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<document id="473DC01CB4E09F0E2CD8EADD569A715B" ID-CLB-Dataset="27392" ID-DOI="10.5852/ejt.2019.512" ID-GBIF-Dataset="85c6241d-5e5a-4022-945f-e207d31458ef" ID-Zenodo-Dep="2628794" ID-ZooBank="urn:lsid:zoobank.org:pub:8DF89CFD-A549-4C69-8FBC-90EC8DEA9A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554393989085" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2019" docId="BE2487B5FF98FFD86D4D7BF24847FB65" docLanguage="en" docName="ejt-512_darby.pdf.imf" docOrigin="European Journal of Taxonomy 512" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium Motschulsky 1855" docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="421DFFCDFF93FFD46F0C7D524D30FFAA" masterDocTitle="New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London" masterLastPageNumber="50" masterPageNumber="1" pageNumber="12" updateTime="1698718664008" updateUser="ExternalLinkService">
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<mods:title id="483BB76ACC6DC63D752A7258D89FEA9F">New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London</mods:title>
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<mods:date id="3FF3F196CA3D8C6C63B44ACDBF9C8174">2019</mods:date>
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<treatment id="BE2487B5FF98FFD86D4D7BF24847FB65" ID-DOI="http://doi.org/10.5281/zenodo.5941272" ID-GBIF-Taxon="154555264" ID-Zenodo-Dep="5941272" LSID="urn:lsid:plazi:treatment:BE2487B5FF98FFD86D4D7BF24847FB65" httpUri="http://treatment.plazi.org/id/BE2487B5FF98FFD86D4D7BF24847FB65" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="7E976528FF98FFDF6D4D7BF24EC3F910" box="[577,1011,1695,1722]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="363236A3FF98FFDF6D4D7BF24EC3F910" blockId="11.[577,1011,1695,1722]" box="[577,1011,1695,1722]" pageId="11" pageNumber="12">
<heading id="6D7A81CFFF98FFDF6D4D7BF24EC3F910" box="[577,1011,1695,1722]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
Genus
<taxonomicName id="F18D4D20FF98FFDF6D9F7BCD4EC3F910" ID-CoL="8X77B" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[659,1011,1695,1722]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="04F9EAB1FF98FFDF6D9F7BCD4E3CF910" bold="true" box="[659,780,1695,1722]" italics="true" pageId="11" pageNumber="12">Cissidium</emphasis>
Motschulsky, 1855
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7E976528FF98FFD86FB17BB34969FE52" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
<paragraph id="363236A3FF98FFD86FB17BB34969FE52" blockId="11.[189,1399,1761,1998]" lastBlockId="12.[189,1399,265,505]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Genus of worldwide distribution particularly in the tropical regions with numerous species, many awaiting description. (Ongoing research by the author proposes to designate several subgenera.) The new species described below share the following features: body convex, pubescent, head finely margined at sides, eyes of male larger and more prominent than those of the female, antennae long, antennomere XI constricted medially producing a ‘dumbell’ or ‘hour–glass’ like figure easily mistaken for two separate antennomeres. Pronota broader than long, sides curved sometimes angulate with clearly marked raised margins, posterior margin with insinuation opposite scutellum. Elytra entire, longer than broad, generally leaving ¾ abdominal sclerites exposed. Prosternum very short anterior to procoxae which are contiguous. Mesosternum with a distinctive central keel reaching between the mesocoxae as a sharp point or wider, hind/side margin crenulate usually in posterior half, without clearly visible perforations [not withstanding that the tribe was erected to contain species with mesoventral perforations]. Metaventrum slightly longer than the mesoventrum, metacoxae separated by ± ¼ the width of the sclerite. Pygidium without teeth at apex. Dissections of the genitalia have not been attempted in some cases because external features are sufficient to determine the species and the risk of destruction to the specimens.</paragraph>
</subSubSection>
<subSubSection id="7E976528FF9FFFD86FB17F724847FB65" pageId="12" pageNumber="13" type="key">
<paragraph id="363236A3FF9FFFD86FB17F724F82FD91" blockId="12.[189,690,544,571]" box="[189,690,544,571]" pageId="12" pageNumber="13">
<heading id="6D7A81CFFF9FFFD86FB17F724F82FD91" bold="true" box="[189,690,544,571]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="04F9EAB1FF9FFFD86FB17F724F82FD91" bold="true" box="[189,690,544,571]" pageId="12" pageNumber="13">
Key to the
<collectingCountry id="4E9A7633FF9FFFD86E497F724C84FD90" box="[325,436,544,570]" name="Malaysia" pageId="12" pageNumber="13">Sarawak</collectingCountry>
species of
<taxonomicName id="F18D4D20FF9FFFD86D357F724F82FD91" ID-CoL="8X77B" authorityName="Motschulsky" authorityYear="1855" box="[569,690,544,571]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="04F9EAB1FF9FFFD86D357F724F82FD91" bold="true" box="[569,690,544,571]" italics="true" pageId="12" pageNumber="13">Cissidium</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<key id="421C6192FF9FFFD86FB17F1D4847FB65" pageId="12" pageNumber="13">
<keyStep id="8D792E06FF9FFFD86FB17F1D4847FD05" pageId="12" pageNumber="13">
<paragraph id="363236A3FF9FFFD86FB17F1D4847FD26" blockId="12.[189,1399,590,687]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FB17F1D4847FD26" pageId="12" pageNumber="13">
1. Thorax without either foveae or distinct foveolae, mentum triangular (
<figureCitation id="AEB62A26FF9FFFD86B2D7F1C494CFDC2" box="[1057,1148,590,616]" captionStart="Fig" captionStartId="14.[189,232,1943,1969]" captionTargetBox="[189,1398,696,1893]" captionTargetId="figure@14.[189,1398,695,1901]" captionTargetPageId="14" captionText="Fig. 6. Cissidium spp., habitus. A. C. marshallae sp. nov. B. C. pauxillum sp. nov. C. C. globulum sp. nov. D. C. subfoveolatum sp. nov. E. C. longum sp. nov. F. C. triangulum sp. nov." httpUri="https://zenodo.org/record/2628808/files/figure.png" pageId="12" pageNumber="13">Figs 6F</figureCitation>
,
<figureCitation id="AEB62A26FF9FFFD86B847F1D499BFDC3" box="[1160,1195,591,617]" captionStart="Fig" captionStartId="15.[189,232,1919,1945]" captionTargetBox="[195,1396,1135,1872]" captionTargetId="figure@15.[189,1399,1131,1882]" captionTargetPageId="15" captionText="Fig. 7. Cissidium spp., pronota.A. C. globulum sp. nov.×450. B. C. longum sp.nov. ×590.C. C.marshallae sp. nov. ×580. D. C. pauxillum sp. nov. ×590. E. C. subfoveolatum sp. nov. ×485. F. C. triangulum sp. nov. ×555." httpUri="https://zenodo.org/record/2628810/files/figure.png" pageId="12" pageNumber="13">7B</figureCitation>
) ........................ .........................................................................................................................
<taxonomicName id="F18D4D20FF9FFFD86B667F204825FD26" ID-CoL="8XXRY" box="[1130,1301,625,652]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="triangulum" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B667F204825FD26" bold="true" box="[1130,1301,625,652]" italics="true" pageId="12" pageNumber="13">C. triangulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A107F204847FD26" box="[1308,1399,626,652]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="363236A3FF9FFFD86FCD7FC74847FD05" blockId="12.[189,1399,590,687]" box="[193,1399,661,687]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FCD7FC74847FD05" box="[193,1399,661,687]" pageId="12" pageNumber="13">– Thorax with either foveae or distinct foveolae, mentum broadly rectangular .................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="8D792E06FF9FFFD86FB17F844847FC9D" pageId="12" pageNumber="13">
<paragraph id="363236A3FF9FFFD86FB17F844847FCBE" blockId="12.[189,1399,726,823]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FB17F844847FCBE" pageId="12" pageNumber="13">
2. Thorax with a series of foveae along the posterior margin the four central ones merging to form two larger ones (
<figureCitation id="AEB62A26FF9FFFD86E777FAB4CE4FCBE" box="[379,468,761,788]" captionStart="Fig" captionStartId="15.[189,232,1919,1945]" captionTargetBox="[195,1396,1135,1872]" captionTargetId="figure@15.[189,1399,1131,1882]" captionTargetPageId="15" captionText="Fig. 7. Cissidium spp., pronota.A. C. globulum sp. nov.×450. B. C. longum sp.nov. ×590.C. C.marshallae sp. nov. ×580. D. C. pauxillum sp. nov. ×590. E. C. subfoveolatum sp. nov. ×485. F. C. triangulum sp. nov. ×555." httpUri="https://zenodo.org/record/2628810/files/figure.png" pageId="12" pageNumber="13">Fig. 7E</figureCitation>
) .................................................................................
<taxonomicName id="F18D4D20FF9FFFD86B4D7FA84825FCBE" ID-CoL="8XXRV" box="[1089,1301,761,788]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subfoveolatum" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B4D7FA84825FCBE" bold="true" box="[1089,1301,761,788]" italics="true" pageId="12" pageNumber="13">C. subfoveolatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A107FA84847FCBE" box="[1308,1399,762,788]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="363236A3FF9FFFD86FCD7E4C4847FC9D" blockId="12.[189,1399,726,823]" box="[193,1399,797,823]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FCD7E4C4847FC9D" box="[193,1399,797,823]" pageId="12" pageNumber="13">
<emphasis id="04F9EAB1FF9FFFD86FCD7E4C4DE0FC9D" box="[193,208,798,823]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Thorax foveolate but without foveae forming large depressions ..................................................... 3
</keyLead>
</paragraph>
</keyStep>
<paragraph id="363236A3FF9FFFD86FB17E0C4847FC36" blockId="12.[189,1399,862,924]" pageId="12" pageNumber="13">
<keyStep id="8D792E06FF9FFFD86FB17E0C4847FC36" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FB17E0C4847FC36" pageId="12" pageNumber="13">
3. Elytra without foveolae, larger more rounded species (
<figureCitation id="AEB62A26FF9FFFD86C657E0C4EF4FCD3" box="[873,964,862,889]" captionStart="Fig" captionStartId="14.[189,232,1943,1969]" captionTargetBox="[189,1398,696,1893]" captionTargetId="figure@14.[189,1398,695,1901]" captionTargetPageId="14" captionText="Fig. 6. Cissidium spp., habitus. A. C. marshallae sp. nov. B. C. pauxillum sp. nov. C. C. globulum sp. nov. D. C. subfoveolatum sp. nov. E. C. longum sp. nov. F. C. triangulum sp. nov." httpUri="https://zenodo.org/record/2628808/files/figure.png" pageId="12" pageNumber="13">Fig. 6C</figureCitation>
) .......................
<taxonomicName id="F18D4D20FF9FFFD86B737E0C4825FCD3" ID-CoL="8XXQ6" box="[1151,1301,862,889]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="globulum" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B737E0C4825FCD3" bold="true" box="[1151,1301,862,889]" italics="true" pageId="12" pageNumber="13">C. globulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A107E0D4847FCD3" box="[1308,1399,863,889]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
– Elytra and pronotum foveolate ......................................................................................................... 4
</keyLead>
</keyStep>
</paragraph>
<keyStep id="8D792E06FF9FFFD86FB17E914847FB8E" pageId="12" pageNumber="13">
<paragraph id="363236A3FF9FFFD86FB17E914847FBAB" blockId="12.[189,1399,963,1060]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FB17E914847FBAB" pageId="12" pageNumber="13">
4. More rounded species, elytral foveolation less clearly marked than pronotum ................................. .........................................................................................................................
<taxonomicName id="F18D4D20FF9FFFD86B677EB44825FBAB" ID-CoL="8XXQR" box="[1131,1301,998,1025]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marshallae" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B677EB44825FBAB" bold="true" box="[1131,1301,998,1025]" italics="true" pageId="12" pageNumber="13">C. marshallae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A107EB54847FBAB" box="[1308,1399,999,1025]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="363236A3FF9FFFD86FCD79584847FB8E" blockId="12.[189,1399,963,1060]" box="[193,1399,1033,1060]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FCD79584847FB8E" box="[193,1399,1033,1060]" pageId="12" pageNumber="13">– Smaller, more parallel sided species ................................................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="8D792E06FF9FFFD86FB179184847FB65" pageId="12" pageNumber="13">
<paragraph id="363236A3FF9FFFD86FB179184847FB22" blockId="12.[189,1399,1098,1231]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FB179184847FB22" pageId="12" pageNumber="13">
5. Lateral margins of mesoventrum straight forming a distinct angle with the posterior margin, mesoventral keel narrow, ± parallel sided (
<figureCitation id="AEB62A26FF9FFFD86DD2793C4E09FB22" box="[734,825,1134,1160]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[200,1390,746,1884]" captionTargetId="figure@18.[189,1399,733,1887]" captionTargetPageId="18" captionText="Fig. 8. Cissidium spp., mesoventra. A. C. globulum sp. nov. ×470. B. C. pauxillum sp. nov ×725. C. C. marshallae sp. nov ×335. D. C. longum sp. nov. ×450. E. C. subfoveolatum sp. nov. ×455. F. C. triangulum sp. nov. ×575." httpUri="https://zenodo.org/record/2628812/files/figure.png" pageId="12" pageNumber="13">Fig. 8D</figureCitation>
) .............................................
<taxonomicName id="F18D4D20FF9FFFD86B9A793C4825FB22" ID-CoL="8XXQL" box="[1174,1301,1133,1160]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="longum" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B9A793C4825FB22" bold="true" box="[1174,1301,1133,1160]" italics="true" pageId="12" pageNumber="13">C. longum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A10793C4847FB22" box="[1308,1399,1134,1160]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="363236A3FF9FFFD86FCD79C04847FB65" blockId="12.[189,1399,1098,1231]" pageId="12" pageNumber="13">
<keyLead id="8D7C9596FF9FFFD86FCD79C04847FB65" pageId="12" pageNumber="13">
– Lateral margins of mesoventrum rounded, thorax densely foveolate (
<figureCitation id="AEB62A26FF9FFFD86B1C79C3495AFB01" box="[1040,1130,1169,1195]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[200,1390,746,1884]" captionTargetId="figure@18.[189,1399,733,1887]" captionTargetPageId="18" captionText="Fig. 8. Cissidium spp., mesoventra. A. C. globulum sp. nov. ×470. B. C. pauxillum sp. nov ×725. C. C. marshallae sp. nov ×335. D. C. longum sp. nov. ×450. E. C. subfoveolatum sp. nov. ×455. F. C. triangulum sp. nov. ×575." httpUri="https://zenodo.org/record/2628812/files/figure.png" pageId="12" pageNumber="13">Fig. 8B</figureCitation>
) ................................. ...........................................................................................................................
<taxonomicName id="F18D4D20FF9FFFD86B7B79E64825FB65" ID-CoL="8XXRB" box="[1143,1301,1204,1231]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pauxillum" status="sp. nov.">
<emphasis id="04F9EAB1FF9FFFD86B7B79E64825FB65" bold="true" box="[1143,1301,1204,1231]" italics="true" pageId="12" pageNumber="13">C. pauxillum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF9FFFD86A1079E74847FB65" box="[1308,1399,1205,1231]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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