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<document id="E5EAAF41822D6E85C762089AB9D97CB7" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFE69D1B4BA505ADDA192128" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsia benedictae Shelley" docType="treatment" docVersion="7" lastPageNumber="98" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="96" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="87C0AD896151468726A32F850BFDC394">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="3C8B75B1207DD013403C0923255C2DEC" type="personal">
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<mods:namePart id="1EEA22C0B2BF078677F92A5E1063B122">Shear, William A.</mods:namePart>
<mods:affiliation id="F9D3DDAC6AAF0E09DA872129D3693DDE">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:namePart id="6E0325FC47523DF3AAFEE07D03017DC3">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="C52032AF9FDB2ECC7564DA8D872968FF" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="6FC6820839810FCA7773255D81A24EC3">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="6D345D66E083A5CAAA6366237C9D0EB9" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:title id="1300046AE0A39274CF0D098DD4B2D826">Zootaxa</mods:title>
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<mods:date id="7A598CB5BAEFB726F1C67B3C6C19D5DC">2021</mods:date>
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<mods:number id="B254E5354E50A908FC47469CEB4CFC89">2021-05-24</mods:number>
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<mods:number id="851952FD2341BDC155E8063E1DAD36A7">4975</mods:number>
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<treatment id="03B18797FFE69D1B4BA505ADDA192128" ID-DOI="http://doi.org/10.5281/zenodo.4923564" ID-GBIF-Taxon="182644907" ID-Zenodo-Dep="4923564" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128" httpUri="http://treatment.plazi.org/id/03B18797FFE69D1B4BA505ADDA192128" lastPageId="18" lastPageNumber="98" pageId="15" pageNumber="96">
<subSubSection id="C302650AFFE69D064BA505ADD8B220FC" box="[151,497,151,178]" pageId="15" pageNumber="96" type="nomenclature">
<paragraph id="8BA73681FFE69D064BA505ADD8B220FC" blockId="15.[151,497,151,213]" box="[151,497,151,178]" pageId="15" pageNumber="96">
<heading id="D0EF81EDFFE69D064BA505ADD8B220FC" bold="true" box="[151,497,151,178]" fontSize="11" level="1" pageId="15" pageNumber="96" reason="1">
<taxonomicName id="4C184D02FFE69D064BA505ADD8B220FC" ID-CoL="6X47F" authority="Shelley" authorityName="Shelley" box="[151,497,151,178]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="96" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFE69D064BA505ADD8B220FC" bold="true" box="[151,497,151,178]" pageId="15" pageNumber="96">
<emphasis id="B96CEA93FFE69D064BA505ADD8D320FF" bold="true" box="[151,400,151,178]" italics="true" pageId="15" pageNumber="96">Retrorsia benedictae</emphasis>
Shelley
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE69D064BA50581D80F2098" box="[151,332,187,213]" pageId="15" pageNumber="96" type="description">
<paragraph id="8BA73681FFE69D064BA50581D80F2098" blockId="15.[151,497,151,213]" box="[151,332,187,213]" pageId="15" pageNumber="96">
<figureCitation id="13232A04FFE69D064BA50581D8542098" box="[151,279,187,213]" captionStart-0="FIGURES 2931" captionStart-1="FIGURES 3236" captionStart-2="FIGURES 3739" captionStartId-0="15.[151,264,1927,1951]" captionStartId-1="16.[151,264,1903,1927]" captionStartId-2="17.[151,264,1714,1738]" captionTargetBox-0="[153,1436,376,1903]" captionTargetBox-1="[183,1403,181,1878]" captionTargetBox-2="[151,1436,416,1690]" captionTargetId-0="figure-43@15.[151,1436,376,1903]" captionTargetId-1="figure-24@16.[183,1403,181,1879]" captionTargetId-2="figure-137@17.[151,1436,416,1690]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURES 2931. Retrorsia benedictae Shelley. 29. Male, lateral view. 30. Head and anterior rings, dorsal view. 31. Pygidium, lateral view. SEM stub WS32-14." captionText-1="FIGURES 3236. Retrorsia benedictae Shelley. 32. Midbody ring, dorsal view. 33. Limbus. 34. Accessory sensory area on sixth segment of antenna, lateral view. 35. Gonopods in situ, posterior view. 36. Left gonopod, lateral view. SEM stub WS32-14. Abbreviations: ac, acropodite; cx, coxa; p, pulvillus; pf, prefemorite; rn, rounded notch distal to pulvillus; rp, retrorse process; tz, terminal zone." captionText-2="FIGURES 3739. Retrorsia benedictae Shelley. 37. Pulvillus of gonopod, posterior view. 38. Suspected fungal theca on antenna. 39. Hypopis of unidentified mite on fifth body ring. SEM stub 32-14." figureDoi-0="http://doi.org/10.5281/zenodo.4804943" figureDoi-1="http://doi.org/10.5281/zenodo.4804951" figureDoi-2="http://doi.org/10.5281/zenodo.4804957" httpUri-0="https://zenodo.org/record/4804943/files/figure.png" httpUri-1="https://zenodo.org/record/4804951/files/figure.png" httpUri-2="https://zenodo.org/record/4804957/files/figure.png" pageId="15" pageNumber="96">Figs 2939</figureCitation>
,
<figureCitation id="13232A04FFE69D064A100581D80F2098" box="[290,332,187,213]" captionStart="FIGURES 121126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="15" pageNumber="96">122</figureCitation>
</paragraph>
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<subSubSection id="C302650AFFE69D064BA5043ADB4E2155" box="[151,525,256,280]" pageId="15" pageNumber="96" type="reference_group">
<paragraph id="8BA73681FFE69D064BA5043ADB4E2155" blockId="15.[151,525,256,280]" box="[151,525,256,280]" pageId="15" pageNumber="96">
<treatmentCitationGroup id="AB0811AFFFE69D064BA5043ADB4E2155" box="[151,525,256,280]" pageId="15" pageNumber="96">
<treatmentCitation id="0AB91090FFE69D064BA5043ADB4E2155" author="Shelley, R. M." box="[151,525,256,280]" page="7" pageId="15" pageNumber="96" year="2003">
<taxonomicName id="4C184D02FFE69D064BA5043ADB4E2155" ID-CoL="6X47F" authority="Shelley, 2003: 7" authorityName="Shelley" authorityPageNumber="7" authorityYear="2003" box="[151,525,256,280]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="96" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFE69D064BA5043AD8292155" box="[151,362,256,280]" italics="true" pageId="15" pageNumber="96">Retrorsia benedictae</emphasis>
<bibRefCitation id="EF894B70FFE69D064A42043ADB4E2155" author="Shelley, R. M." box="[368,525,256,280]" pageId="15" pageNumber="96" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003:7</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="C302650AFFE69D064BA5047CDDD0212D" box="[151,1171,325,353]" pageId="15" pageNumber="96" type="materials_examined">
<paragraph id="8BA73681FFE69D064BA5047CDDD0212D" blockId="15.[151,1171,325,353]" box="[151,1171,325,353]" pageId="15" pageNumber="96">
<materialsCitation id="3B703CDCFFE69D064BA5047CD89E212D" ID-GBIF-Occurrence="3128519329" box="[151,477,325,352]" location="Along" pageId="15" pageNumber="96" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<emphasis id="B96CEA93FFE69D064BA5047CD8752112" bold="true" box="[151,310,325,352]" pageId="15" pageNumber="96">
<typeStatus id="54A38823FFE69D064BA5047CD991212D" box="[151,210,326,352]" pageId="15" pageNumber="96">Type</typeStatus>
locality.
</emphasis>
<location id="8EC7605AFFE69D064A0E047CD8C1212D" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFE69D064A0E047CD8C1212D" box="[316,386,326,352]" name="Along" pageId="15" pageNumber="96" stateProvince="Oregon">Along</location>
<collectingRegion id="49DCF863FFE69D064ABB047CD89E212D" box="[393,477,326,352]" country="United States of America" name="Oregon" pageId="15" pageNumber="96">Oregon</collectingRegion>
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<materialsCitation id="3B703CDCFFE69D064AD6047CDDD3212D" ID-GBIF-Occurrence="3128519306" box="[484,1168,326,353]" county="Clatsop County" location="Olney" pageId="15" pageNumber="96" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
Rt. 202, 3 mi (
<quantity id="4CE09B64FFE69D0649B4047CDB9B212C" box="[646,728,326,353]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="96" unit="km" value="4.8">4.8 km</quantity>
) SE of
<location id="8EC7605AFFE69D06481E047CDA32212D" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFE69D06481E047CDA32212D" box="[812,881,326,352]" county="Clatsop County" name="Olney" pageId="15" pageNumber="96" stateProvince="Oregon">Olney</location>
,
<collectingCounty id="62C64E0DFFE69D06484E047CDD6E212D" box="[892,1069,326,352]" pageId="15" pageNumber="96">Clatsop County</collectingCounty>
,
<collectingRegion id="49DCF863FFE69D064F0A047CDDD3212D" box="[1080,1168,326,352]" country="United States of America" name="Oregon" pageId="15" pageNumber="96">Oregon</collectingRegion>
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.
</paragraph>
</subSubSection>
<caption id="DF676609FFE69D064BA502BDD8AF278E" ID-DOI="http://doi.org/10.5281/zenodo.4804943" ID-Zenodo-Dep="4804943" httpUri="https://zenodo.org/record/4804943/files/figure.png" pageId="15" pageNumber="96" startId="15.[151,264,1927,1951]" targetBox="[153,1436,376,1903]" targetPageId="15">
<paragraph id="8BA73681FFE69D064BA502BDD8AF278E" blockId="15.[151,1437,1927,1987]" pageId="15" pageNumber="96">
<emphasis id="B96CEA93FFE69D064BA502BDD80E27D2" bold="true" box="[151,333,1927,1951]" pageId="15" pageNumber="96">FIGURES 2931</emphasis>
.
<taxonomicName id="4C184D02FFE69D064A6B02B2DBC027D2" authority="Shelley." authorityName="Shelley." box="[345,643,1927,1951]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="96" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFE69D064A6B02B2DB6F27D2" box="[345,556,1927,1951]" italics="true" pageId="15" pageNumber="96">Retrorsia benedictae</emphasis>
Shelley.
</taxonomicName>
<emphasis id="B96CEA93FFE69D0649BA02BDDBE227D2" bold="true" box="[648,673,1927,1951]" pageId="15" pageNumber="96">29</emphasis>
. Male, lateral view.
<emphasis id="B96CEA93FFE69D06485D02BDDACB27D2" bold="true" box="[879,904,1927,1951]" pageId="15" pageNumber="96">30</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFE69D064E3C02BDDC6427D2" bold="true" box="[1294,1319,1927,1951]" pageId="15" pageNumber="96">31</emphasis>
. Pygidium, lateral view. SEM stub WS32-14.
</paragraph>
</caption>
<caption id="DF676609FFF99D194BA50255D80A27BE" ID-DOI="http://doi.org/10.5281/zenodo.4804951" ID-Zenodo-Dep="4804951" httpUri="https://zenodo.org/record/4804951/files/figure.png" pageId="16" pageNumber="97" startId="16.[151,264,1903,1927]" targetBox="[183,1403,181,1878]" targetPageId="16">
<paragraph id="8BA73681FFF99D194BA50255D80A27BE" blockId="16.[151,1437,1903,2036]" pageId="16" pageNumber="97">
<emphasis id="B96CEA93FFF99D194BA50255D80C27CA" bold="true" box="[151,335,1903,1927]" pageId="16" pageNumber="97">FIGURES 3236</emphasis>
.
<taxonomicName id="4C184D02FFF99D194A6C024ADBCE27CA" authority="Shelley." authorityName="Shelley." box="[350,653,1903,1927]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFF99D194A6C024ADB7027CA" box="[350,563,1903,1927]" italics="true" pageId="16" pageNumber="97">Retrorsia benedictae</emphasis>
Shelley.
</taxonomicName>
<emphasis id="B96CEA93FFF99D1949A60255DBEE27CA" bold="true" box="[660,685,1903,1927]" pageId="16" pageNumber="97">32</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFF99D1948EC0255DAB427CA" bold="true" box="[990,1015,1903,1927]" pageId="16" pageNumber="97">33</emphasis>
. Limbus.
<emphasis id="B96CEA93FFF99D194F500255DD3827CA" bold="true" box="[1122,1147,1903,1927]" pageId="16" pageNumber="97">34</emphasis>
. Accessory sensory area on sixth segment of antenna, lateral view.
<emphasis id="B96CEA93FFF99D19492F02A9DB7527E6" bold="true" box="[541,566,1939,1963]" pageId="16" pageNumber="97">35</emphasis>
. Gonopods
<emphasis id="B96CEA93FFF99D19499D02AEDBAE27E6" box="[687,749,1940,1963]" italics="true" pageId="16" pageNumber="97">in situ</emphasis>
, posterior view.
<emphasis id="B96CEA93FFF99D1948A102A9DAEF27E6" bold="true" box="[915,940,1939,1963]" pageId="16" pageNumber="97">36</emphasis>
. Left gonopod, lateral view. SEM stub WS32-14. Abbreviations:
<emphasis id="B96CEA93FFF99D194A060282D80F279D" bold="true" box="[308,332,1976,2000]" pageId="16" pageNumber="97">ac</emphasis>
, acropodite;
<emphasis id="B96CEA93FFF99D194AE20282D8AB279D" bold="true" box="[464,488,1976,2000]" pageId="16" pageNumber="97">cx</emphasis>
, coxa;
<emphasis id="B96CEA93FFF99D1949030282DB7C279D" bold="true" box="[561,575,1976,2000]" pageId="16" pageNumber="97">p</emphasis>
, pulvillus;
<emphasis id="B96CEA93FFF99D194983028DDB8B2782" bold="true" box="[689,712,1975,1999]" pageId="16" pageNumber="97">pf</emphasis>
, prefemorite;
<emphasis id="B96CEA93FFF99D1948650282DA33279D" bold="true" box="[855,880,1976,2000]" pageId="16" pageNumber="97">rn</emphasis>
, rounded notch distal to pulvillus;
<emphasis id="B96CEA93FFF99D194FE20282DDAA279D" bold="true" box="[1232,1257,1976,2000]" pageId="16" pageNumber="97">rp</emphasis>
, retrorse process;
<emphasis id="B96CEA93FFF99D194BA502E6D9E827B9" bold="true" box="[151,171,2012,2036]" pageId="16" pageNumber="97">tz</emphasis>
, terminal zone.
</paragraph>
</caption>
<subSubSection id="C302650AFFF89D184BF505ACDDFD2129" pageId="17" pageNumber="98" type="diagnosis">
<paragraph id="8BA73681FFF89D184BF505ACDD6220B5" blockId="17.[151,1436,150,393]" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D184BF505ACD80120FD" bold="true" box="[199,322,150,176]" pageId="17" pageNumber="98">Diagnosis.</emphasis>
The semicircular notch in the gonopod is prominent (
<emphasis id="B96CEA93FFF89D1848A005ACDAF620FD" bold="true" box="[914,949,150,176]" pageId="17" pageNumber="98">rn,</emphasis>
<figureCitation id="13232A04FFF89D18488905ACDD4920FC" box="[955,1034,150,177]" captionStart="FIGURES 3236" captionStartId="16.[151,264,1903,1927]" captionTargetBox="[183,1403,181,1878]" captionTargetId="figure-24@16.[183,1403,181,1879]" captionTargetPageId="16" captionText="FIGURES 3236. Retrorsia benedictae Shelley. 32. Midbody ring, dorsal view. 33. Limbus. 34. Accessory sensory area on sixth segment of antenna, lateral view. 35. Gonopods in situ, posterior view. 36. Left gonopod, lateral view. SEM stub WS32-14. Abbreviations: ac, acropodite; cx, coxa; p, pulvillus; pf, prefemorite; rn, rounded notch distal to pulvillus; rp, retrorse process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804951" httpUri="https://zenodo.org/record/4804951/files/figure.png" pageId="17" pageNumber="98">Fig. 36</figureCitation>
) but separated from the pulvillus (
<emphasis id="B96CEA93FFF89D184EB405ACDCDF20FD" bold="true" box="[1414,1436,150,176]" pageId="17" pageNumber="98">p,</emphasis>
<figureCitation id="13232A04FFF89D184BA50580D9A42098" box="[151,231,186,213]" captionStart="FIGURES 3236" captionStartId="16.[151,264,1903,1927]" captionTargetBox="[183,1403,181,1878]" captionTargetId="figure-24@16.[183,1403,181,1879]" captionTargetPageId="16" captionText="FIGURES 3236. Retrorsia benedictae Shelley. 32. Midbody ring, dorsal view. 33. Limbus. 34. Accessory sensory area on sixth segment of antenna, lateral view. 35. Gonopods in situ, posterior view. 36. Left gonopod, lateral view. SEM stub WS32-14. Abbreviations: ac, acropodite; cx, coxa; p, pulvillus; pf, prefemorite; rn, rounded notch distal to pulvillus; rp, retrorse process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804951" httpUri="https://zenodo.org/record/4804951/files/figure.png" pageId="17" pageNumber="98">Fig. 35</figureCitation>
) by half the length of the terminal zone (see also
<figureCitation id="13232A04FFF89D18483D0580DA2C2099" box="[783,879,186,212]" captionStart="FIGURES 121126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="17" pageNumber="98">Fig. 121</figureCitation>
). There are 16 or 18 setae in the anterior marginal row of the collum (
<figureCitation id="13232A04FFF89D184A5C05E4D88020B5" box="[366,451,222,248]" captionStart="FIGURES 2931" captionStartId="15.[151,264,1927,1951]" captionTargetBox="[153,1436,376,1903]" captionTargetId="figure-43@15.[151,1436,376,1903]" captionTargetPageId="15" captionText="FIGURES 2931. Retrorsia benedictae Shelley. 29. Male, lateral view. 30. Head and anterior rings, dorsal view. 31. Pygidium, lateral view. SEM stub WS32-14." figureDoi="http://doi.org/10.5281/zenodo.4804943" httpUri="https://zenodo.org/record/4804943/files/figure.png" pageId="17" pageNumber="98">Fig. 30</figureCitation>
). A detailed description is available in
<bibRefCitation id="EF894B70FFF89D18484605E4DD5E20B5" author="Shelley, R. M." box="[884,1053,222,248]" pageId="17" pageNumber="98" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
.
</paragraph>
<paragraph id="8BA73681FFF89D184BF50438DDFD2129" blockId="17.[151,1436,150,393]" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D184BF50438D8522151" bold="true" box="[199,273,258,284]" pageId="17" pageNumber="98">Notes.</emphasis>
The male specimen we used for SEM examination (
<figureCitation id="13232A04FFF89D1848640438DA952151" box="[854,982,258,284]" captionStart-0="FIGURES 2931" captionStart-1="FIGURES 3236" captionStart-2="FIGURES 3739" captionStartId-0="15.[151,264,1927,1951]" captionStartId-1="16.[151,264,1903,1927]" captionStartId-2="17.[151,264,1714,1738]" captionTargetBox-0="[153,1436,376,1903]" captionTargetBox-1="[183,1403,181,1878]" captionTargetBox-2="[151,1436,416,1690]" captionTargetId-0="figure-43@15.[151,1436,376,1903]" captionTargetId-1="figure-24@16.[183,1403,181,1879]" captionTargetId-2="figure-137@17.[151,1436,416,1690]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURES 2931. Retrorsia benedictae Shelley. 29. Male, lateral view. 30. Head and anterior rings, dorsal view. 31. Pygidium, lateral view. SEM stub WS32-14." captionText-1="FIGURES 3236. Retrorsia benedictae Shelley. 32. Midbody ring, dorsal view. 33. Limbus. 34. Accessory sensory area on sixth segment of antenna, lateral view. 35. Gonopods in situ, posterior view. 36. Left gonopod, lateral view. SEM stub WS32-14. Abbreviations: ac, acropodite; cx, coxa; p, pulvillus; pf, prefemorite; rn, rounded notch distal to pulvillus; rp, retrorse process; tz, terminal zone." captionText-2="FIGURES 3739. Retrorsia benedictae Shelley. 37. Pulvillus of gonopod, posterior view. 38. Suspected fungal theca on antenna. 39. Hypopis of unidentified mite on fifth body ring. SEM stub 32-14." figureDoi-0="http://doi.org/10.5281/zenodo.4804943" figureDoi-1="http://doi.org/10.5281/zenodo.4804951" figureDoi-2="http://doi.org/10.5281/zenodo.4804957" httpUri-0="https://zenodo.org/record/4804943/files/figure.png" httpUri-1="https://zenodo.org/record/4804951/files/figure.png" httpUri-2="https://zenodo.org/record/4804957/files/figure.png" pageId="17" pageNumber="98">Figs 2939</figureCitation>
) had some interesting commensals/parasites, including a fungal capilliconidia (amphoromorph) of the genus
<taxonomicName id="4C184D02FFF89D184893041CDC3A210D" authority="(Enghoff &amp; Reboleira 2017)" baseAuthorityName="Enghoff &amp; Reboleira" baseAuthorityYear="2017" box="[929,1401,294,320]" family="Basidiobolaceae" genus="Basidiobolus" kingdom="Fungi" order="Basidiobolales" pageId="17" pageNumber="98" phylum="Zygomycota" rank="genus">
<emphasis id="B96CEA93FFF89D184893041CDD70210D" box="[929,1075,294,320]" italics="true" pageId="17" pageNumber="98">Basidiobolus</emphasis>
(
<bibRefCitation id="EF894B70FFF89D184F70041CDC31210D" author="Enghoff, H. &amp; Reboleira, A. S. P. S." box="[1090,1394,294,320]" pageId="17" pageNumber="98" pagination="130 - 137" refId="ref19602" refString="Enghoff, H. &amp; Reboleira, A. S. P. S. (2017) Diversity of non - Laboulbenialean fungi on millipedes. Studies in Fungi, 2, 130 - 137. https: // doi. org / 10.5943 / sif / 2 / 1 / 15" type="journal article" year="2017">Enghoff &amp; Reboleira 2017</bibRefCitation>
)
</taxonomicName>
on an antenna (
<figureCitation id="13232A04FFF89D184A2D0470D8332129" box="[287,368,330,356]" captionStart="FIGURES 3739" captionStartId="17.[151,264,1714,1738]" captionTargetBox="[151,1436,416,1690]" captionTargetId="figure-137@17.[151,1436,416,1690]" captionTargetPageId="17" captionText="FIGURES 3739. Retrorsia benedictae Shelley. 37. Pulvillus of gonopod, posterior view. 38. Suspected fungal theca on antenna. 39. Hypopis of unidentified mite on fifth body ring. SEM stub 32-14." figureDoi="http://doi.org/10.5281/zenodo.4804957" httpUri="https://zenodo.org/record/4804957/files/figure.png" pageId="17" pageNumber="98">Fig. 38</figureCitation>
) and the hypopis of an unidentified mite attached to the fifth ring (
<figureCitation id="13232A04FFF89D184F6F0470DDF22129" box="[1117,1201,330,356]" captionStart="FIGURES 3739" captionStartId="17.[151,264,1714,1738]" captionTargetBox="[151,1436,416,1690]" captionTargetId="figure-137@17.[151,1436,416,1690]" captionTargetPageId="17" captionText="FIGURES 3739. Retrorsia benedictae Shelley. 37. Pulvillus of gonopod, posterior view. 38. Suspected fungal theca on antenna. 39. Hypopis of unidentified mite on fifth body ring. SEM stub 32-14." figureDoi="http://doi.org/10.5281/zenodo.4804957" httpUri="https://zenodo.org/record/4804957/files/figure.png" pageId="17" pageNumber="98">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF89D1B4BF50454DA192128" lastPageId="18" lastPageNumber="99" pageId="17" pageNumber="98" type="distribution">
<paragraph id="8BA73681FFF89D184BF50454DC7821C4" blockId="17.[151,1436,150,393]" box="[199,1339,366,393]" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D184BF50454D82221C5" bold="true" box="[199,353,366,392]" pageId="17" pageNumber="98">Distribution.</emphasis>
Clatsop Co.,
<collectingRegion id="49DCF863FFF89D184AC80454DB0D21C5" box="[506,590,366,392]" country="United States of America" name="Oregon" pageId="17" pageNumber="98">Oregon</collectingRegion>
(
<bibRefCitation id="EF894B70FFF89D18496E0454DBB621C5" author="Shelley, R. M." box="[604,757,366,392]" pageId="17" pageNumber="98" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley 2003</bibRefCitation>
), and Pacific Co.,
<collectingRegion id="49DCF863FFF89D1848F00454DD0421C5" box="[962,1095,366,392]" country="United States of America" name="Washington" pageId="17" pageNumber="98">Washington</collectingRegion>
(new records below).
</paragraph>
<caption id="DF676609FFF89D184BA50388DAD926A3" ID-DOI="http://doi.org/10.5281/zenodo.4804957" ID-Zenodo-Dep="4804957" httpUri="https://zenodo.org/record/4804957/files/figure.png" pageId="17" pageNumber="98" startId="17.[151,264,1714,1738]" targetBox="[151,1436,416,1690]" targetPageId="17">
<paragraph id="8BA73681FFF89D184BA50388DAD926A3" blockId="17.[151,1436,1714,1774]" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D184BA50388D80C2687" bold="true" box="[151,335,1714,1738]" pageId="17" pageNumber="98">FIGURES 3739</emphasis>
.
<taxonomicName id="4C184D02FFF89D184A6F0389DBC82687" authority="Shelley." authorityName="Shelley." box="[349,651,1714,1738]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFF89D184A6F0389DB712687" box="[349,562,1714,1738]" italics="true" pageId="17" pageNumber="98">Retrorsia benedictae</emphasis>
Shelley.
</taxonomicName>
<emphasis id="B96CEA93FFF89D1849A00388DBE82687" bold="true" box="[658,683,1714,1738]" pageId="17" pageNumber="98">37</emphasis>
. Pulvillus of gonopod, posterior view.
<emphasis id="B96CEA93FFF89D184F0D0388DD1B2687" bold="true" box="[1087,1112,1714,1738]" pageId="17" pageNumber="98">38</emphasis>
. Suspected fungal theca on antenna.
<emphasis id="B96CEA93FFF89D184BE903ECD9B726A3" bold="true" box="[219,244,1750,1774]" pageId="17" pageNumber="98">39</emphasis>
. Hypopis of unidentified mite on fifth body ring. SEM stub 32-14.
</paragraph>
</caption>
<paragraph id="8BA73681FFF89D1B4BF50227DA192128" blockId="17.[151,1437,1820,2027]" lastBlockId="18.[151,1437,151,357]" lastPageId="18" lastPageNumber="99" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D184BF50227D824277A" bold="true" box="[199,359,1821,1847]" pageId="17" pageNumber="98">New records.</emphasis>
<materialsCitation id="3B703CDCFFF89D184A400227DADA2717" ID-GBIF-Occurrence="3128519326" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<collectingRegion id="49DCF863FFF89D184A400227DB72277A" box="[370,561,1821,1847]" country="United States of America" name="Washington" pageId="17" pageNumber="98">WASHINGTON</collectingRegion>
:
<collectingCounty id="62C64E0DFFF89D18490D0227DB83277A" box="[575,704,1821,1847]" pageId="17" pageNumber="98">
<emphasis id="B96CEA93FFF89D18490D0227DBF9277A" box="[575,698,1821,1847]" italics="true" pageId="17" pageNumber="98">Pacific Co</emphasis>
.
</collectingCounty>
: 1.0 mi on
<location id="8EC7605AFFF89D18487D0226DAB6277B" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D18487D0226DAB6277B" box="[847,1013,1820,1846]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Walberg Road" pageId="17" pageNumber="98" stateProvince="Washington">Walberg Road</location>
,
<quantity id="4CE09B64FFF89D184F360227DD13277B" box="[1028,1104,1820,1847]" metricMagnitude="2" metricUnit="m" metricValue="6.437376" pageId="17" pageNumber="98" unit="mi" value="0.4">0.4 mi</quantity>
west of
<location id="8EC7605AFFF89D184F880227DCDB277B" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184F880227DCDB277B" box="[1210,1432,1820,1847]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Icwaco Water Park" pageId="17" pageNumber="98" stateProvince="Washington">Icwaco Water Park</location>
, 46.3236°, -123.9417°,
<date id="FFA61041FFF89D184AAB027BDB0B2716" box="[409,584,1856,1883]" pageId="17" pageNumber="98" value="2005-01-02">
<collectingDate id="EFE2E9A9FFF89D184AAB027BDB0B2716" box="[409,584,1856,1883]" pageId="17" pageNumber="98" value="2005-01-02">2 January 2005</collectingDate>
</date>
,
<collectorName id="26ED5357FFF89D184966027BDB982717" box="[596,731,1856,1883]" pageId="17" pageNumber="98">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFF89D1849D5027BDA1E2717" box="[743,861,1856,1883]" pageId="17" pageNumber="98">C. Richart</collectorName>
, m f;
</materialsCitation>
<materialsCitation id="3B703CDCFFF89D184892027BDA9F2732" ID-GBIF-Occurrence="3128519305" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<quantity id="4CE09B64FFF89D184892027BDAAA2717" box="[928,1001,1856,1883]" metricMagnitude="3" metricUnit="m" metricValue="1.7702784" pageId="17" pageNumber="98" unit="mi" value="1.1">1.1 mi</quantity>
south of SR6
<collectingMethod id="52594E96FFF89D184FBE027BDDA72716" box="[1164,1252,1857,1883]" pageId="17" pageNumber="98">on Trap</collectingMethod>
<location id="8EC7605AFFF89D184FD9027ADC372717" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184FD9027ADC372717" box="[1259,1396,1856,1882]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Creek Road" pageId="17" pageNumber="98" stateProvince="Washington">Creek Road</location>
,
<location id="8EC7605AFFF89D184EB2027BD9872733" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184EB2027BD9872733" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Bline" pageId="17" pageNumber="98" stateProvince="Washington">Bline</location>
, 46.5432°-123.6152°,
<date id="FFA61041FFF89D184AF0025FDBDD2732" box="[450,670,1892,1919]" pageId="17" pageNumber="98" value="2005-11-19">
<collectingDate id="EFE2E9A9FFF89D184AF0025FDBDD2732" box="[450,670,1892,1919]" pageId="17" pageNumber="98" value="2005-11-19">19 November 2005</collectingDate>
</date>
,
<collectorName id="26ED5357FFF89D18499B025FDA732733" box="[681,816,1892,1919]" pageId="17" pageNumber="98">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFF89D18480E025FDAF12733" box="[828,946,1892,1919]" pageId="17" pageNumber="98">C. Richart</collectorName>
, m;
</materialsCitation>
<materialsCitation id="3B703CDCFFF89D1848D1025FDD5627EF" ID-GBIF-Occurrence="3128519323" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<quantity id="4CE09B64FFF89D1848D1025FDD542733" box="[995,1047,1892,1919]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="17" pageNumber="98" unit="mi" value="2.0">2 mi</quantity>
west of
<location id="8EC7605AFFF89D184F44025EDC5F2733" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184F44025EDC5F2733" box="[1142,1308,1892,1918]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Astoria Bridge" pageId="17" pageNumber="98" stateProvince="Washington">Astoria Bridge</location>
on
<collectingCountry id="F30F7611FFF89D184E75025FDC2D2732" box="[1351,1390,1893,1919]" name="United States of America" pageId="17" pageNumber="98">US</collectingCountry>
101, 10 ft asl,
<geoCoordinate id="EE2C5046FFF89D184A3002B2D82327EE" box="[258,352,1928,1955]" degrees="46.2569" direction="north" orientation="latitude" pageId="17" pageNumber="98" precision="5" value="46.2569">46.2569</geoCoordinate>
,
<geoCoordinate id="EE2C5046FFF89D184A5902B2D8A027EE" box="[363,483,1928,1955]" degrees="123.9250" direction="west" orientation="longitude" pageId="17" pageNumber="98" precision="5" value="-123.925">-123.9250</geoCoordinate>
°,
<date id="FFA61041FFF89D184ACA02B3DBE427EE" box="[504,679,1928,1955]" pageId="17" pageNumber="98" value="2005-01-02">
<collectingDate id="EFE2E9A9FFF89D184ACA02B3DBE427EE" box="[504,679,1928,1955]" pageId="17" pageNumber="98" value="2005-01-02">2 January 2005</collectingDate>
</date>
,
<collectorName id="26ED5357FFF89D18498102B3DA7A27EF" box="[691,825,1928,1955]" pageId="17" pageNumber="98">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFF89D18487702B3DAFF27EF" box="[837,956,1928,1955]" pageId="17" pageNumber="98">C. Richart</collectorName>
, mm ff;
</materialsCitation>
<materialsCitation id="3B703CDCFFF89D184F2F02B2DDC5278A" ID-GBIF-Occurrence="3128519303" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<quantity id="4CE09B64FFF89D184F2F02B2DD2527EF" box="[1053,1126,1928,1955]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="17" pageNumber="98" unit="mi" value="3.5">3.5 mi</quantity>
south of
<location id="8EC7605AFFF89D184FE202B2DC6727EF" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184FE202B2DC6727EF" box="[1232,1316,1928,1954]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Naselle" pageId="17" pageNumber="98" stateProvince="Washington">Naselle</location>
on SR401 at
<location id="8EC7605AFFF89D184B860297D818278B" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184B860297D818278B" box="[180,347,1964,1991]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Cement Creek" pageId="17" pageNumber="98" stateProvince="Washington">Cement Creek</location>
, 46.3341°, -123.8003°,
<date id="FFA61041FFF89D18495E0296DA6F278B" box="[620,812,1964,1991]" pageId="17" pageNumber="98" value="2006-01-15">
<collectingDate id="EFE2E9A9FFF89D18495E0296DA6F278B" box="[620,812,1964,1991]" pageId="17" pageNumber="98" value="2006-01-15">15 January 2006</collectingDate>
</date>
,
<collectorName id="26ED5357FFF89D18480A0297DA83278B" box="[824,960,1964,1991]" pageId="17" pageNumber="98">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFF89D1848FC0297DD05278B" box="[974,1094,1964,1991]" pageId="17" pageNumber="98">C. Richart</collectorName>
, mm;
</materialsCitation>
<materialsCitation id="3B703CDCFFF89D184FBD0296DC0927A7" ID-GBIF-Occurrence="3128519327" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<quantity id="4CE09B64FFF89D184FBD0296DD99278B" box="[1167,1242,1964,1991]" metricMagnitude="3" metricUnit="m" metricValue="7.242048" pageId="17" pageNumber="98" unit="mi" value="4.5">4.5 mi</quantity>
north of SR4 on
<collectingCountry id="F30F7611FFF89D184BA502EBD9FD27A6" box="[151,190,2001,2027]" name="United States of America" pageId="17" pageNumber="98">US</collectingCountry>
101, at
<location id="8EC7605AFFF89D184A2002EADB4627A7" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFF89D184A2002EADB4627A7" box="[274,517,2000,2026]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Middle Nemah River" pageId="17" pageNumber="98" stateProvince="Washington">Middle Nemah River</location>
, 46.4874°, -123.8865°,
<date id="FFA61041FFF89D18482402EADA9627A7" box="[790,981,2000,2027]" pageId="17" pageNumber="98" value="2006-01-15">
<collectingDate id="EFE2E9A9FFF89D18482402EADA9627A7" box="[790,981,2000,2027]" pageId="17" pageNumber="98" value="2006-01-15">15 January 2006</collectingDate>
</date>
,
<collectorName id="26ED5357FFF89D1848D002EBDD2927A7" box="[994,1130,2000,2027]" pageId="17" pageNumber="98">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFF89D184F4502EBDDAD27A7" box="[1143,1262,2000,2027]" pageId="17" pageNumber="98">C. Richart</collectorName>
, mm ff;
</materialsCitation>
<materialsCitation id="3B703CDCFFF89D1B4E6102EADB622098" ID-GBIF-Occurrence="3128519316" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" lastPageId="18" lastPageNumber="99" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="17" pageNumber="98" specimenCount="1" stateProvince="Washington">
<quantity id="4CE09B64FFF89D184E6102EADCDE27A7" box="[1363,1437,2000,2027]" metricMagnitude="3" metricUnit="m" metricValue="9.4951296" pageId="17" pageNumber="98" unit="mi" value="5.9">5.9 mi</quantity>
south of SR6
<collectingMethod id="52594E96FFFB9D1B4A0A05ADD8D220FC" box="[312,401,151,177]" pageId="18" pageNumber="99">on Trap</collectingMethod>
<collectorName id="26ED5357FFFB9D1B4AA805ADDB6620FC" box="[410,549,151,177]" pageId="18" pageNumber="99">Creek Road</collectorName>
,
<location id="8EC7605AFFFB9D1B490305ADDBFE20FC" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFFB9D1B490305ADDBFE20FC" box="[561,701,151,177]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Alder Creek" pageId="18" pageNumber="99" stateProvince="Washington">Alder Creek</location>
drainage of
<location id="8EC7605AFFFB9D1B486305ADDAAD20FC" LSID="urn:lsid:plazi:treatment:03B18797FFE69D1B4BA505ADDA192128:8EC7605AFFFB9D1B486305ADDAAD20FC" box="[849,1006,151,177]" country="United States of America" county="Pacific Co." latitude="46.2569" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" name="Naselle River" pageId="18" pageNumber="99" stateProvince="Washington">Naselle River</location>
, 46.4973°, -123.6439°,
<date id="FFA61041FFFB9D1B4E3305ADD9912098" pageId="18" pageNumber="99" value="2005-11-19">
<collectingDate id="EFE2E9A9FFFB9D1B4E3305ADD9912098" pageId="18" pageNumber="99" value="2005-11-19">19 November 2005</collectingDate>
</date>
,
<collectorName id="26ED5357FFFB9D1B4BD30581D8292098" box="[225,362,187,213]" pageId="18" pageNumber="99">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFFB9D1B4A480581D8B02098" box="[378,499,187,213]" pageId="18" pageNumber="99">C. Richart</collectorName>
, m;
</materialsCitation>
<materialsCitation id="3B703CDCFFFB9D1B49190581DA0720B4" ID-GBIF-Occurrence="3128519304" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="18" pageNumber="99" specimenCount="1" stateProvince="Washington">
east side of
<collectingMunicipality id="6BC3ACFBFFFB9D1B498E0581DA392098" box="[700,890,187,213]" pageId="18" pageNumber="99">Ellsworth Creek</collectingMunicipality>
,
<collectorName id="26ED5357FFFB9D1B48BA0581DC552098" box="[904,1302,187,213]" pageId="18" pageNumber="99">The Nature Conservency Preserve</collectorName>
,
<elevation id="0035D1B2FFFB9D1B4E170581DCDB2098" box="[1317,1432,187,213]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="18" pageNumber="99" unit="ft" value="100.0">
<quantity id="4CE09B64FFFB9D1B4E170581DC282098" box="[1317,1387,187,213]" metricMagnitude="1" metricUnit="m" metricValue="3.048" pageId="18" pageNumber="99" unit="ft" value="100.0">100 ft</quantity>
asl
</elevation>
, 46.3838°, -123.8679°,
<date id="FFA61041FFFB9D1B4AAB05E5DB1B20B4" box="[409,600,223,249]" pageId="18" pageNumber="99" value="2004-01-12">
<collectingDate id="EFE2E9A9FFFB9D1B4AAB05E5DB1B20B4" box="[409,600,223,249]" pageId="18" pageNumber="99" value="2004-01-12">12 January 2004</collectingDate>
</date>
,
<collectorName id="26ED5357FFFB9D1B495105E5DBA920B4" box="[611,746,223,249]" pageId="18" pageNumber="99">W. Leonard</collectorName>
, mm ff;
</materialsCitation>
<materialsCitation id="3B703CDCFFFB9D1B487E05E5DA852150" ID-GBIF-Occurrence="3128519309" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="18" pageNumber="99" specimenCount="1" stateProvince="Washington">
<collectorName id="26ED5357FFFB9D1B487E05E5DA9A20B4" box="[844,985,223,249]" pageId="18" pageNumber="99">Long Beach</collectorName>
, 30th
<collectorName id="26ED5357FFFB9D1B4F2D05E5DD7C20B4" box="[1055,1087,223,249]" pageId="18" pageNumber="99">St.</collectorName>
,
<quantity id="4CE09B64FFFB9D1B4F7E05E5DDD620B4" box="[1100,1173,223,249]" metricMagnitude="2" metricUnit="m" metricValue="3.218688" pageId="18" pageNumber="99" unit="mi" value="0.2">0.2 mi</quantity>
from
<collectingCountry id="F30F7611FFFB9D1B4FEE05E5DC4020B4" box="[1244,1283,223,249]" name="United States of America" pageId="18" pageNumber="99">US</collectingCountry>
101, 50 ft asl, 46.3239°, -124.0598°,
<date id="FFA61041FFFB9D1B4AAB0438DB142150" box="[409,599,258,285]" pageId="18" pageNumber="99" value="2006-01-15">
<collectingDate id="EFE2E9A9FFFB9D1B4AAB0438DB142150" box="[409,599,258,285]" pageId="18" pageNumber="99" value="2006-01-15">15 January 2006</collectingDate>
</date>
,
<collectorName id="26ED5357FFFB9D1B49500439DBAA2150" box="[610,745,259,285]" pageId="18" pageNumber="99">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFFB9D1B49C70439DA2F2150" box="[757,876,259,285]" pageId="18" pageNumber="99">C. Richart</collectorName>
, mm ff;
</materialsCitation>
<materialsCitation id="3B703CDCFFFB9D1B48FC0439DD6F210C" ID-GBIF-Occurrence="3128519340" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard &amp; C. Richart" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="18" pageNumber="99" specimenCount="1" stateProvince="Washington">
<collectorName id="26ED5357FFFB9D1B48FC0439DD192150" box="[974,1114,259,285]" pageId="18" pageNumber="99">Long Beach</collectorName>
,
<collectorName id="26ED5357FFFB9D1B4F570439DC4A2150" box="[1125,1289,259,285]" pageId="18" pageNumber="99">Willows Road</collectorName>
,
<quantity id="4CE09B64FFFB9D1B4E270439DC1D2150" box="[1301,1374,259,285]" metricMagnitude="2" metricUnit="m" metricValue="9.656064" pageId="18" pageNumber="99" unit="mi" value="0.6">0.6 mi</quantity>
from 30th
<collectorName id="26ED5357FFFB9D1B4BE3041DD9B2210C" box="[209,241,295,321]" pageId="18" pageNumber="99">St.</collectorName>
, 46.3229°, -124.0575°,
<date id="FFA61041FFFB9D1B4932041DDBFD210C" box="[512,702,295,321]" pageId="18" pageNumber="99" value="2006-01-16">
<collectingDate id="EFE2E9A9FFFB9D1B4932041DDBFD210C" box="[512,702,295,321]" pageId="18" pageNumber="99" value="2006-01-16">16 January 2006</collectingDate>
</date>
,
<collectorName id="26ED5357FFFB9D1B49FB041DDA13210C" box="[713,848,295,321]" pageId="18" pageNumber="99">W. Leonard</collectorName>
,
<collectorName id="26ED5357FFFB9D1B486E041DDA90210C" box="[860,979,295,321]" pageId="18" pageNumber="99">C. Richart</collectorName>
, mm ff;
</materialsCitation>
<materialsCitation id="3B703CDCFFFB9D1B4F06041DDA152128" ID-GBIF-Occurrence="3128519313" collectingDate="2004-01-12" collectingDateMax="2006-01-16" collectingDateMin="2004-01-12" collectorName="W. Leonard" country="United States of America" county="Pacific Co." elevation="30" latitude="46.2569" location="Walberg Road" longLatPrecision="6" longitude="-123.925" municipality="Ellsworth Creek" pageId="18" pageNumber="99" specimenCount="1" stateProvince="Washington">
SR401
<quantity id="4CE09B64FFFB9D1B4FBB041DDDFE210C" box="[1161,1213,295,321]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="18" pageNumber="99" unit="mi" value="8.0">8 mi</quantity>
south of SR4, 40 ft asl, 46.2752°, -123.8158°,
<date id="FFA61041FFFB9D1B4AF60471DB312128" box="[452,626,330,357]" pageId="18" pageNumber="99" value="2005-01-02">
<collectingDate id="EFE2E9A9FFFB9D1B4AF60471DB312128" box="[452,626,330,357]" pageId="18" pageNumber="99" value="2005-01-02">2 January 2005</collectingDate>
</date>
,
<collectorName id="26ED5357FFFB9D1B494F0471DA412128" box="[637,770,331,357]" pageId="18" pageNumber="99">W. Leonard</collectorName>
, mm ff
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>