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<document id="25349EA920A47CACDBBEFE90388DD41B" ID-DOI="10.5281/zenodo.12808371" ID-ISSN="1942-1354" ID-Zenodo-Dep="12808371" ID-ZooBank="0D2107D1-BD4D-426C-ABCF-C4CC3EEA060F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721837453179" checkinUser="felipe" docAuthor="Skelley, Paul E." docDate="2021" docId="03F687B7FFE19568FF56FF7AF73AFAD0" docLanguage="en" docName="InsectaMundi.2021.0892.1-7.pdf" docOrigin="Insecta Mundi 2021 (892)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Leptorhyparus gilli Howden 2003" docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="FFCFFFCFFFE4956DFF96FFAFF47BFF9A" masterDocTitle="A new species of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) in amber from the Dominican Republic, with comments on extant species" masterLastPageNumber="7" masterPageNumber="1" pageNumber="4" updateTime="1721840401121" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="2E96FEC031BC7E7042EE0E29277149D9">A new species of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) in amber from the Dominican Republic, with comments on extant species</mods:title>
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<treatment id="03F687B7FFE19568FF56FF7AF73AFAD0" LSID="urn:lsid:plazi:treatment:03F687B7FFE19568FF56FF7AF73AFAD0" httpUri="http://treatment.plazi.org/id/03F687B7FFE19568FF56FF7AF73AFAD0" lastPageNumber="4" pageId="5" pageNumber="4">
<subSubSection id="C345652AFFE19568FF56FF7AF658FF68" box="[192,547,213,242]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BE036A1FFE19568FF56FF7AF658FF68" blockId="5.[192,557,213,275]" box="[192,547,213,242]" pageId="5" pageNumber="4">
<heading id="D0A881CDFFE19568FF56FF7AF658FF68" box="[192,547,213,242]" fontSize="12" level="2" pageId="5" pageNumber="4" reason="2">
<taxonomicName id="4C5F4D22FFE19568FF56FF7AF658FF68" ID-CoL="6PH8Y" authority="Howden" authorityName="Howden" authorityYear="2003" box="[192,547,213,242]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE19568FF56FF7AF658FF68" bold="true" box="[192,547,213,242]" pageId="5" pageNumber="4">
<emphasis id="B92BEAB3FFE19568FF56FF7AF5CBFF68" bold="true" box="[192,432,213,242]" italics="true" pageId="5" pageNumber="4">Leptorhyparus gilli</emphasis>
Howden
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C345652AFFE19568FF56FF56F656FE89" box="[192,557,249,275]" pageId="5" pageNumber="4" type="description">
<paragraph id="8BE036A1FFE19568FF56FF56F656FE89" blockId="5.[192,557,213,275]" box="[192,557,249,275]" pageId="5" pageNumber="4">
<figureCitation id="13642A24FFE19568FF56FF56F567FE89" box="[192,284,249,275]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="5" pageNumber="4">Figure 2</figureCitation>
(see also
<bibRefCitation id="EFCE4B50FFE19568FE11FF56F65EFE89" author="Howden HF" box="[391,549,249,275]" pageId="5" pageNumber="4" pagination="391 - 402" refId="ref4096" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden 2003</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE19568FF76FE8EF690FEA0" box="[224,747,289,314]" pageId="5" pageNumber="4" type="reference_group">
<paragraph id="8BE036A1FFE19568FF76FE8EF690FEA0" blockId="5.[224,747,289,314]" box="[224,747,289,314]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="AB4F118FFFE19568FF76FE8EF690FEA0" box="[224,747,289,314]" pageId="5" pageNumber="4">
<treatmentCitation id="0AFE10B0FFE19568FF76FE8EF6EBFEA0" author="Howden HF" box="[224,656,289,314]" page="401" pageId="5" pageNumber="4" year="2003">
<taxonomicName id="4C5F4D22FFE19568FF76FE8EF6EBFEA0" ID-CoL="6PH8Y" authority="Howden 2003: 400 - 401" authorityName="Howden" authorityPageNumber="400 - 401" authorityYear="2003" box="[224,656,289,314]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE19568FF76FE8EF5E1FEA3" box="[224,410,289,313]" italics="true" pageId="5" pageNumber="4">Leptorhyparus gilli</emphasis>
<bibRefCitation id="EFCE4B50FFE19568FE36FE8EF6EBFEA0" author="Howden HF" box="[416,656,289,314]" pageId="5" pageNumber="4" pagination="391 - 402" refId="ref4096" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden 2003: 400401</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 47.
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</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE19568FF56FEE6F688FDB7" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BE036A1FFE19568FF56FEE6F688FDB7" blockId="5.[191,1440,329,557]" pageId="5" pageNumber="4">
<emphasis id="B92BEAB3FFE19568FF56FEE6F547FEF9" bold="true" box="[192,316,329,355]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="4C5F4D22FFE19568FED7FEE6F675FEF9" authorityName="Howden" authorityYear="2003" box="[321,526,329,355]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE19568FED7FEE6F675FEF9" box="[321,526,329,355]" italics="true" pageId="5" pageNumber="4">Leptorhyparus gilli</emphasis>
</taxonomicName>
is readily distinguished from the other members of the genus by small body (length
<quantity id="4CA79B44FFE19568FF29FEC4F53CFE1F" box="[191,327,363,389]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="5" pageNumber="4" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.32.6 mm</quantity>
); head lacking distinct tufts of setae; sides of clypeus straight to genae; pronotum having a shortened groove in the posterior interval between the paramedian and discolateral costae that does not reach the anterior fovea, the area between the groove and fovea not raised allowing these costae to appear separated (
<figureCitation id="13642A24FFE19568FB68FE01F144FE52" box="[1278,1343,430,456]" captionStart="Figures 18" captionStartId="4.[192,279,1550,1576]" captionTargetBox="[192,1440,213,1520]" captionTargetId="figure-126@4.[192,1440,213,1520]" captionTargetPageId="4" captionText="Figures 18. Leptorhyparus characters. 13) Head and pronotum.1) L. brasiliensis. 2) L. gilli. 3) L. peruanus. 45) Elytra and pygidium, caudal view. 4) L. brasiliensis, 5) L. peruanus. 68) Body parts of L. brasiliensis, from French Guiana. 6) Epipharynx. 7) Male genitalia, caudal view. 8) Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12808373" httpUri="https://zenodo.org/record/12808373/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
); pronotum having a small anterior lobe of the discolateral costa; elytra with postdiscal bulbs reduced; elytra with caudal bulbs smaller and not distinctly visible from ventral view; median impression of metaventrite a narrow line entire length; and it occurs in southern Central America.
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE19568FF56FD93F723FCD8" pageId="5" pageNumber="4" type="description">
<paragraph id="8BE036A1FFE19568FF56FD93F723FCD8" blockId="5.[192,1442,572,834]" pageId="5" pageNumber="4">
<emphasis id="B92BEAB3FFE19568FF56FD93F54DFDCC" bold="true" box="[192,310,572,598]" pageId="5" pageNumber="4">Variation.</emphasis>
As
<bibRefCitation id="EFCE4B50FFE19568FEC8FD93F676FDCD" author="Howden HF" box="[350,525,572,599]" pageId="5" pageNumber="4" pagination="391 - 402" refId="ref4096" refString="Howden HF. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42 (2): 391 - 402." type="journal article" year="2003">Howden (2003)</bibRefCitation>
discussed for
<taxonomicName id="4C5F4D22FFE19568FD3CFD92F695FDCC" authorityName="Howden" authorityYear="2003" box="[682,750,572,599]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B92BEAB3FFE19568FD3CFD92F6C2FDCC" box="[682,697,573,598]" italics="true" pageId="5" pageNumber="4">L</emphasis>
.
<emphasis id="B92BEAB3FFE19568FD53FD93F695FDCC" box="[709,750,572,598]" italics="true" pageId="5" pageNumber="4">gilli</emphasis>
</taxonomicName>
and
<bibRefCitation id="EFCE4B50FFE19568FCB2FD93F7AFFDCD" author="Minkina, L." box="[804,980,572,599]" pageId="5" pageNumber="4" pagination="34 - 39" refId="ref4351" refString="Minkina, L. 2020. Two new species of Leptorhyparus Howden, 2003 from South America. Folia Heyrovskyana series A 28 (2): 34 - 39." type="journal article" year="2020">Minkina (2020)</bibRefCitation>
suspected for
<taxonomicName id="4C5F4D22FFE19568FBE4FD92F17DFDCC" authorityName="Minkina" authorityYear="2020" box="[1138,1286,572,599]" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B92BEAB3FFE19568FBE4FD92F0FAFDCC" box="[1138,1153,573,598]" italics="true" pageId="5" pageNumber="4">L</emphasis>
.
<emphasis id="B92BEAB3FFE19568FB18FD93F17DFDCC" box="[1166,1286,572,598]" italics="true" pageId="5" pageNumber="4">brasiliensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D22FFE19568FAADFD92F49CFDE2" authorityName="Minkina" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Leptorhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="peruanus">
<emphasis id="B92BEAB3FFE19568FAADFD92F131FDCC" box="[1339,1354,573,598]" italics="true" pageId="5" pageNumber="4">L</emphasis>
.
<emphasis id="B92BEAB3FFE19568FAC1FD92F49CFDE2" italics="true" pageId="5" pageNumber="4">peruanus</emphasis>
</taxonomicName>
, there are variation in details related to the development of the paramedian and discolateral costae of the pronotum, along with the area between them, puncture rows with punctures varying in size and distance, elytral intervals appearing flat or convex depending on the individual, and general size and distinctness of dorsal and ventral punctation. Some of the differences are actual variation between individuals. In others it appears to relate to the visual quality of the structures relating to dirt remnants or angle of lighting. Caution needs to be taken to recognize species differences that hold for series of specimens and not individuals. New species descriptions based on subtle differences of singletons should be avoided.
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE19568FF56FCFEF675FB21" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BE036A1FFE19568FF56FCFEF675FB21" blockId="5.[191,1442,849,1212]" pageId="5" pageNumber="4">
<emphasis id="B92BEAB3FFE19568FF56FCFEF5DAFCF1" bold="true" box="[192,417,849,875]" pageId="5" pageNumber="4">Material examined.</emphasis>
(
<materialsCitation id="3B373CFCFFE19568FE3BFCFEF675FB21" collectingDate="1977-08-28" collectingDateMax="2012-08-20" collectingDateMin="1977-08-28" collectionCode="CSCA, FSCA, CMNC, SEMC, CEMT" collectorName="M. Ferro &amp; J. S. Ashe &amp; Z. Falin &amp; I. Hinojosa &amp; R. S. Anderson &amp; H. Wolda &amp; Rio Indo Lodge, N. &amp; J. Heppner &amp; Santa Fe &amp; Alto Piedra &amp; J. B. Heppner" country="Costa Rica" county="Panamanian" elevation="1330" latitude="8.6629715" location="Estacion Biologica" longLatPrecision="1" longitude="-80.118866" municipality="Estacion Biologica LaSelva" pageId="5" pageNumber="4" specimenCode="SEMC1113155" specimenCount="2" stateProvince="Chiriqui" typeStatus="holotype">
20)
<typeStatus id="54E48803FFE19568FE4EFCFEF63BFCF1" box="[472,576,849,875]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
and 3
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<typeStatus id="54E48803FFE19568FC8EFCFDF7FFFCF6" box="[792,900,850,876]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
in the CMNC plus the following:
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<collectingCountry id="F3487631FFE19568FB63FCFEF1E1FCF1" box="[1269,1434,849,875]" name="Costa Rica" pageId="5" pageNumber="4">COSTA RICA</collectingCountry>
: Heredia
</emphasis>
:
<collectingMunicipality id="6B84ACDBFFE19568FEBDFCDCF62CFC17" box="[299,599,883,909]" pageId="5" pageNumber="4">
<location id="8E80607AFFE19568FEBDFCDCF582FC17" LSID="urn:lsid:plazi:treatment:03F687B7FFE19568FF56FF7AF73AFAD0:8E80607AFFE19568FEBDFCDCF582FC17" box="[299,505,883,909]" country="Costa Rica" county="Panamanian" latitude="8.6629715" longLatPrecision="1" longitude="-80.118866" municipality="Estacion Biologica LaSelva" name="Estacion Biologica" pageId="5" pageNumber="4" stateProvince="Chiriqui">Estación Biológica</location>
LaSelva
</collectingMunicipality>
,
<geoCoordinate id="EE6B5066FFE19568FDF4FCDCF6C5FC17" box="[610,702,883,911]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="5" pageNumber="4" precision="925" value="10.433333">10°26N</geoCoordinate>
,
<geoCoordinate id="EE6B5066FFE19568FD5FFCDCF753FC17" box="[713,808,883,911]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="5" pageNumber="4" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<collectingMethod id="521E4EB6FFE19568FCA4FCDCF070FC17" box="[818,1035,883,909]" pageId="5" pageNumber="4">flight intercept trap</collectingMethod>
#5,
<date id="FFE11061FFE19568FBAFFCDCF0C2FC14" box="[1081,1209,883,910]" pageId="5" pageNumber="4" value="2005-07-08">
<collectingDate id="EFA5E989FFE19568FBAFFCDCF0C2FC14" box="[1081,1209,883,910]" pageId="5" pageNumber="4" value="2005-07-08">8-VII-2005</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FB55FCDCF15DFC17" box="[1219,1318,883,909]" pageId="5" pageNumber="4">M. Ferro</collectorName>
(1
<collectionCode id="ED4EAE64FFE19568FADCFCDCF1EFFC17" box="[1354,1428,883,909]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="5" pageNumber="4" type="Museum">CSCA</collectionCode>
);
<emphasis id="B92BEAB3FFE19568FF56FC3AF536FC35" bold="true" box="[192,333,917,943]" pageId="5" pageNumber="4">Puntarenas:</emphasis>
<location id="8E80607AFFE19568FEC3FC3BF6CBFC35" LSID="urn:lsid:plazi:treatment:03F687B7FFE19568FF56FF7AF73AFAD0:8E80607AFFE19568FEC3FC3BF6CBFC35" box="[341,688,916,943]" country="Costa Rica" county="Panamanian" latitude="8.6629715" longLatPrecision="1" longitude="-80.118866" municipality="Estacion Biologica LaSelva" name="Estacion Biologica Las Cruces" pageId="5" pageNumber="4" stateProvince="Chiriqui">Estación Biológica Las Cruces</location>
,
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</quantity>
,
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,
<geoCoordinate id="EE6B5066FFE19568FC39FC3AF04BFC35" box="[943,1072,916,945]" degrees="82" direction="west" minutes="57.58" orientation="longitude" pageId="5" pageNumber="4" precision="9" value="-82.95966">82°57.58W</geoCoordinate>
,
<date id="FFE11061FFE19568FBABFC3AF0CDFC35" box="[1085,1206,917,943]" pageId="5" pageNumber="4" value="2004-05-30">
<collectingDate id="EFA5E989FFE19568FBABFC3AF0CDFC35" box="[1085,1206,917,943]" pageId="5" pageNumber="4" value="2004-05-30">30-V-2004</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FB52FC3AF148FC34" box="[1220,1331,916,943]" pageId="5" pageNumber="4">J. S. Ashe</collectorName>
,
<collectorName id="26AA5377FFE19568FAD6FC3AF1E7FC34" box="[1344,1436,916,943]" pageId="5" pageNumber="4">Z. Falin</collectorName>
,
<collectorName id="26AA5377FFE19568FF56FC18F53BFC4A" box="[192,320,950,977]" pageId="5" pageNumber="4">I. Hinojosa</collectorName>
, at uv light, CR1AFH04 058, (2
<collectionCode id="ED4EAE64FFE19568FD26FC19F68EFC4A" box="[688,757,950,976]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
: barcode KUNHM-ENT SM060993, SM0609981);
<locationDeviation id="9A522F83FFE19568FAA3FC19F668FC68" location="Estacion Biologica Las Cruces" pageId="5" pageNumber="4">
<quantity id="4CA79B44FFE19568FAA3FC19F10BFC4A" box="[1333,1392,950,977]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="4" unit="km" value="5.0">5 km</quantity>
SW Estación Biológica Las Cruces
</locationDeviation>
,
<quantity id="4CA79B44FFE19568FD8BFC77F608FC68" box="[541,627,984,1010]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="4" unit="m" value="1400.0">
<elevation id="0072D192FFE19568FD8BFC77F608FC68" box="[541,627,984,1010]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<geoCoordinate id="EE6B5066FFE19568FDEBFC77F68AFC68" box="[637,753,984,1012]" degrees="8" direction="north" minutes="47" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="13" value="8.786945">8°4713″N</geoCoordinate>
,
<geoCoordinate id="EE6B5066FFE19568FD6AFC78F7F9FC68" box="[764,898,983,1012]" degrees="82" direction="west" minutes="59" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="13" value="-82.98694">82°5913″W</geoCoordinate>
,
<date id="FFE11061FFE19568FC1DFC77F074FC68" box="[907,1039,984,1010]" pageId="5" pageNumber="4" value="1998-06-22">
<collectingDate id="EFA5E989FFE19568FC1DFC77F074FC68" box="[907,1039,984,1010]" pageId="5" pageNumber="4" value="1998-06-22">22-VI-1998</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FB8CFC77F0C6FC68" box="[1050,1213,984,1010]" pageId="5" pageNumber="4">R.S. Anderson</collectorName>
, wet cloud forest litter (3
<collectionCode id="ED4EAE64FFE19568FE90FC56F525FB89" box="[262,350,1017,1043]" pageId="5" pageNumber="4">CMNC</collectionCode>
).
<emphasis id="B92BEAB3FFE19568FEE5FC56F6F4FB89" bold="true" box="[371,655,1017,1043]" pageId="5" pageNumber="4">
<collectingCountry id="F3487631FFE19568FEE5FC56F660FB89" box="[371,539,1017,1043]" name="Nicaragua" pageId="5" pageNumber="4">NICARAGUA</collectingCountry>
: Jinotega:
</emphasis>
Parque Nacional Cero Saslaya,
<geoCoordinate id="EE6B5066FFE19568FC61FC56F020FB8E" box="[1015,1115,1017,1044]" degrees="13.77165" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="13.77165">13.77165</geoCoordinate>
<geoCoordinate id="EE6B5066FFE19568FBF2FC56F0A0FB8E" box="[1124,1243,1017,1044]" degrees="85.01184" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.01184">85.01184</geoCoordinate>
, ±
<elevation id="0072D192FFE19568FB60FC56F1E6FB8E" box="[1270,1437,1017,1044]" isEnumeration="true" pageId="5" pageNumber="4">
<quantity id="4CA79B44FFE19568FB60FC56F145FB8E" box="[1270,1342,1017,1044]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="4" unit="m" value="150.0">150 m</quantity>
<quantity id="4CA79B44FFE19568FAD1FC56F1E6FB8E" box="[1351,1437,1017,1044]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="5" pageNumber="4" unit="m" value="1110.0">1110 m</quantity>
</elevation>
,
<date id="FFE11061FFE19568FF54FBB4F540FBAF" box="[194,315,1051,1077]" pageId="5" pageNumber="4" value="2011-05-12">
<collectingDate id="EFA5E989FFE19568FF54FBB4F540FBAF" box="[194,315,1051,1077]" pageId="5" pageNumber="4" value="2011-05-12">12-V-2011</collectingDate>
</date>
, ex sifted leaf litter, ridgetop cloud forest, LLAMA11 Wa-D-03-1-all (1
<collectionCode id="ED4EAE64FFE19568FBD3FBB4F0EBFBAF" box="[1093,1168,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="4" type="Museum">SEMC</collectionCode>
: barcode KUNHM-ENT
<specimenCode id="DBF99EDAFFE19568FF29FB93F510FBCD" box="[191,363,1084,1111]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="4" type="Museum">SEMC1113155</specimenCode>
).
<emphasis id="B92BEAB3FFE19568FE16FB93F609FBCC" bold="true" box="[384,626,1084,1110]" pageId="5" pageNumber="4">
<collectingCountry id="F3487631FFE19568FE16FB93F583FBCC" box="[384,504,1084,1110]" name="Panama" pageId="5" pageNumber="4">PANAMA</collectingCountry>
:
<collectingRegion id="499BF843FFE19568FD90FB93F616FBCC" box="[518,621,1084,1110]" country="Panama" name="Chiriqui" pageId="5" pageNumber="4">Chiriqui</collectingRegion>
:
</emphasis>
Fortuna,
<quantity id="4CA79B44FFE19568FD73FB93F746FBCD" box="[741,829,1084,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="4" unit="m" value="1050.0">
<elevation id="0072D192FFE19568FD73FB93F746FBCD" box="[741,829,1084,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="4" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<geoCoordinate id="EE6B5066FFE19568FCDDFB92F7E3FBCD" box="[843,920,1084,1113]" degrees="8" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="4" precision="925" value="8.75">8°45N</geoCoordinate>
<geoCoordinate id="EE6B5066FFE19568FC34FB92F074FBCD" box="[930,1039,1084,1113]" degrees="082" direction="west" minutes="15" orientation="longitude" pageId="5" pageNumber="4" precision="925" value="-82.25">082°15W</geoCoordinate>
,
<date id="FFE11061FFE19568FB8AFB92F0E5FBCD" box="[1052,1182,1085,1111]" pageId="5" pageNumber="4" value="1978-09-19">
<collectingDate id="EFA5E989FFE19568FB8AFB92F0E5FBCD" box="[1052,1182,1085,1111]" pageId="5" pageNumber="4" value="1978-09-19">19-IX-1978</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FB3AFB92F166FBCC" box="[1196,1309,1084,1111]" pageId="5" pageNumber="4">H. Wolda</collectorName>
, Lt. trap (2
<collectionCode id="ED4EAE64FFE19568FF56FBF1F57CFBE2" box="[192,263,1118,1144]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
); Fortuna,
<date id="FFE11061FFE19568FEE9FBF1F66DFBE2" box="[383,534,1118,1144]" pageId="5" pageNumber="4" value="1977-08-28">
<collectingDate id="EFA5E989FFE19568FEE9FBF1F66DFBE2" box="[383,534,1118,1144]" pageId="5" pageNumber="4" value="1977-08-28">28-VIII-1977</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FDB7FBF0F6F6FBE2" box="[545,653,1118,1145]" pageId="5" pageNumber="4">H. Wolda</collectorName>
(1
<collectionCode id="ED4EAE64FFE19568FD27FBF1F686FBE2" box="[689,765,1118,1144]" pageId="5" pageNumber="4">CEMT</collectionCode>
, 4
<collectionCode id="ED4EAE64FFE19568FC8AFBF1F718FBE2" box="[796,867,1118,1144]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
);
<emphasis id="B92BEAB3FFE19568FCE3FBF1F7C4FBE2" bold="true" box="[885,959,1118,1144]" pageId="5" pageNumber="4">
<collectingRegion id="499BF843FFE19568FCE3FBF1F7C1FBE2" box="[885,954,1118,1144]" country="Panama" name="Cocle" pageId="5" pageNumber="4">Cocle</collectingRegion>
:
</emphasis>
<collectorName id="26AA5377FFE19568FC53FBF1F0E7FBE3" box="[965,1180,1118,1145]" pageId="5" pageNumber="4">
<location id="8E80607AFFE19568FC53FBF1F00CFBE2" LSID="urn:lsid:plazi:treatment:03F687B7FFE19568FF56FF7AF73AFAD0:8E80607AFFE19568FC53FBF1F00CFBE2" box="[965,1143,1118,1144]" country="Costa Rica" county="Panamanian" latitude="8.6629715" longLatPrecision="1" longitude="-80.118866" municipality="Estacion Biologica LaSelva" name="Rio Indo Lodge" pageId="5" pageNumber="4" stateProvince="Chiriqui">Rio Indo Lodge</location>
, N.
</collectorName>
El Valle,
<geoCoordinate id="EE6B5066FFE19568FA93FBF1F1E7FBE3" box="[1285,1436,1118,1146]" degrees="08" direction="north" minutes="39" orientation="latitude" pageId="5" pageNumber="4" precision="1" seconds="46.7" value="8.6629715">08°3946.7″N</geoCoordinate>
,
<geoCoordinate id="EE6B5066FFE19568FF56FB2FF521FB00" box="[192,346,1152,1180]" degrees="80" direction="west" minutes="07" orientation="longitude" pageId="5" pageNumber="4" precision="1" seconds="07.9" value="-80.118866">80°0707.9″W</geoCoordinate>
,
<quantity id="4CA79B44FFE19568FEF5FBD0F5D1FB00" box="[355,426,1151,1178]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="5" pageNumber="4" unit="m" value="575.0">
<elevation id="0072D192FFE19568FEF5FBD0F5D1FB00" box="[355,426,1151,1178]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="5" pageNumber="4" unit="m" value="575.0">575 m</elevation>
</quantity>
,
<date id="FFE11061FFE19568FE22FB2FF60FFB00" box="[436,628,1152,1178]" pageId="5" pageNumber="4" value="2012-08-18" valueMax="2012-08-20" valueMin="2012-08-18">
<collectingDate id="EFA5E989FFE19568FE22FB2FF60FFB00" box="[436,628,1152,1178]" pageId="5" pageNumber="4" value="2012-08-18" valueMax="2012-08-20" valueMin="2012-08-18">1820-VIII-2012</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FDE8FB2FF68EFB00" box="[638,757,1152,1178]" pageId="5" pageNumber="4">J. Heppner</collectorName>
(1
<collectionCode id="ED4EAE64FFE19568FC81FB2FF725FB00" box="[791,862,1152,1178]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
);
<emphasis id="B92BEAB3FFE19568FCF8FB2FF79AFB00" bold="true" box="[878,993,1152,1178]" pageId="5" pageNumber="4">
<collectingRegion id="499BF843FFE19568FCF8FB2FF7A7FB00" box="[878,988,1152,1178]" country="Panama" name="Veraguas" pageId="5" pageNumber="4">Veraguas</collectingRegion>
:
</emphasis>
<collectorName id="26AA5377FFE19568FC70FB2FF03EFB00" box="[998,1093,1152,1178]" pageId="5" pageNumber="4">Santa Fe</collectorName>
,
<collectorName id="26AA5377FFE19568FBD9FB2FF0ABFB00" box="[1103,1232,1152,1178]" pageId="5" pageNumber="4">Alto Piedra</collectorName>
,
<quantity id="4CA79B44FFE19568FB4CFBD0F15AFB00" box="[1242,1313,1151,1178]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="4" unit="m" value="850.0">
<elevation id="0072D192FFE19568FB4CFBD0F15AFB00" box="[1242,1313,1151,1178]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="4" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="FFE11061FFE19568FABCFB2FF483FB21" pageId="5" pageNumber="4" value="2011-06-16" valueMax="2011-06-18" valueMin="2011-06-16">
<collectingDate id="EFA5E989FFE19568FABCFB2FF483FB21" pageId="5" pageNumber="4" value="2011-06-16" valueMax="2011-06-18" valueMin="2011-06-16">1618-VI-2011</collectingDate>
</date>
,
<collectorName id="26AA5377FFE19568FE95FB0DF5ECFB26" box="[259,407,1186,1212]" pageId="5" pageNumber="4">J. B. Heppner</collectorName>
(1
<collectionCode id="ED4EAE64FFE19568FE2DFB0EF679FB21" box="[443,514,1185,1211]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345652AFFE19568FF56FB64F73AFAD0" pageId="5" pageNumber="4" type="discussion">
<paragraph id="8BE036A1FFE19568FF56FB64F73AFAD0" blockId="5.[192,1442,1227,1354]" pageId="5" pageNumber="4">
<emphasis id="B92BEAB3FFE19568FF56FB64F533FB7F" bold="true" box="[192,328,1227,1253]" pageId="5" pageNumber="4">Comments.</emphasis>
There is some variation in the distance between the end of the pronotal groove and the anterior foveae. Lighting must be correct to fully illuminate these structures. Sometimes the interval is more depressed making it appear as if the groove is longer. However, the groove at the bottom of this depressed interval is deep, sharply defined and always well separated from the fovea.
</paragraph>
</subSubSection>
</treatment>
</document>