390 lines
48 KiB
XML
390 lines
48 KiB
XML
<document id="9D185E58EF696CC83C362AF01D4270C1" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE53B63FDD9E43E9FEC2039F" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius" docType="key" docVersion="12" lastPageNumber="116" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="115" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="02CE109585D99C8E4E831A6F610EB5BB">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<mods:namePart id="07A15E3F61D5A4040FCBBE398420992E">Tarasov, Sergei</mods:namePart>
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<mods:title id="B65CBED2C452F48B1A706D350E9AA702">Zootaxa</mods:title>
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<mods:part id="4D67E6403423E64B79A0560257ECE0BB">
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<mods:date id="A0443CC85F3814ECA0984EC8C078362C">2017</mods:date>
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<mods:number id="049A93BCF2FB214A1FF7070F2148B3D3">2017-10-04</mods:number>
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<mods:number id="3779830605D7833F403EDF9C2E1EFCC1">4329</mods:number>
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<mods:identifier id="47C5331706542F8ACD1D834B273C2011" type="DOI">10.11646/zootaxa.4329.2.1</mods:identifier>
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<treatment id="03C187E1AE53B63FDD9E43E9FEC2039F" ID-DOI="http://doi.org/10.5281/zenodo.6010806" ID-GBIF-Taxon="135106116" ID-Zenodo-Dep="6010806" LSID="urn:lsid:plazi:treatment:03C187E1AE53B63FDD9E43E9FEC2039F" httpUri="http://treatment.plazi.org/id/03C187E1AE53B63FDD9E43E9FEC2039F" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
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<subSubSection id="C372657CAE53B63EDD9E43E9FEAF0700" pageId="14" pageNumber="115" type="nomenclature">
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<paragraph id="8BD736F7AE53B63EDD9E43E9FDD706CF" blockId="14.[151,524,1189,1216]" box="[151,524,1189,1216]" pageId="14" pageNumber="115">
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<heading id="D09F819BAE53B63EDD9E43E9FDD706CF" bold="true" box="[151,524,1189,1216]" fontSize="11" level="1" pageId="14" pageNumber="115" reason="1">
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<emphasis id="B91CEAE5AE53B63EDD9E43E9FDD706CF" bold="true" box="[151,524,1189,1216]" pageId="14" pageNumber="115">
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Key to females of
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<taxonomicName id="4C684D74AE53B63EDC7F43EAFDD706CF" ID-CoL="63J2L" box="[374,524,1189,1215]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE53B63EDC7F43EAFDD706CF" bold="true" box="[374,524,1189,1215]" italics="true" pageId="14" pageNumber="115">Parachorius</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD736F7AE53B63EDD9E43A3FEAF0700" blockId="14.[151,1437,1260,1393]" pageId="14" pageNumber="115">
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Twelve out of 19
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<taxonomicName id="4C684D74AE53B63EDC7B43A3FE250775" box="[370,510,1260,1285]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEAE5AE53B63EDC7B43A3FE250775" box="[370,510,1260,1285]" italics="true" pageId="14" pageNumber="115">Parachorius</emphasis>
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</taxonomicName>
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species cannot be reliably identified using females. Therefore, the key below contains some couplets that refer to several species at the same time. In such ambiguous cases the distribution criterion should be used to separate species. This criterion is not always reliable as many species have overlapping distributional areas.
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</paragraph>
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</subSubSection>
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<subSubSection id="C372657CAE53B63FDD9E42EFFEC2039F" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115" type="key">
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<keyStep id="309C2E52AE53B63EDD9E42EFFA4707A2" pageId="14" pageNumber="115">
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<paragraph id="8BD736F7AE53B63EDD9E42EFFA4707C5" blockId="14.[151,1437,1440,2018]" box="[151,1436,1440,1461]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E42EFFA4707C5" box="[151,1436,1440,1461]" pageId="14" pageNumber="115">
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1 Clypeus with two teeth, outer margin of each tooth not notched basally (
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<figureCitation id="13532A72AE53B63EDE9F42EFFC2007C5" box="[918,1019,1440,1461]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4F–G</figureCitation>
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)................................2
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</keyLead>
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</paragraph>
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<paragraph id="8BD736F7AE53B63EDD9E42F2FA4707A2" blockId="14.[151,1437,1440,2018]" box="[151,1436,1469,1490]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E42F2FA4707A2" box="[151,1436,1469,1490]" pageId="14" pageNumber="115">
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- Clypeus with two teeth, outer margin of each tooth distinctly or slightly notched basally (
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<figureCitation id="13532A72AE53B63ED93142F2FB7A07A2" box="[1080,1185,1469,1490]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4D–E</figureCitation>
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)...................6
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="309C2E52AE53B63EDD9E4294FA40047D" pageId="14" pageNumber="115">
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<paragraph id="8BD736F7AE53B63EDD9E4294FA400780" blockId="14.[151,1437,1440,2018]" box="[151,1435,1499,1520]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E4294FA400780" box="[151,1435,1499,1520]" pageId="14" pageNumber="115">
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2 Clypeus with two distinctly produced teeth (
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<figureCitation id="13532A72AE53B63EDF834294FD0E0780" box="[650,725,1499,1520]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4G</figureCitation>
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)......................................................... 3
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</keyLead>
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</paragraph>
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<paragraph id="8BD736F7AE53B63EDD9E42B7FA40047D" blockId="14.[151,1437,1440,2018]" box="[151,1435,1528,1549]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E42B7FA40047D" box="[151,1435,1528,1549]" pageId="14" pageNumber="115">
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- Clypeus with two less produced teeth (
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<figureCitation id="13532A72AE53B63EDF5D42B7FD46047D" box="[596,669,1528,1549]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4F</figureCitation>
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).............................................................. 4
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="309C2E52AE53B63EDD9E415AFA400416" pageId="14" pageNumber="115">
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<paragraph id="8BD736F7AE53B63EDD9E415AFA40045B" blockId="14.[151,1437,1440,2018]" box="[151,1435,1557,1579]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E415AFA40045B" box="[151,1435,1557,1579]" pageId="14" pageNumber="115">
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3 Body black; head triangularly shaped (
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<figureCitation id="13532A72AE53B63EDF5D415AFD40045A" box="[596,667,1557,1578]" captionStart="FIGURE 21" captionStartId="37.[151,250,1909,1931]" captionTargetBox="[151,1436,973,1878]" captionTargetId="figure@37.[151,1436,973,1888]" captionTargetPageId="37" captionText="FIGURE 21. Morphology of Parachorius schuelkei. (A) aedeagus, lateral view (paratype). (B) aedeagus, lateral view (holotype). (C) aedeagus lateral view. (D) habitus (male, holotype). (E) habitus (female, paratype). (F) protibia (female, paratype). (G) left metafemur (male, holotype)." httpUri="https://zenodo.org/record/1001959/files/figure.png" pageId="14" pageNumber="115">Fig. 21</figureCitation>
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); length
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<quantity id="4C909B12AE53B63EDFE6415AFCBE045A" box="[751,869,1557,1578]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="14" pageNumber="115" unit="mm" value="6.65" valueMax="6.8" valueMin="6.5">6.5–6.8 mm</quantity>
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; southern China......
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<taxonomicName id="4C684D74AE53B63ED954415AFAC4045A" ID-CoL="6TNLX" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1117,1311,1557,1578]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
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<emphasis id="B91CEAE5AE53B63ED954415AFB14045A" box="[1117,1231,1557,1578]" italics="true" pageId="14" pageNumber="115">P. schuelkei</emphasis>
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Tarasov
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</taxonomicName>
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<taxonomicNameLabel id="A22F579EAE53B63ED82C415AFA40045B" box="[1317,1435,1557,1579]" pageId="14" pageNumber="115" rank="species">
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<emphasis id="B91CEAE5AE53B63ED82C415AFA40045B" bold="true" box="[1317,1435,1557,1579]" pageId="14" pageNumber="115">new species</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="8BD736F7AE53B63EDD9E417CFA400416" blockId="14.[151,1437,1440,2018]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E417CFA400416" pageId="14" pageNumber="115">
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- Body reddish-brown, rarely dark brown; head more rounded not as distinctly triangular as in
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<taxonomicName id="4C684D74AE53B63ED954417CFB150438" ID-CoL="6TNLX" authorityName="S. Tarasov" authorityYear="2017" box="[1117,1230,1587,1608]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="schuelkei">
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<emphasis id="B91CEAE5AE53B63ED954417CFB150438" box="[1117,1230,1587,1608]" italics="true" pageId="14" pageNumber="115">P. schuelkei</emphasis>
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</taxonomicName>
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; length
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<quantity id="4C909B12AE53B63ED814417CFAB70438" box="[1309,1388,1587,1608]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="14" pageNumber="115" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
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||
..............................................
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<taxonomicName id="4C684D74AE53B63EDFD8411FFC210415" ID-CoL="6TNM6" authority="(Boucomont, 1914)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Boucomont" baseAuthorityYear="1914" box="[721,1018,1616,1637]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="javanus">
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<emphasis id="B91CEAE5AE53B63EDFD8411FFCE30415" box="[721,824,1616,1637]" italics="true" pageId="14" pageNumber="115">P. javanus</emphasis>
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(
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<bibRefCitation id="EFF94B06AE53B63EDE4C411FFC280415" author="Boucomont" box="[837,1011,1616,1637]" pageId="14" pageNumber="115" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel Malais. Annales de la Societe Entomologique de France, 83, 238 - 350." type="journal article" year="1914">Boucomont, 1914</bibRefCitation>
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)
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||
</taxonomicName>
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and
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<taxonomicName id="4C684D74AE53B63ED921411FFAC40415" ID-CoL="75NM8" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1064,1311,1616,1637]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="pseudojavanus" status="sp. nov.">
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<emphasis id="B91CEAE5AE53B63ED921411FFB150415" box="[1064,1230,1616,1637]" italics="true" pageId="14" pageNumber="115">P. pseudojavanus</emphasis>
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Tarasov
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</taxonomicName>
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<taxonomicNameLabel id="A22F579EAE53B63ED82C411FFA400416" box="[1317,1435,1616,1638]" pageId="14" pageNumber="115" rank="species">
|
||
<emphasis id="B91CEAE5AE53B63ED82C411FFA400416" bold="true" box="[1317,1435,1616,1638]" pageId="14" pageNumber="115">new species</emphasis>
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</taxonomicNameLabel>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="309C2E52AE53B63EDD9E4122FA4004AA" pageId="14" pageNumber="115">
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<paragraph id="8BD736F7AE53B63EDD9E4122FA4004D0" blockId="14.[151,1437,1440,2018]" pageId="14" pageNumber="115">
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<keyLead id="309995C2AE53B63EDD9E4122FA4004D0" pageId="14" pageNumber="115">
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4 Elytra covered with dense contiguous and rugose punctures (
|
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<figureCitation id="13532A72AE53B63EDE2E4122FCA604F2" box="[807,893,1645,1666]" captionStart="FIGURE 10" captionStartId="22.[151,250,1898,1920]" captionTargetBox="[151,1428,799,1834]" captionTargetId="figure@22.[151,1436,799,1851]" captionTargetPageId="22" captionText="FIGURE 10. Morphology of Parachorius globosus. (A – C) aedeagus. (D) habitus (male). (E) habitus (female, holotype). (F) disc of left elytron. (G) protibia (female)" httpUri="https://zenodo.org/record/1001937/files/figure.png" pageId="14" pageNumber="115">Fig. 10F</figureCitation>
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); pronotum almost as long as elytra; wings reduced (
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<figureCitation id="13532A72AE53B63ED87C4122FF2704D0" captionStart="FIGURE 10" captionStartId="22.[151,250,1898,1920]" captionTargetBox="[151,1428,799,1834]" captionTargetId="figure@22.[151,1436,799,1851]" captionTargetPageId="22" captionText="FIGURE 10. Morphology of Parachorius globosus. (A – C) aedeagus. (D) habitus (male). (E) habitus (female, holotype). (F) disc of left elytron. (G) protibia (female)" httpUri="https://zenodo.org/record/1001937/files/figure.png" pageId="14" pageNumber="115">Fig. 10</figureCitation>
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);
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||
<quantity id="4C909B12AE53B63EDC0441C4FE8204D0" box="[269,345,1675,1696]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="14" pageNumber="115" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7mm</quantity>
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; northeastern
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<collectingCountry id="F37F7667AE53B63EDCE941C4FDCE04D0" box="[480,533,1675,1696]" name="India" pageId="14" pageNumber="115">India</collectingCountry>
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.......................................................
|
||
<taxonomicName id="4C684D74AE53B63ED9B941C4FA4004D0" ID-CoL="6TNLT" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1200,1435,1675,1696]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="globosus">
|
||
<emphasis id="B91CEAE5AE53B63ED9B941C4FAC604D0" box="[1200,1309,1675,1696]" italics="true" pageId="14" pageNumber="115">P. globosus</emphasis>
|
||
<bibRefCitation id="EFF94B06AE53B63ED82A41C4FA4004D0" author="Arrow" box="[1315,1435,1675,1696]" pageId="14" pageNumber="115" refString="Arrow, G. J. (1931) Coleoptera Lamellicornia part. III. (Coprinae). Taylor and Francis, London, 428 pp., 13 pls." type="book" year="1931">Arrow, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE53B63EDD9E41E7FA4004AA" blockId="14.[151,1437,1440,2018]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E41E7FA4004AA" pageId="14" pageNumber="115">
|
||
- Elytra covered with sparse (not contiguous) punctures (
|
||
<figureCitation id="13532A72AE53B63EDFFB41E7FC9304CD" box="[754,840,1704,1725]" captionStart="FIGURE 24" captionStartId="41.[151,250,1951,1973]" captionTargetBox="[168,1425,1036,1927]" captionTargetId="figure@41.[168,1432,1036,1939]" captionTargetPageId="41" captionText="FIGURE 24. Morphology of Parachorius thomsoni. (A) aedeagus, lateral view (P. thomsoni holotype, male) (B) aedeagus, lateral view (P. lannathai, holotype, male). (C – D) aedeagus, lateral view. (E) left metafemur (P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (P. lannathai, holotype, male). (H) habitus (P. thomsoni, holotype, male)." httpUri="https://zenodo.org/record/1001965/files/figure.png" pageId="14" pageNumber="115">Fig. 24F</figureCitation>
|
||
); pronotum notably shorter than elytra; wings normally devel- oped............................................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE53B63EDD9E41ACFD55051D" pageId="14" pageNumber="115">
|
||
<paragraph id="8BD736F7AE53B63EDD9E41ACFA400488" blockId="14.[151,1437,1440,2018]" box="[151,1435,1763,1784]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E41ACFA400488" box="[151,1435,1763,1784]" pageId="14" pageNumber="115">
|
||
5 Length
|
||
<quantity id="4C909B12AE53B63EDC2241ACFEB90488" box="[299,354,1763,1784]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="115" unit="mm" value="6.0">6 mm</quantity>
|
||
(
|
||
<figureCitation id="13532A72AE53B63EDC6641ACFE6C0488" box="[367,439,1763,1784]" captionStart="FIGURE 12" captionStartId="25.[151,250,1061,1083]" captionTargetBox="[151,1426,303,1030]" captionTargetId="figure@25.[151,1436,303,1040]" captionTargetPageId="25" captionText="FIGURE 12. Morphology of Parachorius hookeri. (A) aedeagus, lateral view (holotype). (B) aedeagus lateral view. (C) left metafemur (male, holotype). (D) habitus (male, holotype)" httpUri="https://zenodo.org/record/1001941/files/figure.png" pageId="14" pageNumber="115">Fig. 12</figureCitation>
|
||
); northern
|
||
<collectingCountry id="F37F7667AE53B63EDF2941ACFD8A0488" box="[544,593,1763,1784]" name="India" pageId="14" pageNumber="115">India</collectingCountry>
|
||
..................................................
|
||
<taxonomicName id="4C684D74AE53B63ED9B541ACFA400488" ID-CoL="75NLT" authority="Arrow, 1931" authorityName="Arrow M. E. Bacchus" authorityYear="1931" box="[1212,1435,1763,1784]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="hookeri">
|
||
<emphasis id="B91CEAE5AE53B63ED9B541ACFAC60488" box="[1212,1309,1763,1784]" italics="true" pageId="14" pageNumber="115">P. hookeri</emphasis>
|
||
<bibRefCitation id="EFF94B06AE53B63ED82A41ACFA400488" author="Arrow" box="[1315,1435,1763,1784]" pageId="14" pageNumber="115" refString="Arrow, G. J. (1931) Coleoptera Lamellicornia part. III. (Coprinae). Taylor and Francis, London, 428 pp., 13 pls." type="book" year="1931">Arrow, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE53B63EDD9E404FFD55051D" blockId="14.[151,1437,1440,2018]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E404FFD55051D" pageId="14" pageNumber="115">
|
||
- Length 6.5–9.0 mm. This couplet contains four species with morphologically undistinguishable females. Their identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details).....................
|
||
<taxonomicName id="4C684D74AE53B63EDDE14074FE150520" ID-CoL="6TNLW" authority="Tarasov" authorityName="Tarasov" authorityYear="2017" box="[232,462,1851,1872]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="newthayerae" status="sp. nov.">
|
||
<emphasis id="B91CEAE5AE53B63EDDE14074FEA00520" box="[232,379,1851,1872]" italics="true" pageId="14" pageNumber="115">P. newthayerae</emphasis>
|
||
Tarasov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F579EAE53B63EDCDF4075FD970520" box="[470,588,1850,1872]" pageId="14" pageNumber="115" rank="species">
|
||
<emphasis id="B91CEAE5AE53B63EDCDF4075FD970520" bold="true" box="[470,588,1850,1872]" pageId="14" pageNumber="115">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C684D74AE53B63EDF524074FC560520" ID-CoL="75NLV" authority="Masumoto, Ochi" authorityName="Masumoto, Ochi" box="[603,909,1851,1872]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="maruyamai">
|
||
<emphasis id="B91CEAE5AE53B63EDF524074FD3A0520" box="[603,737,1851,1872]" italics="true" pageId="14" pageNumber="115">P. maruyamai</emphasis>
|
||
Masumoto, Ochi
|
||
</taxonomicName>
|
||
, & Sakchoowong, 2012,
|
||
<taxonomicName id="4C684D74AE53B63ED98C4074FAA40520" ID-CoL="4CPV4" authority="Harold, 1873" authorityName="Harold" authorityYear="1873" box="[1157,1407,1851,1872]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="thomsoni">
|
||
<emphasis id="B91CEAE5AE53B63ED98C4074FB2D0520" box="[1157,1270,1851,1872]" italics="true" pageId="14" pageNumber="115">P. thomsoni</emphasis>
|
||
<bibRefCitation id="EFF94B06AE53B63ED9F44074FAA40520" author="Harold" box="[1277,1407,1851,1872]" pageId="14" pageNumber="115" refString="Harold, E. (1873) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 11, 102 - 105." type="journal article" year="1873">Harold, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C684D74AE53B63ED8834074FD55051D" ID-CoL="75NM4" authority="Kryzhanovsky & Medvedev, 1966" authorityName="Kryzhanovsky & Medvedev" authorityYear="1966" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="fungorum">
|
||
<emphasis id="B91CEAE5AE53B63ED8834074FEE5051D" italics="true" pageId="14" pageNumber="115">P. fungorum</emphasis>
|
||
<bibRefCitation id="EFF94B06AE53B63EDC4D4017FD55051D" author="Kryzhanovsky" box="[324,654,1880,1901]" pageId="14" pageNumber="115" refString="Kryzhanovsky, O. L. & Medvedev, S. I. (1966) Dung beetles of the subfamily Coprinae in Yunnan province of South-Western China. Part 1. Entomologicheskoe Obozrenie, Revue d'entomologie de l'URSS, 45, 387 - 400." type="journal article" year="1966">Kryzhanovsky & Medvedev, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE53B63EDD9E403AFA4005D8" pageId="14" pageNumber="115">
|
||
<paragraph id="8BD736F7AE53B63EDD9E403AFA4705FA" blockId="14.[151,1437,1440,2018]" box="[151,1436,1909,1930]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E403AFA4705FA" box="[151,1436,1909,1930]" pageId="14" pageNumber="115">
|
||
6 Outer margin of each clypeal tooth slightly notched basally (
|
||
<figureCitation id="13532A72AE53B63EDE2F403AFCAA05FA" box="[806,881,1909,1930]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4E</figureCitation>
|
||
).............................................7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE53B63EDD9E40DCFA4005D8" blockId="14.[151,1437,1440,2018]" box="[151,1435,1939,1960]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E40DCFA4005D8" box="[151,1435,1939,1960]" pageId="14" pageNumber="115">
|
||
- Outer margin of each clypeal tooth distinctly notched basally (
|
||
<figureCitation id="13532A72AE53B63EDE3E40DCFC5905D8" box="[823,898,1939,1960]" captionStart="FIGURE 4" captionStartId="11.[151,250,1607,1629]" captionTargetBox="[170,1436,833,1574]" captionTargetId="figure@11.[151,1436,833,1585]" captionTargetPageId="11" captionText="FIGURE 4. Morphological elements of Parachorius. (A – C) Endophallic sclerites; aedeagus icon in blue circle indicates position of aedeagus in relation to endophallic sclerite. (D – G) Head in dorsal view. (A, D) Parachorius longipenis. (B) Parachorius fungorum. (C, G) Parachorius pseudojavanus. (E) Parachorius gotoi. (F) Parachorius globosus." httpUri="https://zenodo.org/record/1001923/files/figure.png" pageId="14" pageNumber="115">Fig. 4D</figureCitation>
|
||
).......................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE53B63EDD9E40FFFA400592" pageId="14" pageNumber="115">
|
||
<paragraph id="8BD736F7AE53B63EDD9E40FFFA4705B5" blockId="14.[151,1437,1440,2018]" box="[151,1436,1968,1989]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E40FFFA4705B5" box="[151,1436,1968,1989]" pageId="14" pageNumber="115">
|
||
7 Clypeal teeth acute; length
|
||
<quantity id="4C909B12AE53B63EDCEE40FFFDED05B5" box="[487,566,1968,1989]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="115" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
(
|
||
<figureCitation id="13532A72AE53B63EDF4A40FFFD5205B5" box="[579,649,1968,1989]" captionStart="FIGURE 11" captionStartId="23.[151,250,1984,2006]" captionTargetBox="[151,1410,826,1961]" captionTargetId="figure@23.[151,1436,826,1969]" captionTargetPageId="23" captionText="FIGURE 11. Morphology of Parachorius gotoi. (A, C) aedeagus. (B) left metafemur (male). (D) protibia (male). (E) habitus (male)." httpUri="https://zenodo.org/record/1001939/files/figure.png" pageId="14" pageNumber="115">Fig. 11</figureCitation>
|
||
);
|
||
<collectingCountry id="F37F7667AE53B63EDF9540FFFD3905B5" box="[668,738,1968,1989]" name="Taiwan" pageId="14" pageNumber="115">Taiwan</collectingCountry>
|
||
...................................
|
||
<taxonomicName id="4C684D74AE53B63ED99340FFFA4705B5" ID-CoL="6TNM5" authority="(Masumoto, 1986)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Masumoto" baseAuthorityYear="1986" box="[1178,1436,1968,1989]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="B91CEAE5AE53B63ED99340FFFB3805B5" box="[1178,1251,1968,1989]" italics="true" pageId="14" pageNumber="115">P. gotoi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B06AE53B63ED9F940FFFA4E05B5" author="Masumoto" box="[1264,1429,1968,1989]" pageId="14" pageNumber="115" refString="Masumoto, K. (1986) New coprophagous Lamellicornia from Japan and Formosa, 3. (Col., Scarabaeidae). Entomological Review of Japan, 41, 85 - 87." type="journal article" year="1986">Masumoto, 1986</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE53B63EDD9E4082FA400592" blockId="14.[151,1437,1440,2018]" box="[151,1435,1997,2018]" pageId="14" pageNumber="115">
|
||
<keyLead id="309995C2AE53B63EDD9E4082FA400592" box="[151,1435,1997,2018]" pageId="14" pageNumber="115">
|
||
- Clypeal teeth rounded; length
|
||
<quantity id="4C909B12AE53B63EDF0A4082FD590592" box="[515,642,1997,2018]" metricMagnitude="-3" metricUnit="m" metricValue="9.399999999999999" metricValueMax="10.6" metricValueMin="8.2" pageId="14" pageNumber="115" unit="mm" value="9.399999999999999" valueMax="10.6" valueMin="8.2">8.2–10.6 mm</quantity>
|
||
(
|
||
<figureCitation id="13532A72AE53B63EDF864082FD0C0592" box="[655,727,1997,2018]" captionStart="FIGURE 22" captionStartId="39.[151,250,1152,1174]" captionTargetBox="[151,1430,195,1127]" captionTargetId="figure@39.[151,1436,195,1131]" captionTargetPageId="39" captionText="FIGURE 22. Morphology of Parachorius semsanganus. (A) aedeagus, lateral view (paratype). (B) aedeagus, dorsal view (paratype). (C) aedeagus, apical view (paratype). (D) aedeagus lateral view. (E) right metafemur (male, paratype). (F) habitus (male, holotype)." httpUri="https://zenodo.org/record/1001961/files/figure.png" pageId="14" pageNumber="115">Fig. 22</figureCitation>
|
||
); northern Laos..............
|
||
<taxonomicName id="4C684D74AE53B63ED92D4082FA400592" ID-CoL="6TNM9" authority="Tarasov & Keith, 2011" authorityName="Tarasov & Keith" authorityYear="2011" box="[1060,1435,1997,2018]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="semsanganus">
|
||
<emphasis id="B91CEAE5AE53B63ED92D4082FB600592" box="[1060,1211,1997,2018]" italics="true" pageId="14" pageNumber="115">P. semsanganus</emphasis>
|
||
<bibRefCitation id="EFF94B06AE53B63ED9C84082FA400592" author="Tarasov" box="[1217,1435,1997,2018]" pageId="14" pageNumber="115" refString="Tarasov, S. & Keith, D. (2011) Parachorius semsanganus sp. n. (Coleoptera, Scarabaeidae, Scarabaeinae) from Laos and its significance in the phylogeny of Oriental Deltochilini. Zookeys, 111, 51 - 57. https: // doi. org / 10.3897 / zookeys. 111.1221" type="journal article" year="2011">Tarasov & Keith, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE52B63FDD9E47D8FA400297" pageId="15" pageNumber="116">
|
||
<paragraph id="8BD736F7AE52B63FDD9E47D8FA4002DC" blockId="15.[151,1436,151,495]" box="[151,1435,151,172]" pageId="15" pageNumber="116">
|
||
<materialsCitation id="3B003CAAAE52B63FDD9E47D8FA4002DC" ID-GBIF-Occurrence="1658411528" box="[151,1435,151,172]" country="India" location="Humeral area" pageId="15" pageNumber="116" specimenCount="1">
|
||
<keyLead id="309995C2AE52B63FDD9E47D8FA4002DC" box="[151,1435,151,172]" pageId="15" pageNumber="116">
|
||
8
|
||
<location id="8EB7602CAE52B63FDDE847D8FEB902DC" LSID="urn:lsid:plazi:treatment:03C187E1AE53B63FDD9E43E9FEC2039F:8EB7602CAE52B63FDDE847D8FEB902DC" box="[225,354,151,172]" country="India" name="Humeral area" pageId="15" pageNumber="116">Humeral area</location>
|
||
of elytron usually with a yellow spot, sometimes the yellow spot is absent and elytra uniformly colored;
|
||
<collectingCountry id="F37F7667AE52B63FD85847D8FA5002DC" box="[1361,1419,151,172]" name="India" pageId="15" pageNumber="116">India</collectingCountry>
|
||
.9
|
||
</keyLead>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE52B63FDD9E47FAFA400297" blockId="15.[151,1436,151,495]" pageId="15" pageNumber="116">
|
||
<keyLead id="309995C2AE52B63FDD9E47FAFA400297" pageId="15" pageNumber="116">
|
||
-
|
||
<materialsCitation id="3B003CAAAE52B63FDDE847FAFA400297" ID-GBIF-Occurrence="1658411533" country="China" location="Humeral area" pageId="15" pageNumber="116" specimenCount="1" stateProvince="Sumatra">
|
||
<location id="8EB7602CAE52B63FDDE847FAFEBF02BA" LSID="urn:lsid:plazi:treatment:03C187E1AE53B63FDD9E43E9FEC2039F:8EB7602CAE52B63FDDE847FAFEBF02BA" box="[225,356,181,202]" country="China" name="Humeral area" pageId="15" pageNumber="116" stateProvince="Sumatra">Humeral area</location>
|
||
of elytron without yellow spot, elytra uniformly colored; southeastern
|
||
<collectingCountry id="F37F7667AE52B63FD91E47FAFB8802BA" box="[1047,1107,181,202]" name="China" pageId="15" pageNumber="116">China</collectingCountry>
|
||
, Indochina, Malay Peninsula, and
|
||
<collectingRegion id="49ACF815AE52B63FDDE8479DFEEA0297" box="[225,305,210,231]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="116">Sumatra</collectingRegion>
|
||
............................................................................................10
|
||
</materialsCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE52B63FDD9E47A0FA470352" pageId="15" pageNumber="116">
|
||
<paragraph id="8BD736F7AE52B63FDD9E47A0FA400375" blockId="15.[151,1436,151,495]" box="[151,1435,239,261]" pageId="15" pageNumber="116">
|
||
<keyLead id="309995C2AE52B63FDD9E47A0FA400375" box="[151,1435,239,261]" pageId="15" pageNumber="116">
|
||
9 Species from southern and central India (
|
||
<figureCitation id="13532A72AE52B63FDF6247A0FD7D0374" box="[619,678,239,260]" captionStart="FIGURE 6" captionStartId="16.[151,250,1359,1381]" captionTargetBox="[151,1436,193,1325]" captionTargetId="figure@16.[151,1436,193,1338]" captionTargetPageId="16" captionText="FIGURE 6. Morphology of Parachorius asymmetricus. (A) aedeagus, left lateral view (holotype). (B) aedeagus, left lateral view (paratype). (C) aedeagus, right lateral view (holotype). (D) aedeagus, dorsal view (holotype). (E) habitus (male, holotype). (F) right metafemur (male, holotype). (G) protibia (putative female). (H) protibia (male, holotype)." httpUri="https://zenodo.org/record/1001927/files/figure.png" pageId="15" pageNumber="116">Fig. 6</figureCitation>
|
||
); length
|
||
<quantity id="4C909B12AE52B63FDFF347A0FCB60374" box="[762,877,239,260]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="15" pageNumber="116" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5–4.7 mm</quantity>
|
||
................
|
||
<taxonomicName id="4C684D74AE52B63FD92447A0FAC40374" ID-CoL="6TNLS" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[1069,1311,239,260]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
|
||
<emphasis id="B91CEAE5AE52B63FD92447A0FB150374" box="[1069,1230,239,260]" italics="true" pageId="15" pageNumber="116">P. asymmetricus</emphasis>
|
||
Tarasov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F579EAE52B63FD82C47A0FA400375" box="[1317,1435,239,261]" pageId="15" pageNumber="116" rank="species">
|
||
<emphasis id="B91CEAE5AE52B63FD82C47A0FA400375" bold="true" box="[1317,1435,239,261]" pageId="15" pageNumber="116">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE52B63FDD9E4642FA470352" blockId="15.[151,1436,151,495]" box="[151,1436,269,290]" pageId="15" pageNumber="116">
|
||
<materialsCitation id="3B003CAAAE52B63FDD9E4642FA470352" ID-GBIF-Occurrence="1658411529" box="[151,1436,269,290]" country="India" location="Sikkim" pageId="15" pageNumber="116" specimenCount="1" stateProvince="Sikkim">
|
||
<keyLead id="309995C2AE52B63FDD9E4642FA470352" box="[151,1436,269,290]" pageId="15" pageNumber="116">
|
||
- Species from northeastern
|
||
<collectingCountry id="F37F7667AE52B63FDCE84642FDC90352" box="[481,530,269,290]" name="India" pageId="15" pageNumber="116">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="49ACF815AE52B63FDF164642FDB30352" box="[543,616,269,290]" country="India" name="Sikkim" pageId="15" pageNumber="116">Sikkim</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ACF815AE52B63FDF784642FD6F0352" box="[625,692,269,290]" country="India" name="Assam" pageId="15" pageNumber="116">Assam</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ACF815AE52B63FDFB64642FCF50352" box="[703,814,269,290]" country="India" name="Meghalaya" pageId="15" pageNumber="116">Meghalaya</collectingRegion>
|
||
); length 3.8–5.0 mm (
|
||
<figureCitation id="13532A72AE52B63FD9094642FB9C0352" box="[1024,1095,269,290]" captionStart="FIGURE 13" captionStartId="26.[151,250,1259,1281]" captionTargetBox="[151,1416,193,1211]" captionTargetId="figure@26.[151,1436,193,1237]" captionTargetPageId="26" captionText="FIGURE 13. Morphology of Parachorius humeralis. (A – B) aedeagus (Darjeeling). (C) aedeagus. (D) habitus (male, holotype). (E) habitus (male). (F – H) right metafemur (male). (I) protibia (female). (J) protibia (male)." httpUri="https://zenodo.org/record/1001943/files/figure.png" pageId="15" pageNumber="116">Fig. 13</figureCitation>
|
||
)......
|
||
<taxonomicName id="4C684D74AE52B63FD99C4642FA470352" ID-CoL="75NM5" authority="(Arrow, 1907)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" box="[1173,1436,269,290]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="humeralis">
|
||
<emphasis id="B91CEAE5AE52B63FD99C4642FAD60352" box="[1173,1293,269,290]" italics="true" pageId="15" pageNumber="116">P. humeralis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B06AE52B63FD8104642FA4E0352" author="Arrow" box="[1305,1429,269,290]" pageId="15" pageNumber="116" refString="Arrow, G. J. (1907) Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 2). The Annals and Magazine of Natural History, including Zoology, Botany and Geology, 7 (19), 416 - 439." type="journal article" year="1907">Arrow, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="309C2E52AE52B63FDD9E4665FEC2039F" pageId="15" pageNumber="116">
|
||
<paragraph id="8BD736F7AE52B63FDD9E4665FA47032C" blockId="15.[151,1436,151,495]" pageId="15" pageNumber="116">
|
||
<keyLead id="309995C2AE52B63FDD9E4665FA47032C" pageId="15" pageNumber="116">
|
||
10 Body shape rectangular and less convex (
|
||
<figureCitation id="13532A72AE52B63FDF674665FD18034F" box="[622,707,298,319]" captionStart="FIGURE 7" captionStartId="17.[151,250,1493,1515]" captionTargetBox="[153,1407,193,1475]" captionTargetId="figure@17.[151,1436,193,1479]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of Parachorius bolavensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype). (C) aedeagus. (D) habitus (male, holotype). (E) protibia (male, paratype). (F) protibia (female, paratype). (G) left metafemur (male, paratype)." httpUri="https://zenodo.org/record/1001929/files/figure.png" pageId="15" pageNumber="116">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="13532A72AE52B63FDFC44665FD32034F" box="[717,745,298,319]" captionStart="FIGURE 8" captionStartId="19.[151,250,1935,1957]" captionTargetBox="[151,1419,233,1853]" captionTargetId="figure@19.[151,1436,233,1866]" captionTargetPageId="19" captionText="FIGURE 8. Morphology of Parachorius fukiensis. (A) aedeagus, lateral view (holotype). (B) aedeagus, lateral view (paratype, Laos, Phonsavan). (C) aedeagus, lateral view (paratype, Vietnam, Donghoi). (D) left metafemur (male, paratype, Laos, Phonsavan). (E) left metafemur (male, paratype, Vietnam, Donghoi). (F) habitus (male, holotype). (G) aedeagus." httpUri="https://zenodo.org/record/1001931/files/figure.png" pageId="15" pageNumber="116">8F</figureCitation>
|
||
); length
|
||
<quantity id="4C909B12AE52B63FDE344665FC6E034F" box="[829,949,298,319]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.6" metricValueMin="3.5" pageId="15" pageNumber="116" unit="mm" value="4.05" valueMax="4.6" valueMin="3.5">3.5–4.6 mm</quantity>
|
||
.......................................................................................
|
||
<taxonomicName id="4C684D74AE52B63FDE054608FC00032C" ID-CoL="6TNM4" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[780,987,327,348]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
|
||
<emphasis id="B91CEAE5AE52B63FDE054608FC52032C" box="[780,905,327,348]" italics="true" pageId="15" pageNumber="116">P. bolavensis</emphasis>
|
||
Tarasov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F579EAE52B63FDEE94608FB8D032D" box="[992,1110,327,349]" pageId="15" pageNumber="116" rank="species">
|
||
<emphasis id="B91CEAE5AE52B63FDEE94608FB8D032D" bold="true" box="[992,1110,327,349]" pageId="15" pageNumber="116">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C684D74AE52B63FD98C4608FA47032C" ID-CoL="75NLS" authority="(Balthasar, 1960)" authorityName="Balthasar S. Tarasov" authorityYear="2017" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[1157,1436,327,348]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="fukiensis">
|
||
<emphasis id="B91CEAE5AE52B63FD98C4608FB2B032C" box="[1157,1264,327,348]" italics="true" pageId="15" pageNumber="116">P. fukiensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B06AE52B63FD9F44608FA4E032C" author="Balthasar" box="[1277,1429,327,348]" pageId="15" pageNumber="116" refString="Balthasar, V. (1960) Einige neue Arten der Familie Scarabaeidae. Entomologische Blatter, 56, 88 - 94." type="journal article" year="1960">Balthasar, 1960</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BD736F7AE52B63FDD9E462AFEC2039F" blockId="15.[151,1436,151,495]" pageId="15" pageNumber="116">
|
||
<keyLead id="309995C2AE52B63FDD9E462AFEC2039F" pageId="15" pageNumber="116">
|
||
- Body shape round and distinctly convex (
|
||
<figureCitation id="13532A72AE52B63FDF67462AFD18030A" box="[622,707,357,378]" captionStart="FIGURE 15" captionStartId="29.[151,250,1247,1269]" captionTargetBox="[151,1436,267,1223]" captionTargetId="figure@29.[151,1436,267,1225]" captionTargetPageId="29" captionText="FIGURE 15. Morphology of Parachorius longipenis. (A) aedeagus, lateral view (paratype, Vietnam). (B) aedeagus, lateral view (paratype, Thailnd). (C) aedeagus, lateral view (paratype, Laos) (D) aedeagus, lateral view (holotype) (E) aedeagus. (F) habitus (male, holotype). (G) right metafemur (male, holotype). (H) right metafemur (male, paratype, Vietnam). (I) right metafemur (male, paratype, Thailand). (J) left protibia (flipped from original, male, holotype). (K) protibia (female, paratype, Laos)." httpUri="https://zenodo.org/record/1001947/files/figure.png" pageId="15" pageNumber="116">Fig. 15F</figureCitation>
|
||
); length
|
||
<quantity id="4C909B12AE52B63FDE1F462AFC57030A" box="[790,908,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.4" metricValueMin="3.8" pageId="15" pageNumber="116" unit="mm" value="4.6" valueMax="5.4" valueMin="3.8">3.8–5.4 mm</quantity>
|
||
. This couplet comprises four species with largely overlapping distributional areas and undistinguishable females. Nevertheless, species identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details)............................................
|
||
<taxonomicName id="4C684D74AE52B63FDDEA46F2FE6B03A2" ID-CoL="6TNM7" authority="Tarasov" authorityName="S. Tarasov" authorityYear="2017" box="[227,432,445,466]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="longipenis" status="sp. nov.">
|
||
<emphasis id="B91CEAE5AE52B63FDDEA46F2FE8503A2" box="[227,350,445,466]" italics="true" pageId="15" pageNumber="116">P. longipenis</emphasis>
|
||
Tarasov
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F579EAE52B63FDCBC46F3FDF103A2" box="[437,554,444,466]" pageId="15" pageNumber="116" rank="species">
|
||
<emphasis id="B91CEAE5AE52B63FDCBC46F3FDF103A2" bold="true" box="[437,554,444,466]" pageId="15" pageNumber="116">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C684D74AE52B63FDF3F46F2FCFF03A2" ID-CoL="4CPV3" authority="Tarasov" authorityName="Tarasov" authorityYear="2017" box="[566,804,445,466]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="solodovnikovi" status="sp. nov.">
|
||
<emphasis id="B91CEAE5AE52B63FDF3F46F2FD0903A2" box="[566,722,445,466]" italics="true" pageId="15" pageNumber="116">P. solodovnikovi</emphasis>
|
||
Tarasov
|
||
</taxonomicName>
|
||
<emphasis id="B91CEAE5AE52B63FDE2046F3FC7E03A2" bold="true" box="[809,933,444,466]" pageId="15" pageNumber="116">
|
||
<taxonomicNameLabel id="A22F579EAE52B63FDE2046F3FC7B03A2" box="[809,928,444,466]" pageId="15" pageNumber="116" rank="species">new species</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C684D74AE52B63FDEA346F2FB5D03A2" ID-CoL="6TNM8" authority="(Sharp, 1875)" authorityName="S. Tarasov" authorityYear="2017" baseAuthorityName="Sharp" baseAuthorityYear="1875" box="[938,1158,445,466]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="nudus">
|
||
<emphasis id="B91CEAE5AE52B63FDEA346F2FC2103A2" box="[938,1018,445,466]" italics="true" pageId="15" pageNumber="116">P. nudus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B06AE52B63FD90E46F2FBA403A2" author="Sharp" box="[1031,1151,445,466]" pageId="15" pageNumber="116" refString="Sharp, D. (1875) Descriptions of some new species of Scarabaeidae from Tropical Asia and Malasia. Part I. Coleopterologische Hefte, 13, 33 - 54." type="journal article" year="1875">Sharp, 1875</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C684D74AE52B63FD9B146F2FEC2039F" ID-CoL="75NLW" authority="(Arrow, 1907)" authorityName="Tarasov" authorityYear="2017" baseAuthorityName="Arrow" baseAuthorityYear="1907" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="peninsularis">
|
||
<emphasis id="B91CEAE5AE52B63FD9B146F2FA9D03A2" box="[1208,1350,445,466]" italics="true" pageId="15" pageNumber="116">P. peninsularis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF94B06AE52B63FD85846F2FEC9039F" author="Arrow" pageId="15" pageNumber="116" refString="Arrow, G. J. (1907) Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 2). The Annals and Magazine of Natural History, including Zoology, Botany and Geology, 7 (19), 416 - 439." type="journal article" year="1907">Arrow, 1907</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |