A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data Author Tarasov, Sergei text Zootaxa 2017 2017-10-04 4329 2 101 149 journal article 31902 10.11646/zootaxa.4329.2.1 7e33ec7d-211b-4042-8d07-344364ee805d 1175-5326 1001915 8226E27D-E3A9-481A-B51B-558643Bbb03A Key to females of Parachorius Twelve out of 19 Parachorius species cannot be reliably identified using females. Therefore, the key below contains some couplets that refer to several species at the same time. In such ambiguous cases the distribution criterion should be used to separate species. This criterion is not always reliable as many species have overlapping distributional areas. 1 Clypeus with two teeth, outer margin of each tooth not notched basally ( Fig. 4F–G )................................2 - Clypeus with two teeth, outer margin of each tooth distinctly or slightly notched basally ( Fig. 4D–E )...................6 2 Clypeus with two distinctly produced teeth ( Fig. 4G )......................................................... 3 - Clypeus with two less produced teeth ( Fig. 4F ).............................................................. 4 3 Body black; head triangularly shaped ( Fig. 21 ); length 6.5–6.8 mm ; southern China...... P. schuelkei Tarasov new species - Body reddish-brown, rarely dark brown; head more rounded not as distinctly triangular as in P. schuelkei ; length 6–7 mm .............................................. P. javanus ( Boucomont, 1914 ) and P. pseudojavanus Tarasov new species 4 Elytra covered with dense contiguous and rugose punctures ( Fig. 10F ); pronotum almost as long as elytra; wings reduced ( Fig. 10 ); 6–7mm ; northeastern India ....................................................... P. globosus Arrow, 1931 - Elytra covered with sparse (not contiguous) punctures ( Fig. 24F ); pronotum notably shorter than elytra; wings normally devel- oped............................................................................................... 5 5 Length 6 mm ( Fig. 12 ); northern India .................................................. P. hookeri Arrow, 1931 - Length 6.5–9.0 mm. This couplet contains four species with morphologically undistinguishable females. Their identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details)..................... P. newthayerae Tarasov new species , P. maruyamai Masumoto, Ochi , & Sakchoowong, 2012, P. thomsoni Harold, 1873 , P. fungorum Kryzhanovsky & Medvedev, 1966 6 Outer margin of each clypeal tooth slightly notched basally ( Fig. 4E ).............................................7 - Outer margin of each clypeal tooth distinctly notched basally ( Fig. 4D ).......................................... 8 7 Clypeal teeth acute; length 5–6 mm ( Fig. 11 ); Taiwan ................................... P. gotoi ( Masumoto, 1986 ) - Clypeal teeth rounded; length 8.2–10.6 mm ( Fig. 22 ); northern Laos.............. P. semsanganus Tarasov & Keith, 2011 8 Humeral area of elytron usually with a yellow spot, sometimes the yellow spot is absent and elytra uniformly colored; India .9 - Humeral area of elytron without yellow spot, elytra uniformly colored; southeastern China , Indochina, Malay Peninsula, and Sumatra ............................................................................................10 9 Species from southern and central India ( Fig. 6 ); length 4.5–4.7 mm ................ P. asymmetricus Tarasov new species - Species from northeastern India ( Sikkim , Assam , Meghalaya ); length 3.8–5.0 mm ( Fig. 13 )...... P. humeralis ( Arrow, 1907 ) 10 Body shape rectangular and less convex ( Figs. 7D , 8F ); length 3.5–4.6 mm ....................................................................................... P. bolavensis Tarasov new species and P. fukiensis ( Balthasar, 1960 ) - Body shape round and distinctly convex ( Fig. 15F ); length 3.8–5.4 mm . This couplet comprises four species with largely overlapping distributional areas and undistinguishable females. Nevertheless, species identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details)............................................ P. longipenis Tarasov new species , P. solodovnikovi Tarasov new species , P. nudus ( Sharp, 1875 ) , and P. peninsularis ( Arrow, 1907 )