treatments-xml/data/BE/79/96/BE7996D3D0BD5936A0A3858F13329CB6.xml

<document ID-DOI="http://dx.doi.org/10.3897/nl.47.114772" ID-Pensoft-Pub="2367-5365-47-57" ID-Pensoft-UUID="4A77F35768A45842A1B27D54D5B50724" ID-ZooBank="18BDAA9F5AE5479ABC3D1998C8193791" ModsDocID="2367-5365-47-57" checkinTime="1710546606512" checkinUser="pensoft" docAuthor="Shovkoon, Dmitry F. &amp; Trofimova, Tatyana A." docDate="2024" docId="BE7996D3D0BD5936A0A3858F13329CB6" docLanguage="en" docName="NotaLepidopt 47: 57-79" docOrigin="Nota Lepidopterologica 47" docPubDate="2024-03-15" docSource="http://dx.doi.org/10.3897/nl.47.114772" docTitle="Carriola witti Shovkoon &amp; Trofimova 2024, sp. nov." docType="treatment" docUuid="BC60E0B5-0FAC-4097-9B62-EEF66EEB8B0C" docUuidSource="ZooBank" docVersion="1" id="4A77F35768A45842A1B27D54D5B50724" lastPageNumber="57" masterDocId="4A77F35768A45842A1B27D54D5B50724" masterDocTitle="Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species" masterLastPageNumber="79" masterPageNumber="57" pageNumber="57" updateTime="1710546606512" updateUser="pensoft">
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<mods:title id="16930AD9FF79844D1A0D49BB588029E0">Review of the genus Carriola Swinhoe, 1922 (Lepidoptera, Erebidae, Lymantriinae), with descriptions of four new species</mods:title>
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<mods:namePart id="E1DCC1D1F203BDF05DD550371E62CA71">Shovkoon, Dmitry F.</mods:namePart>
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<mods:affiliation id="632AC4FD72778DFBEE325D9694E219C3">Faculty of Science, University of South Bohemia, Branisovska 31, 370 05 Ceske Budejovice, Czech Republic; shovkoon. d @ googlemail. com &amp; Institute of Entomology, Biology Centre CAS, Branisovska 1760, 370 05 Ceske Budejovice, Czech Republic</mods:affiliation>
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<mods:namePart id="35EDB1B72A1ACCA4621F4E1ACD3EB830">Trofimova, Tatyana A.</mods:namePart>
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<mods:affiliation id="E73661AC05B7B7DA98363750F0C04D8F">Laboratory of Animal Systematic and Faunistic Samara State University, ul. Ac. Pavlova 1, Samara, 443011, Russia; apamea @ mail. ru</mods:affiliation>
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<mods:title id="EF5B4E7B708E2F33EEC18D97B7B18C5F">Nota Lepidopterologica</mods:title>
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<mods:date id="31C815D2A344B23F97B9F231A122855B">2024</mods:date>
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<subSubSection id="56A415DBA7B1E5489F29001027F56ACA" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="D279AA2DBB27C073A25788573A81BD4F" pageId="0" pageNumber="57">
<taxonomicName id="1D067723FF8658C7968AB802D9C8C00E" LSID="https://zoobank.org/BC60E0B5-0FAC-4097-9B62-EEF66EEB8B0C" authority="Shovkoon &amp; Trofimova, 2024" authorityName="Shovkoon &amp; Trofimova" authorityYear="2024" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola witti" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="witti" status="sp. nov.">Carriola witti</taxonomicName>
<taxonomicNameLabel id="D0814A4CE5C1D7895C269DAA6E99C18F" pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
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<subSubSection id="B15ABE7FADF48228B16F29A8BB8457E3" pageId="0" pageNumber="57" type="description">
<paragraph id="6F5AE5965A632DAC8464DEE915C48AA5" pageId="0" pageNumber="57">
<figureCitation id="2AD688DB6B8650E86B9A66FDC2F45D21" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">Figs 12</figureCitation>
<figureCitation id="BBC8166D376AF7C9C3559CB2A75BDB74" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">, 21</figureCitation>
<figureCitation id="0FD671B8DCA5D5C6F0D1AB3FA89C1D86" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">, 30</figureCitation>
<figureCitation id="7F18F6BC48F6CFE1F979816A47F01F7B" captionStart="Figures 37–44" captionStartId="F6" captionText="Figures 37 - 44. Female genitalia of Carriola species. 37. C. ecnomoda (Swinhoe, 1907) (GU 26.699; ZSM); 38. C. thyridophora (Hampson, [1893]) (GU 28.073; ZSM); 39. C. witti sp. nov. PT (GU 25.853; ZSM); 40. C. seminsula (Strand, 1914) (GU 27.445; ZSM); 41. C. saturnioides (Snellen, 1879) (GU 25.850; ZSM); 42. C. polyakovi sp. nov. PT (GU 28.780; ZSM); 43. C. shorokhovi sp. nov. PT (GU 28.784; ZSM); 44. C. fenestrata (Hampson, [1893]) (GU 27.436; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures37-44" httpUri="https://binary.pensoft.net/fig/1003325" pageId="0" pageNumber="57">, 39</figureCitation>
<figureCitation id="609D9801A2789B70EA4A990BD486FB6F" captionStart="Figures 45–53" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">, 47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="1A8E8F6FD9B16BC2FD32F19D04F825BD" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="9C171E0752A19987BD3A0ADAF8E0D398" pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph id="96768DD62AD584A23A7BB5C55D0AF4A4" pageId="0" pageNumber="57">
<materialsCitation id="625000AA3F0909B2355C55D530FA8219" collectingDate="2005-04-01" collectingDateMax="2005-04-14" collectingDateMin="2005-04-01" collectorName="Martini" country="Malaysia" county="Borneo" elevation="1200" location="Trus Madi" municipality="Borneo" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="Holotype">
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<typeStatus id="8A20B7F1368C1787A5DFF4750C010940" type="holotype">Holotype</typeStatus>
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: 
<collectingCountry id="84683743FFFF114949EA3148F81B2330" name="Malaysia">Malaysia</collectingCountry>
: 
<collectingCounty id="56EFC9ADD4820C477AFBB8B4348A50F4">Borneo</collectingCounty>
• 
<specimenCount id="A5A4BF09A6DB6CE3F12B8A51598ADB10" count="1" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="5A74238BB3088DF9F6B03BBFDC2963DD">Borneo</collectingMunicipality>
, 
<collectingRegion id="553E01DD2B250659484FEC95745E3768" country="Malaysia" name="Sabah">Sabah</collectingRegion>
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<location id="C827725C1C224386AF5BC7EC97ECE80A" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:C827725C1C224386AF5BC7EC97ECE80A" country="Malaysia" county="Borneo" municipality="Borneo" name="Trus Madi" stateProvince="Sabah">Trus Madi</location>
, 
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<elevation id="13395CA1008311B34E03FED6FABA44FE" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
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<collectingDate id="7D5A5E5CB5127B44C2EB8E25554B536B" value="2005-04-01" valueMax="2005-04-14" valueMin="2005-04-01">1.-14.iv.2005</collectingDate>
, leg. 
<collectorName id="6EDD5C2592AD2489280B3FDA27A49A47">Martini</collectorName>
(GU 25.852); ZSM.
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</paragraph>
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<materialsCitation id="81590C7E32AF19657F684808323D7E53" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Borneo" country="Malaysia" location="Malaysia" specimenCount="69" specimenCount-female="29" specimenCount-male="40" typeStatus="Paratypes">
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<typeStatus id="24B6130366AA8DC1C6B5B964C8F1CC63" type="paratype">Paratypes</typeStatus>
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[
<specimenCount id="5ECBE82087E028517688AA3D668784F0" count="40" type="male">40 ♂</specimenCount>
, 
<specimenCount id="95161056C6071B76777D848779AEB1FB" count="28" type="female">28 ♀</specimenCount>
]: 
<collectingCountry id="4AF313BF7513678545464F29CC707951" name="Malaysia">Malaysia</collectingCountry>
: 
<collectorName id="BC170998DDBC94F8F181E13AC1569595">Borneo</collectorName>
• 
<specimenCount id="5AB4B659DFD2220A24AEA3BC84E26E76" count="1" type="female">1 ♀</specimenCount>
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; 
<materialsCitation id="E9D75A9509F110E4934832A5FF496272" collectingDate="2005-04-01" collectingDateMax="2005-04-14" collectingDateMin="2005-04-01" collectorName="Martini" country="Malaysia" elevation="1200" location="Mount Trus Madi" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Sabah" typeStatus="paratype">
<collectingRegion id="6F3356FC6B2A23DBA9D03605605A38BC" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, 
<location id="C3AE227FB1B9D2A0DCBC4D4474F8BC57" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:C3AE227FB1B9D2A0DCBC4D4474F8BC57" country="Malaysia" name="Mount Trus Madi" stateProvince="Sabah">Mount Trus Madi</location>
, 
<quantity id="6BB753AAA634D056B9E036D7A76CF1EC" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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. 
<collectingDate id="631E57823A0D91E61076908EF19211CF" value="2005-04-01" valueMax="2005-04-14" valueMin="2005-04-01">1.-14.iv.2005</collectingDate>
, leg. 
<collectorName id="41A129C2130079B2BDAFD927455FC919">Martini</collectorName>
; ZSM • 
<specimenCount id="C456FE330676179895D966A5E4F160A3" count="4" type="male">4 ♂</specimenCount>
, 
<specimenCount id="499F43DD4517ADCB0BA353119E2F89C1" count="4" type="female">4 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="DE371FD30101D2DF94659B87FCCDFEB3" collectingDate="1996-04-09" collectingDateMax="1996-04-14" collectingDateMin="1996-04-09" collectorName="Martini" country="Malaysia" elevation="1150" location="Mount Trus Madi Hoehe" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<location id="785E23B57A4E9B015F0490E8B9F9BCD4" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:785E23B57A4E9B015F0490E8B9F9BCD4" country="Malaysia" name="Mount Trus Madi Hoehe">
Mount Trus Madi 
<normalizedToken id="DB8FF25872798348420D30BBC1489CD8" originalValue="Höhe">Hoehe</normalizedToken>
</location>
, 
<quantity id="7DFB8CD9DBAFBA4FFD6EAE3615F0C700" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<elevation id="4FBED2F2A5DEDDA8451F3F1761FCB128" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</elevation>
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, 
<collectingDate id="BD50FC4BC37B467A089B9A9507984529" value="1996-04-09" valueMax="1996-04-14" valueMin="1996-04-09">9.-14.iv.1996</collectingDate>
, leg. 
<collectorName id="ED844D29A27828A9305DDF1F3E8E439C">Martini</collectorName>
; ZSM • 
<specimenCount id="87B05759880F6B38CACB9883D66C1744" count="2" type="female">2 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="DFDE96DAA297E141B5FFB54DBB3CFDCF" collectingDate="2004-03" collectorName="Martini" country="Malaysia" elevation="1200" location="Mount Trus Madi" specimenCount="16" specimenCount-female="8" specimenCount-male="8" stateProvince="Sabah" typeStatus="paratype">
<collectingRegion id="E2A3B0BAC1A8419A7B9176AF9A4AD92B" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, 
<location id="EE7A22071680E6E32CFAE0389754999C" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:EE7A22071680E6E32CFAE0389754999C" country="Malaysia" name="Mount Trus Madi" stateProvince="Sabah">Mount Trus Madi</location>
, 
<quantity id="B2AA28222F3BFBE1EA83068D07CF7C8A" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="328314030F21A5BE8CB98BEFCF54EB29" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
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, 
<collectingDate id="34A3852D9680068D406388076291AC2C" value="2004-03">III.2004</collectingDate>
, leg. 
<collectorName id="A85D3747FB5B2506236D2CBF5C986286">Martini</collectorName>
; ZSM • 
<specimenCount id="E3C9D405A6B7B5C1D24C3F9E2B58F793" count="8" type="male">8 ♂</specimenCount>
, 
<specimenCount id="ACF3AD54B5564DE7A31B6F989B904BC8" count="8" type="female">8 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="7BE30DD5BB59C6FFFA939AC08B3448B6" collectingDate="1997-04-01" collectingDateMax="1997-04-07" collectingDateMin="1997-04-01" collectorName="B. &amp; K. Martini" country="Malaysia" elevation="1175" location="Mount Trus Madi" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<location id="D18567AB38FA6DD7A55CF150E03555EC" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:D18567AB38FA6DD7A55CF150E03555EC" country="Malaysia" name="Mount Trus Madi">Mount Trus Madi</location>
, 
<quantity id="5126A22052E002C292FD2A44D9B0B7BD" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">
<elevation id="016464A69C3FE5FD395B8D6C0567A70B" metricMagnitude="3" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" unit="m" value="1175.0" valueMax="1200.0" valueMin="1150.0">1150-1200 m</elevation>
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<collectingDate id="1E798343CE412AA719269A02BCED0DA6" value="1997-04-01" valueMax="1997-04-07" valueMin="1997-04-01">1.-7.iv.1997</collectingDate>
, leg. 
<collectorName id="ED90C65BC7451035F667640BFB094DC5">B. &amp; K. Martini</collectorName>
; ZSM • 
<specimenCount id="65347B3130172AC8AD86670E77F4BE32" count="3" type="male">3 ♂</specimenCount>
, 
<specimenCount id="CBCB489B0346388CBD67420A39FA3B9F" count="2" type="female">2 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="09467DC6068FF912224C7BBC5971168B" collectingDate="2005-03-20" collectingDateMax="2005-04-18" collectingDateMin="2005-03-20" collectorName="B. &amp; K. Martini." country="Malaysia" elevation="1150" location="Mt. Trus Madi" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<collectingRegion id="31DF5826DEA6BFD218CB783F915BB44C" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, 
<location id="8843106B201029CA68A1D0BBCCA2BB27" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:8843106B201029CA68A1D0BBCCA2BB27" country="Malaysia" name="Mt. Trus Madi" stateProvince="Sabah">Mt. Trus Madi</location>
, 
<quantity id="06BEA648AED57F99C42220106619FCE8" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<elevation id="1B35A8B1128A7D2D6F4865E70914251C" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, 
<collectingDate id="2AA701DEA28C8617297A392ED997FFAC" value="2005-03-20" valueMax="2005-04-18" valueMin="2005-03-20">20.iii.-18.iv.2005</collectingDate>
, leg. 
<collectorName id="8BD9EC111B67B629CC9DEE620FF2C82F">B. &amp; K. Martini.</collectorName>
; SSU • 
<specimenCount id="1B54A75F10251575EBAB6B062B178BB0" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="AFFD93E891178BFA30D9A1478082CAB5" count="3" type="female">3 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="75176C11ACAC4EA3DAAAE6A915ECA7AE" collectingDate="1998-03" collectorName="B. &amp; C. Martini" country="Malaysia" elevation="1450" location="Apin Apin" municipality="Mt. Trus Madi" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<collectingRegion id="6EB94E8E8818185A9206887EAA2B4B13" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, 
<collectingMunicipality id="3F4ADCB3407038B45D5D09EAEFFDFDD4">Mt. Trus Madi</collectingMunicipality>
, bei 
<location id="733BDC81D1D80A31B92D5043F566C211" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:733BDC81D1D80A31B92D5043F566C211" country="Malaysia" municipality="Mt. Trus Madi" name="Apin Apin" stateProvince="Sabah">Apin Apin</location>
, 
<quantity id="978B1F90D3CD56FFC38BDC080E83C7B1" metricMagnitude="3" metricUnit="m" metricValue="1.45" unit="m" value="1450.0">
<elevation id="49616183297220C4790F110047B8951B" metricMagnitude="3" metricUnit="m" metricValue="1.45" unit="m" value="1450.0">1450 m</elevation>
</quantity>
, 
<collectingDate id="9D007E4A5F1C9217BD75123AEF947B7E" value="1998-03">iii.1998</collectingDate>
, leg. 
<collectorName id="535EADF5FEB52ABAEB1D3193491FBE00">B. &amp; C. Martini</collectorName>
; ZSM • 
<specimenCount id="F30B448DAAFE3278B1AFD2DEEDDCD5D7" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="5253E0072E120B92A8D0838A3D0FF6F5" collectingDate="1993-04-17" collectorName="Martini." country="Malaysia" elevation="950" location="Apin Apin" municipality="Mt. Trus Madi" specimenCount="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<collectingRegion id="B1E4D4B22C33EE29D2409F063B5C61AE" country="Malaysia" name="Sabah">Sabah</collectingRegion>
, 
<collectingMunicipality id="7FF1274DAACC6367F0309701BF675B39">Mt. Trus Madi</collectingMunicipality>
, 
<location id="1F2A9A7955F8EB77E9E4B4E9D6946503" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:1F2A9A7955F8EB77E9E4B4E9D6946503" country="Malaysia" municipality="Mt. Trus Madi" name="Apin Apin" stateProvince="Sabah">Apin Apin</location>
, 
<quantity id="5F015F649BD80DF7966DDD4D72AF6974" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
<elevation id="D7854BBB89943CF7BC5BE1637F493796" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</elevation>
</quantity>
, 
<collectingDate id="25D5944CA59DF3DC7FC05EC80F1FF534" value="1993-04-17">17.iv.1993</collectingDate>
, leg. 
<collectorName id="9D1D3BA48D96B20C3AB07B16D06A29F6">Martini.</collectorName>
; (GU 25.853); ZSM • 
<specimenCount id="FA6957E08CFD4B6F43E9A51DB883508F" count="3" type="male">3 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="84AEDE9977C1FAB5FD2D81BA316E12B9" collectingDate="1997-09-03" collectingDateMax="1997-09-22" collectingDateMin="1997-09-03" collectorName="E. Kandangan, Rainforest, Jakl" country="Malaysia" elevation="1100" latitude="2.8666668" location="Kalimantan Selatan" longLatPrecision="1307" longitude="115.63333" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="3DA111FB2C52C4AF14ED0BEDE743630D" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:3DA111FB2C52C4AF14ED0BEDE743630D" country="Malaysia" latitude="2.8666668" longLatPrecision="1307" longitude="115.63333" name="Kalimantan Selatan">Kalimantan Selatan</location>
, 
<quantity id="FAE94A8D2707A8D5FD8AAEF66C17F0CF" metricMagnitude="4" metricUnit="m" metricValue="3.0" unit="km" value="30.0">30 km</quantity>
<collectorName id="75FB0E6F94025F2989D6307785FBEBF2">E. Kandangan</collectorName>
, 
<collectorName id="B0C57CAA4B3CFD8D1F39DD7B500FBDC0">Rainforest</collectorName>
, 
<locationDeviation id="C257B920686D4515408AD0A957C353E9" location="Loksado">
<quantity id="74811BC39C3D2DAB9D33C73C126BFBA8" metricMagnitude="4" metricUnit="m" metricValue="1.5" unit="km" value="15.0">15 km</quantity>
NE of Loksado
</locationDeviation>
, 
<quantity id="1CEFB41979D73D9A7E20EC0D65787B19" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation id="E8AC42226FB33036F1C31A4E1F78D96D" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<geoCoordinate id="E9A25FE64D76172ABC885E8640A4613F" degrees="2" direction="north" minutes="52" orientation="latitude" precision="925" value="2.8666668">2°52'N</geoCoordinate>
, 
<geoCoordinate id="2A50A4824FF500373D5D68669E908826" degrees="115" direction="east" minutes="38" orientation="longitude" precision="925" value="115.63333">115°38'E</geoCoordinate>
, 
<collectingDate id="E33066D6436AAA1572B4E6162510DB61" value="1997-09-03" valueMax="1997-09-22" valueMin="1997-09-03">3.-22.ix.1997</collectingDate>
, leg. 
<collectorName id="FE513BF0C9F4D5B50C72D11A24C65839">Jakl</collectorName>
; ZSM • 
<specimenCount id="C3FCF5D9BB05BB6B76D16A5C04A308B3" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="E68A4E662ACC9F753C912DDED36FBD5B" collectingDate="2007-07-22" collectorName="A. Pashenkov" country="Malaysia" location="Tambunan" specimenCount="1" typeStatus="paratype">
<location id="AFF36F17544DDE74EB1987D9009E73AA" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:AFF36F17544DDE74EB1987D9009E73AA" country="Malaysia" name="Tambunan">Tambunan</location>
, 5°40'86&quot;N, 116°'22'52&quot;E, 
<collectingDate id="905E7578E636F0BBE670E927B4C25234" value="2007-07-22">22.vii.2007</collectingDate>
. 
<collectorName id="2206915B19809A8AA2EC1BD443790AB7">A. Pashenkov</collectorName>
; ZISP
</materialsCitation>
.
</paragraph>
<paragraph id="61995EAF71BD2BD62762CCB81818F150" pageId="0" pageNumber="57">
<materialsCitation id="87580202DEFC498F68AA08BF45CB0098" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Island" country="Philippines" location="Palawan" specimenCount="4" specimenCount-male="4" stateProvince="Palawan">
<collectingCountry id="065AD671984818B45CAFE50923F810CC" name="Philippines">Philippines</collectingCountry>
: 
<collectingRegion id="C6D5A0E243987141CC04988C7CB112B1" country="Philippines" name="Palawan">Palawan</collectingRegion>
<collectorName id="47E441939FA025CAA054AD2F5995A5A7">Island</collectorName>
• 
<specimenCount id="55ED5FF082D8ABEEA03770556C49248B" count="4" type="male">4 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="D168D3F92B52CEC3DECF996040D0C799" collectingDate="2007-12-06" collectorName="Lourens" country="Philippines" elevation="10" latitude="8.453067" location="Gangob" longLatPrecision="1" longitude="117.47027" municipality="Tarusan Sandobal" specimenCount="1" specimenCount-male="1" stateProvince="Palawan">
<collectingRegion id="7E67EF157EF69661DDCB36373257C2DB" country="Philippines" name="Palawan">Palawan</collectingRegion>
, E of 
<collectingMunicipality id="C69EA3C513EF063998BA0DEAE1E66E11">Tarusan Sandobal</collectingMunicipality>
, 
<location id="57E37675613B070EEBA6CAEA4AE277CA" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:57E37675613B070EEBA6CAEA4AE277CA" country="Philippines" latitude="8.453067" longLatPrecision="1" longitude="117.47027" municipality="Tarusan Sandobal" name="Gangob" stateProvince="Palawan">Gangob</location>
limestone hill forest, 
<quantity id="1B59A17B4D0E82B5310F3880B7F51F1C" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<elevation id="C465E09986C0DB206ADE50153E4C5B1E" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</elevation>
</quantity>
, 
<geoCoordinate id="C69D393C9A55AB3A15974383DFF87D9A" degrees="08" direction="north" minutes="27.184" orientation="latitude" precision="1" value="8.453067">08°27.184'N</geoCoordinate>
, 
<geoCoordinate id="A8A78058A8E7C971F23A564C2F23ADCB" degrees="117" direction="east" minutes="28.216" orientation="longitude" precision="1" value="117.47027">117°28.216'E</geoCoordinate>
, 
<collectingDate id="0B11CFD6856C53F7E2C39DB752280A90" value="2007-12-06">06.xii.2007</collectingDate>
, leg. JH. 
<collectorName id="55A603A8B7051CC83A957B03FA0D4D5E">Lourens</collectorName>
; ZSM • 
<specimenCount id="56351C69266D3D090BDC0674EFFEB4C6" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="E4791555E1DB4308E82A58E232C42F4D" collectingDate="2006-03-03" collectingDateMax="2006-03-04" collectingDateMin="2006-03-03" collectorName="Lourens" country="Philippines" county="Mantalingajan" elevation="50" latitude="8.85" location="Culasian" longLatPrecision="1300" longitude="117.85" municipality="Pinagar" specimenCount="1" specimenCount-male="1" stateProvince="Palawan">
S. 
<collectingRegion id="AABD6C6B1E210172DFCC707A934EB986" country="Philippines" name="Palawan">Palawan</collectingRegion>
, 
<collectingRegion id="175DD398D53804FA29E942E9EBB888CD" country="Philippines" name="Rizal">Rizal prov.</collectingRegion>
<collectingCounty id="4343473152F5CA88C016A9F6BD440C07">Mantalingajan</collectingCounty>
range 
<collectingMunicipality id="08257CE1CC155F689E3CF088B5C19943">Pinagar</collectingMunicipality>
/ 
<location id="BF2FF9ECAFFCFC1756311CBD1AB14146" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:BF2FF9ECAFFCFC1756311CBD1AB14146" country="Philippines" county="Mantalingajan" latitude="8.85" longLatPrecision="1300" longitude="117.85" municipality="Pinagar" name="Culasian" stateProvince="Palawan">Culasian</location>
, 
<quantity id="5C45FBDFDA48024D64721AD71E1255E5" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation id="8A9E01DB6D26C3165695806DDEB77CE3" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
</quantity>
, primary forest, 
<geoCoordinate id="F289EB139D5B18C1DBFCCFC072E7F2FE" degrees="08" direction="north" minutes="51" orientation="latitude" precision="925" value="8.85">08°51'N</geoCoordinate>
, 
<geoCoordinate id="8B44B5CD9C5306407B54646AB5468765" degrees="117" direction="east" minutes="51" orientation="longitude" precision="925" value="117.85">117°51'E</geoCoordinate>
, 
<collectingDate id="722F7756849E2CD2F995B51A26BD7C98" value="2006-03-03" valueMax="2006-03-04" valueMin="2006-03-03">3.-4.iii.2006</collectingDate>
, leg. JH. 
<collectorName id="51765044673CBBD473989090FAABC8C3">Lourens</collectorName>
; ZSM • 
<specimenCount id="DD10617CF9DB7985707D57DBF7C2FA41" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="008FF9F4B03A30078985E1F7440E7344" collectingDate="2006-03-07" collectorName="Lourens" country="Philippines" elevation="70" latitude="10.433333" location="Mt. Ilian" longLatPrecision="1298" longitude="119.566666" specimenCount="1" specimenCount-female="1" stateProvince="Palawan">
NE. 
<collectingRegion id="BB618827AC20F7E034FA7CCE25FCB171" country="Philippines" name="Palawan">Palawan</collectingRegion>
, foot 
<location id="DBB8318E6CFAAC3BC14D49205B88D587" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:DBB8318E6CFAAC3BC14D49205B88D587" country="Philippines" latitude="10.433333" longLatPrecision="1298" longitude="119.566666" name="Mt. Ilian" stateProvince="Palawan">Mt. Ilian</location>
, 
<quantity id="2B09F9F91BBEBDD2A8972705F1BE893B" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">
<elevation id="EA1D61490DFF61827C47F289E0B1428B" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">70 m</elevation>
</quantity>
, 
<locationDeviation id="19205C5CC83853B6C0CDA3E7A012EBC4" location="Bagon Bayan">
<quantity id="3F7AA67DFF1B6E9E740A65EAF5FA8A45" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
W of Bagon Bayan
</locationDeviation>
, 
<geoCoordinate id="64908F31423C685BE1339BEE4F829C8C" degrees="10" direction="north" minutes="26" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
, 
<geoCoordinate id="A029BC792A820D18188330803231BFF3" degrees="119" direction="east" minutes="34" orientation="longitude" precision="925" value="119.566666">119°34'E</geoCoordinate>
, 
<collectingDate id="B756325B07951CC15772225FA965C489" value="2006-03-07">7.iii.2006</collectingDate>
, leg. JH. 
<collectorName id="9650C455B7A05F95B56B4B6CD48F7C18">Lourens</collectorName>
; ZSM • 
<specimenCount id="5F26A5148E3B9BB93B02390B6A5AEDE2" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="4538D7138CB307F392A5D3E7D509CB1E" collectingDate="2006-03-12" collectorName="Lourens" country="Philippines" elevation="250" latitude="9.833333" location="Salacot Falls" longLatPrecision="1298" longitude="118.61667" specimenCount="4" specimenCount-male="4" stateProvince="Palawan">
C. 
<collectingRegion id="575E5725B053BAB6913F240237B7A4F1" country="Philippines" name="Palawan">Palawan</collectingRegion>
, E of napsan, prim. for. edge 
<location id="85CB7CBA273D0C353A79DBA8A5887AC2" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:85CB7CBA273D0C353A79DBA8A5887AC2" country="Philippines" latitude="9.833333" longLatPrecision="1298" longitude="118.61667" name="Salacot Falls" stateProvince="Palawan">Salacot Falls</location>
, 
<quantity id="73246F4E7605935B372FDDAFC1C1EA9E" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">
<elevation id="B2A2500140E4D61AF0C74214F5B9A05A" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</elevation>
</quantity>
, 
<geoCoordinate id="9BE8B94E8E3C77DEB2AABBA4F9E20154" degrees="09" direction="north" minutes="50" orientation="latitude" precision="925" value="9.833333">09°50'N</geoCoordinate>
, 
<geoCoordinate id="0A57B1BF6A6B4C94F44125F5D9301486" degrees="118" direction="east" minutes="37" orientation="longitude" precision="925" value="118.61667">118°37'E</geoCoordinate>
, 
<collectingDate id="D96D62CC20A47B887C64EA1867C4D0D7" value="2006-03-12">12.iii.2006</collectingDate>
, leg. JH. 
<collectorName id="3360E4B330ACE591643672214E266FE8">Lourens</collectorName>
; ZSM • 
<specimenCount id="8D8B3E4706249372158B555CD8595F12" count="4" type="male">4 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="D0D1898AB3DB44FF978C7B92D520827C" collectingDate="1997-08-05" country="Philippines" elevation="50" location="Irawan" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
<location id="23C7481BD353C0B46D787A0513748C2E" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:23C7481BD353C0B46D787A0513748C2E" country="Philippines" name="Irawan">Irawan</location>
, 
<quantity id="A2B3601740BA77DA73476BFCE2555796" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<elevation id="9D15EAD06A90C0847190CECD8A9F46B2" metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</elevation>
</quantity>
, 
<collectingDate id="1FF5147BDC40C2EE0AC4C964E11ADF81" value="1997-08-05">5.viii.1997</collectingDate>
; ZSM • 
<specimenCount id="F2F2D5D1DBB5A7904D47BCBE57514B99" count="4" type="male">4 ♂</specimenCount>
, 
<specimenCount id="E6A2FB26B34900335F63AE73F10AE9CD" count="2" type="female">2 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="7C4A066EA602B542D0A011D48F0B2AD1" collectingDate="1988-01-12" collectingDateMax="1988-01-17" collectingDateMin="1988-01-12" collectorName="Cerny, Schintlmeister" country="Philippines" elevation="400" latitude="10.35" location="Vicente" longLatPrecision="1298" longitude="119.166664" municipality="Nord" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Palawan">
<collectingMunicipality id="CED8236357EFEB807CE19A557488EA51">Nord</collectingMunicipality>
<collectingRegion id="C30F0DBA7A651E6AA71B0ABB7054866B" country="Philippines" name="Palawan">Palawan</collectingRegion>
, S. 
<location id="9529ED510E0DBB6F6B2B424B8DCDCF62" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:9529ED510E0DBB6F6B2B424B8DCDCF62" country="Philippines" latitude="10.35" longLatPrecision="1298" longitude="119.166664" municipality="Nord" name="Vicente" stateProvince="Palawan">Vicente</location>
, 
<locationDeviation id="02A086FEC07958F87B0AFF53CD729C4F" location="Roxas">
<quantity id="7955BA40D5225A815663AEFE93E9EB0A" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
NEE Roxas
</locationDeviation>
, 
<geoCoordinate id="968DB529B93C0045F47A93D5F0462759" degrees="10" direction="north" minutes="21" orientation="latitude" precision="925" value="10.35">10°21'N</geoCoordinate>
, 
<geoCoordinate id="D36BC9248D0289A116F01548E11F29C6" degrees="119" direction="east" minutes="10" orientation="longitude" precision="925" value="119.166664">119°10'E</geoCoordinate>
, 
<quantity id="05414429E81354B852EDA6640708CB2A" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="1065427F2EE5C56121299309DDB9D677" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
, 
<collectingDate id="C0C5B6186DF910EDE3615837FB52B07E" value="1988-01-12" valueMax="1988-01-17" valueMin="1988-01-12">12.-17.i.1988</collectingDate>
, leg. 
<collectorName id="29457066D3CEA3375F98E6BF615E3F9E">Cerny</collectorName>
&amp; 
<collectorName id="004B52A2E314D58ADF72A8730688283D">Schintlmeister</collectorName>
(GU 26.688); ZSM • 
<specimenCount id="271393ED514EE0A9027DA1B079BC3C67" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="0F4B03F56CDB5A6D775CF95A37A5ED81" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="E976FB24D8F29B6C9F687380FF757386" collectingDate="1997-12" collectorName="Bal" country="Philippines" elevation="800" location="Mt. Mantalingahan" specimenCount="1" specimenCount-male="1">
<location id="A24E49E88A4578646153AFE6658BF67A" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:A24E49E88A4578646153AFE6658BF67A" country="Philippines" name="Mt. Mantalingahan">Mt. Mantalingahan</location>
, 
<quantity id="FDB31A24699735EA86DFE9C5E8346112" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation id="61B8303C7A29983BEBED5818DF891ADD" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
, 
<collectingDate id="FDFDF674E9EDE28B0490A3298ADBC68C" value="1997-12">xii.1997</collectingDate>
, leg. 
<collectorName id="7570D83291971A0CBBE1027913B20F7D">Bal</collectorName>
(GU 26.690); ZSM • 
<specimenCount id="E34CDBFBE3FD4CE8C8DF845605719C6C" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="8358323CFE0336F1B2E2CB5C6A988341" collectingDate="2001-05-23" collectingDateMax="2001-05-30" collectingDateMin="2001-05-23" collectorName="Stroehle, A. &amp; S. Zwick" country="Philippines" elevation="950" latitude="8.783055" location="Kibyawon" longLatPrecision="1" longitude="117.70186" municipality="Mt. Mantalingahan" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingMunicipality id="620326A70ED803158CCE9A37D6386458">Mt. Mantalingahan</collectingMunicipality>
, 
<quantity id="93FA03A23875130C8BBECE35468DE421" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">
<elevation id="233C389F016B3EB998C9B41D72C18797" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</elevation>
</quantity>
(
<location id="9B62E03241A4A03F8402201EC28406BE" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:9B62E03241A4A03F8402201EC28406BE" country="Philippines" latitude="8.783055" longLatPrecision="1" longitude="117.70186" municipality="Mt. Mantalingahan" name="Kibyawon">Kibyawon</location>
) in prim. rainforest 
<geoCoordinate id="5FA113C646BBF601C4E133644F02D562" degrees="8" direction="north" minutes="46" orientation="latitude" precision="1" seconds="59,0" value="8.783055">8°46'59,0&quot;N</geoCoordinate>
, 
<geoCoordinate id="39042CC9B55CF9D62FFD92067F17D6EF" degrees="117" direction="east" minutes="42" orientation="longitude" precision="1" seconds="06,7" value="117.70186">117°42'06,7&quot;E</geoCoordinate>
<collectingDate id="ABD7728BEE96D988EB8073EDB79036DE" value="2001-05-23" valueMax="2001-05-30" valueMin="2001-05-23">23.-30.v.2001</collectingDate>
, MV-lamp coll. 
<normalizedToken id="45C8590F82FE6ED49D193E355DB8745F" originalValue="Ströhle">
<collectorName id="3934E32295C2B6DA93755C113C2D5FA2">Stroehle</collectorName>
</normalizedToken>
leg. 
<collectorName id="9203AA1A17B682F4BF0A3A3F01E1AAE6">A.&amp;S. Zwick</collectorName>
; MSW • 
<specimenCount id="213795CCC9D997C7E867E814804FC076" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="1DF96496E212801D4131574F61B6C0D4" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="E0EE0AA00F9038886BA282D92C978346" collectingDate="1996-10-03" collectingDateMax="1996-10-06" collectingDateMin="1996-10-03" collectorName="Bal" country="Philippinen, Insel" county="Palawan" location="Mainit Brooke's" municipality="Magcasaw" specimenCount="1" specimenCount-female="1">
<collectingCountry id="69916AC6C8728651F6ED09AE66F55C38" isEnumeration="true">Philippinen, Insel</collectingCountry>
<collectingCounty id="1250250237E36CDFC30B97A36F2EB20C">Palawan</collectingCounty>
, 
<collectingMunicipality id="96054160ACDEBA30D5896450E79699A6">Magcasaw</collectingMunicipality>
, 
<location id="DB009E5A485C4BA332B852E015092B37" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:DB009E5A485C4BA332B852E015092B37" country="Philippinen, Insel" county="Palawan" municipality="Magcasaw" name="Mainit Brooke's">Mainit Brooke's</location>
point, 
<collectingDate id="C77E8C87BBFCDABAB045A24D9738C4F6" value="1996-10-03" valueMax="1996-10-06" valueMin="1996-10-03">
3.-6.x. 
<quantity id="826F4F776C5AE6BC75F598D90588F65C" metricMagnitude="6" metricUnit="m" metricValue="1.9969" unit="m" value="1996900.0">1996, 900 m</quantity>
</collectingDate>
, leg. 
<collectorName id="2C595E1837C4439A296E88103FDB89B7">Bal</collectorName>
; ZSM 
<specimenCount id="75A06880601D6A9C7259832B5C27E83C" count="1" type="female">1 ♀</specimenCount>
</materialsCitation>
; 
<materialsCitation id="B465AEEED458A2222D39F271AFA36C3C" collectingDate="2000-02-10" collectingDateMax="2000-02-27" collectingDateMin="2000-02-10" collectorName="Gorbatshev, Siniaev" country="Insel" elevation="800" latitude="9.85" location="Mt. Salacot Res." longLatPrecision="1298" longitude="118.63333" specimenCount="1" specimenCount-male="1">
<location id="F98D960337254EC0F11DA01D3BE0F8CA" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:F98D960337254EC0F11DA01D3BE0F8CA" country="Insel" latitude="9.85" longLatPrecision="1298" longitude="118.63333" name="Mt. Salacot Res.">Mt. Salacot Res.</location>
, 
<quantity id="CD74023FD95240625F2EE8D0BD02945F" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation id="C078A9F5C4E3BEAC0A14C9AD2E7AECF3" metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
, 
<geoCoordinate id="331D4571A64FCD9D365740C37F131B65" degrees="9" direction="north" minutes="51" orientation="latitude" precision="925" value="9.85">9°51'N</geoCoordinate>
, 
<geoCoordinate id="66013F3452BAEAB7341F02480E3F67F1" degrees="118" direction="east" minutes="38" orientation="longitude" precision="925" value="118.63333">118°38'E</geoCoordinate>
<collectingDate id="FF1EB950F0B1182C32FB61748911F00B" value="2000-02-10" valueMax="2000-02-27" valueMin="2000-02-10">10.-27.ii.2000</collectingDate>
, leg. 
<collectorName id="5A7C2479373C26139DF0D27E95BA2336">Gorbatshev</collectorName>
&amp; 
<collectorName id="9A7FD9D614A5C5114FD2C664FA4F0690">Siniaev</collectorName>
; ZSM • 
<specimenCount id="00ACA4D72C7137925C828936B16C68E2" count="1" type="male">1 ♂</specimenCount>
</materialsCitation>
; 
<materialsCitation id="11C7E30C2A5A93B9BFAAB6E60A3109D1" collectingDate="1991-12-10" collectingDateMax="1991-12-21" collectingDateMin="1991-12-10" collectorName="Cerny" country="Insel" county="Calabayong" elevation="200" location="Tanabag river" municipality="Bar Tanabag" specimenCount="1">
<collectingCounty id="AF7683EE18AE34FE584D58A98CECF312">Calabayong</collectingCounty>
, 
<collectingMunicipality id="C7C2D759B8E90A9F56CA9D5955A55CA3">Bar Tanabag</collectingMunicipality>
, 
<location id="55AA9BC3223303E0652C704FE22FA469" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:55AA9BC3223303E0652C704FE22FA469" country="Insel" county="Calabayong" municipality="Bar Tanabag" name="Tanabag river">Tanabag river</location>
valley, 
<quantity id="53D1B23A5A76AB00D4B9D6C84A38A98F" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation id="4CA821A869BEA287F9FF1F4C555CAB3A" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
, forest. 
<collectingDate id="23CB864D5022A2EFD999894376544D5D" value="1991-12-10" valueMax="1991-12-21" valueMin="1991-12-10">10.-21xii.1991</collectingDate>
, leg. 
<collectorName id="633198D86092E6E2F6CB1BDA11DE8D9E">Cerny</collectorName>
; ZSM
</materialsCitation>
.
</paragraph>
<paragraph id="AC9B684AB3F9020E21AC1F4BEA117D38" pageId="0" pageNumber="57">
<materialsCitation id="AD89F1BC05ADD5C10EC45A161EDC3F85" collectingDate="1961-11-11" country="Island" county="Batu Batu" latitude="26.693" location="Tawi-Tawi" longLatPrecision="74" longitude="28.078" municipality="Tarawakan" specimenCount="2" specimenCount-male="2">
<location id="B5C282D42D1951180D8B6A604D1EE23A" LSID="urn:lsid:plazi:treatment:BE7996D3D0BD5936A0A3858F13329CB6:B5C282D42D1951180D8B6A604D1EE23A" country="Island" county="Batu Batu" latitude="26.693" longLatPrecision="74" longitude="28.078" municipality="Tarawakan" name="Tawi-Tawi">Tawi-Tawi</location>
<collectingCountry id="C615A13D52AA86E5612DEBB792275E24" name="Iceland">Island</collectingCountry>
• 
<specimenCount id="10F8F06084B4BF979559BCE24031BBEF" count="2" type="male">2 ♂</specimenCount>
; 
<collectingMunicipality id="AFC9486C3FAE15A3D4D4735FB050D253">Tarawakan</collectingMunicipality>
, north of 
<collectingCounty id="79EC4E02F5A282C659998562D4FBB998">Batu Batu</collectingCounty>
, 
<collectingDate id="6F4D0188E06FE57DB2BC6D97E63EE510" value="1961-11-11">11.xi.1961</collectingDate>
(GU 
<geoCoordinate id="012B344F93FBC744D9E75DC85BD1EE84" degrees="26.693" direction="north" orientation="latitude" precision="55" value="26.693">26.693</geoCoordinate>
, 
<geoCoordinate id="FDFB56D98E06BF4D476324440DA049BF" degrees="28.078" direction="east" orientation="longitude" precision="55" value="28.078">28.078</geoCoordinate>
); ZSM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="01A050352D30DA503B3E1CD624270F89" pageId="0" pageNumber="57" type="diagnosis">
<paragraph id="2814089C9DE29D82ACC66EBE539AF138" pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="6F6BEBF71B10FEEC1DB7F3F11C46ADFE" pageId="0" pageNumber="57">
Externally (Figs 
<figureCitation id="2BE26326C60C3CFFF90E96230380039A" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">12</figureCitation>
, 
<figureCitation id="4F111F798CB7F83D07B79A22FE121743" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21</figureCitation>
) 
<taxonomicName id="101815C37B2E8B5AA6D60134C4510316" lsidName="C. witti" pageId="0" pageNumber="57" rank="species" species="witti">
<emphasis id="999ECF38C72553B3186DB7B68AFF941C" italics="true" pageId="0" pageNumber="57">C. witti</emphasis>
</taxonomicName>
sp. nov. can be confused with 
<taxonomicName id="4FD7BDAB8B96B70D053FD9729EE2360A" lsidName="C. seminsula" pageId="0" pageNumber="57" rank="species" species="seminsula">
<emphasis id="101A206D6FB0A11F2C97277CBAC8B880" italics="true" pageId="0" pageNumber="57">C. seminsula</emphasis>
</taxonomicName>
and 
<taxonomicName id="9F4C8BFF9C3A97298281008AA2309337" lsidName="C. thyridophora" pageId="0" pageNumber="57" rank="species" species="thyridophora">
<emphasis id="031910180862D7C321DF3B156BF528BC" italics="true" pageId="0" pageNumber="57">C. thyridophora</emphasis>
</taxonomicName>
, and any identification must be confirmed by examination of the genital structures. The male genitalia differ in the shape of the valva. As in 
<taxonomicName id="C150BCD84CCE2E062FA9B41AE219C31E" lsidName="C. thyridophora" pageId="0" pageNumber="57" rank="species" species="thyridophora">
<emphasis id="560EAF70CAAC474B8BA4E8998D5A7448" italics="true" pageId="0" pageNumber="57">C. thyridophora</emphasis>
</taxonomicName>
, it is isosceles trapezoidal, but more elongated and pointed. The base of the trapezium is in a ratio of 2.5:1 to the apex and the length of the valva is in a ratio of 3:1 to the base of the valva (Fig. 
<figureCitation id="DD71151C54344078D4C4D660DFE0873F" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">30a</figureCitation>
). The female genitalia differ from those of related species by the scobinate heart-shaped signum, which is about one third the size of the bursa and has poorly separated lobes, each as long as 
<normalizedToken id="6E1D4863E6CEA5738334860804200C06" originalValue="½">1/2</normalizedToken>
the width of the signum.
</paragraph>
<paragraph id="194E586833B1B8CE7F3516FB7BAE102B" pageId="0" pageNumber="57">
The female genitalia differ from those of the related species by the scobinate heart-shaped signum, which is as large as a third of the bursa, with poorly separated lobes that are as long as 
<normalizedToken id="CA56F07D1342B2F0575BC55A02FE1F17" originalValue="½">1/2</normalizedToken>
the width of the signum (Fig. 
<figureCitation id="3FA53B8B262621CD4CF13FEA0A58BA27" captionStart="Figures 37–44" captionStartId="F6" captionText="Figures 37 - 44. Female genitalia of Carriola species. 37. C. ecnomoda (Swinhoe, 1907) (GU 26.699; ZSM); 38. C. thyridophora (Hampson, [1893]) (GU 28.073; ZSM); 39. C. witti sp. nov. PT (GU 25.853; ZSM); 40. C. seminsula (Strand, 1914) (GU 27.445; ZSM); 41. C. saturnioides (Snellen, 1879) (GU 25.850; ZSM); 42. C. polyakovi sp. nov. PT (GU 28.780; ZSM); 43. C. shorokhovi sp. nov. PT (GU 28.784; ZSM); 44. C. fenestrata (Hampson, [1893]) (GU 27.436; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures37-44" httpUri="https://binary.pensoft.net/fig/1003325" pageId="0" pageNumber="57">39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="106E9115571711748465EDAC592907CB" pageId="0" pageNumber="57" type="description">
<paragraph id="78D4D6415F887181A15313384FD32ADD" pageId="0" pageNumber="57">Description.</paragraph>
<paragraph id="2E96FC4D41D876216416FDAB0DC0B577" pageId="0" pageNumber="57">
<emphasis id="1B857DADA85BAE5519163977734A5B05" bold="true" italics="true" pageId="0" pageNumber="57">External appearance.</emphasis>
Frons and vertex orange and covered with dense, drooping scales, with small tufts at the base of antennae. Eyes large, round. Labial palpi bent obliquely upwards, densely pubescent, on the underside of the same pale olive colour as abdomen, orange above. Patagia, tegulae and thorax golden pale brown. 
<emphasis id="41FC709619BE1B70955ACF92BB2DEF04" bold="true" italics="true" pageId="0" pageNumber="57">Male forewings</emphasis>
(Figs 
<figureCitation id="43D8579CF028E0E14FEAD9038286740A" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">12</figureCitation>
, 
<figureCitation id="AAC492AB6B219676F13381A9032A87B1" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21</figureCitation>
): triangular with rounded apex, length of forewing 10.6-15.3 mm (14 mm in holotype). The ratio between the length of forewing margin and the costa is about 1.2:1 in both sexes. Ground colour of upper side beige-green and typically with two hyaline windows (Figs 
<figureCitation id="510125769527A08D008D97B7A4DCFCBE" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">12</figureCitation>
, 
<figureCitation id="E8FD6D97DF2B81039893526BCD7DC011" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21a, b</figureCitation>
). Central hyaline window occupies central cell up to half the length of cells M2-M3, M3-CuA1, CuA1-CuA2, and CuA2-1+2A and bordered by clearly visible antemedial and subterminal lines. Hyaline satellite oval and located between RS4-M1, and bordered by postmedial and subterminal lines. Discal spot forms triangle with postmedial line. Underside of forewing with lighter monotonous background, hyaline windows with black borders (Fig. 
<figureCitation id="EEF5CCFCC587608635ECD1E7D9C08594" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21b</figureCitation>
). 
<emphasis id="EF535EF3BC166D54D21B58B47AADF0A7" bold="true" italics="true" pageId="0" pageNumber="57">Male hindwings</emphasis>
(Figs 
<figureCitation id="D47B6336EE68ECCFAD01DFC5B45091BD" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">12</figureCitation>
, 
<figureCitation id="E70B998F70090A0AC241147E40F6B1CC" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21</figureCitation>
): triangular with rounded tip, central hyaline window occupies central part of wing, from ⅓ of area of central cell to the discal veins and to 
<normalizedToken id="8081039529C051A83698FE0E1AE1B802" originalValue="½">1/2</normalizedToken>
of the area of cells M2-M3, M3-CuA1, CuA1-CuA2 and CuA2-1+2A. First triangular hyaline satellite window between veins Sc+R1 and Rs, and second between the veins Rs and M1 (Fig. 
<figureCitation id="2FEED4D15223F674500BAFBC72F59CB0" captionStart="Figures 10–18" captionStartId="F3" captionText="Figures 10 - 18. The wing venation and hyaline window mapping of Carriola species. 10. C. ecnomoda (Swinhoe, 1907); 11. C. thyridophora (Hampson, [1893]); 12. C. witti sp. nov.; 13. C. seminsula (Strand, 1914); 14. C. saturnioides (Snellen, 1879); 15. C. polyakovi sp. nov.; 16. C. shorokhovi sp. nov.; 17. C. fenestrata (Hampson, [1893]); 18. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures10-18" httpUri="https://binary.pensoft.net/fig/1003322" pageId="0" pageNumber="57">12</figureCitation>
). 
<emphasis id="441DEFDE0DE030D11F032BF2E687E4E3" bold="true" italics="true" pageId="0" pageNumber="57">Female forewings</emphasis>
of olive colour with large hyaline window bordered by clearly visible terminal line (Fig. 
<figureCitation id="B2B6D6C662028CBC76EAFAD89A3205A6" captionStart="Figures 19–27" captionStartId="F4" captionText="Figures 19 - 27. Adults of Carriola species. a. Male; b. Underside wings of male; c. Female. 19. C. ecnomoda (Swinhoe, 1907); a, b. East Java, Mt. Baluran (GU 26.698; ZSM); c. East Java, Mt. Baluran, (GU 26.699; ZSM); 20. C. thyridophora (Hampson, [1893]); a, b. NE India. Assam (GU 27.442; ZSM); c. SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 21 C. witti sp. nov.; a, b. HT (GU 25.852; ZSM); c. PT (GU 25.853; ZSM); 22. C. seminsula (Strand, 1914); a, b. Sud Vietnam, Bao Loc (GU 27.431; ZSM); c. Nord Vietnam, Mai Chau (GU 27.445; ZSM); 23. C. saturnioides (Snellen, 1879); a, b. N. Sulawesi, (GU 21.966; ZSM); c. Sulawesi, Namo (GU 25.850; ZSM); 24. C. polyakovi sp. nov.; a, b. HT (GU 28.779; ZSM); c. PT (GU 28.780; ZSM); 25. C. shorokhovi sp. nov.; a, b. HT (GU 26.961; ZSM); c. PT (GU 28.784; ZSM); 26. C. fenestrata (Hampson, [1893]); a, b. Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); c. Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 27. C. zolotuhini sp. nov.; a, b. HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale." figureDoi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57">21c</figureCitation>
). Length of forewing 14.5-16.5 mm. 
<emphasis id="F30D4871F4DD906E744AD48AE9DF14DB" bold="true" italics="true" pageId="0" pageNumber="57">Female hindwings</emphasis>
triangular with rounded tip. The general outline of pattern, colour and the type of arrangement of hyaline window repeat the forewing. 
<emphasis id="22D51D46C3201AB94A5F8BC4A52C245E" bold="true" italics="true" pageId="0" pageNumber="57">Male genitalia</emphasis>
(Figs 
<figureCitation id="60E02E8E76580AA9F44E00DEC65CFCEB" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">30a-c</figureCitation>
). Superuncus rectangular, slightly divided into two lobes at the apex (Fig. 
<figureCitation id="B7E0DCBE6CBED67DBC68C17FFF8BB8F7" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">30c</figureCitation>
). Uncus (Fig. 
<figureCitation id="6F08631F8ADB5BECE016F0AA9259F92E" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">30a</figureCitation>
) looks like a short, apically rounded lobe. Valva trapezoidal, elongate and pointed. Base of the trapezium is in a ratio of 2.5:1 to the apex, the length of valva/width of its base in a ratio of about 3:1. Juxta U-shaped. Aedeagus short, strongly bevelled at its anterior and posterior end (Fig. 
<figureCitation id="540C95C94956D4072BE77EC91EF2C31E" captionStart="Figures 28–36" captionStartId="F5" captionText="Figures 28 - 36. Male genitalia of Carriola species. a. Genitalia; b. Aedeagus with everted vesica; c. Superuncus (VIII tergum). 28. C. ecnomoda (Swinhoe, 1907) (GU 26.698; ZSM); 29. C. thyridophora (Hampson, [1893]) (GU 27.442; ZSM); 30. C. witti sp. nov. HT (GU 25.852; ZSM); 31. C. seminsula (Strand, 1914): HT (prep. NO 597; SMF); 32. C. saturnioides (Snellen, 1879) (GU 21.966; ZSM); 33. C. polyakovi sp. nov. HT (GU 28.779; ZSM); 34. C. shorokhovi sp. nov. HT (GU 26.961; ZSM); 35. C. fenestrata (Hampson, [1893]) (GU 27.435; ZSM); 36. C. zolotuhini sp. nov. HT (GU 28.069; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures28-36" httpUri="https://binary.pensoft.net/fig/1003324" pageId="0" pageNumber="57">30b</figureCitation>
). 
<emphasis id="8F294441538D2E7E2132ED69D57ACCA1" bold="true" italics="true" pageId="0" pageNumber="57">Female genitalia</emphasis>
(Fig. 
<figureCitation id="ACDC5959CBB31E11FFD13EAA379A5059" captionStart="Figures 37–44" captionStartId="F6" captionText="Figures 37 - 44. Female genitalia of Carriola species. 37. C. ecnomoda (Swinhoe, 1907) (GU 26.699; ZSM); 38. C. thyridophora (Hampson, [1893]) (GU 28.073; ZSM); 39. C. witti sp. nov. PT (GU 25.853; ZSM); 40. C. seminsula (Strand, 1914) (GU 27.445; ZSM); 41. C. saturnioides (Snellen, 1879) (GU 25.850; ZSM); 42. C. polyakovi sp. nov. PT (GU 28.780; ZSM); 43. C. shorokhovi sp. nov. PT (GU 28.784; ZSM); 44. C. fenestrata (Hampson, [1893]) (GU 27.436; ZSM). Scale bar represents 1 mm, all genitalia to scale." figureDoi="10.3897/nl.47.114772.figures37-44" httpUri="https://binary.pensoft.net/fig/1003325" pageId="0" pageNumber="57">39</figureCitation>
). Papillae anales weakly sclerotised, broad and rounded, pseudopapillae small and narrow, membranous. Posterior apophyses slender and as long as two thirds of lobes of papillae anales. Antrum cup-shaped, small. Ductus bursae membranous, strongly constricted before antrum and gradually widened before corpus bursae. Corpus bursae rounded, rather short and as long as ductus. Signum scobinate heart-shaped, as large as one third of bursa, with the lobes poorly separated and as long as 
<normalizedToken id="C86C0C5727FF863E806FF2372A9F41CB" originalValue="½">1/2</normalizedToken>
the width of signum.
</paragraph>
<caption id="C21807DAD1C6815B1C35C5B9147051F3" doi="10.3897/nl.47.114772.figures19-27" httpUri="https://binary.pensoft.net/fig/1003323" pageId="0" pageNumber="57" start="Figures 19–27" startId="F4">
<paragraph id="F88D9F0D1390F5FB869B014B98F5ABB7" pageId="0" pageNumber="57">
<emphasis id="C573BA165ABF59D589B93BF28489699F" bold="true" pageId="0" pageNumber="57">Figures 19-27.</emphasis>
Adults of 
<taxonomicName id="32814A0BB050149971D291C8C48E6AE9" authorityName="Swinhoe" authorityYear="1922" class="Insecta" family="Erebidae" genus="Carriola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carriola" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="D398907AF032523F531B0F8C4D0F35A0" italics="true" pageId="0" pageNumber="57">Carriola</emphasis>
</taxonomicName>
species. 
<emphasis id="95262313BAD7EDF767F95C941E5364FC" bold="true" pageId="0" pageNumber="57">a.</emphasis>
Male; 
<emphasis id="6D963F56F9B1ABB1FBBBABC4926C7F3D" bold="true" pageId="0" pageNumber="57">b.</emphasis>
Underside wings of male; 
<emphasis id="B6BB6B36261E1838E0489C4775EB85C6" bold="true" pageId="0" pageNumber="57">c</emphasis>
. Female. 
<emphasis id="66DD63DECB7F921A301C84EFDEC1F107" bold="true" pageId="0" pageNumber="57">19.</emphasis>
<taxonomicName id="AD9F3662C646A06E64AAA0E24987F6B0" lsidName="C. ecnomoda" pageId="0" pageNumber="57" rank="species" species="ecnomoda">
<emphasis id="2CB28EDDCC14936F1073671A0D4682B3" italics="true" pageId="0" pageNumber="57">C. ecnomoda</emphasis>
</taxonomicName>
(Swinhoe, 1907); 
<emphasis id="9687F6B90897BBBF249D022D65FFFC0B" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
East Java, Mt. Baluran (GU 26.698; ZSM); 
<emphasis id="3943E40493A762E9180E7904BDE30A76" bold="true" pageId="0" pageNumber="57">c.</emphasis>
East Java, Mt. Baluran, (GU 26.699; ZSM); 
<emphasis id="3A2AB2FCF6DE7BF5272AB97672704000" bold="true" pageId="0" pageNumber="57">20.</emphasis>
<taxonomicName id="8768ED428560A875D9DC2D43CDF2EB84" lsidName="C. thyridophora" pageId="0" pageNumber="57" rank="species" species="thyridophora">
<emphasis id="FC802B3354DEF20241B8A955C2A0D354" italics="true" pageId="0" pageNumber="57">C. thyridophora</emphasis>
</taxonomicName>
(Hampson, [1893]); 
<emphasis id="EFE10C3EFB8CDB9CFA03D09481324C6A" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
NE India. Assam (GU 27.442; ZSM); 
<emphasis id="8984547DA3F0CC907C4005CF5DBA225E" bold="true" pageId="0" pageNumber="57">c.</emphasis>
SW Kampuchea, Kirirom Nat. Park (GU 28.073; ZSM); 
<emphasis id="B89DC348C25971C7498A1D3FC4F1E38C" bold="true" pageId="0" pageNumber="57">21</emphasis>
<taxonomicName id="159AB62146CB07F6CB5D70346A5ACE37" lsidName="C. witti" pageId="0" pageNumber="57" rank="species" species="witti">
<emphasis id="623B8572BABD69D2C69C7C1A5911C6F6" italics="true" pageId="0" pageNumber="57">C. witti</emphasis>
</taxonomicName>
sp. nov.; 
<emphasis id="EDF4450F9C4C323A16E9A0096B6E4201" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
HT (GU 25.852; ZSM); 
<emphasis id="72AABA8931B210C6E9495A54297FB589" bold="true" pageId="0" pageNumber="57">c.</emphasis>
PT (GU 25.853; ZSM); 
<emphasis id="AAEB659B400CB567FD677139DFA61CF0" bold="true" pageId="0" pageNumber="57">22.</emphasis>
<taxonomicName id="63F446D4124BAA71B06F9EE4AEF45D21" lsidName="C. seminsula" pageId="0" pageNumber="57" rank="species" species="seminsula">
<emphasis id="A4157A6B5E0C79514EF9FE7116777C6A" italics="true" pageId="0" pageNumber="57">C. seminsula</emphasis>
</taxonomicName>
(Strand, 1914); 
<emphasis id="7912557DB288C835CA5C4C0B71BFAF8D" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
Sud Vietnam, Bao Loc (GU 27.431; ZSM); 
<emphasis id="DB07A6F872DA3951953A516F01F6317D" bold="true" pageId="0" pageNumber="57">c.</emphasis>
Nord Vietnam, Mai Chau (GU 27.445; ZSM); 
<emphasis id="92D786313B63777CC218C3C0FCBA98AA" bold="true" pageId="0" pageNumber="57">23.</emphasis>
<taxonomicName id="3F19C254FBBBB1C9A075CFF82683EF94" lsidName="C. saturnioides" pageId="0" pageNumber="57" rank="species" species="saturnioides">
<emphasis id="899AC427E5A5C672107B0C45196B44CF" italics="true" pageId="0" pageNumber="57">C. saturnioides</emphasis>
</taxonomicName>
(Snellen, 1879); 
<emphasis id="47B0DF85347EED6E57AD854225A12EAE" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
N. Sulawesi, (GU 21.966; ZSM); 
<emphasis id="45058DEE6A6E912740FCE0B4DB0A9344" bold="true" pageId="0" pageNumber="57">c.</emphasis>
Sulawesi, Namo (GU 25.850; ZSM); 
<emphasis id="8673BC3EA3BC126E871A60171D0FEFFF" bold="true" pageId="0" pageNumber="57">24.</emphasis>
<taxonomicName id="99C2AD4CD84D27527F0C732A8578A760" lsidName="C. polyakovi" pageId="0" pageNumber="57" rank="species" species="polyakovi">
<emphasis id="7D6FDF90A059FBB4FD068E9C7926ABB3" italics="true" pageId="0" pageNumber="57">C. polyakovi</emphasis>
</taxonomicName>
sp. nov.; 
<emphasis id="CB3E41F1405B472FD140D88D2CFC2BE8" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
HT (GU 28.779; ZSM); 
<emphasis id="7AD4395D3CD41BE1E1A9AADAFBA14D8B" bold="true" pageId="0" pageNumber="57">c.</emphasis>
PT (GU 28.780; ZSM); 
<emphasis id="13534939DA289BC0B21A2F88DAE8B6C7" bold="true" pageId="0" pageNumber="57">25.</emphasis>
<taxonomicName id="2425028F2E3CAA0FF0A549000E68FD9B" lsidName="C. shorokhovi" pageId="0" pageNumber="57" rank="species" species="shorokhovi">
<emphasis id="4186F3D959944BD22038BC99E8CEDDD7" italics="true" pageId="0" pageNumber="57">C. shorokhovi</emphasis>
</taxonomicName>
sp. nov.; 
<emphasis id="BDE6FB75817D5496A764159170B29156" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
HT (GU 26.961; ZSM); 
<emphasis id="DFADEEA3242C42D7545891FD8AECC311" bold="true" pageId="0" pageNumber="57">c.</emphasis>
PT (GU 28.784; ZSM); 
<emphasis id="E94DDECA52D718DB36FAB2F6AC436562" bold="true" pageId="0" pageNumber="57">26.</emphasis>
<taxonomicName id="E69031CAFD03C69D24142CD5E443D21D" lsidName="C. fenestrata" pageId="0" pageNumber="57" rank="species" species="fenestrata">
<emphasis id="1210D02D5C803CEBAFC94DB747D78E40" italics="true" pageId="0" pageNumber="57">C. fenestrata</emphasis>
</taxonomicName>
(Hampson, [1893]); 
<emphasis id="10735D5DD7D288EE89F62C59DE709EAD" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
Vietnam, prov. Thai Nguyen (GU 27.435; ZSM); 
<emphasis id="7ABF98C11DE7C3839BBF92FD4E9BCBC9" bold="true" pageId="0" pageNumber="57">c.</emphasis>
Vietnam, prov. Thai Nguyen, (GU 27.436; ZSM); 
<emphasis id="8D9F0A6F7DFA069E7A1487E23EF7B50F" bold="true" pageId="0" pageNumber="57">27.</emphasis>
<taxonomicName id="01A7EB1ACF5817ADA2C95FB7C9E37B58" lsidName="C. zolotuhini" pageId="0" pageNumber="57" rank="species" species="zolotuhini">
<emphasis id="7CD0648E0C20C63217C3B20BA85E4786" italics="true" pageId="0" pageNumber="57">C. zolotuhini</emphasis>
</taxonomicName>
sp. nov.; 
<emphasis id="00F7B9BC62D5A62098AFCDB9B1966B8C" bold="true" pageId="0" pageNumber="57">a, b.</emphasis>
HT (GU 28.069; ZSM). Scale bar represents 10 mm, all specimens to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C11080AADDB97CEFEC67AF78E4A3316D" pageId="0" pageNumber="57" type="biology_ecology">
<paragraph id="3086263F871F7D40E2283012A81A030A" pageId="0" pageNumber="57">Biology.</paragraph>
<paragraph id="F3A3E2FF3A306D68E0B671800BA89D94" pageId="0" pageNumber="57">Unknown. Adults were observed from January to December in the forests at elevations up to 1450 m.a.s.l.</paragraph>
</subSubSection>
<subSubSection id="5FC22A607379F80F56B056CC2DF56285" pageId="0" pageNumber="57" type="distribution">
<paragraph id="501FFC1140F3616559C617811BE81576" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="5855275924C8ACE254BC7005BCD79DA6" pageId="0" pageNumber="57">
(Fig. 
<figureCitation id="841621267502C376137346788E733736" captionStart="Figures 45–53" captionStartId="F7" captionText="Figures 45 - 53. Distribution maps of Carriola species. 45. C. ecnomoda (Swinhoe, 1907); 46. C. thyridophora (Hampson, [1893]); 47. C. witti sp. nov.; 48. C. seminsula (Strand, 1914); 49. C. saturnioides (Snellen, 1879); 50. C. polyakovi sp. nov.; 51. C. shorokhovi sp. nov.; 52. C. fenestrata (Hampson, [1893]); 53. C. zolotuhini sp. nov." figureDoi="10.3897/nl.47.114772.figures45-53" httpUri="https://binary.pensoft.net/fig/1003326" pageId="0" pageNumber="57">47</figureCitation>
). Borneo, and the nearest Philippine Islands of Palawan and Tawi-Tawi.
</paragraph>
</subSubSection>
<subSubSection id="6D00CDA71D91844269712019C26E3E5E" pageId="0" pageNumber="57" type="etymology">
<paragraph id="F0AD26662D480C4B04A98F64C7300521" pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph id="3A48F65C41CAEBBDBB6ED749641FF238" pageId="0" pageNumber="57">The species is dedicated to the memory of Dr. Thomas J. Witt, a famous lepidopterist and founder of the Witt Museum in Munich, whose collection fund made this research possible.</paragraph>
</subSubSection>
</treatment>
</document>