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<document id="B38407B0740C79F4EB62F5A231895468" ID-DOI="10.1649/1176.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4924356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623349980529" checkinUser="carolina" docAuthor="Liebherr, James K." docDate="2009" docId="03A987B8FFFFEF286251FA03FD70FCFF" docLanguage="en" docName="TheColeopBull.63.4.383-411.pdf.imf" docOrigin="The Coleopterists Bulletin 63 (4)" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Blackburnia (Metromenus) filipes" docType="treatment" docVersion="3" lastPageNumber="401" masterDocId="FF90FFC0FFEFEF3A632AFFC5FFEAFFE5" masterDocTitle="Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island" masterLastPageNumber="411" masterPageNumber="383" pageNumber="399" updateTime="1698962403354" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="EE6873B57B83A0A2C7C8C256A5DBABB5">
<mods:title id="CDD2B75F7374B5F6E123A559FB345AF1">Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island</mods:title>
</mods:titleInfo>
<mods:name id="45DA5F1BE2182B461FEFF55F8902569E" type="personal">
<mods:role id="41354CD50E6C619AF9F4083FD6FC8DBC">
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<mods:namePart id="1D1D281EC5F623DA86AAF31A1E35B7D2">Liebherr, James K.</mods:namePart>
<mods:affiliation id="AB06519099916EAA8A91A48941FCC7E3">Department of Entomology, John H. and Anna B. Comstock Hall Cornell University Ithaca, NY 14853 - 2601, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="B222CFBDE02A0108EA1D7D689E21486E" type="email">JKL5@cornell.edu</mods:nameIdentifier>
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<mods:title id="4E7378E0F3D25863319F9B3AAB0A2F90">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="BF87AE487E57BD3E161AEC4C442750B9">
<mods:date id="A10524F5EECAA89EA4D83DDA378E96BC">2009</mods:date>
<mods:detail id="FBB0701B7ECBFCE3ABD91345E89A50C5" type="pubDate">
<mods:number id="A9B8739A0D2ECDAD1E8720E2F78292F8">2009-12-29</mods:number>
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<mods:detail id="33C3C17F1655DE9D9BBDD59977765789" type="volume">
<mods:number id="C0A9EE337D9774A8C925C626EA47BF5F">63</mods:number>
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<mods:detail id="09FDE5B0D1E942EC4C8C2635F33B7A60" type="issue">
<mods:number id="BE919228210F8C77D728E623702573EE">4</mods:number>
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<mods:classification id="33307869D5E4A3731D47B044A49DA9E1">journal article</mods:classification>
<mods:identifier id="D15C65B63E8C246ACF95520C231523AC" type="DOI">10.1649/1176.1</mods:identifier>
<mods:identifier id="6481B65377436CFCCE06B8B5668BD1BB" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="2756F0C0175934E4B8486EA1A0153A4A" type="Zenodo-Dep">4924356</mods:identifier>
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<treatment id="03A987B8FFFFEF286251FA03FD70FCFF" LSID="urn:lsid:plazi:treatment:03A987B8FFFFEF286251FA03FD70FCFF" httpUri="http://treatment.plazi.org/id/03A987B8FFFFEF286251FA03FD70FCFF" lastPageId="18" lastPageNumber="401" pageId="16" pageNumber="399">
<subSubSection id="C31A6525FFFFEF2A6251FA03FCCAFA39" box="[379,800,1478,1500]" pageId="16" pageNumber="399" type="nomenclature">
<paragraph id="8BBF36AEFFFFEF2A6251FA03FCCAFA39" blockId="16.[379,800,1478,1500]" box="[379,800,1478,1500]" pageId="16" pageNumber="399">
<heading id="D0F781C2FFFFEF2A6251FA03FCCAFA39" box="[379,800,1478,1500]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="399" reason="2">
<taxonomicName id="4C004D2DFFFFEF2A6251FA03FCCAFA39" ID-CoL="LYNB" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[379,800,1478,1500]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="399" phylum="Arthropoda" rank="species" species="filipes" subGenus="Metromenus">
<emphasis id="B974EABCFFFFEF2A6251FA03FCCAFA39" bold="true" box="[379,800,1478,1500]" pageId="16" pageNumber="399">
<emphasis id="B974EABCFFFFEF2A6251FA03FE1EFA39" bold="true" box="[379,500,1478,1500]" italics="true" pageId="16" pageNumber="399">Blackburnia</emphasis>
(
<emphasis id="B974EABCFFFFEF2A612FFA03FD68FA39" bold="true" box="[517,642,1478,1500]" italics="true" pageId="16" pageNumber="399">Metromenus</emphasis>
)
<emphasis id="B974EABCFFFFEF2A61B8FA03FD27FA39" bold="true" box="[658,717,1478,1500]" italics="true" pageId="16" pageNumber="399">filipes</emphasis>
(Sharp)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A6525FFFFEF2863EDFA37FD70FCFF" lastPageId="18" lastPageNumber="401" pageId="16" pageNumber="399" type="distribution">
<paragraph id="8BBF36AEFFFFEF2A63EDFA37FC06F9C7" blockId="16.[175,1004,1522,1570]" pageId="16" pageNumber="399">
<emphasis id="B974EABCFFFFEF2A63EDFA37FEACF9ED" bold="true" box="[199,326,1522,1544]" pageId="16" pageNumber="399">Distribution.</emphasis>
This species was collected by Perkins from Kaiholena Gulch behind Koele at
<quantity id="4CF89B4BFFFFEF2A6277F9C9FE77F9C7" box="[349,413,1548,1570]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="399" unit="m" value="600.0">600 m</quantity>
el. (lot ‘‘79. Lanai, Lanai Gulch Carabid 2 black legged,
</paragraph>
<caption id="DF7F6626FFFEEF2B638DFAD9FF0CFA92" ID-DOI="http://doi.org/10.5281/zenodo.4924370" ID-Zenodo-Dep="4924370" httpUri="https://zenodo.org/record/4924370/files/figure.png" pageId="17" pageNumber="400" startId="17.[167,198,1308,1327]" targetBox="[172,949,194,1286]" targetPageId="17">
<paragraph id="8BBF36AEFFFEEF2B638DFAD9FF0CFA92" blockId="17.[146,975,1308,1399]" pageId="17" pageNumber="400">
<emphasis id="B974EABCFFFEEF2B638DFAD9FF0CFACA" bold="true" box="[167,230,1308,1327]" pageId="17" pageNumber="400">Fig. 6.</emphasis>
Known distributions of carabid beetle species on Lanai. A.
<taxonomicName id="4C004D2DFFFEEF2B604EFAD9FEE8FAA2" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="lanaihalensis">
<emphasis id="B974EABCFFFEEF2B604EFAD9FEE8FAA2" italics="true" pageId="17" pageNumber="400">Blackburnia lanaihalensis</emphasis>
</taxonomicName>
. B.
<taxonomicName id="4C004D2DFFFEEF2B6205FAF1FE7DFAA2" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[303,407,1332,1351]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="depressa">
<emphasis id="B974EABCFFFEEF2B6205FAF1FE7DFAA2" box="[303,407,1332,1351]" italics="true" pageId="17" pageNumber="400">B. depressa</emphasis>
</taxonomicName>
. C.
<taxonomicName id="4C004D2DFFFEEF2B62EFFAF1FDFDFAA2" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[453,535,1332,1351]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="filipes">
<emphasis id="B974EABCFFFEEF2B62EFFAF1FDFDFAA2" box="[453,535,1332,1351]" italics="true" pageId="17" pageNumber="400">B. filipes</emphasis>
</taxonomicName>
. D.
<taxonomicName id="4C004D2DFFFEEF2B616DFAF1FCCEFAA2" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[583,804,1332,1351]" class="Insecta" family="Carabidae" genus="Metacolpodes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="buchanani">
<emphasis id="B974EABCFFFEEF2B616DFAF1FCCEFAA2" box="[583,804,1332,1351]" italics="true" pageId="17" pageNumber="400">Metacolpodes buchanani</emphasis>
</taxonomicName>
. E.
<taxonomicName id="4C004D2DFFFEEF2B607BFAF1FF27FABA" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="B974EABCFFFEEF2B607BFAF1FF27FABA" italics="true" pageId="17" pageNumber="400">Gnathaphanus picipes</emphasis>
</taxonomicName>
(small circles),
<taxonomicName id="4C004D2DFFFEEF2B6247FA89FE23FABA" box="[365,457,1356,1375]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="B974EABCFFFEEF2B6247FA89FE23FABA" box="[365,457,1356,1375]" italics="true" pageId="17" pageNumber="400">G. picipes</emphasis>
</taxonomicName>
plus
<taxonomicName id="4C004D2DFFFEEF2B612DFA89FD40FABA" box="[519,682,1356,1375]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="multipunctatus">
<emphasis id="B974EABCFFFEEF2B612DFA89FD40FABA" box="[519,682,1356,1375]" italics="true" pageId="17" pageNumber="400">G. multipunctatus</emphasis>
</taxonomicName>
(larger bullseye). F.
<taxonomicName id="4C004D2DFFFEEF2B6055FA89FF0AFA92" baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B974EABCFFFEEF2B6055FA89FF0AFA92" italics="true" pageId="17" pageNumber="400">Perigona nigriceps</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BBF36AEFFFEEF2863B8FA58FEF7FEE8" blockId="17.[146,975,1437,1568]" lastBlockId="18.[175,1005,194,484]" lastPageId="18" lastPageNumber="401" pageId="17" pageNumber="400">
under bark, about
<quantity id="4CF89B4BFFFEEF2B6270FA58FE5BFA56" box="[346,433,1437,1459]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="17" pageNumber="400" unit="ft" value="2.0">2.000 ft.</quantity>
I + II ’94’’; Anonymous N. D.), and also from the summit of Haalelepaakai (lot ‘‘134. Lanai, Halepaakai, VII. ’94’’; Anonymous N. D.). Recent records are restricted to the summit ridge at
<quantity id="4CF89B4BFFFEEF2B6066FA16FC25FA0C" box="[844,975,1491,1513]" metricMagnitude="3" metricUnit="m" metricValue="1.005" metricValueMax="1.02" metricValueMin="0.99" pageId="17" pageNumber="400" unit="m" value="1005.0" valueMax="1020.0" valueMin="990.0">990–1,020 m</quantity>
elevation, well within the confines of the cloud forest community (
<figureCitation id="133B2A2BFFFEEF2B6016FA2AFC92F9E0" box="[828,888,1519,1541]" captionStart="Fig" captionStartId="3.[167,198,915,934]" captionTargetBox="[181,941,194,893]" captionTargetId="figure-200@3.[181,941,194,893]" captionTargetPageId="3" captionText="Fig. 1. Pre-Polynesian vegetation communities on the island of Lanai (after Hobdy 1993): 1) cloud forest; 2) mesic forest; 3) dry forest; 4) arid grassland and shrubland; 5) coast and strand." figureDoi="http://doi.org/10.5281/zenodo.4924358" httpUri="https://zenodo.org/record/4924358/files/figure.png" pageId="17" pageNumber="400">Fig. 1</figureCitation>
). Cloud forest localities include Lanaihale, Haalelepaakai, and Waikeakua. As with
<taxonomicName id="4C004D2DFFFEEF28609BF9CFFF01FF32" baseAuthorityName="Sharp" baseAuthorityYear="1903" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" lastPageId="18" lastPageNumber="401" order="Coleoptera" pageId="17" pageNumber="400" phylum="Arthropoda" rank="species" species="filipes">
<emphasis id="B974EABCFFFEEF28609BF9CFFF01FF32" italics="true" lastPageId="18" lastPageNumber="401" pageId="17" pageNumber="400">M. filipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D2DFFFDEF286202FF07FE75FF32" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[296,415,194,215]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="depressa">
<emphasis id="B974EABCFFFDEF286202FF07FE75FF32" box="[296,415,194,215]" italics="true" pageId="18" pageNumber="401">B. depressa</emphasis>
</taxonomicName>
, a more extensive distribution based on all specimens (
<figureCitation id="133B2A2BFFFDEF28639DFF18FEEDFF16" box="[183,263,221,243]" captionStart="Fig" captionStartId="17.[167,198,1308,1327]" captionTargetBox="[172,949,194,1286]" captionTargetId="figure-66@17.[171,949,194,1286]" captionTargetPageId="17" captionText="Fig. 6. Known distributions of carabid beetle species on Lanai. A. Blackburnia lanaihalensis. B. B. depressa. C. B. filipes. D. Metacolpodes buchanani. E. Gnathaphanus picipes (small circles), G. picipes plus G. multipunctatus (larger bullseye). F. Perigona nigriceps." figureDoi="http://doi.org/10.5281/zenodo.4924370" httpUri="https://zenodo.org/record/4924370/files/figure.png" pageId="18" pageNumber="401">Fig. 6C</figureCitation>
) has been diminished through historical loss of habitats at lower elevations.
</paragraph>
<paragraph id="8BBF36AEFFFDEF2863EDFED6FEB4FE01" blockId="18.[175,1005,194,484]" pageId="18" pageNumber="401">
<emphasis id="B974EABCFFFDEF2863EDFED6FEF1FECC" bold="true" box="[199,283,275,297]" pageId="18" pageNumber="401">Habitat.</emphasis>
Beetles of this species have been beaten from native substrates including mossy ohia, soft understory ferns, and
<taxonomicName id="4C004D2DFFFDEF286196FEEBFCC2FEA6" box="[700,808,302,323]" class="Liliopsida" family="Pandanaceae" genus="Freycinetia" kingdom="Plantae" order="Pandanales" pageId="18" pageNumber="401" phylum="Tracheophyta" rank="species" species="arborea">
<emphasis id="B974EABCFFFDEF286196FEEBFCC2FEA6" box="[700,808,302,323]" italics="true" pageId="18" pageNumber="401">F. arborea</emphasis>
</taxonomicName>
. Within the cloud forest at Lanaihale and Waiakeakua these beetles have been collected on nonnative plants, including on the trunk of a live ‘‘
<collectingCountry id="F317763EFFFDEF2861B3FEA6FCDDFE9C" box="[665,823,355,377]" name="Norfolk Island" pageId="18" pageNumber="401">Norfolk Island</collectingCountry>
pine’’ (
<taxonomicName id="4C004D2DFFFDEF2860A2FEA1FEF1FE76" class="Insecta" family="Delphacidae" genus="Auracaria" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="columnaris">
<emphasis id="B974EABCFFFDEF2860A2FEA1FEF1FE76" italics="true" pageId="18" pageNumber="401">Auracaria columnaris</emphasis>
</taxonomicName>
[G. Forster]) (Little and Skolmen 1989), and on downed
<taxonomicName id="4C004D2DFFFDEF286054FEBBFC06FE76" box="[894,1004,382,403]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="18" pageNumber="401" phylum="Tracheophyta" rank="genus">
<emphasis id="B974EABCFFFDEF286054FEBBFC06FE76" box="[894,1004,382,403]" italics="true" pageId="18" pageNumber="401">Eucalyptus</emphasis>
</taxonomicName>
logs. As mentioned under
<taxonomicName id="4C004D2DFFFDEF2862FBFE5CFDDCFE4B" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[465,566,409,430]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="filipes">
<emphasis id="B974EABCFFFDEF2862FBFE5CFDDCFE4B" box="[465,566,409,430]" italics="true" pageId="18" pageNumber="401">M. filipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C004D2DFFFDEF286163FE5CFD4CFE4B" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[585,678,409,430]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="filipes">
<emphasis id="B974EABCFFFDEF286163FE5CFD4CFE4B" box="[585,678,409,430]" italics="true" pageId="18" pageNumber="401">B. filipes</emphasis>
</taxonomicName>
was collected by beating soft understory ferns along with the natives
<taxonomicName id="4C004D2DFFFDEF286161FE71FD54FE2C" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[587,702,436,457]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="depressa">
<emphasis id="B974EABCFFFDEF286161FE71FD54FE2C" box="[587,702,436,457]" italics="true" pageId="18" pageNumber="401">B. depressa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C004D2DFFFDEF2861DEFE71FCBFFE2C" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[756,853,436,457]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="filipes">
<emphasis id="B974EABCFFFDEF2861DEFE71FCBFFE2C" box="[756,853,436,457]" italics="true" pageId="18" pageNumber="401">M. filipes</emphasis>
</taxonomicName>
, plus the nonnative
<taxonomicName id="4C004D2DFFFDEF2863DFFE0AFEB2FE01" box="[245,344,463,484]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="picipes">
<emphasis id="B974EABCFFFDEF2863DFFE0AFEB2FE01" box="[245,344,463,484]" italics="true" pageId="18" pageNumber="401">G. picipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBF36AEFFFDEF286296FDC0FD35FDFE" blockId="18.[444,735,517,539]" box="[444,735,517,539]" pageId="18" pageNumber="401">
<emphasis id="B974EABCFFFDEF286296FDC0FD35FDFE" bold="true" box="[444,735,517,539]" pageId="18" pageNumber="401">Alien Carabid Beetle Species</emphasis>
</paragraph>
<paragraph id="8BBF36AEFFFDEF2863EDFDE8FCE8FD01" blockId="18.[175,1005,557,740]" pageId="18" pageNumber="401">
Four non-native carabid species are herein recorded from Lanai, three for the first time. These include
<taxonomicName id="4C004D2DFFFDEF286289FD8DFDC3FDB8" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[419,553,584,605]" class="Insecta" family="Carabidae" genus="Metacolpodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="buchanani">
<emphasis id="B974EABCFFFDEF286289FD8DFDC3FDB8" box="[419,553,584,605]" italics="true" pageId="18" pageNumber="401">M. buchanani</emphasis>
</taxonomicName>
, two species of the genus
<taxonomicName id="4C004D2DFFFDEF286003FD8DFEE5FD9C" authority="W. S. MacLeay" authorityName="W. S. MacLeay" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="genus">
<emphasis id="B974EABCFFFDEF286003FD8DFC5CFDB8" box="[809,950,584,605]" italics="true" pageId="18" pageNumber="401">Gnathaphanus</emphasis>
W.S. MacLeay
</taxonomicName>
(tribe
<taxonomicName id="4C004D2DFFFDEF286279FDA6FE52FD9C" box="[339,440,611,633]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini</taxonomicName>
), and
<taxonomicName id="4C004D2DFFFDEF2862DDFDA6FCE6FD9C" authority="(Dejean)" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[503,780,611,633]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B974EABCFFFDEF2862DDFDA6FD47FD9D" box="[503,685,611,632]" italics="true" pageId="18" pageNumber="401">Perigona nigriceps</emphasis>
(Dejean)
</taxonomicName>
(tribe
<taxonomicName id="4C004D2DFFFDEF28607AFDA6FC57FD9C" box="[848,957,611,633]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="tribe" tribe="Perigonini">Perigonini</taxonomicName>
). All can be identified to tribe using Liebherr and Zimmerman (2000). As there are several genera of
<taxonomicName id="4C004D2DFFFDEF286275FD5DFE2BFD4B" box="[351,449,664,686]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini</taxonomicName>
that include adventive representatives in
<collectingRegion id="49C4F84CFFFDEF28604BFD5DFC47FD4B" box="[865,941,664,686]" country="United States of America" name="Hawaii" pageId="18" pageNumber="401">Hawaii</collectingRegion>
, a key to the six species of
<taxonomicName id="4C004D2DFFFDEF2862BCFD76FDA3FD2C" authority="known" authorityName="Known" box="[406,585,691,713]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini known</taxonomicName>
to have been introduced to
<collectingRegion id="49C4F84CFFFDEF2860ADFD76FC3AFD2C" box="[903,976,691,713]" country="United States of America" name="Hawaii" pageId="18" pageNumber="401">Hawaii</collectingRegion>
is presented under treatment of that tribe’s species on Lanai.
</paragraph>
<paragraph id="8BBF36AEFFFDEF28612BFCC1FD70FCFF" blockId="18.[513,666,772,794]" box="[513,666,772,794]" pageId="18" pageNumber="401">
<emphasis id="B974EABCFFFDEF28612BFCC1FD70FCFF" bold="true" box="[513,666,772,794]" pageId="18" pageNumber="401">
Tribe
<taxonomicName id="4C004D2DFFFDEF286115FCC1FD70FCFF" authorityName="Bonelli" authorityYear="1810" box="[575,666,772,794]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="401" phylum="Arthropoda" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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