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<document id="425836B977F44DCDBE26825B57B381C2" ID-CLB-Dataset="27392" ID-DOI="10.5852/ejt.2019.512" ID-GBIF-Dataset="85c6241d-5e5a-4022-945f-e207d31458ef" ID-Zenodo-Dep="2628794" ID-ZooBank="urn:lsid:zoobank.org:pub:8DF89CFD-A549-4C69-8FBC-90EC8DEA9A66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554393989085" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2019" docId="BE2487B5FF8BFFCE6D6778954FF0FD3F" docLanguage="en" docName="ejt-512_darby.pdf.imf" docOrigin="European Journal of Taxonomy 512" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Niptella gutta Darby, 2019, gen. et sp. nov." docType="treatment" docUuid="B6E16C2B-3B88-418C-9536-6E235936248C" docUuidSource="ZooBank" docVersion="9" lastPageNumber="27" masterDocId="421DFFCDFF93FFD46F0C7D524D30FFAA" masterDocTitle="New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London" masterLastPageNumber="50" masterPageNumber="1" pageNumber="25" updateTime="1698718664008" updateUser="ExternalLinkService">
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<mods:title id="BB1768253F01E97A29B8A61BFD48E2C9">New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London</mods:title>
</mods:titleInfo>
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<mods:namePart id="A8304083DA7B1A20880F511F14987888">Darby, Michael</mods:namePart>
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<mods:typeOfResource id="E106B247863A36846AEFF313655CD907">text</mods:typeOfResource>
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<mods:title id="94E0F3C09836644ED9EBADB1EAF528EB">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="37F12357F6DDB1B64CF8BCCB59252BC1">
<mods:date id="1E1F75F7DFB611F6B51453959DFE9E16">2019</mods:date>
<mods:detail id="4AF2682E35ADB04B29FCA011C8CCDC84" type="pubDate">
<mods:number id="E4B7E28082D5B9C689ADA40955D56831">2019-04-04</mods:number>
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<mods:detail id="E358D5624C403AA4DA30985119724157" type="volume">
<mods:number id="A728649EA6E2171F96193EBB94384C4C">512</mods:number>
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<mods:classification id="772F7125594CC5C0F57066A272B02943">journal article</mods:classification>
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<mods:identifier id="BD186F364E09FD40A21C821112B363BB" type="DOI">10.5852/ejt.2019.512</mods:identifier>
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</mods:mods>
<treatment id="BE2487B5FF8BFFCE6D6778954FF0FD3F" ID-DOI="http://doi.org/10.5281/zenodo.5941298" ID-GBIF-Taxon="154555279" ID-Zenodo-Dep="5941298" LSID="urn:lsid:zoobank.org:act:B6E16C2B-3B88-418C-9536-6E235936248C" httpUri="http://treatment.plazi.org/id/BE2487B5FF8BFFCE6D6778954FF0FD3F" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="7E976528FF8BFFCC6D677895498CF9AC" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="363236A3FF8BFFCC6D6778954EF8FA48" blockId="24.[375,1212,1479,1576]" box="[619,968,1479,1506]" pageId="24" pageNumber="25">
<heading id="6D7A81CFFF8BFFCC6D6778954EF8FA48" box="[619,968,1479,1506]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="F18D4D20FF8BFFCC6D6778954E21FA48" ID-CoL="8Y6PB" box="[619,785,1479,1506]" class="Insecta" family="Ptiliidae" genus="Niptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gutta" status="gen. et sp. nov.">
<emphasis id="04F9EAB1FF8BFFCC6D6778954E21FA48" bold="true" box="[619,785,1479,1506]" italics="true" pageId="24" pageNumber="25">Niptella gutta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF8BFFCC6C14789A4EF8FA48" box="[792,968,1480,1506]" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="363236A3FF8BFFCC6E7B78B9498CF9AC" blockId="24.[375,1212,1479,1576]" box="[375,1212,1515,1542]" pageId="24" pageNumber="25">
<uri id="421C3AA1FF8BFFCC6E7B78B9498CF9AC" box="[375,1212,1515,1542]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act:
<uuid id="422B0C76FF8BFFCC6D9B78B9498CF9AC" box="[663,1212,1515,1542]" pageId="24" pageNumber="25">B6E16C2B-3B88-418C-9536-6E235936248C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="7E976528FF8BFFCC6DFC7B5C4E73F982" box="[752,835,1550,1576]" pageId="24" pageNumber="25" type="description">
<paragraph id="363236A3FF8BFFCC6DFC7B5C4E73F982" blockId="24.[375,1212,1479,1576]" box="[752,835,1550,1576]" pageId="24" pageNumber="25">
<figureCitation id="AEB62A26FF8BFFCC6DFC7B5C4E73F982" box="[752,835,1550,1576]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="24" pageNumber="25">Fig. 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7E976528FF8BFFCC6FB17B1D4FC9F932" box="[189,761,1615,1688]" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="363236A3FF8BFFCC6FB17B1D4C06F9C3" blockId="24.[189,310,1615,1641]" box="[189,310,1615,1641]" pageId="24" pageNumber="25">
<heading id="6D7A81CFFF8BFFCC6FB17B1D4C06F9C3" bold="true" box="[189,310,1615,1641]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="04F9EAB1FF8BFFCC6FB17B1D4C06F9C3" bold="true" box="[189,310,1615,1641]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="363236A3FF8BFFCC6FB17B2C4FC9F932" blockId="24.[189,761,1662,1688]" box="[189,761,1662,1688]" pageId="24" pageNumber="25">Determined by the form of the mesoventral keel.</paragraph>
</subSubSection>
<subSubSection id="7E976528FF8BFFCC6FB17B9249D5F8A2" box="[189,1253,1728,1801]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="363236A3FF8BFFCC6FB17B924C74F970" blockId="24.[189,324,1728,1754]" box="[189,324,1728,1754]" pageId="24" pageNumber="25">
<heading id="6D7A81CFFF8BFFCC6FB17B924C74F970" bold="true" box="[189,324,1728,1754]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="04F9EAB1FF8BFFCC6FB17B924C74F970" bold="true" box="[189,324,1728,1754]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="363236A3FF8BFFCC6FB17BBD49D5F8A2" blockId="24.[189,1253,1774,1801]" box="[189,1253,1774,1801]" pageId="24" pageNumber="25">
From the Latin
<taxonomicName id="F18D4D20FF8BFFCC6E727BBD4C8AF8A2" box="[382,442,1775,1800]" class="Insecta" family="Ptiliidae" genus="Niptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gutta">
<emphasis id="04F9EAB1FF8BFFCC6E727BBD4C8AF8A2" box="[382,442,1775,1800]" italics="true" pageId="24" pageNumber="25">gutta</emphasis>
</taxonomicName>
, meaning drop, and referring to the shape of the mesoventral keel.
</paragraph>
</subSubSection>
<subSubSection id="7E976528FF8BFFCD6FB17A624C2BFE7F" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="363236A3FF8BFFCC6FB17A624C5CF8E3" blockId="24.[189,364,1839,1866]" box="[189,364,1839,1866]" pageId="24" pageNumber="25">
<heading id="6D7A81CFFF8BFFCC6FB17A624C5CF8E3" bold="true" box="[189,364,1839,1866]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="04F9EAB1FF8BFFCC6FB17A624C5CF8E3" bold="true" box="[189,364,1839,1866]" pageId="24" pageNumber="25">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="363236A3FF8BFFCC6FE67A0C492DF815" blockId="24.[189,1398,1886,1983]" pageId="24" pageNumber="25">
<materialsCitation id="86E53CFEFF8BFFCC6FE67A0C492DF815" ID-GBIF-Occurrence="2234828129" collectingDate="1978-08" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="1930" location="Gunung Mulu National Park" pageId="24" pageNumber="25" specimenCode="BM 1978-49" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="E9368801FF8BFFCC6FE67A0C4C6CF8D2" box="[234,348,1886,1912]" pageId="24" pageNumber="25" type="holotype">
<emphasis id="04F9EAB1FF8BFFCC6FE67A0C4C6CF8D2" bold="true" box="[234,348,1886,1912]" pageId="24" pageNumber="25">Holotype</emphasis>
</typeStatus>
MALAYSIA-BORNEO •
<specimenCount id="208BFD2AFF8BFFCC6EFF7AD04F3AF836" box="[499,522,1922,1948]" pageId="24" pageNumber="25" type="female"></specimenCount>
;
<collectingCountry id="4E9A7633FF8BFFCC6D167AD34FB2F831" box="[538,642,1921,1947]" name="Malaysia" pageId="24" pageNumber="25">Sarawak</collectingCountry>
, 4
<superScript id="C1F89BEBFF8BFFCC6D927A2D4F9BF827" attach="left" box="[670,683,1919,1933]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
division,
<location id="33526078FF8BFFCC6C2F7AD3494BF831" LSID="urn:lsid:plazi:treatment:BE2487B5FF8BFFCE6D6778954FF0FD3F:33526078FF8BFFCC6C2F7AD3494BF831" box="[803,1147,1921,1947]" country="Malaysia" name="Gunung Mulu National Park" pageId="24" pageNumber="25">Gunung Mulu National Park</location>
;
<elevation id="BDA0D190FF8BFFCC6B8B7AD34813F831" box="[1159,1315,1921,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="24" pageNumber="25" unit="m" value="1930.0">
<quantity id="F1759B46FF8BFFCC6B8B7AD349D2F836" box="[1159,1250,1921,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="24" pageNumber="25" unit="m" value="1930.0">1930 m</quantity>
a.s.l.
</elevation>
; May
<collectingDate id="5277E98BFF8BFFCC6FB17AF74C0CF815" box="[189,316,1957,1983]" pageId="24" pageNumber="25" value="1978-08">
<date id="42331063FF8BFFCC6FB17AF74C0CF815" box="[189,316,1957,1983]" pageId="24" pageNumber="25" value="1978-08">Aug. 1978</date>
</collectingDate>
;
<collectorName id="9B785375FF8BFFCC6E447AF74F31F814" box="[328,513,1956,1983]" pageId="24" pageNumber="25">P.M. Hammond</collectorName>
and
<collectorName id="9B785375FF8BFFCC6D367AF74FE6F814" box="[570,726,1956,1983]" pageId="24" pageNumber="25">J.E. Marshall</collectorName>
leg.,
<specimenCode id="662B9ED8FF8BFFCC6C1B7AF74E82F815" box="[791,946,1957,1983]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="24" pageNumber="25">BM 1978-49</specimenCode>
;
<collectionCode id="509CAE66FF8BFFCC6CB17AF74929F815" box="[957,1049,1957,1983]" pageId="24" pageNumber="25">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="363236A3FF8AFFCD6FE67C594C57FE8F" blockId="25.[189,1398,267,469]" box="[234,359,267,293]" pageId="25" pageNumber="26">
<typeStatus id="E9368801FF8AFFCD6FE67C594C57FE8F" box="[234,359,267,293]" pageId="25" pageNumber="26" type="paratype">
<emphasis id="04F9EAB1FF8AFFCD6FE67C594C57FE8F" bold="true" box="[234,359,267,293]" pageId="25" pageNumber="26">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="363236A3FF8AFFCD6FB17C7C4C2BFE7F" blockId="25.[189,1398,267,469]" pageId="25" pageNumber="26">
MALAYSIA-BORNEO •
<materialsCitation id="86E53CFEFF8AFFCD6EE37C7D4E9DFEE3" ID-GBIF-Occurrence="2234828183" box="[495,941,302,329]" collectingDate="1978-08" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="1930" location="Gunung Mulu National Park" pageId="25" pageNumber="26" specimenCount="1" typeStatus="paratype">
♂; same data as for holotype;
<collectionCode id="509CAE66FF8AFFCD6C5B7C7D4E9DFEE3" box="[855,941,303,329]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="86E53CFEFF8AFFCD6CC07C7C4F75FEC6" ID-GBIF-Occurrence="2234828194" collectingDate="1978-08" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="1850" location="Gunung Mulu National Park" pageId="25" pageNumber="26" specimenCount="1" typeStatus="paratype">
2 specs; same data as for preceding;
<elevation id="BDA0D190FF8AFFCD6FB17C034CA9FEC1" box="[189,409,337,364]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="25" pageNumber="26" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<quantity id="F1759B46FF8AFFCD6FB17C034C68FEC6" box="[189,344,337,364]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="25" pageNumber="26" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800-1900 m</quantity>
a.s.l.
</elevation>
; litter;
<collectionCode id="509CAE66FF8AFFCD6EE37C004F75FEC6" box="[495,581,338,364]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="86E53CFEFF8AFFCD6D687C034CF6FE18" ID-GBIF-Occurrence="2234828128" collectionCode="BMNH" collectorName="M. Hammond &amp; J. E. Marshall" country="Malaysia" location="Gunung Mulu National Park" municipality="May-Aug." pageId="25" pageNumber="26" specimenCode="BM 1978-49" specimenCount="1" typeStatus="paratype">
1 spec. (mounted disarticulated on the slide);
<collectingCountry id="4E9A7633FF8AFFCD6B707C0349D1FEC1" box="[1148,1249,337,363]" name="Malaysia" pageId="25" pageNumber="26">Sarawak</collectingCountry>
, 4
<superScript id="C1F89BEBFF8AFFCD6BF17C02483AFEF4" attach="left" box="[1277,1290,336,350]" fontSize="6" pageId="25" pageNumber="26">th</superScript>
division,
<location id="33526078FF8AFFCD6FB17C274F37FE24" LSID="urn:lsid:plazi:treatment:BE2487B5FF8BFFCE6D6778954FF0FD3F:33526078FF8AFFCD6FB17C274F37FE24" box="[189,519,372,399]" country="Malaysia" municipality="May-Aug." name="Gunung Mulu National Park" pageId="25" pageNumber="26">Gunung Mulu National Park</location>
;
<collectingMunicipality id="D656ACD9FF8AFFCD6D1E7C274FBDFE25" box="[530,653,373,399]" pageId="25" pageNumber="26">MayAug.</collectingMunicipality>
1978; P.
<collectorName id="9B785375FF8AFFCD6DFD7C274EA1FE24" box="[753,913,372,399]" pageId="25" pageNumber="26">M. Hammond</collectorName>
and
<collectorName id="9B785375FF8AFFCD6CC57C274950FE24" box="[969,1120,372,399]" pageId="25" pageNumber="26">J.E. Marshall</collectorName>
leg.,
<specimenCode id="662B9ED8FF8AFFCD6B937C274805FE25" box="[1183,1333,373,399]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="25" pageNumber="26">BM 1978-49</specimenCode>
; litter below comp. 4;
<collectionCode id="509CAE66FF8AFFCD6E7C7CCA4CF6FE18" box="[368,454,408,434]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="86E53CFEFF8AFFCD6ED07CCA4C27FE7F" ID-GBIF-Occurrence="2234828178" collectingDate="1978-03-13" collectionCode="BMNH" collectorName="N. M. Collins" country="Malaysia" location="Mulu" pageId="25" pageNumber="26" specimenCount="1" typeStatus="paratype">
1 spec.;
<collectingCountry id="4E9A7633FF8AFFCD6D3B7CCA4FACFE18" box="[567,668,408,434]" name="Malaysia" pageId="25" pageNumber="26">Sarawak</collectingCountry>
,
<location id="33526078FF8AFFCD6DA97CCA4FD6FE18" LSID="urn:lsid:plazi:treatment:BE2487B5FF8BFFCE6D6778954FF0FD3F:33526078FF8AFFCD6DA97CCA4FD6FE18" box="[677,742,408,434]" country="Malaysia" name="Mulu" pageId="25" pageNumber="26">Mulu</location>
;
<collectingDate id="5277E98BFF8AFFCD6DFD7CCA4EBAFE18" box="[753,906,408,434]" pageId="25" pageNumber="26" value="1978-03-13">
<date id="42331063FF8AFFCD6DFD7CCA4EBAFE18" box="[753,906,408,434]" pageId="25" pageNumber="26" value="1978-03-13">13 Mar. 1978</date>
</collectingDate>
;
<collectorName id="9B785375FF8AFFCD6C987CCA4918FE18" box="[916,1064,408,434]" pageId="25" pageNumber="26">N.M. Collins</collectorName>
leg.; pitfalls in alluvial forest;
<collectionCode id="509CAE66FF8AFFCD6FB17CE94C27FE7F" box="[189,279,443,469]" pageId="25" pageNumber="26">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7E976528FF8AFFCE6FB17CAE4FF0FD3F" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="363236A3FF8AFFCD6FB17CAE4C7EFDBC" blockId="25.[189,334,508,534]" box="[189,334,508,534]" pageId="25" pageNumber="26">
<heading id="6D7A81CFFF8AFFCD6FB17CAE4C7EFDBC" bold="true" box="[189,334,508,534]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="04F9EAB1FF8AFFCD6FB17CAE4C7EFDBC" bold="true" box="[189,334,508,534]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="363236A3FF8AFFCD6FB17F794F9AFDEF" blockId="25.[189,682,555,582]" box="[189,682,555,582]" pageId="25" pageNumber="26">
SIZE. Habitus (
<figureCitation id="AEB62A26FF8AFFCD6E6B7F794CE4FDEF" box="[359,468,555,581]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
), length
<quantity id="F1759B46FF8AFFCD6D367F794F95FDEF" box="[570,677,555,582]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="26" unit="mm" value="0.65">0.65 mm</quantity>
.
</paragraph>
<paragraph id="363236A3FF8AFFCD6FB17F3E497EFD2C" blockId="25.[189,1102,620,647]" box="[189,1102,620,647]" pageId="25" pageNumber="26">COLOUR. Yellowish brown shining, closely covered with fine pale pubescence.</paragraph>
<paragraph id="363236A3FF8AFFCD6FB17FFC4F5DFCA4" blockId="25.[189,1399,686,783]" pageId="25" pageNumber="26">
HEAD. With row of postulate setae behind the eyes (
<figureCitation id="AEB62A26FF8AFFCD6C467FFC4E88FD62" box="[842,952,686,712]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="25" pageNumber="26">Fig. 12F</figureCitation>
). Antennae very pale yellow almost translucent, antennomeres IIIXI, length
<quantity id="F1759B46FF8AFFCD6DBE7F834E12FD41" box="[690,802,721,747]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="25" pageNumber="26" unit="mm" value="0.28">0.28 mm</quantity>
. Eyes prominent, distance across eyes
<quantity id="F1759B46FF8AFFCD6A0E7F834842FD41" box="[1282,1394,721,747]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="mm" value="0.24">0.24 mm</quantity>
. Mentum and submentum (
<figureCitation id="AEB62A26FF8AFFCD6EFE7FA64F50FCA4" box="[498,608,756,782]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="25" pageNumber="26">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="363236A3FF8AFFCD6FB17E644E7DFCFA" blockId="25.[189,845,821,848]" box="[189,845,821,848]" pageId="25" pageNumber="26">
PRONOTUM. Length
<quantity id="F1759B46FF8AFFCD6E927E644F3AFCFA" box="[414,522,822,848]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="25" pageNumber="26" unit="mm" value="0.19">0.19 mm</quantity>
, width
<quantity id="F1759B46FF8AFFCD6D537E644FF8FCFA" box="[607,712,822,848]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="26" unit="mm" value="0.3">0.30 mm</quantity>
(
<figureCitation id="AEB62A26FF8AFFCD6DD47E674E70FCFA" box="[728,832,821,848]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="25" pageNumber="26">Fig. 12F</figureCitation>
).
</paragraph>
<caption id="62F2662BFF8AFFCD6FB17A214C78F87E" httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1907,1933]" targetBox="[194,1398,949,1839]" targetPageId="25">
<paragraph id="363236A3FF8AFFCD6FB17A214C78F87E" blockId="25.[189,1399,1907,2004]" pageId="25" pageNumber="26">
<emphasis id="04F9EAB1FF8AFFCD6FB17A214C2DF824" bold="true" box="[189,285,1907,1934]" pageId="25" pageNumber="26">Fig. 12.</emphasis>
<taxonomicName id="F18D4D20FF8AFFCD6E257A264CFFF824" box="[297,463,1908,1934]" class="Insecta" family="Ptiliidae" genus="Niptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="gutta" status="gen. et sp. nov.">
<emphasis id="04F9EAB1FF8AFFCD6E257A264CFFF824" box="[297,463,1908,1934]" italics="true" pageId="25" pageNumber="26">Niptella gutta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1FCA57CAFF8AFFCD6ED67A264FA7F824" box="[474,663,1908,1934]" pageId="25" pageNumber="26" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="04F9EAB1FF8AFFCD6DAE7A214F87F827" bold="true" box="[674,695,1907,1933]" pageId="25" pageNumber="26">A</emphasis>
. Habitus.
<emphasis id="04F9EAB1FF8AFFCD6C347A264E7CF824" bold="true" box="[824,844,1908,1934]" pageId="25" pageNumber="26">B</emphasis>
. Spermatheca.
<emphasis id="04F9EAB1FF8AFFCD6B047A21492DF827" bold="true" box="[1032,1053,1907,1933]" pageId="25" pageNumber="26">C</emphasis>
. Aedeagus.
<emphasis id="04F9EAB1FF8AFFCD6BB97A2649FAF824" bold="true" box="[1205,1226,1908,1934]" pageId="25" pageNumber="26">D</emphasis>
. Mentum and submentum ×860.
<emphasis id="04F9EAB1FF8AFFCD6E917AC54C81F81B" bold="true" box="[413,433,1943,1969]" pageId="25" pageNumber="26">E</emphasis>
. Pygidium.
<emphasis id="04F9EAB1FF8AFFCD6D497AC54F67F81B" bold="true" box="[581,599,1943,1969]" pageId="25" pageNumber="26">F</emphasis>
. Pronotum ×540.
<emphasis id="04F9EAB1FF8AFFCD6C3E7AC54E79F81B" bold="true" box="[818,841,1943,1969]" pageId="25" pageNumber="26">G</emphasis>
. Meso- and metaventra ×470.
<emphasis id="04F9EAB1FF8AFFCD6BB07AC549E3F81B" bold="true" box="[1212,1235,1943,1969]" pageId="25" pageNumber="26">H</emphasis>
. Mesoventral keel ×1060.
</paragraph>
</caption>
<paragraph id="363236A3FF89FFCE6FB17C5E4E29FE8C" blockId="26.[189,793,267,294]" box="[189,793,267,294]" pageId="26" pageNumber="27">
ELYTRA. Length
<quantity id="F1759B46FF89FFCE6E767C594CD6FE8C" box="[378,486,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="27" unit="mm" value="0.39">0.39 mm</quantity>
, width
<quantity id="F1759B46FF89FFCE6D377C594F97FE8C" box="[571,679,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="mm" value="0.35">0.35 mm</quantity>
. truncate.
</paragraph>
<paragraph id="363236A3FF89FFCE6FB17C1F4E1EFE7A" blockId="26.[189,1399,333,465]" pageId="26" pageNumber="27">
VENTRUM. Proventrum wide before contiguous procoxae, reticulate, with scattered setae. Mesoventrum (
<figureCitation id="AEB62A26FF89FFCE6FCA7C224C07FE20" box="[198,311,368,394]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12G</figureCitation>
), collar not extending onto sharply toothed humeri, keel in shape of a peardrop stopping short of collar and narrowing to a point between the mesocoxae (
<figureCitation id="AEB62A26FF89FFCE6CCB7CC14905FE07" box="[967,1077,403,429]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12H</figureCitation>
). Metaventrum (
<figureCitation id="AEB62A26FF89FFCE6BF77CC14859FE07" box="[1275,1385,403,429]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12G</figureCitation>
), metacoxae separated by half the width of the sclerite.
</paragraph>
<paragraph id="363236A3FF89FFCE6FB17CAA4F03FDB8" blockId="26.[189,563,504,530]" box="[189,563,504,530]" pageId="26" pageNumber="27">
ABDOMEN. Pygidium (
<figureCitation id="AEB62A26FF89FFCE6EB07CAA4F16FDB8" box="[444,550,504,530]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12E</figureCitation>
).
</paragraph>
<paragraph id="363236A3FF89FFCE6FB17F68499AFDF9" blockId="26.[189,1194,569,596]" box="[189,1194,569,596]" pageId="26" pageNumber="27">
FEMALE GENITALIA. Spermatheca globular, very small and difficult to detect (
<figureCitation id="AEB62A26FF89FFCE6B3E7F6B49ADFDF9" box="[1074,1181,569,595]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12B</figureCitation>
).
</paragraph>
<paragraph id="363236A3FF89FFCE6FB17F294FF0FD3F" blockId="26.[189,704,635,661]" box="[189,704,635,661]" pageId="26" pageNumber="27">
MALE GENITALIA. Aedeagus large (
<figureCitation id="AEB62A26FF89FFCE6D497F294F83FD3F" box="[581,691,635,661]" captionStart="Fig" captionStartId="25.[189,232,1907,1933]" captionTargetBox="[194,1398,949,1839]" captionTargetId="figure@25.[189,1399,949,1842]" captionTargetPageId="25" captionText="Fig. 12. Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060." httpUri="https://zenodo.org/record/2628821/files/figure.png" pageId="26" pageNumber="27">Fig. 12G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>