,974,1021,1041]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B9567019644BFF93FCDFEEEDD3FFFBF5" box="[851,974,1021,1041]" italics="true" pageId="34" pageNumber="35">Novispathodus</emphasis>
</taxonomicName>
cf,
<taxonomicName id="4C22D788644BFF93FC66EEEDD4ACFBF5" authority="(Orchard)" baseAuthorityName="Orchard" box="[1002,1181,1021,1041]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FC66EEEDD476FBF5" box="[1002,1095,1021,1041]" italics="true" pageId="34" pageNumber="35">brevissimus</emphasis>
(Orchard)
</taxonomicName>
;
<emphasis id="B9567019644BFF93FB28EEEDD480FBF5" bold="true" box="[1188,1201,1021,1041]" pageId="34" pageNumber="35">C</emphasis>
LIL513A, PIMUZ 39196;
<emphasis id="B9567019644BFF93FAF7EEEDD5B7FBF5" bold="true" box="[1403,1414,1021,1041]" pageId="34" pageNumber="35">F</emphasis>
, LIL514A, PIMUZ 39198
</paragraph>
</caption>
<paragraph id="8B9DAC0B644BFF93FF1BE97FD26FFB43" blockId="34.[151,775,1135,1447]" pageId="34" pageNumber="35">
2018
<taxonomicName id="4C22D788644BFF93FF58E97FD244FB63" authority="(Orchard)" baseAuthorityName="Orchard" box="[212,629,1135,1159]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FF58E97FD1CCFB63" box="[212,509,1135,1159]" italics="true" pageId="34" pageNumber="35">Novispathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
; Maekawa in Maekawa et al., p. 34, figs. 19.4, 19.5, 19.7.
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BE9BFD130FB03" blockId="34.[151,775,1135,1447]" pageId="34" pageNumber="35">
2019
<taxonomicName id="4C22D788644BFF93FF59E9BFD208FB23" authority="Orchard" authorityName="Orchard" box="[213,569,1199,1223]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="curtatus">
<emphasis id="B9567019644BFF93FF59E9BFD1E3FB23" box="[213,466,1199,1223]" italics="true" pageId="34" pageNumber="35">Neospathodus curtatus</emphasis>
Orchard
</taxonomicName>
; Chen et al., fig. 6 nr. 13, 14.
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BE9FFD124FAC3" blockId="34.[151,775,1135,1447]" pageId="34" pageNumber="35">
2019
<taxonomicName id="4C22D788644BFF93FF56E9FFD25AFAE3" authority="Orchard" authorityName="Orchard" box="[218,619,1263,1287]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FF56E9FFD1CEFAE3" box="[218,511,1263,1287]" italics="true" pageId="34" pageNumber="35">Neospathodus brevissimus</emphasis>
Orchard
</taxonomicName>
; Chen et al., fig. 6 nr. 15.
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BE83FD1B8FA83" blockId="34.[151,775,1135,1447]" pageId="34" pageNumber="35">
2019a
<taxonomicName id="4C22D788644BFF93FF6FE83FD279FAA3" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[227,584,1327,1351]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimimus">
<emphasis id="B9567019644BFF93FF6FE83FD279FAA3" box="[227,584,1327,1351]" italics="true" pageId="34" pageNumber="35">Triassospathodus brevissimimus</emphasis>
</taxonomicName>
[sic.] (Orchard); Wu et al., fig. 4, nr. 36.
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BE87FD15BFA43" blockId="34.[151,775,1135,1447]" pageId="34" pageNumber="35">
2019
<taxonomicName id="4C22D788644BFF93FF58E87FD244FA63" authority="(Orchard)" baseAuthorityName="Orchard" box="[212,629,1391,1415]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FF58E87FD1CCFA63" box="[212,509,1391,1415]" italics="true" pageId="34" pageNumber="35">Novispathodus brevissimus</emphasis>
(Orchard)
</taxonomicName>
; Liu et al., p. 14, pl. 4, figs. 15, 18
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BE8DFD18AFA03" blockId="34.[151,443,1487,1511]" box="[151,443,1487,1511]" pageId="34" pageNumber="35">
<emphasis id="B9567019644BFF93FF1BE8DFD14CFA03" box="[151,381,1487,1511]" italics="true" pageId="34" pageNumber="35">Number of specimens</emphasis>
.&gt; 50.
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BEB00D1B7F92C" blockId="34.[151,775,1552,1928]" pageId="34" pageNumber="35">
<emphasis id="B9567019644BFF93FF1BEB00D15BF9CC" box="[151,362,1552,1576]" italics="true" pageId="34" pageNumber="35">Original diagnosis</emphasis>
. “Species characterized by small, short, and high segminate elements with a length:height ratio of 1:1, about 812 largely fused, generally upright denticles atop a deep blade, and a large basal cavity with a subcircular basal outline that occupies most of the lower side” (
<bibRefCitation id="EFB3D1FA644BFF93FF56EBA0D146F92C" author="Orchard, M. J." box="[218,375,1712,1736]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8B9DAC0B644BFF93FF1BEBC0D472FB43" blockId="34.[151,775,1552,1928]" lastBlockId="34.[813,1437,1135,1927]" pageId="34" pageNumber="35">
<emphasis id="B9567019644BFF93FF1BEBC0D0CDF90C" box="[151,252,1744,1768]" italics="true" pageId="34" pageNumber="35">Remarks.</emphasis>
Characteristic of this species is the posterior lateral margin which is slightly to strongly curved. Subcircular to subtriangular outline of the large basal cavity in the lower view. Te denticles are fused and compressed with an erect to radial outline (bouquet-like). Te carina is relatively high. In lateral view, the upper edge is usually straight to slightly arched. Cusp is usually undistinguishable from other denticles.
</paragraph>
<paragraph id="8B9DAC0B644BFF92FCA1E9A0D254FEC7" blockId="34.[813,1437,1135,1927]" lastBlockId="35.[151,775,235,515]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Te here illustrated adult specimens are a bit larger with more denticles than the
<typeStatus id="549912A9644BFF93FBC2E9DFD49FFB03" box="[1102,1198,1231,1255]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFB3D1FA644BFF93FB4FE9DFD556FB03" author="Orchard, M. J." box="[1219,1383,1231,1255]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
; pp. 117, Pl. 3, fig. 22), but the denticulation and the convex posterior edge of the process is very distinct in this species. Terefore, the
<typeStatus id="549912A9644BFF93FBBBE83FD4A6FAA3" box="[1079,1175,1327,1351]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFB3D1FA644BFF93FB26E83FD576FAA3" author="Orchard, M. J." box="[1194,1351,1327,1351]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard 1995</bibRefCitation>
) might represent a younger, juvenile morphotype of this taxon.
<taxonomicName id="4C22D788644BFF93FCA1E87FD441FA63" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[813,1136,1391,1415]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="eotriangularis">
<emphasis id="B9567019644BFF93FCA1E87FD441FA63" box="[813,1136,1391,1415]" italics="true" pageId="34" pageNumber="35">Novispathodus eotriangularis</emphasis>
</taxonomicName>
(Zhao &amp; Orchard in
<bibRefCitation id="EFB3D1FA644BFF93FAEFE87FD394FA43" author="Zhao, L. &amp; Orchard, M. J. &amp; Tong, J. &amp; Sun, Z. &amp; Zuo, J. &amp; Zhang, S. &amp; Yun, A." pageId="34" pageNumber="35" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al., 2007</bibRefCitation>
) has a more triangle-shaped basal cavity and the anterior half of the carina is gradually declining in height, whereas
<taxonomicName id="4C22D788644BFF93FC7AE8DFD519FA03" box="[1014,1320,1487,1511]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FC7AE8DFD519FA03" box="[1014,1320,1487,1511]" italics="true" pageId="34" pageNumber="35">Novispathodus brevissimus</emphasis>
</taxonomicName>
shows a more abrupt decline at the posterior end. Compared to
<taxonomicName id="4C22D788644BFF93FCDFEB1FD52EF9C3" authority="(Orchard 1995)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[851,1311,1551,1575]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="clinatus">
<emphasis id="B9567019644BFF93FCDFEB1FD46DF9C3" box="[851,1116,1551,1575]" italics="true" pageId="34" pageNumber="35">Novispathodus clinatus</emphasis>
(
<bibRefCitation id="EFB3D1FA644BFF93FBF8EB1FD527F9C3" author="Orchard, M. J." box="[1140,1302,1551,1575]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard 1995</bibRefCitation>
)
</taxonomicName>
, this species shows more numerous, short and upright denticles.
<taxonomicName id="4C22D788644BFF93FCA1EB5FD464F983" box="[813,1109,1615,1639]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644BFF93FCA1EB5FD464F983" box="[813,1109,1615,1639]" italics="true" pageId="34" pageNumber="35">Novispathodus brevissimus</emphasis>
</taxonomicName>
shares most similarities with
<taxonomicName id="4C22D788644BFF93FCA1EB7FD4C5F963" authority="(Orchard, 1995)" baseAuthorityName="Orchard" baseAuthorityYear="1995" box="[813,1268,1647,1671]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="curtatus">
<emphasis id="B9567019644BFF93FCA1EB7FD404F963" box="[813,1077,1647,1671]" italics="true" pageId="34" pageNumber="35">Novispathodus curtatus</emphasis>
(
<bibRefCitation id="EFB3D1FA644BFF93FBC4EB7FD4DBF963" author="Orchard, M. J." box="[1096,1258,1647,1671]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
)
</taxonomicName>
, but the latter has a relatively smaller basal cavity, is longer and bears larger and more reclined denticles. Some specimens (e.g.
<figureCitation id="1319B08E644BFF93FCA1EBDFD3B9F903" box="[813,904,1743,1767]" captionStart="Fig" captionStartId="30.[169,200,1700,1720]" captionTargetBox="[153,1439,238,1918]" captionTargetId="figure-16@30.[229,1363,254,1687]" captionTargetPageId="30" captionText="Fig. 19 Novispathodinae from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A Novispathodus cf.brevissimus (Orchard); LIL515C,PIMUZ 39199. B, F Novispathodus brevissimus (Orchard); B LIL515B, PIMUZ 39200; F BAN1, PIMUZ 39201. C Triassospathodus homeri (Bender); LIL515D,PIMUZ 39317. D, H Novispathodus?brevissimus (Orchard); D LIL515D,PIMUZ 39202; H LIL514A, PIMUZ 39203. E Triassospathodus cf.homeri (Bender); LIL515C, PIMUZ 39318. G, K, OQ Novispathodus n. sp. A; G LIL504,PIMUZ 39251; K LIL504,PIMUZ 39252; O LIL505, PIMUZ 39253; P LIL505, PIMUZ 39254;Q LIL505,PIMUZ 39255. I, L, M, Triassospathodus symmetricus (Orchard); I QIA141, PIMUZ 39336; L LIL512, PIMUZ 39337; M LIL514A, PIMUZ 39338. J Novispathodus ex gr.pingdingshanensis (Zhao &amp; Orchard); QIA138, PIMUZ 39237. N Triassospathodus cf. symmetricus; LIL514B, PIMUZ 39339.R Novispathodus praebrevissimus n.sp; LIL505, PIMUZ 39289" figureDoi="http://doi.org/10.5281/zenodo.12000347" httpUri="https://zenodo.org/record/12000347/files/figure.png" pageId="34" pageNumber="35">Fig. 19F</figureCitation>
) bear the postero-lateral pinching that is usually considered diagnostic of
<taxonomicName id="4C22D788644BFF93FBF7EBE0D5A9F8E3" authority="(Bender)" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[1147,1432,1775,1799]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="triangularis">
<emphasis id="B9567019644BFF93FBF7EBE0D51DF8E3" box="[1147,1324,1775,1799]" italics="true" pageId="34" pageNumber="35">Nv. triangularis</emphasis>
(Bender)
</taxonomicName>
, as revised by
<bibRefCitation id="EFB3D1FA644BFF93FC41EA1FD44AF8C3" author="Orchard, M. J." box="[973,1147,1807,1831]" pageId="34" pageNumber="35" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard (1995)</bibRefCitation>
.
<bibRefCitation id="EFB3D1FA644BFF93FB07EA1FD56AF8C3" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[1163,1371,1807,1831]" pageId="34" pageNumber="35" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
illustrated very similar specimens from a higher horizon (their figs. 7.9, 7.10). Contrary to true
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<emphasis id="B9567019644BFF93FB64EA40D5A6F883" box="[1256,1431,1871,1895]" italics="true" pageId="34" pageNumber="35">Nv. triangularis</emphasis>
</taxonomicName>
, our specimens have more rounded denticles tips and the pinching is less conspicuous, suggesting they can still be retained within
<taxonomicName id="4C22D788644AFF92FEC8EC1CD1DDFEC7" box="[324,492,267,291]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="brevissimus">
<emphasis id="B9567019644AFF92FEC8EC1CD1DDFEC7" box="[324,492,267,291]" italics="true" pageId="35" pageNumber="36">Nv. brevissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62644AFF92FE7EEC1CD251FEC7" box="[498,608,267,291]" pageId="35" pageNumber="36" sensu="lato">sensu lato</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8B9DAC0B644AFF92FF21EC3CD26EFDE7" blockId="35.[151,775,235,515]" pageId="35" pageNumber="36">
<emphasis id="B9567019644AFF92FF21EC3CD11FFEA7" box="[173,302,300,323]" italics="true" pageId="35" pageNumber="36">Occurrence.</emphasis>
<collectingCountry id="F335EC9B644AFF92FECBEC3BD1BFFEA7" box="[327,398,299,323]" name="Oman" pageId="35" pageNumber="36">Oman</collectingCountry>
, Jabal Safra, lower Spathian (
<bibRefCitation id="EFB3D1FA644AFF92FF13EC5BD175FE87" author="Orchard, M. J." box="[159,324,331,355]" pageId="35" pageNumber="36" pagination="110 - 122" refId="ref45513" refString="Orchard, M. J. (1995). Taxonomy and correlation of Lower Triassic (Spathian) segminate conodonts from Oman and revision of some species of Neospathodus. Journal of Paleontology, 69 (1), 110 - 122. https: // doi. org / 10.1017 / S 0022336000026962" type="journal article" year="1995">Orchard, 1995</bibRefCitation>
);
<collectingCountry id="F335EC9B644AFF92FEEEEC5CD191FE87" box="[354,416,332,355]" name="Japan" pageId="35" pageNumber="36">Japan</collectingCountry>
, Taho Limestone, occurs with
<taxonomicName id="4C22D788644AFF92FF1BEC7CD25CFE67" authority=", Tahogawa Member" authorityName="Tahogawa Member" box="[151,621,363,387]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B9567019644AFF92FF1BEC7CD087FE67" box="[151,182,364,387]" italics="true" pageId="35" pageNumber="36">Nv</emphasis>
.
<emphasis id="B9567019644AFF92FF49EC7BD1B8FE67" box="[197,393,363,387]" italics="true" pageId="35" pageNumber="36">pingdingshanensis</emphasis>
, Tahogawa Member
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA644AFF92FDF1EC7BD331FE67" author="Koike, T." box="[637,768,363,387]" pageId="35" pageNumber="36" pagination="53 - 69" refId="ref42668" refString="Koike, T. (1981). Biostratigraphy of Triassic conodonts in Japan. Science Reports of the Yokohama National University, Section 2, (38), 53 - 69." type="journal article" year="1981">Koike, 1981</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644AFF92FF1BEC9BD1B2FE47" author="Maekawa, T. &amp; Komatsu, T. &amp; Koike, T." box="[151,387,395,419]" pageId="35" pageNumber="36" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
); Nanpanjiang basin, South
<collectingCountry id="F335EC9B644AFF92FD49EC9BD336FE47" box="[709,775,395,419]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
and North
<collectingCountry id="F335EC9B644AFF92FE99ECBBD145FE27" box="[277,372,427,451]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
, together with
<taxonomicName id="4C22D788644AFF92FD93ECBBD249FE27" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[543,632,427,451]" class="Cephalopoda" family="Columbitidae" genus="Tirolites" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B9567019644AFF92FD93ECBBD249FE27" box="[543,632,427,451]" italics="true" pageId="35" pageNumber="36">Tirolites</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62644AFF92FD0DECBCD2E7FE27" box="[641,726,428,451]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C22D788644AFF92FF1BECDBD194FE07" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[151,421,459,483]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019644AFF92FF1BECDBD194FE07" box="[151,421,459,483]" italics="true" pageId="35" pageNumber="36">Icriospathodus collinsoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA644AFF92FE3FECDBD2A5FE07" author="Komatsu, T. &amp; Takashima, R. &amp; Shigeta, Y. &amp; Maekawa, T. &amp; Tran, H. D. &amp; Cong, T. D. &amp; Sakata, S. &amp; Dinh, H. D. &amp; Takahashi, O." box="[435,660,459,483]" pageId="35" pageNumber="36" pagination="65 - 74" refId="ref43220" refString="Komatsu, T., Takashima, R., Shigeta, Y., Maekawa, T., Tran, H. D., Cong, T. D., Sakata, S., Dinh, H. D., &amp; Takahashi, O. (2016). Carbon isotopic excursions and detailed ammonoid and conodont biostratigraphies around Smithian-Spathian boundary in the Bac Thuy Formation, Vietnam. Palaeogeography, Palaeoclimatology, Palaeoecology, 454, 65 - 74. https: // doi. org / 10.1016 / j. palaeo. 2016.04. 017" type="journal article" year="2016">Komatsu et al., 2016</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644AFF92FD2DECDBD0FAFDE7" author="Liu, S. &amp; Sun, Z. &amp; Ji, C. &amp; Zhou, M. &amp; Jiang, D." pageId="35" pageNumber="36" refId="ref44210" refString="Liu, S., Sun, Z., Ji, C., Zhou, M., &amp; Jiang, D. (2019). Conodont biostratigraphy and age of the early Triassic fish-bearing-nodule levels from Nanjing and Jurong, Jiangsu Province, South China. Journal of Earth Science. https: // doi. org / 10.1007 / s 12583 - 019 - 1232 - y" type="book" year="2019">Liu et al., 2019</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644AFF92FF54ECFBD193FDE7" author="Shigeta, Y. &amp; Komatsu, T. &amp; Maekawa, T. &amp; Huyen, D. T." box="[216,418,491,515]" pageId="35" pageNumber="36" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA644AFF92FE22ECFCD260FDE7" author="Yan, C. &amp; Jiang, H. &amp; Lai, X. &amp; Sun, Y. &amp; Yang, B. &amp; Wang, L." box="[430,593,491,515]" pageId="35" pageNumber="36" refId="ref48390" refString="Yan, C., Jiang, H., Lai, X., Sun, Y., Yang, B., &amp; Wang, L. (2015). The relationship between the &quot; Green-Bean Rock &quot; layers and conodont Chiosella timorensis and implications on defining the Early-Middle Triassic boundary" type="book" year="2015">Yan et al., 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,229 @@
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Z Orchard, 2007
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<figureCitation id="1319B08E6476FFAEFF1BEB3BD0EEF9A7" box="[151,223,1579,1603]" captionStart="Fig" captionStartId="31.[169,200,1715,1735]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-16@32.[229,1363,254,1763]" captionTargetPageId="32" captionText="Fig. 20 Novispathodinae from Qiakong,Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AG, KM, Q, X Novispathodus ex gr. abruptus (Orchard); A LAR212, PIMUZ 39213; B LAR207, PIMUZ 39214; C LAR205,PIMUZ 39215; D LAR202, PIMUZ 39216;E LAR202, PIMUZ 39217; F LIL506, PIMUZ 39218; G LAR210, PIMUZ 39219; K LIL505,PIMUZ 39220; L LIL506, PIMUZ 39221; M LIL505,PIMUZ 39222; Q LAR203,PIMUZ 39223; X QIA136,PIMUZ 39224. H, V Novispathodus sp. indet.; H QIA136, PIMUZ 39291; V, QIA134, PIMUZ 39292. I, J, P, AA Novispathodus n. sp. Z; I QIA138, PIMUZ 39247;J QIA138,PIMUZ 39248; P LIL506, PIMUZ 39249; AA QIA136, PIMUZ 39250. N, O, S Novispathodus ex gr. pingdingshanensis (Zhao &amp; Orchard); N LIL506, PIMUZ 39234; O LIL505, PIMUZ 39235; S QIA136, PIMUZ 39236. R sp.indet.; LIL511, PIMUZ 39296. T, Y, Z Novispathodus n. sp. A; T QIA134,PIMUZ 39256,Y QIA135, PIMUZ 39257; Z QIA134, PIMUZ 39258. U Novispathodus expansus (Zhao &amp; Orchard); QIA134, PIMUZ 39242. W Novispathodus gryphus n. sp.; QIA135, PIMUZ 39246" figureDoi="http://doi.org/10.5281/zenodo.12000362" httpUri="https://zenodo.org/record/12000362/files/figure.png" pageId="31" pageNumber="32">Fig. 20</figureCitation>
, I, J, P, AA
</paragraph>
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*2007
<emphasis id="B95670196476FFAEFCFDEDFCD3BCFEE7" box="[881,909,236,259]" italics="true" pageId="31" pageNumber="32">Ns</emphasis>
. n. sp. Z; Orchard, p. 96, fig. 2.
</paragraph>
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2016
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<emphasis id="B95670196476FFAEFCE5EC1BD43AFEC7" box="[873,1035,267,291]" italics="true" pageId="31" pageNumber="32">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D7886476FFAEFBD9EC1BD51EFEC7" authority="(Orchard)" baseAuthorityName="Orchard" box="[1109,1327,267,291]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFBD9EC1BD486FEC7" box="[1109,1207,267,291]" italics="true" pageId="31" pageNumber="32">abruptus</emphasis>
(Orchard)
</taxonomicName>
; Komatsu et al., p. 77, figs. 5.6ac.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1EC5BD3F8FE67" blockId="31.[813,1437,235,515]" pageId="31" pageNumber="32">
2018
<taxonomicName id="4C22D7886476FFAEFCE4EC5BD4D8FE87" authority="(Orchard)" baseAuthorityName="Orchard" box="[872,1257,331,355]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFCE4EC5BD443FE87" box="[872,1138,331,355]" italics="true" pageId="31" pageNumber="32">Novispathodus abruptus</emphasis>
(Orchard)
</taxonomicName>
; Maekawa et al., p. 33, fig. 18.8.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1EC9BD516FE27" blockId="31.[813,1437,235,515]" pageId="31" pageNumber="32">
2019
<taxonomicName id="4C22D7886476FFAEFC0DEC9BD3A5FE27" authority="(Zhao &amp; Orchard)" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFC0DEC9BD536FE47" box="[897,1287,395,419]" italics="true" pageId="31" pageNumber="32">Novispathodus pingdingshanensis</emphasis>
(Zhao &amp; Orchard)
</taxonomicName>
; Liu et al., p. 13, pl. 3, figs. 3, 6 (only).
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1ECDBD3F9FDE7" blockId="31.[813,1437,235,515]" pageId="31" pageNumber="32">
2019
<taxonomicName id="4C22D7886476FFAEFCFCECDBD57FFE07" authority="? (Orchard)" baseAuthorityName="Orchard" box="[880,1358,459,483]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="symmetricus">
<emphasis id="B95670196476FFAEFCFCECDBD4F4FE07" box="[880,1221,459,483]" italics="true" pageId="31" pageNumber="32">Triassospathodus symmetricus</emphasis>
? (Orchard)
</taxonomicName>
; Chen et al., fig. 4.15.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1EF3BD469FDA7" blockId="31.[813,1437,555,1475]" box="[813,1112,555,579]" pageId="31" pageNumber="32">
<emphasis id="B95670196476FFAEFCA1EF3BD3A5FDA7" box="[813,916,555,579]" italics="true" pageId="31" pageNumber="32">Material.</emphasis>
ca.
<specimenCount id="9D2467826476FFAEFC32EF3BD464FDA7" box="[958,1109,555,579]" count="15" pageId="31" pageNumber="32" type="generic">15 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1EF5BD4ECFB27" blockId="31.[813,1437,555,1475]" pageId="31" pageNumber="32">
<emphasis id="B95670196476FFAEFCA1EF5BD3BAFD87" box="[813,907,587,611]" italics="true" pageId="31" pageNumber="32">Remarks</emphasis>
. One small terminal denticle behind the cusp is present. Te denticles are mostly fused, with subtriangular tips and increasingly reclined towards the posterior end. A moderately deep, conical, subcircular shaped basal cavity is present. Te overall shape of the segminateP
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element is reminiscent of that of the sub-coeval
<taxonomicName id="4C22D7886476FFAEFCA1EE1CD42CFCC7" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[813,1053,779,803]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFCA1EE1CD42CFCC7" box="[813,1053,779,803]" italics="true" pageId="31" pageNumber="32">Nv. pingdingshanensis</emphasis>
</taxonomicName>
or
<taxonomicName id="4C22D7886476FFAEFBC4EE1CD4E7FCC7" box="[1096,1238,779,803]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFBC4EE1CD4E7FCC7" box="[1096,1238,779,803]" italics="true" pageId="31" pageNumber="32">Nv. abruptus</emphasis>
</taxonomicName>
except that, contrary to
<taxonomicName id="4C22D7886476FFAEFC05EE3CD449FCA7" baseAuthorityName="Zhao &amp; Orchard" baseAuthorityYear="2007" box="[905,1144,811,835]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFC05EE3CD449FCA7" box="[905,1144,811,835]" italics="true" pageId="31" pageNumber="32">Nv. pingdingshanensis</emphasis>
</taxonomicName>
, the denticles are reclined not recurved, and contrary to
<taxonomicName id="4C22D7886476FFAEFBF6EE5CD536FC87" box="[1146,1287,843,867]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="abruptus">
<emphasis id="B95670196476FFAEFBF6EE5CD536FC87" box="[1146,1287,843,867]" italics="true" pageId="31" pageNumber="32">Nv. abruptus</emphasis>
</taxonomicName>
, there is only one small denticle behind the cusp, the latter being also conspicuously broader than adjacent denticles. Our specimens from
<collectingCountry id="F335EC9B6476FFAEFC3FEEBBD3C4FC27" box="[947,1013,939,963]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
resemble the specimens from Panthalassa California (Darwin material), suggesting that this species may be important for worldwide correlations. Orchard (2007) mentioned, that
<taxonomicName id="4C22D7886476FFAEFB1BE91BD3EBFBA7" authority="(Koike 1973)" baseAuthorityName="Koike" baseAuthorityYear="1973" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="kedahensis">
<emphasis id="B95670196476FFAEFB1BE91BD376FBA7" italics="true" pageId="31" pageNumber="32">Neospathodus kedahensis</emphasis>
(
<bibRefCitation id="EFB3D1FA6476FFAEFCD9E93BD3E1FBA7" author="Koike, T." box="[853,976,1067,1091]" pageId="31" pageNumber="32" pagination="91 - 113" refId="ref42638" refString="Koike, T. (1973). Triassic conodonts from Kedah and Pahang, Malaysia. Geology and Paleontology of Southeast Asia, 12, 91 - 113." type="journal article" year="1973">Koike 1973</bibRefCitation>
)
</taxonomicName>
might be an available name for his
<emphasis id="B95670196476FFAEFAD2E93CD54BFBA7" box="[1374,1402,1068,1091]" italics="true" pageId="31" pageNumber="32">Ns</emphasis>
. n. sp. Z (Orchard 2007). However,
<taxonomicName id="4C22D7886476FFAEFB1AE95BD3EFFB67" authority="(Koike 1973)" baseAuthorityName="Koike" baseAuthorityYear="1973" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="kedahensis">
<emphasis id="B95670196476FFAEFB1AE95BD376FB67" italics="true" pageId="31" pageNumber="32">Neospathodus kedahensis</emphasis>
(
<bibRefCitation id="EFB3D1FA6476FFAEFCDBE97BD3E5FB67" author="Koike, T." box="[855,980,1131,1155]" pageId="31" pageNumber="32" pagination="91 - 113" refId="ref42638" refString="Koike, T. (1973). Triassic conodonts from Kedah and Pahang, Malaysia. Geology and Paleontology of Southeast Asia, 12, 91 - 113." type="journal article" year="1973">Koike 1973</bibRefCitation>
)
</taxonomicName>
a Middle-to-Late Triassic species, lacks a conspicuous cusp and its posterior part is composed of slightly inclined and subequal denticles.
</paragraph>
<paragraph id="8B9DAC0B6476FFAEFCA1E9DCD5A9FA27" blockId="31.[813,1437,555,1475]" pageId="31" pageNumber="32">
<emphasis id="B95670196476FFAEFCA1E9DCD399FB07" box="[813,936,1228,1251]" italics="true" pageId="31" pageNumber="32">Occurrence</emphasis>
. Taho Formation,
<collectingCountry id="F335EC9B6476FFAEFB01E9DCD4F8FB07" box="[1165,1225,1228,1251]" name="Japan" pageId="31" pageNumber="32">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6476FFAEFB6CE9DBD350FAE7" author="Maekawa, T. &amp; Komatsu, T. &amp; Koike, T." pageId="31" pageNumber="32" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
). North America (Orchard 2007, Goudemand et al., in prep.).
<collectingCountry id="F335EC9B6476FFAEFC24E81BD3C1FAC7" box="[936,1008,1291,1315]" name="Oman" pageId="31" pageNumber="32">Oman</collectingCountry>
: Radio Tower section, UAZ
<subScript id="17A6AE4E6476FFAEFACCE807D578FACC" attach="left" box="[1344,1353,1303,1320]" fontSize="7" pageId="31" pageNumber="32">5</subScript>
(
<bibRefCitation id="EFB3D1FA6476FFAEFAECE81BD397FAA7" author="Chen, Y. &amp; Richoz, S. &amp; Krystyn, L. &amp; Zhang, Z." pageId="31" pageNumber="32" refId="ref39612" refString="Chen, Y., Richoz, S., Krystyn, L., &amp; Zhang, Z. (2019). Quantitative stratigraphic correlation of Tethyan conodonts across the Smithian-Spathian (Early Triassic) extinction event. Earth-Science Reviews. https: // doi. org / 10. 1016 / j. earscirev. 2019.03. 004" type="book" year="2019">Chen et al., 2019</bibRefCitation>
), South
<collectingCountry id="F335EC9B6476FFAEFB8BE83BD47AFAA7" box="[1031,1099,1323,1347]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
; Nanpanjiang basin: Bac Tuy Formation,
<taxonomicName id="4C22D7886476FFAEFC23E85CD4D1FA87" authority="zone" authorityName="Zone" box="[943,1248,1355,1379]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFC23E85CD3FFFA87" box="[943,974,1356,1379]" italics="true" pageId="31" pageNumber="32">Nv</emphasis>
.
<emphasis id="B95670196476FFAEFC52E85BD493FA87" box="[990,1186,1355,1379]" italics="true" pageId="31" pageNumber="32">pingdingshanensis</emphasis>
zone
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA6476FFAEFB7DE85BD350FA67" author="Komatsu, T. &amp; Takashima, R. &amp; Shigeta, Y. &amp; Maekawa, T. &amp; Tran, H. D. &amp; Cong, T. D. &amp; Sakata, S. &amp; Dinh, H. D. &amp; Takahashi, O." pageId="31" pageNumber="32" pagination="65 - 74" refId="ref43220" refString="Komatsu, T., Takashima, R., Shigeta, Y., Maekawa, T., Tran, H. D., Cong, T. D., Sakata, S., Dinh, H. D., &amp; Takahashi, O. (2016). Carbon isotopic excursions and detailed ammonoid and conodont biostratigraphies around Smithian-Spathian boundary in the Bac Thuy Formation, Vietnam. Palaeogeography, Palaeoclimatology, Palaeoecology, 454, 65 - 74. https: // doi. org / 10.1016 / j. palaeo. 2016.04. 017" type="journal article" year="2016">Komatsu et al., 2016</bibRefCitation>
); Qinglong Formation,
<taxonomicName id="4C22D7886476FFAEFBEAE87CD5A9FA67" authority="Zone" authorityName="Zone" box="[1126,1432,1387,1411]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B95670196476FFAEFBEAE87CD564FA67" box="[1126,1365,1387,1411]" italics="true" pageId="31" pageNumber="32">Nv. pingdingshanensis</emphasis>
Zone
</taxonomicName>
,
<collectingRegion id="49E662E96476FFAEFCA1E89BD3D8FA47" box="[813,1001,1419,1443]" country="China" name="Jiangsu" pageId="31" pageNumber="32">Jiangsu Province</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6476FFAEFC71E89BD482FA47" author="Liu, S. &amp; Sun, Z. &amp; Ji, C. &amp; Zhou, M. &amp; Jiang, D." box="[1021,1203,1419,1443]" pageId="31" pageNumber="32" refId="ref44210" refString="Liu, S., Sun, Z., Ji, C., Zhou, M., &amp; Jiang, D. (2019). Conodont biostratigraphy and age of the early Triassic fish-bearing-nodule levels from Nanjing and Jurong, Jiangsu Province, South China. Journal of Earth Science. https: // doi. org / 10.1007 / s 12583 - 019 - 1232 - y" type="book" year="2019">Liu et al., 2019</bibRefCitation>
) Luolou Formation, Qiakong section, Southern
<collectingRegion id="49E662E96476FFAEFBDAE8BBD4F4FA27" box="[1110,1221,1451,1475]" country="China" name="Guizhou" pageId="31" pageNumber="32">Gouizhou</collectingRegion>
,
<collectingCountry id="F335EC9B6476FFAEFB43E8BBD520FA27" box="[1231,1297,1451,1475]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
(this study).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,163 @@
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Genus
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GLADIGONDOLELLA
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1968
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<emphasis id="B95670196478FFA0FF5EEA24D179F8A8" box="[210,328,1844,1868]" italics="true" pageId="17" pageNumber="18">Dichodella</emphasis>
Mosher
</taxonomicName>
, p.
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923
<emphasis id="B95670196478FFA0FCA1E825D47FFAA8" box="[813,1102,1332,1356]" italics="true" pageId="17" pageNumber="18">
Type species.
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Polygnathus
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<emphasis id="B95670196478FFA0FBD7E824D481FAA8" box="[1115,1200,1332,1356]" italics="true" pageId="17" pageNumber="18">tethydis</emphasis>
<bibRefCitation id="EFB3D1FA6478FFA0FB30E824D546FAA8" author="Huckriede, R." box="[1212,1399,1332,1356]" pageId="17" pageNumber="18" pagination="141 - 175" refId="ref42031" refString="Huckriede, R. (1958). Die Conodonten der Mediterranen Trias und ihr stratigraphischer Wert. Palaontologische Zeitschrift, 32, 141 - 175." type="journal article" year="1958">Huckriede, 1958</bibRefCitation>
, p. 157
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158, pl. 2, fig. 38ab.
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<materialsCitation id="3B4AA6566478FFA0FCA1E865D528FA48" country="Austria" county="Rothelstein" location="Trachyceras" municipality="Feuerkogel" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549912A96478FFA0FCA1E865D350FA68" box="[813,865,1397,1420]" pageId="17" pageNumber="18">Type</typeStatus>
stratum and locality.
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austriacum
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bed (Julian),
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near
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,
<collectingCountry id="F335EC9B6478FFA0FB49E884D528FA48" box="[1221,1305,1428,1452]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
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.
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<emphasis id="B95670196478FFA0FCA1E8C4D3BAFA08" box="[813,907,1492,1516]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
. Tis genus was first introduced by
<bibRefCitation id="EFB3D1FA6478FFA0FADFE8C4D342F9E8" author="MUller, K. J." pageId="17" pageNumber="18" pagination="109 - 117" refId="ref45393" refString="MUller, K. J. (1962). Zur systematischen Einteilung der Conodontophorida. Palaontologische Zeitschrift, 36 (1 - 2), 109 - 117. https: // doi. org / 10.1007 / BF 02989634" type="journal article" year="1962">Müller (1962)</bibRefCitation>
for a Middle Triassic carminiplanate P
<subScript id="17A6AE4E6478FFA0FABDEB10D50BF9F5" attach="left" box="[1329,1338,1536,1553]" fontSize="7" pageId="17" pageNumber="18">1</subScript>
element with a relatively short posterior process and corresponding keel posteriorly of the pit, a narrow platform and a low carina. As the
<typeStatus id="549912A96478FFA0FB85EB45D409F989" box="[1033,1080,1621,1645]" pageId="17" pageNumber="18">type</typeStatus>
species may have been significantly different from some Spathian forms, the phylogenetic relationships of
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<emphasis id="B95670196478FFA0FBC4EB84D4C4F948" box="[1096,1269,1684,1708]" italics="true" pageId="17" pageNumber="18">Gladigondolella</emphasis>
</taxonomicName>
are uncertain but the rootstock might lie within the evolution from
<taxonomicName id="4C22D7886478FFA0FCA1EBC4D477F908" authority="(Orchard, 2007)" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[813,1094,1748,1772]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B95670196478FFA0FCA1EBC4D3A0F908" box="[813,913,1748,1772]" italics="true" pageId="17" pageNumber="18">Borinella</emphasis>
(Orchard, 2007)
</taxonomicName>
.
</paragraph>
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@ -0,0 +1,262 @@
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<emphasis id="B95670196478FFA0FCA1EA04D435F8C8" box="[813,1028,1812,1836]" italics="true" pageId="17" pageNumber="18">Gladigondolella laii</emphasis>
Chen et al.
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morphotype A
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<figureCitation id="1319B08E6478FFA0FCA1EA24D3F2F8A8" box="[813,963,1844,1868]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.12000264" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="17" pageNumber="18">Fig. 13B, C, G</figureCitation>
?
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<paragraph id="8B9DAC0B647BFFA3FF1BEDFBD2B6FEE7" blockId="18.[151,775,235,387]" box="[151,647,235,259]" pageId="18" pageNumber="19">
2013
<taxonomicName id="4C22D788647BFFA3FF5EEDFBD1A3FEE7" box="[210,402,235,259]" genus="Neogondolella" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019647BFFA3FF5EEDFBD15DFEE7" box="[210,364,235,259]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
sp.
</taxonomicName>
; Yan et al., fig. 6, nr. X.
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<paragraph id="8B9DAC0B647BFFA3FF1BEC1BD120FE67" blockId="18.[151,775,235,387]" pageId="18" pageNumber="19">
2015
<taxonomicName id="4C22D788647BFFA3FF5EEC1BD15DFEC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[210,364,267,291]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FF5EEC1BD15DFEC7" box="[210,364,267,291]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. A; Chen et al., figs. 9.5
<date id="FF9C8ACB647BFFA3FD31EC1BD0FAFEA7" pageId="18" pageNumber="19" value="2019-08-09">9.8. 2019</date>
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<emphasis id="B9567019647BFFA3FF5EEC3BD15DFEA7" box="[210,364,299,323]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. A; Chen et al., figs. 6.8,
<date id="FF9C8ACB647BFFA3FD35EC3BD0FAFE87" pageId="18" pageNumber="19" value="2021-09-06">6.9. 2021</date>
<taxonomicName id="4C22D788647BFFA3FF58EC5BD19FFE87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[212,430,331,355]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii" status="sp. nov.">
<emphasis id="B9567019647BFFA3FF58EC5BD19FFE87" box="[212,430,331,355]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FE3AEC5CD23DFE87" box="[438,524,332,355]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
; Chen et al., figs. 4.20, 4.21 (only).
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF1BECBBD1F2FE27" blockId="18.[151,451,427,451]" box="[151,451,427,451]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF1BECBBD0C9FE27" box="[151,248,427,451]" italics="true" pageId="18" pageNumber="19">Material</emphasis>
. ca.
<specimenCount id="9D246782647BFFA3FEA5ECBBD18EFE27" box="[297,447,427,451]" count="15" pageId="18" pageNumber="19" type="generic">15 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF1BECFCD274FD87" blockId="18.[151,776,491,1667]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF1BECFCD139FDE7" box="[151,264,492,515]" italics="true" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Segminiplanate P
<subScript id="17A6AE4E647BFFA3FE60ECE7D1C4FDEC" attach="left" box="[492,501,503,520]" fontSize="7" pageId="18" pageNumber="19">1</subScript>
element. Platform is arched, thickened and the platform margins are oriented orally. Denticles slightly fused at mid-length, larger and more discrete towards the anterior end.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF21EF7CD335FC67" blockId="18.[151,776,491,1667]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF21EF7CD101FD67" box="[173,304,620,643]" italics="true" pageId="18" pageNumber="19">Description.</emphasis>
Te element is arched in lateral view. In oral view, the margins of the thickened platform are subparallel at mid-length, and taper anteriorly. Te platform is V-shaped, and the margin is directed orally. Te cusp may be conspicuous in well preserved specimens. A posterior process may be present, which bears additional posterior denticles, otherwise the posterior margin is rounded. Te carina denticles are partly fused at mid-length, taller and more discrete towards the anterior end.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF21EE9BD4FCFD47" blockId="18.[151,776,491,1667]" lastBlockId="18.[813,1437,235,803]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF21EE9BD1C0FC47" box="[173,497,907,931]" italics="true" pageId="18" pageNumber="19">
Remarks.
<taxonomicName id="4C22D788647BFFA3FE94EE9BD1C0FC47" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[280,497,907,931]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">Gladigondolella laii</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C22D788647BFFA3FD0DEE9BD2D4FC47" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[641,741,907,931]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FD0DEE9BD2D4FC47" box="[641,741,907,931]" italics="true" pageId="18" pageNumber="19">Borinella</emphasis>
</taxonomicName>
by having a more fused carina and an arched and thick platform whose margin are oriented orally. In comparison with the
<typeStatus id="549912A9647BFFA3FF75EEFCD119FBE0" box="[249,296,1004,1028]" pageId="18" pageNumber="19">type</typeStatus>
species of
<taxonomicName id="4C22D788647BFFA3FE2EEEFBD27EFBE7" authorityName="Muller" authorityYear="1962" box="[418,591,1003,1027]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FE2EEEFBD27EFBE7" box="[418,591,1003,1027]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, a Middle Triassic species, the posterior process is much less developed and less arcuated in the carinal axis.
<bibRefCitation id="EFB3D1FA647BFFA3FDB0E93BD336FBA7" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[572,775,1067,1091]" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
rejected the assignment of their
<emphasis id="B9567019647BFFA3FE75E95CD226FB87" box="[505,535,1100,1123]" italics="true" pageId="18" pageNumber="19">Ng</emphasis>
. n. sp. A to
<taxonomicName id="4C22D788647BFFA3FD16E95BD0D0FB67" authorityName="Muller" authorityYear="1962" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FD16E95BD0D0FB67" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
on the basis that posterior denticles are never present behind the cusp. In our material (e.g.
<figureCitation id="1319B08E647BFFA3FD2CE99BD2CEFB47" box="[672,767,1163,1187]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.12000264" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="18" pageNumber="19">Fig. 13C</figureCitation>
) small denticles are present occasionally. With the exception of these additional posterior denticles, our specimens resemble those of
<bibRefCitation id="EFB3D1FA647BFFA3FE20E9FBD246FAE7" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[428,631,1259,1283]" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
very closely, hence they are synonymized here. Implicitly we assume there is variation within the species, a small posterior process being present or not. Tis taxon is thought to be intermediary between
<taxonomicName id="4C22D788647BFFA3FE02E87BD1C3FA67" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[398,498,1387,1411]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FE02E87BD1C3FA67" box="[398,498,1387,1411]" italics="true" pageId="18" pageNumber="19">Borinella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FDA4E87BD2E4FA67" authorityName="Muller" authorityYear="1962" box="[552,725,1387,1411]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FDA4E87BD2E4FA67" box="[552,725,1387,1411]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, but through the shared derived characteristic of the posterior cusp we determine it as the former genus. Chen et al., (2019; figs. 6.8, 6.9) illustrated similar and coeval (associated with
<taxonomicName id="4C22D788647BFFA3FEADE8FCD19DF9E7" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[289,428,1515,1539]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FEADE8FCD19DF9E7" box="[289,428,1515,1539]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
) elements from
<collectingCountry id="F335EC9B647BFFA3FDE2E8FBD284F9E7" box="[622,693,1515,1539]" name="Oman" pageId="18" pageNumber="19">Oman</collectingCountry>
, which they assigned to
<taxonomicName id="4C22D788647BFFA3FEDDEB1BD1DAF9C7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[337,491,1547,1571]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B9567019647BFFA3FEDDEB1BD1DAF9C7" box="[337,491,1547,1571]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FE7FEB1CD21FF9C7" box="[499,558,1548,1571]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
A. Teir illustrated specimens more closely resemble
<taxonomicName id="4C22D788647BFFA3FD9EEB3BD174F987" authority="(Huckriede)" baseAuthorityName="Huckriede" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="tethydis">
<emphasis id="B9567019647BFFA3FD9EEB3BD087F987" italics="true" pageId="18" pageNumber="19">Gladigondolella tethydis</emphasis>
(Huckriede)
</taxonomicName>
and were later assigned to a new species of
<taxonomicName id="4C22D788647BFFA3FF69EB7BD1A3F967" authorityName="Muller" authorityYear="1962" box="[229,402,1643,1667]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FF69EB7BD1A3F967" box="[229,402,1643,1667]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
:
<taxonomicName id="4C22D788647BFFA3FE2CEB7BD35AFEE7" authority="(Chen et al., 2021)" baseAuthorityName="Chen" baseAuthorityYear="2021" box="[416,875,235,1667]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FE2CEB7BD248F967" box="[416,633,1643,1667]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
(
<bibRefCitation id="EFB3D1FA647BFFA3FD04EB7BD350FEE7" author="Chen, Y. &amp; Joachimski, M. M. &amp; Richoz, S. &amp; Krystyn, L. &amp; Kolar-Jurkovsek, T. &amp; Lai, X. &amp; Zhang, Z." box="[648,865,235,1667]" pageId="18" pageNumber="19" refId="ref39445" refString="Chen, Y., Joachimski, M. M., Richoz, S., Krystyn, L., AljinoviC, D., SmirCiC, D., Kolar-Jurkovsek, T., Lai, X., &amp; Zhang, Z. (2021). Smithian and Spathian (Early Triassic) conodonts from Oman and Croatia and their depth habitat revealed. Global and Planetary Change, 196, 103362. https: // doi. org / 10. 1016 / j. gloplacha. 2020.103362" type="journal volume" year="2021">Chen et al., 2021</bibRefCitation>
)
</taxonomicName>
. To date
<taxonomicName id="4C22D788647BFFA3FC43EDFBD44DFEE7" authorityName="Muller" authorityYear="1962" box="[975,1148,235,259]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FC43EDFBD44DFEE7" box="[975,1148,235,259]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
is mostly known from the Middle Triassic (e.g.
<bibRefCitation id="EFB3D1FA647BFFA3FB94EC1BD501FEC7" author="Goudemand, N. &amp; Orchard, M. J. &amp; Bucher, H. &amp; Jenks, J." box="[1048,1328,267,291]" pageId="18" pageNumber="19" pagination="199 - 207" refId="ref41061" refString="Goudemand, N., Orchard, M. J., Bucher, H., &amp; Jenks, J. (2012 a). The elusive origin of Chiosella timorensis (Conodont Triassic). Geobios, 45 (2), 199 - 207." type="journal article" year="2012">Goudemand et al., 2012a</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA647BFFA3FACCEC1BD382FEA7" author="Orchard, M. J. &amp; Gradinaru, E. &amp; Nicora, A." pageId="18" pageNumber="19" refId="ref45799" refString="Orchard, M. J., Gradinaru, E., &amp; Nicora, A. (2007 a). A summary of the conodont succession around the Olenekian-Anisian boundary at Desli Caira, North Dobrogea, Romania." type="book" year="2007">Orchard et al., 2007a</bibRefCitation>
,
<bibRefCitation id="EFB3D1FA647BFFA3FC4FEC3BD434FEA7" author="Orchard, M. J. &amp; Lehrmann, D. J. &amp; Wei, J. &amp; Wang, H. &amp; Taylor, H. J." box="[963,1029,299,323]" pageId="18" pageNumber="19" pagination="347 - 354" refId="ref45840" refString="Orchard, M. J., Lehrmann, D. J., Wei, J., Wang, H., &amp; Taylor, H. J. (2007 b). Conodonts from the Olenekian-Anisian boundary beds, Guandao, Guizhou Province, China. New Mexico Museum of Natural History and Science Bulletin, 41, 347 - 354." type="journal article" year="2007">2007b</bibRefCitation>
). If, as it seems, the here illustrated specimens and/or those from Chen et al., (2019, 2021) do indeed belong to
<taxonomicName id="4C22D788647BFFA3FBAEEC7BD4FEFE67" authorityName="Muller" authorityYear="1962" box="[1058,1231,363,387]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FBAEEC7BD4FEFE67" box="[1058,1231,363,387]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, then they represent their oldest known representatives of this genus so far. A few unpublished specimens from middle Smithian rocks of
<collectingCountry id="F335EC9B647BFFA3FC01ECDBD3E3FE07" box="[909,978,459,483]" name="Oman" pageId="18" pageNumber="19">Oman</collectingCountry>
may also belong to this genus (Leu et al., submitted; L. Dudit, pers. comm.), which further question the origin and age of
<taxonomicName id="4C22D788647BFFA3FBC0EF1BD4C8FDC7" authorityName="Muller" authorityYear="1962" box="[1100,1273,523,547]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FBC0EF1BD4C8FDC7" box="[1100,1273,523,547]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
. Its preference for deeper, colder habitats (
<bibRefCitation id="EFB3D1FA647BFFA3FBD5EF3BD517FDA7" author="Trotter, J. A. &amp; Williams, I. S. &amp; Nicora, A. &amp; Mazza, M. &amp; Rigo, M." box="[1113,1318,555,579]" pageId="18" pageNumber="19" pagination="165 - 174" refId="ref47635" refString="Trotter, J. A., Williams, I. S., Nicora, A., Mazza, M., &amp; Rigo, M. (2015). Long-term cycles of Triassic climate change: A new δ 18 O record from conodont apatite. Earth and Planetary Science Letters, 415, 165 - 174. https: // doi. org / 10.1016 / j. epsl. 2015.01. 038" type="journal article" year="2015">Trotter et al., 2015</bibRefCitation>
) may then explain its spotty occurrences within the Early Triassic, preferably during relatively colder intervals and/or in deeper, colder refuge areas like
<collectingCountry id="F335EC9B647BFFA3FB0DEF9BD4F8FD47" box="[1153,1225,651,675]" name="Oman" pageId="18" pageNumber="19">Oman</collectingCountry>
.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCCEEFBCD47DFCC7" blockId="18.[813,1437,235,803]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCCEEFBCD38CFD27" box="[834,957,684,707]" italics="true" pageId="18" pageNumber="19">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B647BFFA3FB99EFBBD468FD27" box="[1045,1113,683,707]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
;
<taxonomicName id="4C22D788647BFFA3FBEAEFBCD4DEFD27" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[1126,1263,683,707]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FBEAEFBCD4DEFD27" box="[1126,1263,683,707]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FAA5EFBCD5ADFD27" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[1321,1436,683,707]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B9567019647BFFA3FAA5EFBCD5ADFD27" box="[1321,1436,683,707]" italics="true" pageId="18" pageNumber="19">Tr. homeri</emphasis>
</taxonomicName>
zone, Jiarong (
<bibRefCitation id="EFB3D1FA647BFFA3FC40EFDBD4B8FD07" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[972,1161,715,739]" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
). Bed 54, Luolou formation,
<taxonomicName id="4C22D788647BFFA3FCEAEFFCD3DCFCE7" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[870,1005,747,771]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FCEAEFFCD3DCFCE7" box="[870,1005,747,771]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FBAFEFFCD4A4FCE7" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[1059,1173,747,771]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B9567019647BFFA3FBAFEFFCD4A4FCE7" box="[1059,1173,747,771]" italics="true" pageId="18" pageNumber="19">Tr. homeri</emphasis>
</taxonomicName>
zone, Bianyang section,
<collectingRegion id="49E662E9647BFFA3FCA1EE1BD3BDFCC7" box="[813,908,779,803]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou</collectingRegion>
(
<bibRefCitation id="EFB3D1FA647BFFA3FC15EE1BD40FFCC7" author="Yan, C. &amp; Wang, L. &amp; Jiang, H. &amp; Wignall, P. B. &amp; Sun, Y. &amp; Chen, Y. &amp; Lai, X." box="[921,1086,779,803]" pageId="18" pageNumber="19" pagination="509 - 522" refId="ref48484" refString="Yan, C., Wang, L., Jiang, H., Wignall, P. B., Sun, Y., Chen, Y., &amp; Lai, X. (2013). Uppermost Permian to Lower Triassic conodonts at Bianyang section, Guizhou province, South China. Palaios, 28 (8), 509 - 522. https: // doi. org / 10.2110 / palo. 2012. p 12 - 077 r" type="journal article" year="2013">Yan et al., 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="740A658F03C78F87520FBC3230EBA155">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="8F1AEF808CA5A9896260865D02D583E4">Leu, Marc</mods:namePart>
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<subSubSection id="C338FF80647BFFA3FCA1EE5BD4D8FC87" box="[813,1257,843,867]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B9DAC0B647BFFA3FCA1EE5BD4D8FC87" blockId="18.[813,1257,843,899]" box="[813,1257,843,867]" pageId="18" pageNumber="19">
<heading id="D0D51B67647BFFA3FCA1EE5BD4D8FC87" box="[813,1257,843,867]" fontSize="9" level="2" pageId="18" pageNumber="19" reason="8">
<taxonomicName id="4C22D788647BFFA3FCA1EE5BD477FC87" authority="Chen" authorityName="Chen" box="[813,1094,843,867]" family="Gondolellidae" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FCA1EE5BD435FC87" box="[813,1028,843,867]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
Chen
</taxonomicName>
morphotype B
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA5FCA1EE7BD0EBFD07" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="8B9DAC0B647BFFA3FCA1EE7BD3E0FC67" blockId="18.[813,1257,843,899]" box="[813,977,875,899]" pageId="18" pageNumber="19">
<figureCitation id="1319B08E647BFFA3FCA1EE7BD3E0FC67" box="[813,977,875,899]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.12000264" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="18" pageNumber="19">
Fig. 13E,
<bibRefCitation id="EFB3D1FA647BFFA3FC1CEE7CD3E0FC67" author="Jenks, J. F. &amp; Monnet, C. &amp; Balini, M. &amp; Brayard, A. &amp; Meier, M." box="[912,977,876,899]" pageId="18" pageNumber="19" pagination="329 - 388" refId="ref42107" refString="Jenks, J. F., Monnet, C., Balini, M., Brayard, A., &amp; Meier, M. (2015). Biostratigraphy of Triassic ammonoids. In C. Klug, D. Korn, K. De Baets, I. Kruta, &amp; R. H. Mapes (Eds.), Ammonoid paleobiology: From macroevolution to paleogeography (pp. 329 - 388). Dordrecht: Springer." type="book chapter" year="2015">F, H, I</bibRefCitation>
</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1EEBBD5B2FC27" blockId="18.[813,1436,939,1155]" box="[813,1411,939,963]" pageId="18" pageNumber="19">
2015
<taxonomicName id="4C22D788647BFFA3FCEBEEBBD430FC27" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[871,1025,939,963]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FCEBEEBBD430FC27" box="[871,1025,939,963]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. B; Chen et al., figs. 9.99.10.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1EEDBD38AFBE7" blockId="18.[813,1436,939,1155]" pageId="18" pageNumber="19">
2019
<taxonomicName id="4C22D788647BFFA3FCE4EEDBD433FC07" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[872,1026,971,995]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FCE4EEDBD433FC07" box="[872,1026,971,995]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. A Chen; Liu et al., p. 14, pl. 4, fig. 19 (only).
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1E91BD359FBA7" blockId="18.[813,1436,939,1155]" pageId="18" pageNumber="19">
2019
<taxonomicName id="4C22D788647BFFA3FCE5E91BD432FBC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[873,1027,1035,1059]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FCE5E91BD432FBC7" box="[873,1027,1035,1059]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. B Chen; Liu et al., p. 14, pl. 4, fig. 9.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1E95BD428FB67" blockId="18.[813,1436,939,1155]" pageId="18" pageNumber="19">
2021
<taxonomicName id="4C22D788647BFFA3FCE7E95BD477FB87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[875,1094,1099,1123]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii" status="sp. nov.">
<emphasis id="B9567019647BFFA3FCE7E95BD477FB87" box="[875,1094,1099,1123]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FBDCE95CD499FB87" box="[1104,1192,1100,1123]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
; Chen et al., figs. 4.5, 4.22, 4.25, 4.26 (only).
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1E9BBD469FB27" blockId="18.[813,1437,1195,1668]" box="[813,1112,1195,1219]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCA1E9BBD3BFFB27" box="[813,910,1195,1219]" italics="true" pageId="18" pageNumber="19">Material</emphasis>
. ca.
<specimenCount id="9D246782647BFFA3FC32E9BBD465FB27" box="[958,1108,1195,1219]" count="20" pageId="18" pageNumber="19" type="generic">20 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1E9DCD357FAA7" blockId="18.[813,1437,1195,1668]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCA1E9DCD3ACFB07" box="[813,925,1228,1251]" italics="true" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Segminiplanate P
<subScript id="17A6AE4E647BFFA3FBE5E9C7D443FB0C" attach="left" box="[1129,1138,1239,1256]" fontSize="7" pageId="18" pageNumber="19">1</subScript>
element, narrow platform tapers towards the anterior, denticles moderately discrete. Very large, distinct, terminal, strongly reclined cusp.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCA1E85CD52BF9E7" blockId="18.[813,1437,1195,1668]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCA1E85CD381FA87" box="[813,944,1356,1379]" italics="true" pageId="18" pageNumber="19">Description.</emphasis>
Te platform is narrow, subparallel, slightly upturned and tapers towards the anterior end. Te posterior margin is rounded in a narrow platform brim around the large terminal cusp. Te cusp is massive and inclined posteriorly. Te carina is moderately high with pointed denticles which are almost uniform in height.
</paragraph>
<paragraph id="8B9DAC0B647BFFA5FCA1EB1BD2B3FD60" blockId="18.[813,1437,1195,1668]" lastBlockId="20.[151,775,235,739]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCA1EB1BD3A0F9C7" box="[813,913,1547,1571]" italics="true" pageId="18" pageNumber="19">Remarks.</emphasis>
Te large terminal cusp differs from
<taxonomicName id="4C22D788647BFFA3FAB4EB1BD5ADF9C7" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[1336,1436,1547,1571]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FAB4EB1BD5ADF9C7" box="[1336,1436,1547,1571]" italics="true" pageId="18" pageNumber="19">Borinella</emphasis>
</taxonomicName>
whose denticles are also higher and more discrete, but resembles that of
<taxonomicName id="4C22D788647BFFA3FC7DEB5BD496F987" authorityName="Kozur" authorityYear="1990" box="[1009,1191,1611,1635]" family="Gondolellidae" genus="Scythogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FC7DEB5BD496F987" box="[1009,1191,1611,1635]" italics="true" pageId="18" pageNumber="19">Scythogondolella</emphasis>
</taxonomicName>
. Yet,
<taxonomicName id="4C22D788647BFFA3FB6AEB5BD503F987" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1254,1330,1611,1635]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FB6AEB5BD503F987" box="[1254,1330,1611,1635]" italics="true" pageId="18" pageNumber="19">Gl. laii</emphasis>
</taxonomicName>
morphotype B lacks a prominent rounded basal loop surrounding a small pit, a distinctive feature of
<taxonomicName id="4C22D788647DFFA5FD86EDFBD2F1FEE7" authorityName="Kozur" authorityYear="1990" box="[522,704,235,259]" family="Gondolellidae" genus="Scythogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FD86EDFBD2F1FEE7" box="[522,704,235,259]" italics="true" pageId="20" pageNumber="21">Scythogondolella</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB3D1FA647DFFA5FD40EDFBD12EFEC7" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." pageId="20" pageNumber="21" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
suggests that their
<emphasis id="B9567019647DFFA5FD8DEC1CD22EFEC7" box="[513,543,268,291]" italics="true" pageId="20" pageNumber="21">Ng</emphasis>
. n. sp. B could be a juvenile stage of
<emphasis id="B9567019647DFFA5FEC7EC3CD158FEA7" box="[331,361,300,323]" italics="true" pageId="20" pageNumber="21">Ng</emphasis>
. n. sp. A (reassigned here and in
<bibRefCitation id="EFB3D1FA647DFFA5FD40EC3BD12DFE87" author="Chen, Y. &amp; Joachimski, M. M. &amp; Richoz, S. &amp; Krystyn, L. &amp; Kolar-Jurkovsek, T. &amp; Lai, X. &amp; Zhang, Z." pageId="20" pageNumber="21" refId="ref39445" refString="Chen, Y., Joachimski, M. M., Richoz, S., Krystyn, L., AljinoviC, D., SmirCiC, D., Kolar-Jurkovsek, T., Lai, X., &amp; Zhang, Z. (2021). Smithian and Spathian (Early Triassic) conodonts from Oman and Croatia and their depth habitat revealed. Global and Planetary Change, 196, 103362. https: // doi. org / 10. 1016 / j. gloplacha. 2020.103362" type="journal volume" year="2021">Chen et al. (2021)</bibRefCitation>
to
<taxonomicName id="4C22D788647DFFA5FEC8EC5BD22FFE87" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[324,542,331,355]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647DFFA5FEC8EC5BD22FFE87" box="[324,542,331,355]" italics="true" pageId="20" pageNumber="21">Gladigondolella laii</emphasis>
</taxonomicName>
see above) based on size considerations and the fact that they are both found in the same interval. Yet, they were not found in the same samples and the small forms (
<taxonomicName id="4C22D788647DFFA5FE50ECBBD216FE27" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[476,551,427,451]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647DFFA5FE50ECBBD216FE27" box="[476,551,427,451]" italics="true" pageId="20" pageNumber="21">Gl. laii</emphasis>
</taxonomicName>
morphotype B) lack a posterior process, although the juvenile forms of true gladigondolellids usually bear a posterior process. Hence, we tentatively keep them as separate morphotypes. Nevertheless, in
<bibRefCitation id="EFB3D1FA647DFFA5FEA8EF3BD1DCFDA7" author="Chen, Y. &amp; Joachimski, M. M. &amp; Richoz, S. &amp; Krystyn, L. &amp; Kolar-Jurkovsek, T. &amp; Lai, X. &amp; Zhang, Z." box="[292,493,555,579]" pageId="20" pageNumber="21" refId="ref39445" refString="Chen, Y., Joachimski, M. M., Richoz, S., Krystyn, L., AljinoviC, D., SmirCiC, D., Kolar-Jurkovsek, T., Lai, X., &amp; Zhang, Z. (2021). Smithian and Spathian (Early Triassic) conodonts from Oman and Croatia and their depth habitat revealed. Global and Planetary Change, 196, 103362. https: // doi. org / 10. 1016 / j. gloplacha. 2020.103362" type="journal volume" year="2021">Chen et al. (2021)</bibRefCitation>
, both forms are found in the same sample (e.g. figs. 4.23 and 4.24) and can therefore be considered as different growth stages.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF25EBA5D1B2F92D" blockId="18.[169,387,1717,1737]" box="[169,387,1717,1737]" pageId="18" pageNumber="19">
<subSection id="E2ADB5E0647BFFA3FF25EBA5D1B1F92D" box="[169,384,1717,1737]" pageId="18" pageNumber="19" type="multiple">(See figure on next page.</subSection>
)
</paragraph>
<caption id="DF5DFC83647BFFA3FF25EBDFD375F867" pageId="18" pageNumber="19" startId="18.[169,200,1743,1763]" targetBox="[190,1402,249,1755]" targetPageId="19">
<paragraph id="8B9DAC0B647BFFA3FF25EBDFD54FF907" blockId="18.[169,1415,1742,1923]" box="[169,1406,1742,1763]" pageId="18" pageNumber="19">
<figureCitation id="1319B08E647BFFA3FF25EBDFD0D9F907" box="[169,232,1743,1763]" captionStart="Fig" captionStartId="19.[169,200,1776,1796]" captionTargetBox="[190,1402,249,1755]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000264" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF25EBDFD0D9F907" bold="true" box="[169,232,1743,1763]" pageId="18" pageNumber="19">Fig. 13</emphasis>
</figureCitation>
<taxonomicName id="4C22D788647BFFA3FF7EEBDFD1A5F907" authorityName="Hirsch" authorityYear="1994" box="[242,404,1743,1763]" pageId="18" pageNumber="18" rank="subFamily" subFamily="Gladigondolellinae">Gladigondolellinae</taxonomicName>
,
<taxonomicName id="4C22D788647BFFA3FE17EBDFD202F907" authorityName="Hirsch" authorityYear="1994" box="[411,563,1743,1763]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="21" phylum="Chordata" rank="subFamily" subFamily="Neogondolellinae">Neogondolellinae</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FDD2EBDFD2E3F907" authorityName="Orchard" authorityYear="2005" box="[606,722,1743,1763]" pageId="18" pageNumber="50" rank="subFamily" subFamily="Cornudininae">Cornudininae</taxonomicName>
from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF25EBFAD5B4F91A" blockId="18.[169,1415,1742,1923]" box="[169,1413,1770,1794]" pageId="18" pageNumber="19">
is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E647BFFA3FDA3EBE4D207F8E6" attach="left" box="[559,566,1780,1794]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B9567019647BFFA3FC5FEBFAD3D0F91A" bold="true" box="[979,993,1770,1790]" pageId="18" pageNumber="19">A</emphasis>
<taxonomicName id="4C22D788647BFFA3FC6AEBFAD48EF91A" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[998,1215,1770,1790]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647BFFA3FC6AEBFAD44CF91A" box="[998,1149,1770,1790]" italics="true" pageId="18" pageNumber="19">Borinella buurensis</emphasis>
(Dagis)
</taxonomicName>
, LAR212, PIMUZ 39103.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF25EA15D375F867" blockId="18.[169,1415,1742,1923]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF25EA15D084F8FD" bold="true" box="[169,181,1797,1817]" pageId="18" pageNumber="19">B</emphasis>
,
<emphasis id="B9567019647BFFA3FF32EA15D0FAF8FD" bold="true" box="[190,203,1797,1817]" pageId="18" pageNumber="19">C</emphasis>
,
<emphasis id="B9567019647BFFA3FF5FEA15D0D0F8FD" bold="true" box="[211,225,1797,1817]" pageId="18" pageNumber="19">G</emphasis>
<taxonomicName id="4C22D788647BFFA3FF69EA15D1B2F8FD" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[229,387,1797,1817]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FF69EA15D1B2F8FD" box="[229,387,1797,1817]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
</taxonomicName>
(morphotype A);
<emphasis id="B9567019647BFFA3FD94EA15D215F8FD" bold="true" box="[536,548,1797,1817]" pageId="18" pageNumber="19">B</emphasis>
LAR231C, PIMUZ 39132;
<emphasis id="B9567019647BFFA3FD74EA15D334F8FD" bold="true" box="[760,773,1797,1817]" pageId="18" pageNumber="19">C</emphasis>
BAN2, PIMUZ 39133;
<emphasis id="B9567019647BFFA3FC36EA15D3F9F8FD" bold="true" box="[954,968,1797,1817]" pageId="18" pageNumber="19">G</emphasis>
LAR227C, PIMUZ 39134.
<emphasis id="B9567019647BFFA3FB10EA15D49AF8FD" bold="true" box="[1180,1195,1797,1817]" pageId="18" pageNumber="19">D</emphasis>
,
<emphasis id="B9567019647BFFA3FB3FEA15D4C4F8FD" box="[1203,1269,1797,1817]" italics="true" pageId="18" pageNumber="19">sp.indet</emphasis>
.(P
<subScript id="17A6AE4E647BFFA3FA82EA1ED524F8F8" attach="left" box="[1294,1301,1806,1820]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
element?), SHA319, PIMUZ 39293.
<emphasis id="B9567019647BFFA3FEE2EA0FD148F8D7" bold="true" box="[366,377,1823,1843]" pageId="18" pageNumber="19">E</emphasis>
,
<emphasis id="B9567019647BFFA3FE0DEA0FD1BDF8D7" bold="true" box="[385,396,1823,1843]" pageId="18" pageNumber="19">F</emphasis>
,
<emphasis id="B9567019647BFFA3FE18EA0FD193F8D7" bold="true" box="[404,418,1823,1843]" pageId="18" pageNumber="19">H</emphasis>
,
<emphasis id="B9567019647BFFA3FE27EA0FD180F8D7" bold="true" box="[427,433,1823,1843]" pageId="18" pageNumber="19">I</emphasis>
<taxonomicName id="4C22D788647BFFA3FE39EA0FD262F8D7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[437,595,1823,1843]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FE39EA0FD262F8D7" box="[437,595,1823,1843]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
</taxonomicName>
(morphotype B);
<emphasis id="B9567019647BFFA3FD6BEA0FD2C3F8D7" bold="true" box="[743,754,1823,1843]" pageId="18" pageNumber="19">E</emphasis>
LIL510,PIMUZ 39135;
<emphasis id="B9567019647BFFA3FC23EA0FD38BF8D7" bold="true" box="[943,954,1823,1843]" pageId="18" pageNumber="19">F</emphasis>
LIL512, PIMUZ 39136;
<emphasis id="B9567019647BFFA3FBF4EA0FD4B7F8D7" bold="true" box="[1144,1158,1823,1843]" pageId="18" pageNumber="19">H</emphasis>
QIA140, PIMUZ 39137;
<emphasis id="B9567019647BFFA3FAC7EA0FD560F8D7" bold="true" box="[1355,1361,1823,1843]" pageId="18" pageNumber="19">I</emphasis>
LIL510,PIMUZ 39138.
<emphasis id="B9567019647BFFA3FEEEEA2AD1B1F8AA" bold="true" box="[354,384,1850,1870]" pageId="18" pageNumber="19">JL</emphasis>
,
<emphasis id="B9567019647BFFA3FE04EA2AD1A6F8AA" bold="true" box="[392,407,1850,1870]" pageId="18" pageNumber="19">O</emphasis>
,
<taxonomicName id="4C22D788647BFFA3FE2CEA2AD2B9F8AA" authority="(Sweet)" baseAuthorityName="Sweet" box="[416,648,1850,1870]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B9567019647BFFA3FE2CEA2AD273F8AA" box="[416,578,1850,1870]" italics="true" pageId="18" pageNumber="19">Spathicuspus spathi</emphasis>
(Sweet)
</taxonomicName>
;
<emphasis id="B9567019647BFFA3FD03EA2AD2A6F8AA" bold="true" box="[655,663,1850,1870]" pageId="18" pageNumber="19">J</emphasis>
QIA141, PIMUZ 39300;
<emphasis id="B9567019647BFFA3FCD1EA2AD358F8AA" bold="true" box="[861,873,1850,1870]" pageId="18" pageNumber="19">K</emphasis>
QIA144, PIMUZ 39301;
<emphasis id="B9567019647BFFA3FBA3EA2AD40BF8AA" bold="true" box="[1071,1082,1850,1870]" pageId="18" pageNumber="19">L</emphasis>
BAN1, PIMUZ 39302;
<emphasis id="B9567019647BFFA3FB63EA2AD4CFF8AA" bold="true" box="[1263,1278,1850,1870]" pageId="18" pageNumber="19">O</emphasis>
SHA322,PIMUZ 39303.
<emphasis id="B9567019647BFFA3FF6AEA45D0C9F88D" bold="true" box="[230,248,1877,1897]" pageId="18" pageNumber="19">M</emphasis>
,
<emphasis id="B9567019647BFFA3FE8CEA45D13DF88D" bold="true" box="[256,268,1877,1897]" pageId="18" pageNumber="19">P</emphasis>
<taxonomicName id="4C22D788647BFFA3FE9CEA45D14DF88D" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[272,380,1877,1897]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B9567019647BFFA3FE9CEA45D14DF88D" box="[272,380,1877,1897]" italics="true" pageId="18" pageNumber="19">Spathicuspus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FE0DEA45D19FF88D" box="[385,430,1877,1897]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
A;
<emphasis id="B9567019647BFFA3FE4AEA45D1E9F88D" bold="true" box="[454,472,1877,1897]" pageId="18" pageNumber="19">M</emphasis>
QIA144, PIMUZ 39297;
<emphasis id="B9567019647BFFA3FD11EA45D298F88D" bold="true" box="[669,681,1877,1897]" pageId="18" pageNumber="19">P</emphasis>
, QIA143, PIMUZ 39298.
<emphasis id="B9567019647BFFA3FCFEEA45D3B1F88D" bold="true" box="[882,896,1877,1897]" pageId="18" pageNumber="19">N</emphasis>
,
<emphasis id="B9567019647BFFA3FC05EA45D3A4F88D" bold="true" box="[905,917,1877,1897]" pageId="18" pageNumber="19">R</emphasis>
<taxonomicName id="4C22D788647BFFA3FC15EA45D40DF88D" box="[921,1084,1877,1897]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="spathi">
<emphasis id="B9567019647BFFA3FC15EA45D40DF88D" box="[921,1084,1877,1897]" italics="true" pageId="18" pageNumber="19">Spathicuspus spathi</emphasis>
</taxonomicName>
(juvenile) (Sweet);
<emphasis id="B9567019647BFFA3FB51EA45D4DAF88D" bold="true" box="[1245,1259,1877,1897]" pageId="18" pageNumber="19">N</emphasis>
, LIL515D, PIMUZ 39304;
<emphasis id="B9567019647BFFA3FF6AEA7FD0C3F867" bold="true" box="[230,242,1903,1923]" pageId="18" pageNumber="19">R</emphasis>
QIA143,PIMUZ 39305.
<emphasis id="B9567019647BFFA3FE3BEA7FD1F7F867" bold="true" box="[439,454,1903,1923]" pageId="18" pageNumber="19">Q</emphasis>
<taxonomicName id="4C22D788647BFFA3FE47EA7FD206F867" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[459,567,1903,1923]" genus="Spathicuspus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B9567019647BFFA3FE47EA7FD206F867" box="[459,567,1903,1923]" italics="true" pageId="18" pageNumber="19">Spathicuspus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FDB0EA7FD258F867" box="[572,617,1903,1923]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
B; LAR226C, PIMUZ 39299
</paragraph>
</caption>
<caption id="DF5DFC83647AFFA2FF25EBE0D1C8F8E0" ID-DOI="http://doi.org/10.5281/zenodo.12000264" ID-Zenodo-Dep="12000264" box="[169,505,1776,1796]" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="19" pageNumber="20" startId="19.[169,200,1776,1796]" targetBox="[190,1402,249,1755]" targetPageId="19" targetType="figure">
<paragraph id="8B9DAC0B647AFFA2FF25EBE0D1C8F8E0" blockId="19.[169,505,1776,1796]" box="[169,505,1776,1796]" pageId="19" pageNumber="20">
<emphasis id="B9567019647AFFA2FF25EBE0D0D6F8E0" bold="true" box="[169,231,1776,1796]" pageId="19" pageNumber="20">Fig. 13</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<paragraph id="8B9DAC0B647DFFA5FF21EF9CD0EBFD07" blockId="20.[151,775,235,739]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FF21EF9CD119FD47" box="[173,296,652,675]" italics="true" pageId="20" pageNumber="21">Occurrence</emphasis>
.
<taxonomicName id="4C22D788647DFFA5FEBAEF9CD1F1FD47" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[310,448,651,675]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647DFFA5FEBAEF9CD1F1FD47" box="[310,448,651,675]" italics="true" pageId="20" pageNumber="21">Ic. collinsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647DFFA5FE76EF9CD25FFD47" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[506,622,651,675]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B9567019647DFFA5FE76EF9CD25FFD47" box="[506,622,651,675]" italics="true" pageId="20" pageNumber="21">Tr. homeri</emphasis>
</taxonomicName>
zone, Luolou formation, Jiarong,
<collectingRegion id="49E662E9647DFFA5FEF8EFBBD1E4FD27" box="[372,469,683,707]" country="China" name="Guizhou" pageId="20" pageNumber="21">Guizhou</collectingRegion>
, South
<collectingCountry id="F335EC9B647DFFA5FDA2EFBBD241FD27" box="[558,624,683,707]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647DFFA5FD0EEFBBD0FAFD07" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." pageId="20" pageNumber="21" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,134 @@
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<mods:title id="9D47D98C50F45C6FF1A0495914E98D5D">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="D013F186A4DE731E3A9F2A2A5066539B">Bucher, Hugo</mods:namePart>
<mods:affiliation id="5897FC0B12AAD829C748EDD2E586142C">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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NEOSPATHODUS
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, pl. 10, fig. 15.
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stratum and locality
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(
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<subSubSection id="C338FF80647DFFA5FF1BE87BD4CEF9E7" pageId="20" pageNumber="21" type="description">
<paragraph id="8B9DAC0B647DFFA5FF1BE87BD0C3FA67" blockId="20.[151,512,1355,1411]" box="[151,242,1387,1411]" pageId="20" pageNumber="21">
<figureCitation id="1319B08E647DFFA5FF1BE87BD0C3FA67" box="[151,242,1387,1411]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.12000264" httpUri="https://zenodo.org/record/12000264/files/figure.png" pageId="20" pageNumber="21">Fig. 13A</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BE8BBD1B0FA07" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
1978
<taxonomicName id="4C22D788647DFFA5FF5EE8BBD233FA27" authority="Sweet" authorityName="Sweet" box="[210,514,1451,1475]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="jubata">
<emphasis id="B9567019647DFFA5FF5EE8BBD188FA27" box="[210,441,1451,1475]" italics="true" pageId="20" pageNumber="21">Neogondolella jubata</emphasis>
Sweet
</taxonomicName>
; Weitschat &amp; Lehmann, p. 98, pl. 13, figs. 16.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BE8FBD170F9A7" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
*1984
<taxonomicName id="4C22D788647DFFA5FF52E8FBD1D8F9E7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[222,489,1515,1539]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis" status="sp. nov.">
<emphasis id="B9567019647DFFA5FF52E8FBD1D8F9E7" box="[222,489,1515,1539]" italics="true" pageId="20" pageNumber="21">Neogondolella buurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647DFFA5FE7EE8FCD21EF9E7" box="[498,559,1516,1539]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
; Dagis, p. 12, pl. 2, figs. 615, pl. 3, fig. 12, pl. 11, figs. 14, pl. 12, figs. 1, 2, pl. 16, figs. 14.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEB5BD151F967" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
1984
<taxonomicName id="4C22D788647DFFA5FF5AEB5BD215F987" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[214,548,1611,1635]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="elongata">
<emphasis id="B9567019647DFFA5FF5AEB5BD1E6F987" box="[214,471,1611,1635]" italics="true" pageId="20" pageNumber="21">Neogondolella elongata</emphasis>
Sweet
</taxonomicName>
; Hatleberg &amp; Clark, Pl. 1, fig. 14 (only).
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEB9BD17DF927" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
2005
<taxonomicName id="4C22D788647DFFA5FF5FEB9BD290F947" authority="Kozur &amp; Mostler" authorityName="Kozur &amp; Mostler" box="[211,673,1675,1699]" genus="Neogondolella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sweeti">
<emphasis id="B9567019647DFFA5FF5FEB9BD15CF947" box="[211,365,1675,1699]" italics="true" pageId="20" pageNumber="21">Neogondolella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FE10EB9CD1ECF947" box="[412,477,1676,1699]" italics="true" pageId="20" pageNumber="21">sweeti</emphasis>
Kozur &amp; Mostler
</taxonomicName>
; Zhao, p. 131, pl. 13, fig. 1.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEBDBD12BF8E7" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
2007
<taxonomicName id="4C22D788647DFFA5FF59EBDBD235F907" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[213,516,1739,1763]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FF59EBDBD19BF907" box="[213,426,1739,1763]" italics="true" pageId="20" pageNumber="21">Borinella buurensis</emphasis>
(Dagis)
</taxonomicName>
; Orchard, p. 113, pl. 1, nr. 7, 15, 27.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEA1BD0C2F8A7" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
2008
<taxonomicName id="4C22D788647DFFA5FF59EA1BD237F8C7" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[213,518,1803,1827]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FF59EA1BD19AF8C7" box="[213,427,1803,1827]" italics="true" pageId="20" pageNumber="21">Borinella buurensis</emphasis>
(Dagis)
</taxonomicName>
; Nakrem et al., p. 528, fig. 4.19.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEA5BD155F867" blockId="20.[151,775,1451,1923]" pageId="20" pageNumber="21">
2008
<taxonomicName id="4C22D788647DFFA5FF5BEA5BD207F887" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[215,566,1867,1891]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FF5BEA5BD10AF887" box="[215,315,1867,1891]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FEFFEA5BD1EAF887" box="[371,475,1867,1891]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagis)
</taxonomicName>
; Nakrem et al., p. 528, fig. 4.174.18.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EDFBD3F2FEC7" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2008
<taxonomicName id="4C22D788647DFFA5FCF9EDFBD48BFEE7" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[885,1210,235,259]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCF9EDFBD464FEE7" box="[885,1109,235,259]" italics="true" pageId="20" pageNumber="21">Borinella buurensis</emphasis>
(Dagis)
</taxonomicName>
; Orchard, p. 400, figs. 5.95.13.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EC3BD571FE87" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2012a
<taxonomicName id="4C22D788647DFFA5FCF7EC3BD4D3FEA7" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[891,1250,299,323]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCF7EC3BD3EEFEA7" box="[891,991,299,323]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FB97EC3BD4B2FEA7" box="[1051,1155,299,323]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagis)
</taxonomicName>
; Goudemand &amp; Orchard in Goudemand et al., p. 1032, figs. 2AA.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EC7BD4B2FE27" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2015
<taxonomicName id="4C22D788647DFFA5FCEBEC7BD486FE67" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[871,1207,363,387]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCEBEC7BD3FAFE67" box="[871,971,363,387]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FC75EC7BD450FE67" box="[1017,1121,363,387]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagis)
</taxonomicName>
; Chen et al., fig.
<date id="FF9C8ACB647DFFA5FAE5EC7BD350FE47" pageId="20" pageNumber="21" value="2018-09-11">9.11. 2018</date>
<taxonomicName id="4C22D788647DFFA5FCFAEC9BD4C3FE47" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[886,1266,395,419]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCFAEC9BD3EBFE47" box="[886,986,395,419]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FBA9EC9BD4BCFE47" box="[1061,1165,395,419]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagis)
</taxonomicName>
; Maekawa in Maekawa et al., p. 45, fig. 29.27.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1ECDBD3CEFDE7" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2019
<taxonomicName id="4C22D788647DFFA5FCE5ECDBD436FE07" baseAuthorityName="Orchard" baseAuthorityYear="2008" box="[873,1031,459,483]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FCE5ECDBD436FE07" box="[873,1031,459,483]" italics="true" pageId="20" pageNumber="21">Neogondolella</emphasis>
</taxonomicName>
ex. gr.
<emphasis id="B9567019647DFFA5FBD5ECDBD4F9FE07" box="[1113,1224,459,483]" italics="true" pageId="20" pageNumber="21">jakutensis</emphasis>
Dagis; Chen et al., figs. 4.3, 4.5 (only).
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EF1BD3EBFDA7" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2019
<taxonomicName id="4C22D788647DFFA5FCE5EF1BD4F4FDC7" authority="(Dagis)" baseAuthorityName="Dagis" baseAuthorityYear="1984" box="[873,1221,523,547]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCE5EF1BD3E1FDC7" box="[873,976,523,547]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FB8CEF1BD45AFDC7" box="[1024,1131,523,547]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagis)
</taxonomicName>
; Liu et al., p. 13, pl. 3, fig. 10 (only).
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EF5BD3F7FD67" blockId="20.[813,1437,235,643]" pageId="20" pageNumber="21">
2021
<taxonomicName id="4C22D788647DFFA5FCFFEF5BD4C5FD87" authority="(Dagys)" baseAuthorityName="Dagys" box="[883,1268,587,611]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FCFFEF5BD3EBFD87" box="[883,986,587,611]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
aff.
<emphasis id="B9567019647DFFA5FB93EF5BD4BBFD87" box="[1055,1162,587,611]" italics="true" pageId="20" pageNumber="21">buurensis</emphasis>
(Dagys)
</taxonomicName>
; Chen et al., fig. 5.9 (only).
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EFBBD402FD27" blockId="20.[813,1075,683,707]" box="[813,1075,683,707]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FCA1EFBBD429FD27" box="[813,1048,683,707]" italics="true" pageId="20" pageNumber="21">Number of specimens</emphasis>
. 5
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCA1EFFCD44BFB67" blockId="20.[813,1437,747,1539]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FCA1EFFCD39FFCE7" box="[813,942,748,771]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
. Subsymmetrical segminiplanate P
<subScript id="17A6AE4E647DFFA5FAD4EFE7D550FCEC" attach="left" box="[1368,1377,759,776]" fontSize="7" pageId="20" pageNumber="21">1</subScript>
element whose platform extends as a brim around the posterior edge, but not to the anteriormost quarter of the unit. In the posterior half, the platform margins are subparallel in oral view. Te cusp varies in height and may be conspicuous. A smaller posterior denticle may be present in some specimens, in which case, it tends to be offset from the main axis. As in any
<taxonomicName id="4C22D788647DFFA5FB73EEDBD36BFBE7" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FB73EEDBD557FC07" box="[1279,1382,971,995]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
species
</taxonomicName>
, the denticles become gradually more discrete and taller towards the anterior end. As illustrated by the figured specimen, whose denticles at mid-length are almost as high as the anteriormost ones, this denticulation gradient may be variable.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCCEE99BD4F5FAC7" blockId="20.[813,1437,747,1539]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FCCEE99BD393FB47" box="[834,930,1163,1187]" italics="true" pageId="20" pageNumber="21">Remarks</emphasis>
. Orchard (2007) suggested that the cusp of theP
<subScript id="17A6AE4E647DFFA5FCEBE9A7D341FB2C" attach="left" box="[871,880,1207,1224]" fontSize="7" pageId="20" pageNumber="21">1</subScript>
of
<taxonomicName id="4C22D788647DFFA5FC18E9BCD42CFB27" box="[916,1053,1195,1219]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FC18E9BCD42CFB27" box="[916,1053,1195,1219]" italics="true" pageId="20" pageNumber="21">B. buurensis</emphasis>
</taxonomicName>
is typically weak, contrary to that of
<taxonomicName id="4C22D788647DFFA5FCC1E9DCD381FB07" box="[845,944,1228,1251]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sweeti">
<emphasis id="B9567019647DFFA5FCC1E9DCD381FB07" box="[845,944,1228,1251]" italics="true" pageId="20" pageNumber="21">B. sweeti</emphasis>
</taxonomicName>
. Our collections suggest instead it is variable, as illustrated here. In our view it is difficult to differentiate
<taxonomicName id="4C22D788647DFFA5FC12E81CD3CFFAC7" box="[926,1022,1292,1315]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sweeti">
<emphasis id="B9567019647DFFA5FC12E81CD3CFFAC7" box="[926,1022,1292,1315]" italics="true" pageId="20" pageNumber="21">B. sweeti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647DFFA5FBBAE81CD48FFAC7" box="[1078,1214,1291,1315]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FBBAE81CD48FFAC7" box="[1078,1214,1291,1315]" italics="true" pageId="20" pageNumber="21">B. buurensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FCCEE83CD4CEF9E7" blockId="20.[813,1437,747,1539]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FCCEE83CD3F6FAA7" box="[834,967,1324,1347]" italics="true" pageId="20" pageNumber="21">Occurrence.</emphasis>
South
<collectingCountry id="F335EC9B647DFFA5FBBAE83BD44DFAA7" box="[1078,1148,1323,1347]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
;
<taxonomicName id="4C22D788647DFFA5FB1BE83CD3E9FA80" authority="Zone, Tsoteng" authorityName="Zone, Tsoteng" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B9567019647DFFA5FB1BE83CD5ACFAA7" box="[1175,1437,1323,1347]" italics="true" pageId="20" pageNumber="21">Nv. pingdingshanensis</emphasis>
Zone, Tsoteng
</taxonomicName>
section (
<bibRefCitation id="EFB3D1FA647DFFA5FBDFE85BD5BCFA87" author="Goudemand, N. &amp; Orchard, M. J. &amp; Tafforeau, P. &amp; Urdy, S. &amp; Bruehwiler, T. &amp; Brayard, A. &amp; BrUhwiler, T. &amp; Bucher, H." box="[1107,1421,1355,1379]" pageId="20" pageNumber="21" pagination="1021 - 1034" refId="ref41110" refString="Goudemand, N., Orchard, M. J., Tafforeau, P., Urdy, S., Bruehwiler, T., Brayard, A., BrUhwiler, T., &amp; Bucher, H. (2012 b). Early Triassic conodont clusters from South China: revision of the architecture of the 15 element apparatuses of the superfamily Gondolelloidea. Palaeontology, 55 (5), 1021 - 1034. https: // doi. org / 10.1111 / j. 1475 - 4983.2012.01174. x" type="journal article" year="2012">Goudemand et al., 2012b</bibRefCitation>
), Jiarong section,
<collectingRegion id="49E662E9647DFFA5FC6EE87BD472FA67" box="[994,1091,1387,1411]" country="China" name="Guizhou" pageId="20" pageNumber="21">Guizhou</collectingRegion>
(
<bibRefCitation id="EFB3D1FA647DFFA5FBDFE87BD520FA67" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[1107,1297,1387,1411]" pageId="20" pageNumber="21" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
): Southwest
<collectingCountry id="F335EC9B647DFFA5FCA1E89CD35AFA47" box="[813,875,1420,1443]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647DFFA5FCF5E89BD457FA47" author="Maekawa, T. &amp; Komatsu, T. &amp; Koike, T." box="[889,1126,1419,1443]" pageId="20" pageNumber="21" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
): Boreal region;
<taxonomicName id="4C22D788647DFFA5FAADE89BD397FA27" authority="zone" authorityName="Zone" class="Cephalopoda" family="Hedenstroemiidae" genus="Hedenstroemia" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647DFFA5FAADE89BD355FA27" italics="true" pageId="20" pageNumber="21">Hedenstroemia</emphasis>
zone
</taxonomicName>
, Siberia (
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),
<emphasis id="B9567019647DFFA5FB35E8BBD536FA27" box="[1209,1287,1451,1475]" italics="true" pageId="20" pageNumber="21">Tardus</emphasis>
zone, Canadian Arctic (
<bibRefCitation id="EFB3D1FA647DFFA5FC5DE8DBD4B1FA07" author="Orchard, M. J." box="[977,1152,1483,1507]" pageId="20" pageNumber="21" pagination="393 - 412" refId="ref45653" refString="Orchard, M. J. (2008). Lower Triassic conodonts from the Canadian Arctic, their intercalibration with ammonoid-based stages and a comparison with other North American Olenekian faunas. Polar Research, 27 (3), 393 - 412. https: // doi. org / 10.1111 / j. 1751 - 8369.2008.00072. x" type="journal article" year="2008">Orchard, 2008</bibRefCitation>
), Spitsbergen (
<bibRefCitation id="EFB3D1FA647DFFA5FACCE8DBD394F9E7" author="Nakrem, H. A. &amp; Orchard, M. J. &amp; Weitschat, W. &amp; Hounslow, M. W. &amp; Beatty, T. W. &amp; MOrk, A." pageId="20" pageNumber="21" pagination="523 - 539" refId="ref45434" refString="Nakrem, H. A., Orchard, M. J., Weitschat, W., Hounslow, M. W., Beatty, T. W., &amp; MOrk, A. (2008). Triassic conodonts from Svalbard and their Boreal correlations. Polar Research, 27 (3), 523 - 539. https: // doi. org / 10.1111 / j. 1751 - 8369.2008.00076. x" type="journal article" year="2008">Nakrem et al., 2008</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA647DFFA5FC3EE8FBD4C1F9E7" author="Weitschat, W. &amp; Lehmann, U." box="[946,1264,1515,1539]" pageId="20" pageNumber="21" pagination="85 - 100" refId="ref48068" refString="Weitschat, W., &amp; Lehmann, U. (1978). Biostratigraphy of the uppermost part of the Smithian Stage (Lower Triassic) at the Botneheia, W-Spitsbergen. Mitteilungen Geologisch- Palaontologischen Institut Universitat Hamburg, Heft, 48, 85 - 100." type="journal article" year="1978">Weitschat &amp; Lehmann, 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,186 @@
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<subSubSection id="C338FF80647DFFA5FF1BEE3BD245FCA7" box="[151,628,811,835]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B9DAC0B647DFFA5FF1BEE3BD245FCA7" blockId="20.[151,689,779,835]" box="[151,628,811,835]" pageId="20" pageNumber="21">
<heading id="D0D51B67647DFFA5FF1BEE3BD245FCA7" box="[151,628,811,835]" fontSize="36" level="2" pageId="20" pageNumber="21" reason="3">
Genus
<taxonomicName id="4C22D788647DFFA5FF68EE3BD245FCA7" authority="Budurov &amp; Sudar 1994" authorityName="Budurov &amp; Sudar" authorityYear="1994" box="[228,628,811,835]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">BORINELLA Budurov &amp; Sudar 1994</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647DFFA5FF1BEE7BD2E1FC07" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8B9DAC0B647DFFA5FF1BEE7BD21BFC67" blockId="20.[151,720,875,995]" box="[151,554,875,899]" pageId="20" pageNumber="21">
1988
<taxonomicName id="4C22D788647DFFA5FF5EEE7BD1E6FC67" authority="Kozur" authorityName="Kozur" authorityYear="1988" box="[210,471,875,899]" genus="Pseudogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FF5EEE7BD1BFFC67" box="[210,398,875,899]" italics="true" pageId="20" pageNumber="21">Pseudogondolella</emphasis>
Kozur
</taxonomicName>
, p. 244.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEE9BD272FC47" blockId="20.[151,720,875,995]" box="[151,579,907,931]" pageId="20" pageNumber="21">
1993
<taxonomicName id="4C22D788647DFFA5FF5EEE9BD1CFFC47" authority="Budurov &amp; Sudar" authorityName="Budurov &amp; Sudar" authorityYear="1993" box="[210,510,907,931]" family="Gondolellidae" genus="Kozurella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FF5EEE9BD10AFC47" box="[210,315,907,931]" italics="true" pageId="20" pageNumber="21">Kozurella</emphasis>
Budurov &amp; Sudar
</taxonomicName>
, p. 24.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEEBBD2A0FC27" blockId="20.[151,720,875,995]" box="[151,657,939,963]" pageId="20" pageNumber="21">
*1994 (June)
<taxonomicName id="4C22D788647DFFA5FEA8EEBBD27DFC27" authority="Budurov &amp; Sudar" authorityName="Budurov &amp; Sudar" authorityYear="1994" box="[292,588,939,963]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FEA8EEBBD1B9FC27" box="[292,392,939,963]" italics="true" pageId="20" pageNumber="21">Borinella</emphasis>
Budurov &amp; Sudar
</taxonomicName>
, p. 30.
</paragraph>
<paragraph id="8B9DAC0B647DFFA5FF1BEEDBD2E1FC07" blockId="20.[151,720,875,995]" box="[151,720,971,995]" pageId="20" pageNumber="21">
1994 (September)
<taxonomicName id="4C22D788647DFFA5FED3EEDBD208FC07" authority="Kozur" authorityName="Kozur" authorityYear="1994" box="[351,569,971,995]" genus="Chengyuania" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FED3EEDBD1C1FC07" box="[351,496,971,995]" italics="true" pageId="20" pageNumber="21">Chengyuania</emphasis>
Kozur
</taxonomicName>
, pp. 529530.
</paragraph>
</subSubSection>
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<emphasis id="B9567019647DFFA5FF1BE91CD110FBC7" box="[151,289,1036,1059]" italics="true" pageId="20" pageNumber="21">
<typeStatus id="549912A9647DFFA5FF1BE91CD0FAFBC7" box="[151,203,1036,1059]" pageId="20" pageNumber="21">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C22D788647DFFA5FEABE91BD287FBC7" authority="Dagis, 1984" authorityName="Dagis" authorityYear="1984" box="[295,694,1035,1059]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="buurensis">
<emphasis id="B9567019647DFFA5FEABE91BD21EFBC7" box="[295,559,1035,1059]" italics="true" pageId="20" pageNumber="21">Neogondolella buurensis</emphasis>
<bibRefCitation id="EFB3D1FA647DFFA5FDB9E91BD287FBC7" author="Dagis, A. A." box="[565,694,1035,1059]" pageId="20" pageNumber="21" pagination="1 - 69" refId="ref39946" refString="Dagis, A. A. (1984). Early Triassic conodonts of northern Middle Siberia. Academy of Sciences of the USSR, Siberian Branch. Institute of Geology and Geophysics Transactions, 554, 1 - 69." type="journal article" year="1984">Dagis, 1984</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<paragraph id="8B9DAC0B647DFFA5FF1BE93CD0D5FB67" blockId="20.[151,775,1035,1315]" pageId="20" pageNumber="21">
<materialsCitation id="3B4AA656647DFFA5FF1BE93CD0D1FB67" country="Russia" county="Early Triassic" location="Buur River Basin" municipality="Siberia" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<emphasis id="B9567019647DFFA5FF1BE93CD1FCFBA7" box="[151,461,1067,1091]" italics="true" pageId="20" pageNumber="21">
<typeStatus id="549912A9647DFFA5FF1BE93CD0FAFBA7" box="[151,203,1068,1091]" pageId="20" pageNumber="21">Type</typeStatus>
stratum and locality.
</emphasis>
<location id="8EFDFAD0647DFFA5FE50E93CD296FBA7" LSID="urn:lsid:plazi:treatment:038B1D1D647DFFA5FF1BEE3BD27EFAC7:8EFDFAD0647DFFA5FE50E93CD296FBA7" box="[476,679,1067,1091]" country="Russia" county="Early Triassic" municipality="Siberia" name="Buur River Basin" pageId="20" pageNumber="21">
<location id="8EFDFAD0647DFFA5FE50E93CD26BFBA7" LSID="urn:lsid:plazi:treatment:038B1D1D647DFFA5FF1BEE3BD27EFAC7:8EFDFAD0647DFFA5FE50E93CD26BFBA7" box="[476,602,1067,1091]" country="Russia" county="Early Triassic" municipality="Siberia" name="Buur River" pageId="20" pageNumber="21">
Buur
<location id="8EFDFAD0647DFFA5FDADE93BD296FBA7" LSID="urn:lsid:plazi:treatment:038B1D1D647DFFA5FF1BEE3BD27EFAC7:8EFDFAD0647DFFA5FDADE93BD296FBA7" box="[545,679,1067,1091]" country="Russia" county="Early Triassic" municipality="Siberia" name="River Basin" pageId="20" pageNumber="21">River Basin</location>
</location>
</location>
,
<location id="8EFDFAD0647DFFA5FD36E93BD08DFB87" LSID="urn:lsid:plazi:treatment:038B1D1D647DFFA5FF1BEE3BD27EFAC7:8EFDFAD0647DFFA5FD36E93BD08DFB87" country="Russia" county="Early Triassic" municipality="Siberia" name="Northern" pageId="20" pageNumber="21">Northern</location>
middle
<collectingMunicipality id="6BF93671647DFFA5FE90E95BD15BFB87" box="[284,362,1099,1123]" pageId="20" pageNumber="21">Siberia</collectingMunicipality>
,
<collectingCounty id="62FCD487647DFFA5FEF4E95BD23FFB87" box="[376,526,1099,1123]" pageId="20" pageNumber="21">Early Triassic</collectingCounty>
<taxonomicName id="4C22D788647DFFA5FD95E95BD332FB87" authority="Zone" authorityName="Zone" box="[537,771,1099,1123]" class="Cephalopoda" family="Hedenstroemiidae" genus="Hedenstroemia" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647DFFA5FD95E95BD28CFB87" box="[537,701,1099,1123]" italics="true" pageId="20" pageNumber="21">Hedenstroemia</emphasis>
Zone
</taxonomicName>
,
<collectingCountry id="F335EC9B647DFFA5FF1BE97BD0D1FB67" box="[151,224,1131,1155]" name="Russia" pageId="20" pageNumber="21">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647DFFA5FF1BE99BD27EFAC7" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B9DAC0B647DFFA5FF1BE99BD27EFAC7" blockId="20.[151,775,1035,1315]" pageId="20" pageNumber="21">
<emphasis id="B9567019647DFFA5FF1BE99BD0CDFB47" box="[151,252,1163,1187]" italics="true" pageId="20" pageNumber="21">Remarks.</emphasis>
Te multi-element apparatus of this genus appears to be essentially the same as that of
<taxonomicName id="4C22D788647DFFA5FD21E9BCD1A6FB07" authority="(Orchard, 2008)" baseAuthorityName="Orchard" baseAuthorityYear="2008" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9567019647DFFA5FD21E9BCD0D0FB07" italics="true" pageId="20" pageNumber="21">Neogondolella</emphasis>
(
<bibRefCitation id="EFB3D1FA647DFFA5FF63E9DBD1BCFB07" author="Orchard, M. J." box="[239,397,1227,1251]" pageId="20" pageNumber="21" pagination="393 - 412" refId="ref45653" refString="Orchard, M. J. (2008). Lower Triassic conodonts from the Canadian Arctic, their intercalibration with ammonoid-based stages and a comparison with other North American Olenekian faunas. Polar Research, 27 (3), 393 - 412. https: // doi. org / 10.1111 / j. 1751 - 8369.2008.00072. x" type="journal article" year="2008">Orchard, 2008</bibRefCitation>
)
</taxonomicName>
. Te P
<subScript id="17A6AE4E647DFFA5FE6CE9C7D1D8FB0C" attach="left" box="[480,489,1239,1256]" fontSize="7" pageId="20" pageNumber="21">1</subScript>
elements of the species of this genus all have in common discrete blade-carinal denticles that lengthen towards the anterior.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="007A6C301DB0682C2F7C289204811147">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<subSubSection id="C338FF80647DFFA5FCA1EBFBD4AEF8E7" box="[813,1183,1771,1795]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B9DAC0B647DFFA5FCA1EBFBD4AEF8E7" blockId="20.[813,1183,1771,1827]" box="[813,1183,1771,1795]" pageId="20" pageNumber="21">
<heading id="D0D51B67647DFFA5FCA1EBFBD4AEF8E7" box="[813,1183,1771,1795]" fontSize="9" level="2" pageId="20" pageNumber="21" reason="8">
<taxonomicName id="4C22D788647DFFA5FCA1EBFBD4AEF8E7" authority="Sweet, 1970" authorityName="Sweet" authorityYear="1970" box="[813,1183,1771,1795]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647DFFA5FCA1EBFBD429F8E7" box="[813,1048,1771,1795]" italics="true" pageId="20" pageNumber="21">Neospathodus dieneri</emphasis>
<bibRefCitation id="EFB3D1FA647DFFA5FB92EBFBD4AEF8E7" author="Sweet, W. C." box="[1054,1183,1771,1795]" pageId="20" pageNumber="21" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
</taxonomicName>
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<subSubSection id="C338FF80647DFFA5FCA1EA1BD3BAF8C7" box="[813,907,1803,1827]" pageId="20" pageNumber="21" type="description">
<paragraph id="8B9DAC0B647DFFA5FCA1EA1BD3BAF8C7" blockId="20.[813,1183,1771,1827]" box="[813,907,1803,1827]" pageId="20" pageNumber="21">
<figureCitation id="1319B08E647DFFA5FCA1EA1BD3BAF8C7" box="[813,907,1803,1827]" captionStart="Fig" captionStartId="21.[169,200,1283,1303]" captionTargetBox="[153,1438,238,1422]" captionTargetId="figure-168@21.[229,1363,254,1271]" captionTargetPageId="21" captionText="Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181" figureDoi="http://doi.org/10.5281/zenodo.12000270" httpUri="https://zenodo.org/record/12000270/files/figure.png" pageId="20" pageNumber="21">Fig. 14H</figureCitation>
</paragraph>
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<footnote id="E839B005647DFFA5FCA1EA5BD3F4F867" pageId="20" pageNumber="21">
<paragraph id="8B9DAC0B647DFFA5FCA1EA5BD3F4F867" blockId="20.[813,1436,1867,1923]" pageId="20" pageNumber="21">
*1970
<taxonomicName id="4C22D788647DFFA5FCF9EA5BD455F887" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[885,1124,1867,1891]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="dieneri" status="sp. nov.">
<emphasis id="B9567019647DFFA5FCF9EA5BD455F887" box="[885,1124,1867,1891]" italics="true" pageId="20" pageNumber="21">Neospathodus dieneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647DFFA5FBE2EA5CD49DF887" box="[1134,1196,1868,1891]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
; Sweet, pp. 249251, pl. 1, figs. 1, 4.
</paragraph>
</footnote>
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<paragraph id="8B9DAC0B647CFFA4FF25E813D57BFA83" blockId="21.[169,1405,1282,1410]" pageId="21" pageNumber="22">
<emphasis id="B9567019647CFFA4FF25E813D0D9FAF3" bold="true" box="[169,232,1283,1303]" pageId="21" pageNumber="22">Fig. 14</emphasis>
<taxonomicName id="4C22D788647CFFA4FF7EE813D1BDFAF3" authorityName="Hirsch" authorityYear="1994" box="[242,396,1283,1303]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="21" phylum="Chordata" rank="subFamily" subFamily="Neogondolellinae">Neogondolellinae</taxonomicName>
,
<taxonomicName id="4C22D788647CFFA4FE1FE813D215FAF3" authorityName="Orchard" authorityYear="2005" box="[403,548,1283,1303]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Novispathodinae">Novispathodinae</taxonomicName>
and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E647CFFA4FEC5E838D161FAD2" attach="left" box="[329,336,1320,1334]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B9567019647CFFA4FD60E80ED2CBFAD6" bold="true" box="[748,762,1310,1330]" pageId="21" pageNumber="22">A</emphasis>
,
<taxonomicName id="4C22D788647CFFA4FC8EE80ED412FAD6" authority="(Staesche)" baseAuthorityName="Staesche" box="[770,1059,1310,1330]" family="Gondolellidae" genus="Eurygnathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="costatus">
<emphasis id="B9567019647CFFA4FC8EE80ED3F7FAD6" box="[770,966,1310,1330]" italics="true" pageId="21" pageNumber="22">Eurygnathodus costatus</emphasis>
(Staesche)
</taxonomicName>
; SHA325, PIMUZ 39130.
<emphasis id="B9567019647CFFA4FB7CE80ED4CDFAD6" bold="true" box="[1264,1276,1310,1330]" pageId="21" pageNumber="22">B</emphasis>
<taxonomicName id="4C22D788647CFFA4FA8DE80ED105FAA9" authority="(Koike)" baseAuthorityName="Koike" baseAuthorityYear="1982" family="Gondolellidae" genus="Eurygnathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="hamadai">
<emphasis id="B9567019647CFFA4FA8DE80ED0C5FAA9" italics="true" pageId="21" pageNumber="22">Eurygnathodus hamadai</emphasis>
(Koike)
</taxonomicName>
; SHA326, PIMUZ 39131.
<emphasis id="B9567019647CFFA4FD8CE829D23CFAA9" bold="true" box="[512,525,1337,1357]" pageId="21" pageNumber="22">C</emphasis>
,
<emphasis id="B9567019647CFFA4FD99E829D215FAA9" bold="true" box="[533,548,1337,1357]" pageId="21" pageNumber="22">D</emphasis>
,
<emphasis id="B9567019647CFFA4FDA0E829D206FAA9" bold="true" box="[556,567,1337,1357]" pageId="21" pageNumber="22">F</emphasis>
,
<emphasis id="B9567019647CFFA4FDB3E829D27CFAA9" bold="true" box="[575,589,1337,1357]" pageId="21" pageNumber="22">G</emphasis>
<taxonomicName id="4C22D788647CFFA4FDDEE829D2FCFAA9" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[594,717,1337,1357]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B9567019647CFFA4FDDEE829D2FCFAA9" box="[594,717,1337,1357]" italics="true" pageId="21" pageNumber="22">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788647CFFA4FC8FE829D3A1FAA9" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[771,912,1337,1357]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019647CFFA4FC8FE829D37BFAA9" box="[771,842,1337,1357]" italics="true" pageId="21" pageNumber="22">waageni</emphasis>
(Sweet)
</taxonomicName>
;
<emphasis id="B9567019647CFFA4FC1BE829D395FAA9" bold="true" box="[919,932,1337,1357]" pageId="21" pageNumber="22">C</emphasis>
SHA328,PIMUZ 39238;
<emphasis id="B9567019647CFFA4FBE1E829D44DFAA9" bold="true" box="[1133,1148,1337,1357]" pageId="21" pageNumber="22">D</emphasis>
SHA328, PIMUZ 39239;
<emphasis id="B9567019647CFFA4FACAE829D560FAA9" bold="true" box="[1350,1361,1337,1357]" pageId="21" pageNumber="22">F</emphasis>
SHA325, PIMUZ 39240;
<emphasis id="B9567019647CFFA4FEE2E843D14DFA83" bold="true" box="[366,380,1363,1383]" pageId="21" pageNumber="22">G</emphasis>
SHA326, PIMUZ 39241.
<emphasis id="B9567019647CFFA4FDCAE843D260FA83" bold="true" box="[582,593,1363,1383]" pageId="21" pageNumber="22">E</emphasis>
<taxonomicName id="4C22D788647CFFA4FDDAE843D2FDFA83" authorityName="Mosher" authorityYear="1968" box="[598,716,1363,1383]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="B9567019647CFFA4FDDAE843D2FDFA83" box="[598,716,1363,1383]" italics="true" pageId="21" pageNumber="22">Neospathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788647CFFA4FC8FE843D38BFA83" authority="(Huckriede)" baseAuthorityName="Huckriede" baseAuthorityYear="1958" box="[771,954,1363,1383]" family="Prioniodontidae" genus="Spathognathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647CFFA4FC8FE843D360FA83" box="[771,849,1363,1383]" italics="true" pageId="21" pageNumber="22">cristagalli</emphasis>
(Huckriede)
</taxonomicName>
; SHA330, PIMUZ 39182.
<emphasis id="B9567019647CFFA4FB09E843D4A2FA83" bold="true" box="[1157,1171,1363,1383]" pageId="21" pageNumber="22">H</emphasis>
<taxonomicName id="4C22D788647CFFA4FB14E843D57BFA83" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1176,1354,1363,1383]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FB14E843D57BFA83" box="[1176,1354,1363,1383]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF25E87ED19FFA66" blockId="21.[169,1405,1282,1410]" box="[169,430,1390,1410]" pageId="21" pageNumber="22">(Sweet); SHA338, PIMUZ 39181</paragraph>
</caption>
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<paragraph id="8B9DAC0B647CFFA4FF1BE8F0D177F9FC" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB328B25647CFFA4FF1BE8F0D177F9FC" pageId="21" pageNumber="22">
1982
<taxonomicName id="4C22D788647CFFA4FF58E8F0D222FA1C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[212,531,1504,1528]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF58E8F0D1F7FA1C" box="[212,454,1504,1528]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
;
<treatmentCitation id="0A838A1A647CFFA4FD92E8F0D333FA1C" author="Matsuda, T." box="[542,770,1504,1528]" page="90" pageId="21" pageNumber="22" year="1982">
<bibRefCitation id="EFB3D1FA647CFFA4FD92E8F0D28EFA1C" author="Matsuda, T." box="[542,703,1504,1528]" pageId="21" pageNumber="22" pagination="87 - 102" refId="ref44946" refString="Matsuda, T. (1982). Early triassic conodonts from Kashmir, India part 2: Neospathodus 1. Journal of Geosciences Osaka City University, 25, 87 - 102." type="journal article" year="1982">Matsuda 1982</bibRefCitation>
, p. 90
</treatmentCitation>
, pl. 2, figs. 111.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF1BEB30D126F9B3" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
1982
<taxonomicName id="4C22D788647CFFA4FF54EB30D22EF9DC" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[216,543,1568,1592]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF54EB30D1FEF9DC" box="[216,463,1568,1592]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Koike, p. 37, pl. 6, figs. 1521.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF1BEB70D0CCF97C" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
1984
<taxonomicName id="4C22D788647CFFA4FF54EB70D22EF99C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[216,543,1632,1656]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF54EB70D1FEF99C" box="[216,463,1632,1656]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Dagis, p. 27, pl. 6, figs. 47.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF1BEBB0D120F93C" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
1991
<taxonomicName id="4C22D788647CFFA4FF54EBB0D22FF95C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[216,542,1696,1720]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF54EBB0D1FFF95C" box="[216,462,1696,1720]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Beyers &amp; Orchard, pl. 5, fig. 4.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF1BEBF0D104F8F3" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
2007
<taxonomicName id="4C22D788647CFFA4FF5EEBF0D239F91C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[210,520,1760,1784]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF5EEBF0D18FF91C" box="[210,446,1760,1784]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Orchard &amp; Krystyn, p. 33, figs. 3, 6, 7.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FF1BEA30D252F8BC" blockId="21.[151,775,1504,1880]" pageId="21" pageNumber="22">
2007
<taxonomicName id="4C22D788647CFFA4FF5BEA30D227F8DC" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[215,534,1824,1848]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FF5BEA30D1F9F8DC" box="[215,456,1824,1848]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Zhao &amp; Orchard in Zhao et al., p. 35, pl. 1, figs. 9AC, 12AB.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FCA1E8F0D486F9FC" blockId="21.[813,1437,1504,1784]" pageId="21" pageNumber="22">
2009
<taxonomicName id="4C22D788647CFFA4FCE4E8F0D4AFFA1C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[872,1182,1504,1528]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FCE4E8F0D465FA1C" box="[872,1108,1504,1528]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Igo in Shigeta et al., p. 186, figs. 151.6151.16, 152.8 (only).
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FCA1EB30D4F7F9BC" blockId="21.[813,1437,1504,1784]" pageId="21" pageNumber="22">
2009
<taxonomicName id="4C22D788647CFFA4FCFCEB0FD487F9DC" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[880,1206,1567,1592]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FCFCEB0FD455F9D3" box="[880,1124,1567,1591]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Orchard &amp; Zonneveld., pp. 782784, fig. 14, parts 14.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FCA1EB70D3F2F97C" blockId="21.[813,1437,1504,1784]" pageId="21" pageNumber="22">
2016
<taxonomicName id="4C22D788647CFFA4FCEBEB70D4ADF99C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[871,1180,1632,1656]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FCEBEB70D462F99C" box="[871,1107,1632,1656]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Maekawa et al., p. 199, figs. 4.84.10.
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FCA1EBB0D475F91C" blockId="21.[813,1437,1504,1784]" pageId="21" pageNumber="22">
2018
<taxonomicName id="4C22D788647CFFA4FCE4EBB0D4AFF95C" authority="Sweet" authorityName="Sweet" authorityYear="1970" box="[872,1182,1696,1720]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647CFFA4FCE4EBB0D465F95C" box="[872,1108,1696,1720]" italics="true" pageId="21" pageNumber="22">Neospathodus dieneri</emphasis>
Sweet
</taxonomicName>
; Maekawa in Maekawa et al., pp. 2528, figs. 15.815.9, 15.1215.14, 15.18 15.19, 15.2115.28 (only).
</paragraph>
<paragraph id="8B9DAC0B647CFFA4FCA1EA30D469F8DC" blockId="21.[813,1437,1824,1912]" box="[813,1112,1824,1848]" pageId="21" pageNumber="22">
<emphasis id="B9567019647CFFA4FCA1EA30D3A5F8DC" box="[813,916,1824,1848]" italics="true" pageId="21" pageNumber="22">Material.</emphasis>
ca.
<specimenCount id="9D246782647CFFA4FC32EA30D464F8DC" box="[958,1109,1824,1848]" count="40" pageId="21" pageNumber="22" type="generic">40 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647CFFA7FCCEEA50D257FE87" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8B9DAC0B647CFFA7FCCEEA50D257FE87" blockId="21.[813,1437,1824,1912]" lastBlockId="22.[151,776,235,995]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B9567019647CFFA4FCCEEA50D382F8B3" box="[834,947,1856,1879]" italics="true" pageId="21" pageNumber="22">Diagnosis.</emphasis>
See
<bibRefCitation id="EFB3D1FA647CFFA4FC63EA50D4B0F8BC" author="Sweet, W. C." box="[1007,1153,1856,1880]" pageId="21" pageNumber="22" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet (1970)</bibRefCitation>
, pp. 249251. Te basal cavity is symmetrically rounded with a circular outline. Te posterior margin is upturned. Te denticles are round (subcircular in cross-section) with a pointy end, discrete and slightly recurved posteriorly. Basal groove runs from the basal pit to the anterior end.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647FFFA7FF21EC7BD260FC07" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8B9DAC0B647FFFA7FF21EC7BD1C8FCE7" blockId="22.[151,776,235,995]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FF21EC7BD125FE67" box="[173,276,363,387]" italics="true" pageId="22" pageNumber="23">Remarks.</emphasis>
<bibRefCitation id="EFB3D1FA647FFFA7FE97EC7BD1D4FE67" author="Zhao, L. &amp; Orchard, M. J. &amp; Tong, J. &amp; Sun, Z. &amp; Zuo, J. &amp; Zhang, S. &amp; Yun, A." box="[283,485,363,387]" pageId="22" pageNumber="23" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al. (2007)</bibRefCitation>
distinguished three morphotypes of
<taxonomicName id="4C22D788647FFFA7FEA5EC9BD211FE47" authorityName="Sweet" authorityYear="1970" box="[297,544,395,419]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647FFFA7FEA5EC9BD211FE47" box="[297,544,395,419]" italics="true" pageId="22" pageNumber="23">Neospathodus dieneri</emphasis>
</taxonomicName>
on the basis of the length of the terminal cusp relative to the other denticles. Teir illustrated morphotype 3 (
<bibRefCitation id="EFB3D1FA647FFFA7FDCCECDBD330FE07" author="Zhao, L. &amp; Orchard, M. J. &amp; Tong, J. &amp; Sun, Z. &amp; Zuo, J. &amp; Zhang, S. &amp; Yun, A." box="[576,769,459,483]" pageId="22" pageNumber="23" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al., 2007</bibRefCitation>
,
<figureCitation id="1319B08E647FFFA7FF1BECFBD128FDE7" box="[151,281,491,515]" captionStart="Fig" captionStartId="16.[169,200,1273,1293]" captionTargetBox="[153,1438,238,1330]" captionTargetId="figure-295@16.[285,1305,254,1260]" captionTargetPageId="16" captionText="Fig. 11 Faunal turnover rate for all the relying conodont UAZs from South China during the Smithian and Spathian interval. Calculated from the optimal solution given in Fig. 8. Note the early Smithian and early Spathian radiation and the late Smithian and middle/late Spathian extinction" figureDoi="http://doi.org/10.5281/zenodo.12000230" httpUri="https://zenodo.org/record/12000230/files/figure.png" pageId="22" pageNumber="23">Fig. 11AC</figureCitation>
) however does not, in our opinion, belongs to
<emphasis id="B9567019647FFFA7FF36EF1CD0FCFDC7" box="[186,205,524,547]" italics="true" pageId="22" pageNumber="23">N</emphasis>
. ex gr.
<taxonomicName id="4C22D788647FFFA7FEA0EF1BD14BFDC7" authorityName="Sweet" authorityYear="1970" box="[300,378,523,547]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647FFFA7FEA0EF1BD14BFDC7" box="[300,378,523,547]" italics="true" pageId="22" pageNumber="23">dieneri</emphasis>
</taxonomicName>
but is more closely related to
<taxonomicName id="4C22D788647FFFA7FD62EF1CD137FDA7" baseAuthorityName="Huckriede" baseAuthorityYear="1958" family="Prioniodontidae" genus="Spathognathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FD62EF1CD137FDA7" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
</taxonomicName>
on the basis of its laterally flattened denticles and S-shaped posterior margin. Maekawa and Igo (2014, in
<bibRefCitation id="EFB3D1FA647FFFA7FE8FEF7BD1E8FD67" author="Shigeta, Y. &amp; Komatsu, T. &amp; Maekawa, T. &amp; Huyen, D. T." box="[259,473,619,643]" pageId="22" pageNumber="23" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
) assigned specimens to
<taxonomicName id="4C22D788647FFFA7FD62EF7CD0D4FD47" authorityName="Sweet" authorityYear="1970" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647FFFA7FD62EF7CD0D4FD47" italics="true" pageId="22" pageNumber="23">N. dieneri</emphasis>
</taxonomicName>
which is laterally flattened, blade-shaped denticles look very different from the
<typeStatus id="549912A9647FFFA7FDAAEFBBD2BDFD27" box="[550,652,683,707]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
. Tey may instead belong to another species or even to another genus (possibly to
<taxonomicName id="4C22D788647FFFA7FEEBEFFBD1EEFCE7" authorityName="Orchard" authorityYear="2005" box="[359,479,747,771]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B9567019647FFFA7FEEBEFFBD1EEFCE7" box="[359,479,747,771]" italics="true" pageId="22" pageNumber="23">Discretella</emphasis>
</taxonomicName>
?).
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF21EE1CD260FC07" blockId="22.[151,776,235,995]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FF21EE1CD11AFCC7" box="[173,299,780,803]" italics="true" pageId="22" pageNumber="23">Occurrence</emphasis>
. Tis species has been reported worldwide from the Dienerian and early Smithian. Tis includes South
<collectingCountry id="F335EC9B647FFFA7FEC9EE5BD1B9FC87" box="[325,392,843,867]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FE1BEE5BD262FC87" author="Zhao, L. &amp; Orchard, M. J. &amp; Tong, J. &amp; Sun, Z. &amp; Zuo, J. &amp; Zhang, S. &amp; Yun, A." box="[407,595,843,867]" pageId="22" pageNumber="23" pagination="24 - 38" refId="ref49075" refString="Zhao, L., Orchard, M. J., Tong, J., Sun, Z., Zuo, J., Zhang, S., &amp; Yun, A. (2007). Lower Triassic conodont sequence in Chaohu, Anhui Province, China and its global correlation. Palaeogeography, Palaeoclimatology, Palaeoecology, 252, 24 - 38." type="journal article" year="2007">Zhao et al., 2007</bibRefCitation>
and this study),
<collectingCountry id="F335EC9B647FFFA7FF1BEE7BD0CBFC67" box="[151,250,875,899]" name="Malaysia" pageId="22" pageNumber="23">Malaysia</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FE83EE7BD1AAFC67" author="Koike, T." box="[271,411,875,899]" pageId="22" pageNumber="23" pagination="9 - 51" refId="ref42702" refString="Koike, T. (1982). Triassic conodont biostratigraphy in Kedah, West Malaysia. Contribution to the geology and palaeontology of Southeast Asia, 220. Geology and Palaeontology of Southeast Asia, 23, 9 - 51." type="journal article" year="1982">Koike, 1982</bibRefCitation>
), the Northern Indian margin (
<bibRefCitation id="EFB3D1FA647FFFA7FF13EE9BD179FC47" author="Matsuda, T." box="[159,328,907,931]" pageId="22" pageNumber="23" pagination="87 - 102" refId="ref44946" refString="Matsuda, T. (1982). Early triassic conodonts from Kashmir, India part 2: Neospathodus 1. Journal of Geosciences Osaka City University, 25, 87 - 102." type="journal article" year="1982">Matsuda, 1982</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA647FFFA7FEDBEE9BD1EEFC47" author="Sweet, W. C." box="[343,479,907,931]" pageId="22" pageNumber="23" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
), South Primorye in
<collectingCountry id="F335EC9B647FFFA7FD5FEE9CD084FC27" name="Russia" pageId="22" pageNumber="23">Russia</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FF44EEBBD19BFC27" author="Shigeta, Y. &amp; Zakharov, Y. D. &amp; Maeda, H. &amp; Popov, A. M." box="[200,426,939,963]" pageId="22" pageNumber="23" refId="ref46464" refString="Shigeta, Y., Zakharov, Y. D., Maeda, H., &amp; Popov, A. M. (Eds.). 2009. The Lower Triassic System in the Abrek Bay Area, South Primorye, Russia" type="book" year="2009">Shigeta et al., 2009</bibRefCitation>
),
<collectingCountry id="F335EC9B647FFFA7FE48EEBBD229FC27" box="[452,536,939,963]" name="Canada" pageId="22" pageNumber="23">Canada</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FDA7EEBBD0FCFC07" author="Beyers, J. M. &amp; Orchard, M. J." pageId="22" pageNumber="23" refId="ref38524" refString="Beyers, J. M., &amp; Orchard, M. J. (1991). Ordovician to Triassic conodont paleontology of the Canadian Cordillera. Ottawa: Geological Survey of Canada." type="book" year="1991">Beyers &amp; Orchard, 1991</bibRefCitation>
) and
<collectingCountry id="F335EC9B647FFFA7FE81EEDCD17AFC07" box="[269,331,972,995]" name="Japan" pageId="22" pageNumber="23">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FED5EEDBD272FC07" author="Maekawa, T. &amp; Komatsu, T. &amp; Koike, T." box="[345,579,971,995]" pageId="22" pageNumber="23" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,216 @@
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<mods:title id="853DF4EF47174D449C638D7202C72687">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="9846959633B1AA8E91301E5ABF55E674">Bagherpour, Borhan</mods:namePart>
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<mods:title id="B62E2B94E962235FC9CCFBBD8B036E1A">Swiss Journal of Palaeontology</mods:title>
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<mods:part id="9BBA7EF4E561BE73214FF3F654F7557C">
<mods:date id="D6FD9C8E5839988F6B614FBF2F52C255">2022</mods:date>
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<mods:number id="C3C7454CD70999012F0D0A07292507AB">141</mods:number>
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<treatment id="038B1D1D647FFFA7FF1BE91BD543FDA7" LSID="urn:lsid:plazi:treatment:038B1D1D647FFFA7FF1BE91BD543FDA7" httpUri="http://treatment.plazi.org/id/038B1D1D647FFFA7FF1BE91BD543FDA7" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C338FF80647FFFA7FF1BE91BD2ABFBC7" box="[151,666,1035,1060]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8B9DAC0B647FFFA7FF1BE91BD2ABFBC7" blockId="22.[151,666,1035,1091]" box="[151,666,1035,1060]" pageId="22" pageNumber="23">
<heading id="D0D51B67647FFFA7FF1BE91BD2ABFBC7" box="[151,666,1035,1060]" fontSize="9" level="2" pageId="22" pageNumber="23" reason="8">
<taxonomicName id="4C22D788647FFFA7FF1BE91BD2ABFBC7" authority="Huckriede, 1958" authorityName="Mosher" authorityYear="1968" box="[151,666,1035,1060]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B9567019647FFFA7FF1BE91BD101FBC7" box="[151,304,1035,1059]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
ex gr.
<emphasis id="B9567019647FFFA7FEFAE91BD1D3FBC7" box="[374,482,1035,1059]" italics="true" pageId="22" pageNumber="23">cristagalli</emphasis>
<bibRefCitation id="EFB3D1FA647FFFA7FE64E91BD2ABFBC7" author="Huckriede, R." box="[488,666,1035,1059]" pageId="22" pageNumber="23" pagination="141 - 175" refId="ref42031" refString="Huckriede, R. (1958). Die Conodonten der Mediterranen Trias und ihr stratigraphischer Wert. Palaontologische Zeitschrift, 32, 141 - 175." type="journal article" year="1958">Huckriede, 1958</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647FFFA7FF1BE93BD543FDA7" pageId="22" pageNumber="23" type="description">
<paragraph id="8B9DAC0B647FFFA7FF1BE93BD0C0FBA7" blockId="22.[151,666,1035,1091]" box="[151,241,1067,1091]" pageId="22" pageNumber="23">
<figureCitation id="1319B08E647FFFA7FF1BE93BD0C0FBA7" box="[151,241,1067,1091]" captionStart="Fig" captionStartId="21.[169,200,1283,1303]" captionTargetBox="[153,1438,238,1422]" captionTargetId="figure-168@21.[229,1363,254,1271]" captionTargetPageId="21" captionText="Fig. 14 Neogondolellinae, Novispathodinae and uncertain from Shanggang. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, Eurygnathodus costatus (Staesche); SHA325, PIMUZ 39130. B Eurygnathodus hamadai (Koike); SHA326, PIMUZ 39131. C, D, F, G Novispathodus ex gr. waageni (Sweet); C SHA328,PIMUZ 39238; D SHA328, PIMUZ 39239; F SHA325, PIMUZ 39240;G SHA326, PIMUZ 39241.E Neospathodus ex gr. cristagalli (Huckriede); SHA330, PIMUZ 39182.H Neospathodus dieneri (Sweet); SHA338, PIMUZ 39181" figureDoi="http://doi.org/10.5281/zenodo.12000270" httpUri="https://zenodo.org/record/12000270/files/figure.png" pageId="22" pageNumber="23">Fig. 14E</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE97BD17BFB47" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
1970
<taxonomicName id="4C22D788647FFFA7FF59E97BD26EFB67" authority="Huckriede" authorityName="Huckriede" box="[213,607,1131,1155]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF59E97BD1D2FB67" box="[213,483,1131,1155]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Sweet, p. 346, pl. 1, figs. 14, 15.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE9BBD170FB07" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
1982
<taxonomicName id="4C22D788647FFFA7FF5FE9BBD26BFB27" authority="Huckriede" authorityName="Huckriede" box="[211,602,1195,1219]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF5FE9BBD1EEFB27" box="[211,479,1195,1219]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Matsuda p. 92, pl. 3, figs. 112.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE9FBD123FAC7" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
2005
<taxonomicName id="4C22D788647FFFA7FF59E9FBD260FAE7" authority="Sweet" authorityName="Sweet" box="[213,593,1259,1283]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF59E9FBD15FFAE7" box="[213,366,1259,1283]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
cf.
<emphasis id="B9567019647FFFA7FE15E9FBD234FAE7" box="[409,517,1259,1283]" italics="true" pageId="22" pageNumber="23">cristagalli</emphasis>
Sweet
</taxonomicName>
; Orchard, p. 89, text-fig. 14.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE83BD2A6FA87" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
2014
<taxonomicName id="4C22D788647FFFA7FF55E83BD25CFAA7" authority="Huckriede" authorityName="Huckriede" box="[217,621,1323,1347]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF55E83BD1DDFAA7" box="[217,492,1323,1347]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., p. 223, figs. 161.10161.12.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE87BD22AFA47" blockId="22.[151,775,1131,1443]" pageId="22" pageNumber="23">
2015
<taxonomicName id="4C22D788647FFFA7FF55E87BD25DFA67" authority="Huckriede" authorityName="Huckriede" box="[217,620,1387,1411]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF55E87BD1DAFA67" box="[217,491,1387,1411]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Maekawa in Maekawa et al., p. 315, figs. 5.45.6.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE8DBD1F2FA07" blockId="22.[151,775,1483,1891]" box="[151,451,1483,1507]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FF1BE8DBD0CEFA07" box="[151,255,1483,1507]" italics="true" pageId="22" pageNumber="23">Material.</emphasis>
ca.
<specimenCount id="9D246782647FFFA7FEA5E8DBD18EFA07" box="[297,447,1483,1507]" count="30" pageId="22" pageNumber="23" type="generic">30 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FF1BE8FBD111F887" blockId="22.[151,775,1483,1891]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FF1BE8FBD0CDF9E7" box="[151,252,1515,1539]" italics="true" pageId="22" pageNumber="23">Remarks.</emphasis>
In comparison with the
<typeStatus id="549912A9647FFFA7FDA8E8FBD2B5F9E7" box="[548,644,1515,1539]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C22D788647FFFA7FD3EE8FCD0EDF9C7" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FD3EE8FCD0EDF9C7" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA647FFFA7FF7DEB1BD19DF9C7" author="Huckriede, R." box="[241,428,1547,1571]" pageId="22" pageNumber="23" pagination="141 - 175" refId="ref42031" refString="Huckriede, R. (1958). Die Conodonten der Mediterranen Trias und ihr stratigraphischer Wert. Palaontologische Zeitschrift, 32, 141 - 175." type="journal article" year="1958">Huckriede, 1958</bibRefCitation>
, Pl. 10, fig. 15), this element is much shorter, the basal cavity is oval, posteriorly rounded and only partly inverted, and the posterior triangular cusp is not conspicuously separated from the other denticles. Sweet still included such forms within
<taxonomicName id="4C22D788647FFFA7FF1BEBBCD112F927" box="[151,291,1707,1731]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FF1BEBBCD112F927" box="[151,291,1707,1731]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
</taxonomicName>
, but excluded similar forms with a rounded basal cavity and rounded, pointy denticles and assigned them to the then new species
<taxonomicName id="4C22D788647FFFA7FE69EBFCD263F8E7" authorityName="Sweet" authorityYear="1970" box="[485,594,1771,1795]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="dieneri">
<emphasis id="B9567019647FFFA7FE69EBFCD263F8E7" box="[485,594,1771,1795]" italics="true" pageId="22" pageNumber="23">N. dieneri</emphasis>
</taxonomicName>
. In our opinion, such short elements where the cusp is not separated from the other denticles would deserve to be differentiated as a new species.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EDFCD543FDA7" blockId="22.[813,1437,235,579]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EDFCD399FEE7" box="[813,936,236,259]" italics="true" pageId="22" pageNumber="23">Occurrence</emphasis>
.
<taxonomicName id="4C22D788647FFFA7FC37EDFCD47FFEE7" box="[955,1102,235,259]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FC37EDFCD47FFEE7" box="[955,1102,235,259]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
</taxonomicName>
is known worldwide in the Dienerian and earliest Smithian. It is not clear yet whether the present form is younger than
<taxonomicName id="4C22D788647FFFA7FA82EC3CD5ACFEA7" box="[1294,1437,299,323]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FA82EC3CD5ACFEA7" box="[1294,1437,299,323]" italics="true" pageId="22" pageNumber="23">N. cristagalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647FFFA7FCA1EC5CD38DFE87" box="[813,956,331,355]" pageId="22" pageNumber="23" sensu="stricto">sensu stricto</taxonomicNameLabel>
and occurs only close to the Dienerian Smithian (InduanOlenekian) boundary: Toad Formation, British Columbia (
<bibRefCitation id="EFB3D1FA647FFFA7FBC3EC9BD4C9FE47" author="Orchard, M. J." box="[1103,1272,395,419]" pageId="22" pageNumber="23" pagination="73 - 101" refId="ref45568" refString="Orchard, M. J. (2005). Multielement conodont apparatuses of Triassic Gondolelloidea. Special Papers in Palaeontology, 73 (73), 73 - 101." type="journal article" year="2005">Orchard, 2005</bibRefCitation>
). Salt Range,
<collectingCountry id="F335EC9B647FFFA7FCA1ECBBD3B8FE27" box="[813,905,427,451]" name="Pakistan" pageId="22" pageNumber="23">Pakistan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA647FFFA7FC10ECBBD415FE27" author="Sweet, W. C." box="[924,1060,427,451]" pageId="22" pageNumber="23" refId="ref47176" refString="Sweet, W. C. (1970). Uppermost Permian and Lower Triassic conodonts of" type="book" year="1970">Sweet, 1970</bibRefCitation>
). Guryul Ravine, Kashmir (
<bibRefCitation id="EFB3D1FA647FFFA7FAEAECBBD395FE07" author="Matsuda, T." pageId="22" pageNumber="23" pagination="87 - 102" refId="ref44946" refString="Matsuda, T. (1982). Early triassic conodonts from Kashmir, India part 2: Neospathodus 1. Journal of Geosciences Osaka City University, 25, 87 - 102." type="journal article" year="1982">Matsuda, 1982</bibRefCitation>
). Nanpanjiang basin; north-eastern
<collectingCountry id="F335EC9B647FFFA7FAB3ECDBD5ADFE07" box="[1343,1436,459,483]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
<taxonomicName id="4C22D788647FFFA7FCA1ECFBD3D4FDE7" authority="beds" authorityName="Beds" box="[813,997,491,515]" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="22" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647FFFA7FCA1ECFBD398FDE7" box="[813,937,491,515]" italics="true" pageId="22" pageNumber="23">Flemingites</emphasis>
beds
</taxonomicName>
within the
<taxonomicName id="4C22D788647FFFA7FBE0ECFBD53FFDE7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1132,1294,491,515]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B9567019647FFFA7FBE0ECFBD53FFDE7" box="[1132,1294,491,515]" italics="true" pageId="22" pageNumber="23">Novispathodus</emphasis>
</taxonomicName>
ex. gr.
<taxonomicName id="4C22D788647FFFA7FAE9ECFCD3ACFDC7" authority="Zone" authorityName="Zone" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019647FFFA7FAE9ECFCD368FDC7" italics="true" pageId="22" pageNumber="23">waageni</emphasis>
Zone
</taxonomicName>
(
<bibRefCitation id="EFB3D1FA647FFFA7FC23EF1BD493FDC7" author="Maekawa, T. &amp; Komatsu, T. &amp; Shigeta, Y. &amp; Huyen, D. T. &amp; Tien, D. C." box="[943,1186,523,547]" pageId="22" pageNumber="23" pagination="312 - 320" refId="ref44838" refString="Maekawa, T., Komatsu, T., Shigeta, Y., Huyen, D. T., &amp; Tien, D. C. (2015). First occurrence of Early Triassic conodonts from the Lang Son Formation, northeastern Vietnam. Paleontological Research, 19 (4), 312 - 320. https: // doi. org / 10.2517 / 2015 PR 014" type="journal article" year="2015">Maekawa et al., 2015</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA647FFFA7FB38EF1BD5BCFDC7" author="Shigeta, Y. &amp; Komatsu, T. &amp; Maekawa, T. &amp; Huyen, D. T." box="[1204,1421,523,547]" pageId="22" pageNumber="23" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
). Luolou Formation,
<taxonomicName id="4C22D788647FFFA7FC72EF3BD44BFDA7" box="[1022,1146,555,579]" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647FFFA7FC72EF3BD44BFDA7" box="[1022,1146,555,579]" italics="true" pageId="22" pageNumber="23">Flemingites</emphasis>
</taxonomicName>
limestone (this study).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3B1F9669980065F80AD55FB4E7F6C982">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<paragraph id="8B9DAC0B647FFFA7FCA1EF7BD45BFD67" blockId="22.[813,1130,619,675]" box="[813,1130,619,643]" pageId="22" pageNumber="23">
<heading id="D0D51B67647FFFA7FCA1EF7BD45BFD67" box="[813,1130,619,643]" fontSize="9" level="2" pageId="22" pageNumber="23" reason="8">
<taxonomicName id="4C22D788647FFFA7FCA1EF7BD41AFD67" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[813,1067,619,643]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="bevelledi" status="sp. nov.">
<emphasis id="B9567019647FFFA7FCA1EF7BD41AFD67" box="[813,1067,619,643]" italics="true" pageId="22" pageNumber="23">Neospathodus bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647FFFA7FBBDEF7CD45BFD67" box="[1073,1130,620,643]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C338FF80647FFFA9FCA1EF9BD445FA27" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="8B9DAC0B647FFFA7FCA1EF9BD43CFD47" blockId="22.[813,1130,619,675]" box="[813,1037,651,675]" pageId="22" pageNumber="23">
<figureCitation id="1319B08E647FFFA7FCA1EF9BD383FD47" box="[813,946,651,675]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="22" pageNumber="23">Figs. 15A, D</figureCitation>
,
<figureCitation id="1319B08E647FFFA7FC31EF9BD438FD47" box="[957,1033,651,675]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.12000298" httpUri="https://zenodo.org/record/12000298/files/figure.png" pageId="22" pageNumber="23">16PR</figureCitation>
.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EFDBD473FCE7" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
1990
<taxonomicName id="4C22D788647FFFA7FCE4EFDBD44FFD07" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[872,1150,715,739]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pamirensis" status="sp. nov.">
<emphasis id="B9567019647FFFA7FCE4EFDBD44FFD07" box="[872,1150,715,739]" italics="true" pageId="22" pageNumber="23">Neospathodus pamirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647FFFA7FB09EFDCD4F1FD07" box="[1157,1216,716,739]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
; Dagis, p. 7980, pl. 4, fig. 8, pl. 6, fig. 9 (only).
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EE1BD359FCA7" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2007
<taxonomicName id="4C22D788647FFFA7FCE2EE1BD436FCC7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[878,1031,779,803]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B9567019647FFFA7FCE2EE1BD436FCC7" box="[878,1031,779,803]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647FFFA7FB9FEE1CD462FCC7" box="[1043,1107,780,803]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
<collectionCode id="ED3334CE647FFFA7FBD3EE1CD440FCC7" box="[1119,1137,780,803]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="23" type="Museum">V</collectionCode>
Orchard; Orchard, p. 96, fig. 2.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EE5BD3E1FC67" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2013
<taxonomicName id="4C22D788647FFFA7FCE0EE5BD4CAFC87" authority="Huckriede" authorityName="Huckriede" box="[876,1275,843,867]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FCE0EE5BD44CFC87" box="[876,1149,843,867]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Yan et al., p. 516, fig. 6, IL.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EE9BD407FBE7" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2014
<taxonomicName id="4C22D788647FFFA7FCE1EE9BD487FC47" authority="Goel" authorityName="Goel" authorityYear="1977" box="[877,1206,907,931]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B9567019647FFFA7FCE1EE9BD445FC47" box="[877,1140,907,931]" italics="true" pageId="22" pageNumber="23">Neospathodus spitiensis</emphasis>
Goel
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., p. 233236, figs. 167.17167.30, 168.7 168.9, 168.16168.27, 168.34168.36, 169.4169.9, 169.21169.32, 170.13.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E91BD4B6FBA7" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2014
<taxonomicName id="4C22D788647FFFA7FCE6E91BD41BFBC7" box="[874,1066,1035,1059]" family="Gondolellidae" genus="Neospathodus" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019647FFFA7FCE6E91BD432FBC7" box="[874,1027,1035,1059]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
sp.
</taxonomicName>
indet A Maekawa &amp; Igo; Shigeta et al., p. 179, fig. 170.10170.33.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E95BD3E3FB47" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2014
<taxonomicName id="4C22D788647FFFA7FCFDE95BD422FB87" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[881,1043,1099,1123]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="B9567019647FFFA7FCFDE95BD422FB87" box="[881,1043,1099,1123]" italics="true" pageId="22" pageNumber="23">Novispathodus</emphasis>
</taxonomicName>
ex gr.
<taxonomicName id="4C22D788647FFFA7FBF9E95CD510FB87" authority="Sweet" authorityName="Sweet" box="[1141,1313,1099,1123]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019647FFFA7FBF9E95CD4FFFB87" box="[1141,1230,1100,1123]" italics="true" pageId="22" pageNumber="23">waageni</emphasis>
Sweet
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., p. 244, figs. 179.4179.6, 179.13179.15.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E9BBD42EFB07" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2015
<taxonomicName id="4C22D788647FFFA7FCE6E9BBD4C5FB27" authority="Huckriede" authorityName="Huckriede" box="[874,1268,1195,1219]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FCE6E9BBD446FB27" box="[874,1143,1195,1219]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Chen et al., p. 112, fig. 8.1213, 8.15.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E9FBD387FAC7" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2016
<taxonomicName id="4C22D788647FFFA7FCE7E9FBD41CFAE7" box="[875,1069,1259,1283]" family="Gondolellidae" genus="Neospathodus" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019647FFFA7FCE7E9FBD435FAE7" box="[875,1028,1259,1283]" italics="true" pageId="22" pageNumber="23">Neospathodus</emphasis>
sp.
</taxonomicName>
indet A; Maekawa et al. p. 200, figs. 5.25.7.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E83BD4F2FA87" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2018
<taxonomicName id="4C22D788647FFFA7FCE2E83BD530FAA7" authority="Huckriede" authorityName="Huckriede" box="[878,1281,1323,1347]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B9567019647FFFA7FCE2E83BD4B1FAA7" box="[878,1152,1323,1347]" italics="true" pageId="22" pageNumber="23">Neospathodus cristagalli</emphasis>
Huckriede
</taxonomicName>
; Maekawa in Maekawa et al., p.22, fig. 14.24 (only).
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E87BD448FA47" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
?2019a
<taxonomicName id="4C22D788647FFFA7FC18E87BD3BAFA47" authority="Zhao &amp; Orchard" authorityName="Zhao &amp; Orchard" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pingdingshanensis">
<emphasis id="B9567019647FFFA7FC18E87BD526FA67" box="[916,1303,1387,1411]" italics="true" pageId="22" pageNumber="23">Novispathodus pingdingshanensis</emphasis>
Zhao &amp; Orchard
</taxonomicName>
; Wu et al. figs. 3234.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1E8BBD342FA07" blockId="22.[813,1437,715,1507]" pageId="22" pageNumber="23">
2021
<taxonomicName id="4C22D788647FFFA7FCE5E8BBD4F7FA27" authority="(Sweet)" baseAuthorityName="Sweet" baseAuthorityYear="1970" box="[873,1222,1451,1475]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B9567019647FFFA7FCE5E8BBD45AFA27" box="[873,1131,1451,1475]" italics="true" pageId="22" pageNumber="23">Novispathodus waageni</emphasis>
(Sweet)
</taxonomicName>
; Sun et al., fig. 5.22 (only).
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EB1BD4A8F987" blockId="22.[813,1437,1547,1859]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EB1BD3AFF9C7" box="[813,926,1547,1571]" italics="true" pageId="22" pageNumber="23">Etymology</emphasis>
: named after the Latin (and British English) word bevelled, which refers to the wedge-like cessation of the denticles of the P
<subScript id="17A6AE4E647FFFA7FBA7EB47D405F98C" attach="left" box="[1067,1076,1623,1640]" fontSize="7" pageId="22" pageNumber="23">1</subScript>
element.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EB7BD4C2F967" blockId="22.[813,1437,1547,1859]" box="[813,1267,1643,1667]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EB7BD3A5F967" box="[813,916,1643,1667]" italics="true" pageId="22" pageNumber="23">
<typeStatus id="549912A9647FFFA7FCA1EB7BD3BFF967" box="[813,910,1643,1667]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
:
</emphasis>
specimen illustrated in
<figureCitation id="1319B08E647FFFA7FB14EB7BD4C2F967" box="[1176,1267,1643,1667]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="22" pageNumber="23">Fig. 15A</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EB9CD4C2F947" blockId="22.[813,1437,1547,1859]" box="[813,1267,1675,1699]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EB9CD3A5F947" box="[813,916,1676,1699]" italics="true" pageId="22" pageNumber="23">
<typeStatus id="549912A9647FFFA7FCA1EB9CD3BEF947" box="[813,911,1676,1699]" pageId="22" pageNumber="23" type="paratype">Paratype</typeStatus>
:
</emphasis>
specimen illustrated in
<figureCitation id="1319B08E647FFFA7FB15EB9BD4C2F947" box="[1177,1267,1675,1699]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="22" pageNumber="23">Fig. 15D</figureCitation>
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EBBCD474F907" blockId="22.[813,1437,1547,1859]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EBBCD3F4F927" box="[813,965,1707,1731]" italics="true" pageId="22" pageNumber="23">Type locality:</emphasis>
Shanggang road cut, Luolou formation,
<collectingRegion id="49E662E9647FFFA7FCA1EBDBD3C3F907" box="[813,1010,1739,1763]" country="China" name="Guangxi" pageId="22" pageNumber="23">Guangxi Province</collectingRegion>
,
<collectingCountry id="F335EC9B647FFFA7FC70EBDBD470F907" box="[1020,1089,1739,1763]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
.
</paragraph>
<paragraph id="8B9DAC0B647FFFA7FCA1EBFCD3DBF8A7" blockId="22.[813,1437,1547,1859]" pageId="22" pageNumber="23">
<emphasis id="B9567019647FFFA7FCA1EBFCD3A8F8E7" box="[813,921,1771,1795]" italics="true" pageId="22" pageNumber="23">Type level</emphasis>
: Luolou Formation, within the early-to-middle Smithian limestones (
<taxonomicName id="4C22D788647FFFA7FB95EA1CD44CF8C7" authorityName="Hyatt &amp; Smith" authorityYear="1905" box="[1049,1149,1804,1827]" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647FFFA7FB95EA1CD44CF8C7" box="[1049,1149,1804,1827]" italics="true" pageId="22" pageNumber="23">Owenites</emphasis>
</taxonomicName>
beds),
<quantity id="4CDA01EE647FFFA7FB47EA1BD520F8C7" box="[1227,1297,1803,1827]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="22" pageNumber="23" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">23 m</quantity>
above
<taxonomicName id="4C22D788647FFFA7FAD1EA1BD342F8A7" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
<emphasis id="B9567019647FFFA7FAD1EA1BD342F8A7" italics="true" pageId="22" pageNumber="23">Flemingites</emphasis>
</taxonomicName>
limestone.
</paragraph>
<caption id="DF5DFC83647EFFA6FF25EBE0D501F86D" ID-DOI="http://doi.org/10.5281/zenodo.12000286" ID-Zenodo-Dep="12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="23" pageNumber="24" startId="23.[169,200,1776,1796]" targetBox="[152,1438,238,1940]" targetPageId="23" targetType="figure">
<paragraph id="8B9DAC0B647EFFA6FF25EBE0D501F86D" blockId="23.[169,1423,1775,1929]" pageId="23" pageNumber="24">
<emphasis id="B9567019647EFFA6FF25EBE0D0D6F8E0" bold="true" box="[169,231,1776,1796]" pageId="23" pageNumber="24">Fig. 15</emphasis>
<taxonomicName id="4C22D788647EFFA6FF7DEBE0D1B8F8E0" authorityName="Hirsch" authorityYear="1994" box="[241,393,1776,1796]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="21" phylum="Chordata" rank="subFamily" subFamily="Neogondolellinae">Neogondolellinae</taxonomicName>
and
<taxonomicName id="4C22D788647EFFA6FE38EBE0D23FF8E0" authorityName="Orchard" authorityYear="2005" box="[436,526,1776,1796]" family="Gondolellidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="45" phylum="Chordata" rank="subFamily" subFamily="Mullerinae">Mullerinae</taxonomicName>
from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P
<subScript id="17A6AE4E647EFFA6FEC4EA04D17EF8C6" attach="left" box="[328,335,1812,1826]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
elements if not specifically identified otherwise.
<emphasis id="B9567019647EFFA6FD60EA1BD2CBF8FB" bold="true" box="[748,762,1803,1823]" pageId="23" pageNumber="24">A</emphasis>
,
<emphasis id="B9567019647EFFA6FC8EEA1BD320F8FB" bold="true" box="[770,785,1803,1823]" pageId="23" pageNumber="24">D</emphasis>
<taxonomicName id="4C22D788647EFFA6FC99EA1BD3E7F8FB" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[789,982,1803,1823]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="bevelledi" status="sp. nov.">
<emphasis id="B9567019647EFFA6FC99EA1BD3E7F8FB" box="[789,982,1803,1823]" italics="true" pageId="23" pageNumber="24">Neospathodus bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647EFFA6FC57EA1BD439F8FB" box="[987,1032,1803,1823]" pageId="23" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B9567019647EFFA6FB82EA1BD42DF8FB" bold="true" box="[1038,1052,1803,1823]" pageId="23" pageNumber="24">A</emphasis>
SHA333,PIMUZ 39176;
<emphasis id="B9567019647EFFA6FB69EA1BD4C5F8FB" bold="true" box="[1253,1268,1803,1823]" pageId="23" pageNumber="24">D</emphasis>
, SHA332, PIMUZ 39177.
<emphasis id="B9567019647EFFA6FF69EA35D0C0F8DD" bold="true" box="[229,241,1829,1849]" pageId="23" pageNumber="24">B</emphasis>
,
<emphasis id="B9567019647EFFA6FF76EA35D133F8DD" bold="true" box="[250,258,1829,1849]" pageId="23" pageNumber="24">J</emphasis>
<taxonomicName id="4C22D788647EFFA6FE8BEA35D18AF8DD" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[263,443,1829,1849]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B9567019647EFFA6FE8BEA35D16AF8DD" box="[263,347,1829,1849]" italics="true" pageId="23" pageNumber="24">Discretella</emphasis>
aff.
<emphasis id="B9567019647EFFA6FEF0EA35D18AF8DD" box="[380,443,1829,1849]" italics="true" pageId="23" pageNumber="24">discreta</emphasis>
</taxonomicName>
(MÜller);
<emphasis id="B9567019647EFFA6FD86EA35D227F8DD" bold="true" box="[522,534,1829,1849]" pageId="23" pageNumber="24">B</emphasis>
SHA304, PIMUZ 39104;
<emphasis id="B9567019647EFFA6FD6CEA35D2D9F8DD" bold="true" box="[736,744,1829,1849]" pageId="23" pageNumber="24">J</emphasis>
SHA334,PIMUZ 39105.
<emphasis id="B9567019647EFFA6FC3EEA35D38EF8DD" bold="true" box="[946,959,1829,1849]" pageId="23" pageNumber="24">C</emphasis>
,
<emphasis id="B9567019647EFFA6FC4BEA35D3E3F8DD" bold="true" box="[967,978,1829,1849]" pageId="23" pageNumber="24">L</emphasis>
<taxonomicName id="4C22D788647EFFA6FC5AEA35D484F8DD" authority="(MUller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[982,1205,1829,1849]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B9567019647EFFA6FC5AEA35D45CF8DD" box="[982,1133,1829,1849]" italics="true" pageId="23" pageNumber="24">Discretella discreta</emphasis>
(MÜller)
</taxonomicName>
;
<emphasis id="B9567019647EFFA6FB37EA35D4F9F8DD" bold="true" box="[1211,1224,1829,1849]" pageId="23" pageNumber="24">C</emphasis>
SHA334, PIMUZ 39107;
<emphasis id="B9567019647EFFA6FF25EA50D085F8B0" bold="true" box="[169,180,1856,1876]" pageId="23" pageNumber="24">L</emphasis>
SHA332, PIMUZ 39108.
<emphasis id="B9567019647EFFA6FEF1EA50D1B9F8B0" bold="true" box="[381,392,1856,1876]" pageId="23" pageNumber="24">E</emphasis>
<taxonomicName id="4C22D788647EFFA6FE00EA50D2BDF8B0" authority="(MUller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[396,652,1856,1876]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B9567019647EFFA6FE00EA50D275F8B0" box="[396,580,1856,1876]" italics="true" pageId="23" pageNumber="24">Guangxidella bransoni</emphasis>
(MÜller)
</taxonomicName>
; SHA304, PIMUZ 39139.
<emphasis id="B9567019647EFFA6FCDBEA50D353F8B0" bold="true" box="[855,866,1856,1876]" pageId="23" pageNumber="24">F</emphasis>
,
<emphasis id="B9567019647EFFA6FCE6EA50D349F8B0" bold="true" box="[874,888,1856,1876]" pageId="23" pageNumber="24">H</emphasis>
,
<emphasis id="B9567019647EFFA6FC0DEA50D3BCF8B0" bold="true" box="[897,909,1856,1876]" pageId="23" pageNumber="24">K</emphasis>
,
<emphasis id="B9567019647EFFA6FC1AEA50D399F8B0" bold="true" box="[918,936,1856,1876]" pageId="23" pageNumber="24">M</emphasis>
,
<emphasis id="B9567019647EFFA6FC3CEA50D38DF8B0" bold="true" box="[944,956,1856,1876]" pageId="23" pageNumber="24">P</emphasis>
<taxonomicName id="4C22D788647EFFA6FC4DEA50D4BBF8B0" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[961,1162,1856,1876]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="pseudodieneri" status="sp. nov.">
<emphasis id="B9567019647EFFA6FC4DEA50D4BBF8B0" box="[961,1162,1856,1876]" italics="true" pageId="23" pageNumber="24">Discretella pseudodieneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647EFFA6FB03EA50D48CF8B0" box="[1167,1213,1856,1876]" pageId="23" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B9567019647EFFA6FB4EEA50D4FCF8B0" bold="true" box="[1218,1229,1856,1876]" pageId="23" pageNumber="24">F</emphasis>
QIA124, PIMUZ 39120;
<emphasis id="B9567019647EFFA6FF25EA4BD086F88B" bold="true" box="[169,183,1883,1903]" pageId="23" pageNumber="24">H</emphasis>
QIA124,PIMUZ SQL54990;
<emphasis id="B9567019647EFFA6FE12EA4BD19BF88B" bold="true" box="[414,426,1883,1903]" pageId="23" pageNumber="24">K</emphasis>
QIA121, PIMUZ 39121;
<emphasis id="B9567019647EFFA6FDFCEA4BD2B3F88B" bold="true" box="[624,642,1883,1903]" pageId="23" pageNumber="24">M</emphasis>
QIA120, PIMUZ 39122;
<emphasis id="B9567019647EFFA6FCCBEA4BD362F88B" bold="true" box="[839,851,1883,1903]" pageId="23" pageNumber="24">P</emphasis>
LAR232, PIMUZ 39123.
<emphasis id="B9567019647EFFA6FB96EA4BD419F88B" bold="true" box="[1050,1064,1883,1903]" pageId="23" pageNumber="24">G</emphasis>
<taxonomicName id="4C22D788647EFFA6FBA0EA4BD4EAF88B" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[1068,1243,1883,1903]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B9567019647EFFA6FBA0EA4BD4B1F88B" box="[1068,1152,1883,1903]" italics="true" pageId="23" pageNumber="24">Discretella</emphasis>
cf.
<emphasis id="B9567019647EFFA6FB10EA4BD4EAF88B" box="[1180,1243,1883,1903]" italics="true" pageId="23" pageNumber="24">discreta</emphasis>
</taxonomicName>
(MÜller); QIA120, PIMUZ 39109.
<emphasis id="B9567019647EFFA6FEAEEA65D119F86D" bold="true" box="[290,296,1909,1929]" pageId="23" pageNumber="24">I</emphasis>
sp. indet.; QIA120, PIMUZ 39294.
<emphasis id="B9567019647EFFA6FDCEEA65D261F86D" bold="true" box="[578,592,1909,1929]" pageId="23" pageNumber="24">N</emphasis>
<taxonomicName id="4C22D788647EFFA6FDD9EA65D298F86D" authorityName="Orchard" authorityYear="2005" box="[597,681,1909,1929]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B9567019647EFFA6FDD9EA65D298F86D" box="[597,681,1909,1929]" italics="true" pageId="23" pageNumber="24">Discretella</emphasis>
</taxonomicName>
? n. sp. B; LAR232, PIMUZ 39112.
<emphasis id="B9567019647EFFA6FC36EA65D3F8F86D" bold="true" box="[954,969,1909,1929]" pageId="23" pageNumber="24">O</emphasis>
<taxonomicName id="4C22D788647EFFA6FC42EA65D413F86D" authorityName="Orchard" authorityYear="2005" box="[974,1058,1909,1929]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B9567019647EFFA6FC42EA65D413F86D" box="[974,1058,1909,1929]" italics="true" pageId="23" pageNumber="24">Discretella</emphasis>
</taxonomicName>
? n. sp. C; SHA342,PIMUZ 39114
</paragraph>
</caption>
<paragraph id="8B9DAC0B6471FFA9FF1BEDFBD181FEE7" blockId="24.[151,776,235,1667]" box="[151,432,235,259]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF1BEDFBD0CEFEE7" box="[151,255,235,259]" italics="true" pageId="24" pageNumber="25">Material.</emphasis>
&gt;
<specimenCount id="9D2467826471FFA9FE9AEDFBD19DFEE7" box="[278,428,235,259]" count="10" pageId="24" pageNumber="25" type="generic">10 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF21EC1CD245FD87" blockId="24.[151,776,235,1667]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF21EC1CD126FEC7" box="[173,279,268,291]" italics="true" pageId="24" pageNumber="25">Diagnosis</emphasis>
. A species with a segminate-to-carminate P
<subScript id="17A6AE4E6471FFA9FD72EC07D336FECC" attach="left" box="[766,775,279,296]" fontSize="7" pageId="24" pageNumber="25">1</subScript>
element with distinctively wedge-shaped, bevelled denticle tips. Cusp is either terminal or in front of one or two posterior denticles. Te shallow basal cavity is posteriorly elongated and tapered. In lateral view the lower margin is straight or upturned anteriorly, conspicuously upturned at mid-length, in front of the cusp, and straight or upturned posteriorly. Te denticle directly in front of the cusp and above the kick in the lower margin may get broader towards its wedge-shaped tip and is usually conspicuously broader than the other denticles.
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF21EF7CD0F7FA67" blockId="24.[151,776,235,1667]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF21EF7CD11BFD67" box="[173,298,620,643]" italics="true" pageId="24" pageNumber="25">Description</emphasis>
. Te P
<subScript id="17A6AE4E6471FFA9FEF3EF67D1B9FD6C" attach="left" box="[383,392,631,648]" fontSize="7" pageId="24" pageNumber="25">1</subScript>
element is segminate to carminate (the posterior process bearing one or two denticles) and has a rigid cockscomb-like form. In lateral view, the lower margin is conspicuously bent or upturned at mid-length, the anterior and posterior parts of the lower margin making an angle of 15° to 40° and being often offset. Te anterior lower margin is straight or slightly upturned, the posterior lower margin is usually more upturned. Te upper margin is arcuate, with a peak in height slightly behind mid-length. Te length-to-height ratio of the element is about 11.2:1. Te moderately fused and laterally flattened denticles seem to radiate from a point that is located below the element, slightly anterior of the lower margin kick. Te denticles are usually slightly recurved posteriorly. Te cusp is located directly behind the lower margin kick and is as high as, or smaller than the up to 2 posterior denticles. Te denticle directly in front of the cusp is usually conspicuously larger and broader than the other denticles. It may get broader towards its characteristically wedge-shaped tip. In lower/aboral view, the shallow basal cavity is subtriangular, being elongated and tapered posteriorly. Te posterior end of the basal cavity can be either pointy or sub-rounded (as in the
<typeStatus id="549912A96471FFA9FD14E83BD2CDFAA7" box="[664,764,1323,1347]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
). A groove runs from the basal cavity pit to the anterior end.
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FF21E89BD3F8FAE7" blockId="24.[151,776,235,1667]" lastBlockId="24.[813,1437,235,1476]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FF21E89BD120FA47" box="[173,273,1419,1443]" italics="true" pageId="24" pageNumber="25">Remarks.</emphasis>
Orchard (2007) was the first to differentiate such forms on the basis of their wedge-shaped denticle tips (his
<taxonomicName id="4C22D7886471FFA9FEBBE8DBD1E1FA07" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[311,464,1483,1507]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196471FFA9FEBBE8DBD1E1FA07" box="[311,464,1483,1507]" italics="true" pageId="24" pageNumber="25">Neospathodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FE50E8DCD22BFA07" box="[476,538,1484,1507]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
<collectionCode id="ED3334CE6471FFA9FDA9E8DBD20AFA07" box="[549,571,1483,1507]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="24" pageNumber="25" type="Museum">V</collectionCode>
). Te
<typeStatus id="549912A96471FFA9FD0AE8DBD2D7FA07" box="[646,742,1483,1507]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C22D7886471FFA9FF1BE8FBD262F9E7" authority="Dagis (1990)" authorityName="Dagis" authorityYear="1990" box="[151,595,1515,1539]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pamirensis">
<emphasis id="B95670196471FFA9FF1BE8FBD185F9E7" box="[151,436,1515,1539]" italics="true" pageId="24" pageNumber="25">Neospathodus pamirensis</emphasis>
<bibRefCitation id="EFB3D1FA6471FFA9FE4EE8FBD262F9E7" author="Dagis, A. A." box="[450,595,1515,1539]" pageId="24" pageNumber="25" refId="ref39987" refString="Dagis, A. A. (1990). Lower Triassic Conodonta from the Southeastern Pamirs: fossil plates and explanatory text, p. 73 - 89. In A. S. Dagis (Ed.), Trias Sibiri (p. 767). Novosibirsk: Trudy Instituta Geologii i Geofiziki." type="book" year="1990">Dagis (1990)</bibRefCitation>
</taxonomicName>
appears to fall within the scope of
<taxonomicName id="4C22D7886471FFA9FEF4EB1CD233F9C7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[376,514,1547,1571]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FEF4EB1CD1A5F9C7" box="[376,404,1548,1571]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
.
<emphasis id="B95670196471FFA9FE2FEB1BD233F9C7" box="[419,514,1547,1571]" italics="true" pageId="24" pageNumber="25">bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FD87EB1CD276F9C7" box="[523,583,1548,1571]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
but Dagis differentiated
<taxonomicName id="4C22D7886471FFA9FF75EB3CD1ABF9A7" authorityName="Dagis" authorityYear="1990" box="[249,410,1580,1603]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pamirensis">
<emphasis id="B95670196471FFA9FF75EB3CD1ABF9A7" box="[249,410,1580,1603]" italics="true" pageId="24" pageNumber="25">Ns. pamirensis</emphasis>
</taxonomicName>
on the basis of a “sharply angular lower margin” of the P
<subScript id="17A6AE4E6471FFA9FE39EB47D18FF98C" attach="left" box="[437,446,1623,1640]" fontSize="7" pageId="24" pageNumber="25">1</subScript>
element and “cut-off ends” of the denticles (our rough translation from Russian), which based on his illustrations seem to correspond to sharply
<emphasis id="B95670196471FFA9FCA1EC1BD347FEC7" box="[813,886,267,291]" italics="true" pageId="24" pageNumber="25">broken</emphasis>
denticles, not to the wedge-shaped natural end that we use as a diagnostic feature of
<taxonomicName id="4C22D7886471FFA9FB47EC3CD565FEA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1227,1364,299,323]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FB47EC3CD4D6FEA7" box="[1227,1255,300,323]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
.
<emphasis id="B95670196471FFA9FB79EC3BD565FEA7" box="[1269,1364,299,323]" italics="true" pageId="24" pageNumber="25">bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FAD0EC3CD5A9FEA7" box="[1372,1432,300,323]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
, a feature that cannot be observed in his
<typeStatus id="549912A96471FFA9FB7FEC5BD562FE87" box="[1267,1363,331,355]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
of
<emphasis id="B95670196471FFA9FAF6EC5CD39BFE67" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C22D7886471FFA9FAF6EC5CD397FE67" authorityName="Dagis" authorityYear="1990" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="pamirensis">Ns. pamirensis</taxonomicName>
.
</emphasis>
Moreover, the “sharply angular lower margin” does not differentiate his elements from those of
<taxonomicName id="4C22D7886471FFA9FCA1ECBCD38AFE27" authorityName="Goel" authorityYear="1977" box="[813,955,428,451]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FCA1ECBCD38AFE27" box="[813,955,428,451]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C22D7886471FFA9FC47ECBCD467FE27" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[971,1110,427,451]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FC47ECBCD467FE27" box="[971,1110,427,451]" italics="true" pageId="24" pageNumber="25">Ns. bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FBECECBCD4ACFE27" box="[1120,1181,428,451]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
compares most to
<taxonomicName id="4C22D7886471FFA9FAF6ECBCD3A8FE07" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B95670196471FFA9FAF6ECBCD3A8FE07" italics="true" pageId="24" pageNumber="25">Ns. cristagalli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D7886471FFA9FC42ECDCD46BFE07" authorityName="Goel" authorityYear="1977" box="[974,1114,460,483]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FC42ECDCD46BFE07" box="[974,1114,460,483]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
but none of the latter display the mid-length located, broad, large and bevelled denticle that is characteristic of
<taxonomicName id="4C22D7886471FFA9FBE8EF1CD4C1FDC7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1124,1264,523,547]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FBE8EF1CD4C1FDC7" box="[1124,1264,523,547]" italics="true" pageId="24" pageNumber="25">Ns. bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FB76EF1CD506FDC7" box="[1274,1335,524,547]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
Furthermore,
<taxonomicName id="4C22D7886471FFA9FCFBEF3CD436FDA7" authorityName="Goel" authorityYear="1977" box="[887,1031,556,579]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FCFBEF3CD436FDA7" box="[887,1031,556,579]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
, as defined and illustrated by
<bibRefCitation id="EFB3D1FA6471FFA9FAE4EF3BD342FD87" author="Goel, R. K." pageId="24" pageNumber="25" pagination="1085 - 1101" refId="ref40976" refString="Goel, R. K. (1977). Triassic conodonts from Spiti (Himachal Pradesh), India. Journal of Paleontology, 51, 1085 - 1101." type="journal article" year="1977">Goel (1977)</bibRefCitation>
has a similarly extended basal cavity and hence a similar lower margin but a much larger length-to-height ratio of about 1.8:1, its denticles being essentially smaller and more reclined posteriorly than in
<taxonomicName id="4C22D7886471FFA9FB5DEFBCD56AFD27" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1233,1371,683,707]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FB5DEFBCD56AFD27" box="[1233,1371,683,707]" italics="true" pageId="24" pageNumber="25">Ns. bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FAEEEFBCD5ACFD27" box="[1378,1437,684,707]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
Maekawa and Igo (in
<bibRefCitation id="EFB3D1FA6471FFA9FB92EFDBD4DFFD07" author="Shigeta, Y. &amp; Komatsu, T. &amp; Maekawa, T. &amp; Huyen, D. T." box="[1054,1262,715,739]" pageId="24" pageNumber="25" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
) assigned similar elements from
<collectingCountry id="F335EC9B6471FFA9FC71EFFBD46BFCE7" box="[1021,1114,747,771]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
either to
<taxonomicName id="4C22D7886471FFA9FB41EFFCD56AFCE7" authorityName="Goel" authorityYear="1977" box="[1229,1371,748,771]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FB41EFFCD56AFCE7" box="[1229,1371,748,771]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
or to their
<emphasis id="B95670196471FFA9FCE0EE1CD3B9FCC7" box="[876,904,780,803]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
. sp. indet. A. Tey differentiated
<emphasis id="B95670196471FFA9FB79EE1CD520FCC7" box="[1269,1297,780,803]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
. sp. indet. A on the basis that the P
<subScript id="17A6AE4E6471FFA9FBA6EE27D402FCAC" attach="left" box="[1066,1075,823,840]" fontSize="7" pageId="24" pageNumber="25">1</subScript>
elements are higher and display fewer denticles. In our view, most of the elements they assigned to
<taxonomicName id="4C22D7886471FFA9FC20EE7CD406FC67" authorityName="Goel" authorityYear="1977" box="[940,1079,876,899]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FC20EE7CD406FC67" box="[940,1079,876,899]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
are already too high to belong to
<taxonomicName id="4C22D7886471FFA9FCA1EE9CD388FC47" authorityName="Goel" authorityYear="1977" box="[813,953,908,931]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FCA1EE9CD388FC47" box="[813,953,908,931]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
and most of them display the characteristic bevelled denticles that none of the original
<taxonomicName id="4C22D7886471FFA9FA83EEBCD5ADFC27" authorityName="Goel" authorityYear="1977" box="[1295,1436,940,963]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FA83EEBCD5ADFC27" box="[1295,1436,940,963]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
</taxonomicName>
possessed. Hence, they probably belong to
<taxonomicName id="4C22D7886471FFA9FA9DEEDCD5ADFC07" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1297,1436,971,995]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FA9DEEDCD5ADFC07" box="[1297,1436,971,995]" italics="true" pageId="24" pageNumber="25">Ns. bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FCA1EEFCD359FBE7" box="[813,872,1004,1027]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
instead. Teir material, however, shows a possible somewhat continuous transition between
<taxonomicName id="4C22D7886471FFA9FA87E91CD350FBA7" authority="Goel" authorityName="Goel" authorityYear="1977" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spitiensis">
<emphasis id="B95670196471FFA9FA87E91CD5ADFBC7" box="[1291,1436,1036,1059]" italics="true" pageId="24" pageNumber="25">Ns. spitiensis</emphasis>
Goel
</taxonomicName>
and
<taxonomicName id="4C22D7886471FFA9FC15E93CD412FBA7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[921,1059,1067,1091]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FC15E93CD412FBA7" box="[921,1059,1067,1091]" italics="true" pageId="24" pageNumber="25">Ns. bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FBA6E93CD454FBA7" box="[1066,1125,1068,1091]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
, suggesting a close relationship between both taxa. In comparison with
<taxonomicName id="4C22D7886471FFA9FAB2E95CD343FB67" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="cristagalli">
<emphasis id="B95670196471FFA9FAB2E95CD56BFB87" box="[1342,1370,1100,1123]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
.
<emphasis id="B95670196471FFA9FAE1E95CD343FB67" italics="true" pageId="24" pageNumber="25">cristagalli</emphasis>
</taxonomicName>
, the P
<subScript id="17A6AE4E6471FFA9FC35E967D3F3FB6C" attach="left" box="[953,962,1143,1160]" fontSize="7" pageId="24" pageNumber="25">1</subScript>
elements of
<taxonomicName id="4C22D7886471FFA9FBDDE97CD4EBFB67" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[1105,1242,1131,1155]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="bevelledi" status="n. sp.">
<emphasis id="B95670196471FFA9FBDDE97CD45CFB67" box="[1105,1133,1132,1155]" italics="true" pageId="24" pageNumber="25">Ns</emphasis>
.
<emphasis id="B95670196471FFA9FBF7E97BD4EBFB67" box="[1147,1242,1131,1155]" italics="true" pageId="24" pageNumber="25">bevelledi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626471FFA9FB6EE97CD52CFB67" box="[1250,1309,1132,1155]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
are shorter and higher and although some elements display a posteriormost denticle that is triangular, it is neither smaller nor separated from the other denticles as is characteristic of
<taxonomicName id="4C22D7886471FFA9FCA1E9FCD3F2FAE7" box="[813,963,1259,1283]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="cristagalli">
Ns.
<emphasis id="B95670196471FFA9FCDBE9FBD3F2FAE7" box="[855,963,1259,1283]" italics="true" pageId="24" pageNumber="25">cristagalli</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9DAC0B6471FFA9FCCEE81CD445FA27" blockId="24.[813,1437,235,1476]" pageId="24" pageNumber="25">
<emphasis id="B95670196471FFA9FCCEE81CD38CFAC7" box="[834,957,1292,1315]" italics="true" pageId="24" pageNumber="25">Occurrence</emphasis>
.
<collectingCountry id="F335EC9B6471FFA9FC58E81BD428FAC7" box="[980,1049,1291,1315]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
: Bianyang and Jiarong sections, Nanpanjiang basin,
<collectingRegion id="49E662E96471FFA9FB8BE83BD4FDFAA7" box="[1031,1228,1323,1347]" country="China" name="Guizhou" pageId="24" pageNumber="25">Guizhou province</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6471FFA9FB50E83BD5A7FAA7" author="Chen, Y. &amp; Jiang, H. &amp; Lai, X. &amp; Yan, C. &amp; Richoz, S. &amp; Liu, X. &amp; Wang, L." box="[1244,1430,1323,1347]" pageId="24" pageNumber="25" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA6471FFA9FCA1E85CD3E2FA87" author="Yan, C. &amp; Wang, L. &amp; Jiang, H. &amp; Wignall, P. B. &amp; Sun, Y. &amp; Chen, Y. &amp; Lai, X." box="[813,979,1355,1379]" pageId="24" pageNumber="25" pagination="509 - 522" refId="ref48484" refString="Yan, C., Wang, L., Jiang, H., Wignall, P. B., Sun, Y., Chen, Y., &amp; Lai, X. (2013). Uppermost Permian to Lower Triassic conodonts at Bianyang section, Guizhou province, South China. Palaios, 28 (8), 509 - 522. https: // doi. org / 10.2110 / palo. 2012. p 12 - 077 r" type="journal article" year="2013">Yan et al., 2013</bibRefCitation>
);
<collectingCountry id="F335EC9B6471FFA9FC65E85CD419FA87" box="[1001,1064,1356,1379]" name="Japan" pageId="24" pageNumber="25">Japan</collectingCountry>
: Taho Formation (
<bibRefCitation id="EFB3D1FA6471FFA9FB7DE85BD350FA67" author="Maekawa, T. &amp; Komatsu, T. &amp; Koike, T." pageId="24" pageNumber="25" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
);
<collectingCountry id="F335EC9B6471FFA9FCF1E87BD3ECFA67" box="[893,989,1387,1411]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
: Bac Tuy Formation, Smithian
<taxonomicName id="4C22D7886471FFA9FAD1E87BD342FA47" class="Cephalopoda" family="Flemingitidae" genus="Flemingites" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
<emphasis id="B95670196471FFA9FAD1E87BD342FA47" italics="true" pageId="24" pageNumber="25">Flemingites</emphasis>
</taxonomicName>
to
<taxonomicName id="4C22D7886471FFA9FC2EE89CD437FA47" authorityName="Hyatt &amp; Smith" authorityYear="1905" box="[930,1030,1420,1443]" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
<emphasis id="B95670196471FFA9FC2EE89CD437FA47" box="[930,1030,1420,1443]" italics="true" pageId="24" pageNumber="25">Owenites</emphasis>
</taxonomicName>
beds (
<bibRefCitation id="EFB3D1FA6471FFA9FBD4E89BD507FA47" author="Shigeta, Y. &amp; Komatsu, T. &amp; Maekawa, T. &amp; Huyen, D. T." box="[1112,1334,1419,1443]" pageId="24" pageNumber="25" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
);
<collectingCountry id="F335EC9B6471FFA9FADDE89BD5A8FA47" box="[1361,1433,1419,1443]" name="Oman" pageId="24" pageNumber="25">Oman</collectingCountry>
: Smithian age (Orchard, 2007).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,65 +1,67 @@
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@ -133,7 +135,7 @@
<heading id="2F319CD9CB9FB7D8760340BEFB3EF481" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="0A5731EBBB8CFA8207A8B52CE655063D">
<materialsCitation id="7291451BD4827CB7DAE4CFA73ACFFF80" collectingDate="1956-10-24" collectingDateMax="1956-10-25" collectingDateMin="1956-10-24" collectionCode="LACM" collectorName="Second &amp; Clarke Scholtz's &amp; Third &amp; Fig." country="Argentina" location="Rio Negro" specimenCode="ENT 160322" specimenCount="1" specimenCount-male="1" stateProvince="El Bolson" typeStatus="holotype">
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<typeStatus id="660A141DC886A0C85EC26312BC6644B4" type="holotype">Holotype</typeStatus>
, by original designation
@ -160,17 +162,15 @@
<emphasis id="1DAC39DE40BEC947955EDA1EE51E4557" italics="true">nocheles</emphasis>
</taxonomicName>
/ 1988 / Scholtz / HOLOTYPE ”. Third label [white, printed]: “ [QR Code] /
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” (Fig.
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<emphasis id="1B6A793ADDCE4BC43968060F57ED6724" bold="true">Type locality</emphasis>
: “
<collectingCountry id="208CEC9249AD0CED1235E162D91BF78A">Argentina</collectingCountry>
,
: “ Argentina,
<collectingRegion id="D579A93BF9A73911F1FF14825D35EE6E">El Bolsón</collectingRegion>
,
<location id="5C577A1F25662EAB142D59A5AA54B48A" LSID="urn:lsid:plazi:treatment:A182B442C86E55DAB2E2411DCF6DE621:5C577A1F25662EAB142D59A5AA54B48A" country="Argentina" name="Rio Negro" stateProvince="El Bolson">Río Negro</location>

View file

@ -0,0 +1,167 @@
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<mods:title id="F131E67DE0E6BBEEF987E91563A28D31">New species and a fascinating diversity of Chironomidae (Diptera, Insecta) in and around an overlooked urban vernal pool</mods:title>
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<mods:affiliation id="49712A004A879680BD568D49AD04B549">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran</mods:affiliation>
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<subSubSection id="1BA60350CA83B2BAFD3C54FAF31B9C5C" type="nomenclature">
<paragraph id="85E5EEC3E499EACB343374AF10DDB8BF">
<taxonomicName id="5E8572FDF52ED6A2F5D252B14C39DA2C" ID-CoL="63MXN" authorityName="Kieffer" authorityYear="1912" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C327ED614DEE709C7C7FD20D09283341" italics="true">Polypedilum</emphasis>
(s. s.) sp.
</taxonomicName>
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<paragraph id="805F3140BE72CA04E7BE0053123C9240">
<heading id="CD793BB7080FD5AA42C9E74E2B9F7719" reason="title">Remarks.</heading>
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<paragraph id="A14D56123A766CBCD0C4E64C19578F4B">
The adult males of the
<taxonomicName id="96E83598941C277C5292D92C3F9185DC" authorityName="Kieffer" authorityYear="1912" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="0A59CBF66FFF3907A687934B9522E01D" italics="true">Polypedilum</emphasis>
(
<emphasis id="F0DE2D840249D5FA61A8416339562AC4" italics="true">s. s.</emphasis>
)
</taxonomicName>
specimens that we collected from Pond A resemble
<taxonomicName id="A54CEBD763FD113683CFB1DBF90374C3" authority="Townes, 1945" authorityName="Townes" authorityYear="1945" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="trigonus">
<emphasis id="C3479AB73F8271F42281B12B57E58F49" italics="true">Polypedilum</emphasis>
(
<emphasis id="61AE985F3AA8CAA30734B8A457EC743D" italics="true">s. s.</emphasis>
)
<emphasis id="15E435C222E9032983E56401786D7114" italics="true">trigonus</emphasis>
, Townes, 1945
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. However, these adults did not key out to any known Nearctic adult male in the
<emphasis id="40921CCED17FB4DA8681174F52AC9944" italics="true">
subgenus
<taxonomicName id="7D46845198AFC6864F644E8A7DB7864E" authorityName="Kieffer" authorityYear="1912" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">Polypedilum</taxonomicName>
</emphasis>
based on
<bibRefCitation id="7BB93B9DAD47B06B291D3774FE04CD94" author="Maschwitz DE &amp; Cook EF" firstAuthor="Maschwitz" journalOrPublisher="Ohio, Hilliard" refId="B34" refString="Maschwitz DE, Cook EF (2000) Revision of the Nearctic species of the genus Polypedilum Kieffer (Diptera: Chironomidae) in the subgenera P. (Polypedilum) Kieffer and P. (Uresipedilum) Oyewo and Sæther. Ohio Biological Survey Bulletin New Series Vol. 12. Ohio, Hilliard, 135 pp." title="Revision of the Nearctic species of the genus Polypedilum Kieffer (Diptera: Chironomidae) in the subgenera P. (Polypedilum) Kieffer and P. (Uresipedilum) Oyewo and Sæther. Ohio Biological Survey Bulletin New Series Vol. 12." year="2000">Maschwitz and Cook (2000)</bibRefCitation>
. The adult male of this species is distinguished by the following characteristics: head, thorax, and abdomen uniformly dark brown, legs stramineous; AR 1.8; wing unmarked; anal point extending to or slightly higher than inferior volsella; superior volsella without tubercle, sickle-shaped, robust at the base narrowing and bent at apex; inferior volsella tubular and narrow. Based on the molecular data obtained, the species does not match any known
<taxonomicName id="8570E1A7D67179F55E943E20C38CB1A9">
<emphasis id="BB7DECED38CDF6091A83A668AF4A604D" italics="true">Polypedilum</emphasis>
species
</taxonomicName>
in GenBank or
<collectionCode id="5C999E1580C1BA926E198C37FDB13269">BOLD</collectionCode>
, only to sequences identified only as
<taxonomicName id="2545E94D5664453A9E8AE4F5EAFA4B53" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A1EEB414CFC232F0084AB21F43063306" italics="true">Polypedilum</emphasis>
sp.
</taxonomicName>
, accession numbers
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<accessionNumber id="674707390EAECCFC7756AB71FCD57F8E" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ982463">HQ 982463</accessionNumber>
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and
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</ext-link>
. The adult males obtained in this study possibly represent a new species. However, we need to examine more materials, including those of the related species, before we can make the decision on the status of this species. The hypopygium of the adult male is shown in Fig.
<specimenCount id="840DEE46B224FFB5A29C6F823B35F81D" count="6" type="generic">
<figureCitation id="285473684207B41F43CFBB34491BF068" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Gallery of Chironomidae species collected from Pond A, 202223 A. Labrundinia pilosella (Loew, 1866) B Allocladius nanseni (Kieffer, 1926) C, D Cricotopus (Isocladius) intersectus (Staeger, 1839) E Diplosmittia harrisoni Sæther, 1981 F Smittia aterima (Meigen, 1818) G, H Smittia terrestris Goetghebuer, 1941 I Chironomus (s. s.) acidophilus Keyl, 1960 J Chironomus (s. s.) bifurcatus Wülker et al., 2009 K Chironomus sp. butleri of Martin (2023) L Chironomus (s. s.) maturus Johannsen, 1908 M Chironomus (s. s.) atrella (Townes, 1945) N Chironomus (Lobochironomus) dorsalis Meigen, 1818 O Kiefferulus dux (Johannsen, 1905) P Polypedilum (s. s.) sp. Q Tanytarsus guerlus (Roback, 1957) R Tribelos jucundum (Walker, 1858)., B, E, F, I R male hypopygium C female sternites and genitalia D, G female antenna H female wing. Scale bars: 50 μm." figureDoi="10.3897/zookeys.1208.124495.figure6" httpUri="https://binary.pensoft.net/fig/1103617">6 P</figureCitation>
.
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</paragraph>
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