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<document id="E612F94D0D2CA8D825321999FCF47BCD" ID-DOI="10.5281/zenodo.13155283" ID-ISSN="0068-547X" ID-Zenodo-Dep="13155283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538343316" checkinUser="felipe" docAuthor="Deuve, Thierry, Kavanaugh, David H. &amp; Liang, Hongbin" docDate="2016" docId="03D6879DFF8CFF953C2C644AFC5F712D" docLanguage="en" docName="ProcCalifAcadSci.63.12.341-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (12)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Queinnectrechus griswoldi Deuve &amp; Kavanaugh &amp; Liang 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="369" masterDocId="FFEFFFE5FF95FF893C576071FFCE7750" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species." masterLastPageNumber="455" masterPageNumber="341" pageNumber="366" updateTime="1722545240047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="DD937913E3021C8535775D78DCB2D85E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ABED210C7F68133F4FE11D2E376036B4">
<mods:title id="25358D0B5D18A0D541AD091D7EA67194">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species.</mods:title>
</mods:titleInfo>
<mods:name id="A130E545D558E167E1B637F7D1F20F82" type="personal">
<mods:role id="F3F86D4E1D05CDAB0A06432FCB434CBB">
<mods:roleTerm id="614AD811E9F9F536EE01FB25E3A4834C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB57460AAB0D5B472900450C99A03B5D">Deuve, Thierry</mods:namePart>
<mods:affiliation id="E62D56929F0807C8ADB32BDECA39970B">Institut de Systématique, Evolution, Biodiversité, ISYEB - UMR 7205 - MNHN, CNRS, UPMC, EPHE, Muséum National d’Histoire Naturelle, Sorbonne-Universités, 57 rue Cuvier, CP 50, F- 75231 Paris cedex 05, France.</mods:affiliation>
</mods:name>
<mods:name id="08FD5FFD81209B86D29D4CE87B0E1E6D" type="personal">
<mods:role id="4EE1B5DFF3F25BDD14688FADAACF299D">
<mods:roleTerm id="D8E0DFBC4E13B73ECB0F883B9B1F47D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB57C5E8ED9B3969FAF2FA296D8B0C22">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="910E2572C24543349F68B1DDDB3D98E6">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="A6820F4950169F31486A50CE45711D9D" type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name id="1050D20A069F39A3A94DB8DF5AFC3A64" type="personal">
<mods:role id="27B471146B3DD77AC2C97A0F1556E6FC">
<mods:roleTerm id="F886B6D93F03A558A6AE0511DDF90BBE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="443C038F7B28DCD170016C9343027466">Liang, Hongbin</mods:namePart>
<mods:affiliation id="5DC634AEB9168B63EE7F2E3DC06F25CD">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China * Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1A4F64E35D8EBBE4B098D3B1C96B68E3">text</mods:typeOfResource>
<mods:relatedItem id="C347CF63A86FE0588B7B3B1B6B690E92" type="host">
<mods:titleInfo id="6E2ED4C79D0B5F707CC9FE169B1F8957">
<mods:title id="5F0BFADB9131962AC5E8BCF279947AB4">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="2290C410134273456D300BA5A5C73BCF">
<mods:date id="DD4A6671C2000F94E55A5CF1FDBBB9DC">2016</mods:date>
<mods:detail id="E55ABD48A2EB4B5777EB8F8807D5B827" type="pubDate">
<mods:number id="1CA94BE67E5F861C3383B03C9BE62AE1">2016-10-14</mods:number>
</mods:detail>
<mods:detail id="A98ACB29CAC1FBEB8B4E1B7A4953E6FE" type="volume">
<mods:number id="9C2E86B7BA722FA3890067AD661D54CA">63</mods:number>
</mods:detail>
<mods:detail id="C777C21412570FD66F11693DA8D92C04" type="issue">
<mods:number id="93FC78F289243090ED2D3047ADCA81A1">12</mods:number>
</mods:detail>
<mods:extent id="DC4C117C91CB89CC9507174ACDA49893" unit="page">
<mods:start id="F1CF02729DACA78A1200D95E3AC6F81E">341</mods:start>
<mods:end id="C1C5024215671DC6C2592DB46FD243AA">455</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B2430F9A817F5FC8B4DE95759908311F">journal article</mods:classification>
<mods:identifier id="373887049E1BCF80BA3FE03DC7C2C42B" type="DOI">10.5281/zenodo.13155283</mods:identifier>
<mods:identifier id="4FD584855F9DF6F72D816ACC6DF4A601" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="14DB03673FE06300534642A2D5980A7E" type="Zenodo-Dep">13155283</mods:identifier>
<mods:identifier id="40B2899A864983AEB3D628A7065514BC" type="ZooBank">4C790FE0-B735-4592-8827-EEF83C663CB1</mods:identifier>
</mods:mods>
<treatment id="03D6879DFF8CFF953C2C644AFC5F712D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D" httpUri="http://treatment.plazi.org/id/03D6879DFF8CFF953C2C644AFC5F712D" lastPageId="28" lastPageNumber="369" pageId="25" pageNumber="366">
<subSubSection id="C3656500FF8CFF903C2C644AFCA87304" box="[123,870,1083,1108]" pageId="25" pageNumber="366" type="nomenclature">
<paragraph id="8BC0368BFF8CFF903C2C644AFCA87304" blockId="25.[123,870,1083,1138]" box="[123,870,1083,1108]" pageId="25" pageNumber="366">
<heading id="D08881E7FF8CFF903C2C644AFCA87304" box="[123,870,1083,1108]" fontSize="10" level="2" pageId="25" pageNumber="366" reason="4">
<emphasis id="B90BEA99FF8CFF903C2C644AFCA87304" bold="true" box="[123,870,1083,1108]" pageId="25" pageNumber="366">
<taxonomicName id="4C7F4D08FF8CFF903C2C644AFD8A7304" ID-CoL="8MCS6" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[123,580,1083,1108]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="366" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8CFF903C2C644AFEF97303" bold="true" box="[123,311,1083,1107]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8CFF903D10644DFE4F7303" bold="true" box="[327,385,1084,1107]" italics="true" pageId="25" pageNumber="366">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8CFF903DC6644AFE387303" bold="true" box="[401,502,1083,1107]" italics="true" pageId="25" pageNumber="366">griswoldi</emphasis>
Deuve
</taxonomicName>
and Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8CFF903F47644DFCA87304" box="[784,870,1084,1108]" pageId="25" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8CFF903C2C642DFE2D7321" box="[123,483,1115,1138]" pageId="25" pageNumber="366" type="description">
<paragraph id="8BC0368BFF8CFF903C2C642DFE2D7321" blockId="25.[123,870,1083,1138]" box="[123,483,1115,1138]" pageId="25" pageNumber="366">
(
<figureCitation id="13442A0EFF8CFF903CD5642DFF1D7321" box="[130,211,1116,1138]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="25" pageNumber="366">Figs. 12</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903C8A642DFECD7322" box="[221,259,1116,1138]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="25" pageNumber="366">37b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903D59642DFEAD7322" box="[270,355,1116,1138]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="25" pageNumber="366">38a, 38b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903D39642DFE5A7322" box="[366,404,1116,1138]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="25" pageNumber="366">39b</figureCitation>
,
<figureCitation id="13442A0EFF8CFF903DC9642AFE127321" box="[414,476,1115,1137]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStartId-0="107.[171,183,1629,1648]" captionStartId-1="108.[171,183,1359,1378]" captionStartId-2="109.[171,183,1628,1647]" captionTargetBox-0="[120,1176,245,1580]" captionTargetBox-1="[120,1176,228,1340]" captionTargetBox-2="[197,1099,216,1598]" captionTargetId-0="figure-14@107.[120,1176,216,1608]" captionTargetId-1="figure-12@108.[120,1176,228,1340]" captionTargetId-2="figure-14@109.[197,1099,216,1598]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionTargetPageId-2="109" captionText-0="FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region." captionText-1="FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region." captionText-2="FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region." figureDoi-0="http://doi.org/10.5281/zenodo.13155378" figureDoi-1="http://doi.org/10.5281/zenodo.13155380" figureDoi-2="http://doi.org/10.5281/zenodo.13155382" httpUri-0="https://zenodo.org/record/13155378/files/figure.png" httpUri-1="https://zenodo.org/record/13155380/files/figure.png" httpUri-2="https://zenodo.org/record/13155382/files/figure.png" pageId="25" pageNumber="366">46–48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8CFF923CFC64F4FD4375E6" lastPageId="27" lastPageNumber="368" pageId="25" pageNumber="366" type="materials_examined">
<paragraph id="8BC0368BFF8CFF903CFC64F4FB5B71ED" blockId="25.[123,1174,1156,1726]" pageId="25" pageNumber="366">
<emphasis id="B90BEA99FF8CFF903CFC64F4FE9473CB" bold="true" box="[171,346,1157,1181]" pageId="25" pageNumber="366">
<smallCapsWord id="8D26A057FF8CFF903CFC64F4FF2A73CB" baselines="1175,1174" box="[171,228,1157,1181]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="25" pageNumber="366">TYPE</smallCapsWord>
<smallCapsWord id="8D26A057FF8CFF903CBA64F9FE9473CB" baselines="1174" box="[237,346,1160,1179]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="25" pageNumber="366">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="3B173CD6FF8CFF903DD464F5FE2A724D" collectingDate="2006-08-13" collectorName="J. A. Miller" country="China" county="Gongshan County" elevation="3720" latitude="27.76978" location="Cikai Township" longLatPrecision="1" longitude="98.44681" pageId="25" pageNumber="366" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54C48829FF8CFF903DD464F5FE2773CC" box="[387,489,1156,1180]" pageId="25" pageNumber="366" type="holotype">Holotype</typeStatus>
, a male, in IOZ, labeled: “CASENT 1026334”/ “
<collectingCountry id="F368761BFF8CFF903FAA64F4FB9873CD" box="[1021,1110,1157,1181]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF90383564F4FF6A73EC" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903CE364D5FEB273ED" box="[180,380,1188,1213]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<location id="8EA06050FF8CFF903DDC64D5FDF373EC" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903DDC64D5FDF373EC" box="[395,573,1188,1212]" country="China" county="Gongshan County" latitude="27.76978" longLatPrecision="1" longitude="98.44681" name="Cikai Township" pageId="25" pageNumber="366" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FF8CFF903E1B64D4FC4B73EC" box="[588,901,1188,1213]" location="Heipu Yakou" pageId="25" pageNumber="366">
<quantity id="4C879B6EFF8CFF903E1B64D4FD5573EC" box="[588,667,1188,1213]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="366" unit="km" value="0.1">0.1 km</quantity>
SE of Heipu Yakou
</locationDeviation>
in valley below tunnel,
<geoCoordinate id="EE4B504CFF8CFF903C2C64B5FF32738C" box="[123,252,1220,1244]" degrees="27.76978" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.76978">N27.76978°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903D5C64B5FE40738C" box="[267,398,1220,1244]" degrees="98.44681" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.44681">E98.44681°</geoCoordinate>
,”/ “
<quantity id="4C879B6EFF8CFF903DE264B4FDC5738C" box="[437,523,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">
<elevation id="0052D1B8FF8CFF903DE264B4FDC5738C" box="[437,523,1221,1245]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,
<date id="FFC1104BFF8CFF903E4064B4FD0A738C" box="[535,708,1220,1245]" pageId="25" pageNumber="366" value="2006-08-13">
<collectingDate id="EF85E9A3FF8CFF903E4064B4FD0A738C" box="[535,708,1220,1245]" pageId="25" pageNumber="366" value="2006-08-13">13 August 2006</collectingDate>
</date>
, Stop #DHK-2006-073 D.H.
<taxonomicName id="4C7F4D08FF8CFF90385764B4FBB7738D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1024,1145,1221,1245]" pageId="25" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFF8CFF903C2C6494FF2173AD" box="[123,239,1253,1277]" pageId="25" pageNumber="366">J.A. Miller</collectorName>
collectors”/ “
<typeStatus id="54C48829FF8CFF903D286494FDC073AD" box="[383,526,1253,1277]" pageId="25" pageNumber="366" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7F4D08FF8CFF903E426494FC1E73AC" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[533,976,1253,1277]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="366" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8CFF903E426494FD0773AD" box="[533,713,1253,1277]" italics="true" pageId="25" pageNumber="366">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8CFF903E8F6494FCDE73AC" box="[728,784,1253,1276]" italics="true" pageId="25" pageNumber="366">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8CFF903F496494FC4D73AD" box="[798,899,1253,1277]" italics="true" pageId="25" pageNumber="366">griswoldi</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8CFF9038206494FF66724C" pageId="25" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [red label]
</materialsCitation>
.
<materialsCitation id="3B173CD6FF8CFF903DB86574FBC4720D" collectionCode="CAS, IOZ, MNHN" country="China" pageId="25" pageNumber="366" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54C48829FF8CFF903DB86574FD99724D" box="[495,599,1285,1309]" pageId="25" pageNumber="366" type="paratype">Paratypes</typeStatus>
(a total of 20):
<specimenCount id="9D79FD02FF8CFF903F556575FC9B724D" box="[770,853,1284,1309]" count="5" pageId="25" pageNumber="366" type="male">5 males</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF903FDC6574FC3D724D" box="[907,1011,1285,1309]" count="3" pageId="25" pageNumber="366" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF9038776574FB98724D" box="[1056,1110,1285,1309]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF9038356574FB5F724D" box="[1122,1169,1285,1309]" pageId="25" pageNumber="366">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903C2C6554FF00726D" box="[123,206,1317,1341]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="366">MNHN</collectionCode>
) labeled same as
<typeStatus id="54C48829FF8CFF903DDD6554FE29726D" box="[394,487,1317,1341]" pageId="25" pageNumber="366" type="holotype">holotype</typeStatus>
except first label “CASENT 1026333”, “CASENT 1026335” to “CASENT 1026338” and “CASENT 1026330” to “CASENT 1026332”, respectively
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF9038426534FC3972ED" collectingDate="2002-10-01" collectingDateMax="2002-10-02" collectingDateMin="2002-10-01" collectionCode="IOZ" collectorName="P. E. Marek" country="China" county="Gongshan County" elevation="3370" latitude="27.77032" location="Cikai Township" longLatPrecision="1" longitude="98.44661" pageId="25" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF9038426534FBBC720D" box="[1045,1138,1349,1373]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903C2C6514FF67722D" box="[123,169,1381,1405]" pageId="25" pageNumber="366">IOZ</collectionCode>
) labeled “CASENT 1010344”/ “
<collectingCountry id="F368761BFF8CFF903E476514FDA7722D" box="[528,617,1381,1405]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903E216514FD04722C" box="[630,714,1381,1404]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E8F6514FC50722D" box="[728,926,1381,1405]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<location id="8EA06050FF8CFF903FFC6514FB94722D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903FFC6514FB94722D" box="[939,1114,1381,1405]" country="China" county="Gongshan County" latitude="27.77032" longLatPrecision="1" longitude="98.44661" name="Cikai Township" pageId="25" pageNumber="366" stateProvince="Yunnan">Cikai Township</location>
,
<locationDeviation id="9A722FA9FF8CFF9038306514FDA372CD" location="Gongshan on Dulong Valley Road" pageId="25" pageNumber="366">
<quantity id="4C879B6EFF8CFF9038306514FF5372CD" metricMagnitude="4" metricUnit="m" metricValue="5.26" pageId="25" pageNumber="366" unit="km" value="52.6">52.6 km</quantity>
W of Gongshan on Dulong Valley Road
</locationDeviation>
,
<quantity id="4C879B6EFF8CFF903E2B65F4FCDF72CD" box="[636,785,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="25" pageNumber="366" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">
<elevation id="0052D1B8FF8CFF903E2B65F4FCDF72CD" box="[636,785,1413,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="25" pageNumber="366" unit="m" value="3370.0" valueMax="3380.0" valueMin="3360.0">3360-3380 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFF8CFF903F6A65F4FC0F72CD" box="[829,961,1413,1437]" degrees="27.77032" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.77032">N27.77032°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903F8365F4FBAD72CD" box="[980,1123,1413,1437]" degrees="098.44661" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.44661">E098.44661°</geoCoordinate>
,
<date id="FFC1104BFF8CFF90382565F4FEDC72ED" pageId="25" pageNumber="366" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">
<collectingDate id="EF85E9A3FF8CFF90382565F4FEDC72ED" pageId="25" pageNumber="366" value="2002-10-01" valueMax="2002-10-02" valueMin="2002-10-01">1-2 October 2002</collectingDate>
</date>
, Stop #DHK-2002-034, D.H.
<taxonomicName id="4C7F4D08FF8CFF903E0A65D4FD1872ED" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[605,726,1445,1469]" pageId="25" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
&amp;
<collectorName id="268A535DFF8CFF903EA965D4FCB972ED" box="[766,887,1445,1469]" pageId="25" pageNumber="366">P.E. Marek</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF90385365D4FC62716D" collectingDate="2006-08-20" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; P. Hu &amp; D. Z. Dong &amp; J. Wang" country="China" county="Gongshan County" latitude="27.98206" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" pageId="25" pageNumber="366" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF90385365D4FB8172ED" box="[1028,1103,1445,1469]" count="1" pageId="25" pageNumber="366" type="male">1 male</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF9038DF65D4FF00728D" count="3" pageId="25" pageNumber="366" type="female">3 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903D5365B4FEF4728D" box="[260,314,1477,1501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903D1D65B4FEB6728D" box="[330,376,1477,1501]" pageId="25" pageNumber="366">IOZ</collectionCode>
) labeled “CASENT 1024862” and “CASENT 1024863” to “CASENT 1024865”, respectively/ “
<collectingCountry id="F368761BFF8CFF903DDD6594FE2D72AD" box="[394,483,1509,1533]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903DB96597FD8C72AD" box="[494,578,1510,1533]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E1A6594FCDE72AD" box="[589,784,1509,1533]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8CFF903F4D6594FBEA72AD" box="[794,1060,1509,1533]" pageId="25" pageNumber="366">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8CFF903CC06674FE9F714D" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903CC06674FE9F714D" box="[151,337,1541,1565]" country="China" county="Gongshan County" latitude="27.98206" longLatPrecision="1" longitude="98.48027" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="25" pageNumber="366" stateProvince="Yunnan">Kawakarpu Shan</location>
, on slope
<locationDeviation id="9A722FA9FF8CFF903DEC6677FD54714D" box="[443,666,1541,1565]" location="Chukuai Lake" pageId="25" pageNumber="366">NE of Chukuai Lake</locationDeviation>
, 3950,”/”
<geoCoordinate id="EE4B504CFF8CFF903EA96674FC4F714D" box="[766,897,1541,1565]" degrees="27. 98206" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.98206">N27.98206°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903FD86674FBD0714D" box="[911,1054,1541,1565]" degrees="098.48027" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.48027">E098.48027°</geoCoordinate>
,
<date id="FFC1104BFF8CFF90387E6674FF7D716D" pageId="25" pageNumber="366" value="2006-08-20">
<collectingDate id="EF85E9A3FF8CFF90387E6674FF7D716D" pageId="25" pageNumber="366" value="2006-08-20">20 August 2006</collectingDate>
</date>
, Stop #DHK-2006-086
<collectorName id="268A535DFF8CFF903DF96657FE3D716D" box="[430,499,1573,1597]" pageId="25" pageNumber="366">Y. Liu</collectorName>
,
<collectorName id="268A535DFF8CFF903DAB6657FDF6716D" box="[508,568,1574,1597]" pageId="25" pageNumber="366">P. Hu</collectorName>
,
<collectorName id="268A535DFF8CFF903E156657FD79716E" box="[578,695,1574,1598]" pageId="25" pageNumber="366">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFF8CFF903E8C6657FCFE716E" box="[731,816,1574,1598]" pageId="25" pageNumber="366">J. Wang</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF903FE16654FC8071ED" collectingDate="2006-08-18" collectionCode="CAS, IOZ, MNHN" collectorName="J. A. Miller &amp; D. Z. Dong &amp; Y. Liu" country="China" county="Gongshan County" elevation="3720" latitude="27.98121" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" pageId="25" pageNumber="366" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF903FE16654FBC9716D" box="[950,1031,1573,1597]" count="3" pageId="25" pageNumber="366" type="male">3 males</specimenCount>
and
<specimenCount id="9D79FD02FF8CFF90386D6654FB58716D" box="[1082,1174,1573,1597]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903CF36634FF14710D" box="[164,218,1605,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903CBD6634FED7710D" box="[234,281,1605,1629]" pageId="25" pageNumber="366">IOZ</collectionCode>
,
<collectionCode id="ED6EAE4EFF8CFF903D7E6634FEB2710D" box="[297,380,1605,1629]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="366">MNHN</collectionCode>
) labeled “CASENT 1026813” to “CASENT 1026815” and “CASENT 1026815”, respectively/ “
<collectingCountry id="F368761BFF8CFF903DDD6614FE2D712D" box="[394,483,1637,1661]" name="China" pageId="25" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8CFF903DB96617FD8C712D" box="[494,578,1638,1661]" country="China" name="Yunnan" pageId="25" pageNumber="366">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8CFF903E1A6614FCDE712D" box="[589,784,1637,1661]" pageId="25" pageNumber="366">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8CFF903F4D6614FBEA712D" box="[794,1060,1637,1661]" pageId="25" pageNumber="366">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8CFF903CC066F4FE8371CD" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903CC066F4FE8371CD" box="[151,333,1669,1693]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="25" pageNumber="366" stateProvince="Yunnan">Kawakarpu Shan</location>
at
<location id="8EA06050FF8CFF903D3966F4FDC871CD" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8CFF903D3966F4FDC871CD" box="[366,518,1669,1693]" country="China" county="Gongshan County" latitude="27.98121" longLatPrecision="1" longitude="98.4758" municipality="Bingzhongluo Township" name="Chukuai Lake" pageId="25" pageNumber="366" stateProvince="Yunnan">Chukuai Lake</location>
,
<quantity id="4C879B6EFF8CFF903E4666F4FDAD71CD" box="[529,611,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">
<elevation id="0052D1B8FF8CFF903E4666F4FDAD71CD" box="[529,611,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.72" pageId="25" pageNumber="366" unit="m" value="3720.0">3720 m</elevation>
</quantity>
,”/ “
<geoCoordinate id="EE4B504CFF8CFF903EDC66F4FCC071CD" box="[651,782,1669,1693]" degrees="27.98121" direction="north" orientation="latitude" pageId="25" pageNumber="366" precision="1" value="27.98121">N27.98121°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8CFF903F4A66F4FC6271CD" box="[797,940,1669,1693]" degrees="098.47580" direction="east" orientation="longitude" pageId="25" pageNumber="366" precision="1" value="98.4758">E098.47580°</geoCoordinate>
,
<date id="FFC1104BFF8CFF903FE166F4FB9071CD" box="[950,1118,1669,1694]" pageId="25" pageNumber="366" value="2006-08-18">
<collectingDate id="EF85E9A3FF8CFF903FE166F4FB9071CD" box="[950,1118,1669,1694]" pageId="25" pageNumber="366" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-079
<collectorName id="268A535DFF8CFF903D6866D7FE7271ED" box="[319,444,1701,1726]" pageId="25" pageNumber="366">J.A. Miller</collectorName>
,
<collectorName id="268A535DFF8CFF903D9966D7FD8571EE" box="[462,587,1702,1726]" pageId="25" pageNumber="366">D.Z. Dong</collectorName>
, &amp;
<collectorName id="268A535DFF8CFF903E2866D7FD0771ED" box="[639,713,1701,1725]" pageId="25" pageNumber="366">Y. Liu</collectorName>
collectors”
</materialsCitation>
;
<materialsCitation id="3B173CD6FF8CFF903F3766D4FB5B71ED" box="[864,1173,1701,1725]" collectionCode="CAS" country="China" pageId="25" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D79FD02FF8CFF903F3766D4FC0A71ED" box="[864,964,1701,1725]" count="1" pageId="25" pageNumber="366" type="female">1 female</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8CFF903FA966D4FBFD71ED" box="[1022,1075,1701,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="366" type="Museum">CAS</collectionCode>
) labeled
</materialsCitation>
</paragraph>
<caption id="DF006603FF8FFF933CFC6640FD547126" ID-DOI="http://doi.org/10.5281/zenodo.13155305" ID-Zenodo-Dep="13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="26" pageNumber="367" startId="26.[171,183,1585,1604]" targetBox="[147,1176,216,1569]" targetPageId="26" targetType="figure">
<paragraph id="8BC0368BFF8FFF933CFC6640FD547126" blockId="26.[123,1173,1584,1655]" pageId="26" pageNumber="367">
<smallCapsWord id="8D26A057FF8FFF933CFC6640FF267112" baselines="1599,1598" box="[171,232,1585,1604]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="26" pageNumber="367">FIGURE</smallCapsWord>
12.
<taxonomicName id="4C7F4D08FF8FFF933D446641FD8C7113" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[275,578,1584,1603]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="367" phylum="Arthropoda" rank="species" species="griswoldi" status="sp. nov.">
<emphasis id="B90BEA99FF8FFF933D446641FE6D7113" box="[275,419,1584,1603]" italics="true" pageId="26" pageNumber="367">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8FFF933DE56640FE2D7113" box="[434,483,1585,1603]" italics="true" pageId="26" pageNumber="367">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8FFF933DA66641FD8C7113" box="[497,578,1584,1603]" italics="true" pageId="26" pageNumber="367">griswoldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF8FFF933E1D6640FD5C7114" box="[586,658,1585,1604]" pageId="26" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for
<taxonomicName id="4C7F4D08FF8FFF933FB8663BFB92710D" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[1007,1116,1610,1629]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="367" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8FFF933FB8663BFB92710D" box="[1007,1116,1610,1629]" italics="true" pageId="26" pageNumber="367">Q. griswoldi</emphasis>
</taxonomicName>
in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
</paragraph>
</caption>
<paragraph id="8BC0368BFF8EFF923C2C60AAFCEA7684" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
“
<materialsCitation id="3B173CD6FF8EFF923CD360AAFCD17684" collectingDate="2006-08-18" collectingDateMax="2006-08-19" collectingDateMin="2006-08-18" collectionCode="IOZ" collectorName="Y. Liu &amp; All" country="China" county="Gongshan County" elevation="3750" latitude="27.97686" location="Kawakarpu Shan" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" pageId="27" pageNumber="368" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
CASENT 1025813”/ “
<collectingCountry id="F368761BFF8EFF923D2360AAFE0377A3" box="[372,461,219,243]" name="China" pageId="27" pageNumber="368">CHINA</collectingCountry>
,
<collectingRegion id="49BBF869FF8EFF923D8060AAFDE577A2" box="[471,555,219,242]" country="China" name="Yunnan" pageId="27" pageNumber="368">Yunnan</collectingRegion>
,
<collectingCounty id="62A14E07FF8EFF923E6260AAFD3977A3" box="[565,759,219,243]" pageId="27" pageNumber="368">Gongshan County</collectingCounty>
,
<collectingMunicipality id="6BA4ACF1FF8EFF923F5660AAFBC777A3" box="[769,1033,219,243]" pageId="27" pageNumber="368">Bingzhongluo Township</collectingMunicipality>
, SW slope of
<location id="8EA06050FF8EFF923C2C608AFEFF7643" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923C2C608AFEFF7643" box="[123,305,251,275]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Kawakarpu Shan" pageId="27" pageNumber="368" stateProvince="Yunnan">Kawakarpu Shan</location>
<locationDeviation id="9A722FA9FF8EFF923D60608AFDAB7643" box="[311,613,251,275]" location="Chukuai Lake" pageId="27" pageNumber="368">
<quantity id="4C879B6EFF8EFF923D60608AFE4E7643" box="[311,384,251,275]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="368" unit="km" value="0.3">0.3 km</quantity>
SW of Chukuai Lake
</locationDeviation>
at campsite,”/, “
<geoCoordinate id="EE4B504CFF8EFF923F43608AFC597643" box="[788,919,251,275]" degrees="27.97686" direction="north" orientation="latitude" pageId="27" pageNumber="368" precision="1" value="27.97686">N27.97686°</geoCoordinate>
/
<geoCoordinate id="EE4B504CFF8EFF923FF1608AFBFA7643" box="[934,1076,251,275]" degrees="098.47799" direction="east" orientation="longitude" pageId="27" pageNumber="368" precision="1" value="98.47799">E098.47799°</geoCoordinate>
,
<quantity id="4C879B6EFF8EFF923868608AFB5F7643" box="[1087,1169,251,275]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">
<elevation id="0052D1B8FF8EFF923868608AFB5F7643" box="[1087,1169,251,275]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="27" pageNumber="368" unit="m" value="3750.0">3750 m</elevation>
</quantity>
,
<date id="FFC1104BFF8EFF923C2C616DFEE47663" box="[123,298,283,308]" pageId="27" pageNumber="368" value="2006-08-18">
<collectingDate id="EF85E9A3FF8EFF923C2C616DFEE47663" box="[123,298,283,308]" pageId="27" pageNumber="368" value="2006-08-18">18 August 2006</collectingDate>
</date>
Stop #DHK-2006-078 D.H.
<taxonomicName id="4C7F4D08FF8EFF923E39616AFD297663" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[622,743,283,307]" pageId="27" pageNumber="372" rank="subGenus" status="subgen. nov." subGenus="Kavanaugh">Kavanaugh</taxonomicName>
collector”;
<specimenCount id="9D79FD02FF8EFF923F3B616DFC197663" box="[876,983,283,308]" count="2" pageId="27" pageNumber="368" type="female">2 females</specimenCount>
(in
<collectionCode id="ED6EAE4EFF8EFF92385E616DFBF97664" box="[1033,1079,284,308]" pageId="27" pageNumber="368">IOZ</collectionCode>
) labeled “CASENT 1025836” and “CASENT 1025837”, respectively / “
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,
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,
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,
<location id="8EA06050FF8EFF923CCC612DFE6B7624" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923CCC612DFE6B7624" box="[155,421,348,372]" country="China" county="Gongshan County" latitude="27.97686" longLatPrecision="1" longitude="98.47799" municipality="Bingzhongluo Township" name="Bingzhongluo Township" pageId="27" pageNumber="368" stateProvince="Yunnan">Bingzhongluo Township</location>
, SW slope of Kawakarpu Shan,
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SW of Chukuai Lake
</locationDeviation>
at campsite,”/ “
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/
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,
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,
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<collectingDate id="EF85E9A3FF8EFF923E00610DFCCD76C4" box="[599,771,380,404]" pageId="27" pageNumber="368" value="2006-08-19">19 August 2006</collectingDate>
</date>
, Stop #DHK-2006-082
<collectorName id="268A535DFF8EFF923854610CFB8876C4" box="[1027,1094,380,404]" pageId="27" pageNumber="368">Y. Liu</collectorName>
collector”.
<collectorName id="268A535DFF8EFF923CE661EDFF1D76E4" box="[177,211,412,436]" pageId="27" pageNumber="368">All</collectorName>
<typeStatus id="54C48829FF8EFF923C8C61ECFE8F76E5" box="[219,321,413,437]" pageId="27" pageNumber="368" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="54C48829FF8EFF923EC361EDFCD576E4" box="[660,795,412,436]" pageId="27" pageNumber="368" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C7F4D08FF8EFF923F7461EDFF0F7684" authority="Deuve" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="368" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8EFF923F7461EDFC1976E4" box="[803,983,412,436]" italics="true" pageId="27" pageNumber="368">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF8EFF923FB061ECFBEF76E4" box="[999,1057,413,436]" italics="true" pageId="27" pageNumber="368">s. str.</emphasis>
)
<emphasis id="B90BEA99FF8EFF92386661EDFB5876E4" box="[1073,1174,412,436]" italics="true" pageId="27" pageNumber="368">griswoldi</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh,
<taxonomicNameLabel id="A23857E2FF8EFF923D3C61CCFE717684" box="[363,447,445,468]" pageId="27" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
designated 2016” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC61ACFC807545" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<materialsCitation id="3B173CD6FF8EFF923CFC61ACFC847545" country="China" county="Gongshan County" elevation="3270" latitude="27.77437" location="Cikai Township" longLatPrecision="1" longitude="98.44793" pageId="27" pageNumber="368" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54C48829FF8EFF923CFC61ACFF2A76A4" box="[171,228,477,501]" pageId="27" pageNumber="368">TYPE</typeStatus>
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</emphasis>
.—
<collectingCountry id="F368761BFF8EFF923DD661ACFE0D76A5" box="[385,451,477,501]" name="China" pageId="27" pageNumber="368">China</collectingCountry>
,
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,
<collectingCounty id="62A14E07FF8EFF923E6761ACFD3B76A5" box="[560,757,477,501]" pageId="27" pageNumber="368">Gongshan County</collectingCounty>
,
<location id="8EA06050FF8EFF923F5661ACFC6076A5" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF8EFF923F5661ACFC6076A5" box="[769,942,477,501]" country="China" county="Gongshan County" latitude="27.77437" longLatPrecision="1" longitude="98.44793" name="Cikai Township" pageId="27" pageNumber="368" stateProvince="Yunnan">Cikai Township</location>
,
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<quantity id="4C879B6EFF8EFF923FED61ACFBC876A5" box="[954,1030,477,501]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="27" pageNumber="368" unit="km" value="0.1">0.1 km</quantity>
SE of Heipu Yakou
</locationDeviation>
, in valley below tunnel,
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/
<geoCoordinate id="EE4B504CFF8EFF923E0B618CFD247545" box="[604,746,509,533]" degrees="098.44793" direction="east" orientation="longitude" pageId="27" pageNumber="368" precision="1" value="98.44793">E098.44793°</geoCoordinate>
,
<quantity id="4C879B6EFF8EFF923EA1618CFC847545" box="[758,842,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="27" pageNumber="368" unit="m" value="3270.0">
<elevation id="0052D1B8FF8EFF923EA1618CFC847545" box="[758,842,509,533]" metricMagnitude="3" metricUnit="m" metricValue="3.27" pageId="27" pageNumber="368" unit="m" value="3270.0">3270 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC626CFD4375E6" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<emphasis id="B90BEA99FF8EFF923CFC626CFE217564" bold="true" box="[171,495,541,565]" pageId="27" pageNumber="368">
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<smallCapsWord id="8D26A057FF8EFF923DE56250FE217564" baselines="559" box="[434,495,545,564]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="27" pageNumber="368">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4C7F4D08FF8EFF923EA3626CFC977565" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[756,857,541,565]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="368" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF8EFF923EA3626CFC977565" box="[756,857,541,565]" italics="true" pageId="27" pageNumber="368">griswoldi</emphasis>
</taxonomicName>
, is the Latinized form (in the genitive case) of the surname of Charles E. Griswold, now Curator Emeritus and former Schlinger Chair of Arachnology at the
<collectingRegion id="49BBF869FF8EFF923DE0622FFDED7526" box="[439,547,606,630]" country="United States of America" name="California" pageId="27" pageNumber="368">California</collectingRegion>
Academy of Sciences, who participated in several of the expeditions to the study area and helped collect many carabid specimens for this project. We are pleased to name this elegant species in his honor.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8EFF923CFC62CFFD917488" pageId="27" pageNumber="368" type="diagnosis">
<paragraph id="8BC0368BFF8EFF923CFC62CFFD917488" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
<smallCapsWord id="8D26A057FF8EFF923CFC62CFFEEE7585" baselines="720,720" box="[171,288,702,726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="27" pageNumber="368">
<emphasis id="B90BEA99FF8EFF923CFC62CFFEEE7585" bold="true" box="[171,288,702,726]" pageId="27" pageNumber="368">DIAGNOSIS</emphasis>
</smallCapsWord>
.— Adults of this species (
<figureCitation id="13442A0EFF8EFF923E1062CFFD6D7587" box="[583,675,702,727]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="27" pageNumber="368">Fig. 12a</figureCitation>
) can be distinguished from those of all other species in the region by the following combination of character states: size medium (BL =
<quantity id="4C879B6EFF8EFF92380062AEFF1F7447" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="27" pageNumber="368" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3 to 4.8 mm</quantity>
), body dark reddish brown, very shiny; eyes small and convex, tempora slightly convex; pronotum markedly narrowed posteriorly, markedly convex, globulose, narrowly cordate, disc smooth, basal angles with distict digitorm projections, each side with a single midlateral and basolateral seta; elytra markedly convex, tear-shaped, discal striae effaced, with three discal setae in a row along the presumed location of stria 3, preapical seta absent; median lobe of male aedeagus (
<figureCitation id="13442A0EFF8EFF923CD463D1FF1074E8" box="[131,222,928,952]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="27" pageNumber="368">Fig. 12b</figureCitation>
) large, elongate, broadest at mid-shaft, with apex long and recurved dorsally, endophallus with internal sclerites acuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF8EFF953CFC6391FC797526" lastPageId="28" lastPageNumber="369" pageId="27" pageNumber="368" type="description">
<paragraph id="8BC0368BFF8EFF923CFC6391FF107369" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
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<emphasis id="B90BEA99FF8EFF923CFC6391FEF474A7" bold="true" box="[171,314,992,1016]" pageId="27" pageNumber="368">DESCRIPTION</emphasis>
</smallCapsWord>
.— Size medium, BL =
<quantity id="4C879B6EFF8EFF923E146390FD2C74A8" box="[579,738,993,1017]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="27" pageNumber="368" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3 to 4.8 mm</quantity>
. Color of body, antennae and legs dark reddish brown, palpi paler, yellowish tan, body surface very shiny, micaceous, smooth and glabrous.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC6430FBDE72CB" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">
Head. Moderate in size, eyes small and convex, their convexity greater than and their diameter about as long as tempora, the latter only slightly convex and glabrous. Frons with frontal furrows deep, rounded, and not interrupted posteriorly; two suporaorbital setae present, the anterior inserted opposite midpoint of eye, the posterior inserted in postocular groove. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible tridentate. Left mandible with a small, minutely tridentate process.
<collectingRegion id="49BBF869FF8EFF923EFF6492FCCC73AA" box="[680,770,1251,1274]" country="Myanmar" name="Chin" pageId="27" pageNumber="368">Mentum</collectingRegion>
and submentum fused.
<collectingRegion id="49BBF869FF8EFF9238536492FB9073AA" box="[1028,1118,1251,1274]" country="Myanmar" name="Chin" pageId="27" pageNumber="368">Mentum</collectingRegion>
with medial tooth wide, bifid or truncate, one half the length of the lateral lobes. Submentum with six setae anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae of moderate length, extended posteriorly almost to or slightly beyond basal one-fourth of elytra, with 3.5 antennomeres in females and 4.5 antennomeres in males extended beyond basal pronotal margin, antennomeres slightly broadened, antennomere 3 slightly longer than antennomeres 2 or 4.
</paragraph>
<paragraph id="8BC0368BFF8EFF923CFC65D5FE6371ED" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="368">Pronotum. Shape narrowly cordiform, only slightly transverse (ratio PW/PL = 1.1), widest at the anterior one-fourth, markedly narrowed posteriorly, lateral margins distinctly sinuate anterior to basal angles, the latter extended posteriorly as slender, pointed, digitiform processes. Disc markedly convex, globulose, smooth and glabrous, with median longitudinal impression very faintly impressed or effaced, basal fovea small and smooth; median basal area smooth; basal margin slightly convex and rounded. Lateral border of pronotum slender, distinctly defined only in anteri- or one-third to one-half, effaced in posterior one-half to two –thirds. Single midlateral setae on each side inserted at anterior one-third; single basolateral seta on each side, inserted on basal angle at base of digitform process.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC60AAFEE776E4" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Elytra. Elytral silhouette slightly tear-shaped, with humeri effaced, disc markedly convex, smooth and glabrous. All discal striae effaced except for faintly impressed parascutellar striole and a short, faintly impressed recurrent stria. Basal setiferous pore present. Three discal setae present, aligned along presumed track of stria 3. Preapicale seta absent, one or two apicoangular setules present. Umbilicate setal series with setae of humeral group equidistance from each other, with the first slightly more medially inserted than the others, setae of median group inserted distinctly posterior to middle.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC61CCFBFB76A5" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Legs. Moderately long but slender. Protibiae with longitudinal furrow, sparely pubescent apically on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC618CFBE07565" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of males apically with one pair of paramedial setae, females with two pairs.</paragraph>
<paragraph id="8BC0368BFF89FF953CFC624CFC797526" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
Male aedeagus. Median lobe (
<figureCitation id="13442A0EFF89FF953DB8624CFD847505" box="[495,586,573,597]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="28" pageNumber="369">Fig. 12b</figureCitation>
) large, elongate, broadest at mid-shaft, with apex long and recurved dorsally; endophallus with internal sclerites acuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C3656500FF89FF953CFC620FFC5F712D" pageId="28" pageNumber="369" type="discussion">
<paragraph id="8BC0368BFF89FF953CFC620FFD917407" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
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<emphasis id="B90BEA99FF89FF953CFC620FFEE675C4" bold="true" box="[171,296,638,662]" pageId="28" pageNumber="369">COMMENTS</emphasis>
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.— Based on similarities in the form of the aedeagus of males, this species is closely related to
<taxonomicName id="4C7F4D08FF89FF953D5362EFFDBE75E6" authority="Deuve" authorityName="Deuve" box="[260,624,670,694]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D5362EFFDEC75E6" box="[260,546,670,694]" italics="true" pageId="28" pageNumber="369">Queinnectrechus jiuhecola</emphasis>
Deuve
</taxonomicName>
&amp; Kavanaugh (
<bibRefCitation id="EFEE4B7AFF89FF953F4B62EFFC2E75E6" author="DEUVE, T. &amp; D. KAVANAUGH &amp; H. B. LIANG" box="[796,992,670,694]" pageId="28" pageNumber="369" pagination="171 - 178" refId="ref59155" refString="DEUVE, T., D. KAVANAUGH, AND H. B. LIANG. 2015. Trois Trechini nouveaux du Mont Laojun, pres de Lijiang, dans le Yunnan, Chine (Coleoptera, Caraboidea). Coleopteres 21: 171 - 178." type="journal article" year="2015">Deuve et al. 2015</bibRefCitation>
), described from Lijiang County, northwestern
<collectingRegion id="49BBF869FF89FF953D9562CEFDD97586" box="[450,535,703,726]" country="China" name="Yunnan" pageId="28" pageNumber="369">Yunnan</collectingRegion>
. However, members of this new species are distinguished from those of the latter by their much smaller size (BL =
<quantity id="4C879B6EFF89FF953EB762AFFCB875A6" box="[736,886,734,758]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.8" metricValueMin="5.3" pageId="28" pageNumber="369" unit="mm" value="5.55" valueMax="5.8" valueMin="5.3">5.3 to 5.8 mm</quantity>
in
<taxonomicName id="4C7F4D08FF89FF953FCE62AEFBD375A7" box="[921,1053,735,759]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953FCE62AEFBD375A7" box="[921,1053,735,759]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members), more elongate body form, more convex elytra, discal striae fully effaced (at least three medial striae evident in
<taxonomicName id="4C7F4D08FF89FF953D45636EFE567467" box="[274,408,799,823]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D45636EFE567467" box="[274,408,799,823]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members) and, most significantly, preapical seta absent (present near stria 2 subapically in
<taxonomicName id="4C7F4D08FF89FF953D366331FE287407" box="[353,486,831,855]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953D366331FE287407" box="[353,486,831,855]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
members).
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC6311FEB573C9" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
In a recent paper, Belousov and Kabak (2016) established a new genus,
<taxonomicName id="4C7F4D08FF89FF953F9F6311FBE97427" authorityName="Belousov and Kabak" authorityYear="2016" box="[968,1063,864,887]" class="Insecta" family="Carabidae" genus="Uenoites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953F9F6311FBE97427" box="[968,1063,864,887]" italics="true" pageId="28" pageNumber="369">Uenoites</emphasis>
</taxonomicName>
, in which they placed
<taxonomicName id="4C7F4D08FF89FF953CA963F1FE4C74C8" box="[254,386,896,920]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF953CA963F1FE4C74C8" box="[254,386,896,920]" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
, as well as three other species previously included in
<taxonomicName id="4C7F4D08FF89FF953F9C63F1FBF474C8" authorityName="Jeannel" authorityYear="1935" box="[971,1082,896,920]" class="Insecta" family="Carabidae" genus="Kozlovites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953F9C63F1FBF474C8" box="[971,1082,896,920]" italics="true" pageId="28" pageNumber="369">Kozlovites</emphasis>
</taxonomicName>
or
<taxonomicName id="4C7F4D08FF89FF95383763F1FEB174E8" authority="Ueno (1995)" authorityName="Ueno" authorityYear="1995" class="Insecta" family="Carabidae" genus="Deuveotrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF95383763F1FF2074E8" italics="true" pageId="28" pageNumber="369">Deuveotrechus</emphasis>
<bibRefCitation id="EFEE4B7AFF89FF953CA263D1FEB174E8" author="UENO, S. I." box="[245,383,928,952]" pageId="28" pageNumber="369" pagination="13 - 25" refId="ref61208" refString="UENO, S. I. 1995. The Trechinae (Coleoptera, Carabidae) from Northern Vietnam. I. Two new species from Mt. Tam Dao. Bulletin of the National Science Museum, Ser. A (Zool.) 21: 13 - 25." type="journal article" year="1995">Uéno (1995)</bibRefCitation>
</taxonomicName>
, based on the presence of a preapical seta, more convex tempora, and the median lobe of the male aedeagus not markedly hooked apically. However, all these features are plesiomorphic among trechines, which is problematic for demonstrating phylogenetic affinity of the included taxa. Clearly, phylogenetic relationships among these groups of species remain unresolved. As noted above, the genitalia of
<taxonomicName id="4C7F4D08FF89FF953E716450FD627369" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[550,684,1057,1081]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="griswoldi">
<emphasis id="B90BEA99FF89FF953E716450FD627369" box="[550,684,1057,1081]" italics="true" pageId="28" pageNumber="369">Q. griswoldi</emphasis>
</taxonomicName>
males are very similar to those of
<taxonomicName id="4C7F4D08FF89FF95384A6450FF5E7309" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="jiuhecola">
<emphasis id="B90BEA99FF89FF95384A6450FF5E7309" italics="true" pageId="28" pageNumber="369">Q. jiuhecola</emphasis>
</taxonomicName>
males, wherease those of the new species described below,
<taxonomicName id="4C7F4D08FF89FF953F776430FBA17309" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[800,1135,1089,1113]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953F776430FBA17309" box="[800,1135,1089,1113]" italics="true" pageId="28" pageNumber="369">Queinnectrechus gongshanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF9538206433FF657329" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
, are of a very different form, although both of the new species described here are members of genus
<taxonomicName id="4C7F4D08FF89FF953C9764F0FEBA73C9" authorityName="Deuve" authorityYear="1992" box="[192,372,1153,1177]" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEA99FF89FF953C9764F0FEBA73C9" box="[192,372,1153,1177]" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC64D3FBA672CB" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC64D3FE6A73E8" bold="true" box="[171,420,1186,1210]" pageId="28" pageNumber="369">
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<smallCapsWord id="8D26A057FF89FF953D4764D4FE6A73E8" baselines="1203" box="[272,420,1189,1208]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="28" pageNumber="369">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Members of this species have been found in a variety of microhabitats in the alpine zone near the crest of the Gaoligong
<collectingRegion id="49BBF869FF89FF953E8D64B3FCC1738A" box="[730,783,1218,1242]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
and the eastern slope, at elevations ranging from
<quantity id="4C879B6EFF89FF953D466493FE0F73AA" box="[273,449,1250,1274]" metricMagnitude="3" metricUnit="m" metricValue="3.61" metricValueMax="3.95" metricValueMin="3.27" pageId="28" pageNumber="369" unit="m" value="3610.0" valueMax="3950.0" valueMin="3270.0">3270 to 3950 m</quantity>
. Specimens were collected under stones in moist meadows and on tundra slopes and ridges with sparse to thick herbaceous vegetation (
<figureCitation id="13442A0EFF89FF953F346573FC05724A" box="[867,971,1282,1306]" captionStart="FIGURE" captionStartId="99.[171,183,1602,1621]" captionTargetBox="[133,1163,216,1584]" captionTargetId="figure-15@99.[133,1163,216,1585]" captionTargetPageId="99" captionText="FIGURE 37. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Lushui County, Pianma Township,9.3 km ENE of Pianma along road to Lushui at Changyan He, elevation 2460 m; habitat in which specimens of Trechepaphiopsis unisetosa (Deuve), T. uniporosa sp. nov., and Trechepaphiama gaoligong sp. nov. were collected. b. Gongshan County, Cikai Township, slope S of of Heipu Yakou, elevation 3370 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov., Q. balli sp. nov., and Trechus qiqiensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155358" httpUri="https://zenodo.org/record/13155358/files/figure.png" pageId="28" pageNumber="369">Figs. 37b</figureCitation>
,
<figureCitation id="13442A0EFF89FF953F806572FBF8724A" box="[983,1078,1282,1307]" captionStart="FIGURE" captionStartId="100.[175,187,1625,1644]" captionTargetBox="[130,1166,216,1606]" captionTargetId="figure-11@100.[130,1166,216,1606]" captionTargetPageId="100" captionText="FIGURE 38. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan 0.3 km SW of Chukuai Lake, elevation 3750 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanicus sp. nov. were collected. b. same area, but habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechepaphiopsis monochaeta sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155360" httpUri="https://zenodo.org/record/13155360/files/figure.png" pageId="28" pageNumber="369">38a, 38b</figureCitation>
,
<figureCitation id="13442A0EFF89FF9538146573FBA0724A" box="[1091,1134,1282,1306]" captionStart="FIGURE" captionStartId="101.[171,183,1609,1628]" captionTargetBox="[124,1172,216,1587]" captionTargetId="figure-15@101.[124,1172,216,1588]" captionTargetPageId="101" captionText="FIGURE 39. Photographs of habitats for trechine species in the Gaoligong Shan region. a. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan, 0.3 km NNE of Chukuai Lake, elevation 3745 m; habitat in which specimens of Queinnectrechus gongshanicus sp. nov., Q. balli sp. nov. and Trechus gongshanensis sp. nov. were collected. b. Gongshan County, Bingzhongluo Township, SW slope of Kawakarpu Shan on slope NE of Chukuai Lake, elevation 3950 m; habitat in which specimens of Queinnectrechus griswoldi sp. nov. and Trechus gongshanensis sp. nov. were collected. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.13155362" httpUri="https://zenodo.org/record/13155362/files/figure.png" pageId="28" pageNumber="369">39b</figureCitation>
), at the edges of small streams and seeps from talus slopes, and at the edges of
<taxonomicName id="4C7F4D08FF89FF953FC96552FBF2726B" box="[926,1084,1315,1339]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="28" pageNumber="369" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BEA99FF89FF953FC96552FBF2726B" box="[926,1084,1315,1339]" italics="true" pageId="28" pageNumber="369">Rhododendron</emphasis>
</taxonomicName>
thickets up to
<quantity id="4C879B6EFF89FF953CED6532FF36720A" box="[186,248,1347,1371]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="369" unit="m" value="1.5">1.5 m</quantity>
tall. Members of this species have been found syntopic with specimens of
<taxonomicName id="4C7F4D08FF89FF9538716532FE16722B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Queinnectrechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="balli" status="sp. nov." subGenus="Gaoligongtrechus">
<emphasis id="B90BEA99FF89FF9538716532FF07722B" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF89FF953C8F6512FE56722B" box="[216,408,1379,1403]" italics="true" pageId="28" pageNumber="369">Gaoligongtrechus</emphasis>
)
<emphasis id="B90BEA99FF89FF953DF06512FE16722B" box="[423,472,1379,1403]" italics="true" pageId="28" pageNumber="369">balli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953D886515FDFA722B" box="[479,564,1380,1403]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF953E696512FC12722B" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[574,988,1379,1403]" class="Insecta" family="Carabidae" genus="Queinnectrechus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanicus" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953E696512FD3C722B" box="[574,754,1379,1403]" italics="true" pageId="28" pageNumber="369">Queinnectrechus</emphasis>
(
<emphasis id="B90BEA99FF89FF953F556515FCF4722B" box="[770,826,1380,1403]" italics="true" pageId="28" pageNumber="369">s. str.</emphasis>
)
<emphasis id="B90BEA99FF89FF953F1E6512FC12722B" box="[841,988,1379,1403]" italics="true" pageId="28" pageNumber="369">gongshanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953FB56515FBF9722B" box="[994,1079,1380,1403]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF9538156512FED672CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="gongshanensis" status="sp. nov.">
<emphasis id="B90BEA99FF89FF9538156512FED672CB" italics="true" pageId="28" pageNumber="369">Trechus gongshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953D4865F5FEBB72CB" box="[287,373,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C7F4D08FF89FF953DD765F2FDF772CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[384,569,1411,1435]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="qiqiensis" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953DD765F2FDF772CB" box="[384,569,1411,1435]" italics="true" pageId="28" pageNumber="369">Trechus qiqiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF953E1665F5FD5A72CB" box="[577,660,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C7F4D08FF89FF953E9F65F2FBC372CB" authorityName="Deuve &amp; Kavanaugh &amp; Liang" authorityYear="2016" box="[712,1037,1411,1435]" class="Insecta" family="Carabidae" genus="Trechepaphiopsis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="369" phylum="Arthropoda" rank="species" species="monochaeta" status="sp. nov.">
<emphasis id="B90BEA99FF89FF953E9F65F2FBC372CB" box="[712,1037,1411,1435]" italics="true" pageId="28" pageNumber="369">Trechepaphiopsis monochaeta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857E2FF89FF95384265F5FBA672CB" box="[1045,1128,1412,1435]" pageId="28" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC65D2FF04716D" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC65D2FD4E72EA" bold="true" box="[171,640,1443,1467]" pageId="28" pageNumber="369">
<smallCapsWord id="8D26A057FF89FF953CFC65D2FE9872EA" baselines="1461,1461" box="[171,342,1443,1467]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="28" pageNumber="369">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953D0865D6FE3D72EA" baselines="1461" box="[351,499,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="28" pageNumber="369">DISTRIBUTION</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953DAA65D6FD8272EA" baselines="1461" box="[509,588,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="within" pageId="28" pageNumber="369">WITHIN</smallCapsWord>
<smallCapsWord id="8D26A057FF89FF953E0165D6FD4E72EA" baselines="1461" box="[598,640,1447,1466]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="the" pageId="28" pageNumber="369">THE</smallCapsWord>
</emphasis>
<smallCapsWord id="8D26A057FF89FF953EDD65D2FCC172EA" baselines="1461,1461" box="[650,783,1443,1467]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Gaoligong" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953EDD65D2FCC172EA" bold="true" box="[650,783,1443,1467]" pageId="28" pageNumber="369">GAOLIGONG</emphasis>
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<materialsCitation id="3B173CD6FF89FF953F4E65D2FB9D728C" location="We" pageId="28" pageNumber="369" specimenCount="42" specimenCount-female="11" specimenCount-male="10" stateProvince="Shan">
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<emphasis id="B90BEA99FF89FF953F4E65D2FC9A72EA" bold="true" box="[793,852,1443,1467]" pageId="28" pageNumber="369">
<collectingRegion id="49BBF869FF89FF953F4E65D2FC9A72EA" box="[793,852,1443,1467]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">SHAN</collectingRegion>
</emphasis>
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.—
<figureCitation id="13442A0EFF89FF953FD765D5FC1072EC" box="[896,990,1444,1468]" captionStart="FIGURE" captionStartId="26.[171,183,1585,1604]" captionTargetBox="[147,1176,216,1569]" captionTargetId="figure-11@26.[120,1176,216,1569]" captionTargetPageId="26" captionText="FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q. griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km." figureDoi="http://doi.org/10.5281/zenodo.13155305" httpUri="https://zenodo.org/record/13155305/files/figure.png" pageId="28" pageNumber="369">Fig. 12c</figureCitation>
.
<location id="8EA06050FF89FF953FBC65D5FBC072EB" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF89FF953FBC65D5FBC072EB" box="[1003,1038,1444,1467]" name="We" pageId="28" pageNumber="369" stateProvince="Shan">We</location>
examined a total of
<specimenCount id="9D79FD02FF89FF953C9965B5FE90728C" box="[206,350,1476,1500]" count="21" pageId="28" pageNumber="369" type="generic">21 specimens</specimenCount>
(
<specimenCount id="9D79FD02FF89FF953D3A65B5FE03728C" box="[365,461,1476,1500]" count="10" pageId="28" pageNumber="369" type="male">10 males</specimenCount>
and
<specimenCount id="9D79FD02FF89FF953E5665B5FDB6728C" box="[513,632,1476,1500]" count="11" pageId="28" pageNumber="369" type="female">11 females</specimenCount>
), all from the northern part of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF89FF95380D65B5FC31714C" county="Gongshan County" location="Cikai Townships" municipality="Bingzhongluo" pageId="28" pageNumber="369" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF89FF95380D65B5FB5C728C" box="[1114,1170,1476,1500]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
, in
<collectingMunicipality id="6BA4ACF1FF89FF953CC06595FEE072AC" box="[151,302,1508,1532]" pageId="28" pageNumber="369">Bingzhongluo</collectingMunicipality>
and
<location id="8EA06050FF89FF953D336595FDD572AC" LSID="urn:lsid:plazi:treatment:03D6879DFF8CFF953C2C644AFC5F712D:8EA06050FF89FF953D336595FDD572AC" box="[356,539,1508,1532]" county="Gongshan County" municipality="Bingzhongluo" name="Cikai Townships" pageId="28" pageNumber="369" stateProvince="Shan">Cikai Townships</location>
,
<collectingCounty id="62A14E07FF89FF953E716595FD2772AC" box="[550,745,1508,1532]" pageId="28" pageNumber="369">Gongshan County</collectingCounty>
(see
<typeStatus id="54C48829FF89FF953F756594FC9972AD" box="[802,855,1509,1533]" pageId="28" pageNumber="369">Type</typeStatus>
material above for exact collection data). These localities are all on the crest or eastern slope of the Gaoligong
</materialsCitation>
<materialsCitation id="3B173CD6FF89FF9538506675FF08716D" location="Core Area" pageId="28" pageNumber="369" specimenCount="1" stateProvince="Shan">
<collectingRegion id="49BBF869FF89FF9538506675FBF2714C" box="[1031,1084,1540,1564]" country="Myanmar" name="Shan" pageId="28" pageNumber="369">Shan</collectingRegion>
in
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2
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0368BFF89FF953CFC6635FC5F712D" blockId="28.[123,1174,219,1661]" pageId="28" pageNumber="369">
<emphasis id="B90BEA99FF89FF953CFC6635FDAF710B" bold="true" box="[171,609,1604,1628]" pageId="28" pageNumber="369">
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<smallCapsWord id="8D26A057FF89FF953D4A6639FE0A710B" baselines="1622" box="[285,452,1608,1627]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="geographical" pageId="28" pageNumber="369">GEOGRAPHICAL</smallCapsWord>
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</emphasis>
.— This species currently is known only from the northern part of the Gaoligong Shan, in western
<collectingRegion id="49BBF869FF89FF953ED36614FCF1712D" box="[644,831,1637,1661]" country="China" name="Yunnan" pageId="28" pageNumber="369">Yunnan Province</collectingRegion>
,
<collectingCountry id="F368761BFF89FF953F1D6614FC43712D" box="[842,909,1637,1661]" name="China" pageId="28" pageNumber="369">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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