treatments-xml/data/D6/1B/72/D61B722AD0EE5A56BA6F007362EC9651.xml

<document id="ED27FD230BC9C5076F9FF1E48FFCDE35" ID-DOI="10.3897/zse.100.123213" ID-ZooBank="CF91D7F2-FC99-431C-AD78-6C5CB1DB1E9D" ID-publisher-id="123213" URI-arpha="9D419344-945D-50CF-8F7C-FFC200DF6680" URI-zoobank="http://zoobank.org/CF91D7F2-FC99-431C-AD78-6C5CB1DB1E9D" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1721914003814" checkinUser="pensoft" docAuthor="Jeong, Kyung-Hoon, Harms, Danilo &amp; Yoo, Jung-Sun" docDate="2024" docId="D61B722AD0EE5A56BA6F007362EC9651" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 1053-1060" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/123213/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Nipponogarypus seosanensis Jeong &amp; Harms 2024, sp. nov." docType="treatment" docUuid="2CFFDE30-7D05-4E5B-9006-E57EE69E780F" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="1060" masterDocId="9D419344945D50CF8F7CFFC200DF6680" masterDocTitle="The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea" masterLastPageNumber="1060" masterPageNumber="1053" pageNumber="1053" updateTime="1721920860098" updateUser="admin">
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<mods:title id="C2E23B2999B43F366EAAC1BE3BEA23E9">The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea</mods:title>
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<mods:namePart id="402A09B25F64366E7097F3A8648CE514">Jeong, Kyung-Hoon</mods:namePart>
<mods:affiliation id="ED66B8A07257852EBC0C287002EEE307">Seoul National University, 1, Gwanak-ro, Gwanak-gu, Seoul, 08826, Republic of Korea &amp; National Institute of Biological Resources, Species Diversity Research Division, Environmental Research Complex, Hwangyeong-ro 42, Seo-gu, Incheon, 22689 Republic of Korea &amp; Department of Agricultural Convergence Technology, Jeonbuk National University, Jeonju, Republic of Korea</mods:affiliation>
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<mods:namePart id="D22E755161B46D43EE2C5F032023A0B7">Harms, Danilo</mods:namePart>
<mods:nameIdentifier id="EAF9193FDFABDD310A3BF660A274481B" type="ORCID">0000-0002-7189-5345</mods:nameIdentifier>
<mods:affiliation id="F98963C57F6F05E65CFDE85D55CB16A7">Lab of Insect Phylogenetics &amp; Evolution, Department of Plant Protection &amp; Quarantine, Jeonbuk National University, Jeonju, Republic of Korea &amp; Museum of Nature Hamburg – Zoology, Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King-Platz 3, Hamburg, 20146, Germany &amp; Harry Butler Institute, Murdoch University, Murdoch, Australia</mods:affiliation>
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<mods:namePart id="17071E2F83F9DD424FB2F82F90E4AD0B">Yoo, Jung-Sun</mods:namePart>
<mods:nameIdentifier id="A2556140E75AC685DBD84326C2250E43" type="ORCID">0000-0002-3243-2006</mods:nameIdentifier>
<mods:affiliation id="4FF383479C0BBBD63A190A20CC05420A">National Institute of Biological Resources, Species Diversity Research Division, Environmental Research Complex, Hwangyeong-ro 42, Seo-gu, Incheon, 22689 Republic of Korea</mods:affiliation>
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<mods:title id="654105A1FFE462BDE5623CB9A3B95F95">Zoosystematics and Evolution</mods:title>
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<mods:date id="B79541A9F3F8AB5290C30325513D981D">2024</mods:date>
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<mods:number id="E05B5FA5F73C69DACDF892CDE45CB6A5">100</mods:number>
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<treatment id="D61B722AD0EE5A56BA6F007362EC9651" ID-DOI="http://doi.org/10.5281/zenodo.12820291" ID-Zenodo-Dep="12820291" ID-arpha="D61B722A-D0EE-5A56-BA6F-007362EC9651" ID-zoobank="https://zoobank.org/2CFFDE30-7D05-4E5B-9006-E57EE69E780F" LSID="urn:lsid:zoobank.org:act:2CFFDE30-7D05-4E5B-9006-E57EE69E780F" LSID-ZBK="urn:lsid:zoobank.org:act:2CFFDE30-7D05-4E5B-9006-E57EE69E780F" httpUri="http://treatment.plazi.org/id/D61B722AD0EE5A56BA6F007362EC9651">
<subSubSection id="D748B552F6AE0F112F9449470FB13FF0" type="nomenclature">
<paragraph id="6B216C72845D23347072E54C7BD23B38">
<taxonomicName id="84A3177C1DB33107C47E611F85DC731A" authority="Jeong &amp; Harms" authorityName="Jeong &amp; Harms" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="1A6BDA581341B1C9C33ECBC5CDC74EEB" italics="true">Nipponogarypus seosanensis</emphasis>
Jeong &amp; Harms
</taxonomicName>
<taxonomicNameLabel id="E107B2DD63506B960F37EA46924E8B65" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFLAE" type="type material">
<paragraph id="252A6E6F4A38B772FE31D3BC5E9F8B15">
<heading id="46ED66AC4F05E71C111AA7AFD2DE7D4F" reason="title">Type material.</heading>
</paragraph>
<paragraph id="BE82A98CF6AA8A8AD5B5BA56F0A74D82">
<materialsCitation id="2BC0C3AB9AAFC9D1D8D89C7766300D61" collectingDate="2022-07" collectionCode="NUHGIV" collectorName="KH Jeong" country="South Korea" county="Seosan-si" latitude="36.601543" location="Female" longLatPrecision="1" longitude="128.1746" municipality="Buseok-myeon" specimenCode="NUHGIV 0000001225" specimenCount="1" stateProvince="Chungcheongnam-province" typeStatus="holotype">
<emphasis id="6C6FFDC53CBD78C40A844C75B6A8A3DF" bold="true">
<emphasis id="90AB97F5BD5CA705633E1189E819BE61" bold="true" italics="true">
<typeStatus id="83B98AFE1CD909950935E99B5A68C66A" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<location id="344220BA4E47D4C234BB53202E724088" LSID="urn:lsid:plazi:treatment:D61B722AD0EE5A56BA6F007362EC9651:344220BA4E47D4C234BB53202E724088" country="South Korea" county="Seosan-si" latitude="36.601543" longLatPrecision="1" longitude="128.1746" municipality="Buseok-myeon" name="Female" stateProvince="Chungcheongnam-province">Female</location>
(
<specimenCode id="604AA89A00CE051293F2F4D0CACACF44">NUHGIV 0000001225</specimenCode>
). 
<collectingCountry id="84B2E019BFA3F5B392FA78CF14F58866" name="South Korea">Korea</collectingCountry>
: 
<collectingRegion id="3B4952C0FA02FF404DA66D959301E06D">Chungcheongnam-province</collectingRegion>
: 
<location id="E084E61942A5275764E29DFD1AB26D3C" LSID="urn:lsid:plazi:treatment:D61B722AD0EE5A56BA6F007362EC9651:E084E61942A5275764E29DFD1AB26D3C" country="South Korea" county="Seosan-si" latitude="36.601543" longLatPrecision="1" longitude="128.1746" municipality="Buseok-myeon" name="Ganwoldo" stateProvince="Chungcheongnam-province">Ganwoldo</location>
1 – gil, 
<collectingMunicipality id="817C2EFB488801151AA58AA7AA1BF34B">Buseok-myeon</collectingMunicipality>
, 
<collectingCounty id="E5A2CDB67737AE484F3A92EA5E4462CA">Seosan-si</collectingCounty>
, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETLAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[128.174594,36.601544]}">
<geoCoordinate id="507F3433CA9F2144F1540B90C7BAA20F" degrees="36" direction="north" minutes="36" orientation="latitude" precision="1" seconds="5.56" value="36.601543">36 ° 36 ' 5.56 &quot; N</geoCoordinate>
, 
<geoCoordinate id="E244A61A98D9B2ABE35D5A677A962C56" degrees="128" direction="east" minutes="10" orientation="longitude" precision="1" seconds="28.54" value="128.1746">128 ° 10 ' 28.54 &quot; E</geoCoordinate>
</named-content>
, 03, 
<date id="3EA6C4275847F82269DEE3B5445DD5A8" value="2022-07">
<collectingDate id="1653DBED66E2C850A47DB05443095E08" value="2022-07">Jul 2022</collectingDate>
</date>
, 
<collectorName id="5A293E94076826BC6D52B3A6EA3B4E15">KH Jeong</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="F45A770DC8BBC134F3FED356699A9D65">
<materialsCitation id="D1F52F05211385FA57A7C24A3D180757" collectingDate="2022-07" collectionCode="NUHGIV" collectorName="KH Jeong" country="South Korea" county="Seosan-si" latitude="36.601543" location="Female" longLatPrecision="1" longitude="128.1746" municipality="Buseok-myeon" specimenCode="NUHGIV 0000001228" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chungcheongnam-province" typeStatus="paratype">
<emphasis id="5326C92B36E8624976004FDFD7D303E3" bold="true" italics="true">
<typeStatus id="30B753D78F3340C0299EAC953472487A" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="50D8A312A30150250DADA1063CC5B6CC" count="1" type="female">One female</specimenCount>
and 
<specimenCount id="FA5DAEF2A1B2B300F216F810D3A37548" count="2" type="male">two males</specimenCount>
(
<specimenCode id="782C3A981E9724738E358FE03B28A6DC">NUHGIV 0000001228</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6LAE" type="etymology">
<paragraph id="88D34A8B978336C1FB943ADE23B77726">
<heading id="1154B2CB1AA567A2700ED4BF92D91ADF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="8ABF0DEC977FB5B1714DCACAFBDFB423">
This species is named after the 
<typeStatus id="014BE17A3507231E61BC6F477AF21738">type</typeStatus>
locality, Seosan-si, 
<collectingRegion id="23C51D89E6B1DB41A811656FF4516B68" country="South Korea" name="Chungnam">Chungcheongnam-do</collectingRegion>
, in 
<collectingCountry id="8E3C386988B03BAE85A0D6299A02BBA5" name="South Korea">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEMAE" type="habitat">
<paragraph id="5E6C8CA6171EE9BC611A3FF9FE4BC09C">
<heading id="0D008B2DC435D68B493347A71130BDCD" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="E86A13D89D57395A7F48CC0A00FA265F">
This species was collected from cracks and fissures of moist rocks in the supralittoral zone, right next to the shoreline (Fig. 
<figureCitation id="E8FE98F63281D6B563BE6499C02ADA04" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of Nipponogarypus seosanensis sp. nov. in the littoral zone of Seosan-si, Chungcheongnam-do, South Korea." figureDoi="10.3897/zse.100.123213.figure2" httpUri="https://binary.pensoft.net/fig/1102003">2</figureCitation>
).
</paragraph>
<caption id="D50F953EE9AA19A05454D038BA89CF0D" ID-DOI="10.3897/zse.100.123213.figure2" ID-arpha="B237468A-4280-5FAB-9836-48C6CC970D2C" httpUri="https://binary.pensoft.net/fig/1102003" startId="F2">
<paragraph id="483D743611E8820B299759D85053E8C1">
<label id="0A5339AAB9975EACFF5E3151531C9160">Figure 2.</label>
</paragraph>
<paragraph id="233ED23CD09201B3406B35BCF51C5E69">
Habitat of 
<taxonomicName id="9F2F0E716D7CD43F48AF366BB08F58D4" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="837EA75560658EA36E3C3F7D7D531842" italics="true">Nipponogarypus seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEE616F5F2BB2434BF3DAFB7B18F2949" rank="species">sp. nov.</taxonomicNameLabel>
in the littoral zone of Seosan-si, Chungcheongnam-do, South Korea.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGNAE" type="diagnosis">
<paragraph id="D4341DEAFCB06A58310A4522B3F3391F">
<heading id="90600F6B56D948BF360B18B36528EF53" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B8C6B85187D73E2A972B79ED921E8880">
This species is most similar to 
<taxonomicName id="387004C2047B562018832EFAFFAC82CE" authorityName="sensu Morikawa" authorityYear="1955" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="enoshimaensis">
<emphasis id="F6E77978FB19350806B58143331C0C4F" italics="true">N. enoshimaensis</emphasis>
</taxonomicName>
by having similar L / W ratios in the pedipalpal femur (0.56–0.61 / 
<quantity id="DDD85D9A7A83DCBAAAEE6B1593659AB4" metricMagnitude="-4" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.7999999999999998" unit="mm" value="0.19" valueMax="0.2" valueMin="0.18">0.18 – 0.20 mm</quantity>
in 
<taxonomicName id="EEFAFF2621C4F7CCA63389A6855B6F07" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="AEC3E6D4E14CED7E5506C7AC30D31DBF" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8C5E142A92EF669B1C51291AE2C796A8" rank="species">sp. nov.</taxonomicNameLabel>
and 0.52–0.59 / 
<quantity id="3E73118A7E360765FFF787AC140D312E" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" unit="mm" value="0.18">0.18 mm</quantity>
in 
<taxonomicName id="D51138F0616593B1F9386325E93B439B" authorityName="sensu Morikawa" authorityYear="1955" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="enoshimaensis">
<emphasis id="41456F7443978028F9745DFAE44F91DC" italics="true">N. enoshimaensis</emphasis>
</taxonomicName>
) and pedipalpal patella (0.53–0.59 / 
<quantity id="172B70D3449FA0465E4A7533E0C284EB" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.21 – 0.23 mm</quantity>
in 
<taxonomicName id="43851C257AF311107747BFF4A18303ED" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="3BD7679F1360F0376A4B2979940F2EA7" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65A61190C144B74B1CF8C6E843C8E343" rank="species">sp. nov.</taxonomicNameLabel>
and 0.57 / 
<quantity id="D7ECFF308FFC46D64381A75672781047" metricMagnitude="-4" metricUnit="m" metricValue="2.2" unit="mm" value="0.22">0.22 mm</quantity>
in 
<taxonomicName id="47A43BBF16F809F7A18882A78B29D8EB" authorityName="sensu Morikawa" authorityYear="1955" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="enoshimaensis">
<emphasis id="54406944B689F06CB46F906DAC51DBAF" italics="true">N. enoshimaensis</emphasis>
</taxonomicName>
). Both species can easily be distinguished by the number of marginal teeth on the fixed chelal finger (
<quantity id="22E703D9BA93035350893A915D730CC4" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.0922" metricValueMin="1.016" unit="in" value="41.5" valueMax="43.0" valueMin="40.0">40–43 in</quantity>
<taxonomicName id="58AB05FDFE3A1CB84327C31909D4E62B" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="EA61242A419DE1335882FCE33310179F" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1C2A74F16F735DC0F656AC36F82E5165" rank="species">sp. nov.</taxonomicNameLabel>
, 
<quantity id="E2C6676A03B63CB74C3DD54503698296" metricMagnitude="0" metricUnit="m" metricValue="1.27" unit="in" value="50.0">50 in</quantity>
<taxonomicName id="EB505380B961AEB1715CA7966E19A14B" authorityName="sensu Morikawa" authorityYear="1955" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="enoshimaensis">
<emphasis id="7DF07D7E7464C7C0785EB5EFE30492CE" italics="true">N. enoshimaensis</emphasis>
</taxonomicName>
). 
<taxonomicName id="332CF67B8530C6167176FA5D1C7C96ED" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="0AD3CBB3F50FC15F8C17AAF9A3F726A1" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="829CAEED7756D41105AA0068C92ACF4B" rank="species">sp. nov.</taxonomicNameLabel>
can further be distinguished from 
<taxonomicName id="235C79B7FFC99AEEF191273AB937A7B3" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" baseAuthorityName="Morikawa" baseAuthorityYear="1960" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="okinoerabensis" status="stat. nov.">
<emphasis id="28405EAF150B512258E7306523EB38C5" italics="true">N. okinoerabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="224E5026FB01E777BCF77FE8F2E03683">stat. nov.</taxonomicNameLabel>
by having a longer pedipalpal femur (
<quantity id="E51234502E89B14FF069312A496AB7D8" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" unit="mm" value="0.585" valueMax="0.61" valueMin="0.56">0.56–0.61 mm</quantity>
in 
<taxonomicName id="01474ADE72AD3F9302CE56BDDD7DCCA0" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="9D2067FE98C7171F145A442E2DF0DB24" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DCF8956D11D017CB51ADA12CE0A19414" rank="species">sp. nov.</taxonomicNameLabel>
and 
<quantity id="F02EFBD7C3C0B1E57B48CD12DDC09CF1" metricMagnitude="-4" metricUnit="m" metricValue="4.8" unit="mm" value="0.48">0.48 mm</quantity>
in 
<taxonomicName id="F37EA3D2B647C2B895516951D863D318" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" baseAuthorityName="Morikawa" baseAuthorityYear="1960" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="okinoerabensis" status="stat. nov.">
<emphasis id="367CCE1F235D0E15EF4A9CB9C342AE5B" italics="true">N. okinoerabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7DD50689D8D81EA061286AEDAD0C7CC7">stat. nov.</taxonomicNameLabel>
) and patella (
<quantity id="58DB1764D11CD0CDD79FA517C260C5BC" metricMagnitude="-4" metricUnit="m" metricValue="5.6000000000000005" metricValueMax="5.8999999999999995" metricValueMin="5.300000000000001" unit="mm" value="0.56" valueMax="0.59" valueMin="0.53">0.53–0.59 mm</quantity>
in 
<taxonomicName id="1128A0C2F3438F5166F1D6C2B6A4033D" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="C1805C1737823C702794F7995D7EC025" italics="true">N. seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0D5D458296030FF682C77201EC35BBC" rank="species">sp. nov.</taxonomicNameLabel>
and 
<quantity id="4E054E6D470760F44E2B6DA3B0A1B2C6" metricMagnitude="-4" metricUnit="m" metricValue="4.5" unit="mm" value="0.45">0.45 mm</quantity>
in 
<taxonomicName id="37E2F2071D2AD40B8D34AD776E1F349C" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" baseAuthorityName="Morikawa" baseAuthorityYear="1960" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="okinoerabensis" status="stat. nov.">
<emphasis id="A23C553C40D8DF8BB06D3D6F89D95294" italics="true">N. okinoerabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50BF46F323B5A2B2AB68465C33294869">stat. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2RAE" type="description">
<paragraph id="C8F6FAA43CA2C6A629CFD84AA03CD2CA">
<heading id="D38DF77A34DDBF986251A90D2B74FC6D" reason="title">Description.</heading>
</paragraph>
<paragraph id="ECAF1E921D0DC33E2DE2F9D340E2A0C8">
Female, adult (
<typeStatus id="83572444A66FAA446954374EB3BB9261" type="holotype">holotype</typeStatus>
) (Fig. 
<figureCitation id="38F918870CA349FB7A41E2B8B8031E3C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Nipponogarypus seosanensis sp. nov. A. Female holotype, dorsal view; B. Female holotype, ventral view; C. Male paratype, dorsal view; D. Male paratype, ventral view." figureDoi="10.3897/zse.100.123213.figure3" httpUri="https://binary.pensoft.net/fig/1102004">3 A, B</figureCitation>
).
</paragraph>
<caption id="4AC20C25F65784A67F9DF972AF14700F" ID-DOI="10.3897/zse.100.123213.figure3" ID-arpha="A008C6C1-979E-585E-A148-F6EB3262C7EA" httpUri="https://binary.pensoft.net/fig/1102004" startId="F3">
<paragraph id="055C0015A9EE59F27D83EF30923524E0">
<label id="62C19A930341D0A1BC71A099C8A32EAB">Figure 3.</label>
</paragraph>
<paragraph id="3AA15879DFFF48BA58A473699DA17224">
Habitus of 
<taxonomicName id="11A7D8E816B4A748A5FEBFC7BA329495" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="06AE9918A4E5B42670C92DA2152FEBB1" italics="true">Nipponogarypus seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="247E23E06ACC8FBE5FB689D6CAFF04E3" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="4C35E15C65A2B57899B6EBB8FCED3385" bold="true">A.</emphasis>
Female holotype, dorsal view; 
<emphasis id="74958237A21296437FC5E2D85C2547FC" bold="true">B.</emphasis>
Female holotype, ventral view; 
<emphasis id="47A0C75F872F07598D8E0F7C6E0CDF2C" bold="true">C.</emphasis>
Male paratype, dorsal view; 
<emphasis id="65691CFA7466E2A4FBCE581DF4FF3A38" bold="true">D.</emphasis>
Male paratype, ventral view.
</paragraph>
</caption>
<paragraph id="93595E0D19F344BBC98CD49871E7E5E2">Color. Blackish-brown, glossy; the ventral surface darker than the end of the body appendages; the coxal region reddish-orange.</paragraph>
<paragraph id="6F856B01E1CD9B43EDD9B7344BFE6515">
Cephalothorax (Fig. 
<figureCitation id="01B5A79BF2A2A603AA436A9CF8D5D3CB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Appendages of Nipponogarypus seosanensis sp. nov., holotype female. A. Carapace; B. Chelicera in dorsal view; C. Left pedipalp in dorsal view; D. Right chela from lateral view; E. Coxa; F. Right leg I; G. Right leg II; H. Female genital sternites; I. Right leg IV; J. Male genital sternites. Scale bars: 0.5 mm (A, C – G, I); 0.2 mm (H, J); 0.1 mm (B)." figureDoi="10.3897/zse.100.123213.figure4" httpUri="https://binary.pensoft.net/fig/1102005">4 A, H</figureCitation>
). Carapace 1.18 times longer than the broad; carapace sub-rectangular; four conspicuous eyes; two transverse furrows on the carapace; ten lyrifissures; first furrow situated in medial position on the carapace, second furrow near the posterior margin of the carapace; carapacal chaetotaxy 4–2: 22; setae short and acuminate. Pedipalpal coxa with 10 setae; coxal chaetotaxy 5: 5: 6: 13; one lyrifissure on coxa I – III, two lyrifissures on coxa IV.
</paragraph>
<caption id="412F2F8F9B0EEC5293167A64A73B4649" ID-DOI="10.3897/zse.100.123213.figure4" ID-arpha="969CEB83-A170-50D6-8047-71CD7E096815" httpUri="https://binary.pensoft.net/fig/1102005" startId="F4">
<paragraph id="93B83F31F14FC76D851EA64E94AEB796">
<label id="C440932B08835A1397C9696823835FAE">Figure 4.</label>
</paragraph>
<paragraph id="40AA795E99CDEEDDBE6F4A52DCAB2726">
Appendages of 
<taxonomicName id="B56B41D1E2D09137E30873CC247DBA8B" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="CA176678461EC9489DFE981DB696547B" italics="true">Nipponogarypus seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3387023A98FC896596E6C43C65443358" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female. 
<emphasis id="956D78330428A61B7452E2713C3030E6" bold="true">A.</emphasis>
Carapace; 
<emphasis id="5B5C56AF5C1FC6B860D68A4E4EDAE68F" bold="true">B.</emphasis>
Chelicera in dorsal view; 
<emphasis id="6CB923BBAD80E09491EAF7E5FAA0EEA3" bold="true">C.</emphasis>
Left pedipalp in dorsal view; 
<emphasis id="45F55EAC8CA879FC6581126133C770DF" bold="true">D.</emphasis>
Right chela from lateral view; 
<emphasis id="FF2D0D64D8B11A062CFEF8F75BDCC188" bold="true">E.</emphasis>
Coxa; 
<emphasis id="42C85D81DF6500D941F74E7CD92B2DF0" bold="true">F.</emphasis>
Right leg I; 
<emphasis id="357989301B6D4037272FAEA5A87FFD04" bold="true">G.</emphasis>
Right leg II; 
<emphasis id="76AF2D373FD35D568C07F6C04854A40C" bold="true">H.</emphasis>
Female genital sternites; 
<emphasis id="3685D5B8E2119AA73AB103980B058A9D" bold="true">I.</emphasis>
Right leg IV; 
<emphasis id="D42F92C8091DF88FC061F0CFEE7E94DF" bold="true">J.</emphasis>
Male genital sternites. Scale bars: 0.5 mm (
<emphasis id="007F9C2512FF5266F96ECEBF92FB2E88" bold="true">A, C</emphasis>
– 
<emphasis id="D78C5C23439AAD0971083D3CE838F96F" bold="true">G, I</emphasis>
); 0.2 mm (
<emphasis id="888D3DE8A73548464DB50FF2C162B0D1" bold="true">H, J</emphasis>
); 0.1 mm (
<emphasis id="500868C4FE6C41140E39AB59E60AD0B0" bold="true">B</emphasis>
).
</paragraph>
</caption>
<paragraph id="571A7A21BD238D7DCB44724EC67130BF">
Chelicera (Figs 
<figureCitation id="495065AFAB5159364C61AABCD9251041" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Appendages of Nipponogarypus seosanensis sp. nov., holotype female. A. Carapace; B. Chelicera in dorsal view; C. Left pedipalp in dorsal view; D. Right chela from lateral view; E. Coxa; F. Right leg I; G. Right leg II; H. Female genital sternites; I. Right leg IV; J. Male genital sternites. Scale bars: 0.5 mm (A, C – G, I); 0.2 mm (H, J); 0.1 mm (B)." figureDoi="10.3897/zse.100.123213.figure4" httpUri="https://binary.pensoft.net/fig/1102005">4 C</figureCitation>
, 
<figureCitation id="55F397FF433F2CB159B3A1A2F4C1A05B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Appendages of Nipponogarypus seosanensis sp. nov., paratype female. A. Galea; B. Left chela, lateral view; C. Left chelal fingers, lateral view, sensory setae in the white box; D. Right leg I; E. Claws and arolium of right leg I; F. Pleural membrane. Scale bars: 0.02 mm (A); 0.3 mm (B); 0.1 mm (C, D, F); 0.05 mm (E)." figureDoi="10.3897/zse.100.123213.figure5" httpUri="https://binary.pensoft.net/fig/1102006">5 A</figureCitation>
). Cheliceral margin smooth; five setae on the cheliceral hand, one seta on the movable finger; galea long and shortly three-branched at the tip in both sexes; serrula exterior with 19 blades, rallum with three blades. Two lyrifissures on the hand.
</paragraph>
<paragraph id="74A9B9DBD6A77A695DB2D4FEF439518F">
Pedipalp (Figs 
<figureCitation id="C5A7E31A820D7BA7376B776BB1531713" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Appendages of Nipponogarypus seosanensis sp. nov., holotype female. A. Carapace; B. Chelicera in dorsal view; C. Left pedipalp in dorsal view; D. Right chela from lateral view; E. Coxa; F. Right leg I; G. Right leg II; H. Female genital sternites; I. Right leg IV; J. Male genital sternites. Scale bars: 0.5 mm (A, C – G, I); 0.2 mm (H, J); 0.1 mm (B)." figureDoi="10.3897/zse.100.123213.figure4" httpUri="https://binary.pensoft.net/fig/1102005">4 B</figureCitation>
, 
<figureCitation id="3ABBBCFBED13326F6DCAA54870D1BB3B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Appendages of Nipponogarypus seosanensis sp. nov., paratype female. A. Galea; B. Left chela, lateral view; C. Left chelal fingers, lateral view, sensory setae in the white box; D. Right leg I; E. Claws and arolium of right leg I; F. Pleural membrane. Scale bars: 0.02 mm (A); 0.3 mm (B); 0.1 mm (C, D, F); 0.05 mm (E)." figureDoi="10.3897/zse.100.123213.figure5" httpUri="https://binary.pensoft.net/fig/1102006">5 B, C</figureCitation>
). Trochanter 1.60, femur 2.95, patella 2.46, chela 3.24 times longer than the broad, movable finger 0.99 times longer than the hand. Two pseudotactile setae on the femur. Sensory setae present between 
<abbrev id="ABBRID0EVVAE" xlink_title="external terminal">
<emphasis id="2940E901BE1CF206D14588EEE3F3971C" italics="true">et</emphasis>
</abbrev>
and 
<emphasis id="9C79EE871A54EE6E9C6AC578D749D173" italics="true">
<abbrev id="ABBRID0E1VAE" xlink_title="internal subbasal">isb</abbrev>
. b
</emphasis>
and 
<abbrev id="ABBRID0EAWAE" xlink_title="subbasal">
<emphasis id="A8B285F2DAFEF7E2558B4692EF1F9C4D" italics="true">sb</emphasis>
</abbrev>
on the basal of the finger; 
<abbrev id="ABBRID0EFWAE" xlink_title="subterminal">
<emphasis id="5F180DF94BC137BD253DD20AB8E77E2C" italics="true">st</emphasis>
</abbrev>
on the middle of the 
<emphasis id="71E05F80F7207AD4069498AB57BB95EA" italics="true">t</emphasis>
and 
<emphasis id="DAB481B461AA46B3DB4EA2F329B84339" italics="true">
b. 
<abbrev id="ABBRID0ENWAE" xlink_title="internal basal">ib</abbrev>
</emphasis>
, 
<abbrev id="ABBRID0ESWAE" xlink_title="external subbasal">
<emphasis id="E33574736A82622B63CE353E79578834" italics="true">esb</emphasis>
</abbrev>
, and 
<abbrev id="ABBRID0EXWAE" xlink_title="external basal">
<emphasis id="363B5B96EA63D3D3A649930E1F9ABCDE" italics="true">eb</emphasis>
</abbrev>
grouped at the base of the finger; 
<abbrev id="ABBRID0E3WAE" xlink_title="external subterminal">
<emphasis id="2DE0F3112D6A7366AFAD09F77CB3C4C0" italics="true">est</emphasis>
</abbrev>
on the middle of 
<abbrev id="ABBRID0EBXAE" xlink_title="external terminal">
<emphasis id="C90D56B4D88C088BDA2AD7BE1F76789C" italics="true">et</emphasis>
</abbrev>
and 
<abbrev id="ABBRID0EGXAE" xlink_title="external subbasal">
<emphasis id="3DAEEF9521EC9CE6F527DFA4B9A2AA02" italics="true">esb</emphasis>
</abbrev>
; 
<abbrev id="ABBRID0ELXAE" xlink_title="internal terminal">
<emphasis id="B98998144CFF46DDADE8DCD50C8B9063" italics="true">it</emphasis>
</abbrev>
and 
<abbrev id="ABBRID0EQXAE" xlink_title="internal basal">
<emphasis id="990132C075F5F4F75FD9ADF65FECA30A" italics="true">ib</emphasis>
</abbrev>
on the dorsal surface of the fixed finger, 
<abbrev id="ABBRID0EVXAE" xlink_title="internal terminal">
<emphasis id="1A92B28FB479C24774A45BAE96E37BBC" italics="true">it</emphasis>
</abbrev>
on the middle of the finger, and 
<abbrev id="ABBRID0E1XAE" xlink_title="internal basal">
<emphasis id="C95AEC2C51A7C32DE0E1EF11F35705B3" italics="true">ib</emphasis>
</abbrev>
on the basal of the finger; 
<abbrev id="ABBRID0E6XAE" xlink_title="internal subterminal">
<emphasis id="1573696B782B6A936C86D612BF2CA8AD" italics="true">ist</emphasis>
</abbrev>
on the paraxial surface of the chela, between 
<abbrev id="ABBRID0EEYAE" xlink_title="internal subbasal">
<emphasis id="70EAF0F654D21D8D1E202C2463DA7638" italics="true">isb</emphasis>
</abbrev>
and 
<abbrev id="ABBRID0EJYAE" xlink_title="external subterminal">
<emphasis id="3E750F300D21ACE77F23DFF08BF3D1AE" italics="true">est</emphasis>
</abbrev>
, but closer to 
<abbrev id="ABBRID0EOYAE" xlink_title="internal subbasal">
<emphasis id="FE1E2F87B81D7AF958DC62441E5F0445" italics="true">isb</emphasis>
</abbrev>
; short venom ducts in both fingers, not extending past trichobothrium 
<abbrev id="ABBRID0ETYAE" xlink_title="external terminal">
<emphasis id="48D73CCDB1FEDBBB2E34A1354DABBEF8" italics="true">et</emphasis>
</abbrev>
.
</paragraph>
<paragraph id="F09F7B03D73D741E634D551238043090">Marginal teeth small and contiguous; triangular teeth on both fingers; fixed finger with 40, movable finger with 43 marginal teeth.</paragraph>
<paragraph id="705D395112A86320FC5932CD02442EF8">
Legs (Figs 
<figureCitation id="9BE87C2F2AF3FD0687D93257543919CB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Appendages of Nipponogarypus seosanensis sp. nov., holotype female. A. Carapace; B. Chelicera in dorsal view; C. Left pedipalp in dorsal view; D. Right chela from lateral view; E. Coxa; F. Right leg I; G. Right leg II; H. Female genital sternites; I. Right leg IV; J. Male genital sternites. Scale bars: 0.5 mm (A, C – G, I); 0.2 mm (H, J); 0.1 mm (B)." figureDoi="10.3897/zse.100.123213.figure4" httpUri="https://binary.pensoft.net/fig/1102005">4 D, E</figureCitation>
, 
<figureCitation id="DB3C393EC939EE49338B674D008580E5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Appendages of Nipponogarypus seosanensis sp. nov., paratype female. A. Galea; B. Left chela, lateral view; C. Left chelal fingers, lateral view, sensory setae in the white box; D. Right leg I; E. Claws and arolium of right leg I; F. Pleural membrane. Scale bars: 0.02 mm (A); 0.3 mm (B); 0.1 mm (C, D, F); 0.05 mm (E)." figureDoi="10.3897/zse.100.123213.figure5" httpUri="https://binary.pensoft.net/fig/1102006">5 D</figureCitation>
). Leg I: trochanter 1.56, femur 2.51, patella 1.80, tibia 3.06, metatarsus 2.08, tarsus 2.33 times longer than the broad; leg IV: trochanter 1.40, femur + patella 3.26, tibia 3.93, metatarsus 2.69, tarsus 2.98 times longer than the broad. Typical for the genus: femur and patella of leg I freely mobile; metatarsus of leg IV shorter than the tarsus; metatarsus of leg I longer than the tarsus; one each pseudotactile seta in the distal position of the tarsus; basal position of the metatarsus; arolium undivided, longer than claws, and simple.
</paragraph>
<paragraph id="0A7D4AA267B4EC65576E7B6742E98906">
Abdomen (Fig. 
<figureCitation id="14D0DEB5F1B327858C1F77821D96AC67" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Appendages of Nipponogarypus seosanensis sp. nov., paratype female. A. Galea; B. Left chela, lateral view; C. Left chelal fingers, lateral view, sensory setae in the white box; D. Right leg I; E. Claws and arolium of right leg I; F. Pleural membrane. Scale bars: 0.02 mm (A); 0.3 mm (B); 0.1 mm (C, D, F); 0.05 mm (E)." figureDoi="10.3897/zse.100.123213.figure5" httpUri="https://binary.pensoft.net/fig/1102006">5 F</figureCitation>
). Pleural membrane striate; all tergites and sternites both undivided; all setae short and acuminate; tergal chaetotaxy 2: 4: 4: 4: 4: 4: 6: 5: 6: 4: 4: 2. Sternal chaetotaxy 10: 4: 6: 8: 6: 4: 6: 6: 4: 4: 2.
</paragraph>
<caption id="5D10A8099FEEB46FE317881149B5D033" ID-DOI="10.3897/zse.100.123213.figure5" ID-arpha="A9410697-9942-5137-9A7A-9124E8006484" httpUri="https://binary.pensoft.net/fig/1102006" startId="F5">
<paragraph id="E5D232AF0283AA6279206DAB78AFD5CE">
<label id="D3CEE2690916B38542F5DA9A71E03577">Figure 5.</label>
</paragraph>
<paragraph id="7D526553DACDFF1F108496009CF9B54A">
Appendages of 
<taxonomicName id="F77E11DE1BCB609446948814C513589F" authorityName="Jeong &amp; Harms &amp; Yoo" authorityYear="2024" class="Arachnida" family="Olpiidae" genus="Nipponogarypus" kingdom="Animalia" order="Pseudoscorpiones" phylum="Arthropoda" rank="species" species="seosanensis" status="sp. nov.">
<emphasis id="AD39AAC2F55D9758BC400DCD11EA0046" italics="true">Nipponogarypus seosanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DC290BCFB817DC81358AF7A61AFC839" rank="species">sp. nov.</taxonomicNameLabel>
, paratype female. 
<emphasis id="3B0C7E73811A360268B0D467AB6709EB" bold="true">A.</emphasis>
Galea; 
<emphasis id="4EA8CE312718583DA0FDE8EB1D063939" bold="true">B.</emphasis>
Left chela, lateral view; 
<emphasis id="5F928083DFF6018B1CF5D2CF969AD126" bold="true">C.</emphasis>
Left chelal fingers, lateral view, sensory setae in the white box; 
<emphasis id="B19D3F57F9B926B0A7C1A9DC856178A2" bold="true">D.</emphasis>
Right leg I; 
<emphasis id="21B4F76D587925B7BA3C7BEFF876623D" bold="true">E.</emphasis>
Claws and arolium of right leg I; 
<emphasis id="D2E892FF73A413DA248C38DF6FA98730" bold="true">F.</emphasis>
Pleural membrane. Scale bars: 0.02 mm (
<emphasis id="558E0B048838968C93D4AAAF8D0433EB" bold="true">A</emphasis>
); 0.3 mm (
<emphasis id="A556586A5CA878E018316BB3296B74FA" bold="true">B</emphasis>
); 0.1 mm (
<emphasis id="A22F56AEBABD694DDB2B6DAECD3AE7F3" bold="true">C</emphasis>
, 
<emphasis id="0F0495BE0E87E0C7FF0A3F3827784C6B" bold="true">D</emphasis>
, 
<emphasis id="68130D8B32458B196E1055A505822E98" bold="true">F</emphasis>
); 0.05 mm (
<emphasis id="63BF6BE0383E2D45822FDCBFE6F6D811" bold="true">E</emphasis>
).
</paragraph>
</caption>
<paragraph id="5C3C2C9B80EF538315DB9A3135D27565">Dimensions (in mm). Body length: 1.93 cephalothorax: carapace 0.63 / 0.54, anterior eye 0.07, posterior eye 0.05; chelicera: total 0.22 / 0.12, movable finger 0.16; pedipalp: trochanter 0.31 / 0.19, femur 0.59 / 0.20, patella 0.58 / 0.23, chela 1.08 / 0.33, movable finger 0.53, hand 0.54; leg I: trochanter 0.15 / 0.10, femur 0.23 / 0.09, patella 0.18 / 0.10, tibia 0.21 / 0.07, metatarsus 0.10 / 0.05, tarsus 0.11 / 0.05; leg IV: trochanter 0.16 / 0.11, femur + patella 0.55 / 0.17, tibia 0.38 / 0.10, metatarsus 0.17 / 0.06, tarsus 0.16 / 0.05.</paragraph>
<paragraph id="D9F981FD6CAED16B89BDEF8EFD1F5CCF">
Variation (
<specimenCount id="A13C34AC7D7CA4F3397E7514BCC1DD3F" count="1" type="female">one female</specimenCount>
, 
<typeStatus id="13516EA48DCD42FA8839EF159090BDEF" type="paratype">paratype</typeStatus>
)
</paragraph>
<paragraph id="3F076AD9B41B78329E044DFA1A01B9E7">Cephalothorax. Carapace 1.16 times longer than broad.</paragraph>
<paragraph id="75183B593C42AF0C4D87A03C90C46F3D">Pedipalp. Trochanter 1.65, femur 3.13, patella 2.64, chela 3.29 times longer than broad, movable finger 0.88 times longer than hand. Fixed finger with 42, movable finger with 43 marginal teeth.</paragraph>
<paragraph id="560B9F28FC37B1CD9A6497A33E59770C">Legs. Leg I: trochanter 1.28, femur 3.29, tibia 3.34, metatarsus 2.38, tarsus 2.89 times longer than broad; leg IV: trochanter 1.50, femur + patella 3.06, tibia 3.08, metatarsus 2.25, tarsus 2.82 times longer than broad.</paragraph>
<paragraph id="F8D6F1A82174392A53A71C13EEC90081">Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 4: 2: 2. Sternal chaetotaxy 8: 4: 6: 6: 6: 6: 6: 4: 4: 4: 2.</paragraph>
<paragraph id="67192D4A2D666021120C0EC0E1F5FCAC">Dimensions (in mm). Body length 1.90; cephalothorax: carapace 0.58 / 0.50, anterior eye 0.06, posterior eye 0.05; chelicera: total 0.20 / 0.12, movable finger 0.16; pedipalp: trochanter 0.33 / 0.20, femur 0.61 / 0.20, patella 0.59 / 0.22, chela 1.04 / 0.32, movable finger 0.49, hand 0.56; leg I: trochanter 0.14 / 0.11, femur 0.24 / 0.10, patella 0.19 / 0.10, tibia 0.25 / 0.08, metatarsus 0.12 / 0.05, tarsus 0.13 / 0.05; leg IV: trochanter 0.19 / 0.13, femur + patella 0.60 / 0.20, tibia 0.34 / 0.11, metatarsus 0.15 / 0.07, tarsus 0.18 / 0.06.</paragraph>
<paragraph id="0183D216A1321C6035C1AB37A99A409F">
Variation (
<specimenCount id="332D0211CB4394B354176EF73D9C3310" count="2" type="male">two males</specimenCount>
, 
<typeStatus id="95073AD967EFA7DE8DF39B630B3C4DB2" type="paratype">paratype</typeStatus>
)
</paragraph>
<paragraph id="919ED6880FAD4D1BD422BF11BDD22067">Cephalothorax. Carapace 1.20–1.23 times longer than broad.</paragraph>
<paragraph id="5A3361A795BD3FC12C42F45DDDCC1391">Pedipalp. Trochanter 1.34–1.59, femur 3.07–3.13, patella 2.6, chela 3.05–3.21 times longer than the broad, movable finger 0.81–0.89 times longer than the hand. Fixed finger with 40–41, movable finger with 41–43 marginal teeth.</paragraph>
<paragraph id="4717E32AD5ABB8FD87C759CF866C2C1E">Legs. Leg I: trochanter 1.20–1.67, femur 2.08–2.45, patella 1.66–1.70, tibia 2.67–3.22, metatarsus 2.62–2.67, tarsus 2.86–3 times longer than broad; leg IV: trochanter 1.25–1.39, femur + patella 2.67–2.74, tibia 3.54–3.63, metatarsus 2.77–3.16, tarsus 2.5–2.7 times longer than broad.</paragraph>
<paragraph id="EAC7CC9AA41B8E9FB5AD5CD6AF688DD9">Abdomen. Tergal chaetotaxy 2: 4: 4: 4: 4: 6: 6: 6: 6: 4: 4: 2. Sternal chaetotaxy 11: 6: 4: 6: 6: 6: 6: 5: 4: 4: 2.</paragraph>
<paragraph id="A07761161E22E778A987F67253C30C4E">
Male genital sternites (Fig. 
<figureCitation id="DA35D22023ACE10DD5ABC2A56149B2DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Appendages of Nipponogarypus seosanensis sp. nov., holotype female. A. Carapace; B. Chelicera in dorsal view; C. Left pedipalp in dorsal view; D. Right chela from lateral view; E. Coxa; F. Right leg I; G. Right leg II; H. Female genital sternites; I. Right leg IV; J. Male genital sternites. Scale bars: 0.5 mm (A, C – G, I); 0.2 mm (H, J); 0.1 mm (B)." figureDoi="10.3897/zse.100.123213.figure4" httpUri="https://binary.pensoft.net/fig/1102005">4 G</figureCitation>
). Nine lyrifissures on sternites II and III; 11 setae on the genital opening area; six setae on the sternite III.
</paragraph>
<paragraph id="16831186F9A46073678B7688287B677F">Dimensions (in mm). Body length: 1.83–1.84; cephalothorax: carapace 0.61 / 0.50–0.51, anterior eye 0.06, posterior eye: 0.04–0.05; chelicera: total 0.20 / 0.10–0.11 movable finger 0.14; pedipalp: trochanter 0.25–0.32 / 0.19 – 0.20, femur 0.56–0.60 / 0.18 – 0.19, patella 0.53–0.57 / 0.21 – 0.22, chela 0.91–0.92 / 0.21 – 0.29, movable finger 0.41–0.43, z 0.49–0.50; leg I: trochanter 0.12 / 0.07–0.10, femur 0.18–0.24 / 0.09 – 0.10, patella 0.15–0.16 / 0.09 – 0.10, tibia 0.17–0.23 / 0.06 – 0.07, metatarsus 0.12–0.13 / 0.05, tarsus 0.13 / 0.04; leg IV: trochanter 0.17–0.18 / 0.13, femur + patella 0.51 / 0.19, tibia 0.37 / 0.10, metatarsus 0.15–0.16 / 0.05, tarsus 0.12–0.16 / 0.05 – 0.06.</paragraph>
</subSubSection>
</treatment>
</document>