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<document id="5A5FF74235771B30B39C080EA12B87F6" ID-DOI="10.1649/0010-065X-76.4.612" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252933" ID-ZooBank="urn:lsid:zoobank.org:pub:6DA30E51-0FD4-48A5-B7B7-744472F20B9F" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723003944909" checkinUser="felipe" docAuthor="Skelley, Paul E. &amp; Vaz-de-Mello, Fernando Z." docDate="2022" docId="038C6452FFC9C34EA22B48692E906E39" docLanguage="en" docName="TheColeopBull.76.4.612-618.pdf" docOrigin="The Coleopterists Bulletin 76 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-76.4.612" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Batesiana CHALUMEAU 1983" docType="treatment" docVersion="2" lastPageNumber="613" masterDocId="FFB51C2AFFCBC34CA3394C732A396D31" masterDocTitle="A Second Species of Batesiana Chalumeau, 1983 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="618" masterPageNumber="612" pageNumber="613" updateTime="1723006876623" updateUser="ExternalLinkService">
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<mods:title id="067FA3E5ECAA7BA7569A42A40D82BCB6">A Second Species of Batesiana Chalumeau, 1983 (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:namePart id="22050B937D9BBC4D2E1A9031EF8267FD">Skelley, Paul E.</mods:namePart>
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<mods:namePart id="4D84D4183E3D6D9A7D948C33F5ECA7D4">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="DE0420F4D2D4BFA91EEBBDE68B6EBC59">The Coleopterists Bulletin</mods:title>
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<mods:date id="60D9DAAB0AC205BB5054DAAC80C6CB5E">2022</mods:date>
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<mods:identifier id="F2C0C64439F14E8BA08B0E56D1DE97CC" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="C33F86CFFFC9C34EA22B48692817691F" box="[274,558,1049,1071]" pageId="2" pageNumber="613" type="nomenclature">
<paragraph id="8B9AD544FFC9C34EA22B48692817691F" blockId="2.[274,558,1049,1071]" box="[274,558,1049,1071]" pageId="2" pageNumber="613">
<heading id="D0D26228FFC9C34EA22B48692817691F" box="[274,558,1049,1071]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="613" reason="2">
<taxonomicName id="4C25AEC7FFC9C34EA22B48692817691F" ID-CoL="38HL" authority="Chalumeau, 1983" authorityName="CHALUMEAU" authorityYear="1983" box="[274,558,1049,1071]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA22B48692817691F" bold="true" box="[274,558,1049,1071]" pageId="2" pageNumber="613">
<emphasis id="B9510956FFC9C34EA22B48692B4C691E" bold="true" box="[274,373,1050,1071]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
Chalumeau, 1983
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F86CFFFC9C34EA39448362B186945" pageId="2" pageNumber="613" type="reference_group">
<paragraph id="8B9AD544FFC9C34EA39448362B186945" blockId="2.[173,659,1092,1140]" pageId="2" pageNumber="613">
<taxonomicName id="4C25AEC7FFC9C34EA3944836281E6968" ID-CoL="38HL" authority="Chalumeau 1983: 143 - 144" authorityName="CHALUMEAU" authorityPageNumber="143 - 144" authorityYear="1983" box="[173,551,1092,1114]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA39448362B34696B" box="[173,269,1093,1114]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
Chalumeau 1983: 143–144
</taxonomicName>
. Stebnicka 2009: 22.
</paragraph>
</subSubSection>
<subSubSection id="C33F86CFFFC9C34EA3FC48FC2B9B69F1" pageId="2" pageNumber="613" type="type_taxon">
<paragraph id="8B9AD544FFC9C34EA3FC48FC2B9B69F1" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
<emphasis id="B9510956FFC9C34EA3FC48FC2B766994" bold="true" box="[197,335,1167,1189]" pageId="2" pageNumber="613">
<typeStatus id="549E6BE6FFC9C34EA3FC48FC2ACE6995" box="[197,247,1167,1188]" pageId="2" pageNumber="613">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C25AEC7FFC9C34EA26F48E328AA6995" ID-CoL="3CJ35" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[342,659,1167,1189]" class="Insecta" family="Scarabaeidae" genus="Euparia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B9510956FFC9C34EA26F48E328236995" box="[342,538,1167,1189]" italics="true" pageId="2" pageNumber="613">Euparia tuberculata</emphasis>
Bates, 1887
</taxonomicName>
(by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C33F86CFFFC9C34EA3FC48B42BDA6B09" pageId="2" pageNumber="613" type="diagnosis">
<paragraph id="8B9AD544FFC9C34EA3FC48B42BDA6B09" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
<emphasis id="B9510956FFC9C34EA3FC48B42B0969EC" bold="true" box="[197,304,1223,1245]" pageId="2" pageNumber="613">Diagnosis.</emphasis>
<taxonomicName id="4C25AEC7FFC9C34EA20348B42BA169EC" authorityName="Schmidt" authorityYear="1910" box="[314,408,1223,1245]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="tribe" tribe="Eupariini">Eupariini</taxonomicName>
, body elongate-oval, robust. Head medially gibbous; ventral clypeal and prothoracic indentations form cavity to hold appendages and prothoracic legs when retracted. Pronotum strongly transverse; lateral margin short, straight, crenate; broadly explanate laterally, lacking fringe of setae; posterior margin lacking marginal line. Elytral humeral angle distinctly dentate, not prolonged anteriorly; intervals irregularly tuberculate laterally, some with minute orifices. Protibia with small teeth in apical quarter. Tarsomeres with dense ventral pads of setae in addition to those on margin. Apical meso- and metatibial spurs short, reduced.
</paragraph>
</subSubSection>
<subSubSection id="C33F86CFFFC9C34EA3FC4A4C2B7D6B41" pageId="2" pageNumber="613" type="distribution">
<paragraph id="8B9AD544FFC9C34EA3FC4A4C2B7D6B41" blockId="2.[173,660,1167,1730]" pageId="2" pageNumber="613">
<emphasis id="B9510956FFC9C34EA3FC4A4C2B706B64" bold="true" box="[197,329,1599,1621]" pageId="2" pageNumber="613">Distribution.</emphasis>
Central America and northern South America.
</paragraph>
</subSubSection>
<subSubSection id="C33F86CFFFC9C34EA3FC4A042E906E39" pageId="2" pageNumber="613" type="discussion">
<paragraph id="8B9AD544FFC9C34EA3FC4A0429146C9C" blockId="2.[173,660,1167,1730]" lastBlockId="2.[706,1193,194,776]" pageId="2" pageNumber="613">
<emphasis id="B9510956FFC9C34EA3FC4A042B1D6BBD" bold="true" box="[197,292,1655,1676]" pageId="2" pageNumber="613">Remarks.</emphasis>
<taxonomicName id="4C25AEC7FFC9C34EA2134A0B2BBC6BBC" authorityName="CHALUMEAU" authorityYear="1983" box="[298,389,1656,1677]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA2134A0B2BBC6BBC" box="[298,389,1656,1677]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25AEC7FFC9C34EA2A84A042AE66B99" authority="Kozhantshikov, 1916" authorityName="Kozhantshikov" authorityYear="1916" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA2A84A042BC66BBD" box="[401,511,1655,1676]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
Kozhantshikov, 1916
</taxonomicName>
, and
<taxonomicName id="4C25AEC7FFC9C34EA22A4AE12BCD6B96" authority="Stebnicka, 1999" authorityName="Stebnicka" authorityYear="1999" box="[275,500,1682,1704]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA22A4AE12B6A6B96" box="[275,339,1682,1703]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
Stebnicka, 1999
</taxonomicName>
form a group of genera readily distinguished from all other New World
<taxonomicName id="4C25AEC7FFC9C34EA03E4CB129596DE9" authorityName="Schmidt" authorityYear="1910" box="[775,864,194,216]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="tribe" tribe="Eupariini">Eupariini</taxonomicName>
by the basal meso- and metatarsomeres being broadened and distinctly longer than the associated spurs. In addition, the ventral surfaces of these tarsomeres are usually covered with a dense pad of setae (other genera have a marginal fringe of setae only).
<taxonomicName id="4C25AEC7FFC9C34EA05A4D3B29FC6C6C" authorityName="CHALUMEAU" authorityYear="1983" box="[867,965,328,349]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA05A4D3B29FC6C6C" box="[867,965,328,349]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C25AEC7FFC9C34EA1FA4D1129006C46" authorityName="Kozhantshikov" authorityYear="1916" box="[707,825,354,375]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA1FA4D1129006C46" box="[707,825,354,375]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
</taxonomicName>
by the tuberculate elytra and from
<taxonomicName id="4C25AEC7FFC9C34EA1FA4D0E293C6CA3" authorityName="Stebnicka" authorityYear="1999" box="[707,773,381,402]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA1FA4D0E293C6CA3" box="[707,773,381,402]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
</taxonomicName>
by the widely explanate lateral pronotal margin.
</paragraph>
<paragraph id="8B9AD544FFC9C34EA1E24DC12E906E39" blockId="2.[706,1193,194,776]" pageId="2" pageNumber="613">
The genus
<taxonomicName id="4C25AEC7FFC9C34EA07C4DC1298C6CF6" authorityName="Kozhantshikov" authorityYear="1916" box="[837,949,434,455]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA07C4DC1298C6CF6" box="[837,949,434,455]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
</taxonomicName>
has 19 described species and several undescribed species that are morphologically diverse (Stebnicka 2009; P. Skelley, personal observation). Some of these species show intermediate character states among some members of
<taxonomicName id="4C25AEC7FFC9C34EA1FA4E44290A6F7D" authorityName="Kozhantshikov" authorityYear="1916" box="[707,819,567,588]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA1FA4E44290A6F7D" box="[707,819,567,588]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C25AEC7FFC9C34EA07F4E4B299A6F7C" authorityName="CHALUMEAU" authorityYear="1983" box="[838,931,568,589]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA07F4E4B299A6F7C" box="[838,931,568,589]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C25AEC7FFC9C34EA0DF4E4B2E1D6F7C" authorityName="Stebnicka" authorityYear="1999" box="[998,1060,568,589]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA0DF4E4B2E1D6F7C" box="[998,1060,568,589]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
</taxonomicName>
. The genera
<taxonomicName id="4C25AEC7FFC9C34EA1FA4E2129196F56" authorityName="CHALUMEAU" authorityYear="1983" box="[707,800,594,615]" class="Insecta" family="Scarabaeidae" genus="Batesiana" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA1FA4E2129196F56" box="[707,800,594,615]" italics="true" pageId="2" pageNumber="613">Batesiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C25AEC7FFC9C34EA06A4E2129AB6F56" authorityName="Stebnicka" authorityYear="1999" box="[851,914,594,615]" class="Insecta" family="Scarabaeidae" genus="Napoa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA06A4E2129AB6F56" box="[851,914,594,615]" italics="true" pageId="2" pageNumber="613">Napoa</emphasis>
</taxonomicName>
have a few autapomorphies, most notably the tuberculate elytra that distinguish them from
<taxonomicName id="4C25AEC7FFC9C34EA0114EF429A16FAD" authorityName="Kozhantshikov" authorityYear="1916" box="[808,920,647,668]" class="Insecta" family="Scarabaeidae" genus="Odontolytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="613" phylum="Arthropoda" rank="genus">
<emphasis id="B9510956FFC9C34EA0114EF429A16FAD" box="[808,920,647,668]" italics="true" pageId="2" pageNumber="613">Odontolytes</emphasis>
</taxonomicName>
. The validity of these generic limits, along with the potential paraphyly of these genera, needs to be reviewed.However, generic level analysis of this complex is not the purpose of this paper.For now, we maintain the current generic limits.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="74383F08B75B530ECE3C00161738978D" ID-DOI="10.1649/0010-065X-76.4.584" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252893" ID-ZooBank="urn:lsid:zoobank.org:pub:68F3E048-5DDE-4FA0-936F-AF39E750FAB2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723003841931" checkinUser="felipe" docAuthor="Aswathi, S. B., Thomas, Sabu K. &amp; Aswathi, P." docDate="2022" docId="03D387D0FFC8FFDFFF0FA2D0FD45AC7C" docLanguage="en" docName="TheColeopBull.76.4.584-588.pdf" docOrigin="The Coleopterists Bulletin 76 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-76.4.584" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Parathlibops FROM" docType="key" docVersion="2" lastPageNumber="585" masterDocId="FFEAFFA8FFCAFFDDFFCAA221FFD7A80F" masterDocTitle="A New Apterous Species of the Genus Parathlibops Basilewsky (Coleoptera: Carabidae: Scaritinae) from the Western Ghats, a Global Hotspot of Biodiversity in Southwest India" masterLastPageNumber="588" masterPageNumber="584" pageNumber="585" updateTime="1723006804009" updateUser="ExternalLinkService">
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<mods:title id="82BAE89D1899015435BB5B4DC9B8FDF1">A New Apterous Species of the Genus Parathlibops Basilewsky (Coleoptera: Carabidae: Scaritinae) from the Western Ghats, a Global Hotspot of Biodiversity in Southwest India</mods:title>
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<mods:namePart id="7668BF3AB2E5267104D2D447E731C9BC">Aswathi, S. B.</mods:namePart>
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<mods:title id="790E30556AF43092016FEBCF4F40B139">The Coleopterists Bulletin</mods:title>
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<mods:date id="F492BF4D8ADD28D83AF87949E4E982C9">2022</mods:date>
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1. Pronotum longer than wide............................
<emphasis id="B90EEAD4FFC8FFDFFD4DA31FFD44A95C" bold="true" box="[647,659,318,339]" pageId="2" pageNumber="585">2</emphasis>
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1′. Pronotum stout (clearly wider than long) ........ ...................................
<taxonomicName id="4C7A4D45FFC8FFDFFE65A352FD45A987" ID-CoL="8M9ZW" authority="Fedorenko" authorityName="Fedorenko" authorityYear="2016" box="[431,658,371,392]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="bulirschi">
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<emphasis id="B90EEAD4FFC8FFDFFE65A352FDC9A987" bold="true" box="[431,542,371,392]" italics="true" pageId="2" pageNumber="585">P. BuLIRScHI</emphasis>
Fedorenko
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2. Pronotum with coarse punctures; elytral striae crossing the basal declivity............................
<emphasis id="B90EEAD4FFC8FFDFFD4DA389FD44A9B2" bold="true" box="[647,659,424,445]" pageId="2" pageNumber="585">4</emphasis>
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2′. Pronotum without coarse punctures; elytral striae not crossing the basal declivity ........................
<emphasis id="B90EEAD4FFC8FFDFFD4DA3FFFD44A9FC" bold="true" box="[647,659,478,499]" pageId="2" pageNumber="585">3</emphasis>
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3. Pronotal basal fovea lightly impressed ......................................
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<emphasis id="B90EEAD4FFC8FFDFFE0AA032FDC0AA27" bold="true" box="[448,535,531,552]" italics="true" pageId="2" pageNumber="585">P. gLABeR</emphasis>
(Andrewes)
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3′. Pronotal basal fovea deeply impressed..........
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4. Pronotum punctate laterally, basal fovea deep; elytral striae punctate.....................................
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4′. Pronotum punctate near front angles only, basal fovea shallow; elytral striae impunctate on basal three-fourths, punctate on apical fourth .................................
<taxonomicName id="4C7A4D45FFC8FFDFFE69A0EFFD45AAEC" ID-CoL="8MGRT" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[419,658,717,739]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="filiformis">
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<emphasis id="B90EEAD4FFC8FFDFFE69A0EFFDC1AAEC" bold="true" box="[419,534,717,739]" italics="true" pageId="2" pageNumber="585">P. fILIfORmIS</emphasis>
(Andrewes)
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5. Pronotal disc slightly wrinkled, transverse impression impunctate, sulcus faintly impressed, anterior angles distinctly emarginate; submentum asetose..........................
<taxonomicName id="4C7A4D45FFC8FFDFFE2AA118FD44AB41" ID-CoL="8MLMH" authority="Casale" authorityName="Casale" authorityYear="1980" box="[480,659,824,846]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="wittmeri">
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<emphasis id="B90EEAD4FFC8FFDFFE2AA118FD9FAB41" bold="true" box="[480,584,825,846]" italics="true" pageId="2" pageNumber="585">P. wITTmeRI</emphasis>
Casale
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5′. Pronotal disc evidently wrinkled, transverse impression clearly punctate, sulcus deeply impressed, anterior angles slightly emarginate; submentum bisetose................
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<taxonomicName id="4C7A4D45FFC8FFDFFE30A182FD45ABB6" authorityName="Aswathi, Thomas, and Aswathi" authorityYear="2022" box="[506,658,931,953]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="devagiriensis">
<emphasis id="B90EEAD4FFC8FFDFFE30A182FD45ABB6" bold="true" box="[506,658,931,953]" italics="true" pageId="2" pageNumber="585">P. devAgIRIenSIS</emphasis>
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Aswathi, Thomas, and Aswathi,
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6. Odd elytral intervals markedly wider than even intervals, striae obliterated on basal declivity .................................
<taxonomicName id="4C7A4D45FFC8FFDFFE55A62FFD45AC2C" ID-CoL="8MMBC" authority="Fedorenko" authorityName="Fedorenko" authorityYear="2016" box="[415,658,1038,1059]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="nepalensis">
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<emphasis id="B90EEAD4FFC8FFDFFE55A62FFDC9AC2C" bold="true" box="[415,542,1038,1059]" italics="true" pageId="2" pageNumber="585">P. nepALenSIS</emphasis>
Fedorenko
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6′. Even elytral intervals clearly wider than odd elytral intervals, all striae well impressed on basal declivity................
<taxonomicName id="4C7A4D45FFC8FFDFFE79A67FFD45AC7C" ID-CoL="8MCBY" authority="(Gestro)" baseAuthorityName="Gestro" baseAuthorityYear="1883" box="[435,658,1117,1139]" class="Insecta" family="Carabidae" genus="Parathlibops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="puncticollis">
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<emphasis id="B90EEAD4FFC8FFDFFE79A67FFDEEAC7C" bold="true" box="[435,569,1118,1139]" italics="true" pageId="2" pageNumber="585">P. puncTIcOLLIS</emphasis>
(Gestro)
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<mods:title id="2ACCEDCC45BC05AC4F6426486F2E1ACF">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="08E480091374471D1E1D1EA9228598CA">
<mods:date id="30A2A7369AAB9EA2041F27D90EBC5014">2022</mods:date>
<mods:detail id="BE7AA25C5167127662C673FAC596D84B" type="pubDate">
<mods:number id="39D5DF517A6B000A03C1A87C72D46090">2022-12-20</mods:number>
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<mods:number id="599F33C1C20C2F80C14F4FDEA2D302FD">76</mods:number>
</mods:detail>
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<mods:number id="2B2B7D604A8CA4281EBEDF1ACB6191AA">4</mods:number>
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<mods:start id="C9347A60DDD3BE89E65A3B6016E0CE4A">589</mods:start>
<mods:end id="0E5CF84EA6A0817C2F97A231023171F3">594</mods:end>
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<mods:url id="C3F977829CBEB7B19CA3D8ECCF199197">http://dx.doi.org/10.1649/0010-065x-76.4.589</mods:url>
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<mods:identifier id="DDA35F148A3EBD44DA77D79964DB03F1" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="10EDCF9F5AF7243E3DF6DCFEF2A28203" type="Zenodo-Dep">13252899</mods:identifier>
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<subSubSection id="C350655BFFA4FFB9FF2FFF0E6BC5FEE2" pageId="4" pageNumber="592" type="nomenclature">
<paragraph id="8BF536D0FFA4FFB9FF2FFF0E6BC5FEE2" blockId="4.[161,629,192,325]" pageId="4" pageNumber="592">
<heading id="D0BD81BCFFA4FFB9FF2FFF0E6BBEFF52" centered="true" fontSize="9" level="2" pageId="4" pageNumber="592" reason="2">
<emphasis id="B93EEAC2FFA4FFB9FF2FFF0E6BBEFF52" bold="true" pageId="4" pageNumber="592">
<taxonomicName id="4C4A4D53FFA4FFB9FF2FFF0E6808FF70" authority="Caterino and Vasquez-Velez" authorityName="Caterino and VAsQuez-VElez" authorityYear="2022" box="[161,624,192,215]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="videns" status="sp. nov.">
<emphasis id="B93EEAC2FFA4FFB9FF2FFF0E6B3EFF71" bold="true" box="[161,326,193,215]" italics="true" pageId="4" pageNumber="592">Prespelea videns</emphasis>
Caterino and Vásquez-Vélez
</taxonomicName>
,
<taxonomicNameLabel id="A20D57B9FFA4FFB9FEDEFF106BBEFF52" box="[336,454,222,244]" pageId="4" pageNumber="592" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
zoobank.org/
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urn:lsid:zoobank.org:act:
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<figureCitation id="13712A55FFA4FFB9FED6FEE06BC5FEE2" box="[344,445,303,325]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Fig" captionStartId-0="4.[176,215,1692,1711]" captionStartId-1="5.[197,236,606,625]" captionStartId-2="5.[197,236,1686,1705]" captionStartId-3="6.[176,207,554,573]" captionTargetBox-0="[153,1171,807,1675]" captionTargetBox-1="[177,654,193,577]" captionTargetBox-2="[225,1140,1266,1668]" captionTargetBox-3="[152,638,193,536]" captionTargetId-0="figure-245@4.[153,1171,807,1675]" captionTargetId-1="figure-331@5.[177,654,193,589]" captionTargetId-2="figure-306@5.[224,1142,1265,1669]" captionTargetId-3="figure-361@6.[152,638,193,536]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figs. 4–7. Prespelea videns, new species. Photos: 4) Lateral habitus; 5) Dorsal habitus; 6) Aedeagus, dorsal view." captionText-1="Figs. 8–10. Male secondary sexual characters of Prespelea videns, new species. 8) Metaventral process, lateral view; 9) Metaventral process, posterior view; 10) Metacoxa, metatrochanter, and metafemur, ventral view." captionText-2="Figs. 11–12. Abdomen of female Prespelea videns, new species, tergite VII (pygidium) and ventrite VII." captionText-3="Fig. 13. Map of southern Appalachia showing type locality of Prespelea videns, new species." figureDoi-0="http://doi.org/10.5281/zenodo.13252903" figureDoi-1="http://doi.org/10.5281/zenodo.13252905" figureDoi-2="http://doi.org/10.5281/zenodo.13252907" figureDoi-3="http://doi.org/10.5281/zenodo.13252909" httpUri-0="https://zenodo.org/record/13252903/files/figure.png" httpUri-1="https://zenodo.org/record/13252905/files/figure.png" httpUri-2="https://zenodo.org/record/13252907/files/figure.png" httpUri-3="https://zenodo.org/record/13252909/files/figure.png" pageId="4" pageNumber="592">Figs. 4–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C350655BFFA4FFB9FF3EFEAE690AFEE7" pageId="4" pageNumber="592" type="materials_examined">
<paragraph id="8BF536D0FFA4FFB9FF3EFEAE690AFEE7" blockId="4.[152,638,353,749]" lastBlockId="4.[685,1172,193,749]" pageId="4" pageNumber="592">
<emphasis id="B93EEAC2FFA4FFB9FF3EFEAE6B24FED1" bold="true" box="[176,348,353,375]" pageId="4" pageNumber="592">Type Specimens.</emphasis>
<materialsCitation id="3B223C8DFFA4FFB9FEEBFEAE6803FE47" collectionCode="FMNH" collectorName="F. Etzler &amp; A. Haberski &amp; P. Wooden" country="United States of America" county="Macon Co." latitude="35.2348" location="Macon Co." longLatPrecision="6" longitude="-83.5596" pageId="4" pageNumber="592" specimenCode="CUAC000139302" specimenCount="1" typeStatus="holotype">
<emphasis id="B93EEAC2FFA4FFB9FEEBFEAE687BFED0" bold="true" box="[357,515,353,374]" pageId="4" pageNumber="592">
<typeStatus id="54F18872FFA4FFB9FEEBFEAE6BBAFED0" box="[357,450,353,374]" pageId="4" pageNumber="592" type="holotype">Holotype</typeStatus>
male:
</emphasis>
“
<collectingCountry id="F35D7640FFA4FFB9FD9BFEAE6832FED0" box="[533,586,353,374]" name="United States of America" pageId="4" pageNumber="592">USA</collectingCountry>
: NC:
<collectingCounty id="62944E5CFFA4FFB9FF16FEB36B7CFE37" box="[152,260,380,401]" pageId="4" pageNumber="592">Macon Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA4FFB9FE82FEB36B0FFE34" box="[268,375,380,402]" degrees="35.2348" direction="north" orientation="latitude" pageId="4" pageNumber="592" precision="5" value="35.2348">35.2348°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA4FFB9FEF1FEB36B95FE34" box="[383,493,380,402]" degrees="83.5596" direction="west" orientation="longitude" pageId="4" pageNumber="592" precision="5" value="-83.5596">83.5596°W</geoCoordinate>
, Nantahala NF, 5032’, Copper Ridge Bald, ix.15.2020,
<collectorName id="26BF5306FFA4FFB9FDA0FE586802FE0B" box="[558,634,407,429]" pageId="4" pageNumber="592">F.Etzler</collectorName>
,
<collectorName id="26BF5306FFA4FFB9FF16FE7E6B7EFE61" box="[152,262,433,455]" pageId="4" pageNumber="592">A.Haberski</collectorName>
&amp;
<collectorName id="26BF5306FFA4FFB9FEA4FE7E6BF5FE61" box="[298,397,433,455]" pageId="4" pageNumber="592">P.Wooden</collectorName>
, sifted litter”/“Clemson- ENT
<specimenCode id="DBEC9EABFFA4FFB9FF5BFE036BF2FE47" box="[213,394,460,481]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000139302</specimenCode>
”; deposited in
<collectionCode id="ED5BAE15FFA4FFB9FDBCFE036803FE47" box="[562,635,460,481]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="592" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B223C8DFFA4FFB9FF16FE286837FDEA" collectionCode="CUAC" collectorName="F. Etzler &amp; A. Haberski &amp; P. Wooden" country="United States of America" county="Macon Co." latitude="35.2348" location="Macon Co." longLatPrecision="6" longitude="-83.5596" pageId="4" pageNumber="592" specimenCode="CUAC000135311, CUAC000139306" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B93EEAC2FFA4FFB9FF16FE286B48FE5A" bold="true" box="[152,304,486,508]" pageId="4" pageNumber="592">
<typeStatus id="54F18872FFA4FFB9FF16FE286A87FE5A" box="[152,255,487,508]" pageId="4" pageNumber="592" type="paratype">Paratypes</typeStatus>
(5):
</emphasis>
<specimenCount id="9D4CFD59FFA4FFB9FEB1FE286BD3FE5B" box="[319,427,487,509]" count="2" pageId="4" pageNumber="592" type="male">two males</specimenCount>
, same data as type (
<specimenCode id="DBEC9EABFFA4FFB9FF10FDCE6B37FDB1" box="[158,335,513,535]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000135311</specimenCode>
, DNA Extract MSC-6635, High Appalachia morphospecies CrB.B.323; and
<specimenCode id="DBEC9EABFFA4FFB9FF16FDF86B33FDEB" box="[152,331,567,589]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000139306</specimenCode>
), both deposited in
<collectionCode id="ED5BAE15FFA4FFB9FD87FDF86837FDEA" box="[521,591,567,588]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B223C8DFFA4FFB9FDD7FDF8683DFD4A" collectionCode="CUAC, FMNH" collectorName="Nantahala NF &amp; Copper Ridge Bald &amp; F. Etzler &amp; A. Haberski &amp; P. Wooden" country="United States of America" county="Macon Co." latitude="35.2376" location="Macon Co." longLatPrecision="6" longitude="-83.5594" pageId="4" pageNumber="592" specimenCode="CUAC000135293, CUAC000139289" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D4CFD59FFA4FFB9FDD7FDF86A9EFDC1" count="2" pageId="4" pageNumber="592" type="female">two females</specimenCount>
:
<collectingCountry id="F35D7640FFA4FFB9FF72FD9E6B48FDC0" box="[252,304,593,614]" name="United States of America" pageId="4" pageNumber="592">USA</collectingCountry>
: NC:
<collectingCounty id="62944E5CFFA4FFB9FEF1FD9D6B81FDC0" box="[383,505,593,615]" pageId="4" pageNumber="592">Macon Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA4FFB9FD80FD9E6802FDC1" box="[526,634,593,615]" degrees="35.2376" direction="north" orientation="latitude" pageId="4" pageNumber="592" precision="5" value="35.2376">35.2376°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA4FFB9FF16FDA36B7EFD24" box="[152,262,620,642]" degrees="83.5594" direction="west" orientation="longitude" pageId="4" pageNumber="592" precision="5" value="-83.5594">83.5594°W</geoCoordinate>
,
<collectorName id="26BF5306FFA4FFB9FE9DFDA36BE5FD27" box="[275,413,620,642]" pageId="4" pageNumber="592">Nantahala NF</collectorName>
, 5,068′,
<collectorName id="26BF5306FFA4FFB9FE7AFDA36AB0FD3B" pageId="4" pageNumber="592">Copper Ridge Bald</collectorName>
, ix.15.2020,
<collectorName id="26BF5306FFA4FFB9FEDEFD486BD1FD3B" box="[336,425,647,669]" pageId="4" pageNumber="592">F. Etzler</collectorName>
,
<collectorName id="26BF5306FFA4FFB9FE39FD48684AFD3B" box="[439,562,647,669]" pageId="4" pageNumber="592">A. Haberski</collectorName>
and
<collectorName id="26BF5306FFA4FFB9FDE3FD486A92FD11" pageId="4" pageNumber="592">P. Wooden</collectorName>
, sifted litter (
<specimenCode id="DBEC9EABFFA4FFB9FE0AFD6E684FFD11" box="[388,567,673,695]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000135293</specimenCode>
, DNA Extract MSC-6617, deposited in
<collectionCode id="ED5BAE15FFA4FFB9FD8EFD73683EFD77" box="[512,582,700,721]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC</collectionCode>
; and
<specimenCode id="DBEC9EABFFA4FFB9FF16FD186B31FD4A" box="[152,329,727,748]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000139289</specimenCode>
, deposited in
<collectionCode id="ED5BAE15FFA4FFB9FE7AFD186846FD4A" box="[500,574,727,748]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="592" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B223C8DFFA4FFB9FDD5FD186916FEE7" collectionCode="CUAC" collectorName="Nantahala NF &amp; Copper Ridge Bald &amp; M. Caterino &amp; M. Ferro" country="United States of America" county="Macon Co." latitude="35.2357" location="Macon Co." longLatPrecision="6" longitude="-83.5602" pageId="4" pageNumber="592" specimenCode="CUAC000003213" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4CFD59FFA4FFB9FDD5FD18688AFF7E" box="[603,754,194,748]" count="1" pageId="4" pageNumber="592" type="female">one female</specimenCount>
:
<collectingCountry id="F35D7640FFA4FFB9FC84FF0D6946FF71" box="[778,830,194,215]" name="United States of America" pageId="4" pageNumber="592">USA</collectingCountry>
: NC:
<collectingCounty id="62944E5CFFA4FFB9FC1FFF0D6E74FF71" box="[913,1036,194,215]" pageId="4" pageNumber="592">Macon Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA4FFB9FBAAFF0E6EF7FF71" box="[1060,1167,193,215]" degrees="35.2357" direction="north" orientation="latitude" pageId="4" pageNumber="592" precision="5" value="35.2357">35.2357°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA4FFB9FD23FF136963FF54" box="[685,795,220,242]" degrees="83.5602" direction="west" orientation="longitude" pageId="4" pageNumber="592" precision="5" value="-83.5602">83.5602°W</geoCoordinate>
,
<collectorName id="26BF5306FFA4FFB9FCA7FF1369CAFF54" box="[809,946,220,242]" pageId="4" pageNumber="592">Nantahala NF</collectorName>
, 5,144′,
<collectorName id="26BF5306FFA4FFB9FB87FF1368A5FEAB" pageId="4" pageNumber="592">Copper Ridge Bald</collectorName>
, vii.9.2019, sifted litter,
<collectorName id="26BF5306FFA4FFB9FC41FF386E3DFEAB" box="[975,1093,247,269]" pageId="4" pageNumber="592">M. Caterino</collectorName>
and
<collectorName id="26BF5306FFA4FFB9FBF9FF386898FE81" pageId="4" pageNumber="592">M. Ferro</collectorName>
(
<specimenCode id="DBEC9EABFFA4FFB9FD65FEDD69E3FE81" box="[747,923,274,295]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC000003213</specimenCode>
, DNA Extract MSC-3427) deposited in
<collectionCode id="ED5BAE15FFA4FFB9FCA6FEE36916FEE7" box="[808,878,300,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="4" pageNumber="592" type="University or college">CUAC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C350655BFFA4FFB8FD4BFE886E5FFE07" lastPageId="5" lastPageNumber="593" pageId="4" pageNumber="592" type="diagnosis">
<paragraph id="8BF536D0FFA4FFB9FD4BFE886EEAFD4A" blockId="4.[685,1172,193,749]" pageId="4" pageNumber="592">
<emphasis id="B93EEAC2FFA4FFB9FD4BFE88695FFEFB" bold="true" box="[709,807,327,349]" pageId="4" pageNumber="592">Diagnosis.</emphasis>
HL:
<quantity id="4CB29B35FFA4FFB9FCDCFE8869AEFEFA" box="[850,982,327,348]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="4" pageNumber="592" unit="mm" value="0.35" valueMax="0.37" valueMin="0.33">0.33–0.37 mm</quantity>
;PnL:
<quantity id="4CB29B35FFA4FFB9FB82FE886EE8FEFA" box="[1036,1168,327,348]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="4" pageNumber="592" unit="mm" value="0.35" valueMax="0.37" valueMin="0.33">0.33–0.37 mm</quantity>
; PnW:
<quantity id="4CB29B35FFA4FFB9FD61FEAE69FBFED1" box="[751,899,353,375]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="4" pageNumber="592" unit="mm" value="0.32999999999999996" valueMax="0.35" valueMin="0.31">0.31–0.35 mm</quantity>
; EL:
<quantity id="4CB29B35FFA4FFB9FC4DFEAE6E2FFED1" box="[963,1111,353,375]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.699999999999999" metricValueMin="4.5" pageId="4" pageNumber="592" unit="mm" value="0.51" valueMax="0.57" valueMin="0.45">0.45–0.57 mm</quantity>
; EW:
<quantity id="4CB29B35FFA4FFB9FD23FEB36941FE34" box="[685,825,380,402]" metricMagnitude="-4" metricUnit="m" metricValue="6.85" metricValueMax="7.4" metricValueMin="6.3" pageId="4" pageNumber="592" unit="mm" value="0.685" valueMax="0.74" valueMin="0.63">0.63–0.74 mm</quantity>
; T3L:
<quantity id="4CB29B35FFA4FFB9FCFBFEB36E79FE34" box="[885,1025,380,402]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="4" pageNumber="592" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.45–0.47 mm</quantity>
; AL:
<quantity id="4CB29B35FFA4FFB9FBBDFEB368ADFE0A" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="8.6" metricValueMin="5.8999999999999995" pageId="4" pageNumber="592" unit="mm" value="0.725" valueMax="0.86" valueMin="0.59">0.59–0.86 mm</quantity>
; TL:
<quantity id="4CB29B35FFA4FFB9FC8BFE5869E8FE0A" box="[773,912,407,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.755" metricValueMax="2.14" metricValueMin="1.37" pageId="4" pageNumber="592" unit="mm" value="1.755" valueMax="2.14" valueMin="1.37">1.37–2.14 mm</quantity>
. Distinguishable from other
<taxonomicName id="4C4A4D53FFA4FFB9FD23FE7D692EFE6E" box="[685,854,434,456]" pageId="4" pageNumber="592">
<emphasis id="B93EEAC2FFA4FFB9FD23FE7D6974FE61" box="[685,780,434,455]" italics="true" pageId="4" pageNumber="592">Prespelea</emphasis>
species
</taxonomicName>
only by the following characters: both sexes with strongly narrowed elytral humeri, presumably flightless, male metaventrite strongly projecting anteriad, apically truncate and slightly constricted medially (
<figureCitation id="13712A55FFA4FFB9FC0AFDD3699FFD97" box="[900,999,540,562]" captionStart="Figs" captionStartId="5.[197,236,606,625]" captionTargetBox="[177,654,193,577]" captionTargetId="figure-331@5.[177,654,193,589]" captionTargetPageId="5" captionText="Figs. 8–10. Male secondary sexual characters of Prespelea videns, new species. 8) Metaventral process, lateral view; 9) Metaventral process, posterior view; 10) Metacoxa, metatrochanter, and metafemur, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13252905" httpUri="https://zenodo.org/record/13252905/files/figure.png" pageId="4" pageNumber="592">Figs. 8, 9</figureCitation>
); male eyes with 6–10 ommatidia (
<figureCitation id="13712A55FFA4FFB9FCDCFDF869F2FDEA" box="[850,906,567,589]" captionStart="Figs" captionStartId="4.[176,215,1692,1711]" captionTargetBox="[153,1171,807,1675]" captionTargetId="figure-245@4.[153,1171,807,1675]" captionTargetPageId="4" captionText="Figs. 4–7. Prespelea videns, new species. Photos: 4) Lateral habitus; 5) Dorsal habitus; 6) Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13252903" httpUri="https://zenodo.org/record/13252903/files/figure.png" pageId="4" pageNumber="592">Fig. 4</figureCitation>
); male metatrochanters with distal tooth, acutely projecting anteriorly, basally wide (
<figureCitation id="13712A55FFA4FFB9FD68FDA36953FD27" box="[742,811,620,642]" captionStart="Figs" captionStartId="5.[197,236,606,625]" captionTargetBox="[177,654,193,577]" captionTargetId="figure-331@5.[177,654,193,589]" captionTargetPageId="5" captionText="Figs. 8–10. Male secondary sexual characters of Prespelea videns, new species. 8) Metaventral process, lateral view; 9) Metaventral process, posterior view; 10) Metacoxa, metatrochanter, and metafemur, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13252905" httpUri="https://zenodo.org/record/13252905/files/figure.png" pageId="4" pageNumber="592">Fig. 10</figureCitation>
); tergite 6 with lateral marginal carinae incomplete; female tergite VII (pygidium) shallowly depressed on either side of weak median carina (
<figureCitation id="13712A55FFA4FFB9FD3AFD736957FD77" box="[692,815,700,722]" captionStart="Figs" captionStartId="5.[197,236,1686,1705]" captionTargetBox="[225,1140,1266,1668]" captionTargetId="figure-306@5.[224,1142,1265,1669]" captionTargetPageId="5" captionText="Figs. 11–12. Abdomen of female Prespelea videns, new species, tergite VII (pygidium) and ventrite VII." figureDoi="http://doi.org/10.5281/zenodo.13252907" httpUri="https://zenodo.org/record/13252907/files/figure.png" pageId="4" pageNumber="592">Figs. 11, 12</figureCitation>
); apical margin of sternite VII produced, rugosely sculptured, distinctly dentate at
</paragraph>
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<paragraph id="8BF536D0FFA4FFB9FF3EF9536EECF909" blockId="4.[152,1172,1692,1735]" box="[176,1172,1692,1711]" pageId="4" pageNumber="592">
<emphasis id="B93EEAC2FFA4FFB9FF3EF9536B7EF909" bold="true" box="[176,262,1692,1711]" pageId="4" pageNumber="592">Figs. 4–7.</emphasis>
<taxonomicName id="4C4A4D53FFA4FFB9FE92F9536BD5F909" authorityName="Caterino &amp; Vásquez-Vélez" authorityYear="2022" box="[284,429,1692,1711]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="videns" status="sp. nov.">
<emphasis id="B93EEAC2FFA4FFB9FE92F9536BD5F909" box="[284,429,1692,1711]" italics="true" pageId="4" pageNumber="592">Prespelea videns</emphasis>
</taxonomicName>
,
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<taxonomicNameLabel id="A20D57B9FFA4FFB9FE36F9536859F909" box="[440,545,1692,1711]" pageId="4" pageNumber="592" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Photos:
<emphasis id="B93EEAC2FFA4FFB9FDFCF95368FCF909" bold="true" box="[626,644,1692,1711]" pageId="4" pageNumber="592">4)</emphasis>
Lateral habitus;
<emphasis id="B93EEAC2FFA4FFB9FC98F9536950F909" bold="true" box="[790,808,1692,1711]" pageId="4" pageNumber="592">5)</emphasis>
Dorsal habitus;
<emphasis id="B93EEAC2FFA4FFB9FC3BF95369B0F909" bold="true" box="[949,968,1692,1711]" pageId="4" pageNumber="592">6)</emphasis>
Aedeagus, dorsal view.
</paragraph>
</caption>
<paragraph id="8BF536D0FFA4FFB9FF16F97B6B93F961" blockId="4.[152,1172,1692,1735]" box="[152,491,1716,1735]" pageId="4" pageNumber="592">
<emphasis id="B93EEAC2FFA4FFB9FF16F97B6AD1F961" bold="true" box="[152,169,1716,1735]" pageId="4" pageNumber="592">7)</emphasis>
Aedeagus, dorsal view, line drawing.
</paragraph>
<paragraph id="8BF536D0FFA5FFB8FD4DFF0D6E57FEFB" blockId="5.[707,1192,194,349]" pageId="5" pageNumber="593">eyes, prominent metaventral process, and strongly narrowed elytral humeri. The female pygidium/terminal sternite complex is also Quite distinctive, with the rugose upturned borders and medial tooth of sternite VII, and the median ridge and lateral depressions of tergite VII (pygidium).</paragraph>
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<emphasis id="B93EEAC2FFA5FFB8FC1FFE4469BAFE06" bold="true" box="[913,962,395,419]" pageId="5" pageNumber="593">NEW</emphasis>
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<emphasis id="B93EEAC2FFA5FFB8FC49FE446E5FFE07" bold="true" box="[967,1063,395,419]" pageId="5" pageNumber="593">RECORDS</emphasis>
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</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AB3B1861AFA086990A8EB0BF995B3BC1">A New Species of the Southern Appalachian Genus Prespelea Park (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<paragraph id="8BF536D0FFA5FFB8FD4DFE766ED8FE69" blockId="5.[707,1193,441,570]" box="[707,1184,441,463]" pageId="5" pageNumber="593">
<heading id="D0BD81BCFFA5FFB8FD4DFE766ED8FE69" box="[707,1184,441,463]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="593" reason="2">
<taxonomicName id="4C4A4D53FFA5FFB8FD4DFE766ED8FE69" ID-CoL="4MBDB" authority="Caterino and Vasquez-Velez" authorityName="Caterino and Vasquez-Velez" box="[707,1184,441,463]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B93EEAC2FFA5FFB8FD4DFE766ED8FE69" bold="true" box="[707,1184,441,463]" pageId="5" pageNumber="593">
<emphasis id="B93EEAC2FFA5FFB8FD4DFE76690EFE69" bold="true" box="[707,886,441,463]" italics="true" pageId="5" pageNumber="593">Prespelea carltoni</emphasis>
Caterino and Vásquez-Vélez
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF536D0FFA5FFB8FD55FE1A6EDBFD9C" blockId="5.[707,1193,441,570]" pageId="5" pageNumber="593">
<materialsCitation id="3B223C8DFFA5FFB8FD55FE1A6EE7FD9C" collectionCode="CUAC" collectorName="Nantahala NF &amp; Teyahalee Bald &amp; M. Caterino &amp; M. Ferro" county="Graham Co." latitude="35.2585" location="Graham Co." longLatPrecision="6" longitude="-83.7959" pageId="5" pageNumber="593" specimenCode="CUAC000149743, CUAC000149726" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D4CFD59FFA5FFB8FD55FE1A6929FE4C" box="[731,849,468,490]" count="2" pageId="5" pageNumber="593" type="male">Two males</specimenCount>
(MSC-7472,
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;
<specimenCode id="DBEC9EABFFA5FFB8FD4DFE20690FFDA3" box="[707,887,495,517]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="593" type="University or college">CUAC000149726</specimenCode>
): NC:
<collectingCounty id="62944E5CFFA5FFB8FC38FE206E57FDA2" box="[950,1071,495,517]" pageId="5" pageNumber="593">Graham Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA5FFB8FBB7FE206EDCFDA3" box="[1081,1188,495,517]" degrees="35.2585" direction="north" orientation="latitude" pageId="5" pageNumber="593" precision="5" value="35.2585">35.2585°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA5FFB8FD4DFDC66949FDB9" box="[707,817,521,543]" degrees="83.7959" direction="west" orientation="longitude" pageId="5" pageNumber="593" precision="5" value="-83.7959">83.7959°W</geoCoordinate>
,
<collectorName id="26BF5306FFA5FFB8FCB4FDC569C7FDB9" box="[826,959,522,544]" pageId="5" pageNumber="593">Nantahala NF</collectorName>
, 5,241′,
<collectorName id="26BF5306FFA5FFB8FB81FDC56EDDFD86" box="[1039,1189,522,544]" pageId="5" pageNumber="593">Teyahalee Bald</collectorName>
, vii.9.2019, sifted litter,
<collectorName id="26BF5306FFA5FFB8FC2AFDEA6E60FD9C" box="[932,1048,548,570]" pageId="5" pageNumber="593">M. Caterino</collectorName>
and
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.
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<paragraph id="8BF536D0FFA5FFB8FF4BFD9168E8FD1F" blockId="5.[173,660,606,698]" pageId="5" pageNumber="593">
<emphasis id="B93EEAC2FFA5FFB8FF4BFD916B55FDD7" bold="true" box="[197,301,606,625]" pageId="5" pageNumber="593">Figs. 8–10.</emphasis>
Male secondary sexual characters of
<taxonomicName id="4C4A4D53FFA5FFB8FF23FDB96B39FD2F" authorityName="Caterino &amp; Vásquez-Vélez" authorityYear="2022" box="[173,321,630,649]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="videns" status="sp. nov.">
<emphasis id="B93EEAC2FFA5FFB8FF23FDB96B39FD2F" box="[173,321,630,649]" italics="true" pageId="5" pageNumber="593">Prespelea videns</emphasis>
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,
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.
<emphasis id="B93EEAC2FFA5FFB8FE4BFDB96BAFFD2F" bold="true" box="[453,471,630,649]" pageId="5" pageNumber="593">8)</emphasis>
Metaventral process, lateral view;
<emphasis id="B93EEAC2FFA5FFB8FE90FD416B48FD07" bold="true" box="[286,304,654,673]" pageId="5" pageNumber="593">9)</emphasis>
Metaventral process, posterior view;
<emphasis id="B93EEAC2FFA5FFB8FDF8FD4168EAFD07" bold="true" box="[630,658,654,673]" pageId="5" pageNumber="593">10)</emphasis>
Metacoxa, metatrochanter, and metafemur, ventral view.
</paragraph>
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<subSubSection id="C350655BFFA5FFB8FF23FD256879FC20" pageId="5" pageNumber="593" type="description">
<paragraph id="8BF536D0FFA5FFB8FF23FD256879FC20" blockId="5.[173,659,746,1195]" pageId="5" pageNumber="593">
middle; aedeagus (
<figureCitation id="13712A55FFA5FFB8FEE4FD256BB5FCA6" box="[362,461,746,768]" captionStart="Figs" captionStartId="4.[176,215,1692,1711]" captionTargetBox="[153,1171,807,1675]" captionTargetId="figure-245@4.[153,1171,807,1675]" captionTargetPageId="4" captionText="Figs. 4–7. Prespelea videns, new species. Photos: 4) Lateral habitus; 5) Dorsal habitus; 6) Aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13252903" httpUri="https://zenodo.org/record/13252903/files/figure.png" pageId="5" pageNumber="593">Figs. 6, 7</figureCitation>
) symmetrical, apex emarginate, apicolateral ridges convergent subapically, internal sac with medial sclerotized process, apically bifurcate; parameres with membranous apical lobes, with several long, subapical inwardlypointing setae along lateral margin.
</paragraph>
</subSubSection>
<subSubSection id="C350655BFFA5FFB8FF4BFC456B3CFBD3" pageId="5" pageNumber="593" type="distribution">
<paragraph id="8BF536D0FFA5FFB8FF4BFC456B3CFBD3" blockId="5.[173,659,746,1195]" pageId="5" pageNumber="593">
<emphasis id="B93EEAC2FFA5FFB8FF4BFC456B31FC06" bold="true" box="[197,329,906,928]" pageId="5" pageNumber="593">Distribution.</emphasis>
This new species is known only from Copper Ridge Bald in the Nantahala National Forest of western
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. Despite the name, the
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locality is rather densely forested with deciduous trees,
<taxonomicName id="4C4A4D53FFA5FFB8FEDFFC3A6BD9FBAC" box="[337,417,1013,1034]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAC2FFA5FFB8FEDFFC3A6BD9FBAC" box="[337,417,1013,1034]" italics="true" pageId="5" pageNumber="593">Quercus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4A4D53FFA5FFB8FE34FC3A686EFBAC" authorityName="Dumortier" authorityYear="1829" box="[442,534,1013,1034]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
) and
<taxonomicName id="4C4A4D53FFA5FFB8FDEBFC3A68EAFBAC" box="[613,658,1013,1034]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAC2FFA5FFB8FDEBFC3A68EAFBAC" box="[613,658,1013,1034]" italics="true" pageId="5" pageNumber="593">Acer</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4A4D53FFA5FFB8FF3AFBDF6B49FB80" box="[180,305,1040,1062]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
), with a shrubby understory of
<taxonomicName id="4C4A4D53FFA5FFB8FF23FBE56B43FB99" box="[173,315,1066,1087]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAC2FFA5FFB8FF23FBE56B43FB99" box="[173,315,1066,1087]" italics="true" pageId="5" pageNumber="593">Rhododendron</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4A4D53FFA5FFB8FEC3FBE56BC8FBE6" box="[333,432,1066,1088]" class="Magnoliopsida" family="Ericaceae" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="593" phylum="Tracheophyta" rank="family">Ericaceae</taxonomicName>
). The specimens were sifted from mixed litter around the edges of small rocky outcrops.
</paragraph>
</subSubSection>
<subSubSection id="C350655BFFA5FFB8FF4BFBB468EBFB0D" pageId="5" pageNumber="593" type="discussion">
<paragraph id="8BF536D0FFA5FFB8FF4BFBB468EBFB0D" blockId="5.[173,659,746,1195]" pageId="5" pageNumber="593">
<emphasis id="B93EEAC2FFA5FFB8FF4BFBB46B50FB36" bold="true" box="[197,296,1147,1168]" pageId="5" pageNumber="593">Remarks.</emphasis>
Males of this new species are highly distinctive in the combination of intermediate-sized
</paragraph>
</subSubSection>
</treatment>
</document>

162
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@ -0,0 +1,162 @@
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<mods:title id="5E6FDABBF0332D1D6539C9B5AE0EF6F3">A New Species of the Southern Appalachian Genus Prespelea Park (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<paragraph id="8BF536D0FFA5FFB8FD4DFB936EDCFBD7" blockId="5.[707,1192,1115,1192]" box="[707,1188,1115,1138]" pageId="5" pageNumber="593">
<heading id="D0BD81BCFFA5FFB8FD4DFB936EDCFBD7" box="[707,1188,1115,1138]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="593" reason="2">
<taxonomicName id="4C4A4D53FFA5FFB8FD4DFB936EDCFBD7" ID-CoL="4MBDJ" authority="Caterino and Vasquez-Velez" authorityName="Caterino and Vasquez-Velez" box="[707,1188,1115,1138]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="myersae">
<emphasis id="B93EEAC2FFA5FFB8FD4DFB936EDCFBD7" bold="true" box="[707,1188,1115,1138]" pageId="5" pageNumber="593">
<emphasis id="B93EEAC2FFA5FFB8FD4DFB936902FBD4" bold="true" box="[707,890,1116,1138]" italics="true" pageId="5" pageNumber="593">Prespelea myersae</emphasis>
Caterino and Vásquez-Vélez
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF536D0FFA5FFB8FD55FBB86ED0FB01" blockId="5.[707,1192,1115,1192]" pageId="5" pageNumber="593">
<materialsCitation id="3B223C8DFFA5FFB8FD55FBB86E02FB2B" box="[731,1146,1143,1165]" collectionCode="CUAC" pageId="5" pageNumber="593" specimenCode="CUAC000146101" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4CFD59FFA5FFB8FD55FBB86941FB2B" box="[731,825,1143,1165]" count="1" pageId="5" pageNumber="593" type="male">One male</specimenCount>
(MSC-7473,
<specimenCode id="DBEC9EABFFA5FFB8FC4FFBB86E0CFB2B" box="[961,1140,1143,1165]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="593" type="University or college">CUAC000146101</specimenCode>
)
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,
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<specimenCount id="9D4CFD59FFA5FFB8FB0BFBB86970FB0E" count="1" pageId="5" pageNumber="593" type="female">one female</specimenCount>
: GA:
<collectingCounty id="62944E5CFFA5FFB8FCC8FB5D69CAFB01" box="[838,946,1170,1192]" pageId="5" pageNumber="593">Rabun Co.</collectingCounty>
,
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,
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,
</paragraph>
</subSubSection>
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<emphasis id="B93EEAC2FFA5FFB8FF4BF9596B57F90F" bold="true" box="[197,303,1686,1705]" pageId="5" pageNumber="593">Figs. 11–12.</emphasis>
Abdomen of female
<taxonomicName id="4C4A4D53FFA5FFB8FE7BF95968FDF90F" authorityName="Caterino &amp; Vásquez-Vélez" authorityYear="2022" box="[501,645,1686,1705]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="videns" status="sp. nov.">
<emphasis id="B93EEAC2FFA5FFB8FE7BF95968FDF90F" box="[501,645,1686,1705]" italics="true" pageId="5" pageNumber="593">Prespelea videns</emphasis>
</taxonomicName>
,
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<taxonomicNameLabel id="A20D57B9FFA5FFB8FD1EF9586881F90F" box="[656,761,1686,1706]" pageId="5" pageNumber="593" rank="species">new species</taxonomicNameLabel>
</emphasis>
, tergite VII (pygidium) and ventrite VII.
</paragraph>
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<emphasis id="B93EEAC2FFA5FFB8FF23F9616AB0F967" bold="true" box="[173,200,1710,1729]" pageId="5" pageNumber="593">11)</emphasis>
Posterior view;
<emphasis id="B93EEAC2FFA5FFB8FEDBF9616B09F967" bold="true" box="[341,369,1710,1729]" pageId="5" pageNumber="593">12)</emphasis>
Lateral view.
</paragraph>
<paragraph id="8BF536D0FFA6FFBBFD23FF0E6985FF52" blockId="6.[685,1172,193,458]" pageId="6" pageNumber="594">
<emphasis id="B93EEAC2FFA6FFBBFD23FF0E6985FF52" bold="true" pageId="6" pageNumber="594">Females from distinct localities, which could potentially represent new species</emphasis>
</paragraph>
<paragraph id="8BF536D0FFA6FFBBFD4BFF366955FEF9" blockId="6.[685,1172,193,458]" pageId="6" pageNumber="594">
<materialsCitation id="3B223C8DFFA6FFBBFD4BFF366952FEF9" collectionCode="CUAC" collectorName="Pisgah NF &amp; Mountains-to-Sea Tr. &amp; M. Caterino" county="Haywood Co." latitude="35.3074" location="Haywood Co." longLatPrecision="6" longitude="-82.912" pageId="6" pageNumber="594" specimenCode="CUAC000143455" specimenCount="1">
One (MSC-7476,
<specimenCode id="DBEC9EABFFA6FFBBFCEFFF366E7FFEA9" box="[865,1031,249,271]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000143455</specimenCode>
):NC:
<collectingCounty id="62944E5CFFA6FFBBFBB2FF3668A9FE8F" pageId="6" pageNumber="594">Haywood Co.</collectingCounty>
,
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,
<geoCoordinate id="EE7E5017FFA6FFBBFCD4FEDB69BFFE8F" box="[858,967,276,297]" degrees="82.9120" direction="west" orientation="longitude" pageId="6" pageNumber="594" precision="5" value="-82.912">82.9120°W</geoCoordinate>
,
<collectorName id="26BF5306FFA6FFBBFC59FEDB6E3AFE8F" box="[983,1090,276,298]" pageId="6" pageNumber="594">Pisgah NF</collectorName>
, 5,150′,
<collectorName id="26BF5306FFA6FFBBFD23FEE0690FFEE2" box="[685,887,303,325]" pageId="6" pageNumber="594">Mountains-to-Sea Tr.</collectorName>
, ix.8.2020,
<collectorName id="26BF5306FFA6FFBBFC6DFEE06E2EFEE3" box="[995,1110,303,325]" pageId="6" pageNumber="594">M. Caterino</collectorName>
, sifted litter nr. seep
</materialsCitation>
.
</paragraph>
<paragraph id="8BF536D0FFA6FFBBFD4BFEAB6964FE6C" blockId="6.[685,1172,193,458]" pageId="6" pageNumber="594">
<materialsCitation id="3B223C8DFFA6FFBBFD4BFEAB6960FE6C" collectionCode="CUAC" collectorName="Chattahoochee NF &amp; Brasstown Bald &amp; M. Caterino" county="Towns Co." latitude="34.8766" location="Towns Co." longLatPrecision="6" longitude="-83.8109" pageId="6" pageNumber="594" specimenCode="CUAC000137663" specimenCount="1">
One (MSC-5561,
<specimenCode id="DBEC9EABFFA6FFBBFCE5FEAB6E63FEDC" box="[875,1051,356,378]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000137663</specimenCode>
): GA:
<collectingCounty id="62944E5CFFA6FFBBFBDAFEAB68AAFE32" pageId="6" pageNumber="594">Towns Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA6FFBBFD6EFEB06933FE33" box="[736,843,383,405]" degrees="34.8766" direction="north" orientation="latitude" pageId="6" pageNumber="594" precision="5" value="34.8766">34.8766°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA6FFBBFCD4FEB069B0FE32" box="[858,968,383,404]" degrees="83.8109" direction="west" orientation="longitude" pageId="6" pageNumber="594" precision="5" value="-83.8109">83.8109°W</geoCoordinate>
,
<collectorName id="26BF5306FFA6FFBBFC58FEB06EF7FE32" box="[982,1167,383,405]" pageId="6" pageNumber="594">Chattahoochee NF</collectorName>
,
<collectorName id="26BF5306FFA6FFBBFD23FE556934FE09" box="[685,844,409,431]" pageId="6" pageNumber="594">Brasstown Bald</collectorName>
, 4,519′, vii.2.2020,
<collectorName id="26BF5306FFA6FFBBFB9BFE556EF7FE09" box="[1045,1167,409,431]" pageId="6" pageNumber="594">M. Caterino</collectorName>
, sifted litter
</materialsCitation>
.
</paragraph>
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<paragraph id="8BF536D0FFA6FFBBFF3EFDE56B85FDF3" blockId="6.[152,627,554,597]" pageId="6" pageNumber="594">
<emphasis id="B93EEAC2FFA6FFBBFF3EFDE56A8BFD9B" bold="true" box="[176,243,554,573]" pageId="6" pageNumber="594">Fig. 13.</emphasis>
Map of southern Appalachia showing type locality of
<taxonomicName id="4C4A4D53FFA6FFBBFF7AFD8D6BFCFDF3" authorityName="Caterino &amp; Vásquez-Vélez" authorityYear="2022" box="[244,388,578,597]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="594" phylum="Arthropoda" rank="species" species="videns" status="sp. nov.">
<emphasis id="B93EEAC2FFA6FFBBFF7AFD8D6BFCFDF3" box="[244,388,578,597]" italics="true" pageId="6" pageNumber="594">Prespelea videns</emphasis>
</taxonomicName>
,
<emphasis id="B93EEAC2FFA6FFBBFE01FD8D6B80FDF3" bold="true" box="[399,504,578,597]" pageId="6" pageNumber="594">
<taxonomicNameLabel id="A20D57B9FFA6FFBBFE01FD8D6B80FDF3" box="[399,504,578,597]" pageId="6" pageNumber="594" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BF536D0FFA6FFBBFF16FD6C6850FD72" blockId="6.[152,638,674,724]" pageId="6" pageNumber="594">Chattahoochee NF, Rabun Bald, 4,663′, v.11.2021, sifted litter, M. Caterino and A. Haberski.</paragraph>
</subSubSection>
</treatment>
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176
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<document id="7770790BE40F0AAFAF36B39E0B5A14B9" ID-DOI="10.1649/0010-065X-76.4.589" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252899" ID-ZooBank="urn:lsid:zoobank.org:pub:F5349B29-BF57-4A24-841B-8B0455859504" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723003869626" checkinUser="felipe" docAuthor="Caterino, Michael S. &amp; Vásquez-Vélez, Laura M." docDate="2022" docId="03E387C6FFA6FFBBFF16FD396B00FA82" docLanguage="en" docName="TheColeopBull.76.4.589-594.pdf" docOrigin="The Coleopterists Bulletin 76 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-76.4.589" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Prespelea morsei Caterino and Vasquez-Velez" docType="treatment" docVersion="2" lastPageNumber="594" masterDocId="FFDAFFBEFFA0FFBDFF8EFFCF6A78FFA6" masterDocTitle="A New Species of the Southern Appalachian Genus Prespelea Park (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="594" masterPageNumber="589" pageNumber="594" updateTime="1723006817371" updateUser="ExternalLinkService">
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<mods:title id="F250CEC747E460017204A659DF6D918C">A New Species of the Southern Appalachian Genus Prespelea Park (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<taxonomicName id="4C4A4D53FFA6FFBBFF16FD396813FCAD" ID-CoL="4MBDH" authority="Caterino and Vasquez-Velez" authorityName="Caterino and Vasquez-Velez" box="[152,619,757,780]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="594" phylum="Arthropoda" rank="species" species="morsei">
<emphasis id="B93EEAC2FFA6FFBBFF16FD396813FCAD" bold="true" box="[152,619,757,780]" pageId="6" pageNumber="594">
<emphasis id="B93EEAC2FFA6FFBBFF16FD396B39FCAD" bold="true" box="[152,321,758,780]" italics="true" pageId="6" pageNumber="594">Prespelea morsei</emphasis>
Caterino and Vásquez-Vélez
</emphasis>
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</paragraph>
<paragraph id="8BF536D0FFA6FFBBFF3EFCDE6BDDFC5A" blockId="6.[152,638,757,1020]" pageId="6" pageNumber="594">
<materialsCitation id="3B223C8DFFA6FFBBFF3EFCDE683EFCD1" collectionCode="CUAC" collectorName="Nantahala NF &amp; M. Caterino &amp; A. Haberski &amp; P. Wooden" county="Jackson Co." latitude="35.1321" location="Toxaway Mt." longLatPrecision="6" longitude="-82.9823" pageId="6" pageNumber="594" specimenCode="CUAC000143429" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4CFD59FFA6FFBBFF3EFCDE6B75FC81" box="[176,269,785,807]" count="1" pageId="6" pageNumber="594" type="male">One male</specimenCount>
(MSC-7474,
<specimenCode id="DBEC9EABFFA6FFBBFE1CFCDE683EFC80" box="[402,582,785,806]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000143429</specimenCode>
): NC:
<collectingCounty id="62944E5CFFA6FFBBFF16FCE46B6CFCE6" box="[152,276,811,833]" pageId="6" pageNumber="594">Jackson Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA6FFBBFEAAFCE46BF7FCE7" box="[292,399,811,833]" degrees="35.1321" direction="north" orientation="latitude" pageId="6" pageNumber="594" precision="5" value="35.1321">35.1321°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA6FFBBFE10FCE46875FCE6" box="[414,525,811,832]" degrees="82.9823" direction="west" orientation="longitude" pageId="6" pageNumber="594" precision="5" value="-82.9823">82.9823°W</geoCoordinate>
,
<collectorName id="26BF5306FFA6FFBBFD92FCE46ACFFCFD" pageId="6" pageNumber="594">Nantahala NF</collectorName>
, 4,750′,
<location id="8E95600BFFA6FFBBFE88FC896BF0FCFD" LSID="urn:lsid:plazi:treatment:03E387C6FFA6FFBBFF16FD396B00FA82:8E95600BFFA6FFBBFE88FC896BF0FCFD" box="[262,392,838,859]" county="Jackson Co." latitude="35.1321" longLatPrecision="6" longitude="-82.9823" name="Toxaway Mt." pageId="6" pageNumber="594">Toxaway Mt.</location>
, viii.5.2020,
<collectorName id="26BF5306FFA6FFBBFD8AFC896802FCFA" box="[516,634,838,860]" pageId="6" pageNumber="594">M. Caterino</collectorName>
,
<collectorName id="26BF5306FFA6FFBBFF16FCAE6B6CFCD1" box="[152,276,865,887]" pageId="6" pageNumber="594">A. Haberski</collectorName>
and
<collectorName id="26BF5306FFA6FFBBFEDFFCAE6BB9FCD1" box="[337,449,865,887]" pageId="6" pageNumber="594">P. Wooden</collectorName>
, sifted litter
</materialsCitation>
.
<materialsCitation id="3B223C8DFFA6FFBBFDDBFCAE6BDAFC5A" collectionCode="R" collectorName="Pisgah NF &amp; Hwy &amp; Blue Ridge Parkway &amp; M. Caterino &amp; Kucuk &amp; L. Cushman" county="Transylvania Co." latitude="35.291" location="Transylvania Co." longLatPrecision="6" longitude="-82.9133" pageId="6" pageNumber="594" specimenCode="CUAC000134132" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4CFD59FFA6FFBBFDDBFCAE6ABFFC37" count="1" pageId="6" pageNumber="594" type="male">One male</specimenCount>
(MSC-7475,
<specimenCode id="DBEC9EABFFA6FFBBFEF7FCB46848FC36" box="[377,560,891,912]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000134132</specimenCode>
): NC:
<collectingCounty id="62944E5CFFA6FFBBFF16FC596B3AFC0D" box="[152,322,918,940]" pageId="6" pageNumber="594">Transylvania Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA6FFBBFEC5FC596BC1FC0A" box="[331,441,918,940]" degrees="35.2910" direction="north" orientation="latitude" pageId="6" pageNumber="594" precision="5" value="35.291">35.2910°N</geoCoordinate>
,
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,
<collectorName id="26BF5306FFA6FFBBFDB2FC596AC0FC60" pageId="6" pageNumber="594">Pisgah NF</collectorName>
, 5,122′,
<collectorName id="26BF5306FFA6FFBBFE9BFC7E6B3CFC60" box="[277,324,945,966]" pageId="6" pageNumber="594">Hwy</collectorName>
215,
<quantity id="4CB29B35FFA6FFBBFE09FC7E6BC7FC61" box="[391,447,945,967]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="6" pageNumber="594" unit="mi" value="1.0">1 mi.</quantity>
S. of
<collectorName id="26BF5306FFA6FFBBFD87FC7E6A97FC47" pageId="6" pageNumber="594">Blue Ridge Parkway</collectorName>
, v.8.2018, sifted litter,
<collectorName id="26BF5306FFA6FFBBFE79FC036802FC47" box="[503,634,971,993]" pageId="6" pageNumber="594">M. Caterino</collectorName>
,
<collectionCode id="ED5BAE15FFA6FFBBFF16FC296AD3FC5D" box="[152,171,998,1019]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="594">R</collectionCode>
.
<collectorName id="26BF5306FFA6FFBBFF3BFC296A8EFC5A" box="[181,246,998,1020]" pageId="6" pageNumber="594">Kucuk</collectorName>
and
<collectorName id="26BF5306FFA6FFBBFEA8FC296BDAFC5A" box="[294,418,998,1020]" pageId="6" pageNumber="594">L. Cushman</collectorName>
</materialsCitation>
.
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<taxonomicName id="4C4A4D53FFA6FFBBFF16FBD16BEFFB95" authority="Park" authorityName="Park" box="[152,407,1054,1076]" class="Insecta" family="Staphylinidae" genus="Prespelea" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="594" phylum="Arthropoda" rank="species" species="copelandi">
<emphasis id="B93EEAC2FFA6FFBBFF16FBD16BEFFB95" bold="true" box="[152,407,1054,1076]" pageId="6" pageNumber="594">
<emphasis id="B93EEAC2FFA6FFBBFF16FBD16B26FB92" bold="true" box="[152,350,1054,1076]" italics="true" pageId="6" pageNumber="594">Prespelea copelandi</emphasis>
Park
</emphasis>
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<paragraph id="8BF536D0FFA6FFBBFF3EFBF66B00FA82" blockId="6.[152,639,1054,1316]" pageId="6" pageNumber="594">
<materialsCitation id="3B223C8DFFA6FFBBFF3EFBF66871FB1F" collectionCode="CUAC" collectorName="Nantahala NF &amp; Cowee Bald &amp; M. Caterino &amp; M. Ferro" county="Macon Co." latitude="35.3263" location="Macon Co." longLatPrecision="6" longitude="-83.335" pageId="6" pageNumber="594" specimenCode="CUAC 000110795, CUAC000079340" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D4CFD59FFA6FFBBFF3EFBF66B61FBE9" box="[176,281,1081,1103]" count="1" pageId="6" pageNumber="594" type="male">One male</specimenCount>
,
<specimenCount id="9D4CFD59FFA6FFBBFEA3FBF66BD9FBE9" box="[301,417,1081,1103]" count="1" pageId="6" pageNumber="594" type="female">one female</specimenCount>
(MSC-4151,
<specimenCode id="DBEC9EABFFA6FFBBFDB5FBF66B7BFBCF" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC 000110795</specimenCode>
and MSC-3521,
<specimenCode id="DBEC9EABFFA6FFBBFE4EFB9C680CFBCE" box="[448,628,1107,1128]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000079340</specimenCode>
): NC:
<collectingCounty id="62944E5CFFA6FFBBFF4AFBA16B57FB25" box="[196,303,1134,1155]" pageId="6" pageNumber="594">Macon Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA6FFBBFEB9FBA16BD8FB22" box="[311,416,1134,1156]" degrees="35.3263" direction="north" orientation="latitude" pageId="6" pageNumber="594" precision="5" value="35.3263">35.3263°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA6FFBBFE27FBA1686DFB22" box="[425,533,1134,1156]" degrees="83.3350" direction="west" orientation="longitude" pageId="6" pageNumber="594" precision="5" value="-83.335">83.3350°W</geoCoordinate>
,
<collectorName id="26BF5306FFA6FFBBFD90FBA16ACFFB38" pageId="6" pageNumber="594">Nantahala NF</collectorName>
, 4,925′,
<collectorName id="26BF5306FFA6FFBBFEBDFB466BB9FB39" box="[307,449,1161,1183]" pageId="6" pageNumber="594">Cowee Bald</collectorName>
, vii.9.2019,
<collectorName id="26BF5306FFA6FFBBFDECFB466A93FB1F" pageId="6" pageNumber="594">M. Caterino</collectorName>
and
<collectorName id="26BF5306FFA6FFBBFEA6FB6B6BFEFB1F" box="[296,390,1188,1209]" pageId="6" pageNumber="594">M. Ferro</collectorName>
, sifted litter
</materialsCitation>
.
<materialsCitation id="3B223C8DFFA6FFBBFD97FB6C6B0CFA82" collectionCode="CUAC" collectorName="Nantahala NF &amp; Cowee Bald &amp; F. Etzler &amp; A. Haberski &amp; P. Wooden" county="Macon Co." latitude="35.327" location="Macon Co." longLatPrecision="6" longitude="-83.3359" pageId="6" pageNumber="594" specimenCode="CUAC000135248" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4CFD59FFA6FFBBFD97FB6C6806FB1F" box="[537,638,1187,1209]" count="1" pageId="6" pageNumber="594" type="male">One male</specimenCount>
(MSC-6570,
<specimenCode id="DBEC9EABFFA6FFBBFE96FB716BB4FB72" box="[280,460,1214,1236]" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="6" pageNumber="594" type="University or college">CUAC000135248</specimenCode>
): NC:
<collectingCounty id="62944E5CFFA6FFBBFD82FB716802FB75" box="[524,634,1214,1235]" pageId="6" pageNumber="594">Macon Co.</collectingCounty>
,
<geoCoordinate id="EE7E5017FFA6FFBBFF16FB166B7BFB49" box="[152,259,1241,1263]" degrees="35.3270" direction="north" orientation="latitude" pageId="6" pageNumber="594" precision="5" value="35.327">35.3270°N</geoCoordinate>
,
<geoCoordinate id="EE7E5017FFA6FFBBFE99FB166BFDFB49" box="[279,389,1241,1263]" degrees="83.3359" direction="west" orientation="longitude" pageId="6" pageNumber="594" precision="5" value="-83.3359">83.3359°W</geoCoordinate>
,
<collectorName id="26BF5306FFA6FFBBFE17FB166851FB48" box="[409,553,1241,1263]" pageId="6" pageNumber="594">Nantahala NF</collectorName>
, 4,931′,
<collectorName id="26BF5306FFA6FFBBFF16FB3C6B68FAAF" box="[152,272,1267,1289]" pageId="6" pageNumber="594">Cowee Bald</collectorName>
, ix.15.2020,
<collectorName id="26BF5306FFA6FFBBFE04FB3B6BA3FAAF" box="[394,475,1267,1289]" pageId="6" pageNumber="594">F. Etzler</collectorName>
,
<collectorName id="26BF5306FFA6FFBBFE6DFB3B682EFAAF" box="[483,598,1267,1289]" pageId="6" pageNumber="594">A. Haberski</collectorName>
and
<collectorName id="26BF5306FFA6FFBBFF16FAC16B78FA82" box="[152,256,1294,1316]" pageId="6" pageNumber="594">P. Wooden</collectorName>
, sifted litter
</materialsCitation>
.
</paragraph>
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</treatment>
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151
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<collectingRegion id="4986F80AFFEFFF8DFF77AA22A2D0F9FA" box="[197,397,1543,1565]" country="Canada" name="New Brunswick" pageId="3" pageNumber="443">NEW BRUNSWICK</collectingRegion>
:
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<collectingRegion id="4986F80AFFEFFF8DFE07AA22A15FF9FA" box="[437,514,1543,1565]" country="Australia" name="Victoria" pageId="3" pageNumber="443">Victoria</collectingRegion>
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,
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(
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,
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),
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<collectingDate id="EFB8E9C0FFEFFF8DFE51AA07A124F9D0" box="[483,633,1570,1591]" pageId="3" pageNumber="443" value="2022-01-01">1 January 2022</collectingDate>
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,
<collectorName id="26B7533EFFEFFF8DFD31AA07A258F9B5" pageId="3" pageNumber="443">J. H. Lewis</collectorName>
, dead specimen collected from spiderwebs along trimming of country-house in agricultural area (
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,
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.
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This is a wetland specialist which develops in species of
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<emphasis id="B936EAFAFFEFFF8DFEAAAA82A23DF95B" box="[280,352,1703,1724]" italics="true" pageId="3" pageNumber="443">Scirpus</emphasis>
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L. and
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<emphasis id="B936EAFAFFEFFF8DFE01AA82A2B0F95B" box="[435,493,1703,1724]" italics="true" pageId="3" pageNumber="443">Carex</emphasis>
</taxonomicName>
L. (
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) (
<bibRefCitation id="EFD34B19FFEFFF8DFD7BACE7A068FF30" author="Koch, K." box="[713,821,194,216]" pageId="3" pageNumber="443" refId="ref2307" refString="Koch, K. 1992. Die Kafer Mitteleuropas, Okologie, Volume 3. Goecke &amp; Evers Verlag, Krefeld, Germany, 389 pp." type="book" year="1992">Koch 1992</bibRefCitation>
). This species occurs naturally throughout the Palearctic region, with a range extending from western Europe to the Far East (
<bibRefCitation id="EFD34B19FFEFFF8DFBEBACD2A0CDFEC0" author="Alonso-Zarazaga, M. A. &amp; Barrios, R. &amp; Borovec, P. &amp; Bouchard, R. &amp; Caldara, E. &amp; Colonnelli, L. &amp; Gultekin, P. &amp; Hlava, B. &amp; Korotyaev, C. H. C. &amp; Lyal, A. &amp; Machado, M. &amp; Meregalli, H. &amp; Pierotti, L. &amp; Ren, M. &amp; Sanchez-Ruiz, A. &amp; Sforzi, H. &amp; Silfverberg, J. &amp; Skuhrovec, M. &amp; Tryzna, A. J." pageId="3" pageNumber="443" refId="ref1872" refString="Alonso-Zarazaga, M. A., H. Barrios, R. Borovec, P. Bouchard, R. Caldara, E. Colonnelli, L. Gultekin, P. Hlava, B. Korotyaev, C. H. C. Lyal, A. Machado, M. Meregalli, H. Pierotti, L. Ren, M. Sanchez-Ruiz, A. Sforzi, H. Silfverberg, J. Skuhrovec, M. Tryzna, A. J. Velazquez de Castro, and N. N. Yunakov. 2017. Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias Electronicas SEA 8. Sociedad Entomologica Aragonesa SEA, Zaragoza, Spain, 729 pp." type="book" year="2017">
Alonso-Zarazaga
<emphasis id="B936EAFAFFEFFF8DFC90AD37A008FEC0" box="[802,853,274,295]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
2017
</bibRefCitation>
). The first adventive records of
<taxonomicName id="4C424D6BFFEFFF8DFD6CAD08A06EFEA5" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[734,819,301,322]" class="Insecta" family="Brachyceridae" genus="Notaris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="scirpi">
<emphasis id="B936EAFAFFEFFF8DFD6CAD08A06EFEA5" box="[734,819,301,322]" italics="true" pageId="3" pageNumber="443">N. scirpi</emphasis>
</taxonomicName>
from the Nearctic region are given in
<bibRefCitation id="EFD34B19FFEFFF8DFD71AD62A0F3FEBB" author="Pentinsaari, M. &amp; Anderson, L. &amp; Borowiec, P. &amp; Bouchard, A. &amp; Brunke, H. &amp; Douglas, A. B. T." box="[707,942,327,349]" pageId="3" pageNumber="443" pagination="53 - 150" refId="ref2530" refString="Pentinsaari, M., R. S. Anderson, L. Borowiec, P. Bouchard, A. Brunke, H. Douglas, A. B. T. Smith, and P. D. Hebert. 2019. DNA barcodes reveal 63 overlooked species of Canadian beetles (Insecta, Coleoptera). ZooKeys 894: 53 - 150. doi. org / 10.3897 / zookeys. 894.37862." type="journal article" year="2019">
Pentinsaari
<emphasis id="B936EAFAFFEFFF8DFC86AD6DA03BFEBB" box="[820,870,327,349]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
(2019)
</bibRefCitation>
, which includes
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taken from several localities in
<collectingRegion id="4986F80AFFEFFF8DFBEBAD47A7F8FE9F" box="[1113,1189,354,376]" country="Canada" name="Quebec" pageId="3" pageNumber="443">Quebec</collectingRegion>
,
<collectingCountry id="F3557678FFEFFF8DFD71AD59A053FE75" box="[707,782,380,402]" name="Canada" pageId="3" pageNumber="443">Canada</collectingCountry>
. As noted in
<bibRefCitation id="EFD34B19FFEFFF8DFC14AD59A7F9FE76" author="Pentinsaari, M. &amp; Anderson, L. &amp; Borowiec, P. &amp; Bouchard, A. &amp; Brunke, H. &amp; Douglas, A. B. T." box="[934,1188,380,402]" pageId="3" pageNumber="443" pagination="53 - 150" refId="ref2530" refString="Pentinsaari, M., R. S. Anderson, L. Borowiec, P. Bouchard, A. Brunke, H. Douglas, A. B. T. Smith, and P. D. Hebert. 2019. DNA barcodes reveal 63 overlooked species of Canadian beetles (Insecta, Coleoptera). ZooKeys 894: 53 - 150. doi. org / 10.3897 / zookeys. 894.37862." type="journal article" year="2019">
Pentinsaari
<emphasis id="B936EAFAFFEFFF8DFBACAD58A70BFE76" box="[1054,1110,380,402]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
(2019)
</bibRefCitation>
,
<taxonomicName id="4C424D6BFFEFFF8DFD71ADBDA049FE4A" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[707,788,408,429]" class="Insecta" family="Brachyceridae" genus="Notaris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="scirpi">
<emphasis id="B936EAFAFFEFFF8DFD71ADBDA049FE4A" box="[707,788,408,429]" italics="true" pageId="3" pageNumber="443">N. scirpi</emphasis>
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possesses prominent dense, pale yellow to orange scales on the abdomen and thorax laterally, a character which distinguishes them from other, similar species. The collection of the present specimen in Four Falls,
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, a locality which lies less than 4.5 kilometers away from the Canada-USA border, indicates that this species may also be present in
<collectingRegion id="4986F80AFFEFFF8DFD6EAE77A046FD8F" box="[732,795,594,616]" country="United States of America" name="Maine" pageId="3" pageNumber="443">Maine</collectingRegion>
,
<collectingCountry id="F3557678FFEFFF8DFC97AE77A005FD80" box="[805,856,594,615]" name="United States of America" pageId="3" pageNumber="443">USA</collectingCountry>
. Further field surveying is required to uncover the complete geographic range of
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<emphasis id="B936EAFAFFEFFF8DFD71AEADA04CFD7A" box="[707,785,648,669]" italics="true" pageId="3" pageNumber="443">N. scirpi</emphasis>
</taxonomicName>
in
<collectingRegion id="4986F80AFFEFFF8DFC9EAEA2A09DFD7A" box="[812,960,647,669]" inferredBySuffix="true" pageId="3" pageNumber="443">New Brunswick</collectingRegion>
and surrounding regions.
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187
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<emphasis id="B936EAFAFFEFFF8DFF64AE97A2ACFD20" bold="true" box="[214,497,690,711]" italics="true" pageId="3" pageNumber="443">OTIoRHYnCHUS RUgoSoSTRIaTUS</emphasis>
(Goeze, 1777)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFD36E8FFEFFF8DFF77AEF9A215FCBB" blockId="3.[173,660,732,1474]" pageId="3" pageNumber="443">
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<collectingRegion id="4986F80AFFEFFF8DFF77AEF9A2D0FD15" box="[197,397,732,754]" country="Canada" name="New Brunswick" pageId="3" pageNumber="443">NEW BRUNSWICK</collectingRegion>
:
</emphasis>
<collectingRegion id="4986F80AFFEFFF8DFE07AEF9A15FFD15" box="[437,514,732,754]" country="Australia" name="Victoria" pageId="3" pageNumber="443">Victoria</collectingRegion>
<collectingMunicipality id="6B99AC92FFEFFF8DFDBCAEF9A10BFD16" box="[526,598,732,753]" pageId="3" pageNumber="443">County</collectingMunicipality>
,
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(
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,
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),
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<collectingDate id="EFB8E9C0FFEFFF8DFE51AED2A124FCEB" box="[483,633,759,780]" pageId="3" pageNumber="443" value="2022-01-01">1 January 2022</collectingDate>
</date>
,
<collectorName id="26B7533EFFEFFF8DFD31AED2A257FCCF" pageId="3" pageNumber="443">J. H. Lewis</collectorName>
, dead specimens collected from spiderwebs in private garage in agricultural area (
<specimenCount id="9D44FD61FFEFFF8DFD34AF09A3B4FCBA" count="6" pageId="3" pageNumber="443" type="adult">6 adults</specimenCount>
,
<collectionCode id="ED53AE2DFFEFFF8DFF41AF62A263FCBB" box="[243,318,839,860]" pageId="3" pageNumber="443">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<paragraph id="8BFD36E8FFEFFF8DFF77AF47A1CDFA25" blockId="3.[173,660,732,1474]" pageId="3" pageNumber="443">
This species is native to the western Palearctic region (Warner and Negley 1976), and is adventive in
<collectingCountry id="F3557678FFEFFF8DFF75AFB2A27CFC4A" box="[199,289,919,941]" name="Australia" pageId="3" pageNumber="443">Australia</collectingCountry>
,
<collectingCountry id="F3557678FFEFFF8DFE9CAFB2A2E9FC4A" box="[302,436,919,941]" name="New Zealand" pageId="3" pageNumber="443">New Zealand</collectingCountry>
,
<collectingCountry id="F3557678FFEFFF8DFE72AFB2A2ABFC4A" box="[448,502,919,941]" name="Chile" pageId="3" pageNumber="443">Chile</collectingCountry>
,
<collectingCountry id="F3557678FFEFFF8DFDB0AFB2A139FC4A" box="[514,612,919,941]" name="Argentina" pageId="3" pageNumber="443">Argentina</collectingCountry>
, and the Nearctic region (
<bibRefCitation id="EFD34B19FFEFFF8DFEC7AF97A1D2FC20" author="Alonso-Zarazaga, M. A. &amp; Barrios, R. &amp; Borovec, P. &amp; Bouchard, R. &amp; Caldara, E. &amp; Colonnelli, L. &amp; Gultekin, P. &amp; Hlava, B. &amp; Korotyaev, C. H. C. &amp; Lyal, A. &amp; Machado, M. &amp; Meregalli, H. &amp; Pierotti, L. &amp; Ren, M. &amp; Sanchez-Ruiz, A. &amp; Sforzi, H. &amp; Silfverberg, J. &amp; Skuhrovec, M. &amp; Tryzna, A. J." box="[373,655,946,968]" pageId="3" pageNumber="443" refId="ref1872" refString="Alonso-Zarazaga, M. A., H. Barrios, R. Borovec, P. Bouchard, R. Caldara, E. Colonnelli, L. Gultekin, P. Hlava, B. Korotyaev, C. H. C. Lyal, A. Machado, M. Meregalli, H. Pierotti, L. Ren, M. Sanchez-Ruiz, A. Sforzi, H. Silfverberg, J. Skuhrovec, M. Tryzna, A. J. Velazquez de Castro, and N. N. Yunakov. 2017. Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias Electronicas SEA 8. Sociedad Entomologica Aragonesa SEA, Zaragoza, Spain, 729 pp." type="book" year="2017">
Alonso-Zarazaga
<emphasis id="B936EAFAFFEFFF8DFD91AF97A10BFC20" box="[547,598,946,967]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EFD34B19FFEFFF8DFF1FAFE9A234FC06" author="Del Rio, M. G." box="[173,361,972,994]" pageId="3" pageNumber="443" pagination="101 - 110" refId="ref2104" refString="Del Rio, M. G., P. Klasmer, and A. A. Lanteri. 2010. Weevils (Coleoptera: Curculionidae) harmful for berry fruits in Argentina. Revista de la Sociedad Entomologica Argentina 69: 101 - 110." type="journal article" year="2010">
Del Río
<emphasis id="B936EAFAFFEFFF8DFF4DAFE8A26CFC06" box="[255,305,972,994]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EFD34B19FFEFFF8DFEC1AFE9A2A1FC06" author="Kuschel, G." box="[371,508,972,994]" pageId="3" pageNumber="443" refId="ref2334" refString="Kuschel, G. 1990. Beetles in a Suburban Environment: A New Zealand Case Study. The Identity and Status of Coleoptera in the Natural and Modified Habitats of Lynfield, Auckland (1974 - 1989). DSIR Plant Protection Report No. 3. New Zealand Department of Scientific and Industrial Research, Auckland, New Zealand, 118 pp." type="book" year="1990">Kuschel 1990</bibRefCitation>
). In
<collectingCountry id="F3557678FFEFFF8DFD95AFE9A12CFC05" box="[551,625,972,994]" name="Canada" pageId="3" pageNumber="443">Canada</collectingCountry>
,
<taxonomicName id="4C424D6BFFEFFF8DFDC9AFE8A265FC1A" baseAuthorityName="Goeze" baseAuthorityYear="1777" class="Insecta" family="Curculionidae" genus="Otiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosostriatus">
<emphasis id="B936EAFAFFEFFF8DFDC9AFE8A265FC1A" italics="true" pageId="3" pageNumber="443">O. rugosostriatus</emphasis>
</taxonomicName>
is known from
<collectingRegion id="4986F80AFFEFFF8DFE50AFC2A1D2FC1A" box="[482,655,999,1021]" country="Canada" name="British Columbia" pageId="3" pageNumber="443">British Columbia</collectingRegion>
,
<collectingRegion id="4986F80AFFEFFF8DFF1FA827A3A4FBFF" box="[173,249,1026,1048]" country="Canada" name="Ontario" pageId="3" pageNumber="443">Ontario</collectingRegion>
,
<collectingRegion id="4986F80AFFEFFF8DFEB0A827A213FBFF" box="[258,334,1026,1048]" country="Canada" name="Quebec" pageId="3" pageNumber="443">Quebec</collectingRegion>
, and
<collectingRegion id="4986F80AFFEFFF8DFECDA827A2A8FBFF" box="[383,501,1026,1048]" country="Canada" name="Nova Scotia" pageId="3" pageNumber="443">Nova Scotia</collectingRegion>
(
<bibRefCitation id="EFD34B19FFEFFF8DFDB3A827A382FBD6" author="Bousquet, Y. &amp; Bouchard, A. E." pageId="3" pageNumber="443" refId="ref1996" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska - 2 nd Edition. Pensoft Series Faunistica, No. 109. Pensoft Publishers, Sofia, Bulgaria, 402 pp." type="book" year="2013">
Bousquet
<emphasis id="B936EAFAFFEFFF8DFDD1A827A1C9FBF0" box="[611,660,1026,1047]" italics="true" pageId="3" pageNumber="443">et al.</emphasis>
2013
</bibRefCitation>
), and also occurs widely throughout the
<collectingCountry id="F3557678FFEFFF8DFDE1A839A23FFBAA" name="United States of America" pageId="3" pageNumber="443">United States of America</collectingCountry>
(O’Brien and Wibmer 1982).
<taxonomicName id="4C424D6BFFEFFF8DFF1FA877A2E3FB80" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[173,446,1106,1127]" class="Insecta" family="Curculionidae" genus="Otiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosostriatus">
<emphasis id="B936EAFAFFEFFF8DFF1FA877A2E3FB80" box="[173,446,1106,1127]" italics="true" pageId="3" pageNumber="443">Otiorhynchus rugosostriatus</emphasis>
</taxonomicName>
is easily distinguished from congeners in the region by characters listed in Bright and Bouchard (2008). This species is known to be parthenogenetic across much of its range (
<bibRefCitation id="EFD34B19FFEFFF8DFF06A899A201FB36" author="Dieckmann, L." box="[180,348,1212,1234]" pageId="3" pageNumber="443" pagination="145 - 310" refId="ref2150" refString="Dieckmann, L. 1980. Beitrage zur Insektenfauna der DDR: Coleoptera - Curculionidae (Brachycerinae, Otiorhynchinae, Brachyderinae). Beitrage zur Entomologie 30: 145 - 310." type="journal article" year="1980">Dieckmann 1980</bibRefCitation>
) and feeds on a wide variety of plants, including
<taxonomicName id="4C424D6BFFEFFF8DFED0A8F2A29FFB0B" box="[354,450,1239,1260]" class="Magnoliopsida" family="Primulaceae" genus="Cyclamen" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFED0A8F2A29FFB0B" box="[354,450,1239,1260]" italics="true" pageId="3" pageNumber="443">Cyclamen</emphasis>
</taxonomicName>
L. and
<taxonomicName id="4C424D6BFFEFFF8DFD90A8F2A12CFB0B" box="[546,625,1239,1260]" class="Magnoliopsida" family="Primulaceae" genus="Primula" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFD90A8F2A12CFB0B" box="[546,625,1239,1260]" italics="true" pageId="3" pageNumber="443">Primula</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D6BFFEFFF8DFF06A8D7A273FAEF" box="[180,302,1266,1288]" class="Magnoliopsida" family="Primulaceae" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="family">Primulaceae</taxonomicName>
),
<taxonomicName id="4C424D6BFFEFFF8DFE8EA8D7A2CEFAE0" box="[316,403,1266,1287]" class="Magnoliopsida" family="Rosaceae" genus="Fragaria" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFE8EA8D7A2CEFAE0" box="[316,403,1266,1287]" italics="true" pageId="3" pageNumber="443">Fragaria</emphasis>
</taxonomicName>
L. and
<taxonomicName id="4C424D6BFFEFFF8DFE65A8D7A15BFAE0" box="[471,518,1266,1287]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFE65A8D7A15BFAE0" box="[471,518,1266,1287]" italics="true" pageId="3" pageNumber="443">Rosa</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D6BFFEFFF8DFD98A8D7A1D5FAE0" box="[554,648,1266,1287]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
),
<taxonomicName id="4C424D6BFFEFFF8DFF1FA929A38CFAC6" box="[173,209,1292,1313]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFF1FA929A38CFAC6" box="[173,209,1292,1313]" italics="true" pageId="3" pageNumber="443">Ilex</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D6BFFEFFF8DFF4CA929A2D7FAC5" box="[254,394,1292,1314]" class="Magnoliopsida" family="Aquifoliaceae" kingdom="Plantae" order="Aquifoliales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="family">Aquifoliaceae</taxonomicName>
),
<taxonomicName id="4C424D6BFFEFFF8DFE2EA928A2A2FAC5" box="[412,511,1293,1314]" class="Magnoliopsida" family="Oleaceae" genus="Ligustrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFE2EA928A2A2FAC5" box="[412,511,1293,1314]" italics="true" pageId="3" pageNumber="443">Ligustrum</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D6BFFEFFF8DFD99A929A1D5FAC5" authorityName="von Hoffmannsegg &amp; Link" authorityYear="1809" box="[555,648,1292,1314]" class="Magnoliopsida" family="Oleaceae" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="family">Oleaceae</taxonomicName>
), and
<taxonomicName id="4C424D6BFFEFFF8DFF65A902A276FADB" box="[215,299,1319,1340]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFAFFEFFF8DFF65A902A276FADB" box="[215,299,1319,1340]" italics="true" pageId="3" pageNumber="443">Solanum</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C424D6BFFEFFF8DFEE7A902A29AFADA" box="[341,455,1319,1341]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="443" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
); however, it is most well-known as a pest on strawberries (Warner and Negley 1976). Given the widespread presence of
<taxonomicName id="4C424D6BFFEFFF8DFDCEA978A26AFA6A" baseAuthorityName="Goeze" baseAuthorityYear="1777" class="Insecta" family="Curculionidae" genus="Otiorhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="443" phylum="Arthropoda" rank="species" species="rugosostriatus">
<emphasis id="B936EAFAFFEFFF8DFDCEA978A26AFA6A" italics="true" pageId="3" pageNumber="443">O. rugosostriatus</emphasis>
</taxonomicName>
in Nova Scotia and southern Quebec (Bright and Bouchard 2008), it is unsurprising that this species is also established in New Brunswick.
</paragraph>
</subSubSection>
</treatment>
</document>

142
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<emphasis id="B926EAD9FF88FFF4B6FAEE40FE55420F" bold="true" box="[240,438,1108,1129]" italics="true" pageId="5" pageNumber="573">TeNeBROIDeS cOLLARIS</emphasis>
(Sturm, 1807)
</taxonomicName>
, New State Record
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3486540FF88FFF7B78EEE98FD9E4697" lastPageId="6" lastPageNumber="574" pageId="5" pageNumber="573" type="description">
<paragraph id="8BED36CBFF88FFF4B78EEE98FE5F42C7" blockId="5.[240,592,1108,1186]" box="[388,444,1164,1186]" pageId="5" pageNumber="573">
<figureCitation id="13692A4EFF88FFF4B78EEE98FE5F42C7" box="[388,444,1164,1186]" captionStart="Fig" captionStartId="5.[731,762,1177,1196]" captionTargetBox="[707,1193,193,1159]" captionTargetId="figure-538@5.[707,1193,193,1159]" captionTargetPageId="5" captionText="Fig. 3. Dorsal habitus of Tenebroides collaris collected from a Lindgren funnel trap baited with ETOH and alphapinene, 1 mi. south of Ridge Rd., Carter County, Montana, 8 June to 20 July 2012, by Ivie and Foley (MTEC)." figureDoi="http://doi.org/10.5281/zenodo.13252785" httpUri="https://zenodo.org/record/13252785/files/figure.png" pageId="5" pageNumber="573">Fig. 3</figureCitation>
</paragraph>
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<paragraph id="8BED36CBFF88FFF4B4D1EE8DFB8B4292" blockId="5.[707,1193,1177,1268]" pageId="5" pageNumber="573">
<emphasis id="B926EAD9FF88FFF4B4D1EE8DFCF242CA" bold="true" box="[731,785,1177,1196]" pageId="5" pageNumber="573">Fig. 3.</emphasis>
Dorsal habitus of
<taxonomicName id="4C524D48FF88FFF4B5BFEE8DFBB942CA" box="[949,1114,1177,1196]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="573" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B926EAD9FF88FFF4B5BFEE8DFBB942CA" box="[949,1114,1177,1196]" italics="true" pageId="5" pageNumber="573">Tenebroides collaris</emphasis>
</taxonomicName>
collected from a Lindgren funnel trap baited with ETOH and alphapinene, 1 mi. south of Ridge Rd., Carter County, Montana, 8 June to 20 July 2012, by Ivie and Foley (MTEC).
</paragraph>
</caption>
<paragraph id="8BED36CBFF88FFF4B6CFEEAEFDCD40D6" blockId="5.[173,659,1210,1712]" pageId="5" pageNumber="573">
This unmistakable species is easily recognized by its combination of a depressed body with orange-red head and pronotum, and relatively smooth, black elytra (
<figureCitation id="13692A4EFF88FFF4B6F8EF1EFECF4379" box="[242,300,1290,1312]" captionStart="Fig" captionStartId="5.[731,762,1177,1196]" captionTargetBox="[707,1193,193,1159]" captionTargetId="figure-538@5.[707,1193,193,1159]" captionTargetPageId="5" captionText="Fig. 3. Dorsal habitus of Tenebroides collaris collected from a Lindgren funnel trap baited with ETOH and alphapinene, 1 mi. south of Ridge Rd., Carter County, Montana, 8 June to 20 July 2012, by Ivie and Foley (MTEC)." figureDoi="http://doi.org/10.5281/zenodo.13252785" httpUri="https://zenodo.org/record/13252785/files/figure.png" pageId="5" pageNumber="573">Fig. 3</figureCitation>
). Prior to records presented here, the distribution of
<taxonomicName id="4C524D48FF88FFF4B74FEF31FE4A435F" box="[325,425,1316,1338]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="573" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B926EAD9FF88FFF4B74FEF31FE4A435F" box="[325,425,1316,1338]" italics="true" pageId="5" pageNumber="573">T. collaris</emphasis>
</taxonomicName>
was listed as “Eastern
<collectingCountry id="F345765BFF88FFF4B6A7EF2BFED04332" box="[173,307,1343,1365]" name="United States of America" pageId="5" pageNumber="573">United States</collectingCountry>
, Ontario, west to
<collectingRegion id="4996F829FF88FFF4B7EFEF2BFDA04333" box="[485,579,1343,1365]" country="United States of America" name="Michigan" pageId="5" pageNumber="573">Michigan</collectingRegion>
, eastern
<collectingRegion id="4996F829FF88FFF4B6A7EF4EFF054309" box="[173,230,1370,1391]" country="United States of America" name="Texas" pageId="5" pageNumber="573">Texas</collectingRegion>
” by
<bibRefCitation id="EFC34B3AFF88FFF4B71AEF4EFE7B4309" author="Barron, J. R." box="[272,408,1370,1391]" pageId="5" pageNumber="573" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron (1971)</bibRefCitation>
. The furthest northwestern
<collectingCountry id="F345765BFF88FFF4B6A7EF60FF2F43EF" box="[173,204,1396,1417]" name="United States of America" pageId="5" pageNumber="573">US</collectingCountry>
record documented by
<bibRefCitation id="EFC34B3AFF88FFF4B7C5EF61FDBC43EF" author="Barron, J. R." box="[463,607,1396,1418]" pageId="5" pageNumber="573" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron (1971)</bibRefCitation>
was Alpena, Alpena County,
<collectingRegion id="4996F829FF88FFF4B79DEF9BFE1643C3" box="[407,501,1423,1445]" country="United States of America" name="Michigan" pageId="5" pageNumber="573">Michigan</collectingRegion>
, in the far northeastern portion of that state, while the furthest southwestern record was from Grayburg, Hardin Co.,
<collectingRegion id="4996F829FF88FFF4B6A7EFCBFF0B4392" box="[173,232,1503,1524]" country="United States of America" name="Texas" pageId="5" pageNumber="573">Texas</collectingRegion>
. In
<collectingRegion id="4996F829FF88FFF4B71AEFCBFE8B4392" box="[272,360,1503,1524]" country="United States of America" name="Montana" pageId="5" pageNumber="573">Montana</collectingRegion>
, this species is now known to occur in 12 counties in the eastern half of the state (
<figureCitation id="13692A4EFF88FFF4B6BEEC00FEE2404F" box="[180,257,1556,1578]" captionStart="Fig" captionStartId="4.[176,207,1637,1656]" captionTargetBox="[196,1127,194,1618]" captionTargetId="figure-6@4.[195,1127,193,1619]" captionTargetPageId="4" captionText="Fig. 2. Distribution of 12 species of bark-gnawing beetles by county in Montana.A) Calitys scabra, B) Corticotomus caviceps, C) Temnoscheila chlorodia, D) Tenebroides collaris, E) Tenebroides corticalis, F) Tenebroides crassicornis, G) Tenebroides occidentalis, H) Tenebroides sinuatus, I) Peltis fraterna, J) Peltis pippingskoeldi, K) Peltis septentrionalis, L) Eronyxa angusta." figureDoi="http://doi.org/10.5281/zenodo.13252783" httpUri="https://zenodo.org/record/13252783/files/figure.png" pageId="5" pageNumber="573">Fig. 2D</figureCitation>
) and is represented by
<specimenCount id="9D54FD42FF88FFF4B7E3EC00FD8C404C" box="[489,623,1556,1578]" count="41" pageId="5" pageNumber="573" type="generic">41 specimens</specimenCount>
, all but one collected by Lindgren funnels. Available label data indicate that some funnels were baited with either EtOH or a combination of EtOH and alpha-pinene. The earliest records are from 1990 from both Carter and Rosebud counties.
</paragraph>
<paragraph id="8BED36CBFF88FFF4B4D1EF31FB7A43EC" blockId="5.[707,1192,1317,1712]" pageId="5" pageNumber="573">
<emphasis id="B926EAD9FF88FFF4B4D1EF31FC07435D" bold="true" box="[731,996,1317,1339]" pageId="5" pageNumber="573">
County Records (
<figureCitation id="13692A4EFF88FFF4B580EF31FC34435D" box="[906,983,1317,1339]" captionStart="Fig" captionStartId="4.[176,207,1637,1656]" captionTargetBox="[196,1127,194,1618]" captionTargetId="figure-6@4.[195,1127,193,1619]" captionTargetPageId="4" captionText="Fig. 2. Distribution of 12 species of bark-gnawing beetles by county in Montana.A) Calitys scabra, B) Corticotomus caviceps, C) Temnoscheila chlorodia, D) Tenebroides collaris, E) Tenebroides corticalis, F) Tenebroides crassicornis, G) Tenebroides occidentalis, H) Tenebroides sinuatus, I) Peltis fraterna, J) Peltis pippingskoeldi, K) Peltis septentrionalis, L) Eronyxa angusta." figureDoi="http://doi.org/10.5281/zenodo.13252783" httpUri="https://zenodo.org/record/13252783/files/figure.png" pageId="5" pageNumber="573">Fig. 2D</figureCitation>
):
</emphasis>
Big Horn (1), Carter (13), Dawson (3), Garfield (2), Golden Valley (1), Musselshell (1), Phillips (2), Powder River (4), Prairie (3), Rosebud (2), Treasure (7), Valley (2).
</paragraph>
<paragraph id="8BED36CBFF88FFF7B4D1EF84FD9E4697" blockId="5.[707,1192,1317,1712]" lastBlockId="6.[152,638,194,242]" lastPageId="6" lastPageNumber="574" pageId="5" pageNumber="573">
Additionally, during the course of identifying trogossitid specimens in 2020 for the C. P. Gillette Museum of Arthropod Diversity,
<collectingRegion id="4996F829FF88FFF4B21EEFD1FB4B43BC" box="[1044,1192,1477,1499]" country="United States of America" name="Colorado" pageId="5" pageNumber="573">Colorado State</collectingRegion>
University, Fort Collins,
<collectingRegion id="4996F829FF88FFF4B5A6EFF4FBE44390" box="[940,1031,1504,1526]" country="United States of America" name="Colorado" pageId="5" pageNumber="573">Colorado</collectingRegion>
,
<collectingCountry id="F345765BFF88FFF4B205EFF4FBA24393" box="[1039,1089,1504,1525]" name="United States of America" pageId="5" pageNumber="573">USA</collectingCountry>
, the author discovered a specimen of
<taxonomicName id="4C524D48FF88FFF4B5B2EFEFFBFB4069" box="[952,1048,1530,1552]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="573" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B926EAD9FF88FFF4B5B2EFEFFBFB4069" box="[952,1048,1530,1552]" italics="true" pageId="5" pageNumber="573">T. collaris</emphasis>
</taxonomicName>
collected from Larimer County,
<collectingRegion id="4996F829FF88FFF4B56FEC01FC5D404D" box="[869,958,1557,1579]" country="United States of America" name="Colorado" pageId="5" pageNumber="573">Colorado</collectingRegion>
which constitutes a
<emphasis id="B926EAD9FF88FFF4B275EC01FCA14023" bold="true" pageId="5" pageNumber="573">new state record</emphasis>
. The specimen was collected from
<taxonomicName id="4C524D48FF88FFF4B4C9EC5FFCA24039" box="[707,833,1610,1632]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="573" phylum="Tracheophyta" rank="species" species="ponderosa">
<emphasis id="B926EAD9FF88FFF4B4C9EC5FFCA24039" box="[707,833,1610,1632]" italics="true" pageId="5" pageNumber="573">P. ponderosa</emphasis>
</taxonomicName>
in 1996 (D. Leatherman,
<emphasis id="B926EAD9FF88FFF4B24CEC5FFB614039" box="[1094,1154,1610,1632]" italics="true" pageId="5" pageNumber="573">in litt.</emphasis>
). A second
<collectingRegion id="4996F829FF88FFF4B506EC71FC86401D" box="[780,869,1637,1659]" country="United States of America" name="Colorado" pageId="5" pageNumber="573">Colorado</collectingRegion>
specimen was collected approximately
<quantity id="4CAA9B2EFF88FFF4B51AEC94FCBC40F0" box="[784,863,1664,1686]" metricMagnitude="5" metricUnit="m" metricValue="1.28" pageId="5" pageNumber="573" unit="km" value="128.0">128 km</quantity>
south southeast of the Larimer County record in Douglas County from a
<collectingRegion id="4996F829FF88FFF4B245EC8EFB4B40D6" box="[1103,1192,1690,1712]" country="United States of America" name="Colorado" pageId="5" pageNumber="573">Colorado</collectingRegion>
Department of Agriculture Lindgren funnel baited with UHR EtOH and omega-pinene (C. Harp,
<emphasis id="B926EAD9FF8BFFF7B431EAC9FD924697" box="[571,625,220,242]" italics="true" pageId="6" pageNumber="574">in litt.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>

137
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@ -0,0 +1,137 @@
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Müller, 1764
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<paragraph id="8BED36CBFF8BFFF7B4CFEBFEFC4F44DC" blockId="6.[685,1173,490,1178]" pageId="6" pageNumber="574">
This genus is restricted to the Northern Hemisphere. Three species are known from the
<collectingCountry id="F345765BFF8BFFF7B203E810FB6C447F" box="[1033,1167,516,538]" name="United States of America" pageId="6" pageNumber="574">United States</collectingCountry>
, none of which were recorded as occurring in
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by
<bibRefCitation id="EFC34B3AFF8BFFF7B522E82EFC484429" author="Barron, J. R." box="[808,939,570,591]" pageId="6" pageNumber="574" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron (1971</bibRefCitation>
,
<bibRefCitation id="EFC34B3AFF8BFFF7B5BFE82EFC0A4436" author="Barron, J. R." box="[949,1001,570,592]" pageId="6" pageNumber="574" pagination="193 - 202" refId="ref5257" refString="Barron, J. R. 1996. Review of the Nearctic species of Ostoma (Coleoptera: Cleroidea, Trogositidae). Annals of the Entomological Society of America 89: 193 - 202." type="journal article" year="1996">1996</bibRefCitation>
).
<bibRefCitation id="EFC34B3AFF8BFFF7B5F0E82EFB704429" author="Barron, J. R." box="[1018,1171,570,591]" pageId="6" pageNumber="574" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron’s (1971)</bibRefCitation>
lack of
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records, however, was most likely an artifact of collecting as he indicated that species of this genus are widely distributed in the western
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and
<collectingCountry id="F345765BFF8BFFF7B557E8B0FC4B44DC" box="[861,936,676,698]" name="Canada" pageId="6" pageNumber="574">Canada</collectingCountry>
.
</paragraph>
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<paragraph id="8BED36CBFF8BFFF7B4CFE8ABFC4142FC" blockId="6.[685,1173,490,1178]" pageId="6" pageNumber="574">
<bibRefCitation id="EFC34B3AFF8BFFF7B4CFE8ABFCBB44B3" author="Barron, J. R." box="[709,856,703,725]" pageId="6" pageNumber="574" pagination="193 - 202" refId="ref5257" refString="Barron, J. R. 1996. Review of the Nearctic species of Ostoma (Coleoptera: Cleroidea, Trogositidae). Annals of the Entomological Society of America 89: 193 - 202." type="journal article" year="1996">Barron (1996)</bibRefCitation>
reviewed Nearctic
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<emphasis id="B926EAD9FF8BFFF7B23AE8ABFB8444B2" box="[1072,1127,703,724]" italics="true" pageId="6" pageNumber="574">Peltis</emphasis>
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(as
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<emphasis id="B926EAD9FF8BFFF7B4A7E8CEFD144489" box="[685,759,730,751]" italics="true" pageId="6" pageNumber="574">Ostoma</emphasis>
Laicharting
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), and changed certain names he had used previously (
<bibRefCitation id="EFC34B3AFF8BFFF7B58EE8E0FBE5456F" author="Barron, J. R." box="[900,1030,756,777]" pageId="6" pageNumber="574" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron 1971</bibRefCitation>
): the Nearctic members of his 1971 Holarctic
<taxonomicName id="4C524D48FF8BFFF7B5D7E91BFCF54526" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="574" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B926EAD9FF8BFFF7B5D7E91BFB714542" box="[989,1170,783,804]" italics="true" pageId="6" pageNumber="574">Ostoma ferruginea</emphasis>
(Linnaeus)
</taxonomicName>
were recognized as
<taxonomicName id="4C524D48FF8BFFF7B5E4E93EFCE8453C" authority="(Randall)" authorityName="Randall" authorityYear="1838" baseAuthorityName="Randall" baseAuthorityYear="1838" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="574" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="B926EAD9FF8BFFF7B5E4E93EFB704559" box="[1006,1171,810,831]" italics="true" pageId="6" pageNumber="574">Ostoma fraterna</emphasis>
(Randall)
</taxonomicName>
, while
<taxonomicName id="4C524D48FF8BFFF7B552E951FB80453F" authority="Casey" authorityName="Casey" box="[856,1123,836,858]" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="574" phylum="Arthropoda" rank="species" species="columbiana">
<emphasis id="B926EAD9FF8BFFF7B552E951FBFD453F" box="[856,1054,836,858]" italics="true" pageId="6" pageNumber="574">Ostoma columbiana</emphasis>
Casey
</taxonomicName>
was placed as a junior synonym of
<taxonomicName id="4C524D48FF8BFFF7B5B5E94BFCE945F6" authority="(Randall)" baseAuthorityName="Randall" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="574" phylum="Arthropoda" rank="species" species="septentrionalis">
<emphasis id="B926EAD9FF8BFFF7B5B5E94BFB704512" box="[959,1171,863,884]" italics="true" pageId="6" pageNumber="574">Ostoma septentrionalis</emphasis>
(Randall)
</taxonomicName>
, thus leaving only
<taxonomicName id="4C524D48FF8BFFF7B5B2E96EFCDB45CC" authority="(Mannerheim)" baseAuthorityName="Mannerheim" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="574" phylum="Arthropoda" rank="species" species="pippingskoeldi">
<emphasis id="B926EAD9FF8BFFF7B5B2E96EFB7045E9" box="[952,1171,890,911]" italics="true" pageId="6" pageNumber="574">Ostoma pippingskoeldi</emphasis>
(Mannerheim)
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unchanged among the Nearctic fauna. Although
<bibRefCitation id="EFC34B3AFF8BFFF7B569E9BBFBE345A2" author="Kolibac, J." box="[867,1024,943,965]" pageId="6" pageNumber="574" pagination="1 - 194" refId="ref5962" refString="Kolibac, J. 2013. Trogossitidae: A review of the beetle family, with a catalogue and keys. ZooKeys 366: 1 - 194." type="journal article" year="2013">Kolibáč (2013)</bibRefCitation>
cited
<bibRefCitation id="EFC34B3AFF8BFFF7B25AE9BBFD0C4586" author="Barron, J. R." pageId="6" pageNumber="574" pagination="193 - 202" refId="ref5257" refString="Barron, J. R. 1996. Review of the Nearctic species of Ostoma (Coleoptera: Cleroidea, Trogositidae). Annals of the Entomological Society of America 89: 193 - 202." type="journal article" year="1996">Barron (1996)</bibRefCitation>
, he missed these nomenclatural changes, and used
<bibRefCitation id="EFC34B3AFF8BFFF7B4D4E9F0FC96459F" author="Barron, J. R." box="[734,885,996,1017]" pageId="6" pageNumber="574" pagination="1 - 143" refId="ref5220" refString="Barron, J. R. 1971. A revision of the Trogositidae of America north of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron’s (1971)</bibRefCitation>
interpretation in his checklist. This should be considered a
<emphasis id="B926EAD9FF8BFFF7B5C1E9EBFBB14272" box="[971,1106,1023,1044]" italics="true" pageId="6" pageNumber="574">lapsus calami</emphasis>
, not a nomenclatural change, as
<bibRefCitation id="EFC34B3AFF8BFFF7B5A5EE0EFBA74249" author="Kolibac, J." box="[943,1092,1050,1072]" pageId="6" pageNumber="574" pagination="1 - 194" refId="ref5962" refString="Kolibac, J. 2013. Trogossitidae: A review of the beetle family, with a catalogue and keys. ZooKeys 366: 1 - 194." type="journal article" year="2013">Kolibáč (2013)</bibRefCitation>
did not include
<bibRefCitation id="EFC34B3AFF8BFFF7B4F4EE21FC68422C" author="Barron, J. R." box="[766,907,1076,1098]" pageId="6" pageNumber="574" pagination="193 - 202" refId="ref5257" refString="Barron, J. R. 1996. Review of the Nearctic species of Ostoma (Coleoptera: Cleroidea, Trogositidae). Annals of the Entomological Society of America 89: 193 - 202." type="journal article" year="1996">Barron (1996)</bibRefCitation>
in his synonymical table. I therefore follow
<bibRefCitation id="EFC34B3AFF8BFFF7B564EE5BFBEF4203" author="Barron, J. R." box="[878,1036,1103,1125]" pageId="6" pageNumber="574" pagination="193 - 202" refId="ref5257" refString="Barron, J. R. 1996. Review of the Nearctic species of Ostoma (Coleoptera: Cleroidea, Trogositidae). Annals of the Entomological Society of America 89: 193 - 202." type="journal article" year="1996">Barron’s (1996)</bibRefCitation>
species-level taxonomy here.
<bibRefCitation id="EFC34B3AFF8BFFF7B54BEE7EFC2D4219" author="Kolibac, J." box="[833,974,1130,1152]" pageId="6" pageNumber="574" pagination="1 - 194" refId="ref5962" refString="Kolibac, J. 2013. Trogossitidae: A review of the beetle family, with a catalogue and keys. ZooKeys 366: 1 - 194." type="journal article" year="2013">Kolibáč (2013)</bibRefCitation>
stated that all species of
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<emphasis id="B926EAD9FF8BFFF7B4CDEE90FD1D42FF" box="[711,766,1156,1177]" italics="true" pageId="6" pageNumber="574">Peltis</emphasis>
</taxonomicName>
are fungivorous.
</paragraph>
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85
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<mods:title id="D22DEAF94AC4F08ED2F918EB873E6BE0">The Bark-Gnawing Beetles of Montana, USA (Coleoptera: Trogossitidae, Peltidae, and Lophocateridae), Including New Substantial Distributional Records for Tenebroides collaris (Sturm)</mods:title>
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<emphasis id="B926EAD9FF8BFFF7B6B0EB13FE71477A" bold="true" box="[186,402,263,284]" italics="true" pageId="6" pageNumber="574">TeNeBROIDeS cORtIcALIS</emphasis>
(Melsheimer, 1844)
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species has an extensive distribution throughout
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and the United States southward into
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and
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(
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).
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is possible this species occurs in all of Montana. All but 19 of the
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examined were collected by either funnel or vane traps
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.
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County Records (
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):
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Big Horn (6), Blaine (3), Carter (27), Cascade (1), Custer (8), Fallon (5), Gallatin (4), Hill (19), Jefferson (4), Lewis and Clark (5), Mineral (1), Musselshell (1), Park (1), Phillips (2), Pondera (6), Powder River (2), Powell (5), Richland (24), Roosevelt (1), Rosebud (13), Stillwater (1), Toole (1), Valley (3), Wheatland (3), Wibaux (8), Yellowstone (7).
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280
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<mods:titleInfo id="BB5CD595C3DFA21CCD05F836FE6F37A2">
<mods:title id="0BFD16CE82BCCB40BC8727DFC8B29EC2">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="AA8B9E54192BDF1AF30462E75731D171">
<mods:date id="5F85A437B6EE8911D0016AD14CA1E14B">2022</mods:date>
<mods:detail id="D48B5D6E8F2DC1C91056088DCC745E90" type="pubDate">
<mods:number id="2066DFBC53F614B6A84F9484DC48B29A">2022-12-20</mods:number>
</mods:detail>
<mods:detail id="755D030C9C54B38BC194061557E11C65" type="volume">
<mods:number id="EBE1E5FA7A63EE5A260EA0BF711A5791">76</mods:number>
</mods:detail>
<mods:detail id="D3E5066110FB0444953D1487A3A4E780" type="issue">
<mods:number id="2F7C522E8ADA060AFDA74E677A87D94C">4</mods:number>
</mods:detail>
<mods:extent id="F9812FD36F6EB0E4586444D283AF90D5" unit="page">
<mods:start id="A3634101F3A5BD6BF0E2C38853FCDE22">503</mods:start>
<mods:end id="C56B8DA58F541B80376C995A8CA829C2">509</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="5503D840D233F979D908919868F6FFFA">
<mods:url id="D4D9180B2D784202BBC252347973A3EA">http://dx.doi.org/10.1649/0010-065x-76.4.503</mods:url>
</mods:location>
<mods:classification id="DAEF78932979716BAA768EEFD84A9CED">journal article</mods:classification>
<mods:identifier id="F8E4C9B64902406FEF9D0111506961E9" type="DOI">10.1649/0010-065X-76.4.503</mods:identifier>
<mods:identifier id="8B0DFE21AE30A797DAC83A72A7845F47" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="333431D26F032BBE30C1B9EA792995D7" type="Zenodo-Dep">13252795</mods:identifier>
<mods:identifier id="CBFFB52669FE3978D2799E6DD772F9D0" type="ZooBank">urn:lsid:zoobank.org:pub:7EBD9808-DC89-42FF-9923-555582F9F236</mods:identifier>
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<treatment id="131F116B4D73FFEBFF02F98429D30DCC" LSID="urn:lsid:plazi:treatment:131F116B4D73FFEBFF02F98429D30DCC" httpUri="http://treatment.plazi.org/id/131F116B4D73FFEBFF02F98429D30DCC" lastPageId="6" lastPageNumber="507" pageId="3" pageNumber="505">
<subSubSection id="D3ACF3F64D73FFEEFF02F9842A310E00" box="[227,605,1601,1623]" pageId="3" pageNumber="505" type="nomenclature">
<paragraph id="9B09A07D4D73FFEEFF02F9842A310E00" blockId="3.[227,605,1601,1623]" box="[227,605,1601,1623]" pageId="3" pageNumber="505">
<heading id="C04117114D73FFEEFF02F9842A310E00" bold="true" box="[227,605,1601,1623]" centered="true" fontSize="8" level="1" pageId="3" pageNumber="505" reason="1">
<taxonomicName id="5CB6DBFE4D73FFEEFF02F9842A310E00" authority="(DeGeer, 1774)" baseAuthorityName="DeGeer" baseAuthorityYear="1774" box="[227,605,1601,1623]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="A9C27C6F4D73FFEEFF02F9842A310E00" bold="true" box="[227,605,1601,1623]" pageId="3" pageNumber="505">
<emphasis id="A9C27C6F4D73FFEEFF02F98429D20E00" bold="true" box="[227,446,1602,1623]" italics="true" pageId="3" pageNumber="505">PHOTURIS pENSYLVANICA</emphasis>
(
<bibRefCitation id="FF27DD8C4D73FFEEFE2AF9872A3A0E00" author="DeGeer, C." box="[459,598,1601,1623]" pageId="3" pageNumber="505" refId="ref4145" refString="DeGeer, C. 1774. Memoires pour servir a l'histoire des insectes. Tome quatrieme. Pierre Hesselberg, Stockholm, xii + 456 pp. + 19 pls." type="book" year="1774">DeGeer, 1774</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D3ACF3F64D73FFEEFF4CF9AB2AFF0E91" pageId="3" pageNumber="505" type="reference_group">
<paragraph id="9B09A07D4D73FFEEFF4CF9AB2A0E0ED5" blockId="3.[173,610,1644,1666]" box="[173,610,1644,1666]" pageId="3" pageNumber="505">
<treatmentCitationGroup id="BBA687534D73FFEEFF4CF9AB2A0E0ED5" box="[173,610,1644,1666]" pageId="3" pageNumber="505">
<treatmentCitation id="1A17866C4D73FFEEFF4CF9AB2A310ED5" author="DeGeer, C." box="[173,605,1644,1666]" page="53" pageId="3" pageNumber="505" year="1774">
<taxonomicName id="5CB6DBFE4D73FFEEFF4CF9AB2A310ED5" authority="DeGeer 1774: 52 - 53" authorityName="DeGeer" authorityPageNumber="52 - 53" authorityYear="1774" box="[173,605,1644,1666]" class="Insecta" family="Lampyridae" genus="Lampyris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="A9C27C6F4D73FFEEFF4CF9AB29E70ED6" box="[173,395,1644,1666]" italics="true" pageId="3" pageNumber="505">Lampyris pensylvanica</emphasis>
<bibRefCitation id="FF27DD8C4D73FFEEFE70F9AA2A310ED5" author="DeGeer, C." box="[401,605,1644,1666]" pageId="3" pageNumber="505" refId="ref4145" refString="DeGeer, C. 1774. Memoires pour servir a l'histoire des insectes. Tome quatrieme. Pierre Hesselberg, Stockholm, xii + 456 pp. + 19 pls." type="book" year="1774">DeGeer 1774: 52–53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9B09A07D4D73FFEEFF24F9502AFF0E91" blockId="3.[173,659,1685,1734]" pageId="3" pageNumber="505">
<emphasis id="A9C27C6F4D73FFEEFF24F95029070EFC" bold="true" box="[197,363,1685,1707]" pageId="3" pageNumber="505">TYpe Specimens.</emphasis>
<materialsCitation id="2BDEAA204D73FFEEFE91F9502AFF0E91" location="National Museum of Natural History" pageId="3" pageNumber="505" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<typeStatus id="440D1EDF4D73FFEEFE91F95029AC0EFC" box="[368,448,1686,1707]" pageId="3" pageNumber="505" type="neotype">Neotype</typeStatus>
(
<specimenCount id="8DB06BF44D73FFEEFE2DF95029B30EFC" box="[460,479,1686,1707]" count="1" pageId="3" pageNumber="505" type="male">♂</specimenCount>
),
<emphasis id="A9C27C6F4D73FFEEFE10F9502AE00EFC" bold="true" box="[497,652,1685,1707]" pageId="3" pageNumber="505">here designated</emphasis>
, placed in the
<location id="9E69F6A64D73FFEEFECEF9762AFF0E91" LSID="urn:lsid:plazi:treatment:131F116B4D73FFEBFF02F98429D30DCC:9E69F6A64D73FFEEFECEF9762AFF0E91" box="[303,659,1712,1734]" name="National Museum of Natural History" pageId="3" pageNumber="505">National Museum of Natural History</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3ACF3F64D73FFEEFD22FF042C7E09C6" pageId="3" pageNumber="505" type="description">
<paragraph id="9B09A07D4D73FFEEFD22FF042C7E09C6" blockId="3.[707,1193,194,1736]" pageId="3" pageNumber="505">
(Smithsonian Institution),
<materialsCitation id="2BDEAA204D73FFEEFC27FF042BCA08A6" collectionCode="DC" country="United States of America" location="Washington" pageId="3" pageNumber="505" specimenCount="1" stateProvince="Washington">
<collectingRegion id="59726E9F4D73FFEEFC27FF042C57088F" box="[966,1083,194,216]" country="United States of America" name="Washington" pageId="3" pageNumber="505">Washington</collectingRegion>
,
<collectionCode id="FDA738B84D73FFEEFBA6FF042C070880" box="[1095,1131,194,215]" pageId="3" pageNumber="505">DC</collectionCode>
,
<collectingCountry id="E3A1E0ED4D73FFEEFB99FF042CC40880" box="[1144,1192,194,215]" name="United States of America" pageId="3" pageNumber="505">USA</collectingCountry>
(USNMENT01788002)
</materialsCitation>
:
<materialsCitation id="2BDEAA204D73FFEEFC4FFF1A2C6209C6" collectingDate="2015-07-02" collectionCode="NCCO, DET, COL" collectorName="C. M. Heckscher." country="United States of America" latitude="39.721954" location="Washington" longLatPrecision="1" longitude="-75.561745" pageId="3" pageNumber="505" specimenCount="6" stateProvince="Delaware">
“
<collectingCountry id="E3A1E0ED4D73FFEEFC56FF1A2B8708A6" box="[951,1003,220,241]" name="United States of America" pageId="3" pageNumber="505">USA</collectingCountry>
.
<collectingRegion id="59726E9F4D73FFEEFC13FF1A2C2008A5" box="[1010,1100,220,242]" country="United States of America" name="Delaware" pageId="3" pageNumber="505">Delaware</collectingRegion>
(
<collectionCode id="FDA738B84D73FFEEFBB6FF1A2CF208A6" box="[1111,1182,220,241]" pageId="3" pageNumber="505">NCCO</collectionCode>
). Wilmington Marsh.
<geoCoordinate id="FE82C6BA4D73FFEEFC61FF312B8B095A" box="[896,999,247,269]" degrees="39.721953" direction="north" orientation="latitude" pageId="3" pageNumber="505" precision="1" value="39.721954">39.721953</geoCoordinate>
,
<geoCoordinate id="FE82C6BA4D73FFEEFC11FF312C08095A" box="[1008,1124,247,269]" degrees="75.561746" direction="west" orientation="longitude" pageId="3" pageNumber="505" precision="1" value="-75.561745">–75.561746</geoCoordinate>
.
<date id="EF0886BD4D73FFEEFB8AFF312A990970" pageId="3" pageNumber="505" value="2015-07-02">
<collectingDate id="FF4C7F554D73FFEEFB8AFF312A990970" pageId="3" pageNumber="505" value="2015-07-02">2-VII-2015</collectingDate>
</date>
.
<collectionCode id="FDA738B84D73FFEEFD1DFED42B450970" box="[764,809,274,295]" pageId="3" pageNumber="505">DET</collectionCode>
&amp;
<collectionCode id="FDA738B84D73FFEEFCA4FED42B140970" box="[837,888,274,295]" country="Colombia" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="3" pageNumber="505" type="Herbarium">COL</collectionCode>
:
<collectorName id="3643C5AB4D73FFEEFC9EFED42C73097F" box="[895,1055,274,296]" pageId="3" pageNumber="505">C.M. Heckscher.</collectorName>
” An additional
<specimenCount id="8DB06BF44D73FFEEFD22FEEA2B240915" box="[707,840,300,322]" count="6" pageId="3" pageNumber="505" type="generic">six specimens</specimenCount>
collected from the same locality and on the same date are deposited in the CMH
<taxonomicName id="5CB6DBFE4D73FFEEFD22FEA42B58092F" authorityName="Rafinesque" authorityYear="1815" box="[707,820,354,376]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
collection currently housed at
<collectingRegion id="59726E9F4D73FFEEFBAFFEA42A9F09C6" country="United States of America" name="Delaware" pageId="3" pageNumber="505">Delaware State</collectingRegion>
University, Dover, DE,
<collectingCountry id="E3A1E0ED4D73FFEEFC3AFEBA2C6209C6" box="[987,1038,380,401]" name="United States of America" pageId="3" pageNumber="505">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D3ACF3F64D73FFE8FD3AFE512CF20BE0" lastPageId="5" lastPageNumber="507" pageId="3" pageNumber="505" type="diagnosis">
<paragraph id="9B09A07D4D73FFEEFD3AFE512B420D4A" blockId="3.[707,1193,194,1736]" pageId="3" pageNumber="505">
<emphasis id="A9C27C6F4D73FFEEFD3AFE512B2E09FA" bold="true" box="[731,834,407,429]" pageId="3" pageNumber="505">Diagnosis.</emphasis>
This is a pale and diminutive
<taxonomicName id="5CB6DBFE4D73FFEEFBB6FE512CC409FB" authorityName="Dejean" authorityYear="1833" box="[1111,1192,407,428]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="A9C27C6F4D73FFEEFBB6FE512CC409FB" box="[1111,1192,407,428]" italics="true" pageId="3" pageNumber="505">Photuris</emphasis>
</taxonomicName>
(
<figureCitation id="038DBCF84D73FFEEFD2BFE742B6F0990" box="[714,771,434,456]" captionStart="Fig" captionStartId="4.[176,207,1446,1465]" captionTargetBox="[152,1172,193,1419]" captionTargetId="figure-129@4.[152,1172,193,1419]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Photuris pensylvanica (neotype). Note pale bicolored elytra (anterior one-fourth darker than posterior three-fourths) with pale vittae fading posteriorly." figureDoi="http://doi.org/10.5281/zenodo.13252797" httpUri="https://zenodo.org/record/13252797/files/figure.png" pageId="3" pageNumber="505">Fig. 1</figureCitation>
) associated with freshwater tidal wetlands. Pronotal vitta has a wide base and apex creating an overall hourglass-shaped form. Pronotal orange pigmentation lacks red undertones. Elytra are structurally weak, often resulting in a noticeably curled or arched appearance after desiccation. Elytral tips are pointed, not broadly rounded, in comparison to some congeners. Elytral vittae engulf the humeri and fade posteriorly. Anterior one-fourth of elytra is often darker than posterior three-fourths. Scutellar shield and mesonotal plates are brown. Metacoxae are brown. Mean total length (
<emphasis id="A9C27C6F4D73FFEEFB82FD1E2C030ABA" box="[1123,1135,728,749]" italics="true" pageId="3" pageNumber="505">n</emphasis>
= 5):
<quantity id="5C4E0D984D73FFEEFD22FD372B780B50" box="[707,788,753,775]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="505" unit="mm" value="9.5">9.5 mm</quantity>
± 0.5 SD, range:
<quantity id="5C4E0D984D73FFEEFC3EFD342C330B50" box="[991,1119,754,776]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.9" metricValueMin="8.6" pageId="3" pageNumber="505" unit="mm" value="9.25" valueMax="9.9" valueMin="8.6">8.6–9.9 mm</quantity>
.
<emphasis id="A9C27C6F4D73FFEEFB8FFD342B790B76" bold="true" pageId="3" pageNumber="505">Flash pattern:</emphasis>
A two-part “dot-dash” emission consisting of a single short flash (~0.2 sec) immediately followed by a longer emission (~1.0–1.3 sec). Signal emitted while in flight with the dash typically given on a descent. The species is also thought to have an adjunct flash pattern of a single emission possibly given early in the evening, but this has been brought into question lately considering the confusion surrounding this taxon and the recent discovery of potentially sympatric single-flashing congeners (
<emphasis id="A9C27C6F4D73FFEEFD2AFBDE2A820C7A" box="[715,750,1048,1069]" italics="true" pageId="3" pageNumber="505">e.g.</emphasis>
,
<taxonomicName id="5CB6DBFE4D73FFEEFD18FBD12C200C7A" authority="Heckscher" authorityName="Heckscher" authorityYear="2013" box="[761,1100,1047,1069]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="species" species="mysticalampas">
<emphasis id="A9C27C6F4D73FFEEFD18FBD12B8C0C7B" box="[761,992,1047,1068]" italics="true" pageId="3" pageNumber="505">Photuris mysticalampas</emphasis>
Heckscher
</taxonomicName>
,
<taxonomicName id="5CB6DBFE4D73FFEEFBB4FBD12B0F0C1F" authority="Heckscher" authorityName="Heckscher" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="505" phylum="Arthropoda" rank="species" species="eliza">
<emphasis id="A9C27C6F4D73FFEEFBB4FBD12A9C0C10" italics="true" pageId="3" pageNumber="505">Photuris eliza</emphasis>
Heckscher
</taxonomicName>
).
<emphasis id="A9C27C6F4D73FFEEFC97FBF72BB70C10" bold="true" box="[886,987,1073,1095]" pageId="3" pageNumber="505">Behavior:</emphasis>
Adults emerge after sunset and usually fly low among or slightly above emergent vegetation in coastal freshwater tide marshes or emergent vegetation among tidal swamp forests. May also sparsely inhabit non-tidal freshwater emergent marshes in coastal regions. The “dot-dash” signal is typically given on a descent and sometimes during a tumbling fall through dense vegetation.
</paragraph>
<paragraph id="9B09A07D4D73FFE8FD3AFAE42CF20BE0" blockId="3.[707,1193,194,1736]" lastBlockId="5.[706,1194,877,1059]" lastPageId="5" lastPageNumber="507" pageId="3" pageNumber="505">
<emphasis id="A9C27C6F4D73FFEEFD3AFAE42C6C0D60" bold="true" box="[731,1024,1313,1335]" pageId="3" pageNumber="505">
<typeStatus id="440D1EDF4D73FFEEFD3AFAE42B5C0D60" box="[731,816,1314,1335]" pageId="3" pageNumber="505" type="neotype">NeotYpe</typeStatus>
Description. Head:
</emphasis>
Frons yellow or very slightly infuscate. Mandibles infuscate. Maxillary palpi gray. Clypeus amber, its apical edge black. Clypeal projections rather insignificant, middle projection most prominent. Clypeus bearing prominent setae. Vertex mostly yellow with an ill-defined broad brown crown.
<emphasis id="A9C27C6F4D73FFEEFBAEFA072CC40D81" bold="true" box="[1103,1192,1473,1494]" pageId="3" pageNumber="505">Antenna</emphasis>
(
<figureCitation id="038DBCF84D73FFEEFD2BFA1A2B7C0DA6" box="[714,784,1500,1522]" captionStart="Fig" captionStartId="4.[176,207,1446,1465]" captionTargetBox="[152,1172,193,1419]" captionTargetId="figure-129@4.[152,1172,193,1419]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Photuris pensylvanica (neotype). Note pale bicolored elytra (anterior one-fourth darker than posterior three-fourths) with pale vittae fading posteriorly." figureDoi="http://doi.org/10.5281/zenodo.13252797" httpUri="https://zenodo.org/record/13252797/files/figure.png" pageId="3" pageNumber="505">Figs. 1</figureCitation>
,
<figureCitation id="038DBCF84D73FFEEFCC1FA1A2B430DA6" box="[800,815,1500,1521]" captionStart="Fig" captionStartId="5.[197,228,769,788]" captionTargetBox="[173,1193,193,742]" captionTargetId="figure-313@5.[173,1193,193,742]" captionTargetPageId="5" captionText="Fig. 2. Lateral view of Photuris penyslvanica (neotype). Note long legs, brown metacoxa and comparatively short antenna relative to body length." figureDoi="http://doi.org/10.5281/zenodo.13252799" httpUri="https://zenodo.org/record/13252799/files/figure.png" pageId="3" pageNumber="505">2</figureCitation>
): Filiform. Brown with pale articulations. Rather short, ~0.45× length of body.
<emphasis id="A9C27C6F4D73FFEEFD22F9D72B0F0E70" bold="true" box="[707,867,1553,1575]" pageId="3" pageNumber="505">Scutellar shield</emphasis>
(
<figureCitation id="038DBCF84D73FFEEFC8EF9D42BC70E70" box="[879,939,1554,1576]" captionStart="Fig" captionStartId="5.[731,762,1634,1653]" captionTargetBox="[707,1193,1102,1607]" captionTargetId="figure-335@5.[707,1193,1102,1607]" captionTargetPageId="5" captionText="Fig. 3. Scutellar shield and pronotum of Photuris pensylvanica (neotype). Note brown scutellar shield and hourglass-shaped pronotal vitta with a diffuse apical tip and a rectangular base." figureDoi="http://doi.org/10.5281/zenodo.13252801" httpUri="https://zenodo.org/record/13252801/files/figure.png" pageId="3" pageNumber="505">Fig. 3</figureCitation>
): Scutellar shield and mesonotal plates brown.
<emphasis id="A9C27C6F4D73FFEEFC28F9EA2C5E0E16" bold="true" box="[969,1074,1580,1601]" pageId="3" pageNumber="505">Pronotum</emphasis>
(
<figureCitation id="038DBCF84D73FFEEFBAEF9EA2CF70E16" box="[1103,1179,1580,1602]" captionStart="Fig" captionStartId="5.[731,762,1634,1653]" captionTargetBox="[707,1193,1102,1607]" captionTargetId="figure-335@5.[707,1193,1102,1607]" captionTargetPageId="5" captionText="Fig. 3. Scutellar shield and pronotum of Photuris pensylvanica (neotype). Note brown scutellar shield and hourglass-shaped pronotal vitta with a diffuse apical tip and a rectangular base." figureDoi="http://doi.org/10.5281/zenodo.13252801" httpUri="https://zenodo.org/record/13252801/files/figure.png" pageId="3" pageNumber="505">Fig. 3</figureCitation>
): Pronotum semicircular. Pale margins broad and mostly translucent. Posterior margin of pronotum slightly sinuate, lateral and anterior margins round- ed. Surface evenly punctate and covered uniformly in rather dense setae; pubescence denser and longer toward hind margin. Posterior lateral corners rounded, only slightly produced. Orange pigmentation circular, a rich dark orange without deep red undertones, with broad black central vitta. Vitta with apical tip broadly rounded with sharp demarcation toward outer edge continuous with edge of orange pigmentation and a protruding diffuse terminal tip nearly reaching terminal edge of pronotum. Shaft slightly constricted, serifs and flukes more diffuse. Serifs and flukes not well produced; rather, base of vitta broad, more rectangular in shape than anchor-like. Thus, vitta not arrowlike, more hourglass shaped. Total length of pronotum: 2.0 mm. Length along basal edge between two posterior corners:
<quantity id="5C4E0D984D74FFE9FC74F95E2B8A0EFA" box="[917,998,1688,1710]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="506" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="A9C27C6F4D74FFE9FC12F95E2C5A0EFA" bold="true" box="[1011,1078,1688,1709]" pageId="4" pageNumber="506">ElYtra</emphasis>
(
<figureCitation id="038DBCF84D74FFE9FBA7F95E2CE80EFA" box="[1094,1156,1688,1710]" captionStart="Fig" captionStartId="4.[176,207,1446,1465]" captionTargetBox="[152,1172,193,1419]" captionTargetId="figure-129@4.[152,1172,193,1419]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Photuris pensylvanica (neotype). Note pale bicolored elytra (anterior one-fourth darker than posterior three-fourths) with pale vittae fading posteriorly." figureDoi="http://doi.org/10.5281/zenodo.13252797" httpUri="https://zenodo.org/record/13252797/files/figure.png" pageId="4" pageNumber="506">Fig. 1</figureCitation>
): Pale brown with anterior one-fourth darker than posterior three-fourths. Vittae pale, the distal three-fourths almost imperceptible and extending about two-thirds length of elytra. Vittae engulf the humerus. Lateral margins somewhat broad, not reaching the apical tips, instead fading distally. Inner sutural margins quite pale, reaching both the apical tips and scutellar shield. Apical tips noticeably narrow and pointed. Total length:
<quantity id="5C4E0D984D75FFE8FDE7FBEE2A3A0C6A" box="[518,598,1064,1085]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="5" pageNumber="507" unit="mm" value="7.8">7.8 mm</quantity>
. Total width at one-half length of elytra:
<quantity id="5C4E0D984D75FFE8FE0FFB842A540C0F" box="[494,568,1090,1112]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="507" unit="mm" value="3.9">3.9 mm</quantity>
.
<emphasis id="A9C27C6F4D75FFE8FDDEFB842AFE0C00" bold="true" box="[575,658,1090,1111]" pageId="5" pageNumber="507">Thorax:</emphasis>
Meso- and metaventrites brown.
<emphasis id="A9C27C6F4D75FFE8FE17FB9B2A370C25" bold="true" box="[502,603,1117,1138]" pageId="5" pageNumber="507">Abdomen</emphasis>
(
<figureCitation id="038DBCF84D75FFE8FD8AFB9B28D00CDA" captionStart="Fig" captionStartId="6.[176,207,1049,1068]" captionTargetBox="[152,1172,193,1022]" captionTargetId="figure-326@6.[152,1172,193,1022]" captionTargetPageId="6" captionText="Fig. 4. Ventral view of Photuris pensylvanica (neotype). Note brown metacoxa and pale trochanters." figureDoi="http://doi.org/10.5281/zenodo.13252803" httpUri="https://zenodo.org/record/13252803/files/figure.png" pageId="5" pageNumber="507">Fig. 4</figureCitation>
): First 3 visible ventrites (ventrites 1–3 or true abdominal segments II–IV; see Branham and Archangelsky [2000] for discussion regarding visible ventrite versus true abdominal segment number in
<taxonomicName id="5CB6DBFE4D75FFE8FF0BFB24290E0CAF" authorityName="Rafinesque" authorityYear="1815" box="[234,354,1250,1272]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="507" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
) brown, 4 (V) largely yellow, 5 (VI) and 6 (VII) yellow, 7 (VIII) pale and enclosing the aedeagus. Ventrite 6 (VII) only slightly emarginated.
<emphasis id="A9C27C6F4D75FFE8FEE4FAF4295A0D10" bold="true" box="[261,310,1330,1351]" pageId="5" pageNumber="507">Legs</emphasis>
(
<figureCitation id="038DBCF84D75FFE8FEA6FAF429E30D10" box="[327,399,1330,1352]" captionStart="Fig" captionStartId="5.[197,228,769,788]" captionTargetBox="[173,1193,193,742]" captionTargetId="figure-313@5.[173,1193,193,742]" captionTargetPageId="5" captionText="Fig. 2. Lateral view of Photuris penyslvanica (neotype). Note long legs, brown metacoxa and comparatively short antenna relative to body length." figureDoi="http://doi.org/10.5281/zenodo.13252799" httpUri="https://zenodo.org/record/13252799/files/figure.png" pageId="5" pageNumber="507">Figs. 2</figureCitation>
,
<figureCitation id="038DBCF84D75FFE8FE7CFAF429C00D10" box="[413,428,1330,1351]" captionStart="Fig" captionStartId="6.[176,207,1049,1068]" captionTargetBox="[152,1172,193,1022]" captionTargetId="figure-326@6.[152,1172,193,1022]" captionTargetPageId="6" captionText="Fig. 4. Ventral view of Photuris pensylvanica (neotype). Note brown metacoxa and pale trochanters." figureDoi="http://doi.org/10.5281/zenodo.13252803" httpUri="https://zenodo.org/record/13252803/files/figure.png" pageId="5" pageNumber="507">4</figureCitation>
): Hind legs long for a
<taxonomicName id="5CB6DBFE4D75FFE8FF4CFA8B296D0D35" authorityName="Dejean" authorityYear="1833" box="[173,257,1357,1378]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="A9C27C6F4D75FFE8FF4CFA8B296D0D35" box="[173,257,1357,1378]" italics="true" pageId="5" pageNumber="507">Photuris</emphasis>
</taxonomicName>
, ~0.80× length of body. Procoxa and mesocoxa pale yellow with some brown shading laterally. Metacoxa entirely brown. Trochanters pale yellow. Profemur with basal three-fourths pale, apical one-fourth brown. Meso- and metafemur pale yellow. Protibia mostly brown; meso- and metatibia largely yellow with slight infuscate areas bearing dark brown setae yielding an overall dusted appearance. Meso- and metatibia with two apical spurs of near equal length. Tarsi pale yellow with brown apical tips and blond setae. Claws pale yellow with amber points, anterior claws (of each pair) bifid, posterior claws simple.
<emphasis id="A9C27C6F4D75FFE8FDC7F94B2AFE0EF4" bold="true" box="[550,658,1677,1699]" pageId="5" pageNumber="507">Aedeagus:</emphasis>
Typical for a
<taxonomicName id="5CB6DBFE4D75FFE8FEB7F96E29C60EEA" authorityName="Dejean" authorityYear="1833" box="[342,426,1704,1725]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="A9C27C6F4D75FFE8FEB7F96E29C60EEA" box="[342,426,1704,1725]" italics="true" pageId="5" pageNumber="507">Photuris</emphasis>
</taxonomicName>
with no recognized diagnostic characters; consisting of a basal piece, two lateral lobes, and a central medial lobe with two lateral filaments (
<emphasis id="A9C27C6F4D75FFE8FC8DFC642BE90BE0" box="[876,901,930,951]" italics="true" pageId="5" pageNumber="507">cf.</emphasis>
<bibRefCitation id="FF27DD8C4D75FFE8FC68FC642CFF0BE0" author="LloYd, J. E. &amp; L. A. BallantYne" box="[905,1171,930,952]" pageId="5" pageNumber="507" pagination="464 - 473" refId="ref4527" refString="LloYd, J. E., and L. A. BallantYne. 2003. Taxonomy and behavior of Photuris trivittata sp. n. (Coleoptera: Lampyridae: Cratomorphini). Florida Entomologist 86: 464 - 473." type="journal article" year="2003">Lloyd and Ballantyne 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="CFC9F0F54D74FFE9FF51FA602AE60D86" ID-DOI="http://doi.org/10.5281/zenodo.13252797" ID-Zenodo-Dep="13252797" httpUri="https://zenodo.org/record/13252797/files/figure.png" pageId="4" pageNumber="506" startId="4.[176,207,1446,1465]" targetBox="[152,1172,193,1419]" targetPageId="4" targetType="figure">
<paragraph id="9B09A07D4D74FFE9FF51FA602AE60D86" blockId="4.[152,1171,1446,1489]" pageId="4" pageNumber="506">
<emphasis id="A9C27C6F4D74FFE9FF51FA6028870DEE" bold="true" box="[176,235,1446,1465]" pageId="4" pageNumber="506">Fig. 1.</emphasis>
Habitus of
<taxonomicName id="5CB6DBFE4D74FFE9FE84FA602A490DEE" box="[357,549,1446,1465]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="506" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="A9C27C6F4D74FFE9FE84FA602A490DEE" box="[357,549,1446,1465]" italics="true" pageId="4" pageNumber="506">Photuris pensylvanica</emphasis>
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(neotype). Note pale bicolored elytra (anterior one-fourth darker than posterior three-fourths) with pale vittae fading posteriorly.
</paragraph>
</caption>
<caption id="CFC9F0F54D75FFE8FF24FCC729D00B7B" ID-DOI="http://doi.org/10.5281/zenodo.13252799" ID-Zenodo-Dep="13252799" httpUri="https://zenodo.org/record/13252799/files/figure.png" pageId="5" pageNumber="507" startId="5.[197,228,769,788]" targetBox="[173,1193,193,742]" targetPageId="5" targetType="figure">
<paragraph id="9B09A07D4D75FFE8FF24FCC729D00B7B" blockId="5.[173,1192,769,813]" pageId="5" pageNumber="507">
<emphasis id="A9C27C6F4D75FFE8FF24FCC728920B43" bold="true" box="[197,254,769,788]" pageId="5" pageNumber="507">Fig. 2.</emphasis>
Lateral view of
<taxonomicName id="5CB6DBFE4D75FFE8FE7DFCC72A360B43" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[412,602,769,788]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="507" phylum="Arthropoda" rank="species" species="penyslvanica">
<emphasis id="A9C27C6F4D75FFE8FE7DFCC72A360B43" box="[412,602,769,788]" italics="true" pageId="5" pageNumber="507">Photuris penyslvanica</emphasis>
</taxonomicName>
(neotype). Note long legs, brown metacoxa and comparatively short antenna relative to body length.
</paragraph>
</caption>
<subSubSection id="D3ACF3F64D75FFEBFD3AFC7B29D30DCC" lastPageId="6" lastPageNumber="508" pageId="5" pageNumber="507" type="description">
<paragraph id="9B09A07D4D75FFEBFD3AFC7B29D30DCC" blockId="5.[706,1194,877,1059]" lastBlockId="6.[151,639,1120,1436]" lastPageId="6" lastPageNumber="508" pageId="5" pageNumber="507">
<emphasis id="A9C27C6F4D75FFE8FD3AFC7B2C770B85" bold="true" box="[731,1051,957,979]" pageId="5" pageNumber="507">
Variation Including
<typeStatus id="440D1EDF4D75FFE8FC4AFC7B2B920B85" box="[939,1022,957,978]" pageId="5" pageNumber="507" type="neotype">NeotYpe</typeStatus>
(
<emphasis id="A9C27C6F4D75FFE8FBEFFC7B2C770B85" bold="true" box="[1038,1051,957,978]" italics="true" pageId="5" pageNumber="507">N</emphasis>
</emphasis>
=
<emphasis id="A9C27C6F4D75FFE8FBD8FC7B2C3E0B84" bold="true" box="[1081,1106,957,979]" pageId="5" pageNumber="507">5).</emphasis>
Clypeus varies from amber to yellow. Antenna varies from ~0.35–0.55 length of body.Total length of pronotum:
<quantity id="5C4E0D984D75FFE8FD23FBCB2B660C75" box="[706,778,1037,1058]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="507" unit="mm" value="1.9">1.9 mm</quantity>
± 0.2 SD, range:
<quantity id="5C4E0D984D75FFE8FC4FFBCB2C4F0C75" box="[942,1059,1037,1059]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="5" pageNumber="507" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.6–2.1 mm</quantity>
. Length along basal edge between two posterior corners:
<quantity id="5C4E0D984D76FFEBFDD2FBA62A110C21" box="[563,637,1120,1142]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="508" unit="mm" value="2.4">2.4 mm</quantity>
± 0.1 SD, range:
<quantity id="5C4E0D984D76FFEBFECDFBBD29F60CC7" box="[300,410,1147,1169]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="6" pageNumber="508" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
. Posterior broadened area of vitta varies in width relative to shaft but always more rectangular than anchor-shaped. Although most individuals are slender in appearance due to pointed apical tips, variation in width of lateral margins result in some specimens appearing oval in outline from above (large length-to-width ratio). Elytra in some specimens are somewhat translucent. Elytra total length:
<quantity id="5C4E0D984D76FFEBFEA9FA9629E20D31" box="[328,398,1360,1382]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="508" unit="mm" value="7.4">7.4 mm</quantity>
± 0.36 SD, range:
<quantity id="5C4E0D984D76FFEBFDD6FA9628AC0DD7" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.8" metricValueMin="6.9" pageId="6" pageNumber="508" unit="mm" value="7.35" valueMax="7.8" valueMin="6.9">6.9–7.8 mm</quantity>
. Total width at one-half length of elytra:
<quantity id="5C4E0D984D76FFEBFDD7FAAD2A110DD7" box="[566,637,1387,1408]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="508" unit="mm" value="3.8">3.8 mm</quantity>
± 0.18 SD, range: 3.6–4.0 mm.
</paragraph>
</subSubSection>
</treatment>
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206
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(
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)
<emphasis id="8CE2EAB8E051A770FE19FBFFFE27EF02" bold="true" box="[416,481,1057,1079]" italics="true" pageId="2" pageNumber="578">milleri</emphasis>
Ivie, Fanti, and Ferreira
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,
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<emphasis id="8CE2EAB8E051A770FF09FB69FE8AEFF8" bold="true" box="[176,332,1207,1229]" pageId="2" pageNumber="578">Type Specimen.</emphasis>
<materialsCitation id="0EFE3CF7E051A770FEE8FB69FE30EFD2" collectionCode="WIBF" pageId="2" pageNumber="578" specimenCode="WIBF 049821" specimenCount="1" typeStatus="holotype">
<typeStatus id="612D8808E051A770FEE8FB69FE6AEFF8" box="[337,428,1207,1229]" pageId="2" pageNumber="578" type="holotype">Holotype</typeStatus>
: female, in Dominican amber, accession no.
<specimenCode id="EE309ED1E051A770FEDDFB0CFE37EFD2" box="[356,497,1234,1255]" collectionCode="WIBF" country="USA" httpUri="http://grbio.org/cool/mzn7-y8yj" name="West Indian Beetle Fauna Project Collection" pageId="2" pageNumber="578">WIBF 049821</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="BE2936AAE051A770FF09FB32FED9EE28" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
<materialsCitation id="0EFE3CF7E051A770FF09FB32FEDDEE28" country="Dominican Republic" location="Cordillera Septentrional" pageId="2" pageNumber="578" specimenCount="1" typeStatus="holotype">
<emphasis id="8CE2EAB8E051A770FF09FB32FE87EE37" bold="true" box="[176,321,1260,1282]" pageId="2" pageNumber="578">
<typeStatus id="612D8808E051A770FF09FB32FF24EE34" box="[176,226,1260,1281]" pageId="2" pageNumber="578">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="C681763AE051A770FEFEFB32FDD4EE37" box="[327,530,1260,1282]" name="Dominican Republic" pageId="2" pageNumber="578">Dominican Republic</collectingCountry>
:
<location id="BB496071E051A770FDA5FB32FEDDEE28" LSID="urn:lsid:plazi:treatment:363F87BCE051A776FF07FBFCFDECEA4D:BB496071E051A770FDA5FB32FEDDEE28" country="Dominican Republic" name="Cordillera Septentrional" pageId="2" pageNumber="578">Cordillera Septentrional</location>
</materialsCitation>
.
</paragraph>
<paragraph id="BE2936AAE051A770FF09FAFCFE6CEE58" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
<emphasis id="8CE2EAB8E051A770FF09FAFCFE8DEE02" bold="true" box="[176,331,1313,1335]" pageId="2" pageNumber="578">
<typeStatus id="612D8808E051A770FF09FAFCFF24EE02" box="[176,226,1314,1335]" pageId="2" pageNumber="578">Type</typeStatus>
Horizon.
</emphasis>
Late Early/Middle Miocene; 15–20 million years ago (mya), probably close to 16 mya. La Toca Formation.
</paragraph>
</subSubSection>
<subSubSection id="F68C6521E051A770FF09FAACFDD0ED38" pageId="2" pageNumber="578" type="etymology">
<paragraph id="BE2936AAE051A770FF09FAACFDD0ED38" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
<emphasis id="8CE2EAB8E051A770FF09FAACFEE5EEB2" bold="true" box="[176,291,1394,1415]" pageId="2" pageNumber="578">Etymology.</emphasis>
The species is named after the late Richard Stuart Miller (1945–2021). Rich’s interests were mainly focused on “cantharoids”, most strongly on
<taxonomicName id="79964D29E051A770FF0CFA1CFEC7EEED" box="[181,257,1474,1496]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="family">Lycidae</taxonomicName>
and
<taxonomicName id="79964D29E051A770FE96FA1CFE60EEED" authorityName="Rafinesque" authorityYear="1815" box="[303,422,1474,1496]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
, and his extensive collection was donated to the
<collectingRegion id="7C52F848E051A770FE11FA03FE3BEEC7" box="[424,509,1501,1522]" country="United States of America" name="Montana" pageId="2" pageNumber="578">Montana</collectingRegion>
Entomology Collection at
<collectingRegion id="7C52F848E051A770FEA0FA29FE63ED39" box="[281,421,1527,1548]" country="United States of America" name="Montana" pageId="2" pageNumber="578">Montana State</collectingRegion>
University.
</paragraph>
</subSubSection>
<subSubSection id="F68C6521E051A770FF09F9CFFBE4E9B7" pageId="2" pageNumber="578" type="diagnosis">
<paragraph id="BE2936AAE051A770FF09F9CFFBE4E9B7" blockId="2.[152,638,1207,1709]" lastBlockId="2.[685,1173,194,1709]" pageId="2" pageNumber="578">
<emphasis id="8CE2EAB8E051A770FF09F9CFFEDFED12" bold="true" box="[176,281,1553,1575]" pageId="2" pageNumber="578">Diagnosis.</emphasis>
The generic placement of the species is discussed below. Currently, four other species of short-elytra cantharids are known from Dominican amber, two of which have been described:
<taxonomicName id="79964D29E051A770FD9EF9BCFDB8ED42" authorityName="Poinar and Fanti" authorityYear="2016" box="[551,638,1634,1655]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis id="8CE2EAB8E051A770FD9EF9BCFDB8ED42" box="[551,638,1634,1655]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
</taxonomicName>
and
<taxonomicName id="79964D29E051A770FF7DF9A3FEDCEDA4" authorityName="Fanti and Damgaard" authorityYear="2019" box="[196,282,1660,1682]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
<emphasis id="8CE2EAB8E051A770FF7DF9A3FEDCEDA4" box="[196,282,1660,1682]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
</taxonomicName>
.
<taxonomicName id="79964D29E051A770FE93F9A2FE09EDA4" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[298,463,1660,1681]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8CE2EAB8E051A770FE93F9A2FE09EDA4" box="[298,463,1660,1681]" italics="true" pageId="2" pageNumber="578">Tytthonyx milleri</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="79964D29E051A770FF6AF946FEEBED99" authorityName="Fanti and Damgaard" authorityYear="2019" box="[211,301,1687,1709]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
<emphasis id="8CE2EAB8E051A770FF6AF946FEEBED99" box="[211,301,1687,1709]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
</taxonomicName>
by its smaller size (
<quantity id="796E9B4FE051A770FDB4F949FD98ED99" box="[525,606,1687,1708]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="578" unit="mm" value="2.8">2.8 mm</quantity>
in
<taxonomicName id="79964D29E051A770FD14FF1CFCC3EBE2" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[685,773,194,215]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8CE2EAB8E051A770FD14FF1CFCC3EBE2" box="[685,773,194,215]" italics="true" pageId="2" pageNumber="578">T. milleri</emphasis>
</taxonomicName>
and 6.0 mm in
<taxonomicName id="79964D29E051A770FC24FF1CFC37EBE2" authorityName="Fanti and Damgaard" authorityYear="2019" box="[925,1009,194,215]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
<emphasis id="8CE2EAB8E051A770FC24FF1CFC37EBE2" box="[925,1009,194,215]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
</taxonomicName>
) and the flavous color (
<taxonomicName id="79964D29E051A770FD55FF03FC87EBC4" authorityName="Fanti and Damgaard" authorityYear="2019" box="[748,833,220,242]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
<emphasis id="8CE2EAB8E051A770FD55FF03FD3FEBC7" box="[748,761,221,242]" italics="true" pageId="2" pageNumber="578">T</emphasis>
.
<emphasis id="8CE2EAB8E051A770FCBCFF02FC87EBC4" box="[773,833,220,241]" italics="true" pageId="2" pageNumber="578">stadili</emphasis>
</taxonomicName>
has a black face, elytra, venter, and basal eight antennomeres).
<taxonomicName id="79964D29E051A770FC78FF29FBAEEA39" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[961,1128,247,268]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8CE2EAB8E051A770FC78FF29FBAEEA39" box="[961,1128,247,268]" italics="true" pageId="2" pageNumber="578">Tytthonyx milleri</emphasis>
</taxonomicName>
has somewhat stronger serration of antennomeres 3–5 and a convex head (almost flat in
<taxonomicName id="79964D29E051A770FC51FEF3FBFCEA74" authorityName="Fanti and Damgaard" authorityYear="2019" box="[1000,1082,300,322]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
<emphasis id="8CE2EAB8E051A770FC51FEF3FBFCEA74" box="[1000,1082,300,322]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
</taxonomicName>
although this character state is possibly an artifact of preservation) (Fanti and Damgaard 2019). From
<taxonomicName id="79964D29E051A770FB8EFEBCFB4BEA42" authorityName="Poinar and Fanti" authorityYear="2016" box="[1079,1165,354,375]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis id="8CE2EAB8E051A770FB8EFEBCFB4BEA42" box="[1079,1165,354,375]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
</taxonomicName>
, which is similar in size, it is quite different (see Discussion below). The easiest way to distinguish them is by the length of the antenna, which is as long as the body in
<taxonomicName id="79964D29E051A770FCCBFE13FC0BEAD4" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[882,973,460,482]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8CE2EAB8E051A770FCCBFE13FCB9EAD7" box="[882,895,461,482]" italics="true" pageId="2" pageNumber="578">T</emphasis>
.
<emphasis id="8CE2EAB8E051A770FC34FE12FC0BEAD4" box="[909,973,460,481]" italics="true" pageId="2" pageNumber="578">milleri</emphasis>
</taxonomicName>
and longer than the body in
<taxonomicName id="79964D29E051A770FD40FE36FC95EAC8" authorityName="Poinar and Fanti" authorityYear="2016" box="[761,851,487,509]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis id="8CE2EAB8E051A770FD40FE36FCC0EAC8" box="[761,774,488,509]" italics="true" pageId="2" pageNumber="578">T</emphasis>
.
<emphasis id="8CE2EAB8E051A770FCABFE36FC95EAC8" box="[786,851,488,509]" italics="true" pageId="2" pageNumber="578">geiseri</emphasis>
</taxonomicName>
. The form of the pronotum is also distinctly different, being more rounded anterolaterally (square in
<taxonomicName id="79964D29E051A770FCF3FDC3FC67E907" authorityName="Poinar and Fanti" authorityYear="2016" box="[842,929,541,562]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
<emphasis id="8CE2EAB8E051A770FCF3FDC3FC67E907" box="[842,929,541,562]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
</taxonomicName>
) (Poinar and Fanti 2016). These characters are all potentially somewhat relat- ed to the different sexes of the
<typeStatus id="612D8808E051A770FC74FD8CFBC4E952" box="[973,1026,594,615]" pageId="2" pageNumber="578">types</typeStatus>
, but exceed the variation seen in most modern species.
</paragraph>
</subSubSection>
<subSubSection id="F68C6521E051A776FD7CFD59FDECEA4D" lastPageId="4" lastPageNumber="580" pageId="2" pageNumber="578" type="description">
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<emphasis id="8CE2EAB8E051A770FD7CFD59FC84E9A8" bold="true" box="[709,834,647,669]" pageId="2" pageNumber="578">Description.</emphasis>
Female (based on the wide last abdominal ventrite), adult, alate, entirely flavous. Body length
<emphasis id="8CE2EAB8E051A770FC9FFD63FC85E9E7" box="[806,835,701,722]" italics="true" pageId="2" pageNumber="578">ca.</emphasis>
<quantity id="796E9B4FE051A770FCFEFD62FC54E9E7" box="[839,914,700,722]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="578" unit="mm" value="2.8">2.8 mm</quantity>
. Head completely exposed, elongate, narrowed behind the eyes, convex medially, pubescent and shallowly punctate. Eyes round, inserted laterally. Frontoclypeal region (face) constricted by antennal insertions near front of eyes, with a transverse depression between antennal insertions and a widened frontal margin, frontal margin nearly straight, slightly concave. Mandibles long, broad basal to the median strong retinaculum, thereafter long, thin, falciform. Maxillary palp 4-segmented; first palpomere robust, short; second elongate, cylindrical; third robust, intermediate in length between first two; fourth globular, apically pointed. Labial palp 3-segmented; terminal palpomere globular and apically pointed.Antenna with 11 antennomeres, serrate; relatively short, reaching
<emphasis id="8CE2EAB8E051A770FD14FBB6FD0DEF48" box="[685,715,1128,1149]" italics="true" pageId="2" pageNumber="578">ca.</emphasis>
midpoint of abdomen; scape stout, club-shaped; antennomere 2 filiform, about two-thirds length of first; antennomeres 3–8 (especially 4–7) serrate, subequal in length; antennomere 9 slightly serrate; antennomeres 10–11 filiform with terminal antennomere rounded at apex; all antennomeres pubescent. Pronotum transverse, surface slightly convex, pubescent, shallowly punctate, slightly wider than head, corners round, anterior and posterior margins and lateral beads strongly bordered, sides irregular and slightly enlarged near the anterior margin, lacking secretory pore. Elytra very short, reaching base of first ventrite; slightly dehiscent apically, apices round; setae short. Hind wings fully developed, surpassing last abdominal tergite; costal margin with series of dark bristles; radial cell open; r-m connected at both ends; Mr apparently narrowly erased at base; M
<subScript id="221234EFE051A770FCBCF9E6FCDBED73" attach="left" box="[773,797,1592,1606]" fontSize="6" pageId="2" pageNumber="578">3+4</subScript>
complete; cubital vein not visible. Ventrites transverse, pubescent, last ventrite wide. Legs long, slender; pro- and mesocoxae conical; trochanters elongate and offset, rounded apically; femora and tibiae cylindrical, straight; with small, fine apical spur on pro- and metatibiae, slightly stouter on mesotibia. Tarsal formula 5-5-5; metatarsus with first tarsomere elongate and thin, the second tarsomere about one-third length of first, third tarsomere small, fourth very short and strongly bilobed, fifth tarsomere slender, subequal to second; claws simple and minute. Male unknown.
</paragraph>
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</treatment>
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83
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@ -1,55 +1,56 @@
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. Dorsal color black to dark brown, except clypeus yellow to pale brown, anterolateral angles of pronotum orange to pale brown, and apical two-thirds of elytra yellow, orange or pale brown (
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<figureCitation id="C7C92A6AA605FFEF4ECDFAE780BC1E6A" box="[958,984,1298,1319]" captionStart="Figs" captionStartId="9.[197,236,1617,1636]" captionTargetBox="[201,621,202,1592]" captionTargetId="figure-216@9.[185,632,195,1606]" captionTargetPageId="9" captionText="Figs. 29–32. Pygidia with attached sternite VIII of male Microasydates, new genus in glycerol. 29) M. punctipennis; 30) M. sanclemente, new species; 31) M. santabarbara, new species; 32) M. umbratus. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252827" httpUri="https://zenodo.org/record/13252827/files/figure.png" pageId="19" pageNumber="555">30</figureCitation>
,
<figureCitation id="C7C92A6AA605FFEF4E91FAE780981E6A" box="[994,1020,1298,1319]" captionStart="Figs" captionStartId="11.[197,238,465,484]" captionTargetBox="[176,1196,185,447]" captionTargetId="figure-626@11.[176,1196,185,447]" captionTargetPageId="11" captionText="Figs. 38–41. Female pygidia of Microasydates, new genus showing pubescence. 38) M. punctipennis;" figureDoi="http://doi.org/10.5281/zenodo.13252831" httpUri="https://zenodo.org/record/13252831/files/figure.png" pageId="19" pageNumber="555">39</figureCitation>
,
<figureCitation id="C7C92A6AA605FFEF4975FAE787201E6A" box="[1030,1092,1297,1319]" captionStart="Figs" captionStartId="12.[176,215,1616,1635]" captionTargetBox="[192,1131,211,1589]" captionTargetId="figure-7@12.[192,1131,192,1600]" captionTargetPageId="12" captionText="Figs. 42–53. Median lobe (lateral view) and tegmen (ventral view) for male Microasydates, new genus in glycerol. M. punctipennis: 42) Median lobe; 43) Tegmen. M. sanclemente, new species: 44) Median lobe; 45) Tegmen. M. santabarbara, new species, specimen from Santa Ynez Mountains, CA: 46) Median lobe; 47) Tegmen. M. santabarbara, new species, specimen from Santa Cruz Island, CA: 48) Median lobe; 49) Tegmen. M. umbratus, specimen from Shasta Co., CA: 50) Median lobe; 51) Tegmen. M. umbratus, specimen from Juab Co., UT: 52) Median lobe; 53) Tegmen. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252833" httpUri="https://zenodo.org/record/13252833/files/figure.png" pageId="19" pageNumber="555">44, 45</figureCitation>
,
<figureCitation id="C7C92A6AA605FFEF493DFAE7870C1E65" box="[1102,1128,1298,1320]" captionStart="Figs" captionStartId="13.[197,236,1367,1386]" captionTargetBox="[173,1193,893,1350]" captionTargetId="figure-87@13.[173,1193,893,1350]" captionTargetPageId="13" captionText="Figs. 62–63. Distribution maps for species of Microasydates, new genus. 62) M. punctipennis (green squares), M. sanclemente, new species (red triangles), and M. santabarbara, new species (blue circles); 63) M. umbratus (black circles)." figureDoi="http://doi.org/10.5281/zenodo.13252837" httpUri="https://zenodo.org/record/13252837/files/figure.png" pageId="19" pageNumber="555">62</figureCitation>
,
<figureCitation id="C7C92A6AA605FFEF4901FAE787EF1E65" box="[1138,1163,1298,1320]" captionStart="Fig" captionStartId="13.[198,229,1695,1714]" captionTargetBox="[221,419,1522,1651]" captionTargetId="graphics-126@13.[221,419,1522,1652]" captionTargetPageId="13" captionText="Fig. 64. Seasonalities of adult Microasydates, new genus species by month, based on collection records in the Specimens Examined sections." figureDoi="http://doi.org/10.5281/zenodo.13252839" httpUri="https://zenodo.org/record/13252839/files/figure.png" pageId="19" pageNumber="555">64</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="17E86564A605FFEF4FA8FAC9873E1EBF" pageId="19" pageNumber="555" type="materials_examined">
<paragraph id="5F4D36EFA605FFEF4FA8FAC9873E1EBF" blockId="19.[706,1194,1340,1736]" pageId="19" pageNumber="555">
<emphasis id="6D86EAFDA605FFEF4FA8FAC980E01E1F" bold="true" box="[731,900,1340,1362]" pageId="19" pageNumber="555">Type Specimen.</emphasis>
<materialsCitation id="EF9A3CB2A605FFEF4EE4FAC9873E1EBF" collectionCode="SBMNH" county="Los Angeles Co." location="San Clemente Isl." municipality="Cave Cyn." pageId="19" pageNumber="555" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="8049884DA605FFEF4EE4FAC980891E1F" box="[919,1005,1340,1362]" pageId="19" pageNumber="555" type="holotype">Holotype</typeStatus>
<specimenCount id="49F4FD66A605FFEF4973FAC887771E1F" box="[1024,1043,1341,1362]" count="1" pageId="19" pageNumber="555" type="male">♂</specimenCount>
, deposited in
<collectionCode id="39E3AE2AA605FFEF4FB0FAA280721E21" box="[707,790,1367,1388]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="19" pageNumber="555" type="Museum">SBMNH</collectionCode>
, labeled “CA:
<collectingCounty id="B62C4E63A605FFEF4EE9FAA287541E21" box="[922,1072,1367,1389]" pageId="19" pageNumber="555">Los Angeles Co.</collectingCounty>
\ 32.8734°N, 118.4480°W \
<location id="5A2D6034A605FFEF4E39FA8780941EC5" LSID="urn:lsid:plazi:treatment:D75B87F9A605FFE94FACFB5182D41A0F:5A2D6034A605FFEF4E39FA8780941EC5" box="[842,1008,1394,1416]" county="Los Angeles Co." municipality="Cave Cyn." name="San Clemente Isl." pageId="19" pageNumber="555">San Clemente Isl.</location>
, upper \
<collectingMunicipality id="BF29AC95A605FFEF4933FA8787C21ECA" box="[1088,1190,1394,1415]" pageId="19" pageNumber="555">Cave Cyn.</collectingMunicipality>
, vi.15.2008 \ Caterino&amp;Chatzimanolis [white printed label] // CA BEETLE PROJ \CBP0079731 [blue printed label]//
<typeStatus id="8049884DA605FFEF4E77FA37801E1E9A" box="[772,890,1474,1495]" pageId="19" pageNumber="555" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="49F4FD66A605FFEF4E0EFA3780EB1E9A" box="[893,911,1474,1495]" count="1" pageId="19" pageNumber="555" type="male">♂</specimenCount>
\
<taxonomicName id="98F24D6CA605FFEF4EEFFA3787441E9A" authorityName="Gimmel &amp; Mayor" authorityYear="2022" box="[924,1056,1474,1495]" class="Insecta" family="Melyridae" genus="Microasydates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="555" phylum="Arthropoda" rank="genus">
<emphasis id="6D86EAFDA605FFEF4EEFFA3787441E9A" box="[924,1056,1474,1495]" italics="true" pageId="19" pageNumber="555">Microasydates</emphasis>
</taxonomicName>
\
<taxonomicName id="98F24D6CA605FFEF495DFA3787FA1E9A" authorityName="Gimmel and Mayor" authorityYear="2022" box="[1070,1182,1474,1495]" class="Insecta" family="Melyridae" genus="Microasydates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="555" phylum="Arthropoda" rank="species" species="sanclemente">
<emphasis id="6D86EAFDA605FFEF495DFA3787FA1E9A" box="[1070,1182,1474,1495]" italics="true" pageId="19" pageNumber="555">sanclemente</emphasis>
</taxonomicName>
\ Gimmel &amp; Mayor 2022 [red printed label]”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="17E86564A605FFE84FA8FA02824E1A6A" lastPageId="20" lastPageNumber="556" pageId="19" pageNumber="555" type="diagnosis">
<paragraph id="5F4D36EFA605FFE84FA8FA02824E1A6A" blockId="19.[706,1194,1340,1736]" lastBlockId="20.[152,639,194,1736]" lastPageId="20" lastPageNumber="556" pageId="19" pageNumber="555">
<emphasis id="6D86EAFDA605FFEF4FA8FA0280271D40" bold="true" box="[731,835,1527,1549]" pageId="19" pageNumber="555">Diagnosis.</emphasis>
The body with at least mouthparts, legs, and elytral apices (
<figureCitation id="C7C92A6AA605FFEF4ED3F9E780801D65" box="[928,996,1554,1576]" captionStart="Figs" captionStartId="8.[175,214,884,903]" captionTargetBox="[167,1171,193,862]" captionTargetId="figure-419@8.[151,1171,193,862]" captionTargetPageId="8" captionText="Figs. 25–28. Elytral apices of male Microasydates, new genus. 25) M. punctipennis; 26) M. sanclemente, new species; 27) M. santabarbara, new species; 28) M. umbratus." figureDoi="http://doi.org/10.5281/zenodo.13252823" httpUri="https://zenodo.org/record/13252823/files/figure.png" pageId="19" pageNumber="555">Fig. 26</figureCitation>
) pale yellow-orange, and with both pale and dark setae (
<figureCitation id="C7C92A6AA605FFEF497AF9D987281D0F" box="[1033,1100,1580,1602]" captionStart="Figs" captionStartId="4.[177,216,1579,1598]" captionTargetBox="[151,1171,193,1563]" captionTargetId="figure-8@4.[151,1171,193,1563]" captionTargetPageId="4" captionText="Figs. 5–8. Dorsal habitus of male Microasydates, new genus. 5) M. punctipennis (SBMNHENT0080249); 6) M. sanclemente, new species (holotype; SBMNHENT0079371); 7) M. santabarbara, new species, patterned form; 8) M. umbratus, “umbratus” form (specimen from Shasta Co., CA). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252815" httpUri="https://zenodo.org/record/13252815/files/figure.png" pageId="19" pageNumber="555">Figs. 6</figureCitation>
,
<figureCitation id="C7C92A6AA605FFEF4927F9D9870B1D0C" box="[1108,1135,1580,1601]" captionStart="Figs" captionStartId="6.[177,216,1500,1519]" captionTargetBox="[151,1171,193,1480]" captionTargetId="figure-57@6.[151,1171,193,1480]" captionTargetPageId="6" captionText="Figs. 13–16. Dorsal habitus of female Microasydates, new genus. 13) M. punctipennis (LACMENT540887); 14) M. sanclemente, new species (SBMNHENT0079368); 15) M. santabarbara, new species; 16) M. umbratus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252819" httpUri="https://zenodo.org/record/13252819/files/figure.png" pageId="19" pageNumber="555">14</figureCitation>
); pronotum with a short, tight lateral fringe of pale setae (
<figureCitation id="C7C92A6AA605FFEF4FB9F997805B1D3A" box="[714,831,1634,1656]" captionStart="Figs" captionStartId="7.[197,236,1639,1658]" captionTargetBox="[173,1193,194,1622]" captionTargetId="figure-8@7.[173,1193,194,1622]" captionTargetPageId="7" captionText="Figs. 17–24. Pronota of Microasydates, new genus. M. punctipennis: 17) Male; 18) Female. M. sanclemente, new species: 19) Male; 20) Female. M. santabarbara, new species: 21) Male; 22) Female. M. umbratus: 23) Male; 24) Female." figureDoi="http://doi.org/10.5281/zenodo.13252821" httpUri="https://zenodo.org/record/13252821/files/figure.png" pageId="19" pageNumber="555">Figs. 19–20</figureCitation>
); female pygidium moderately deeply emarginate (as in
<figureCitation id="C7C92A6AA605FFEF4E1AF98980CA1DDF" box="[873,942,1660,1682]" captionStart="Figs" captionStartId="10.[176,215,1377,1396]" captionTargetBox="[153,626,215,1331]" captionTargetId="figure-400@10.[150,636,193,1359]" captionTargetPageId="10" captionText="Figs. 35–37. Female pygidia of Microasydates, new genus in glycerol showing degree of apical emargination. 35) M. punctipennis; 36) M. santabarbara, new species; 37) M. umbratus. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252829" httpUri="https://zenodo.org/record/13252829/files/figure.png" pageId="19" pageNumber="555">Fig. 35</figureCitation>
), with a broad median glabrous area (
<figureCitation id="C7C92A6AA605FFEF4E41F962801E1DE1" box="[818,890,1687,1709]" captionStart="Figs" captionStartId="11.[197,238,465,484]" captionTargetBox="[176,1196,185,447]" captionTargetId="figure-626@11.[176,1196,185,447]" captionTargetPageId="11" captionText="Figs. 38–41. Female pygidia of Microasydates, new genus showing pubescence. 38) M. punctipennis;" figureDoi="http://doi.org/10.5281/zenodo.13252831" httpUri="https://zenodo.org/record/13252831/files/figure.png" pageId="19" pageNumber="555">Fig. 39</figureCitation>
); median lobe of aedeagus with only two ventral teeth and a subapical bulge in addition to the apical hook (
<figureCitation id="C7C92A6AA602FFE84CECFF3782811B9A" box="[415,485,194,216]" captionStart="Figs" captionStartId="12.[176,215,1616,1635]" captionTargetBox="[192,1131,211,1589]" captionTargetId="figure-7@12.[192,1131,192,1600]" captionTargetPageId="12" captionText="Figs. 42–53. Median lobe (lateral view) and tegmen (ventral view) for male Microasydates, new genus in glycerol. M. punctipennis: 42) Median lobe; 43) Tegmen. M. sanclemente, new species: 44) Median lobe; 45) Tegmen. M. santabarbara, new species, specimen from Santa Ynez Mountains, CA: 46) Median lobe; 47) Tegmen. M. santabarbara, new species, specimen from Santa Cruz Island, CA: 48) Median lobe; 49) Tegmen. M. umbratus, specimen from Shasta Co., CA: 50) Median lobe; 51) Tegmen. M. umbratus, specimen from Juab Co., UT: 52) Median lobe; 53) Tegmen. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252833" httpUri="https://zenodo.org/record/13252833/files/figure.png" pageId="20" pageNumber="556">Fig. 44</figureCitation>
); and a distribution on San Clemente Island,
<collectingRegion id="9D36F80DA602FFE84CDDFF29816B1BBF" box="[430,527,220,242]" country="United States of America" name="California" pageId="20" pageNumber="556">California</collectingRegion>
distinguish
<taxonomicName id="98F24D6CA602FFE84DEBFF0D825B1A41" authorityName="Gimmel and Mayor" authorityYear="2022" box="[152,319,247,269]" class="Insecta" family="Melyridae" genus="Microasydates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="556" phylum="Arthropoda" rank="species" species="sanclemente">
<emphasis id="6D86EAFDA602FFE84DEBFF0D825B1A41" box="[152,319,247,269]" italics="true" pageId="20" pageNumber="556">M. sanclemente</emphasis>
</taxonomicName>
from all other species of
<taxonomicName id="98F24D6CA602FFE84DEBFEE782401A6A" authorityName="Gimmel &amp; Mayor" authorityYear="2022" box="[152,292,274,295]" class="Insecta" family="Melyridae" genus="Microasydates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="6D86EAFDA602FFE84DEBFEE782401A6A" box="[152,292,274,295]" italics="true" pageId="20" pageNumber="556">Microasydates</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="17E86564A602FFE84DC3FED980E21FD5" pageId="20" pageNumber="556" type="description">
<paragraph id="5F4D36EFA602FFE84DC3FED980E21FD5" blockId="20.[152,639,194,1736]" lastBlockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84DC3FED9824F1A0F" bold="true" box="[176,299,300,322]" pageId="20" pageNumber="556">Description.</emphasis>
Body (
<figureCitation id="C7C92A6AA602FFE84C1CFED982D61A0F" box="[367,434,300,322]" captionStart="Figs" captionStartId="4.[177,216,1579,1598]" captionTargetBox="[151,1171,193,1563]" captionTargetId="figure-8@4.[151,1171,193,1563]" captionTargetPageId="4" captionText="Figs. 5–8. Dorsal habitus of male Microasydates, new genus. 5) M. punctipennis (SBMNHENT0080249); 6) M. sanclemente, new species (holotype; SBMNHENT0079371); 7) M. santabarbara, new species, patterned form; 8) M. umbratus, “umbratus” form (specimen from Shasta Co., CA). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252815" httpUri="https://zenodo.org/record/13252815/files/figure.png" pageId="20" pageNumber="556">Figs. 6</figureCitation>
,
<figureCitation id="C7C92A6AA602FFE84CC9FED982B61A0C" box="[442,466,300,321]" captionStart="Figs" captionStartId="6.[177,216,1500,1519]" captionTargetBox="[151,1171,193,1480]" captionTargetId="figure-57@6.[151,1171,193,1480]" captionTargetPageId="6" captionText="Figs. 13–16. Dorsal habitus of female Microasydates, new genus. 13) M. punctipennis (LACMENT540887); 14) M. sanclemente, new species (SBMNHENT0079368); 15) M. santabarbara, new species; 16) M. umbratus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252819" httpUri="https://zenodo.org/record/13252819/files/figure.png" pageId="20" pageNumber="556">14</figureCitation>
) small, somewhat stout; shiny with slight bronze luster; male pronotum and elytra with distinct but weak isodiametric microsculpture, weaker still on head; female pronotum with microsculpture variable, similar to male or stronger, distinct, somewhat more weakly indicated on elytra. Length
<quantity id="980A9B0AA602FFE84C0BFE3981661AAF" box="[376,514,460,482]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" metricValueMax="2.14" metricValueMin="1.8" pageId="20" pageNumber="556" unit="mm" value="1.97" valueMax="2.14" valueMin="1.8">1.80–2.14 mm</quantity>
(male mean:
<quantity id="980A9B0AA602FFE84DEBFE12838A1AB1" box="[152,238,487,508]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="20" pageNumber="556" unit="mm" value="1.88">1.88 mm</quantity>
; female mean:
<quantity id="980A9B0AA602FFE84C0FFE1282B71AB1" box="[380,467,487,509]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="20" pageNumber="556" unit="mm" value="2.05">2.05 mm</quantity>
); width
<quantity id="980A9B0AA602FFE84F6DFE1283D9195A" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="9.399999999999999" metricValueMin="8.0" pageId="20" pageNumber="556" unit="mm" value="0.87" valueMax="0.94" valueMin="0.8">0.80–0.94 mm</quantity>
(male mean:
<quantity id="980A9B0AA602FFE84C48FDF782F5195A" box="[315,401,514,535]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="20" pageNumber="556" unit="mm" value="0.81">0.81 mm</quantity>
; female mean:
<quantity id="980A9B0AA602FFE84F6EFDF78110195A" box="[541,628,514,535]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="20" pageNumber="556" unit="mm" value="0.89">0.89 mm</quantity>
). Body (
<figureCitation id="C7C92A6AA602FFE84DABFDE9827F197F" box="[216,283,540,562]" captionStart="Figs" captionStartId="4.[177,216,1579,1598]" captionTargetBox="[151,1171,193,1563]" captionTargetId="figure-8@4.[151,1171,193,1563]" captionTargetPageId="4" captionText="Figs. 5–8. Dorsal habitus of male Microasydates, new genus. 5) M. punctipennis (SBMNHENT0080249); 6) M. sanclemente, new species (holotype; SBMNHENT0079371); 7) M. santabarbara, new species, patterned form; 8) M. umbratus, “umbratus” form (specimen from Shasta Co., CA). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252815" httpUri="https://zenodo.org/record/13252815/files/figure.png" pageId="20" pageNumber="556">Figs. 6</figureCitation>
,
<figureCitation id="C7C92A6AA602FFE84C57FDE98259197C" box="[292,317,540,561]" captionStart="Figs" captionStartId="6.[177,216,1500,1519]" captionTargetBox="[151,1171,193,1480]" captionTargetId="figure-57@6.[151,1171,193,1480]" captionTargetPageId="6" captionText="Figs. 13–16. Dorsal habitus of female Microasydates, new genus. 13) M. punctipennis (LACMENT540887); 14) M. sanclemente, new species (SBMNHENT0079368); 15) M. santabarbara, new species; 16) M. umbratus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252819" httpUri="https://zenodo.org/record/13252819/files/figure.png" pageId="20" pageNumber="556">14</figureCitation>
) entirely black except the following: antennae and legs dark reddish-brown, antennae apically becoming darker, tibiae and tarsi often lighter reddish-brown; elytral apices (
<figureCitation id="C7C92A6AA602FFE84C85FD99815E19CF" box="[502,570,620,642]" captionStart="Figs" captionStartId="8.[175,214,884,903]" captionTargetBox="[167,1171,193,862]" captionTargetId="figure-419@8.[151,1171,193,862]" captionTargetPageId="8" captionText="Figs. 25–28. Elytral apices of male Microasydates, new genus. 25) M. punctipennis; 26) M. sanclemente, new species; 27) M. santabarbara, new species; 28) M. umbratus." figureDoi="http://doi.org/10.5281/zenodo.13252823" httpUri="https://zenodo.org/record/13252823/files/figure.png" pageId="20" pageNumber="556">Fig. 26</figureCitation>
) bright reddish brown in a well-defined lunule across the suture and extending along lateral margins anywhere from one-fifth to one-third elytral length; apex of abdomen around genital opening and sometimes female with apical 2–3 ventrites bright reddish brown. Pubescence moderately dense, completely reclining, mostly rusty-brown to black, but patches of pale silvery setae present over most of head dorsally (occiput with patch of rusty setae), pronotum (large central patch consisting of rusty setae), scutellar shield, and elytra at extreme basal margin and in a broad sub-basal band extending across and along suture to scutellar shield, in certain light giving the elytra a tricolored (black-cream-red) appearance, ventral surface with scattered pale setae; pronotal fringe short and comprised of pale setae, not projecting but somewhat loose; elytral fringe longer, mostly pale except at apex, somewhat loose. Punctation shallow and inconspicuous on head and pronotum, coarser on elytra, punctures separated by about one puncture diameter or less near elytral base but becoming sparser apically.
<emphasis id="6D86EAFDA602FFE84CCCFB4282991F81" bold="true" box="[447,509,1207,1228]" pageId="20" pageNumber="556">Head:</emphasis>
Not conspicuously enlarged, very slightly wider in male, distinctly narrower than pronotum, eyes rounded laterally, bulging; 1.54–1.69 times wider than interocular distance (male mean: 1.64; female mean: 1.56). Antennomeres 5–10 weakly serrate, antennomeres 6 and 8 somewhat reduced, antennomeres 9–11 more symmetrical; male antenna short, reaching to near basal one-fourth of pronotum posteriorly, female antenna slightly shorter.
<emphasis id="6D86EAFDA602FFE84CB5FA52817E1EF1" bold="true" box="[454,538,1447,1468]" pageId="20" pageNumber="556">ThoraX:</emphasis>
Pronotum (
<figureCitation id="C7C92A6AA602FFE84DECFA37826A1E9A" box="[159,270,1474,1496]" captionStart="Figs" captionStartId="7.[197,236,1639,1658]" captionTargetBox="[173,1193,194,1622]" captionTargetId="figure-8@7.[173,1193,194,1622]" captionTargetPageId="7" captionText="Figs. 17–24. Pronota of Microasydates, new genus. M. punctipennis: 17) Male; 18) Female. M. sanclemente, new species: 19) Male; 20) Female. M. santabarbara, new species: 21) Male; 22) Female. M. umbratus: 23) Male; 24) Female." figureDoi="http://doi.org/10.5281/zenodo.13252821" httpUri="https://zenodo.org/record/13252821/files/figure.png" pageId="20" pageNumber="556">Figs. 19, 20</figureCitation>
) wider than long, appearing transverse, 0.69–0.75 times as long as wide (male mean: 0.71; female mean: 0.72); widest at middle; lateral edges straight and diverging to middle in basal half, then straight and converging to anterior margin, not constricted, lateral bead very weakly crenulate; anterior and posterior angles obsolete, broadly rounded. Elytra moderately elongate, broadly oval, more elongate in female, elytron 3.26–3.61 times longer than wide (male mean: 3.38; female mean: 3.48), elytra at humeri slightly wider than pronotum; elytral epipleuron distinct at base, tapering gradually and becoming obsolete near apex of abdominal ventrite
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both sexes, lateral carina barely visible in dorsal view for entire length, not explanate; apical margin broadly rounded, sutural angle absent in male (
<figureCitation id="C7C92A6AA602FFE84F94FE97804F1A35" box="[743,811,354,376]" captionStart="Figs" captionStartId="8.[175,214,884,903]" captionTargetBox="[167,1171,193,862]" captionTargetId="figure-419@8.[151,1171,193,862]" captionTargetPageId="8" captionText="Figs. 25–28. Elytral apices of male Microasydates, new genus. 25) M. punctipennis; 26) M. sanclemente, new species; 27) M. santabarbara, new species; 28) M. umbratus." figureDoi="http://doi.org/10.5281/zenodo.13252823" httpUri="https://zenodo.org/record/13252823/files/figure.png" pageId="20" pageNumber="556">Fig. 26</figureCitation>
), sometimes weakly present in female, margin very weakly serrulate in both sexes.All tibiae with fine, pale setae along external margin; pro- and mesotibiae with very inconspicuous, short, slender and sparse spines arranged along external margin, spines slightly more conspicuous in female; metatibia of male with weak brush of dense, fine, black setae in apical one-third on inner margin, brush virtually absent in female.
<emphasis id="6D86EAFDA602FFE84E9DFDC2873E1901" bold="true" box="[1006,1114,567,588]" pageId="20" pageNumber="556">Abdomen:</emphasis>
Male ventrites 4 and 5 weakly flattened or depressed medially, with transverse field of black spiculate setae (very inconspicuous on the dark abdomen) in apical half medially on ventrite 4, a few black spiculate setae present apicomedially on ventrite 3, black spiculate setae absent from ventrite 5. Pygidium of male with scattered sparse setae, glabrous in ill-defined basal triangle, with apex of pygidium truncate (
<figureCitation id="C7C92A6AA602FFE84FACFCD280411871" box="[735,805,807,829]" captionStart="Figs" captionStartId="9.[197,236,1617,1636]" captionTargetBox="[201,621,202,1592]" captionTargetId="figure-216@9.[185,632,195,1606]" captionTargetPageId="9" captionText="Figs. 29–32. Pygidia with attached sternite VIII of male Microasydates, new genus in glycerol. 29) M. punctipennis; 30) M. sanclemente, new species; 31) M. santabarbara, new species; 32) M. umbratus. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252827" httpUri="https://zenodo.org/record/13252827/files/figure.png" pageId="20" pageNumber="556">Fig. 30</figureCitation>
); female pygidium sparsely setose with wide median glabrous strip (
<figureCitation id="C7C92A6AA602FFE84EBCFCB7877F181A" box="[975,1051,834,856]" captionStart="Figs" captionStartId="11.[197,238,465,484]" captionTargetBox="[176,1196,185,447]" captionTargetId="figure-626@11.[176,1196,185,447]" captionTargetPageId="11" captionText="Figs. 38–41. Female pygidia of Microasydates, new genus showing pubescence. 38) M. punctipennis;" figureDoi="http://doi.org/10.5281/zenodo.13252831" httpUri="https://zenodo.org/record/13252831/files/figure.png" pageId="20" pageNumber="556">Fig. 39</figureCitation>
), apex with small, weak, arcuate emargination. Spiculum gastrale with basal strut short, distinctly less than one-half diameter of ring. Median lobe of aedeagus in lateral view (
<figureCitation id="C7C92A6AA602FFE84E41FC598018188C" box="[818,892,940,962]" captionStart="Figs" captionStartId="12.[176,215,1616,1635]" captionTargetBox="[192,1131,211,1589]" captionTargetId="figure-7@12.[192,1131,192,1600]" captionTargetPageId="12" captionText="Figs. 42–53. Median lobe (lateral view) and tegmen (ventral view) for male Microasydates, new genus in glycerol. M. punctipennis: 42) Median lobe; 43) Tegmen. M. sanclemente, new species: 44) Median lobe; 45) Tegmen. M. santabarbara, new species, specimen from Santa Ynez Mountains, CA: 46) Median lobe; 47) Tegmen. M. santabarbara, new species, specimen from Santa Cruz Island, CA: 48) Median lobe; 49) Tegmen. M. umbratus, specimen from Shasta Co., CA: 50) Median lobe; 51) Tegmen. M. umbratus, specimen from Juab Co., UT: 52) Median lobe; 53) Tegmen. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252833" httpUri="https://zenodo.org/record/13252833/files/figure.png" pageId="20" pageNumber="556">Fig. 44</figureCitation>
) with apical limb distinctly longer than basal limb, apex short, stout, strongly acute and sharply directed ventrally, ventral margin of apical limb with pair of ventrolateral teeth at same level (near proximal end of apical orifice), with slight ventral bulge between paired teeth and apex; tegmen (
<figureCitation id="C7C92A6AA602FFE84E37FBB980EB1F2F" box="[836,911,1100,1122]" captionStart="Figs" captionStartId="12.[176,215,1616,1635]" captionTargetBox="[192,1131,211,1589]" captionTargetId="figure-7@12.[192,1131,192,1600]" captionTargetPageId="12" captionText="Figs. 42–53. Median lobe (lateral view) and tegmen (ventral view) for male Microasydates, new genus in glycerol. M. punctipennis: 42) Median lobe; 43) Tegmen. M. sanclemente, new species: 44) Median lobe; 45) Tegmen. M. santabarbara, new species, specimen from Santa Ynez Mountains, CA: 46) Median lobe; 47) Tegmen. M. santabarbara, new species, specimen from Santa Cruz Island, CA: 48) Median lobe; 49) Tegmen. M. umbratus, specimen from Shasta Co., CA: 50) Median lobe; 51) Tegmen. M. umbratus, specimen from Juab Co., UT: 52) Median lobe; 53) Tegmen. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.13252833" httpUri="https://zenodo.org/record/13252833/files/figure.png" pageId="20" pageNumber="556">Fig. 45</figureCitation>
) relatively elongate, with slight constriction near base of ring. Female internal terminalia not studied.
</paragraph>
</subSubSection>
<subSubSection id="17E86564A602FFE84FB6FB69873A1FA5" pageId="20" pageNumber="556" type="etymology">
<paragraph id="5F4D36EFA602FFE84FB6FB69873A1FA5" blockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84FB6FB69805C1FFC" bold="true" box="[709,824,1180,1201]" pageId="20" pageNumber="556">Etymology.</emphasis>
Named after San Clemente Island of the
<collectingRegion id="9D36F80DA602FFE84FBCFB42804A1F80" box="[719,814,1207,1229]" country="United States of America" name="California" pageId="20" pageNumber="556">California</collectingRegion>
Channel Islands, the
<typeStatus id="8049884DA602FFE84E80FB42877F1F81" box="[1011,1051,1207,1228]" pageId="20" pageNumber="556">type</typeStatus>
locality. The name is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="17E86564A602FFE84FB6FB1987461D11" pageId="20" pageNumber="556" type="biology_ecology">
<paragraph id="5F4D36EFA602FFE84FB6FB1980A01E50" blockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84FB6FB1980D81E4F" bold="true" box="[709,956,1260,1282]" pageId="20" pageNumber="556">Host Plant Associations.</emphasis>
No host plant records are available for this species.
</paragraph>
<paragraph id="5F4D36EFA602FFE84FB6FAD4806C1E1F" blockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84FB6FAD480D61E7A" bold="true" box="[709,946,1313,1335]" pageId="20" pageNumber="556">Seasonal Distribution.</emphasis>
8 May to 15 June (
<figureCitation id="C7C92A6AA602FFE84FC7FAC981981E1F" box="[692,764,1340,1362]" captionStart="Fig" captionStartId="13.[198,229,1695,1714]" captionTargetBox="[221,419,1522,1651]" captionTargetId="graphics-126@13.[221,419,1522,1652]" captionTargetPageId="13" captionText="Fig. 64. Seasonalities of adult Microasydates, new genus species by month, based on collection records in the Specimens Examined sections." figureDoi="http://doi.org/10.5281/zenodo.13252839" httpUri="https://zenodo.org/record/13252839/files/figure.png" pageId="20" pageNumber="556">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="5F4D36EFA602FFE84FB6FAA2872B1EEF" blockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84FB6FAA280A41E20" bold="true" box="[709,960,1367,1389]" pageId="20" pageNumber="556">Geographic Distribution.</emphasis>
<materialsCitation id="EF9A3CB2A602FFE84EB6FAA2872F1EEF" location="Channel Islands" municipality="San Clemente Island" pageId="20" pageNumber="556" specimenCount="1" stateProvince="California">
Known only from
<collectingMunicipality id="BF29AC95A602FFE8491CFAA280301EC5" pageId="20" pageNumber="556">San Clemente Island</collectingMunicipality>
, southernmost of the
<location id="5A2D6034A602FFE84930FA8781951EEF" LSID="urn:lsid:plazi:treatment:D75B87F9A605FFE94FACFB5182D41A0F:5A2D6034A602FFE84930FA8781951EEF" municipality="San Clemente Island" name="Channel Islands" pageId="20" pageNumber="556" stateProvince="California">Channel Islands</location>
off the coast of southern
<collectingRegion id="9D36F80DA602FFE84E95FA79872F1EEF" box="[998,1099,1420,1442]" country="United States of America" name="California" pageId="20" pageNumber="556">California</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="5F4D36EFA602FFE84FB6FA5287461D11" blockId="20.[685,1171,194,1736]" pageId="20" pageNumber="556">
<materialsCitation id="EF9A3CB2A602FFE84FB6FA52877B1D11" collectingDate="2008-06-15" collectingDateMax="2010-05-08" collectingDateMin="2008-06-15" collectionCode="SBMNH" collectorName="Twin Dams Canyon &amp; US Navy &amp; Caterino &amp; Chatzimanolis" county="Los Angeles County" location="San Clemente Island" municipality="San Clemente Island" pageId="20" pageNumber="556" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
<emphasis id="6D86EAFDA602FFE84FB6FA5280651E9A" bold="true" pageId="20" pageNumber="556">
<typeStatus id="8049884DA602FFE84FB6FA5280481EF1" box="[709,812,1447,1468]" pageId="20" pageNumber="556" type="paratype">Paratypes</typeStatus>
(6).
<collectingRegion id="9D36F80DA602FFE84E16FA5287671EF0" box="[869,1027,1447,1469]" country="United States of America" name="California" pageId="20" pageNumber="556">CALIFORNIA</collectingRegion>
:
<collectingCounty id="B62C4E63A602FFE84960FA5281991E9A" pageId="20" pageNumber="556">Los Angeles County</collectingCounty>
:
</emphasis>
<collectingMunicipality id="BF29AC95A602FFE84E7AFA3780B21E95" box="[777,982,1474,1496]" pageId="20" pageNumber="556">San Clemente Island</collectingMunicipality>
, above
<collectorName id="F2075339A602FFE84951FA37819E1EBC" pageId="20" pageNumber="556">Twin Dams Canyon</collectorName>
, 32°54′13.03′′N, 118°28′28.08′′W, loamy grassland,
<date id="2B4C102FA602FFE84E66FA0280EA1D41" box="[789,910,1527,1548]" pageId="20" pageNumber="556" value="2010-05-08">
<collectingDate id="3B08E9C7A602FFE84E66FA0280EA1D41" box="[789,910,1527,1548]" pageId="20" pageNumber="556" value="2010-05-08">8 May 2010</collectingDate>
</date>
,
<collectorName id="F2075339A602FFE84EE9FA0280901D41" box="[922,1012,1527,1548]" pageId="20" pageNumber="556">US Navy</collectorName>
(
<specimenCount id="49F4FD66A602FFE84976FA0287461D41" box="[1029,1058,1527,1548]" count="1" pageId="20" pageNumber="556" type="female">1♀</specimenCount>
,
<collectionCode id="39E3AE2AA602FFE8495CFA0287ED1D41" box="[1071,1161,1527,1548]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="20" pageNumber="556" type="Museum">SBMNH</collectionCode>
);
<location id="5A2D6034A602FFE84FDEF9E780161D65" LSID="urn:lsid:plazi:treatment:D75B87F9A605FFE94FACFB5182D41A0F:5A2D6034A602FFE84FDEF9E780161D65" box="[685,882,1554,1576]" county="Los Angeles County" municipality="San Clemente Island" name="San Clemente Island" pageId="20" pageNumber="556" stateProvince="California">San Clemente Island</location>
,
<location id="5A2D6034A602FFE84E09F9E787531D6A" LSID="urn:lsid:plazi:treatment:D75B87F9A605FFE94FACFB5182D41A0F:5A2D6034A602FFE84E09F9E787531D6A" box="[890,1079,1554,1575]" county="Los Angeles County" municipality="San Clemente Island" name="upper Cave Canyon" pageId="20" pageNumber="556" stateProvince="California">upper Cave Canyon</location>
, 32.873 4, −118.4480,
<date id="2B4C102FA602FFE84E46F9D980861D0C" box="[821,994,1580,1602]" pageId="20" pageNumber="556" value="2008-06-15">
<collectingDate id="3B08E9C7A602FFE84E46F9D980861D0C" box="[821,994,1580,1602]" pageId="20" pageNumber="556" value="2008-06-15">15 June 2008</collectingDate>
</date>
,
<collectorName id="F2075339A602FFE84971F9D987311D0F" box="[1026,1109,1580,1602]" pageId="20" pageNumber="556">Caterino</collectorName>
and
<collectorName id="F2075339A602FFE84FDEF9B2805C1D10" box="[685,824,1607,1629]" pageId="20" pageNumber="556">Chatzimanolis</collectorName>
(
<specimenCount id="49F4FD66A602FFE84E35F9B2801C1D11" box="[838,888,1607,1628]" count="1" pageId="20" pageNumber="556" type="male">1♂♂</specimenCount>
,
<specimenCount id="49F4FD66A602FFE84EF1F9B280D71D11" box="[898,947,1607,1628]" count="4" pageId="20" pageNumber="556" type="female">4♀♀</specimenCount>
,
<collectionCode id="39E3AE2AA602FFE84ECDF9B2877C1D11" box="[958,1048,1607,1628]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" lsid="urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="20" pageNumber="556" type="Museum">SBMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="17E86564A602FFE94FB6F99782D41A0F" lastPageId="21" lastPageNumber="557" pageId="20" pageNumber="556" type="discussion">
<paragraph id="5F4D36EFA602FFE94FB6F99782D41A0F" blockId="20.[685,1171,194,1736]" lastBlockId="21.[173,659,194,322]" lastPageId="21" lastPageNumber="557" pageId="20" pageNumber="556">
<emphasis id="6D86EAFDA602FFE84FB6F997804C1D3A" bold="true" box="[709,808,1634,1655]" pageId="20" pageNumber="556">Remarks.</emphasis>
<taxonomicName id="98F24D6CA602FFE84E44F997872D1D3A" authorityName="Gimmel and Mayor" authorityYear="2022" box="[823,1097,1634,1655]" class="Insecta" family="Melyridae" genus="Microasydates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="556" phylum="Arthropoda" rank="species" species="sanclemente">
<emphasis id="6D86EAFDA602FFE84E44F997872D1D3A" box="[823,1097,1634,1655]" italics="true" pageId="20" pageNumber="556">Microasydates sanclemente</emphasis>
</taxonomicName>
shares many characters with
<taxonomicName id="98F24D6CA602FFE84EE1F98887561DDF" authorityName="LeConte" authorityYear="1866" box="[914,1074,1661,1682]" class="Insecta" family="Melyridae" genus="Pristoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="556" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="6D86EAFDA602FFE84EE1F98887561DDF" box="[914,1074,1661,1682]" italics="true" pageId="20" pageNumber="556">M. punctipennis</emphasis>
</taxonomicName>
of Santa Catalina Island, including the shape of the aedeagal median lobe, but may be readily separated from that species by the presence of pale setae on the pronotum and elytra (pale setae absent in
<taxonomicName id="98F24D6CA603FFE94F78FF2883AD1A40" authorityName="LeConte" authorityYear="1866" class="Insecta" family="Melyridae" genus="Pristoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="557" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="6D86EAFDA603FFE94F78FF2883AD1A40" italics="true" pageId="21" pageNumber="557">M. punctipennis</emphasis>
</taxonomicName>
), the conspicuously pale reddish elytral apices (elytra uniformly dark in
<taxonomicName id="98F24D6CA603FFE94CECFEE7815E1A6A" authorityName="LeConte" authorityYear="1866" box="[415,570,274,295]" class="Insecta" family="Melyridae" genus="Pristoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="557" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="6D86EAFDA603FFE94CECFEE7815E1A6A" box="[415,570,274,295]" italics="true" pageId="21" pageNumber="557">M. punctipennis</emphasis>
</taxonomicName>
), and the more transverse pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>

1693
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<mods:url id="2B8DE96CA090B611218E95D12E71C1FF">http://dx.doi.org/10.1649/0010-065x-76.3.482</mods:url>
</mods:location>
<mods:classification id="3A3736C3615CE481E0ED9B5E18BACEF5">journal article</mods:classification>
<mods:identifier id="1E865D5557A885839983E7783ED07FBA" type="DOI">10.1649/0010-065X-76.3.482</mods:identifier>
<mods:identifier id="B785AC1A861BB969CFD601CB1094B7E8" type="ISSN">1938-4394</mods:identifier>
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</mods:mods>
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<subSubSection id="2DAC81C26E1B05107768FB52224DFB55" box="[787,1093,1181,1203]" pageId="1" pageNumber="482" type="nomenclature">
<paragraph id="6509D2496E1B05107768FB52224DFB55" blockId="1.[787,1093,1181,1203]" box="[787,1093,1181,1203]" pageId="1" pageNumber="482">
<heading id="3E4165256E1B05107768FB52224DFB55" box="[787,1093,1181,1203]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="482" reason="2">
<emphasis id="57C20E5B6E1B05107768FB52224DFB55" bold="true" box="[787,1093,1181,1203]" pageId="1" pageNumber="482">
Genus
<taxonomicName id="A2B6A9CA6E1B05107721FB52224DFB55" ID-CoL="8VXKV" authority="Kraatz, 1859" authorityName="Kraatz" authorityYear="1859" box="[858,1093,1181,1202]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="57C20E5B6E1B05107721FB5225B3FB55" bold="true" box="[858,955,1181,1202]" italics="true" pageId="1" pageNumber="482">EUPIeSTUS</emphasis>
Kraatz, 1859
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2DAC81C26E1B051376D6FB072460FE75" lastPageId="2" lastPageNumber="483" pageId="1" pageNumber="482" type="materials_examined">
<paragraph id="6509D2496E1B051076D6FB072542FB1F" blockId="1.[685,1170,1224,1272]" pageId="1" pageNumber="482">
<typeStatus id="BA0D6CEB6E1B051076D6FB0724D5FB3A" box="[685,733,1224,1245]" pageId="1" pageNumber="482">Type</typeStatus>
species:
<taxonomicName id="A2B6A9CA6E1B0510774DFB07225FFB3A" authority="Kraatz" authorityName="Kraatz" authorityYear="1859" box="[822,1111,1224,1245]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="482" phylum="Arthropoda" rank="species" species="sculpticollis">
<emphasis id="57C20E5B6E1B0510774DFB072206FB3A" box="[822,1038,1224,1245]" italics="true" pageId="1" pageNumber="482">Eupiestus sculpticollis</emphasis>
Kraatz
</taxonomicName>
, fixed by monotypy.
</paragraph>
<paragraph id="6509D2496E1B05107697FAD225D3FA47" blockId="1.[705,1150,1309,1440]" pageId="1" pageNumber="482">
<emphasis id="57C20E5B6E1B05107697FAD225D3FAB7" bold="true" pageId="1" pageNumber="482">
<taxonomicName id="A2B6A9CA6E1B05107697FAD2226FFAD5" authority="Park and Ahn" authorityName="Park and Ahn" authorityYear="2022" box="[748,1127,1309,1331]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="482" phylum="Arthropoda" rank="species" species="rotundifrons">
<emphasis id="57C20E5B6E1B05107697FAD225D8FAD4" bold="true" box="[748,976,1309,1331]" italics="true" pageId="1" pageNumber="482">EUPIeSTUS roTUndIfronS</emphasis>
Park and Ahn
</taxonomicName>
, new species
</emphasis>
zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="1110E89C6E1B051076BAFABF2276FA61" box="[705,1150,1392,1414]" pageId="1" pageNumber="482">9598D8C0-60B5-4C35-8FF1-3FD8356E1277</uuid>
</uri>
(
<figureCitation id="FD8DCECC6E1B05107717FA4525DBFA47" box="[876,979,1418,1440]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[197,236,1687,1706]" captionStartId-1="3.[177,216,1639,1658]" captionTargetBox-0="[185,1189,671,1648]" captionTargetBox-1="[158,1155,502,1612]" captionTargetId-0="figure-236@2.[173,1193,661,1670]" captionTargetId-1="figure-157@3.[151,1171,494,1623]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1–3. Eupiestus rotundifrons, new species. 1) Habitus, holotype, male, 2.8 mm; 2) Superior marginal line" captionText-1="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13252725" figureDoi-1="http://doi.org/10.5281/zenodo.13252727" httpUri-0="https://zenodo.org/record/13252725/files/figure.png" httpUri-1="https://zenodo.org/record/13252727/files/figure.png" pageId="1" pageNumber="482">Figs. 1–13</figureCitation>
)
</paragraph>
<paragraph id="6509D2496E1B051376BEFA7A240EFEEA" blockId="1.[685,1171,1461,1723]" lastBlockId="2.[173,660,194,616]" lastPageId="2" lastPageNumber="483" pageId="1" pageNumber="482">
<emphasis id="57C20E5B6E1B051076BEFA7A257AFA2C" bold="true" box="[709,882,1461,1483]" pageId="1" pageNumber="482">Type Specimens.</emphasis>
<materialsCitation id="D5DED8146E1B05107707FA7A24F1F98D" collectionCode="CNUIC" country="South Korea" location="Mt. Sikjangsan" pageId="1" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Chungnam" typeStatus="holotype">
<typeStatus id="BA0D6CEB6E1B05107707FA7A25D0FA2C" box="[892,984,1461,1483]" pageId="1" pageNumber="482" type="holotype">Holotype</typeStatus>
,
<specimenCount id="73B019C06E1B0510779FFA7925F0FA2C" box="[996,1016,1462,1483]" count="1" pageId="1" pageNumber="482" type="male">♂</specimenCount>
, labeled as follows: “
<collectingCountry id="1DA192D96E1B05107681FA1F255AFA02" box="[762,850,1488,1509]" name="South Korea" pageId="1" pageNumber="482">KOREA</collectingCountry>
:
<collectingRegion id="A7721CAB6E1B0510771FFA1F221BFA02" box="[868,1043,1488,1510]" country="South Korea" name="Chungnam" pageId="1" pageNumber="482">Chungnam Prov.</collectingRegion>
, Daejeon-si, Dong-gu, Secheon-dong,
<location id="606984926E1B051077D4FA242240F9E7" LSID="urn:lsid:plazi:treatment:ED1F635F6E1B05157768FB5226F2FB2A:606984926E1B051077D4FA242240F9E7" box="[943,1096,1514,1536]" country="South Korea" name="Mt. Sikjangsan" pageId="1" pageNumber="482" stateProvince="Chungnam">Mt. Sikjangsan</location>
, 2019– 2020, WJ Jeong, JY Park, GN Jeon, YJ Kim, YJ Lim, YJ Shin, under bark;
<typeStatus id="BA0D6CEB6E1B051077CBF9EF2227F9D2" box="[944,1071,1568,1589]" pageId="1" pageNumber="482" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="A2B6A9CA6E1B0510704EF9EF25F4F9B7" authority="Park and Ahn 2022" authorityName="Park and Ahn" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="482" phylum="Arthropoda" rank="species" species="rotundifrons">
<emphasis id="57C20E5B6E1B0510704EF9EF252BF9A8" italics="true" pageId="1" pageNumber="482">Eupiestus rotundifrons</emphasis>
Park and Ahn 2022
</taxonomicName>
”. Deposited in
<collectionCode id="03A74A8C6E1B051076D6F99A24F1F98D" box="[685,761,1621,1642]" pageId="1" pageNumber="482">CNUIC</collectionCode>
</materialsCitation>
.
<materialsCitation id="D5DED8146E1B0513777AF99A240AFEEA" collectingDate="1998-09-09" collectingDateMax="2010-05-27" collectingDateMin="1998-09-09" collectionCode="CNUIC" collectorName="K. - J. Ahn" country="South Korea" county="Pyeongchang-gun" elevation="770" lastPageId="2" lastPageNumber="483" latitude="37.766193" location="Mt. Odaesan" longLatPrecision="1" longitude="128.57697" municipality="Jinbu-myeon" pageId="1" pageNumber="482" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Gangwon" typeStatus="paratype">
<emphasis id="57C20E5B6E1B0510777AF99A256EF98D" bold="true" box="[769,870,1621,1642]" pageId="1" pageNumber="482">
<typeStatus id="BA0D6CEB6E1B0510777AF99A256EF98D" box="[769,870,1621,1642]" pageId="1" pageNumber="482" type="paratype">Paratypes</typeStatus>
</emphasis>
, 13 exx. (
<collectionCode id="03A74A8C6E1B051077C4F99A2206F98D" box="[959,1038,1621,1642]" pageId="1" pageNumber="482">CNUIC</collectionCode>
):
<specimenCount id="73B019C06E1B05107066F99A2236F98C" box="[1053,1086,1621,1643]" count="8" pageId="1" pageNumber="482" type="male">8♂</specimenCount>
<specimenCount id="73B019C06E1B05107045F99A2251F98C" box="[1086,1113,1621,1643]" count="5" pageId="1" pageNumber="482" type="female">5♀</specimenCount>
, same data as holotype; 1, “
<collectingCountry id="1DA192D96E1B051077F7F9BF25ECF962" box="[908,996,1648,1669]" name="South Korea" pageId="1" pageNumber="482">KOREA</collectingCountry>
:
<collectingRegion id="A7721CAB6E1B05107789F9BF2298F962" box="[1010,1168,1648,1669]" country="South Korea" name="Gangwon" pageId="1" pageNumber="482">Gangwon Prov.</collectingRegion>
,
<collectingCounty id="8C68AAC56E1B051076D6F9452555F947" box="[685,861,1674,1696]" pageId="1" pageNumber="482">Pyeongchang-gun</collectingCounty>
,
<collectingMunicipality id="856D48336E1B05107709F94525FAF947" box="[882,1010,1674,1696]" pageId="1" pageNumber="482">Jinbu-myeon</collectingMunicipality>
,
<location id="606984926E1B0510707CF9442287F947" LSID="urn:lsid:plazi:treatment:ED1F635F6E1B05157768FB5226F2FB2A:606984926E1B0510707CF9442287F947" box="[1031,1167,1674,1696]" country="South Korea" county="Pyeongchang-gun" latitude="37.766193" longLatPrecision="1" longitude="128.57697" municipality="Jinbu-myeon" name="Mt. Odaesan" pageId="1" pageNumber="482" stateProvince="Gangwon">Mt. Odaesan</location>
,
<geoCoordinate id="0082B48E6E1B051076D6F96B252FF95C" box="[685,807,1700,1723]" degrees="37" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="482" precision="1" seconds="58.3" value="37.766193">N37°45′58.3</geoCoordinate>
<geoCoordinate id="0082B48E6E1B05107755F96B25B3F95C" box="[814,955,1700,1723]" degrees="128" direction="east" minutes="34" orientation="longitude" pageId="1" pageNumber="482" precision="1" seconds="37.1" value="128.57697">E128°34′37.1″</geoCoordinate>
<quantity id="A24E7FAC6E1B051077B9F96A25F4F95D" box="[962,1020,1701,1722]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="482" unit="m" value="770.0">
<elevation id="EE9B357A6E1B051077B9F96A25F4F95D" box="[962,1020,1701,1722]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="482" unit="m" value="770.0">770m</elevation>
</quantity>
,
<date id="1108F4896E1B0510707DF96A2265F95D" box="[1030,1133,1701,1723]" pageId="1" pageNumber="482" value="2010-05-27">
<collectingDate id="014C0D616E1B0510707DF96A2265F95D" box="[1030,1133,1701,1723]" pageId="1" pageNumber="482" value="2010-05-27">27 V 2010</collectingDate>
</date>
, KJ Ahn, SG Lee, under bark”; 2 exx., “
<collectingCountry id="1DA192D96E180513764CFF0D2487FF30" box="[567,655,194,215]" name="South Korea" pageId="2" pageNumber="483">KOREA</collectingCountry>
:
<location id="606984926E18051374D6FF122741FF15" LSID="urn:lsid:plazi:treatment:ED1F635F6E1B05157768FB5226F2FB2A:606984926E18051374D6FF122741FF15" box="[173,329,221,242]" country="South Korea" county="Pyeongchang-gun" latitude="37.766193" longLatPrecision="1" longitude="128.57697" municipality="Jinbu-myeon" name="Kangwon Prov." pageId="2" pageNumber="483" stateProvince="Gangwon">Kangwon Prov.</location>
, Hoengsungun, Unduryeng,
<date id="1108F4896E1805137618FF1326F7FEEB" pageId="2" pageNumber="483" value="1998-09-09" valueMax="1998-09-10" valueMin="1998-09-09">
<collectingDate id="014C0D616E1805137618FF1326F7FEEB" pageId="2" pageNumber="483" value="1998-09-09" valueMax="1998-09-10" valueMin="1998-09-09">9–10 IX 1998</collectingDate>
</date>
,
<collectorName id="C843B79F6E1805137572FF382760FEEA" box="[265,360,247,269]" pageId="2" pageNumber="483">K.-J. Ahn</collectorName>
,
<emphasis id="57C20E5B6E180513750AFF37278EFEEA" box="[369,390,248,269]" italics="true" pageId="2" pageNumber="483">ex</emphasis>
under bark”
</materialsCitation>
.
</paragraph>
<paragraph id="6509D2496E18051374BEFEDE2460FE75" blockId="2.[173,660,194,616]" pageId="2" pageNumber="483">
<materialsCitation id="D5DED8146E18051374BEFEDE246CFE75" collectingDate="2020-07-08" collectionCode="CNUIC" country="South Korea" elevation="118" latitude="36.32632" location="Sechondong" longLatPrecision="1" longitude="127.4839" municipality="Dong-gu" pageId="2" pageNumber="483" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Chungnam">
<emphasis id="57C20E5B6E18051374BEFEDE27E1FEC0" bold="true" box="[197,489,273,295]" pageId="2" pageNumber="483">
<typeStatus id="BA0D6CEB6E18051374BEFEDE27EDFEC0" box="[197,485,273,295]" isOtherTypeStatus="true" pageId="2" pageNumber="483">Other Specimens Examined</typeStatus>
.
</emphasis>
9 exx. (
<collectionCode id="03A74A8C6E1805137643FEDD2481FEC0" box="[568,649,274,295]" pageId="2" pageNumber="483">CNUIC</collectionCode>
):
<specimenCount id="73B019C06E18051374D6FEE326C7FEA5" box="[173,207,300,322]" count="3" pageId="2" pageNumber="483" type="male">3♂</specimenCount>
<specimenCount id="73B019C06E18051374B4FEE326E2FEA5" box="[207,234,300,322]" count="5" pageId="2" pageNumber="483" type="female">5♀</specimenCount>
, same data as holotype;
<specimenCount id="73B019C06E1805137586FEE32414FEA6" box="[509,540,300,321]" count="1" pageId="2" pageNumber="483" type="male">1♂</specimenCount>
, “
<collectingCountry id="1DA192D96E180513764CFEE32487FEA6" box="[567,655,300,321]" name="South Korea" pageId="2" pageNumber="483">KOREA</collectingCountry>
:
<collectingRegion id="A7721CAB6E18051374D6FE88275DFEBB" box="[173,341,327,349]" country="South Korea" name="Chungnam" pageId="2" pageNumber="483">Chungnam Prov.</collectingRegion>
, Daejeon- si,
<collectingMunicipality id="856D48336E180513759BFE882430FEBB" box="[480,568,327,348]" pageId="2" pageNumber="483">Dong-gu</collectingMunicipality>
,
<location id="606984926E180513763FFE8826D7FE9F" LSID="urn:lsid:plazi:treatment:ED1F635F6E1B05157768FB5226F2FB2A:606984926E180513763FFE8826D7FE9F" country="South Korea" latitude="36.32632" longLatPrecision="1" longitude="127.4839" municipality="Dong-gu" name="Sechondong" pageId="2" pageNumber="483" stateProvince="Chungnam">Sechondong</location>
,
<geoCoordinate id="0082B48E6E1805137493FEAF2770FE90" box="[232,376,352,375]" degrees="36" direction="north" minutes="19" orientation="latitude" pageId="2" pageNumber="483" precision="1" seconds="34.76" value="36.32632">N36°19′34.76″</geoCoordinate>
<geoCoordinate id="0082B48E6E1805137506FEAF241FFE90" box="[381,535,352,375]" degrees="127" direction="east" minutes="29" orientation="longitude" pageId="2" pageNumber="483" precision="1" seconds="02.05" value="127.4839">E127°29′02.05″</geoCoordinate>
<quantity id="A24E7FAC6E1805137667FEAD2450FE90" box="[540,600,354,375]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="483" unit="m" value="118.0">
<elevation id="EE9B357A6E1805137667FEAD2450FE90" box="[540,600,354,375]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="483" unit="m" value="118.0">118 m</elevation>
</quantity>
,
<date id="1108F4896E180513761AFEAD26D7FE76" pageId="2" pageNumber="483" value="2020-07-08">
<collectingDate id="014C0D616E180513761AFEAD26D7FE76" pageId="2" pageNumber="483" value="2020-07-08">8 VII 2020</collectingDate>
</date>
, JY Park, YJ Kim, YJ Shin, under bark”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2DAC81C26E18051574BEFE5827F0FD36" lastPageId="4" lastPageNumber="485" pageId="2" pageNumber="483" type="description">
<paragraph id="6509D2496E18051374BEFE58249CFDAA" blockId="2.[173,660,194,616]" pageId="2" pageNumber="483">
<emphasis id="57C20E5B6E18051374BEFE58274AFE4A" bold="true" box="[197,322,407,429]" pageId="2" pageNumber="483">Description.</emphasis>
Body length
<quantity id="A24E7FAC6E18051375AFFE582458FE4B" box="[468,592,407,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="2" pageNumber="483" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">2.8–3.1 mm</quantity>
. Body flattened dorso-ventrally. Head black to dark brown; pronotum black to dark brown; elytra dark brown to brown; abdomen black to dark brown; antennomeres 1–11 reddish-brown; mandibles reddish-brown to brown with dark distal area, maxillary and labial palpi and legs reddish-brown to brown.
</paragraph>
<paragraph id="6509D2496E18051374D6FD9D22A0FD8F" blockId="2.[173,660,194,616]" lastBlockId="2.[706,1193,193,616]" pageId="2" pageNumber="483">
<emphasis id="57C20E5B6E18051374D6FD9D26E2FD80" bold="true" box="[173,234,594,615]" pageId="2" pageNumber="483">Head.</emphasis>
Subtriangular with curved anterior margin, about 1.25–1.43 times as wide as long, about 0.83–0.92 times as wide as pronotum; surface glossy with coarse punctures; disc with one pair of relatively large and elevated antennal tubercles, a pair of deep and large depressions located near inner sides of antennal tubercles, a deep and large fovea located in medio-basal region; eye moderate in size, about 0.31–0.39 times as long as head; gular sutures converged at about apical one-third; antennae long, relative length ratio of each antennomere 24:13:12: 9:10:11:11:10:10:10:16; relative width ratio of each antennomere 12:10:10:10:11:12:13:12:13:13:12.
<emphasis id="57C20E5B6E18051376B8FDCD254BFDF0" bold="true" box="[707,835,514,535]" pageId="2" pageNumber="483">Mouthparts.</emphasis>
Labrum slightly transverse, anterior margin more or less rounded, lateral margin almost straight; mandible without tooth; maxillary palpus elongate, palpomere 4 fusiform, longer than 3; labial
</paragraph>
<caption id="31C982C16E18051374BEF95822A0F94D" ID-DOI="http://doi.org/10.5281/zenodo.13252725" ID-Zenodo-Dep="13252725" box="[197,1192,1687,1706]" httpUri="https://zenodo.org/record/13252725/files/figure.png" pageId="2" pageNumber="483" startId="2.[197,236,1687,1706]" targetBox="[185,1189,671,1648]" targetPageId="2" targetType="figure">
<paragraph id="6509D2496E18051374BEF95822A0F94D" blockId="2.[173,1192,1687,1730]" box="[197,1192,1687,1706]" pageId="2" pageNumber="483">
<emphasis id="57C20E5B6E18051374BEF9582716F94D" bold="true" box="[197,286,1687,1706]" pageId="2" pageNumber="483">Figs. 1–3.</emphasis>
<taxonomicName id="A2B6A9CA6E1805137548F95827FFF94D" authorityName="Park &amp; Ahn" authorityYear="2022" box="[307,503,1687,1706]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="483" phylum="Arthropoda" rank="species" species="rotundifrons" status="sp. nov.">
<emphasis id="57C20E5B6E1805137548F95827FFF94D" box="[307,503,1687,1706]" italics="true" pageId="2" pageNumber="483">Eupiestus rotundifrons</emphasis>
</taxonomicName>
,
<emphasis id="57C20E5B6E180513767EF9582478F94D" bold="true" box="[517,624,1687,1706]" pageId="2" pageNumber="483">
<taxonomicNameLabel id="4CF1B3206E180513767EF9582478F94D" box="[517,624,1687,1706]" pageId="2" pageNumber="483" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id="57C20E5B6E1805137605F9582487F94D" bold="true" box="[638,655,1687,1706]" pageId="2" pageNumber="483">1)</emphasis>
Habitus, holotype, male, 2.8 mm;
<emphasis id="57C20E5B6E18051377B2F95825D2F94D" bold="true" box="[969,986,1687,1706]" pageId="2" pageNumber="483">2)</emphasis>
Superior marginal line
</paragraph>
</caption>
<paragraph id="6509D2496E18051374D6F96025C2F925" blockId="2.[173,1192,1687,1730]" box="[173,970,1711,1730]" pageId="2" pageNumber="483">
(dashed line) of pronotum;
<emphasis id="57C20E5B6E18051375ECF96027A0F925" bold="true" box="[407,424,1711,1730]" pageId="2" pageNumber="483">3)</emphasis>
Dorsolateral ridge (dashed line) of elytra. Scale bars =
<quantity id="A24E7FAC6E18051377F9F96025CEF925" box="[898,966,1711,1730]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="483" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="6509D2496E19051574E3FFB127F0FD36" blockId="3.[152,184,126,145]" lastBlockId="4.[173,660,193,1229]" lastPageId="4" lastPageNumber="485" pageId="3" pageNumber="484">
484 palpus elongate, palpomere 3 suboval, longer than glossy with larger and deeper punctures than head; 2; ligula not divided.
<emphasis id="57C20E5B6E1905127526FF1327A5FF16" bold="true" box="[349,429,220,241]" pageId="3" pageNumber="484">Thorax.</emphasis>
Pronotum moderately elytra
<quantity id="A24E7FAC6E1905127692FF132554FF15" box="[745,860,220,242]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="484" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
long,
<quantity id="A24E7FAC6E19051277E3FF132203FF15" box="[920,1035,220,242]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="3" pageNumber="484" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
wide, slightly narrowed anteriad and distinctly narrowed posteri- wider than long, with large and deep punctures, ad, about 1.20–1.38 times as wide as long, widest posterior margin slightly concave; each elytron with at middle; disc with one pair of long and prominent four longitudinal ridges excluding marginal sutures, keels in sublateral region, two pairs of short keels dorsolateral ridge slightly concave (
<figureCitation id="FD8DCECC6E1905127045FE88228EFEBB" box="[1086,1158,327,349]" captionStart="Figs" captionStartId="2.[197,236,1687,1706]" captionTargetBox="[185,1189,671,1648]" captionTargetId="figure-236@2.[173,1193,661,1670]" captionTargetPageId="2" captionText="Figs. 1–3. Eupiestus rotundifrons, new species. 1) Habitus, holotype, male, 2.8 mm; 2) Superior marginal line" figureDoi="http://doi.org/10.5281/zenodo.13252725" httpUri="https://zenodo.org/record/13252725/files/figure.png" pageId="3" pageNumber="484">Fig. 3</figureCitation>
). in apical half and basal half of median region, su-
<emphasis id="57C20E5B6E19051276D6FEAE251FFE91" bold="true" box="[685,791,353,374]" pageId="3" pageNumber="484">Abdomen.</emphasis>
Slightly narrowed posteriad, finely perior marginal line slightly constricted in middle punctate; surface glossy with coarse punctures. (
<figureCitation id="FD8DCECC6E19051274E4FE5826D6FE4B" box="[159,222,407,429]" captionStart="Figs" captionStartId="2.[197,236,1687,1706]" captionTargetBox="[185,1189,671,1648]" captionTargetId="figure-236@2.[173,1193,661,1670]" captionTargetPageId="2" captionText="Figs. 1–3. Eupiestus rotundifrons, new species. 1) Habitus, holotype, male, 2.8 mm; 2) Superior marginal line" figureDoi="http://doi.org/10.5281/zenodo.13252725" httpUri="https://zenodo.org/record/13252725/files/figure.png" pageId="3" pageNumber="484">Fig. 2</figureCitation>
), large foveae located between keels and
<emphasis id="57C20E5B6E19051276D6FE5824EFFE4B" bold="true" box="[685,743,407,428]" pageId="3" pageNumber="484">Male.</emphasis>
Tergite VIII more or less rounded apically between keel and superior marginal line; surface (
<figureCitation id="FD8DCECC6E19051276CEFE7D24F5FE20" box="[693,765,434,456]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="3" pageNumber="484">Fig. 4</figureCitation>
); sternite VIII triangularly produced medio-apically (
<figureCitation id="FD8DCECC6E1E05157530FF0D278FFF30" box="[331,391,193,216]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 5</figureCitation>
); sternite IX slightly asymmetrical at basal part, rounded apically with several apical setae (
<figureCitation id="FD8DCECC6E1E05157553FF38276AFEEA" box="[296,354,247,269]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 6</figureCitation>
); tergite X more or less rounded apically with several apical setae (
<figureCitation id="FD8DCECC6E1E05157587FEDD2430FEC0" box="[508,568,274,296]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 7</figureCitation>
); median lobe of aedeagus rounded at apex, with rounded basal capsule (
<figureCitation id="FD8DCECC6E1E05157543FE882779FEBB" box="[312,369,327,349]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 8</figureCitation>
); median lobe curved dorsally and narrowed apically in lateral view (
<figureCitation id="FD8DCECC6E1E05157653FEAD246CFE90" box="[552,612,354,376]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 9</figureCitation>
); internal sac with one long curved sclerite in middle and two symmetrical spear-like sclerites dorsally on both sides (
<figureCitation id="FD8DCECC6E1E05157564FE7D2776FE20" box="[287,382,434,456]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Figs. 8–9</figureCitation>
); paramere bilobed, slender, rounded at each apex, with several apical setae, transparent hemispherical sclerites on inner sides of apical 1/7, transparent longitudinal sclerites on inner sides from apical 1/7 to 1/2 (
<figureCitation id="FD8DCECC6E1E051575A4FDD32424FDD6" box="[479,556,540,562]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 10</figureCitation>
).
<emphasis id="57C20E5B6E1E05157639FDD3249AFDD6" bold="true" box="[578,658,540,561]" pageId="4" pageNumber="485">Female.</emphasis>
Tergite VIII similar to that of male (
<figureCitation id="FD8DCECC6E1E05157583FDF82434FDAB" box="[504,572,567,589]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 11</figureCitation>
); sternite VIII broadly prominent medio-apically (
<figureCitation id="FD8DCECC6E1E05157647FD9D248FFD80" box="[572,647,594,616]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 12</figureCitation>
); gonocoxites once-coiled with some minute setae (
<figureCitation id="FD8DCECC6E1E051574CFFD4826F3FD7B" box="[180,251,647,669]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 13</figureCitation>
); tergite X with deep and narrow emargination at basal part, more or less rounded apically with several apical setae (
<figureCitation id="FD8DCECC6E1E051575DFFD7327E3FD36" box="[420,491,700,722]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="31C982C16E19051274CAF9A8252EF925" ID-DOI="http://doi.org/10.5281/zenodo.13252727" ID-Zenodo-Dep="13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="3" pageNumber="484" startId="3.[177,216,1639,1658]" targetBox="[158,1155,502,1612]" targetPageId="3" targetType="figure">
<paragraph id="6509D2496E19051274CAF9A8252EF925" blockId="3.[152,1172,1639,1730]" pageId="3" pageNumber="484">
<emphasis id="57C20E5B6E19051274CAF9A82719F99D" bold="true" box="[177,273,1639,1658]" pageId="3" pageNumber="484">Figs. 4–13.</emphasis>
<taxonomicName id="A2B6A9CA6E190512755CF9A827E0F99D" authorityName="Park &amp; Ahn" authorityYear="2022" box="[295,488,1639,1658]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="species" species="rotundifrons" status="sp. nov.">
<emphasis id="57C20E5B6E190512755CF9A827E0F99D" box="[295,488,1639,1658]" italics="true" pageId="3" pageNumber="484">Eupiestus rotundifrons</emphasis>
</taxonomicName>
,
<emphasis id="57C20E5B6E1905127588F9A82454F99D" bold="true" box="[499,604,1639,1658]" pageId="3" pageNumber="484">
<taxonomicNameLabel id="4CF1B3206E1905127588F9A82454F99D" box="[499,604,1639,1658]" pageId="3" pageNumber="484" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Male:
<emphasis id="57C20E5B6E19051276E5F9A824B8F99D" bold="true" box="[670,688,1639,1658]" pageId="3" pageNumber="484">4)</emphasis>
Tergite VIII, dorsal view;
<emphasis id="57C20E5B6E19051277EFF9A82597F99D" bold="true" box="[916,927,1639,1658]" pageId="3" pageNumber="484">5</emphasis>
) Sternite VIII, ventral view;
<emphasis id="57C20E5B6E19051274E3F9B026A2F975" bold="true" box="[152,170,1663,1682]" pageId="3" pageNumber="484">6)</emphasis>
Sternite IX, ventral view;
<emphasis id="57C20E5B6E19051275F1F9B02793F975" bold="true" box="[394,411,1663,1682]" pageId="3" pageNumber="484">7)</emphasis>
Laterotergal sclerites and tergite X, dorsal view;
<emphasis id="57C20E5B6E1905127747F9B02546F975" bold="true" box="[828,846,1663,1682]" pageId="3" pageNumber="484">8)</emphasis>
Aedeagus, ventral view;
<emphasis id="57C20E5B6E190512705FF9B0223EF975" bold="true" box="[1060,1078,1663,1682]" pageId="3" pageNumber="484">9)</emphasis>
Aedeagus, lateral view;
<emphasis id="57C20E5B6E190512756AF9582727F94D" bold="true" box="[273,303,1687,1706]" pageId="3" pageNumber="484">10)</emphasis>
Paramere, ventral view. Female:
<emphasis id="57C20E5B6E1905127618F9582476F94D" bold="true" box="[611,638,1687,1706]" pageId="3" pageNumber="484">11)</emphasis>
Tergite VIII, dorsal view;
<emphasis id="57C20E5B6E190512770DF958259CF94D" bold="true" box="[886,916,1687,1706]" pageId="3" pageNumber="484">12)</emphasis>
Sternite VIII, ventral view;
<emphasis id="57C20E5B6E19051274E3F96026BEF925" bold="true" box="[152,182,1711,1730]" pageId="3" pageNumber="484">13)</emphasis>
Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="2DAC81C26E1E051574BEFD1827D6FD0A" box="[197,478,727,749]" pageId="4" pageNumber="485" type="distribution">
<paragraph id="6509D2496E1E051574BEFD1827D6FD0A" blockId="4.[173,660,193,1229]" box="[197,478,727,749]" pageId="4" pageNumber="485">
<emphasis id="57C20E5B6E1E051574BEFD182741FD0A" bold="true" box="[197,329,727,749]" pageId="4" pageNumber="485">Distribution.</emphasis>
<collectingCountry id="1DA192D96E1E05157534FD182782FD0B" box="[335,394,727,748]" name="South Korea" pageId="4" pageNumber="485">Korea</collectingCountry>
(South).
</paragraph>
</subSubSection>
<subSubSection id="2DAC81C26E1E051574BEFD3D245CFCDA" pageId="4" pageNumber="485" type="etymology">
<paragraph id="6509D2496E1E051574BEFD3D245CFCDA" blockId="4.[173,660,193,1229]" pageId="4" pageNumber="485">
<emphasis id="57C20E5B6E1E051574BEFD3D2731FCE0" bold="true" box="[197,313,754,775]" pageId="4" pageNumber="485">Etymology.</emphasis>
The name is derived from the Latin
<emphasis id="57C20E5B6E1E051574D6FCC326E4FCC6" box="[173,236,780,801]" italics="true" pageId="4" pageNumber="485">rotund</emphasis>
and
<emphasis id="57C20E5B6E1E05157565FCC32746FCC6" box="[286,334,780,801]" italics="true" pageId="4" pageNumber="485">frons</emphasis>
meaning “rounded front”, which refers to the rounded clypeus of the species.
</paragraph>
</subSubSection>
<subSubSection id="2DAC81C26E1E051574BEFC8D26F2FB2A" pageId="4" pageNumber="485" type="discussion">
<paragraph id="6509D2496E1E051574BEFC8D243BFBA0" blockId="4.[173,660,193,1229]" pageId="4" pageNumber="485">
<emphasis id="57C20E5B6E1E051574BEFC8D272DFCB0" bold="true" box="[197,293,834,855]" pageId="4" pageNumber="485">Remarks.</emphasis>
The new species is similar to
<taxonomicName id="A2B6A9CA6E1E05157643FC8D249BFCB0" authorityName="Fauvel" authorityYear="1902" box="[568,659,834,855]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="485" phylum="Arthropoda" rank="species" species="sikkimi">
<emphasis id="57C20E5B6E1E05157643FC8D249BFCB0" box="[568,659,834,855]" italics="true" pageId="4" pageNumber="485">E. sikkimi</emphasis>
</taxonomicName>
in the body form and structure (head subtriangular with curved anterior margin), body length (around 3.0 mm) and the slightly transverse antennomeres 6–10, characters that the other Palearctic species do not possess. The new species can be distinguished from
<taxonomicName id="A2B6A9CA6E1E05157642FC07249BFC3B" authorityName="Fauvel" authorityYear="1902" box="[569,659,967,989]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="485" phylum="Arthropoda" rank="species" species="sikkimi">
<emphasis id="57C20E5B6E1E05157642FC07249BFC3B" box="[569,659,967,989]" italics="true" pageId="4" pageNumber="485">E. sikkimi</emphasis>
</taxonomicName>
by the slightly constricted superior marginal lines of pronotum at middle (
<figureCitation id="FD8DCECC6E1E05157517FC3327ABFBF6" box="[364,419,1020,1042]" captionStart="Figs" captionStartId="2.[197,236,1687,1706]" captionTargetBox="[185,1189,671,1648]" captionTargetId="figure-236@2.[173,1193,661,1670]" captionTargetPageId="2" captionText="Figs. 1–3. Eupiestus rotundifrons, new species. 1) Habitus, holotype, male, 2.8 mm; 2) Superior marginal line" figureDoi="http://doi.org/10.5281/zenodo.13252725" httpUri="https://zenodo.org/record/13252725/files/figure.png" pageId="4" pageNumber="485">Fig. 2</figureCitation>
), the slightly concave dorsolateral ridges of elytra (
<figureCitation id="FD8DCECC6E1E051575F6FBD827C9FBCB" box="[397,449,1047,1069]" captionStart="Figs" captionStartId="2.[197,236,1687,1706]" captionTargetBox="[185,1189,671,1648]" captionTargetId="figure-236@2.[173,1193,661,1670]" captionTargetPageId="2" captionText="Figs. 1–3. Eupiestus rotundifrons, new species. 1) Habitus, holotype, male, 2.8 mm; 2) Superior marginal line" figureDoi="http://doi.org/10.5281/zenodo.13252725" httpUri="https://zenodo.org/record/13252725/files/figure.png" pageId="4" pageNumber="485">Fig. 3</figureCitation>
), and the different form and structure of the aedeagus (
<figureCitation id="FD8DCECC6E1E051575B8FBFD242FFBA0" box="[451,551,1074,1096]" captionStart="Figs" captionStartId="3.[177,216,1639,1658]" captionTargetBox="[158,1155,502,1612]" captionTargetId="figure-157@3.[151,1171,494,1623]" captionTargetPageId="3" captionText="Figs. 4–13. Eupiestus rotundifrons, new species. Male: 4) Tergite VIII, dorsal view; 5) Sternite VIII, ventral view; 6) Sternite IX, ventral view; 7) Laterotergal sclerites and tergite X, dorsal view; 8) Aedeagus, ventral view; 9) Aedeagus, lateral view; 10) Paramere, ventral view. Female: 11) Tergite VIII, dorsal view; 12) Sternite VIII, ventral view; 13) Abdominal segments IX (right) and X; dorsal view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13252727" httpUri="https://zenodo.org/record/13252727/files/figure.png" pageId="4" pageNumber="485">Figs. 8–10</figureCitation>
).
</paragraph>
<paragraph id="6509D2496E1E051574BEFB8326F2FB2A" blockId="4.[173,660,193,1229]" pageId="4" pageNumber="485">
Formerly, the northernmost record of the genus
<taxonomicName id="A2B6A9CA6E1E051574D6FBA72701FB9A" box="[173,265,1128,1149]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="57C20E5B6E1E051574D6FBA72701FB9A" box="[173,265,1128,1149]" italics="true" pageId="4" pageNumber="485">Eupiestus</emphasis>
</taxonomicName>
was
<collectingRegion id="A7721CAB6E1E0515754CFBA82760FB9A" box="[311,360,1127,1149]" country="China" name="Xizang" pageId="4" pageNumber="485">Tibet</collectingRegion>
(
<taxonomicName id="A2B6A9CA6E1E0515750FFBA727F3FB9B" box="[372,507,1127,1149]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="485" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="57C20E5B6E1E0515750FFBA727F3FB9B" box="[372,507,1127,1149]" italics="true" pageId="4" pageNumber="485">E. longipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="A2B6A9CA6E1E0515767DFBA7246AFB9B" authorityName="Fauvel" authorityYear="1902" box="[518,610,1127,1149]" class="Insecta" family="Staphylinidae" genus="Eupiestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="485" phylum="Arthropoda" rank="species" species="sikkimi">
<emphasis id="57C20E5B6E1E0515767DFBA7246AFB9B" box="[518,610,1127,1149]" italics="true" pageId="4" pageNumber="485">E. sikkimi</emphasis>
</taxonomicName>
), but now its distributional range is extended farther north (Korean peninsula) by the discovery of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>