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data/03/B7/87/03B787F6FF889B2962077DB6FCC7FDCC.xml
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@ -1,29 +1,31 @@
<document id="E41807E026D67C9775D2880CE3CBFEB9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634280027544" checkinUser="donat" docAuthor="Fawcett, Susan &amp; Smith, Alan R." docDate="2021" docId="03B787F6FF889B2962077DB6FCC7FDCC" docLanguage="en" docName="2021_ThelypteridaceaeFerns_OACopy3.pdf.imf" docOrigin="Fort Worth, Texas, USA: BRIT Press" docTitle="Strophocaulon" docType="treatment" docVersion="1" lastPageNumber="88" masterDocId="FF8EFF8EFFE99B4B602B7D49FFA7FFBB" masterDocTitle="A Generic Classification of the Thelypteridaceae" pageNumber="87" updateTime="1731345217528" updateUser="felipe" zenodo-license-document="CC-BY-NC-SA-4.0">
<mods:mods id="B3C2CE66DC6FFE8C8B9EDB1114B8C63E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6C1174290D628B0EE62CFB37BE22C565">
<mods:title id="EC8736A1DDBB4EE4CA141769C399A6D7">A Generic Classification of the Thelypteridaceae</mods:title>
<document id="BBD769D3A130160713F441E1BBE34267" ID-DOI="10.17348/jbrit.v15.i2.1206" ID-Zenodo-Dep="14111022" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634280027544" checkinUser="donat" docAuthor="Fawcett, Susan &amp; Smith, Alan R." docDate="2021" docId="03B787F6FF889B2962077DB6FCC7FDCC" docLanguage="en" docName="2021_ThelypteridaceaeFerns_OACopy3.pdf.imf" docOrigin="Fort Worth, Texas, USA: BRIT Press" docTitle="Strophocaulon" docType="treatment" docVersion="3" lastPageNumber="88" masterDocId="FF8EFF8EFFE99B4B602B7D49FFA7FFBB" masterDocTitle="A Generic Classification of the Thelypteridaceae" pageNumber="87" updateTime="1731444927364" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="E6B8601CB4CBCBA50008BAFF84EF441C">
<mods:title id="EF239FC4D58B9752BF83BCA7D2A44959">A Generic Classification of the Thelypteridaceae</mods:title>
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<mods:namePart id="0C64A022333A14233070E98924B0DE39">Fawcett, Susan</mods:namePart>
<mods:namePart id="4371A734BA40ACFAD96DE2BA5B1B755B">Fawcett, Susan</mods:namePart>
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<mods:namePart id="C22888BDFC83F72AC5FB659364207FF4">Smith, Alan R.</mods:namePart>
<mods:namePart id="32EC78D7E8AB4EE0CF84D7B2F7BE598F">Smith, Alan R.</mods:namePart>
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<mods:dateIssued id="81F295237765E330D7CBB800BDC49475">2021</mods:dateIssued>
<mods:publisher id="FF3DBFED534059106B9B0FADE49C1C6A">BRIT Press</mods:publisher>
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<treatment id="03B787F6FF889B2962077DB6FCC7FDCC" LSID="urn:lsid:plazi:treatment:03B787F6FF889B2962077DB6FCC7FDCC" httpUri="http://treatment.plazi.org/id/03B787F6FF889B2962077DB6FCC7FDCC" lastPageId="98" lastPageNumber="88" pageId="97" pageNumber="87">
<subSubSection id="C304656BFF889B2A62077DB6FC8AFEA2" box="[556,813,255,281]" pageId="97" pageNumber="87" type="nomenclature">
@ -91,7 +93,7 @@ Gr.
, twisted cord +
<emphasis id="B96AEAF2FF889B2A62277C3EFDEEFE37" box="[524,585,375,396]" italics="true" pageId="97" pageNumber="87">caulon</emphasis>
, stem, in reference to the tortuous, long-creeping subterranean rhizomes (
<figureCitation id="13252A65FF889B2A60E87CD1FEB5FE16" box="[195,274,408,429]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." pageId="97" pageNumber="87">
<figureCitation id="13252A65FF889B2A60E87CD1FEB5FE16" box="[195,274,408,429]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." figureDoi="http://doi.org/10.5281/zenodo.14111108" httpUri="https://zenodo.org/record/14111108/files/figure.png" pageId="97" pageNumber="87">
<emphasis id="B96AEAF2FF889B2A60E87CD1FEB5FE16" bold="true" box="[195,274,408,429]" pageId="97" pageNumber="87">Fig. 10B</emphasis>
</figureCitation>
), which distinguish it from
@ -106,7 +108,7 @@ Gr.
Plants terrestrial, forming colonies in open sites, medium to large, to&gt; 1.2 m tall;
<emphasis id="B96AEAF2FF889B2A639B7CF1FBA9FE76" bold="true" box="[944,1038,440,461]" pageId="97" pageNumber="87">rhizomes</emphasis>
thick (5+ mm), black, long-creeping, and subterranean, with internodes to 7+ cm long (
<figureCitation id="13252A65FF889B2A62C27C91FC9EFE56" box="[745,825,472,493]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." pageId="97" pageNumber="87">
<figureCitation id="13252A65FF889B2A62C27C91FC9EFE56" box="[745,825,472,493]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." figureDoi="http://doi.org/10.5281/zenodo.14111108" httpUri="https://zenodo.org/record/14111108/files/figure.png" pageId="97" pageNumber="87">
<emphasis id="B96AEAF2FF889B2A62C27C91FC9EFE56" bold="true" box="[745,825,472,493]" pageId="97" pageNumber="87">Fig. 10B</emphasis>
</figureCitation>
), bearing stramineous to brown, lanceolate, sometimes tortuous scales with setose margins;
@ -126,7 +128,7 @@ chartaceous to coriaceous, often bicolorous (drying paler below), often drying r
<emphasis id="B96AEAF2FF889B2A61A57F1EFE43FDD7" box="[398,484,599,620]" italics="true" pageId="97" pageNumber="87">S. unitum</emphasis>
</taxonomicName>
,
<figureCitation id="13252A65FF889B2A61C57F11FD98FDD6" box="[494,575,600,621]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." pageId="97" pageNumber="87">
<figureCitation id="13252A65FF889B2A61C57F11FD98FDD6" box="[494,575,600,621]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." figureDoi="http://doi.org/10.5281/zenodo.14111108" httpUri="https://zenodo.org/record/14111108/files/figure.png" pageId="97" pageNumber="87">
<emphasis id="B96AEAF2FF889B2A61C57F11FD98FDD6" bold="true" box="[494,575,600,621]" pageId="97" pageNumber="87">Fig. 10C</emphasis>
</figureCitation>
), frond base with pinnae abruptly reduced (
@ -144,7 +146,7 @@ chartaceous to coriaceous, often bicolorous (drying paler below), often drying r
sessile to short-petiolulate, pinna bases truncate or broadly cuneate, sometimes slightly dilated; margins sharply dentate to incised to 1/3 towards costae;
<emphasis id="B96AEAF2FF889B2A64877FF1FB46FD76" bold="true" box="[1196,1249,696,717]" pageId="97" pageNumber="87">veins</emphasis>
with at least one pair anastomosing at an angle &lt;90 degrees, with an excurrent veinlet running to the sinus, with several pairs of veins forming areoles, running along a deep, narrow sinus membrane, lowest veins sometimes arising from costae, not from costules (
<figureCitation id="13252A65FF889B2A61527E51FE6DFC96" box="[377,458,792,813]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." pageId="97" pageNumber="87">
<figureCitation id="13252A65FF889B2A61527E51FE6DFC96" box="[377,458,792,813]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." figureDoi="http://doi.org/10.5281/zenodo.14111108" httpUri="https://zenodo.org/record/14111108/files/figure.png" pageId="97" pageNumber="87">
<emphasis id="B96AEAF2FF889B2A61527E51FE6DFC96" bold="true" box="[377,458,792,813]" pageId="97" pageNumber="87">Fig. 10A</emphasis>
</figureCitation>
);
@ -154,7 +156,7 @@ inconspicuous;
of hyaline, spreading, acicular hairs, these restricted to axes, or also present on laminar tissue;
<emphasis id="B96AEAF2FF889B2A62E47E71FC33FCF6" bold="true" box="[719,916,824,845]" pageId="97" pageNumber="87">indument abaxially</emphasis>
of hyaline acicular hairs present on all axes (
<figureCitation id="13252A65FF889B2A60E67E11FEBAFCD6" box="[205,285,856,877]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." pageId="97" pageNumber="87">
<figureCitation id="13252A65FF889B2A60E67E11FEBAFCD6" box="[205,285,856,877]" captionStart="FIG" captionStartId="29.[121,129,1704,1724]" captionTargetBox="[118,1248,161,1657]" captionTargetId="figure-8@29.[115,1248,154,1660]" captionTargetPageId="29" captionText="FIG. 10.Strophocaulon unitum, Fawcett 641 (VT),A.pinna lobes,abaxial view.B. rhizome.C.laminar apex." figureDoi="http://doi.org/10.5281/zenodo.14111108" httpUri="https://zenodo.org/record/14111108/files/figure.png" pageId="97" pageNumber="87">
<emphasis id="B96AEAF2FF889B2A60E67E11FEBAFCD6" bold="true" box="[205,285,856,877]" pageId="97" pageNumber="87">Fig. 10A</emphasis>
</figureCitation>
), frequently abundant on laminar tissue, often forming a tangled mat of whitish hairs along rachises, spherical yellow or brown glands present on laminar surface, restricted to veins, or glands absent;

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data/03/B7/87/03B787F6FF899B2A604A793BFD70FF6C.xml
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@ -1,29 +1,31 @@
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@ -128,7 +130,7 @@ var.
(Sw.) Pic.Serm.
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@ -393,7 +395,7 @@ fidelei
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nearly symmetrical with segments spreading or only slightly oblique; margins entire, pinna bases sessile to short-petiolulate, truncate, acroscopic segment often slightly elongate but not auriculate;
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free, simple or often forked, ending at margins of lobes just above sinuses (
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<figureCitation id="13252A65FF8B9B2964587804FB1DFAD9" box="[1139,1210,1357,1378]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="98" pageNumber="88">
<emphasis id="B96AEAF2FF8B9B2964587804FB1DFAD9" bold="true" box="[1139,1210,1357,1378]" pageId="98" pageNumber="88">Fig. 6D</emphasis>
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);
@ -175,11 +177,11 @@ Diagnosis.—
can be readily distinguished from the superficially similar, and sometimes co-occurring amauropeltoid genera by a combination of characteristics. It has consistently long-creeping, relatively narrow rhizomes (1.5–
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diam.); grows in persistently wet habitats in temperate areas; has largely scaleless rhizomes behind the apex; has persistent ovate scales on the costae abaxially (
<figureCitation id="13252A65FF8A9B28625B7DEBFD1EFF0C" box="[624,697,162,183]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." pageId="99" pageNumber="89">
<figureCitation id="13252A65FF8A9B28625B7DEBFD1EFF0C" box="[624,697,162,183]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="99" pageNumber="89">
<emphasis id="B96AEAF2FF8A9B28625B7DEBFD1EFF0C" bold="true" box="[624,697,162,183]" pageId="99" pageNumber="89">Fig. 6D</emphasis>
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); lacks greatly reduced proximal pinnae; has frequently forked, free veins on the ultimate segments (
<figureCitation id="13252A65FF8A9B2862377D8BFDC5FF6C" box="[540,610,194,215]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." pageId="99" pageNumber="89">
<figureCitation id="13252A65FF8A9B2862377D8BFDC5FF6C" box="[540,610,194,215]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="99" pageNumber="89">
<emphasis id="B96AEAF2FF8A9B2862377D8BFDC5FF6C" bold="true" box="[540,610,194,215]" pageId="99" pageNumber="89">Fig. 6D</emphasis>
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); and is often slightly dimorphic, with fertile fronds taller, and with confluent sori resulting in an almost acrostichoid appearance.
@ -465,7 +467,7 @@ All phylogenetic studies with broad sampling (
,
</emphasis>
as treated here, sister to the rest of Thelypteridoideae (
<figureCitation id="13252A65FF8A9B28632178ABFC99FA4C" box="[778,830,1506,1527]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="99" pageNumber="89">
<figureCitation id="13252A65FF8A9B28632178ABFC99FA4C" box="[778,830,1506,1527]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="99" pageNumber="89">
<emphasis id="B96AEAF2FF8A9B28632178ABFC99FA4C" bold="true" box="[778,830,1506,1527]" pageId="99" pageNumber="89">Fig. 1</emphasis>
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). This subfamily includes 900+ species, and is sister to the smaller subfam.
@ -510,7 +512,7 @@ represents an ancient lineage, diverged from its closest extant relatives by per
(Salisb.) Schott
</taxonomicName>
(
<figureCitation id="13252A65FF8A9B2864067B0BFBD4F9EC" box="[1069,1139,1602,1623]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." pageId="99" pageNumber="89">
<figureCitation id="13252A65FF8A9B2864067B0BFBD4F9EC" box="[1069,1139,1602,1623]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="99" pageNumber="89">
<emphasis id="B96AEAF2FF8A9B2864067B0BFBD4F9EC" bold="true" box="[1069,1139,1602,1623]" pageId="99" pageNumber="89">Fig. 6D</emphasis>
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42
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@ -134,11 +136,11 @@ chartaceous to subcoriaceous, drying greenish, pinnae generally pinnatifid or pi
opposite or nearly so proximally, becoming alternate distally, adaxially with a groove that is not continuous with the rachis groove, shallowly to often deeply pinnatifid;
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usually prominent on both sides, one or sometimes two pairs from adjacent segments connivent at an acute angle at or very slightly below the sinus (
<figureCitation id="13252A65FFA19B03630878F1FCCCFA77" box="[803,875,1463,1485]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="72" pageNumber="62">
<figureCitation id="13252A65FFA19B03630878F1FCCCFA77" box="[803,875,1463,1485]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="72" pageNumber="62">
<emphasis id="B96AEAF2FFA19B03630878F1FCCCFA77" bold="true" box="[803,875,1463,1485]" pageId="72" pageNumber="62">Fig. 7D</emphasis>
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), or the distal one or a pair meeting segment margin just above the sinus (veins not forming areoles) (
<figureCitation id="13252A65FFA19B0362E37891FCAAFA56" box="[712,781,1496,1517]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="72" pageNumber="62">
<figureCitation id="13252A65FFA19B0362E37891FCAAFA56" box="[712,781,1496,1517]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="72" pageNumber="62">
<emphasis id="B96AEAF2FFA19B0362E37891FCAAFA56" bold="true" box="[712,781,1496,1517]" pageId="72" pageNumber="62">Fig. 7C</emphasis>
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), rarely with a single pair truly obtusely united below the sinus and with an excurrent vein running to sinus, vein endings reaching segment margins;

44
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@ -64,7 +66,7 @@ G. Forst., Fl. Ins. Austr.Prodr.
(G. Forst.) H.H. Allan
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(
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<figureCitation id="13252A65FFA19B0364887E19FF74FCC1" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="72" pageNumber="62">
<emphasis id="B96AEAF2FFA19B0364887E19FF74FCC1" bold="true" pageId="72" pageNumber="62">Figs. 7E, 7F</emphasis>
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48
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@ -97,7 +99,7 @@ Gr.
<emphasis id="B96AEAF2FFA29B0063177BD8FC1EF91D" box="[828,953,1681,1702]" italics="true" pageId="75" pageNumber="65">Mesophlebion</emphasis>
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, from which it is distinguished by the basal basiscopic vein never arising far from its costule (
<figureCitation id="13252A65FFA29B0062E97BFBFCA0F97C" box="[706,775,1714,1735]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." pageId="75" pageNumber="65">
<figureCitation id="13252A65FFA29B0062E97BFBFCA0F97C" box="[706,775,1714,1735]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="75" pageNumber="65">
<emphasis id="B96AEAF2FFA29B0062E97BFBFCA0F97C" bold="true" box="[706,775,1714,1735]" pageId="75" pageNumber="65">Fig. 6A</emphasis>
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).
@ -174,7 +176,7 @@ unadorned, or with spherical amber or yellow glands (e.g.,
<taxonomicName id="4C1E4D63FFA59B0762627FEBFD6DFD0D" box="[585,714,673,695]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="76" pageNumber="66" phylum="Tracheophyta" rank="species" species="imbricatum">P.imbricatum</taxonomicName>
,
</emphasis>
<figureCitation id="13252A65FFA59B0762FA7FEBFCA8FD0C" box="[721,783,674,695]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="76" pageNumber="66">
<figureCitation id="13252A65FFA59B0762FA7FEBFCA8FD0C" box="[721,783,674,695]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="76" pageNumber="66">
<emphasis id="B96AEAF2FFA59B0762FA7FEBFCA8FD0C" bold="true" box="[721,783,674,695]" pageId="76" pageNumber="66">Fig. 2I</emphasis>
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), or setulae;
@ -222,9 +224,9 @@ and the western Pacific, differs from the strictly Malesian
<emphasis id="B96AEAF2FFA59B0761217E68FE2FFC8D" box="[266,392,801,822]" italics="true" pageId="76" pageNumber="66">Mesophlebion</emphasis>
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by the proximity of the origin of the basal basiscopic vein to the costule (
<figureCitation id="13252A65FFA59B0764747E6BFB3FFC8C" box="[1119,1176,802,823]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." pageId="76" pageNumber="66">Fig. 6</figureCitation>
<figureCitation id="13252A65FFA59B0764747E6BFB3FFC8C" box="[1119,1176,802,823]" captionStart="FIG" captionStartId="26.[97,105,1778,1798]" captionTargetBox="[102,1218,158,1728]" captionTargetId="figure-8@26.[102,1218,156,1733]" captionTargetPageId="26" captionText="FIG. 6. A. Plesioneuron hopeanum, Fawcett 613 (VT),pinna lobes,adaxial view.Amauropelta (subg.Nibaa) noveboracensis, Fawcett 567 (MICH,VT),B. pinna-lobes, adaxial view.C.habit.D. Thelypterispalustris,Fawcett 569 (VT),pinna lobes abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111064" httpUri="https://zenodo.org/record/14111064/files/figure.png" pageId="76" pageNumber="66">Fig. 6</figureCitation>
) (vs. arising far from it;
<figureCitation id="13252A65FFA59B0761337E0BFEF8FCEC" box="[280,351,834,855]" captionStart="FIG" captionStartId="28.[97,105,1748,1768]" captionTargetBox="[93,1223,156,1702]" captionTargetId="figure-8@28.[92,1230,156,1705]" captionTargetPageId="28" captionText="FIG. 9.Steiropteris deltoidea,Fawcett 464 (VT) A. habit.B. pinna lobes,adaxial view.C.Mesophlebion sp., Karger 1796 (VT),pinna lobes,abaxial view." pageId="76" pageNumber="66">Fig. 9C</figureCitation>
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@ -1,29 +1,31 @@
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@ -90,7 +92,7 @@ Gr.
, air, breath +
<emphasis id="B96AEAF2FFA79B0561E77C52FDA7FE8B" box="[460,512,283,304]" italics="true" pageId="78" pageNumber="68">pteris</emphasis>
, fern. The peg-like pneumatophores (aerophores) extending from the stipe, and the bases of the pinnae (and sometimes the bases of the costules) are diagnostic for this genus (
<figureCitation id="13252A65FFA79B0564207C72FBF6FEEB" box="[1035,1105,315,336]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="78" pageNumber="68">
<figureCitation id="13252A65FFA79B0564207C72FBF6FEEB" box="[1035,1105,315,336]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="78" pageNumber="68">
<emphasis id="B96AEAF2FFA79B0564207C72FBF6FEEB" bold="true" box="[1035,1105,315,336]" pageId="78" pageNumber="68">Fig. 2C</emphasis>
</figureCitation>
).
@ -119,7 +121,7 @@ sessile, with truncate bases, shallowly lobed (generally &lt;
⁄
<subScript id="179A34A5FFA79B0564707F4BFBC5FDB5" attach="left" box="[1115,1122,514,526]" fontSize="5" pageId="78" pageNumber="68">2</subScript>
to costae), with pronounced cartilaginous sinus membranes that are grooved adaxially (
<figureCitation id="13252A65FFA79B0563177F52FCD8FD8B" box="[828,895,539,560]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="78" pageNumber="68">
<figureCitation id="13252A65FFA79B0563177F52FCD8FD8B" box="[828,895,539,560]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="78" pageNumber="68">
<emphasis id="B96AEAF2FFA79B0563177F52FCD8FD8B" bold="true" box="[828,895,539,560]" pageId="78" pageNumber="68">Fig.7A</emphasis>
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) and project out of the plane of the blade abaxially, segment tips generally acute;
@ -127,7 +129,7 @@ to costae), with pronounced cartilaginous sinus membranes that are grooved adaxi
prominent above and below, often paler than lamina, with at least one pair anastomosing below each sinus, forming an excurrent vein to sinus, with more distal veins running to both sides of an elongate sinus membrane, veins closely spaced, (7–)11–12(–15) pairs per segment;
<emphasis id="B96AEAF2FFA79B0563CD7F33FBF0FD34" bold="true" box="[998,1111,634,655]" pageId="78" pageNumber="68">aerophores</emphasis>
prominent, peg-like (
<figureCitation id="13252A65FFA79B0560977FD2FEA4FD0B" box="[188,259,667,688]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="78" pageNumber="68">
<figureCitation id="13252A65FFA79B0560977FD2FEA4FD0B" box="[188,259,667,688]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="78" pageNumber="68">
<emphasis id="B96AEAF2FFA79B0560977FD2FEA4FD0B" bold="true" box="[188,259,667,688]" pageId="78" pageNumber="68">Fig. 2C</emphasis>
</figureCitation>
), projecting through mucilage during early frond development, along stipes, and at pinna bases, frequently also with peg-like or swollen aerophores at the bases of costae;
@ -202,7 +204,7 @@ Diagnosis.—
.
</taxonomicNameLabel>
is distinguished from most other christelloids by having prominent aerophores along the stipe, at the bases of pinnae (
<figureCitation id="13252A65FFA79B0561E57972FDB3FBEB" box="[462,532,1083,1104]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="78" pageNumber="68">
<figureCitation id="13252A65FFA79B0561E57972FDB3FBEB" box="[462,532,1083,1104]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="78" pageNumber="68">
<emphasis id="B96AEAF2FFA79B0561E57972FDB3FBEB" bold="true" box="[462,532,1083,1104]" pageId="78" pageNumber="68">Fig. 2C</emphasis>
</figureCitation>
), and at the bases of costae; pinnae abruptly reduced to vestiges subtending aerophores (not more than a few mm in length); fronds producing mucilage during early development; and veins closely spaced (ca. 12 per segment). The pinna segments of
@ -449,7 +451,7 @@ by
<emphasis id="B96AEAF2FFA69B0461607F28FE1CFDCD" box="[331,443,609,630]" italics="true" pageId="79" pageNumber="69">Reholttumia</emphasis>
</taxonomicName>
;
<figureCitation id="13252A65FFA69B0461EE7F2BFE5DFDCC" box="[453,506,610,631]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="79" pageNumber="69">
<figureCitation id="13252A65FFA69B0461EE7F2BFE5DFDCC" box="[453,506,610,631]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="79" pageNumber="69">
<emphasis id="B96AEAF2FFA69B0461EE7F2BFE5DFDCC" bold="true" box="[453,506,610,631]" pageId="79" pageNumber="69">Fig. 1</emphasis>
</figureCitation>
). To preserve a monophyletic
@ -513,7 +515,7 @@ Holttum
(Brack.) Holttum
</taxonomicName>
(
<figureCitation id="13252A65FFA69B0460F27E6BFE86FC8C" box="[217,289,802,823]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="79" pageNumber="69">
<figureCitation id="13252A65FFA69B0460F27E6BFE86FC8C" box="[217,289,802,823]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="79" pageNumber="69">
<emphasis id="B96AEAF2FFA69B0460F27E6BFE86FC8C" bold="true" box="[217,289,802,823]" pageId="79" pageNumber="69">Fig. 2C</emphasis>
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);
@ -532,7 +534,7 @@ Holttum
(F. Ballard) Holttum
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(
<figureCitation id="13252A65FFA69B0464977E6BFF35FCEC" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="79" pageNumber="69">
<figureCitation id="13252A65FFA69B0464977E6BFF35FCEC" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="79" pageNumber="69">
<emphasis id="B96AEAF2FFA69B0464977E6BFF35FCEC" bold="true" pageId="79" pageNumber="69">Fig. 7A</emphasis>
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44
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@ -191,7 +193,7 @@ is quite distinctive and bears the narrow elongate pinnae with oblique costules
<emphasis id="B96AEAF2FFA89B0A627879BEFCBEFAB0" box="[595,793,1270,1292]" italics="true" pageId="65" pageNumber="55">Menisciopsis boydiae</emphasis>
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<emphasis id="B96AEAF2FFA89B0A630D79B1FCCAFAB7" bold="true" box="[806,877,1271,1293]" pageId="65" pageNumber="55">Fig. 5B</emphasis>
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@ -264,7 +266,7 @@ and
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, a New World genus of 16 species (elevated to generic status herein) (
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@ -88,7 +90,7 @@ Gr.
, middle +
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, vein, in reference to basal basiscopic veins arising from costae between costules (
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<figureCitation id="13252A65FFAB9B0960907E74FEA6FCE9" box="[187,257,829,850]" captionStart="FIG" captionStartId="28.[97,105,1748,1768]" captionTargetBox="[93,1223,156,1702]" captionTargetId="figure-8@28.[92,1230,156,1705]" captionTargetPageId="28" captionText="FIG. 9.Steiropteris deltoidea,Fawcett 464 (VT) A. habit.B. pinna lobes,adaxial view.C.Mesophlebion sp., Karger 1796 (VT),pinna lobes,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111101" httpUri="https://zenodo.org/record/14111101/files/figure.png" pageId="66" pageNumber="56">
<emphasis id="B96AEAF2FFAB9B0960907E74FEA6FCE9" bold="true" box="[187,257,829,850]" pageId="66" pageNumber="56">Fig. 9C</emphasis>
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).
@ -133,7 +135,7 @@ free, rarely weakly united (
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), usually connivent at or just below the sinuses and touching sides of an often hairy ridge below sinus, simple in ultimate segments, ending at segment margins, basal basiscopic veins arising from costa midway between costules (
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<figureCitation id="13252A65FFAB9B09613E79F4FEFBFB69" box="[277,348,1213,1234]" captionStart="FIG" captionStartId="28.[97,105,1748,1768]" captionTargetBox="[93,1223,156,1702]" captionTargetId="figure-8@28.[92,1230,156,1705]" captionTargetPageId="28" captionText="FIG. 9.Steiropteris deltoidea,Fawcett 464 (VT) A. habit.B. pinna lobes,adaxial view.C.Mesophlebion sp., Karger 1796 (VT),pinna lobes,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111101" httpUri="https://zenodo.org/record/14111101/files/figure.png" pageId="66" pageNumber="56">
<emphasis id="B96AEAF2FFAB9B09613E79F4FEFBFB69" bold="true" box="[277,348,1213,1234]" pageId="66" pageNumber="56">Fig. 9C</emphasis>
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), near base of costule, or from costules (in smaller spp., e.g.,
@ -231,7 +233,7 @@ and
<emphasis id="B96AEAF2FFAA9B0863777C28FBD6FECD" box="[860,1137,353,374]" italics="true" pageId="67" pageNumber="57">Sphaerostephanos beccarianus</emphasis>
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group (
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<figureCitation id="13252A65FFAA9B0864977C2BFF35FE2C" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="67" pageNumber="57">
<emphasis id="B96AEAF2FFAA9B0864977C2BFF35FE2C" bold="true" pageId="67" pageNumber="57">Fig. 7A</emphasis>
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). Holttum treated it in
@ -297,7 +299,7 @@ subg.
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), which they superficially resemble in venation, blade architecture, and blade texture (
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<figureCitation id="13252A65FFAA9B0861D87FCBFD8CFD2C" box="[499,555,642,663]" captionStart="FIG" captionStartId="28.[97,105,1748,1768]" captionTargetBox="[93,1223,156,1702]" captionTargetId="figure-8@28.[92,1230,156,1705]" captionTargetPageId="28" captionText="FIG. 9.Steiropteris deltoidea,Fawcett 464 (VT) A. habit.B. pinna lobes,adaxial view.C.Mesophlebion sp., Karger 1796 (VT),pinna lobes,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111101" httpUri="https://zenodo.org/record/14111101/files/figure.png" pageId="67" pageNumber="57">
<emphasis id="B96AEAF2FFAA9B0861D87FCBFD8CFD2C" bold="true" box="[499,555,642,663]" pageId="67" pageNumber="57">Fig. 9</emphasis>
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).

40
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@ -98,7 +100,7 @@ light brown, with brown ovate to lanceolate glabrous scales on stipe bases;
membranaceous to chartaceous, drying dull green, ovate to lanceolate; two to five pairs of proximal pinnae gradually reduced, the lowest 1–4(–6) cm long (never rudimentary or glanduliform);
<emphasis id="B96AEAF2FFAE9B0C6091793BFEA7FB3C" bold="true" box="[186,256,1138,1159]" pageId="71" pageNumber="61">pinnae</emphasis>
opposite or subopposite (
<figureCitation id="13252A65FFAE9B0C61D1793BFD99FB3D" box="[506,574,1137,1159]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="71" pageNumber="61">
<figureCitation id="13252A65FFAE9B0C61D1793BFD99FB3D" box="[506,574,1137,1159]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="71" pageNumber="61">
<emphasis id="B96AEAF2FFAE9B0C61D1793BFD99FB3D" bold="true" box="[506,574,1137,1159]" pageId="71" pageNumber="61">Fig. 7F</emphasis>
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), even towards blade apex, grooved adaxially, (10–)15–30 pairs, lobes oblique, rounded with crenulate margins, incised halfway or a little more to costae, proximal pinnae frequently auricled, with acroscopic auricles often secondarily lobed; costules
@ -112,7 +114,7 @@ absent;
of brown, often lustrous, ovate to deltate scales (to
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wide and about as long) along rachis and costae (
<figureCitation id="13252A65FFAE9B0C619D787BFE5DFAFD" box="[438,506,1329,1351]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="71" pageNumber="61">
<figureCitation id="13252A65FFAE9B0C619D787BFE5DFAFD" box="[438,506,1329,1351]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="71" pageNumber="61">
<emphasis id="B96AEAF2FFAE9B0C619D787BFE5DFAFD" bold="true" box="[438,506,1329,1351]" pageId="71" pageNumber="61">Fig. 7E</emphasis>
</figureCitation>
), hairs sparse and short, unicellular, acicular or capitate on costae, or lacking;
@ -122,7 +124,7 @@ of arching reddish brown hairs on rachis and costae, the hairs directed toward p
absent;
<emphasis id="B96AEAF2FFAE9B0C60CF78DBFEABFA1C" bold="true" box="[228,268,1426,1447]" pageId="71" pageNumber="61">sori</emphasis>
inframedial, exindusiate, the basal pair or two on each segment (larger fronds) often oval, to twice as long as wide (
<figureCitation id="13252A65FFAE9B0C60D078FBFE98FA7D" box="[251,319,1457,1479]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="71" pageNumber="61">
<figureCitation id="13252A65FFAE9B0C60D078FBFE98FA7D" box="[251,319,1457,1479]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="71" pageNumber="61">
<emphasis id="B96AEAF2FFAE9B0C60D078FBFE98FA7D" bold="true" box="[251,319,1457,1479]" pageId="71" pageNumber="61">Fig. 7E</emphasis>
</figureCitation>
);
@ -204,7 +206,7 @@ Fée
), from Africa (
<bibRefCitation id="EF8F4B11FFA19B0360F27DEBFECBFF0C" author="Holttum" box="[217,364,162,183]" firstAuthor="Holttum" journalOrPublisher="J. S. African Bot." pageId="72" pageNumber="62" pagination="123 - 168" part="40" refId="ref109577" refString="HOLTTUM, R. E. 1974 a. Thelypteridaceae of Africa and adjacent islands. J. S. African Bot. 40: 123 - 168." title="Thelypteridaceae of Africa and adjacent islands" type="journal article" year="1974" yearSuffix="a">Holttum 1974a</bibRefCitation>
). Molecular evidence (Fawcett et al. in press) suggests the two species diverged successively from the backbone of the phylogeny and are on long branches (
<figureCitation id="13252A65FFA19B03629D7D8BFD4DFF6C" box="[694,746,194,215]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="72" pageNumber="62">
<figureCitation id="13252A65FFA19B03629D7D8BFD4DFF6C" box="[694,746,194,215]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="72" pageNumber="62">
<emphasis id="B96AEAF2FFA19B03629D7D8BFD4DFF6C" bold="true" box="[694,746,194,215]" pageId="72" pageNumber="62">Fig. 1</emphasis>
</figureCitation>
).
@ -302,7 +304,7 @@ is quite distinct from any extant genus of
) all sensu
<bibRefCitation id="EF8F4B11FFA19B0361477FCBFDA6FD2C" author="Holttum" box="[364,513,642,663]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="72" pageNumber="62" pagination="17 - 52" part="19" refId="ref109463" refString="HOLTTUM, R. E. 1971. Studies in the family Thelypteridaceae III. A new system of genera in the Old World. Blumea 19: 17 - 52." title="Studies in the family Thelypteridaceae III. A new system of genera in the Old World" type="journal article" year="1971">Holttum (1971)</bibRefCitation>
, as well as a half dozen smaller genera (
<figureCitation id="13252A65FFA19B0363547FCBFC12FD2C" box="[895,949,642,663]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="72" pageNumber="62">
<figureCitation id="13252A65FFA19B0363547FCBFC12FD2C" box="[895,949,642,663]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="72" pageNumber="62">
<emphasis id="B96AEAF2FFA19B0363547FCBFC12FD2C" bold="true" box="[895,949,642,663]" pageId="72" pageNumber="62">Fig. 1</emphasis>
</figureCitation>
). The christelloid genera are almost exclusively Old World, primarily restricted to southeastern and southern Asia,

58
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@ -107,7 +109,7 @@ For additional generic synonymy, see
, sphere +
<emphasis id="B96AEAF2FFB09B1261E97CCCFDBBFE21" box="[450,540,389,410]" italics="true" pageId="89" pageNumber="79">stephanos</emphasis>
, crown, in reference to the spherical yellow glands adorning the sori; these may be borne on the receptacles, indusia (
<figureCitation id="13252A65FFB09B1262237CEFFDE9FE00" box="[520,590,422,443]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="89" pageNumber="79">
<figureCitation id="13252A65FFB09B1262237CEFFDE9FE00" box="[520,590,422,443]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="89" pageNumber="79">
<emphasis id="B96AEAF2FFB09B1262237CEFFDE9FE00" bold="true" box="[520,590,422,443]" pageId="89" pageNumber="79">Fig. 2D</emphasis>
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), or sporangia, and are characteristic of the genus.
@ -133,7 +135,7 @@ monomorphic or with fertile fronds smaller and more erect (fronds subdimorphic t
<emphasis id="B96AEAF2FFB09B1262AC7F0CFCAEFDE1" box="[647,777,581,602]" italics="true" pageId="89" pageNumber="79">S. beccarianus</emphasis>
</taxonomicName>
(
<figureCitation id="13252A65FFB09B12633E7F0FFCFEFDE1" box="[789,857,581,603]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="89" pageNumber="79">
<figureCitation id="13252A65FFB09B12633E7F0FFCFEFDE1" box="[789,857,581,603]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="89" pageNumber="79">
<emphasis id="B96AEAF2FFB09B12633E7F0FFCFEFDE1" bold="true" box="[789,857,581,603]" pageId="89" pageNumber="79">Fig. 7B</emphasis>
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) and related species (no simple-bladed or twice-pinnate species known), usually arching;
@ -145,7 +147,7 @@ usually chartaceous, less often thicker and subcoriaceous to coriaceous (species
<emphasis id="B96AEAF2FFB09B1260C87F8CFEC3FD61" box="[227,356,709,730]" italics="true" pageId="89" pageNumber="79">S. lithophyllus</emphasis>
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), usually drying green, lanceolate, ovate to deltate, proximal pinnae usually subabruptly or abruptly reduced (
<figureCitation id="13252A65FFB09B1261077FAFFED4FD40" box="[300,371,742,763]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="89" pageNumber="79">
<figureCitation id="13252A65FFB09B1261077FAFFED4FD40" box="[300,371,742,763]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="89" pageNumber="79">
<emphasis id="B96AEAF2FFB09B1261077FAFFED4FD40" bold="true" box="[300,371,742,763]" pageId="89" pageNumber="79">Fig. 2K</emphasis>
</figureCitation>
), occasionally gradually reduced, often&gt; 6 pairs (to 30+ pairs, e.g., in
@ -207,7 +209,7 @@ generally absent on laminar tissue (present in some spp., e.g.,
<emphasis id="B96AEAF2FFB09B1262C5780CFCD5FAE1" box="[750,882,1349,1370]" italics="true" pageId="89" pageNumber="79">S. beccarianus</emphasis>
</taxonomicName>
,
<figureCitation id="13252A65FFB09B126356780FFC65FAE0" box="[893,962,1350,1371]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="89" pageNumber="79">
<figureCitation id="13252A65FFB09B126356780FFC65FAE0" box="[893,962,1350,1371]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="89" pageNumber="79">
<emphasis id="B96AEAF2FFB09B126356780FFC65FAE0" bold="true" box="[893,962,1350,1371]" pageId="89" pageNumber="79">Fig. 7A</emphasis>
</figureCitation>
);
@ -215,7 +217,7 @@ generally absent on laminar tissue (present in some spp., e.g.,
inframedial, medial, or infrequently submarginal, usually round, less often oblong or elongate along veins in a few spp. (e.g., the
<typeStatus id="54A58842FFB09B126466782FFBDDFAC0" box="[1101,1146,1382,1403]" pageId="89" pageNumber="79">type</typeStatus>
), indusiate or exindusiate, indusia if present usually round-reniform or reduced to a fragment, usually whitish or tan when young, persistent to evanescent, hairy and/or glandular to glabrous (
<figureCitation id="13252A65FFB09B1262F878EFFCE3FA00" box="[723,836,1446,1467]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="89" pageNumber="79">
<figureCitation id="13252A65FFB09B1262F878EFFCE3FA00" box="[723,836,1446,1467]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="89" pageNumber="79">
<emphasis id="B96AEAF2FFB09B1262F878EFFCE3FA00" bold="true" box="[723,836,1446,1467]" pageId="89" pageNumber="79">Fig. 2D, 2E</emphasis>
</figureCitation>
);
@ -359,7 +361,7 @@ are highly localized, endemic to relatively small areas, and found mostly in pri
. heterocarpos
</emphasis>
,
<figureCitation id="13252A65FFB39B1161017EDBFED7FC1C" box="[298,368,914,936]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="90" pageNumber="80">
<figureCitation id="13252A65FFB39B1161017EDBFED7FC1C" box="[298,368,914,936]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="90" pageNumber="80">
<emphasis id="B96AEAF2FFB39B1161017EDBFED7FC1C" bold="true" box="[298,368,914,936]" pageId="90" pageNumber="80">Fig. 2K</emphasis>
</figureCitation>
) occur in disturbed forests, at forest margins, and sometimes in open areas; occasionally they grow on rocks or streamsides, where they tend to
@ -578,7 +580,7 @@ group of related, pinnatifid or weakly divided species with just a few pinna pai
<emphasis id="B96AEAF2FFB29B1061BF7FE4FDB0FD7A" box="[404,535,684,706]" italics="true" pageId="91" pageNumber="81">S. beccarianus</emphasis>
</taxonomicName>
(
<figureCitation id="13252A65FFB29B10620E7FE5FDCDFD7A" box="[549,618,684,706]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="91" pageNumber="81">
<figureCitation id="13252A65FFB29B10620E7FE5FDCDFD7A" box="[549,618,684,706]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="91" pageNumber="81">
<emphasis id="B96AEAF2FFB29B10620E7FE5FDCDFD7A" bold="true" box="[549,618,684,706]" pageId="91" pageNumber="81">Fig. 7B</emphasis>
</figureCitation>
), formerly included by
@ -645,7 +647,7 @@ clades includes many of the pinnate-pinnatifid taxa that tend to be hairy on bot
. heterocarpos
</emphasis>
,
<figureCitation id="13252A65FFB29B1063097EBAFCC0FBB3" box="[802,871,1011,1033]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="91" pageNumber="81">
<figureCitation id="13252A65FFB29B1063097EBAFCC0FBB3" box="[802,871,1011,1033]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="91" pageNumber="81">
<emphasis id="B96AEAF2FFB29B1063097EBAFCC0FBB3" bold="true" box="[802,871,1011,1033]" pageId="91" pageNumber="81">Fig. 2K</emphasis>
</figureCitation>
), while the second
@ -672,7 +674,7 @@ is sister to newly redefined
<taxonomicNameLabel id="A2595789FFB29B1064EF797CFB46FBF1" box="[1220,1249,1077,1098]" pageId="91" pageNumber="81" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
(Fawcett et al. in press;
<figureCitation id="13252A65FFB29B106179791CFE23FBD0" box="[338,388,1109,1131]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="91" pageNumber="81">Fig. 1</figureCitation>
<figureCitation id="13252A65FFB29B106179791CFE23FBD0" box="[338,388,1109,1131]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="91" pageNumber="81">Fig. 1</figureCitation>
). This clade is in turn sister to
<taxonomicName id="4C1E4D63FFB29B106289791CFCB5FBD1" ID-CoL="8RTZR" box="[674,786,1109,1130]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="91" pageNumber="81" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFB29B106289791CFCB5FBD1" box="[674,786,1109,1130]" italics="true" pageId="91" pageNumber="81">Reholttumia</emphasis>

40
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@ -1,29 +1,31 @@
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group,
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<figureCitation id="13252A65FFB79B15604A7D8AFF02FF63" box="[97,165,195,217]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="94" pageNumber="84">
<emphasis id="B96AEAF2FFB79B15604A7D8AFF02FF63" bold="true" box="[97,165,195,217]" pageId="94" pageNumber="84">Fig. 4E</emphasis>
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), forming areoles with or without included veinlets, or regularly anastomosing (
@ -126,7 +128,7 @@ group,
<taxonomicName id="4C1E4D63FFB79B1563987D8DFB95FF62" box="[947,1074,196,217]" class="Polypodiopsida" family="Thelypteridaceae" genus="Stegnogramma" kingdom="Plantae" order="Polypodiales" pageId="94" pageNumber="84" phylum="Tracheophyta" rank="species" species="aspidioides">S. aspidioides</taxonomicName>
,
</emphasis>
<figureCitation id="13252A65FFB79B1564177D8AFB23FF63" box="[1084,1156,195,217]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." pageId="94" pageNumber="84">
<figureCitation id="13252A65FFB79B1564177D8AFB23FF63" box="[1084,1156,195,217]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="94" pageNumber="84">
<emphasis id="B96AEAF2FFB79B1564177D8AFB23FF63" bold="true" box="[1084,1156,195,217]" pageId="94" pageNumber="84">Fig. 4D</emphasis>
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), often with several pairs of veins united below the sinus with excurrent vein running to the sinus; veins reaching laminar margins;
@ -173,7 +175,7 @@ group is quite distinctive within the family, with its irregularly anastomosing
<emphasis id="B96AEAF2FFB79B1561D17F1FFDFCFDD0" box="[506,603,598,619]" italics="true" pageId="94" pageNumber="84">S. wilfordii</emphasis>
</taxonomicName>
,
<figureCitation id="13252A65FFB79B15624D7F1CFD0EFDD1" box="[614,681,597,619]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." pageId="94" pageNumber="84">
<figureCitation id="13252A65FFB79B15624D7F1CFD0EFDD1" box="[614,681,597,619]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="94" pageNumber="84">
<emphasis id="B96AEAF2FFB79B15624D7F1CFD0EFDD1" bold="true" box="[614,681,597,619]" pageId="94" pageNumber="84">Fig. 4E</emphasis>
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), though these grade into somewhat more elongate laminae with up to six free pinna-pairs in the newly described Vietnamese endemic
@ -236,7 +238,7 @@ single species, the
<emphasis id="B96AEAF2FFB79B15616F7EECFD90FC01" box="[324,567,933,954]" italics="true" pageId="94" pageNumber="84">Stegnogramma aspidioides</emphasis>
</taxonomicName>
(
<figureCitation id="13252A65FFB79B15626F7EEDFD2EFC02" box="[580,649,932,954]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." pageId="94" pageNumber="84">
<figureCitation id="13252A65FFB79B15626F7EEDFD2EFC02" box="[580,649,932,954]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="94" pageNumber="84">
<emphasis id="B96AEAF2FFB79B15626F7EEDFD2EFC02" bold="true" box="[580,649,932,954]" pageId="94" pageNumber="84">Fig. 4D</emphasis>
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), extends into Java, Sumatra, and Borneo.
@ -364,7 +366,7 @@ are well-supported as sister to
<emphasis id="B96AEAF2FFB79B1562E17B49FCEFF9AE" box="[714,840,1536,1557]" italics="true" pageId="94" pageNumber="84">Cyclogramma</emphasis>
</taxonomicName>
on a long branch, together forming the stegnogrammoid subclade, which diverged early within the larger cyclosoroid clade (
<figureCitation id="13252A65FFB79B1563837B68FC78F98D" box="[936,991,1569,1591]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." pageId="94" pageNumber="84">
<figureCitation id="13252A65FFB79B1563837B68FC78F98D" box="[936,991,1569,1591]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="94" pageNumber="84">
<emphasis id="B96AEAF2FFB79B1563837B68FC78F98D" bold="true" box="[936,991,1569,1591]" pageId="94" pageNumber="84">Fig. 1</emphasis>
</figureCitation>
). The
@ -428,7 +430,7 @@ and
Blume
</taxonomicName>
(
<figureCitation id="13252A65FFB69B1462EB7DABFCA0FF4C" box="[704,775,226,247]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." pageId="95" pageNumber="85">
<figureCitation id="13252A65FFB69B1462EB7DABFCA0FF4C" box="[704,775,226,247]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="95" pageNumber="85">
<emphasis id="B96AEAF2FFB69B1462EB7DABFCA0FF4C" bold="true" box="[704,775,226,247]" pageId="95" pageNumber="85">Fig. 4D</emphasis>
</figureCitation>
); *
@ -466,7 +468,7 @@ Ching
(Hook.) Seriz
</taxonomicName>
(
<figureCitation id="13252A65FFB69B1462607C6BFD29FE8C" box="[587,654,290,311]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." pageId="95" pageNumber="85">
<figureCitation id="13252A65FFB69B1462607C6BFD29FE8C" box="[587,654,290,311]" captionStart="FIG" captionStartId="24.[97,105,1752,1772]" captionTargetBox="[112,1215,163,1706]" captionTargetId="figure-8@24.[110,1215,160,1709]" captionTargetPageId="24" captionText="FIG.4.A.Hoiokula sandwicensis,Aborn s.n. (VT),pinna lobes,abaxial view.B.Hoiokula pendens,Hobdy2664 (UC),pinna lobes,adaxial view.C.Leptogrammapilosa, Diaz 6621 (UC), abaxial pinna lobes. D. Stegnogramma aspidioides, Palmer 1035 (UC), abaxial pinna. E. Stegnogramma wilfordii, Boufford 20159 (VT), proximal portionof lamina,abaxial view." figureDoi="http://doi.org/10.5281/zenodo.14111054" httpUri="https://zenodo.org/record/14111054/files/figure.png" pageId="95" pageNumber="85">
<emphasis id="B96AEAF2FFB69B1462607C6BFD29FE8C" bold="true" box="[587,654,290,311]" pageId="95" pageNumber="85">Fig. 4E</emphasis>
</figureCitation>
).

48
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@ -172,7 +174,7 @@ Holttum
(Ces.) S.E. Fawc. &amp; A.R. Sm.
</taxonomicName>
(
<figureCitation id="13252A65FFB59B1760F679DAFEF2FB1C" box="[221,341,1170,1192]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." pageId="92" pageNumber="82">
<figureCitation id="13252A65FFB59B1760F679DAFEF2FB1C" box="[221,341,1170,1192]" captionStart="FIG" captionStartId="27.[121,129,1750,1770]" captionTargetBox="[124,1252,168,1700]" captionTargetId="figure-8@27.[119,1252,168,1705]" captionTargetPageId="27" captionText="FIG.7.A.Pneumatopteris parksii,Fawcett 645 (UC),pinna lobes,adaxial view.B.Reholttumia magnifica,Fawcett 676 (VT),pinna lobes,abaxial view.C.Pelazoneuron patens,Fawcett 744 (VT), abaxial pinna-lobes.D. P.serra,Fawcett 1034 (VT), adaxial pinna lobes.E. Pakau pennigera, Given 11842 (VT),pinna lobes,abaxial view. F.Pakau pennigera, Stuart s.n.(VT),pinna." figureDoi="http://doi.org/10.5281/zenodo.14111097" httpUri="https://zenodo.org/record/14111097/files/figure.png" pageId="92" pageNumber="82">
<emphasis id="B96AEAF2FFB59B1760F679DAFEF2FB1C" bold="true" box="[221,341,1170,1192]" pageId="92" pageNumber="82">Figs. 7A, 7B</emphasis>
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);
@ -294,7 +296,7 @@ Holttum
(Copel.) Holttum
</taxonomicName>
(
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<emphasis id="B96AEAF2FFB59B1763C2781AFB8CFAD3" bold="true" box="[1001,1067,1363,1384]" pageId="92" pageNumber="82">Fig. 2E</emphasis>
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);
@ -409,7 +411,7 @@ Holttum
. heterocarpos
</emphasis>
(Blume) Holttum (
<figureCitation id="13252A65FFB59B1760EB7B5AFEA1F993" box="[192,262,1555,1576]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="92" pageNumber="82">
<figureCitation id="13252A65FFB59B1760EB7B5AFEA1F993" box="[192,262,1555,1576]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="92" pageNumber="82">
<emphasis id="B96AEAF2FFB59B1760EB7B5AFEA1F993" bold="true" box="[192,262,1555,1576]" pageId="92" pageNumber="82">Fig. 2K</emphasis>
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@ -740,7 +742,7 @@ Holttum
. polycarpos
</emphasis>
(Blume) Copel. (
<figureCitation id="13252A65FFB49B1662447C8FFD1FFE60" box="[623,696,454,476]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." pageId="93" pageNumber="83">
<figureCitation id="13252A65FFB49B1662447C8FFD1FFE60" box="[623,696,454,476]" captionStart="FIG" captionStartId="22.[97,105,1615,1635]" captionTargetBox="[144,1177,161,1569]" captionTargetId="figure-8@22.[142,1177,161,1570]" captionTargetPageId="22" captionText="FIG. 2. Morphology of the Thelypteridaceae.A. Coryphopteris kolombangarae, SITW07554 (TAIF), glandular indusia.B. Goniopteris yaucoensis, Proctor 43584 (US), stellatehairs along adaxialcosta.C.Pneumatopterisglandulifera,SITW11069 (TAIF),peg-like aerophoresat pinna bases.D.Sphaerostephanospolycarpos,SITW11070 (TAIF),marginally glandular indusia.E.Sphaerostephanos doodioides,SITW05702(TAIF),setoseindusia,spherical glands on indusia,sporangia,and laminar tissue. F. Goniopteris reptans, Wright 813 (US), sori with shriveled, setose indusia, some hairs furcate. G. Macrothelypteris polypodioides, SITW11122 (TAIF), stipe scales. H. Hoiokula pendens, Hobdy 2664 (UC), setulose sporangia. I. Plesioneuron imbricatum, SITW05370 (TAIF), exindusiate sori, sporangia with spherical glands. J. Christella dentata, SITW10525 (TAIF), shriveled,setulose indusia.K. Sphaerostephanos heterocarpos SITW11079 (TAIF),abruptly reduced proximal pinnae,adaxial surface with acicular hairs.L. Coryphopteris subbipinnata, SITW11667 (TAIF),appressed stipe base scales,these marginally setulose.All photos by Cheng-Wei Chen except B,F,and H by Susan Fawcett." figureDoi="http://doi.org/10.5281/zenodo.14111046" httpUri="https://zenodo.org/record/14111046/files/figure.png" pageId="93" pageNumber="83">
<emphasis id="B96AEAF2FFB49B1662447C8FFD1FFE60" bold="true" box="[623,696,454,476]" pageId="93" pageNumber="83">Fig. 2D</emphasis>
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42
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