674 lines
86 KiB
XML
674 lines
86 KiB
XML
<document id="5E821C6971284C8D18F6CAE019681A80" ID-DOI="10.17348/jbrit.v15.i2.1206" ID-Zenodo-Dep="14111022" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634280027544" checkinUser="donat" docAuthor="Fawcett, Susan & Smith, Alan R." docDate="2021" docId="03B787F6FFBA9B1F623F7DB6FE48F9CC" docLanguage="en" docName="2021_ThelypteridaceaeFerns_OACopy3.pdf.imf" docOrigin="Fort Worth, Texas, USA: BRIT Press" docTitle="Pseudocyclosorus" docType="treatment" docVersion="3" lastPageNumber="74" masterDocId="FF8EFF8EFFE99B4B602B7D49FFA7FFBB" masterDocTitle="A Generic Classification of the Thelypteridaceae" pageNumber="73" updateTime="1731444927364" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
|
||
<mods:mods id="D88AADB55573895EA17517CC1491E5EB" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0F34AE0FD2CE7C472723ECF5EDFCCF5C">
|
||
<mods:title id="D3D2A446C3AE49C78BADBDF1DDD49FD7">A Generic Classification of the Thelypteridaceae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DE63EECB7B1187A9FB57673B8CD9B8C6" type="personal">
|
||
<mods:role id="F6C03FAFDE9D543984FA0661CC88DF20">
|
||
<mods:roleTerm id="DA7D6A9C8C52086144CEC2D325212514">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3433E7696367F9B6C0DC3990382A5006">Fawcett, Susan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0F3B6520681B7D02502C8B2D5A7747E4" type="personal">
|
||
<mods:role id="D115AA9362963CD8A82AC0EE74D4F418">
|
||
<mods:roleTerm id="E805CAD7C925032EF5CC5AE82ED969B1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0CA765427E63481B3A6E5E73A37F7579">Smith, Alan R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="2A7341C0AC263334FE78294C63885C13">text</mods:typeOfResource>
|
||
<mods:originInfo id="C17E1BA68C3E4016CE35A34455CA2160">
|
||
<mods:dateIssued id="D9F7D648405A62285F81255319EFC7BA">2021</mods:dateIssued>
|
||
<mods:publisher id="7019A803F2A256A5B12E9266EE43BECD">BRIT Press</mods:publisher>
|
||
<mods:place id="95A7018858DCCF65EF3BAB4CBD8E2919">
|
||
<mods:placeTerm id="641D187F5C29440D69479C8BE631B669">Fort Worth, Texas, USA</mods:placeTerm>
|
||
</mods:place>
|
||
</mods:originInfo>
|
||
<mods:classification id="412F05E517B6828B72E1220E7FE790C5">book</mods:classification>
|
||
<mods:identifier id="E53F59BF98E28B12979461EC61ED40A8" type="DOI">10.17348/jbrit.v15.i2.1206</mods:identifier>
|
||
<mods:identifier id="09E6DEF1CBC0DEF026D5CCBEEF787C8D" type="Zenodo-Dep">14111022</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B787F6FFBA9B1F623F7DB6FE48F9CC" LSID="urn:lsid:plazi:treatment:03B787F6FFBA9B1F623F7DB6FE48F9CC" httpUri="http://treatment.plazi.org/id/03B787F6FFBA9B1F623F7DB6FE48F9CC" lastPageId="84" lastPageNumber="74" pageId="83" pageNumber="73">
|
||
<subSubSection id="C304656BFFBA9B18623F7DB6FCE2FEA2" box="[532,837,255,281]" pageId="83" pageNumber="73" type="nomenclature">
|
||
<paragraph id="8BA136E0FFBA9B18623F7DB6FCE2FEA2" blockId="83.[121,1251,255,369]" box="[532,837,255,281]" pageId="83" pageNumber="73">
|
||
<heading id="D0E9818CFFBA9B18623F7DB6FCE2FEA2" allCaps="true" bold="true" box="[532,837,255,281]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="73" reason="0">
|
||
<taxonomicName id="4C1E4D63FFBA9B18623F7DB6FCE2FEA2" ID-CoL="6ZT7" box="[532,837,255,281]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B18623F7DB6FCE2FEA2" bold="true" box="[532,837,255,281]" pageId="83" pageNumber="73">PSEUDOCYCLOSORUS</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B1860527C6EFD50FE2C" pageId="83" pageNumber="73" type="reference_group">
|
||
<paragraph id="8BA136E0FFBA9B1860527C6EFE47FECA" blockId="83.[121,1251,255,369]" pageId="83" pageNumber="73">
|
||
<treatmentCitation id="0ABF10F1FFBA9B1860527C6EFD70FE86" author="Pseudocyclosorus Ching" box="[121,727,295,317]" journal="Acta Phytotax. Sin." page="322" pageId="83" pageNumber="73" score="3.863100345872848" volume="8" year="1963">
|
||
<taxonomicName id="4C1E4D63FFBA9B1860527C6EFD92FE86" ID-CoL="6ZT7" authority="Ching, Acta Phytotax. Sin." authorityName="Ching, Acta Phytotax. Sin." box="[121,565,295,317]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860527C6EFE88FE87" bold="true" box="[121,303,295,316]" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
Ching, Acta Phytotax. Sin.
|
||
</taxonomicName>
|
||
8:322–324. 1963
|
||
</treatmentCitation>
|
||
.—
|
||
<typeStatus id="54A58842FFBA9B1862C77C62FCABFE80" box="[748,780,299,315]" pageId="83" pageNumber="73">
|
||
<collectionCode id="ED0FAE25FFBA9B1862C77C62FCABFE80" box="[748,780,299,315]" pageId="83" pageNumber="73">TYPE</collectionCode>
|
||
</typeStatus>
|
||
:
|
||
<taxonomicName id="4C1E4D63FFBA9B18633E7C63FBEDFE87" ID-CoL="4NVFP" authority="(Kunze) Ching" authorityName="Ching" baseAuthorityName="Kunze" box="[789,1098,298,316]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBA9B18633E7C63FC74FE80" box="[789,979,298,315]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus tylodes</emphasis>
|
||
(Kunze) Ching
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="4C1E4D63FFBA9B1864497C63FF4CFEEC" ID-CoL="5VV53" authority="Kunze" authorityName="Kunze" class="Polypodiopsida" family="Thelypteridaceae" genus="Aspidium" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBA9B1864497C63FB46FE80" box="[1122,1249,298,315]" italics="true" pageId="83" pageNumber="73">Aspidium tylodes</emphasis>
|
||
Kunze
|
||
</taxonomicName>
|
||
]—
|
||
<taxonomicName id="4C1E4D63FFBA9B18612F7C0CFD3EFEEC" authority="(Ching) Panigrahi" authorityName="Panigrahi" baseAuthorityName="Ching" box="[260,665,325,343]" class="Polypodiopsida" family="Thelypteridaceae" genus="Cyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="subGenus" subGenus="Pseudocyclosorus">
|
||
<emphasis id="B96AEAF2FFBA9B18612F7C0CFEF0FEED" box="[260,343,325,342]" italics="true" pageId="83" pageNumber="73">Cyclosorus</emphasis>
|
||
subg.
|
||
<emphasis id="B96AEAF2FFBA9B1861AD7C0CFDACFEED" box="[390,523,325,342]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
(Ching) Panigrahi
|
||
</taxonomicName>
|
||
—
|
||
<taxonomicName id="4C1E4D63FFBA9B1862877C0CFBF7FEEC" ID-CoL="8H8B" authority="(Ching) Fraser-Jenk." authorityName="Fraser-Jenk." baseAuthorityName="Ching" box="[684,1104,325,343]" class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="section" section="Pseudocyclosorus">
|
||
<emphasis id="B96AEAF2FFBA9B1862877C0CFD58FEED" box="[684,767,325,342]" italics="true" pageId="83" pageNumber="73">Thelypteris</emphasis>
|
||
sect.
|
||
<emphasis id="B96AEAF2FFBA9B1863027C0CFC09FEED" box="[809,942,325,342]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
(Ching) Fraser-Jenk.
|
||
</taxonomicName>
|
||
—
|
||
<taxonomicName id="4C1E4D63FFBA9B18644A7C0CFE47FECA" ID-CoL="8H8B" authority="(Ching) Fraser-Jenk." class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="subGenus" subGenus="Pseudocyclosorus">
|
||
<emphasis id="B96AEAF2FFBA9B18644A7C0CFB13FEED" box="[1121,1204,325,342]" italics="true" pageId="83" pageNumber="73">Thelypteris</emphasis>
|
||
subg.
|
||
<emphasis id="B96AEAF2FFBA9B1860927C29FE99FECA" box="[185,318,352,369]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
(Ching) Fraser-Jenk.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BA136E0FFBA9B1860B27CCCFD50FE2C" blockId="83.[153,759,389,407]" box="[153,759,389,407]" pageId="83" pageNumber="73">
|
||
For synonymy, see
|
||
<bibRefCitation id="EF8F4B11FFBA9B1861037CCFFE00FE2C" author="Holttum" box="[296,423,390,407]" firstAuthor="Holttum" journalOrPublisher="Fern Gaz." pageId="83" pageNumber="73" pagination="55 - 56" part="11" refId="ref109658" refString="HOLTTUM, R. E. 1974 d. The alternative spellings Aspidium tylodes Kze. and A. xylodes Kze. Fern Gaz. 11: 55 - 56." title="The alternative spellings Aspidium tylodes Kze. and A. xylodes Kze" type="journal article" year="1974" yearSuffix="d">Holttum (1974d)</bibRefCitation>
|
||
,Holttum and
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862397CCFFD27FE2C" author="Grimes" box="[530,640,390,407]" firstAuthor="Grimes" journalOrPublisher="Kew Bull." pageId="83" pageNumber="73" pagination="517 - 520" part="34" refId="ref109072" refString="GRIMES, J. W. 1980. Spore morphology in Thelypteridaceae I: Pseudocyclosorus. Kew Bull. 34: 517 - 520." title="Spore morphology in Thelypteridaceae I: Pseudocyclosorus" type="journal article" year="1980">Grimes (1980)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862AC7CCFFD54FE2C" author="Li" box="[647,755,389,407]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="83" pageNumber="73" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al.(2019)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B1860527CE5FE8BFE5A" pageId="83" pageNumber="73" type="etymology">
|
||
<paragraph id="8BA136E0FFBA9B1860527CE5FE8BFE5A" blockId="83.[121,1252,428,1826]" pageId="83" pageNumber="73">
|
||
<emphasis id="B96AEAF2FFBA9B1860527CE5FF52FE7A" box="[121,245,428,449]" italics="true" pageId="83" pageNumber="73">Etymology.—</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBA9B1860DE7CE5FE68FE79" authority="Latin" authorityName="Latin" box="[245,463,428,450]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860DE7CE5FE30FE7A" box="[245,407,428,449]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
Latin
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAF2FFBA9B1861FF7CE5FDB5FE7A" box="[468,530,428,449]" italics="true" pageId="83" pageNumber="73">pseudo</emphasis>
|
||
, false +
|
||
<taxonomicName id="4C1E4D63FFBA9B18624B7CE5FD51FE79" authority="(Gr.)" baseAuthorityName="Gr." box="[608,758,428,450]" class="Polypodiopsida" family="Thelypteridaceae" genus="Cyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B18624B7CE5FD64FE7A" box="[608,707,428,449]" italics="true" pageId="83" pageNumber="73">Cyclosorus</emphasis>
|
||
(Gr.)
|
||
</taxonomicName>
|
||
; a distinct genus segregated from a broadly circumscribed
|
||
<taxonomicName id="4C1E4D63FFBA9B1860EF7C85FE81FE5A" box="[196,294,460,481]" class="Polypodiopsida" family="Thelypteridaceae" genus="Cyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860EF7C85FE81FE5A" box="[196,294,460,481]" italics="true" pageId="83" pageNumber="73">Cyclosorus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B1860837CA4FD26FB19" pageId="83" pageNumber="73" type="description">
|
||
<paragraph id="8BA136E0FFBA9B1860837CA4FD26FB19" blockId="83.[121,1252,428,1826]" pageId="83" pageNumber="73">
|
||
Plants terrestrial, often along streamsides, or shady forest understories, medium sized,
|
||
<quantity id="4CE69B05FFBA9B1863C37CA4FBF4FDBA" box="[1000,1107,492,514]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="83" pageNumber="73" unit="cm" value="95.0" valueMax="150.0" valueMin="40.0">40–150 cm</quantity>
|
||
tall;
|
||
<emphasis id="B96AEAF2FFBA9B1864A87CA4FB46FDB9" bold="true" box="[1155,1249,493,514]" pageId="83" pageNumber="73">rhizomes</emphasis>
|
||
short-creeping or forming erect caudices (e.g.,
|
||
<taxonomicName id="4C1E4D63FFBA9B1862057F44FD27FD9A" box="[558,640,524,546]" class="Polypodiopsida" family="Thelypteridaceae" genus="Aspidium" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBA9B1862057F44FD27FD9A" box="[558,640,524,546]" italics="true" pageId="83" pageNumber="73">P.tylodes</emphasis>
|
||
</taxonomicName>
|
||
); scales light brown, glabrous;
|
||
<emphasis id="B96AEAF2FFBA9B18638E7F44FC4FFD99" bold="true" box="[933,1000,525,546]" pageId="83" pageNumber="73">fronds</emphasis>
|
||
monomorphic, arching to erect;
|
||
<emphasis id="B96AEAF2FFBA9B1860997F64FF4AFDF9" bold="true" box="[178,237,557,578]" pageId="83" pageNumber="73">stipes</emphasis>
|
||
stramineous, castaneous, or dull brown;
|
||
<emphasis id="B96AEAF2FFBA9B1862537F64FD4CFDFA" bold="true" box="[632,747,556,578]" pageId="83" pageNumber="73">stipe scales</emphasis>
|
||
often sparse or caducous, when present, brown, glabrous or with marginal hairs, ovate to lanceolate;
|
||
<emphasis id="B96AEAF2FFBA9B1862767F05FD38FDDA" bold="true" box="[605,671,588,609]" pageId="83" pageNumber="73">blades</emphasis>
|
||
chartaceous, drying green, brown, or olivaceous, gradually reduced towards apex, proximal pinnae abruptly reduced to a series of opposite auriculate pinnae, these often caudate and/or hastate (see fig.
|
||
<quantity id="4CE69B05FFBA9B1861707FC4FE25FD19" box="[347,386,653,674]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="83" pageNumber="73" unit="in" value="2.0">2 in</quantity>
|
||
<bibRefCitation id="EF8F4B11FFBA9B1861A37FC4FDAEFD19" author="Li" box="[392,521,652,674]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="83" pageNumber="73" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. 2019</bibRefCitation>
|
||
), or reduced to a series of peg-like aerophores (
|
||
<taxonomicName id="4C1E4D63FFBA9B1863FC7FC4FB8AFD1A" box="[983,1069,652,674]" class="Polypodiopsida" family="Thelypteridaceae" genus="Aspidium" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBA9B1863FC7FC4FB8AFD1A" box="[983,1069,652,674]" italics="true" pageId="83" pageNumber="73">P. tylodes</emphasis>
|
||
</taxonomicName>
|
||
); proliferous buds absent;
|
||
<emphasis id="B96AEAF2FFBA9B1860E87FE4FEAEFD79" bold="true" box="[195,265,685,706]" pageId="83" pageNumber="73">pinnae</emphasis>
|
||
sessile, or short-petiolulate, bases truncate, margins deeply lobed, lobes nearly perpendicular (e.g.,
|
||
<taxonomicName id="4C1E4D63FFBA9B1864FB7FE4FF6BFD5A" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBA9B1864FB7FE4FF6BFD5A" italics="true" pageId="83" pageNumber="73">P. esquirolii</emphasis>
|
||
</taxonomicName>
|
||
) to strongly oblique (<45 degrees from costae in
|
||
<taxonomicName id="4C1E4D63FFBA9B1862997F84FCB8FD5A" box="[690,799,716,738]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="falcilobus">
|
||
<emphasis id="B96AEAF2FFBA9B1862997F84FCB8FD5A" box="[690,799,716,738]" italics="true" pageId="83" pageNumber="73">P. falcilobus</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<emphasis id="B96AEAF2FFBA9B18631F7F84FCCEFD59" bold="true" box="[820,873,717,738]" pageId="83" pageNumber="73">veins</emphasis>
|
||
free, reaching sinus, translucent sinus membrane often with tuft of hairs;
|
||
<emphasis id="B96AEAF2FFBA9B1861FD7FA5FDEEFCBA" bold="true" box="[470,585,748,769]" pageId="83" pageNumber="73">aerophores</emphasis>
|
||
swollen and conspicuous, protruding from mucilage in developing fronds;
|
||
<emphasis id="B96AEAF2FFBA9B1860EE7E44FE35FC99" bold="true" box="[197,402,781,802]" pageId="83" pageNumber="73">indument abaxially</emphasis>
|
||
of spreading hairs restricted to axes, of various lengths, minute (<0.1 mm) to medium-sized (0.5 mm), stipes often with long tangled hairs; small spherical, sessile or stipitate golden (but never sulfurcolored) glands sometimes present on axes;
|
||
<emphasis id="B96AEAF2FFBA9B1862317E04FD47FCD9" bold="true" box="[538,736,845,866]" pageId="83" pageNumber="73">indument adaxially</emphasis>
|
||
of hyaline acicular hairs, restricted to axes, and sometimes also abundant on laminae between veins (
|
||
<taxonomicName id="4C1E4D63FFBA9B1862727E24FD03FC39" box="[601,676,877,898]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="canus">
|
||
<emphasis id="B96AEAF2FFBA9B1862727E24FD03FC39" box="[601,676,877,898]" italics="true" pageId="83" pageNumber="73">P. canus</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<emphasis id="B96AEAF2FFBA9B1862927E25FCB6FC3A" bold="true" box="[697,785,876,897]" pageId="83" pageNumber="73">pustules</emphasis>
|
||
lacking;
|
||
<emphasis id="B96AEAF2FFBA9B1863467E24FC32FC39" bold="true" box="[877,917,877,898]" pageId="83" pageNumber="73">sori</emphasis>
|
||
round, discrete, rarely confluent, medial, along costules, or inframarginal, sometimes restricted to distal portion of pinnae, always indusiate, indusium typically large, especially evident on young fronds, persistent, glabrous or with short hairs or glands;
|
||
<emphasis id="B96AEAF2FFBA9B1864627EE4FB09FC79" bold="true" box="[1097,1198,941,962]" pageId="83" pageNumber="73">sporangia</emphasis>
|
||
with capsules lacking glands or hairs, capitate hairs sometimes present on sporangial stalks;
|
||
<emphasis id="B96AEAF2FFBA9B18639F7E85FC51FC5A" bold="true" box="[948,1014,972,993]" pageId="83" pageNumber="73">spores</emphasis>
|
||
monolete with cristae or echinae (
|
||
<bibRefCitation id="EF8F4B11FFBA9B1860E57EA4FEEBFBB9" author="Grimes" box="[206,332,1005,1026]" firstAuthor="Grimes" journalOrPublisher="Kew Bull." pageId="83" pageNumber="73" pagination="517 - 520" part="34" refId="ref109072" refString="GRIMES, J. W. 1980. Spore morphology in Thelypteridaceae I: Pseudocyclosorus. Kew Bull. 34: 517 - 520." title="Spore morphology in Thelypteridaceae I: Pseudocyclosorus" type="journal article" year="1980">Grimes 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8F4B11FFBA9B18617F7EA4FE6AFBB9" author="Li" box="[340,461,1004,1026]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="83" pageNumber="73" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. 2019</bibRefCitation>
|
||
);
|
||
<emphasis id="B96AEAF2FFBA9B1861F07EA5FE41FBBA" box="[475,486,1004,1025]" italics="true" pageId="83" pageNumber="73">x</emphasis>
|
||
= 36 or 35, six species counted, diploids, triploids, and tetraploids known.
|
||
<collectionCode id="ED0FAE25FFBA9B1864887EA4FB14FBB9" box="[1187,1203,1005,1026]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="83" pageNumber="73" type="Herbarium">A</collectionCode>
|
||
base number of 35 has been published for three species, based on several counts from multiple authors, and if correct this represents a dysploid change from the ancestral base number 36, which is synapomorphic for the cyclosoroid clade.
|
||
<bibRefCitation id="EF8F4B11FFBA9B1860527904FF54FBD9" author="Loyal" box="[121,243,1101,1122]" firstAuthor="Loyal" journalOrPublisher="Proc. Indian Sci. Congr. Assoc." pageId="83" pageNumber="73" pagination="265 - 266" part="48" refId="ref111578" refString="LOYAL, D. S. 1961. Cytological studies in Himalayan Thelypteridaceae I. Cytology of Thelypteris Schmidel. Proc. Indian Sci. Congr. Assoc. 48: 265 - 266." title="Cytological studies in Himalayan Thelypteridaceae I. Cytology of Thelypteris Schmidel" type="journal article" year="1961">Loyal (1961)</bibRefCitation>
|
||
reported a triploid
|
||
<taxonomicName id="4C1E4D63FFBA9B18619A7905FE5AFBDA" box="[433,509,1100,1121]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="canus">
|
||
<emphasis id="B96AEAF2FFBA9B18619A7905FE5AFBDA" box="[433,509,1100,1121]" italics="true" pageId="83" pageNumber="73">P. canus</emphasis>
|
||
</taxonomicName>
|
||
with irregular meiosis from Darjeeling, representing a possible infrageneric hybrid. Shieh and Tsai (1987) reported a putative hybrid between
|
||
<taxonomicName id="4C1E4D63FFBA9B1862DD7925FBBDFB3A" box="[758,1050,1132,1153]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="erubescens">
|
||
<emphasis id="B96AEAF2FFBA9B1862DD7925FBBDFB3A" box="[758,1050,1132,1153]" italics="true" pageId="83" pageNumber="73">Glaphyropteridopsis erubescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D63FFBA9B1864617925FF6BFB1A" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocylosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBA9B1864617925FF6BFB1A" italics="true" pageId="83" pageNumber="73">Pseudocylosorus esquirolii</emphasis>
|
||
</taxonomicName>
|
||
, a finding that warrants further investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B18608279E5FC13FA79" pageId="83" pageNumber="73" type="diagnosis">
|
||
<paragraph id="8BA136E0FFBA9B18608279E5FC13FA79" blockId="83.[121,1252,428,1826]" pageId="83" pageNumber="73">
|
||
<emphasis id="B96AEAF2FFBA9B18608279E5FEA4FB7A" box="[169,259,1196,1217]" italics="true" pageId="83" pageNumber="73">Diagnosis</emphasis>
|
||
.—The combination of deeply dissected pinnae, swollen aerophores (most easily observed in fiddleheads and fresh material), large indusia, and proximal pinnae abruptly reduced to auricles (or aerophores in
|
||
<taxonomicName id="4C1E4D63FFBA9B1864177985FB29FB5A" box="[1084,1166,1228,1249]" class="Polypodiopsida" family="Thelypteridaceae" genus="Aspidium" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBA9B1864177985FB29FB5A" box="[1084,1166,1228,1249]" italics="true" pageId="83" pageNumber="73">P.tylodes</emphasis>
|
||
</taxonomicName>
|
||
) is diagnostic for
|
||
<taxonomicName id="4C1E4D63FFBA9B1860F779A5FED9FABA" box="[220,382,1260,1281]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860F779A5FED9FABA" box="[220,382,1260,1281]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C1E4D63FFBA9B1861A079A5FDA2FABA" box="[395,517,1260,1281]" class="Polypodiopsida" family="Thelypteridaceae" genus="Trigonospora" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1861A079A5FDA2FABA" box="[395,517,1260,1281]" italics="true" pageId="83" pageNumber="73">Trigonospora</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished by its trilete spores and erect (vs. usually creeping) rhizomes.
|
||
<taxonomicName id="4C1E4D63FFBA9B1860F47845FE9DFA9A" box="[223,314,1292,1313]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860F47845FE9DFA9A" box="[223,314,1292,1313]" italics="true" pageId="83" pageNumber="73">Christella</emphasis>
|
||
</taxonomicName>
|
||
rarely has deeply divided pinnae with free veins, and often bears orangish pear-shaped glands.
|
||
<taxonomicName id="4C1E4D63FFBA9B1860527865FF57FAFA" box="[121,240,1324,1345]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860527865FF57FAFA" box="[121,240,1324,1345]" italics="true" pageId="83" pageNumber="73">Plesioneuron</emphasis>
|
||
</taxonomicName>
|
||
has similar deeply incised pinnae with free veins, and large persistent indusia, but does not overlap in geographic distribution, and often has more thickly chartaceous (and sometimes pustulate) laminae, pinna bases often short-petiolulate and rounded or cuneate (vs. sessile and truncate), and stipe scales usually thick and persistent (vs. thin and caducous).
|
||
<taxonomicName id="4C1E4D63FFBA9B18612578C5FE15FA1A" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[270,434,1420,1441]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B18612578C5FE15FA1A" box="[270,434,1420,1441]" italics="true" pageId="83" pageNumber="73">Sphaerostephanos</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D63FFBA9B1861CA78C5FDC2FA1A" box="[481,613,1420,1441]" class="Polypodiopsida" family="Thelypteridaceae" genus="Strophocaulon" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1861CA78C5FDC2FA1A" box="[481,613,1420,1441]" italics="true" pageId="83" pageNumber="73">Strophocaulon</emphasis>
|
||
</taxonomicName>
|
||
both may have abruptly reduced auriculate pinnae, but often bear yellowish glands, and are largely non-overlapping with
|
||
<emphasis id="B96AEAF2FFBA9B1862AC78E5FCB9FA7A" box="[647,798,1452,1473]" italics="true" pageId="83" pageNumber="73">Pseudocylosorus</emphasis>
|
||
geographically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B1860827885FB81F9F9" pageId="83" pageNumber="73" type="biology_ecology">
|
||
<paragraph id="8BA136E0FFBA9B1860827885FB81F9F9" blockId="83.[121,1252,428,1826]" pageId="83" pageNumber="73">
|
||
<emphasis id="B96AEAF2FFBA9B1860827885FDF5FA5A" box="[169,594,1484,1505]" italics="true" pageId="83" pageNumber="73">
|
||
Biogeography and ecology.—
|
||
<taxonomicName id="4C1E4D63FFBA9B18619A7885FDF5FA5A" box="[433,594,1484,1505]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">Pseudocyclosorus</taxonomicName>
|
||
</emphasis>
|
||
, here circumscribed to include 14 species, is most diverse at middle elevations of the Pan-Himalayan region.
|
||
<emphasis id="B96AEAF2FFBA9B1861D278A5FD72F9BA" box="[505,725,1516,1537]" italics="true" pageId="83" pageNumber="73">
|
||
<taxonomicName id="4C1E4D63FFBA9B1861D278A5FD3EF9BA" box="[505,665,1516,1537]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">Pseudocyclosorus</taxonomicName>
|
||
tylode
|
||
</emphasis>
|
||
s occupies the most extensive range, occurring from
|
||
<collectingCountry id="F3097670FFBA9B1864ED78A4FF11F999" name="Sri Lanka" pageId="83" pageNumber="73">Sri Lanka</collectingCountry>
|
||
to the
|
||
<collectingCountry id="F3097670FFBA9B1860D27B44FECEF999" box="[249,361,1549,1570]" name="Philippines" pageId="83" pageNumber="73">Philippines</collectingCountry>
|
||
. Two species,
|
||
<taxonomicName id="4C1E4D63FFBA9B1861C47B45FDFEF99A" box="[495,601,1548,1569]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocylosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBA9B1861C47B45FDFEF99A" box="[495,601,1548,1569]" italics="true" pageId="83" pageNumber="73">P. esquirolii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B96AEAF2FFBA9B1862AC7B45FD5DF99A" box="[647,762,1548,1569]" italics="true" pageId="83" pageNumber="73">
|
||
<taxonomicName id="4C1E4D63FFBA9B1862AC7B45FD51F99A" box="[647,758,1548,1569]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="falcilobus">P.falcilobus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
reach as far north as
|
||
<collectingCountry id="F3097670FFBA9B1863E37B45FBA6F99A" box="[968,1025,1548,1569]" name="Japan" pageId="83" pageNumber="73">Japan</collectingCountry>
|
||
. Three species occur in Afro-Madagascar. Members of the genus are typically plants of streamsides and forest understories.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBA9B1F60827B05FEDEFD6C" lastPageId="84" lastPageNumber="74" pageId="83" pageNumber="73" type="discussion">
|
||
<paragraph id="8BA136E0FFBA9B1F60827B05FEB4FE2C" blockId="83.[121,1252,428,1826]" lastBlockId="84.[97,1226,161,1655]" lastPageId="84" lastPageNumber="74" pageId="83" pageNumber="73">
|
||
<emphasis id="B96AEAF2FFBA9B1860827B05FE56F9DA" box="[169,497,1612,1633]" italics="true" pageId="83" pageNumber="73">Taxonomic and phylogenetic studies</emphasis>
|
||
.—
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862257B04FD2DF9D9" author="Ching" box="[526,650,1613,1634]" firstAuthor="Ching" journalOrPublisher="Bull. Fan Mem. Inst. Biol." pageId="83" pageNumber="73" pagination="237 - 352" part="6" refId="ref107458" refString="CHING, R. C. 1936. A revision of the Chinese and Sikkim-Himalayan Dryopteris with reference to some species from neighboring regions. Bull. Fan Mem. Inst. Biol. 6: 237 - 352." title="A revision of the Chinese and Sikkim-Himalayan Dryopteris with reference to some species from neighboring regions" type="journal article" year="1936">Ching (1936</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862B87B04FD63F9D9" author="Ching" box="[659,708,1613,1634]" firstAuthor="Ching" journalOrPublisher="Sunyatsenia" pageId="83" pageNumber="73" pagination="201 - 268" part="5" refId="ref107498" refString="CHING, R. C. 1940. On natural classification of the family " Polypodiaceae ". Sunyatsenia 5: 201 - 268." title="On natural classification of the family " Polypodiaceae" type="journal article" year="1940">1940</bibRefCitation>
|
||
) recognized the species now treated in
|
||
<taxonomicName id="4C1E4D63FFBA9B18646A7B05FB46F9DA" box="[1089,1249,1612,1633]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B18646A7B05FB46F9DA" box="[1089,1249,1612,1633]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
as a natural group in early publications, but did not name the genus until later (
|
||
<bibRefCitation id="EF8F4B11FFBA9B1863517B24FC49F939" author="Ching" box="[890,1006,1645,1666]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="83" pageNumber="73" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching 1963</bibRefCitation>
|
||
). Based on similarities in venation,
|
||
<bibRefCitation id="EF8F4B11FFBA9B1860F27BC4FEFDF919" author="Ching" box="[217,346,1677,1698]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="83" pageNumber="73" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching (1963)</bibRefCitation>
|
||
believed
|
||
<taxonomicName id="4C1E4D63FFBA9B18619C7BC5FDD2F91A" box="[439,629,1676,1697]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pelazoneuron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="patens">
|
||
<emphasis id="B96AEAF2FFBA9B18619C7BC5FDD2F91A" box="[439,629,1676,1697]" italics="true" pageId="83" pageNumber="73">Pelazoneuron patens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D63FFBA9B18628D7BC5FD58F91A" box="[678,767,1676,1697]" class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="species" species="kunthii">
|
||
<emphasis id="B96AEAF2FFBA9B18628D7BC5FD58F91A" box="[678,767,1676,1697]" italics="true" pageId="83" pageNumber="73">P. kunthii</emphasis>
|
||
</taxonomicName>
|
||
might also be allied, but this is not supported by molecular evidence (
|
||
<bibRefCitation id="EF8F4B11FFBA9B1861177BE4FDB3F979" author="Smith & Cranfill" box="[316,532,1709,1730]" firstAuthor="Smith" journalOrPublisher="Amer. Fern J." pageId="83" pageNumber="73" pagination="131 - 150" part="92" refId="ref114628" refString="SMITH, A. R. & R. B. CRANFILL. 2002. Intrafamilial relationships of the thelypteroid ferns (Thelypteridaceae). Amer. Fern J. 92: 131 - 150." title="Intrafamilial relationships of the thelypteroid ferns (Thelypteridaceae)" type="journal article" year="2002">Smith & Cranfill 2002</bibRefCitation>
|
||
; Fawcett et al. in press). Species-level taxonomy of
|
||
<taxonomicName id="4C1E4D63FFBA9B1863C77BE5FB2BF97A" box="[1004,1164,1708,1729]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1863C77BE5FB2BF97A" box="[1004,1164,1708,1729]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
is highly variable among authors.
|
||
<taxonomicName id="4C1E4D63FFBA9B1861437B85FD23F959" authority="sensu Ching" authorityName="Ching" box="[360,644,1740,1762]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1861437B85FDE6F95A" box="[360,577,1740,1761]" italics="true" pageId="83" pageNumber="73">Pseudocyclosorus sensu</emphasis>
|
||
Ching
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862BA7B84FCA2F959" author="Ching" box="[657,773,1741,1762]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="83" pageNumber="73" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching 1963</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8F4B11FFBA9B1863247B84FC3CF959" author="Lin" box="[783,923,1740,1762]" etAl="et al." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A." pageId="83" pageNumber="73" pagination="319 - 396" refId="ref111240" refString="LIN, Y. X., Z. Y. LI, K. IWATSUKI, & A. R. SMITH. 2013. Thelypteridaceae. In Z. Y. Wu, P. H. Raven, & D. Y. Hong, eds., Flora of China, Vol. 2 - 3 (Pteridophytes). Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A. Pp. 319 - 396." title="Thelypteridaceae" type="book chapter" volumeTitle="Z. Y. Wu, P. H. Raven, & D. Y. Hong" year="2013">Lin et al. 2013</bibRefCitation>
|
||
) included two Chinese species of
|
||
<taxonomicName id="4C1E4D63FFBA9B1860527BA5FF52F8BA" box="[121,245,1772,1793]" class="Polypodiopsida" family="Thelypteridaceae" genus="Trigonospora" kingdom="Plantae" order="Polypodiales" pageId="83" pageNumber="73" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBA9B1860527BA5FF52F8BA" box="[121,245,1772,1793]" italics="true" pageId="83" pageNumber="73">Trigonospora</emphasis>
|
||
</taxonomicName>
|
||
, a genus that was segregated by
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862167BA4FD71F8B9" author="Holttum" box="[573,726,1773,1794]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="83" pageNumber="73" pagination="17 - 52" part="19" refId="ref109463" refString="HOLTTUM, R. E. 1971. Studies in the family Thelypteridaceae III. A new system of genera in the Old World. Blumea 19: 17 - 52." title="Studies in the family Thelypteridaceae III. A new system of genera in the Old World" type="journal article" year="1971">Holttum (1971)</bibRefCitation>
|
||
based on having trilete spores and lacking abruptly reduced proximal pinnae. Holttum and
|
||
<bibRefCitation id="EF8F4B11FFBA9B1862397A44FD01F899" author="Grimes" box="[530,678,1805,1826]" firstAuthor="Grimes" journalOrPublisher="Kew Bull." pageId="83" pageNumber="73" pagination="517 - 520" part="34" refId="ref109072" refString="GRIMES, J. W. 1980. Spore morphology in Thelypteridaceae I: Pseudocyclosorus. Kew Bull. 34: 517 - 520." title="Spore morphology in Thelypteridaceae I: Pseudocyclosorus" type="journal article" year="1980">Grimes (1980)</bibRefCitation>
|
||
recognized twelve species in their generic treatment of
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A7DE8FEA7FF0D" box="[97,256,161,182]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A7DE8FEA7FF0D" box="[97,256,161,182]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
. Later, the genus was greatly expanded, and 20 new species were described by Lin (in
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F64647DEBFF34FF6C" author="Shing & Lin" firstAuthor="Shing" journalOrPublisher="Flora Reipublicae Popularis Sinicae" pageId="84" pageNumber="74" pagination="1 - 398" part="4" refId="ref114070" refString="SHING, K. & Y. LIN. 1999. Pteridophyta: Hypodematiaceae, Thelypteridaceae. Flora Reipublicae Popularis Sinicae 4: 1 - 398, i-xi." title="Pteridophyta: Hypodematiaceae, Thelypteridaceae" type="journal article" year="1999">Shing & Lin 1999</bibRefCitation>
|
||
); 38 species are recognized in the Flora of
|
||
<collectingCountry id="F3097670FFBD9B1F621F7D8BFDD5FF6C" box="[564,626,194,215]" name="China" pageId="84" pageNumber="74">China</collectingCountry>
|
||
, including the two Chinese species of
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63C17D8BFF3EFF4C" authority="(Lin et al. 2013)" baseAuthorityName="Lin" baseAuthorityYear="2013" class="Polypodiopsida" family="Thelypteridaceae" genus="Trigonospora" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63C17D8BFBC4FF6C" box="[1002,1123,194,215]" italics="true" pageId="84" pageNumber="74">Trigonospora</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F645B7D8BFF35FF4C" author="Lin" etAl="et al." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A." pageId="84" pageNumber="74" pagination="319 - 396" refId="ref111240" refString="LIN, Y. X., Z. Y. LI, K. IWATSUKI, & A. R. SMITH. 2013. Thelypteridaceae. In Z. Y. Wu, P. H. Raven, & D. Y. Hong, eds., Flora of China, Vol. 2 - 3 (Pteridophytes). Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A. Pp. 319 - 396." title="Thelypteridaceae" type="book chapter" volumeTitle="Z. Y. Wu, P. H. Raven, & D. Y. Hong" year="2013">Lin et al. 2013</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, in a recent revision of the species of the Pan-Himalayan region,
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F63437DABFC55FF4C" author="Li" box="[872,1010,226,247]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="84" pageNumber="74" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. (2019)</bibRefCitation>
|
||
recognized only eight species, reducing 21 names to synonymy. It is apparent that
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62847C48FCF7FEAD" box="[687,848,257,278]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62847C48FCF7FEAD" box="[687,848,257,278]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
would benefit from a densely sampled molecular study. For example, the species boundary between
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62857C6BFC9AFE8D" box="[686,829,289,311]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="subochthodes">
|
||
<emphasis id="B96AEAF2FFBD9B1F62857C6BFC9AFE8D" box="[686,829,289,311]" italics="true" pageId="84" pageNumber="74">P. subochthodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63407C6BFC73FE8D" box="[875,980,289,311]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocylosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBD9B1F63407C6BFC73FE8D" box="[875,980,289,311]" italics="true" pageId="84" pageNumber="74">P.esquirolii</emphasis>
|
||
</taxonomicName>
|
||
is unclear (
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F646A7C6BFB18FE8C" author="Li" box="[1089,1215,290,311]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="84" pageNumber="74" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. 2019</bibRefCitation>
|
||
), and phylogenetic studies that have included multiple accessions of
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62FF7C0BFC9BFEED" box="[724,828,321,343]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocylosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBD9B1F62FF7C0BFC9BFEED" box="[724,828,321,343]" italics="true" pageId="84" pageNumber="74">P.esquirolii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F636D7C0BFC55FEEC" author="He & Zhang" box="[838,1010,322,343]" firstAuthor="He" journalOrPublisher="Molec. Phylogen. Evol." pageId="84" pageNumber="74" pagination="757 - 764" part="65" refId="ref109255" refString="HE, L. J. & X. C. ZHANG. 2012. Exploring generic delimitation within the fern family Thelypteridaceae. Molec. Phylogen. Evol. 65: 757 - 764." title="Exploring generic delimitation within the fern family Thelypteridaceae" type="journal article" year="2012">He & Zhang 2012</bibRefCitation>
|
||
; Fawcett et al. in press) have resolved that species as paraphyletic; these results suggest that further study, especially of widespread and variable taxa, is warranted.
|
||
</paragraph>
|
||
<paragraph id="8BA136E0FFBD9B1F60A47CEBFB6FFDAC" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
Herein, we adopt a conservative, morphology-based classification largely following
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F63897CEBFB8EFE0C" author="Li" box="[930,1065,418,439]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="84" pageNumber="74" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. (2019)</bibRefCitation>
|
||
, pending further study and a greatly expanded phylogenetic dataset, preferably including taxa recognized by
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F63FD7C8BFBD6FE6C" author="Lin" box="[982,1137,450,471]" etAl="et al." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A." pageId="84" pageNumber="74" pagination="319 - 396" refId="ref111240" refString="LIN, Y. X., Z. Y. LI, K. IWATSUKI, & A. R. SMITH. 2013. Thelypteridaceae. In Z. Y. Wu, P. H. Raven, & D. Y. Hong, eds., Flora of China, Vol. 2 - 3 (Pteridophytes). Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A. Pp. 319 - 396." title="Thelypteridaceae" type="book chapter" volumeTitle="Z. Y. Wu, P. H. Raven, & D. Y. Hong" year="2013">Lin et al. (2013)</bibRefCitation>
|
||
. An even more conservative taxonomic concept of this group is favored for species of
|
||
<collectingCountry id="F3097670FFBD9B1F63057CABFCC6FE4C" box="[814,865,482,503]" name="India" pageId="84" pageNumber="74">India</collectingCountry>
|
||
by Fraser-Jenkins (1997, 2008b), who considered several species and varieties to be synonymous, and treated
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63287F48FC04FDAD" box="[771,931,513,534]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63287F48FC04FDAD" box="[771,931,513,534]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
as a subgenus of
|
||
<taxonomicName id="4C1E4D63FFBD9B1F64687F48FB0CFDAD" box="[1091,1195,513,534]" class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus" sensu="lato">
|
||
<emphasis id="B96AEAF2FFBD9B1F64687F48FB0CFDAD" box="[1091,1195,513,534]" italics="true" pageId="84" pageNumber="74">Thelypteris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2595789FFBD9B1F64847F48FB6FFDAC" box="[1199,1224,513,535]" pageId="84" pageNumber="74" sensu="lato">
|
||
<emphasis id="B96AEAF2FFBD9B1F64847F48FB64FDAD" box="[1199,1219,513,534]" italics="true" pageId="84" pageNumber="74">s.l</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BA136E0FFBD9B1F60BA7F68FEDEFD6C" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
<taxonomicName id="4C1E4D63FFBD9B1F60BA7F68FE96FD8D" ID-CoL="6ZT7" box="[145,305,545,566]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F60BA7F68FE96FD8D" box="[145,305,545,566]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
is sister to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61B47F6BFDB0FD8C" ID-CoL="823C" box="[415,535,546,567]" class="Polypodiopsida" family="Thelypteridaceae" genus="Trigonospora" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F61B47F6BFDB0FD8C" box="[415,535,546,567]" italics="true" pageId="84" pageNumber="74">Trigonospora</emphasis>
|
||
</taxonomicName>
|
||
plus a clade of two species from
|
||
<collectingCountry id="F3097670FFBD9B1F63717F6BFC69FD8C" box="[858,974,546,567]" name="Madagascar" pageId="84" pageNumber="74">Madagascar</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63FC7F6BFB6FFD8D" ID-CoL="4KKSY" baseAuthorityName="Holttum" baseAuthorityYear="1974" box="[983,1224,545,567]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="humbertii">
|
||
<emphasis id="B96AEAF2FFBD9B1F63FC7F6BFB6FFD8D" box="[983,1224,545,567]" italics="true" pageId="84" pageNumber="74">Pneumatopteris humbertii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B96AEAF2FFBD9B1F60A17F08FE99FDED" box="[138,318,577,598]" italics="true" pageId="84" pageNumber="74">
|
||
<taxonomicName id="4C1E4D63FFBD9B1F60A17F08FEA4FDED" ID-CoL="6WT2" authorityName="sensu Holttum. These" authorityYear="1982" box="[138,259,577,598]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pronephrium" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">Pronephrium</taxonomicName>
|
||
fidelei
|
||
</emphasis>
|
||
. This clade is in turn sister to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62777F08FD6EFDED" ID-CoL="8RG5Z" box="[604,713,577,598]" class="Polypodiopsida" family="Thelypteridaceae" genus="Abacopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62777F08FD6EFDED" box="[604,713,577,598]" italics="true" pageId="84" pageNumber="74">Abacopteris</emphasis>
|
||
</taxonomicName>
|
||
, treated herein as a small genus of Southeast Asia and eastern
|
||
<collectingCountry id="F3097670FFBD9B1F60857F2BFF5CFDCC" box="[174,251,610,631]" name="Malaysia" pageId="84" pageNumber="74">Malesia</collectingCountry>
|
||
. Two species here tentatively maintained in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62807F28FCA1FDCD" ID-CoL="3P92" box="[683,774,609,630]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBD9B1F62807F28FCA1FDCD" box="[683,774,609,630]" italics="true" pageId="84" pageNumber="74">Christella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2595789FFBD9B1F63277F2BFC80FDCC" box="[780,807,610,631]" pageId="84" pageNumber="74" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBD9B1F63277F2BFC86FDCC" box="[780,801,610,631]" italics="true" pageId="84" pageNumber="74">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63187F2BFC03FDCD" ID-CoL="5YGPW" box="[819,932,609,631]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="chaseana">
|
||
<emphasis id="B96AEAF2FFBD9B1F63187F2BFC03FDCD" box="[819,932,609,631]" italics="true" pageId="84" pageNumber="74">C. chaseana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B96AEAF2FFBD9B1F63FE7F2BFBF9FDCC" box="[981,1118,610,631]" italics="true" pageId="84" pageNumber="74">
|
||
<collectionCode id="ED0FAE25FFBD9B1F63FE7F2BFC40FDCC" box="[981,999,610,631]" country="Denmark" name="University of Copenhagen" pageId="84" pageNumber="74" type="Herbarium">C</collectionCode>
|
||
. gueintziana
|
||
</emphasis>
|
||
, resolve in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A7FC8FEA6FD2D" ID-CoL="6ZT7" box="[97,257,641,662]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A7FC8FEA6FD2D" box="[97,257,641,662]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
in some individual gene trees (Fawcett et al. in press), suggesting possible hybrid origin. They are not closely related to other species of
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61897FE8FDBAFD0D" ID-CoL="8RTTY" box="[418,541,673,694]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pelazoneuron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F61897FE8FDBAFD0D" box="[418,541,673,694]" italics="true" pageId="84" pageNumber="74">Pelazoneuron</emphasis>
|
||
</taxonomicName>
|
||
, where
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F62437FEBFCAFFD0C" author="Holttum" box="[616,776,674,695]" firstAuthor="Holttum" journalOrPublisher="J. S. African Bot." pageId="84" pageNumber="74" pagination="123 - 168" part="40" refId="ref109577" refString="HOLTTUM, R. E. 1974 a. Thelypteridaceae of Africa and adjacent islands. J. S. African Bot. 40: 123 - 168." title="Thelypteridaceae of Africa and adjacent islands" type="journal article" year="1974" yearSuffix="a">Holttum (1974a)</bibRefCitation>
|
||
treated them, despite striking superficial similarity (
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F60B47F8BFEC8FD6C" author="Moran & Smith" box="[159,367,706,727]" firstAuthor="Moran" journalOrPublisher="Brittonia" pageId="84" pageNumber="74" pagination="304 - 351" part="53" refId="ref112141" refString="MORAN, R. C. & A. R. SMITH. 2001. Phytogeographic relationships between neotropical and African-Madagascan pteridophytes. Brittonia 53: 304 - 351." title="Phytogeographic relationships between neotropical and African-Madagascan pteridophytes" type="journal article" year="2001">Moran & Smith 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBD9B1F60BA7FA8FBB2FC0C" pageId="84" pageNumber="74" type="taxon_list">
|
||
<paragraph id="8BA136E0FFBD9B1F60BA7FA8FF56FCCC" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
<emphasis id="B96AEAF2FFBD9B1F60BA7FA8FEFEFD4D" box="[145,345,737,758]" italics="true" pageId="84" pageNumber="74">Constituent species.—</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61727FA8FD47FD4C" ID-CoL="78F48" authority="Holttum" authorityName="Holttum" box="[345,736,737,759]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="camerounensis">
|
||
<emphasis id="B96AEAF2FFBD9B1F61727FA8FD20FD4C" box="[345,647,737,759]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus camerounensis</emphasis>
|
||
Holttum
|
||
</taxonomicName>
|
||
; *
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62D97FABFBCBFD4C" ID-CoL="6WF3V" authority="(Baker) Holttum & J. W. Grimes" authorityName="Holttum & J. W. Grimes" baseAuthorityName="Baker" box="[754,1132,738,759]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="canus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62D97FABFD5BFD4C" box="[754,764,738,759]" italics="true" pageId="84" pageNumber="74">P</emphasis>
|
||
.
|
||
<emphasis id="B96AEAF2FFBD9B1F632D7FABFC9CFD4C" box="[774,827,738,759]" italics="true" pageId="84" pageNumber="74">canus</emphasis>
|
||
(Baker) Holttum & J.W. Grimes
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B96AEAF2FFBD9B1F645F7FA8FBD9FD4D" box="[1140,1150,737,758]" italics="true" pageId="84" pageNumber="74">*</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBD9B1F64567FA8FEBCFCAC" authority="(Christ) Ching" authorityName="Ching" baseAuthorityName="Christ" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocylosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="esquirolii">
|
||
<emphasis id="B96AEAF2FFBD9B1F64567FA8FF20FCAD" italics="true" pageId="84" pageNumber="74">P.esquirolii</emphasis>
|
||
(Christ) Ching
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B96AEAF2FFBD9B1F610F7E48FE89FCAD" box="[292,302,769,790]" italics="true" pageId="84" pageNumber="74">*</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61067E48FD89FCAC" ID-CoL="78F4B" authority="(Hook.) Ching" authorityName="Ching" baseAuthorityName="Hook." box="[301,558,769,791]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="falcilobus">
|
||
<emphasis id="B96AEAF2FFBD9B1F61067E48FE3CFCAD" box="[301,411,769,790]" italics="true" pageId="84" pageNumber="74">P.falcilobus</emphasis>
|
||
(Hook.) Ching
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F621C7E4BFCDAFCAC" ID-CoL="7C7R6" authority="Holttum & J. W. Grimes" authorityName="Holttum & J. W. Grimes" box="[567,893,769,791]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="gamblei">
|
||
<emphasis id="B96AEAF2FFBD9B1F621C7E4BFD32FCAD" box="[567,661,769,791]" italics="true" pageId="84" pageNumber="74">P. gamblei</emphasis>
|
||
Holttum & J.W. Grimes
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63AE7E4BFB62FCAC" ID-CoL="6WF4F" authority="Holttum & J. W. Grimes" authorityName="Holttum & J. W. Grimes" box="[901,1221,770,791]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="griseus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63AE7E4BFC7BFCAC" box="[901,988,770,791]" italics="true" pageId="84" pageNumber="74">P. griseus</emphasis>
|
||
Holttum & J.W. Grimes
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A7E6BFE85FC8C" ID-CoL="6WF4G" authority="Holttum" authorityName="Holttum" box="[97,290,801,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="johannae">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A7E6BFF6EFC8D" box="[97,201,801,823]" italics="true" pageId="84" pageNumber="74">P.johannae</emphasis>
|
||
Holttum
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61017E6BFDE3FC8C" ID-CoL="78F4K" authority="(Kunze) Holttum" authorityName="Holttum" baseAuthorityName="Kunze" box="[298,580,801,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="ochthodes">
|
||
<emphasis id="B96AEAF2FFBD9B1F61017E6BFE3EFC8D" box="[298,409,801,823]" italics="true" pageId="84" pageNumber="74">P. ochthodes</emphasis>
|
||
(Kunze) Holttum
|
||
</taxonomicName>
|
||
; *
|
||
<taxonomicName id="4C1E4D63FFBD9B1F627C7E6BFC0FFC8C" ID-CoL="4NVF2" authority="Holttum & J. W. Grimes" authorityName="Holttum & J. W. Grimes" box="[599,936,802,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="ornatipes">
|
||
<emphasis id="B96AEAF2FFBD9B1F627C7E6BFD65FC8C" box="[599,706,802,823]" italics="true" pageId="84" pageNumber="74">P.ornatipes</emphasis>
|
||
Holttum & J.W. Grimes
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F639A7E6BFB62FC8C" ID-CoL="4NVF6" authority="W. M. Chu" authorityName="W. M. Chu" box="[945,1221,801,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="pseudofalcilobus">
|
||
<emphasis id="B96AEAF2FFBD9B1F639A7E6BFBFBFC8D" box="[945,1116,801,823]" italics="true" pageId="84" pageNumber="74">P. pseudofalcilobus</emphasis>
|
||
W.M. Chu
|
||
</taxonomicName>
|
||
; *
|
||
<taxonomicName id="4C1E4D63FFBD9B1F60477E0BFE68FCEC" ID-CoL="4NVF8" authority="(Bory ex Willd.) Holttum" authorityName="Holttum" baseAuthorityName="Willd." box="[108,463,833,855]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="pulcher">
|
||
<emphasis id="B96AEAF2FFBD9B1F60477E0BFF6DFCED" box="[108,202,833,855]" italics="true" pageId="84" pageNumber="74">P. pulcher</emphasis>
|
||
(Bory ex Willd.) Holttum
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61F17E0BFCB3FCEC" ID-CoL="8RDP4" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" box="[474,788,834,855]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="stramineus">
|
||
<emphasis id="B96AEAF2FFBD9B1F61F17E0BFDFFFCEC" box="[474,600,834,855]" italics="true" pageId="84" pageNumber="74">P. stramineus</emphasis>
|
||
Ching ex Y.X. Lin
|
||
</taxonomicName>
|
||
; *
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63017E0BFBFDFCEC" ID-CoL="4NVFF" authority="(Ching) Ching" authorityName="Ching" baseAuthorityName="Ching" box="[810,1114,833,855]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="subochthodes">
|
||
<emphasis id="B96AEAF2FFBD9B1F63017E0BFC19FCED" box="[810,958,833,855]" italics="true" pageId="84" pageNumber="74">P. subochthodes</emphasis>
|
||
(Ching) Ching
|
||
</taxonomicName>
|
||
; *
|
||
<taxonomicName id="4C1E4D63FFBD9B1F645B7E0BFF56FCCC" ID-CoL="5VV53" authority="(Kunze) Ching" authorityName="Ching" baseAuthorityName="Kunze" class="Polypodiopsida" family="Thelypteridaceae" genus="Aspidium" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="tylodes">
|
||
<emphasis id="B96AEAF2FFBD9B1F645B7E0BFBDDFCEC" box="[1136,1146,834,855]" italics="true" pageId="84" pageNumber="74">P</emphasis>
|
||
.
|
||
<emphasis id="B96AEAF2FFBD9B1F64AC7E08FB6EFCED" box="[1159,1225,833,854]" italics="true" pageId="84" pageNumber="74">tylodes</emphasis>
|
||
(Kunze) Ching
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BA136E0FFBD9B1F60BA7ECBFBB2FC0C" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
<emphasis id="B96AEAF2FFBD9B1F60BA7ECBFE16FC2D" box="[145,433,897,919]" italics="true" pageId="84" pageNumber="74">
|
||
Taxa treated in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F610B7EC8FE31FC2D" box="[288,406,897,918]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">Plesioneuron</taxonomicName>
|
||
.—
|
||
</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBD9B1F619A7EC8FCA3FC2C" authority="Holttum" authorityName="Holttum" box="[433,772,897,919]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="caudatus">
|
||
<emphasis id="B96AEAF2FFBD9B1F619A7EC8FD0BFC2D" box="[433,684,897,918]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus caudatus</emphasis>
|
||
Holttum
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63267ECBFBA7FC2C" authority="Holttum. These" authorityName="Holttum. These" box="[781,1024,898,919]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="excisus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63267ECBFCC2FC2C" box="[781,869,898,919]" italics="true" pageId="84" pageNumber="74">P. excisus</emphasis>
|
||
Holttum. These
|
||
</taxonomicName>
|
||
were both described in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F60527EE8FE50FC0C" authority="(Holttum & Roy 1965)" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[121,503,929,951]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F60527EE8FEBEFC0D" box="[121,281,929,950]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F610F7EEBFE48FC0C" author="Holttum & Roy" box="[292,495,930,951]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="84" pageNumber="74" pagination="129 - 139" part="13" refId="ref110091" refString="HOLTTUM, R. E. & S. K. ROY. 1965. Cytological observations on ferns of New Guinea, with descriptions of new species. Blumea 13: 129 - 139." title="Cytological observations on ferns of New Guinea, with descriptions of new species" type="journal article" year="1965">Holttum & Roy 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and later transferred to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62F17EEBFBB6FC0C" authority="(Holttum 1973 a)" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[730,1041,930,951]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62F17EEBFCCCFC0C" box="[730,875,930,951]" italics="true" pageId="84" pageNumber="74">Pneumatopteris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F635D7EEBFBADFC0C" author="Holttum" box="[886,1034,930,951]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="84" pageNumber="74" pagination="293 - 325" part="21" refId="ref109522" refString="HOLTTUM, R. E. 1973 a. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293 - 325." title="Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai" type="journal article" year="1973" yearSuffix="a">Holttum 1973a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBD9B1F60BA7E8BFE48F9CC" pageId="84" pageNumber="74" type="discussion">
|
||
<paragraph id="8BA136E0FFBD9B1F60BA7E8BFE9FFBEC" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
<emphasis id="B96AEAF2FFBD9B1F60BA7E8BFD30FC6D" box="[145,663,961,983]" italics="true" pageId="84" pageNumber="74">
|
||
Names of uncertain status.—
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61BC7E88FD30FC6D" box="[407,663,961,982]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="duclouxii">Pseudocyclosorus duclouxii</taxonomicName>
|
||
</emphasis>
|
||
was treated as dubious by
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F63BC7E8BFB84FC6C" author="Li" box="[919,1059,962,983]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="84" pageNumber="74" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. (2019)</bibRefCitation>
|
||
;
|
||
<emphasis id="B96AEAF2FFBD9B1F64077E8BFB6EFC6D" box="[1068,1225,961,983]" italics="true" pageId="84" pageNumber="74">
|
||
<collectionCode id="ED0FAE25FFBD9B1F64077E8BFB9DFC6C" box="[1068,1082,962,983]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="84" pageNumber="74" type="Herbarium">P</collectionCode>
|
||
. furcovenulosus
|
||
</emphasis>
|
||
<collectionCode id="ED0FAE25FFBD9B1F604A7EABFFD5FC4C" box="[97,114,994,1015]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="84" pageNumber="74" type="Herbarium">Y</collectionCode>
|
||
.X. Lin,
|
||
<taxonomicName id="4C1E4D63FFBD9B1F609C7EABFDAFFC4C" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" box="[183,520,994,1015]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="guangxianensis">
|
||
<emphasis id="B96AEAF2FFBD9B1F609C7EABFEFDFC4C" box="[183,346,994,1015]" italics="true" pageId="84" pageNumber="74">P. guangxianensis</emphasis>
|
||
Ching ex Y.X. Lin
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1E4D63FFBD9B1F623B7EABFD62FC4C" authority="Ching" authorityName="Ching" box="[528,709,994,1015]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="pectinatus">
|
||
<emphasis id="B96AEAF2FFBD9B1F623B7EABFD25FC4C" box="[528,642,994,1015]" italics="true" pageId="84" pageNumber="74">P.pectinatus</emphasis>
|
||
Ching
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62DE7EABFB90FC4C" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" box="[757,1079,993,1015]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="submarginalis">
|
||
<emphasis id="B96AEAF2FFBD9B1F62DE7EABFC2CFC4D" box="[757,907,993,1015]" italics="true" pageId="84" pageNumber="74">P.submarginalis</emphasis>
|
||
Ching ex Y.X. Lin
|
||
</taxonomicName>
|
||
were treated in Flora of
|
||
<collectingCountry id="F3097670FFBD9B1F6084794BFF4AFBAC" box="[175,237,1026,1047]" name="China" pageId="84" pageNumber="74">China</collectingCountry>
|
||
, but are known from few collections that are unavailable for study, and were not included in the revision by
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F6055796BFEADFB8C" author="Li" box="[126,266,1058,1079]" etAl="et al." firstAuthor="Li" journalOrPublisher="Phytotaxa" pageId="84" pageNumber="74" pagination="197 - 216" part="424" refId="ref111093" refString="LI, Z. Y., X. C. ZHANG, Z. L. LIANG, & J. L. LEONARD. 2019. A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus. Phytotaxa 424: 197 - 216." title="A taxonomic revision of fern genus Pseudocyclosorus (Thelypteridaceae) from China and the Pan-Himalaya region, with special reference to the identity of Pseudocyclosorus stramineus" type="journal article" year="2019">Li et al. (2019)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C1E4D63FFBD9B1F61397968FD13FB8C" authority="Ching & Y. X. Lin" authorityName="Ching & Y. X. Lin" box="[274,692,1057,1079]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="tibeticus">
|
||
<emphasis id="B96AEAF2FFBD9B1F61397968FDA2FB8D" box="[274,517,1057,1078]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus tibeticus</emphasis>
|
||
Ching & Y.X. Lin
|
||
</taxonomicName>
|
||
was excluded from the Flora of
|
||
<collectingCountry id="F3097670FFBD9B1F63C1796BFB81FB8C" box="[1002,1062,1058,1079]" name="China" pageId="84" pageNumber="74">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F6419796BFB18FB8C" author="Lin" box="[1074,1215,1058,1079]" etAl="et al." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A." pageId="84" pageNumber="74" pagination="319 - 396" refId="ref111240" refString="LIN, Y. X., Z. Y. LI, K. IWATSUKI, & A. R. SMITH. 2013. Thelypteridaceae. In Z. Y. Wu, P. H. Raven, & D. Y. Hong, eds., Flora of China, Vol. 2 - 3 (Pteridophytes). Science Press, Beijing, China; Missouri Botanical Garden Press, St. Louis, U. S. A. Pp. 319 - 396." title="Thelypteridaceae" type="book chapter" volumeTitle="Z. Y. Wu, P. H. Raven, & D. Y. Hong" year="2013">Lin et al. 2013</bibRefCitation>
|
||
), pending further study.
|
||
</paragraph>
|
||
<paragraph id="8BA136E0FFBD9B1F60BA792BFE48F9CC" blockId="84.[97,1226,161,1655]" pageId="84" pageNumber="74">
|
||
<emphasis id="B96AEAF2FFBD9B1F60BA792BFE80FBCD" box="[145,295,1121,1143]" italics="true" pageId="84" pageNumber="74">Incertae sedis.—</emphasis>
|
||
<taxonomicName id="4C1E4D63FFBD9B1F610C7928FD72FBCC" authority="(Copel.) Holttum" authorityName="Holttum" baseAuthorityName="Copel." box="[295,725,1121,1143]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="petrophila">
|
||
<emphasis id="B96AEAF2FFBD9B1F610C7928FD8DFBCD" box="[295,554,1121,1142]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus petrophila</emphasis>
|
||
(Copel.) Holttum
|
||
</taxonomicName>
|
||
, from New
|
||
<collectingCountry id="F3097670FFBD9B1F636A792BFC20FBCC" box="[833,903,1122,1143]" name="Guinea-Bissau" pageId="84" pageNumber="74">Guinea</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F63B9792BFBF9FBCC" author="Holttum & Roy" box="[914,1118,1122,1143]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="84" pageNumber="74" pagination="129 - 139" part="13" refId="ref110091" refString="HOLTTUM, R. E. & S. K. ROY. 1965. Cytological observations on ferns of New Guinea, with descriptions of new species. Blumea 13: 129 - 139." title="Cytological observations on ferns of New Guinea, with descriptions of new species" type="journal article" year="1965">Holttum & Roy 1965</bibRefCitation>
|
||
), treated as
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A79CBFE5AFB2C" authority="(Copel.) Holttum" authorityName="Holttum" baseAuthorityName="Copel." box="[97,509,1153,1175]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="petrophila">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A79CBFEF3FB2D" box="[97,340,1153,1175]" italics="true" pageId="84" pageNumber="74">Pneumatopteris petrophila</emphasis>
|
||
(Copel.) Holttum
|
||
</taxonomicName>
|
||
in later works, shares the opposite, hastate-auricled reduced proximal pinnae of mainland
|
||
<taxonomicName id="4C1E4D63FFBD9B1F612B79E8FE07FB0D" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[256,416,1185,1206]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F612B79E8FE07FB0D" box="[256,416,1185,1206]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
and free veins shared among all species, but is
|
||
<taxonomicName id="4C1E4D63FFBD9B1F637279E8FC07FB0C" box="[857,928,1185,1207]" genus="The" pageId="84" pageNumber="4" rank="species" species="major">a major</taxonomicName>
|
||
geographic outlier. It may have closer affinities to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F613A7988FE21FB6D" box="[273,390,1217,1238]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F613A7988FE21FB6D" box="[273,390,1217,1238]" italics="true" pageId="84" pageNumber="74">Plesioneuron</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C1E4D63FFBD9B1F618E7988FDEEFB6D" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[421,585,1217,1238]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F618E7988FDEEFB6D" box="[421,585,1217,1238]" italics="true" pageId="84" pageNumber="74">Sphaerostephanos</emphasis>
|
||
</taxonomicName>
|
||
, but we do not believe it is close to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63B0798BFB8BFB6C" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[923,1068,1218,1239]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63B0798BFB8BFB6C" box="[923,1068,1218,1239]" italics="true" pageId="84" pageNumber="74">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
, where
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F645C798BFF0CFB4C" author="Holttum" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="84" pageNumber="74" pagination="293 - 325" part="21" refId="ref109522" refString="HOLTTUM, R. E. 1973 a. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293 - 325." title="Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai" type="journal article" year="1973" yearSuffix="a">Holttum (1973a)</bibRefCitation>
|
||
later treated it.
|
||
<taxonomicName id="4C1E4D63FFBD9B1F616779A8FC2DFB4C" authority="(Houlston & T. Moore) Holttum" authorityName="Holttum" baseAuthorityName="Houlston & T. Moore" box="[332,906,1249,1271]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pronephrium" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="articulatum">
|
||
<emphasis id="B96AEAF2FFBD9B1F616779A8FD98FB4D" box="[332,575,1249,1270]" italics="true" pageId="84" pageNumber="74">Pronephrium articulatum</emphasis>
|
||
(Houlston & T. Moore) Holttum
|
||
</taxonomicName>
|
||
was recovered as nested within
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A7848FEA6FAAD" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[97,257,1281,1302]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A7848FEA6FAAD" box="[97,257,1281,1302]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
in recent Bayesian phylogenetic analyses, with 99% posterior probability (
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F6394784BFBC7FAAC" author="Patel" box="[959,1120,1281,1303]" etAl="et al." firstAuthor="Patel" journalOrPublisher="Int. J. Pl. Sci." pageId="84" pageNumber="74" pagination="1016 - 1035" part="180" refId="ref112729" refString="PATEL, N., S. FAWCETT, M. SUNDUE, & J. BUDKE. 2019 a. Evolution of perine morphology in Thelypteridaceae. Int. J. Pl. Sci. 180: 1016 - 1035." title="Evolution of perine morphology in Thelypteridaceae" type="journal article" year="2019" yearSuffix="a">Patel et al. 2019a</bibRefCitation>
|
||
). However, it differs in having up to four pairs of anastomosing veins below the sinus, proximal sori sometimes elongate, and unusual venation not seen elsewhere in the family: abaxially, veins are prominent until reaching sori, and obscure beyond them. The large, persistent indusia agree well with
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62B57828FC99FACD" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[670,830,1377,1398]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62B57828FC99FACD" box="[670,830,1377,1398]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
but, pending further data confirming its placement in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F60C878C8FE25FA2D" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[227,386,1409,1430]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F60C878C8FE25FA2D" box="[227,386,1409,1430]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
, we refrain from making a new combination.
|
||
<collectionCode id="ED0FAE25FFBD9B1F631678CBFCEAFA2C" box="[829,845,1410,1431]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="84" pageNumber="74" type="Herbarium">A</collectionCode>
|
||
species of the
|
||
<collectingCountry id="F3097670FFBD9B1F63F278CBFB9BFA2C" box="[985,1084,1410,1431]" name="Seychelles" pageId="84" pageNumber="74">Seychelles</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3097670FFBD9B1F644078CBFB6DFA2C" box="[1131,1226,1410,1431]" name="Sao Tome and Principe" pageId="84" pageNumber="74">São Tomé</collectingCountry>
|
||
(see
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F60A578EBFE83FA0C" author="Holttum" box="[142,292,1442,1463]" firstAuthor="Holttum" journalOrPublisher="J. S. African Bot." pageId="84" pageNumber="74" pagination="123 - 168" part="40" refId="ref109577" refString="HOLTTUM, R. E. 1974 a. Thelypteridaceae of Africa and adjacent islands. J. S. African Bot. 40: 123 - 168." title="Thelypteridaceae of Africa and adjacent islands" type="journal article" year="1974" yearSuffix="a">Holttum 1974a</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C1E4D63FFBD9B1F611F78E8FD1AFA0C" authority="(Bojer) Holttum" authorityName="Holttum" baseAuthorityName="Bojer" box="[308,701,1441,1463]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="elatus">
|
||
<emphasis id="B96AEAF2FFBD9B1F611F78E8FDBDFA0D" box="[308,538,1441,1462]" italics="true" pageId="84" pageNumber="74">Sphaerostephanos elatus</emphasis>
|
||
(Bojer) Holttum
|
||
</taxonomicName>
|
||
is well supported as sister to
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63CE78E8FB2FFA0D" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[997,1160,1441,1462]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63CE78E8FB2FFA0D" box="[997,1160,1441,1462]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
in the nDNA analyses of Fawcett et al. (in press), but is well supported as sister to three species we now recognize as
|
||
<taxonomicName id="4C1E4D63FFBD9B1F64777888FB6EFA6D" box="[1116,1225,1473,1494]" class="Polypodiopsida" family="Thelypteridaceae" genus="Abacopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F64777888FB6EFA6D" box="[1116,1225,1473,1494]" italics="true" pageId="84" pageNumber="74">Abacopteris</emphasis>
|
||
</taxonomicName>
|
||
in studies relying primarily on cpDNA (e.g.,
|
||
<bibRefCitation id="EF8F4B11FFBD9B1F620B78ABFD6BFA4C" author="Patel" box="[544,716,1505,1527]" etAl="et al." firstAuthor="Patel" journalOrPublisher="Int. J. Pl. Sci." pageId="84" pageNumber="74" pagination="1016 - 1035" part="180" refId="ref112729" refString="PATEL, N., S. FAWCETT, M. SUNDUE, & J. BUDKE. 2019 a. Evolution of perine morphology in Thelypteridaceae. Int. J. Pl. Sci. 180: 1016 - 1035." title="Evolution of perine morphology in Thelypteridaceae" type="journal article" year="2019" yearSuffix="a">Patel et al. 2019a</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1E4D63FFBD9B1F62F778A8FC63FA4D" box="[732,964,1505,1526]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="elatus">
|
||
<emphasis id="B96AEAF2FFBD9B1F62F778A8FC63FA4D" box="[732,964,1505,1526]" italics="true" pageId="84" pageNumber="74">Sphaerostephanos elatus</emphasis>
|
||
</taxonomicName>
|
||
differs from all species in
|
||
<taxonomicName id="4C1E4D63FFBD9B1F604A7B48FEA6F9AD" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[97,257,1537,1558]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F604A7B48FEA6F9AD" box="[97,257,1537,1558]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
in having several pairs of veins anastomosing below the sinus, but it is alike in having the characteristic proximal pinnae abruptly reduced to triangular auricles, and large, persistent indusia. In our phylogenetic analysis (Fawcett et al. in press) the Afro-Madagascan
|
||
<taxonomicName id="4C1E4D63FFBD9B1F623E7B0BFDC8F9ED" box="[533,623,1601,1623]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="species" species="pulcher">
|
||
<emphasis id="B96AEAF2FFBD9B1F623E7B0BFDC8F9ED" box="[533,623,1601,1623]" italics="true" pageId="84" pageNumber="74">P.pulcher</emphasis>
|
||
</taxonomicName>
|
||
is sister to all other
|
||
<taxonomicName id="4C1E4D63FFBD9B1F63057B08FC6AF9ED" baseAuthorityName="Holttum & Roy" baseAuthorityYear="1965" box="[814,973,1601,1622]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="84" pageNumber="74" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBD9B1F63057B08FC6AF9ED" box="[814,973,1601,1622]" italics="true" pageId="84" pageNumber="74">Pseudocyclosorus</emphasis>
|
||
</taxonomicName>
|
||
, with the core clade of the genus all from the Pan-Himalayan region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |