162 lines
16 KiB
XML
162 lines
16 KiB
XML
<document id="5A5FF74235771B30B39C080EA12B87F6" ID-DOI="10.1649/0010-065X-76.4.612" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252933" ID-ZooBank="urn:lsid:zoobank.org:pub:6DA30E51-0FD4-48A5-B7B7-744472F20B9F" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723003944909" checkinUser="felipe" docAuthor="Skelley, Paul E. & Vaz-de-Mello, Fernando Z." docDate="2022" docId="038C6452FFC9C34EA22B48692E906E39" docLanguage="en" docName="TheColeopBull.76.4.612-618.pdf" docOrigin="The Coleopterists Bulletin 76 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-76.4.612" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Batesiana CHALUMEAU 1983" docType="treatment" docVersion="2" lastPageNumber="613" masterDocId="FFB51C2AFFCBC34CA3394C732A396D31" masterDocTitle="A Second Species of Batesiana Chalumeau, 1983 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="618" masterPageNumber="612" pageNumber="613" updateTime="1723006876623" updateUser="ExternalLinkService">
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<mods:title id="067FA3E5ECAA7BA7569A42A40D82BCB6">A Second Species of Batesiana Chalumeau, 1983 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:namePart id="22050B937D9BBC4D2E1A9031EF8267FD">Skelley, Paul E.</mods:namePart>
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<mods:namePart id="4D84D4183E3D6D9A7D948C33F5ECA7D4">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="5E22286B07235F7DDBDFC4B7521AB784">text</mods:typeOfResource>
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<mods:title id="DE0420F4D2D4BFA91EEBBDE68B6EBC59">The Coleopterists Bulletin</mods:title>
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<mods:part id="66B05B3CB52FE29602F1DA4502E6832F">
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<mods:date id="60D9DAAB0AC205BB5054DAAC80C6CB5E">2022</mods:date>
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<mods:number id="93658A618D906EF91FB78882C57A96C5">2022-12-20</mods:number>
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<mods:number id="6849043B8014B6D3700FB832E95E0DDD">76</mods:number>
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<mods:start id="E0E3BB696FB145DFC5EF45D2A0BF0A18">612</mods:start>
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<mods:url id="5CA644B42EEC566A638FF40A8B673795">http://dx.doi.org/10.1649/0010-065x-76.4.612</mods:url>
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<mods:identifier id="C6C4300AAC7B06BD870258D3C6DE233E" type="DOI">10.1649/0010-065X-76.4.612</mods:identifier>
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<mods:identifier id="F2C0C64439F14E8BA08B0E56D1DE97CC" type="ISSN">1938-4394</mods:identifier>
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<treatment id="038C6452FFC9C34EA22B48692E906E39" ID-DOI="http://doi.org/10.5281/zenodo.13252919" ID-Zenodo-Dep="13252919" LSID="urn:lsid:plazi:treatment:038C6452FFC9C34EA22B48692E906E39" httpUri="http://treatment.plazi.org/id/038C6452FFC9C34EA22B48692E906E39" lastPageNumber="613" pageId="2" pageNumber="613">
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<subSubSection id="C33F86CFFFC9C34EA22B48692817691F" box="[274,558,1049,1071]" pageId="2" pageNumber="613" type="nomenclature">
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<paragraph id="8B9AD544FFC9C34EA22B48692817691F" blockId="2.[274,558,1049,1071]" box="[274,558,1049,1071]" pageId="2" pageNumber="613">
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<heading id="D0D26228FFC9C34EA22B48692817691F" box="[274,558,1049,1071]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="613" reason="2">
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<taxonomicName id="4C25AEC7FFC9C34EA22B48692817691F" ID-CoL="38HL" authority="Chalumeau, 1983" authorityName="CHALUMEAU" authorityYear="1983" box="[274,558,1049,1071]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA22B48692817691F" bold="true" box="[274,558,1049,1071]" pageId="2" pageNumber="613">
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<emphasis id="B9510956FFC9C34EA22B48692B4C691E" bold="true" box="[274,373,1050,1071]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
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Chalumeau, 1983
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F86CFFFC9C34EA39448362B186945" pageId="2" pageNumber="613" type="reference_group">
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<paragraph id="8B9AD544FFC9C34EA39448362B186945" blockId="2.[173,659,1092,1140]" pageId="2" pageNumber="613">
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<taxonomicName id="4C25AEC7FFC9C34EA3944836281E6968" ID-CoL="38HL" authority="Chalumeau 1983: 143 - 144" authorityName="CHALUMEAU" authorityPageNumber="143 - 144" authorityYear="1983" box="[173,551,1092,1114]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA39448362B34696B" box="[173,269,1093,1114]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
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Chalumeau 1983: 143–144
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</taxonomicName>
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. Stebnicka 2009: 22.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F86CFFFC9C34EA3FC48FC2B9B69F1" pageId="2" pageNumber="613" type="type_taxon">
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<paragraph id="8B9AD544FFC9C34EA3FC48FC2B9B69F1" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
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<emphasis id="B9510956FFC9C34EA3FC48FC2B766994" bold="true" box="[197,335,1167,1189]" pageId="2" pageNumber="613">
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<typeStatus id="549E6BE6FFC9C34EA3FC48FC2ACE6995" box="[197,247,1167,1188]" pageId="2" pageNumber="613">Type</typeStatus>
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Species.
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</emphasis>
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<taxonomicName id="4C25AEC7FFC9C34EA26F48E328AA6995" ID-CoL="3CJ35" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[342,659,1167,1189]" class="Insecta" family="Scarabaeidae" genus="Euparia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="species" species="tuberculata">
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<emphasis id="B9510956FFC9C34EA26F48E328236995" box="[342,538,1167,1189]" italics="true" pageId="2" pageNumber="613">Euparia tuberculata</emphasis>
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Bates, 1887
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</taxonomicName>
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(by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F86CFFFC9C34EA3FC48B42BDA6B09" pageId="2" pageNumber="613" type="diagnosis">
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<paragraph id="8B9AD544FFC9C34EA3FC48B42BDA6B09" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
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<emphasis id="B9510956FFC9C34EA3FC48B42B0969EC" bold="true" box="[197,304,1223,1245]" pageId="2" pageNumber="613">Diagnosis.</emphasis>
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<taxonomicName id="4C25AEC7FFC9C34EA20348B42BA169EC" authorityName="Schmidt" authorityYear="1910" box="[314,408,1223,1245]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="tribe" tribe="Eupariini">Eupariini</taxonomicName>
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, body elongate-oval, robust. Head medially gibbous; ventral clypeal and prothoracic indentations form cavity to hold appendages and prothoracic legs when retracted. Pronotum strongly transverse; lateral margin short, straight, crenate; broadly explanate laterally, lacking fringe of setae; posterior margin lacking marginal line. Elytral humeral angle distinctly dentate, not prolonged anteriorly; intervals irregularly tuberculate laterally, some with minute orifices. Protibia with small teeth in apical quarter. Tarsomeres with dense ventral pads of setae in addition to those on margin. Apical meso- and metatibial spurs short, reduced.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F86CFFFC9C34EA3FC4A4C2B7D6B41" pageId="2" pageNumber="613" type="distribution">
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<paragraph id="8B9AD544FFC9C34EA3FC4A4C2B7D6B41" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
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<emphasis id="B9510956FFC9C34EA3FC4A4C2B706B64" bold="true" box="[197,329,1599,1621]" pageId="2" pageNumber="613">Distribution.</emphasis>
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Central America and northern South America.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F86CFFFC9C34EA3FC4A042E906E39" pageId="2" pageNumber="613" type="discussion">
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<paragraph id="8B9AD544FFC9C34EA3FC4A0429146C9C" blockId="2.[173,660,1167,1730]" lastBlockId="2.[706,1193,194,776]" pageId="2" pageNumber="613">
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<emphasis id="B9510956FFC9C34EA3FC4A042B1D6BBD" bold="true" box="[197,292,1655,1676]" pageId="2" pageNumber="613">Remarks.</emphasis>
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<taxonomicName id="4C25AEC7FFC9C34EA2134A0B2BBC6BBC" authorityName="CHALUMEAU" authorityYear="1983" box="[298,389,1656,1677]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA2134A0B2BBC6BBC" box="[298,389,1656,1677]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C25AEC7FFC9C34EA2A84A042AE66B99" authority="Kozhantshikov, 1916" authorityName="Kozhantshikov" authorityYear="1916" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA2A84A042BC66BBD" box="[401,511,1655,1676]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
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Kozhantshikov, 1916
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</taxonomicName>
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, and
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<taxonomicName id="4C25AEC7FFC9C34EA22A4AE12BCD6B96" authority="Stebnicka, 1999" authorityName="Stebnicka" authorityYear="1999" box="[275,500,1682,1704]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA22A4AE12B6A6B96" box="[275,339,1682,1703]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
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Stebnicka, 1999
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</taxonomicName>
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form a group of genera readily distinguished from all other New World
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<taxonomicName id="4C25AEC7FFC9C34EA03E4CB129596DE9" authorityName="Schmidt" authorityYear="1910" box="[775,864,194,216]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="tribe" tribe="Eupariini">Eupariini</taxonomicName>
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by the basal meso- and metatarsomeres being broadened and distinctly longer than the associated spurs. In addition, the ventral surfaces of these tarsomeres are usually covered with a dense pad of setae (other genera have a marginal fringe of setae only).
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<taxonomicName id="4C25AEC7FFC9C34EA05A4D3B29FC6C6C" authorityName="CHALUMEAU" authorityYear="1983" box="[867,965,328,349]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA05A4D3B29FC6C6C" box="[867,965,328,349]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
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</taxonomicName>
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is distinguished from
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<taxonomicName id="4C25AEC7FFC9C34EA1FA4D1129006C46" authorityName="Kozhantshikov" authorityYear="1916" box="[707,825,354,375]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA1FA4D1129006C46" box="[707,825,354,375]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
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</taxonomicName>
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by the tuberculate elytra and from
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<taxonomicName id="4C25AEC7FFC9C34EA1FA4D0E293C6CA3" authorityName="Stebnicka" authorityYear="1999" box="[707,773,381,402]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA1FA4D0E293C6CA3" box="[707,773,381,402]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
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</taxonomicName>
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by the widely explanate lateral pronotal margin.
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</paragraph>
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<paragraph id="8B9AD544FFC9C34EA1E24DC12E906E39" blockId="2.[706,1193,194,776]" pageId="2" pageNumber="613">
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The genus
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<taxonomicName id="4C25AEC7FFC9C34EA07C4DC1298C6CF6" authorityName="Kozhantshikov" authorityYear="1916" box="[837,949,434,455]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA07C4DC1298C6CF6" box="[837,949,434,455]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
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</taxonomicName>
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has 19 described species and several undescribed species that are morphologically diverse (Stebnicka 2009; P. Skelley, personal observation). Some of these species show intermediate character states among some members of
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<taxonomicName id="4C25AEC7FFC9C34EA1FA4E44290A6F7D" authorityName="Kozhantshikov" authorityYear="1916" box="[707,819,567,588]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA1FA4E44290A6F7D" box="[707,819,567,588]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C25AEC7FFC9C34EA07F4E4B299A6F7C" authorityName="CHALUMEAU" authorityYear="1983" box="[838,931,568,589]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA07F4E4B299A6F7C" box="[838,931,568,589]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C25AEC7FFC9C34EA0DF4E4B2E1D6F7C" authorityName="Stebnicka" authorityYear="1999" box="[998,1060,568,589]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA0DF4E4B2E1D6F7C" box="[998,1060,568,589]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
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</taxonomicName>
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. The genera
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<taxonomicName id="4C25AEC7FFC9C34EA1FA4E2129196F56" authorityName="CHALUMEAU" authorityYear="1983" box="[707,800,594,615]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA1FA4E2129196F56" box="[707,800,594,615]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
|
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</taxonomicName>
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and
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<taxonomicName id="4C25AEC7FFC9C34EA06A4E2129AB6F56" authorityName="Stebnicka" authorityYear="1999" box="[851,914,594,615]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA06A4E2129AB6F56" box="[851,914,594,615]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
|
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</taxonomicName>
|
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have a few autapomorphies, most notably the tuberculate elytra that distinguish them from
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<taxonomicName id="4C25AEC7FFC9C34EA0114EF429A16FAD" authorityName="Kozhantshikov" authorityYear="1916" box="[808,920,647,668]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
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<emphasis id="B9510956FFC9C34EA0114EF429A16FAD" box="[808,920,647,668]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
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</taxonomicName>
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. The validity of these generic limits, along with the potential paraphyly of these genera, needs to be reviewed.However, generic level analysis of this complex is not the purpose of this paper.For now, we maintain the current generic limits.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |