722 lines
90 KiB
XML
722 lines
90 KiB
XML
<document id="3C885F4E0ABC0B294255F8968C143783" ID-DOI="10.1093/zoolinnean/zlae054" ID-ISSN="0024-4082" ID-Zenodo-Dep="14764381" ID-ZooBank="49071C9-46D4-42D0-BE6E-6111110C0B09" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1738156038381" checkinUser="plazi" docAuthor="Pereira, Paulo V L G C, Bandeira, Kamila L N, Vidal, Luciano S, Ribeiro, Theo B, Candeiro, Carlos R dos A & Bergqvist, Lilian P" docDate="2024" docId="03D187F7FFE1FFBF5F6134EFFA9D4E39" docLanguage="en" docName="zlae054.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (4)" docSource="https://doi.org/10.1093/zoolinnean/zlae054" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Tiamat valdecii Pereira, Bandeira, Vidal, Ribeiro, Candeiro & Bergqvist, 2024, sp.nov." docType="treatment" docVersion="1" lastPageNumber="7" masterDocId="FFE8FF8FFFE5FFB95E2E353FFFD4486B" masterDocTitle="A new sauropod species from north-western Brazil: biomechanics and the radiation of Titanosauria (Sauropoda: Somphospondyli)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="5" updateTime="1738781995253" updateUser="guilherme">
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<mods:title id="7D45300918878992D8E606889D5540BA">A new sauropod species from north-western Brazil: biomechanics and the radiation of Titanosauria (Sauropoda: Somphospondyli)</mods:title>
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<mods:namePart id="401FC06404CA9D8A42FAB1CD8D8EF36B">Pereira, Paulo V L G C</mods:namePart>
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<mods:namePart id="BAB9C3965A99599F1BD1BAB733850CEF">Bandeira, Kamila L N</mods:namePart>
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<mods:namePart id="971AB95365AE65B5517D519A43150102">Vidal, Luciano S</mods:namePart>
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<mods:namePart id="16B08DF45C02E269B5ADF70872364682">Ribeiro, Theo B</mods:namePart>
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<mods:namePart id="89871354E201D1C47D7860DAC6D65945">Candeiro, Carlos R dos A</mods:namePart>
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<mods:namePart id="A26B92BC33C6C0F06E24082FD9B20CE4">Bergqvist, Lilian P</mods:namePart>
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<mods:title id="2812F7BB735832E636F6E06E8E35B094">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="0BA04CB157B618ADC5FD06BE7C441604">2024</mods:date>
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<mods:number id="834D140D9BDE4C2277F7B748BF4C83F3">2024-05-13</mods:number>
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<mods:number id="1C3BFBD8423DD7861124EDF8D406A1F9">202</mods:number>
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<mods:identifier id="99855051E75537F919A1755641F44D38" type="DOI">10.1093/zoolinnean/zlae054</mods:identifier>
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<mods:identifier id="033CB0721395989DE28D62F045FB1FE4" type="ISSN">0024-4082</mods:identifier>
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<subSubSection id="C362656AFFE1FFBD5F6134EFFDEF4980" box="[335,571,464,491]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8BC736E1FFE1FFBD5F6134EFFDEF4980" blockId="4.[129,778,464,558]" box="[335,571,464,491]" pageId="4" pageNumber="5">
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<emphasis id="B90CEAF3FFE1FFBD5F6134EFFDEF4980" bold="true" box="[335,571,464,491]" pageId="4" pageNumber="5">
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<taxonomicName id="4C784D62FFE1FFBD5F6134EFFE3E4980" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[335,490,464,491]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="valdecii" status="sp. nov.">
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<emphasis id="B90CEAF3FFE1FFBD5F6134EFFE3E4980" bold="true" box="[335,490,464,491]" italics="true" pageId="4" pageNumber="5">Tiamat valdecii</emphasis>
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<taxonomicNameLabel id="A23F5788FFE1FFBD5FC134EEFDEF4980" box="[495,571,465,491]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C362656AFFE1FFBD5EAF34C8FDB54A45" pageId="4" pageNumber="5" type="description">
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<paragraph id="8BC736E1FFE1FFBD5EAF34C8FDB54A45" blockId="4.[129,778,464,558]" pageId="4" pageNumber="5">
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ZooBank LSID for the species:
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<uri id="FFE93AE3FFE1FFBD5FDB34C8FDB54A45" pageId="4" pageNumber="5">
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urn: lsid:zoobank.org:act:
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<uuid id="FFDE0C34FFE1FFBD5EAF3729FDB54A45" box="[129,609,534,558]" pageId="4" pageNumber="5">549071C9-46D4-42D0-BE6E-6111110C0B09</uuid>
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</uri>
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</paragraph>
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<subSubSection id="C362656AFFE1FFBD5EAF376FFE624AAD" pageId="4" pageNumber="5" type="etymology">
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<paragraph id="8BC736E1FFE1FFBD5EAF376FFE624AAD" blockId="4.[128,778,592,710]" pageId="4" pageNumber="5">
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<emphasis id="B90CEAF3FFE1FFBD5EAF376FFF244A03" box="[129,240,592,616]" italics="true" pageId="4" pageNumber="5">Etymology:</emphasis>
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The specific epithet is in honour of Prof. Dr Valdeci dos Santos Júnior of Universidade Estadual do
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<collectingRegion id="49BCF803FFE1FFBD5C403750FF144ACD" country="Brazil" name="Rio Grande do Norte" pageId="4" pageNumber="5">Rio Grande do Norte</collectingRegion>
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, for the discovery of the fossil site and for his fundamental support during the fieldworks.
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</paragraph>
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</subSubSection>
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<subSubSection id="C362656AFFE1FFBD5EAF37D8FD514B36" pageId="4" pageNumber="5" type="materials_examined">
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<paragraph id="8BC736E1FFE1FFBD5EAF37D8FD514B36" blockId="4.[128,777,743,861]" pageId="4" pageNumber="5">
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<materialsCitation id="3B103CBCFFE1FFBD5EAF37D8FDFB4B36" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
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<emphasis id="B90CEAF3FFE1FFBD5EAF37D8FF354A94" box="[129,225,743,767]" italics="true" pageId="4" pageNumber="5">
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<typeStatus id="54C38843FFE1FFBD5EAF37D8FF084A94" box="[129,220,743,767]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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UFRJ-DG 636-R, UFRJ-DG 638-R, UFRJ-DG 606-R (anterior caudal vertebrae), UFRJ-DG 635-R, UFRJ-DG 591-R, UFRJ-DG 527-R, UFRJ-DG 704-R (middle caudal vertebrae), and UFRJ-DG 574-R (partial neural arch)
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</materialsCitation>
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||
(
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<figureCitation id="13432A64FFE1FFBD5C13367AFDA14B36" box="[573,629,837,861]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C362656AFFE1FFBD5EAF3640FE2E4D07" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="8BC736E1FFE1FFBD5EAF3640FE2E4D07" blockId="4.[128,779,894,1388]" pageId="4" pageNumber="5">
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||
<emphasis id="B90CEAF3FFE1FFBD5EAF3640FF334BFD" box="[129,231,895,918]" italics="true" pageId="4" pageNumber="5">Diagnosis:</emphasis>
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A titanosaur sauropod diagnosed by four diagnostic features, all in the caudal vertebrae: marked accessory tuberosity dorsoventrally developed, located above the prezygapophyses in the posteriormost and in middle caudal vertebrae; deeply medioventral excavated articulation facets of the prezygapophysis and post-zygapophysis (see fig.07); presence of hypanthrum–hyposphene articulation; and short middle caudal centra with well-marked articular facets for the haemal arch. Additionally, some of the following combinations of characters assist the placement of
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<taxonomicName id="4C784D62FFE1FFBD5F4431A6FDD54CDB" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[362,513,1176,1200]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="valdecii">
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<emphasis id="B90CEAF3FFE1FFBD5F4431A6FDD54CDB" box="[362,513,1176,1200]" italics="true" pageId="4" pageNumber="5">Tiamat valdecii</emphasis>
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</taxonomicName>
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as a titanosaurian, such as strong procoelous anterior caudal vertebrae with the apex of the posterior condyle positioned on the dorsal half of the centrum; wider than high centra which, progressively widen to the posterior vertebrae; pedicles of the neural arch attached to the anterior half of the centrum and presence of a ventral longitudinal hollow on the caudal vertebral series.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C362656AFFE1FFBF5FA130B2FA9D4E39" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
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||
<paragraph id="8BC736E1FFE1FFBD5FA130B2FE2F4DCD" blockId="4.[127,780,1421,1984]" box="[399,507,1421,1446]" pageId="4" pageNumber="5">
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<emphasis id="B90CEAF3FFE1FFBD5FA130B2FE2F4DCD" box="[399,507,1421,1446]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
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||
</paragraph>
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<paragraph id="8BC736E1FFE1FFBD5EAF308DFB3748AC" blockId="4.[127,780,1421,1984]" lastBlockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
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||
There are three anterior caudal vertebrae, four middle caudal vertebrae, and a neural spine (possibly an anterior one). The anterior caudal vertebrae (UFRJ-DG 636-R, UFRJ-DG 638-R, UFRJ-DG 606-R) are strongly procoelous, with subcircular outline, while one of the middle caudal vertebrae (UFRJ-DG 635-R) is slightly procoelous, and the other middle caudal (UFRJ-DG 591-R, UFRJ-DG 527-R, UFRJ-DG 704-R) are amphicoelous. This condition (transition of procoelous to slightly procoelous) is observed in
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<taxonomicName id="4C784D62FFE1FFBD5F353392FE614EAE" box="[283,437,1709,1733]" class="Reptilia" family="Titanosauridae" genus="Mendozasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
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<emphasis id="B90CEAF3FFE1FFBD5F353392FE614EAE" box="[283,437,1709,1733]" italics="true" pageId="4" pageNumber="5">Mendozasaurus</emphasis>
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||
</taxonomicName>
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||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5FE63392FD4F4EAE" author="Gonzalez-Riga BJ" box="[456,667,1709,1733]" pageId="4" pageNumber="5" pagination="155 - 72" refId="ref12321" refString="Gonzalez-Riga BJ. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza Province, Argentina. Ameghiniana 2003; 40: 155 - 72." year="2003">González-Riga 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5C873392FEF04E8F" author="Gonzalez Riga BJ & Mannion PD & Poropat SF" pageId="4" pageNumber="5" pagination="136 - 81" refId="ref12412" refString="Gonzalez Riga BJ, Mannion PD, Poropat SF et al. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 2018; 184: 136 - 81. https: // doi. org / 10.1093 / zoolinnean / zlx 103" year="2018">
|
||
Gonzàlez Riga
|
||
<emphasis id="B90CEAF3FFE1FFBD5E9933F2FF3D4E8F" box="[183,233,1740,1764]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C784D62FFE1FFBD5F4833F2FD0C4E8F" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" box="[358,728,1740,1764]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5F4833F2FE034E8F" box="[358,471,1741,1764]" italics="true" pageId="4" pageNumber="5">Patagotitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5FC733F3FD184E8F" author="Carballido JL & Pol D & Otero A" box="[489,716,1740,1764]" pageId="4" pageNumber="5" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
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||
Carballido
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||
<emphasis id="B90CEAF3FFE1FFBD5C7033F2FD5E4E8F" box="[606,650,1740,1764]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
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||
; but differs from other taxa such as
|
||
<taxonomicName id="4C784D62FFE1FFBD5F9433D3FF144F48" authority="(Mannion and Calvo 2011)" baseAuthorityName="Mannion and Calvo" baseAuthorityYear="2011" class="Reptilia" family="Titanosauridae" genus="Andesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
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||
<emphasis id="B90CEAF3FFE1FFBD5F9433D3FDF84F6F" box="[442,556,1772,1796]" italics="true" pageId="4" pageNumber="5">Andesaurus</emphasis>
|
||
(
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||
<bibRefCitation id="EFE94B10FFE1FFBD5C1333D3FF604F48" author="Mannion PD & Calvo JO" pageId="4" pageNumber="5" refId="ref13041" refString="Mannion PD, Calvo JO. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albianearly Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 2011; 163: no - no. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00699. x" year="2011">Mannion and Calvo 2011</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
,
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<taxonomicName id="4C784D62FFE1FFBD5EE33234FE834F48" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[205,343,1803,1827]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
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<emphasis id="B90CEAF3FFE1FFBD5EE33234FE834F48" box="[205,343,1803,1827]" italics="true" pageId="4" pageNumber="5">Malawisaurus</emphasis>
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||
</taxonomicName>
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, and
|
||
<taxonomicName id="4C784D62FFE1FFBD5FBC3234FDD14F48" box="[402,517,1803,1827]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5FBC3234FDD14F48" box="[402,517,1803,1827]" italics="true" pageId="4" pageNumber="5">Rukwatitan</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5C393234FD734F48" author="Gomani EM" box="[535,679,1803,1827]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5C9A3234FF384F29" author="Gorscak E & O'Connor PM & Stevens NJ" pageId="4" pageNumber="5" pagination="1133 - 54" refId="ref12567" refString="Gorscak E, O'Connor PM, Stevens NJ et al. The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the Middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 2014; 34: 1133 - 54. https: // doi. org / 10.1080 / 02724634.2014.845568" year="2014">
|
||
Gorscak
|
||
<emphasis id="B90CEAF3FFE1FFBD5EAF3214FF7F4F29" box="[129,171,1834,1858]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), since in these taxa, the anteriormost caudal vertebrae exhibits strongly procoelous condition, while the other anterior caudal vertebrae and, consequently the middle ones, turn to an amphiplatyan condition. The most anterior caudal vertebra recovered for the new Brazilian taxa also differs from
|
||
<taxonomicName id="4C784D62FFE1FFBD5D1735AFFC3848C3" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[825,1004,144,168]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5D1735AFFC3848C3" box="[825,1004,144,168]" italics="true" pageId="4" pageNumber="5">Mnyamawamtuka</emphasis>
|
||
</taxonomicName>
|
||
, since the African taxon presents a ‘mildprocoely’ (
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5D873590FB0648AC" author="Gorscak E & O'Connor PM" box="[937,1234,175,199]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE1FFBD5D7B35F0FA16498A" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||
In lateral view, all vertebrae centra are anteroposteriorly short, bearing a concave lateral surface, which, in turn, is developed in the anteroposterior axis. The preserved neural arches are small and placed slightly anterior to the midpoint of the vertebral body. The vertebrae vary in height and width when observed on anterior and posterior views (
|
||
<tableCitation id="C6FA035AFFE1FFBD5A203454FB8D49E8" box="[1038,1113,363,387]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||
). The centrum of the more anterior caudal vertebra (UFRJ-DG 636-R) is taller than wide (
|
||
<tableCitation id="C6FA035AFFE1FFBD5B4934B4FA6649C8" box="[1383,1458,395,419]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||
), while it progressively dorsoventrally compresses in the middle caudal preserved (e.g.UFRJ-DG 704-R) (
|
||
<figureCitation id="13432A64FFE1FFBD5AFF34F6FACF498A" box="[1233,1307,457,481]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4A</figureCitation>
|
||
<figureCitation id="13432A64FFE1FFBD5B3534F6FA954989" box="[1307,1345,457,482]" captionStart="Figure 1" captionStartId="1.[114,179,1926,1950]" captionTargetBox="[117,1459,1429,1896]" captionTargetId="figure-639@1.[114,1461,1424,1898]" captionTargetPageId="1" captionText="Figure 1. Representation of the recovery process of the middle caudal vertebrae UFRJ-DG 591-R. A, absence of the left prezygapophysis in the vertebra; B, using the right prezygapophysis present in the UFRJ-DG 591-R, it was possible to infer the left prezygapophysis." figureDoi="http://doi.org/10.5281/zenodo.14764385" httpUri="https://zenodo.org/record/14764385/files/figure.png" pageId="4" pageNumber="5">1–A</figureCitation>
|
||
<figureCitation id="13432A64FFE1FFBD5B6F34F6FA9A498A" box="[1345,1358,457,481]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
|
||
,
|
||
<figureCitation id="13432A64FFE1FFBD5B7634F6FAAC498A" box="[1368,1400,457,481]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">4G</figureCitation>
|
||
<figureCitation id="13432A64FFE1FFBD5B5634F6FA71498A" box="[1400,1445,457,481]" captionStart="Figure 1" captionStartId="1.[114,179,1926,1950]" captionTargetBox="[117,1459,1429,1896]" captionTargetId="figure-639@1.[114,1461,1424,1898]" captionTargetPageId="1" captionText="Figure 1. Representation of the recovery process of the middle caudal vertebrae UFRJ-DG 591-R. A, absence of the left prezygapophysis in the vertebra; B, using the right prezygapophysis present in the UFRJ-DG 591-R, it was possible to infer the left prezygapophysis." figureDoi="http://doi.org/10.5281/zenodo.14764385" httpUri="https://zenodo.org/record/14764385/files/figure.png" pageId="4" pageNumber="5">1–G</figureCitation>
|
||
<figureCitation id="13432A64FFE1FFBD5B8B34F6FA66498A" box="[1445,1458,457,481]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE1FFBD5D7B34D6FC654AB7" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||
As in
|
||
<emphasis id="B90CEAF3FFE1FFBD5D8E34D6FBEA4A6B" box="[928,1086,489,512]" italics="true" pageId="4" pageNumber="5">Tastavinasaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5A7934D6FAE54A6A" author="Canudo JI & Royo-Torres R & Cuenca-Bescos G" box="[1111,1329,489,513]" pageId="4" pageNumber="5" pagination="712 - 31" refId="ref11762" refString="Canudo JI, Royo-Torres R, Cuenca-Bescos G. A new sauropod: Tastavinsaurus sanzi gen. et sp. nov. from the Early Cretaceous (Aptian) of Spain. Journal of Vertebrate Paleontology 2008; 28: 712 - 31. https: // doi. org / 10.1671 / 0272 - 4634 (2008) 28 [712: anstsg] 2.0. co; 2" type="journal article" year="2008">
|
||
Canudo
|
||
<emphasis id="B90CEAF3FFE1FFBD5A9634D6FB244A6A" box="[1208,1264,489,513]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
) and in the anteriormost caudal preserved of
|
||
<taxonomicName id="4C784D62FFE1FFBD5A833737FC2B4A54" authority="(Gonzalez Riga et al. 2009)" baseAuthorityName="Gonzalez Riga" baseAuthorityYear="2009" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5A833737FA9E4A4B" box="[1197,1354,520,544]" italics="true" pageId="4" pageNumber="5">Malarguesaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B4C3737FC204A54" author="Gonzalez Riga BJ & Previtera E & Pirrone CA" pageId="4" pageNumber="5" pagination="135 - 48" refId="ref12351" refString="Gonzalez Riga BJ, Previtera E, Pirrone CA. Malarguesaurus florenciae gen. et sp. nov., a new titanosauriform (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina. Cretaceous Research 2009; 30: 135 - 48. https: // doi. org / 10.1016 / j. cretres. 2008.06.006" year="2009">
|
||
González Riga
|
||
<emphasis id="B90CEAF3FFE1FFBD5D563717FC654A54" box="[888,945,551,575]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the ventral surface of the centrum is anteroposteriorly concave, lacking an articular surface for the haemal arch at the anteriormost vertebrae. However, the other caudal vertebrae have a visible articular surface for a haemal arch with very pronounced end, in a similar way to many other titanosaurs.
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE1FFBD5D7B37DBFC324CD9" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||
No signal of the ventrolateral ridges is preserved for
|
||
<taxonomicName id="4C784D62FFE1FFBD5B5737DBFC564B70" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="valdecii">
|
||
<emphasis id="B90CEAF3FFE1FFBD5B5737DBFC564B70" italics="true" pageId="4" pageNumber="5">Tiamat valdecii</emphasis>
|
||
</taxonomicName>
|
||
. The lack of this structure is also observed in several titanosaurs, e.g. in the African titanosaurs
|
||
<taxonomicName id="4C784D62FFE1FFBD5AF8361DFC414B32" authority="(Gorscak et al. 2014)" baseAuthorityName="Gorscak" baseAuthorityYear="2014" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5AF8361DFA9D4B51" box="[1238,1353,802,826]" italics="true" pageId="4" pageNumber="5">Rukwatitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B76361DFC5E4B32" author="Gorscak E & O'Connor PM & Stevens NJ" pageId="4" pageNumber="5" pagination="1133 - 54" refId="ref12567" refString="Gorscak E, O'Connor PM, Stevens NJ et al. The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the Middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 2014; 34: 1133 - 54. https: // doi. org / 10.1080 / 02724634.2014.845568" year="2014">
|
||
Gorscak
|
||
<emphasis id="B90CEAF3FFE1FFBD5B9F361CFC864B32" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C784D62FFE1FFBD5DEB367EFA684B32" authority="(Gorscak and O'Connor 2019)" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[965,1468,833,857]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5DEB367EFBAC4B32" box="[965,1144,833,857]" italics="true" pageId="4" pageNumber="5">Mnyamawamtuka</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5AA6367EFA654B32" author="Gorscak E & O'Connor PM" box="[1160,1457,833,857]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, as well as in
|
||
<taxonomicName id="4C784D62FFE1FFBD5D91365EFAB74B12" authority="(Averianov and Efimov 2018)" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" box="[959,1379,865,889]" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5D91365EFBF74B12" box="[959,1059,865,889]" italics="true" pageId="4" pageNumber="5">Volgatitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5A1B365EFA834B12" author="Averianov A & Efimov V" box="[1077,1367,865,889]" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. But this differs from the well-developed ventrolateral ridges observed in the basal lithostrotians
|
||
<taxonomicName id="4C784D62FFE1FFBD5A0C369FFA9F4BDC" authority="(Wang et al. 2021)" baseAuthorityName="Wang" baseAuthorityYear="2021" box="[1058,1355,927,952]" genus="Hamititan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5A0C369FFB5D4BDC" box="[1058,1161,928,951]" italics="true" pageId="4" pageNumber="5">Hamititan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5AB7369FFAEB4BDC" author="Wang X & Bandeira KLN & Qiu R" box="[1177,1343,927,952]" pageId="4" pageNumber="5" refId="ref14686" refString="Wang X, Bandeira KLN, Qiu R et al. The first dinosaurs from the Early Cretaceous Hami Pterosaur Fauna, China. Scientific Reports 2021; 11: 14962." type="journal volume" year="2021">
|
||
Wang
|
||
<emphasis id="B90CEAF3FFE1FFBD5AF6369FFAD44BDC" box="[1240,1280,927,951]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Also, it differs from the less developed ventrolateral ridges in
|
||
<taxonomicName id="4C784D62FFE1FFBD5B193680FC364B9D" authority="(Gomani 2005)" baseAuthorityName="Gomani" baseAuthorityYear="2005" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5B193680FA154BBC" box="[1335,1473,959,983]" italics="true" pageId="4" pageNumber="5">Malawisaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5D6D36E1FC034B9D" author="Gomani EM" box="[835,983,990,1014]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and in the derived lithostrotian
|
||
<taxonomicName id="4C784D62FFE1FFBD5B6236E1FB1D4C7E" authority="(Santucci and Arruda-Campos 2011)" baseAuthorityName="Santucci and Arruda-Campos" baseAuthorityYear="2011" class="Reptilia" genus="Arrudatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5B6236E1FA164B9D" box="[1356,1474,990,1014]" italics="true" pageId="4" pageNumber="5">Arrudatitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5D6A36C2FB694C7E" author="Santucci RM & Arruda-Campos AC de" box="[836,1213,1021,1045]" pageId="4" pageNumber="5" pagination="1 - 33" refId="ref14368" refString="Santucci RM, Arruda-Campos AC de. A new sauropod (Macronaria, Titanosauria) from the Adamantina Formation, Bauru Group, Upper Cretaceous of Brazil and the phylogenetic relationships of Aeolosaurini. Zootaxa 2011; 3085: 1 - 33." year="2011">Santucci and Arruda-Campos 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Also, there is no ventral longitudinal groove as presented in some titanosauriforms from the Lower Cretaceous, e.g. in
|
||
<taxonomicName id="4C784D62FFE1FFBD5A9A3103FC1D4C18" authority="(Averianov and Efimov 2018)" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5A9A3103FACC4C3F" box="[1204,1304,1084,1108]" italics="true" pageId="4" pageNumber="5">Volgatitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B023103FC694C18" author="Averianov A & Efimov V" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Also ventrally, some few small vascular foramina (‘pneumatopore’) are abundant over the ventral surface of all caudal vertebrae.
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE1FFBD5D7B3186FA6B4D24" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||
One of the features of the new species, the short middle caudal centra (
|
||
<tableCitation id="C6FA035AFFE1FFBD5DA631E6FC074C9A" box="[904,979,1241,1265]" captionStart="Table 1" captionStartId="3.[113,168,143,167]" captionTargetPageId="3" captionText="Table 1. Breakages and wear in the vertebrae of Tiamat resulted from taphonomic processes." pageId="4" pageNumber="5">Table 1</tableCitation>
|
||
) with well-marked articular facets for the haemal arch, is also distinctive from other titanosaurian taxa. This is not observed in other titanosauriforms from the Lower Cretaceous, for example, and even in other taxa from the Upper Cretaceous.
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE1FFBC5D7B3069FF3C4F04" blockId="4.[824,1477,144,1985]" lastBlockId="5.[112,763,1785,1997]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
The transverse processes are relatively short (
|
||
<tableCitation id="C6FA035AFFE1FFBD5B163069FA534D05" box="[1336,1415,1366,1390]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||
) and dorsolaterally projected from the dorsal margin of the vertebral centra, as in many titanosaurs (e.g.
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5A8A30AAFAE64DC7" author="Gomani EM" box="[1188,1330,1428,1453]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B1030AAFBCA4DA7" author="Santucci RM & Arruda-Campos AC de" pageId="4" pageNumber="5" pagination="1 - 33" refId="ref14368" refString="Santucci RM, Arruda-Campos AC de. A new sauropod (Macronaria, Titanosauria) from the Adamantina Formation, Bauru Group, Upper Cretaceous of Brazil and the phylogenetic relationships of Aeolosaurini. Zootaxa 2011; 3085: 1 - 33." year="2011">Santucci and Arruda-Campos 2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5A02308BFA874DA7" author="Averianov A & Efimov V" box="[1068,1363,1460,1484]" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B4C308BFC334D80" author="Gonzalez Riga BJ & Mannion PD & Poropat SF" pageId="4" pageNumber="5" pagination="136 - 81" refId="ref12412" refString="Gonzalez Riga BJ, Mannion PD, Poropat SF et al. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 2018; 184: 136 - 81. https: // doi. org / 10.1093 / zoolinnean / zlx 103" year="2018">
|
||
Gonzàlez Riga
|
||
<emphasis id="B90CEAF3FFE1FFBD5D5D30EBFC7D4D80" box="[883,937,1491,1515]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5DD730ECFAE24D80" author="Gorscak E & O'Connor PM" box="[1017,1334,1491,1515]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B6930EBFCB94E60" author="Wang X & Bandeira KLN & Qiu R" pageId="4" pageNumber="5" refId="ref14686" refString="Wang X, Bandeira KLN, Qiu R et al. The first dinosaurs from the Early Cretaceous Hami Pterosaur Fauna, China. Scientific Reports 2021; 11: 14962." type="journal volume" year="2021">
|
||
Wang
|
||
<emphasis id="B90CEAF3FFE1FFBD5BA330EBFA684D80" box="[1421,1468,1491,1515]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
). This is indicative of the positioning of the caudal vertebrae, due to the development of the transverse processes, especially in the middle caudal vertebrae, where they progressively diminish in size until becoming vestigial (e.g.
|
||
<bibRefCitation id="EFE94B10FFE1FFBD5B4D336EFC5B4EE3" author="Ibiricu LM & Lamanna MC & Martinez RDF" pageId="4" pageNumber="5" pagination="221 - 36" refId="ref12761" refString="Ibiricu LM, Lamanna MC, Martinez RDF et al. A novel form of postcranial skeletal pneumaticity in a sauropod dinosaur: Implications for the paleobiology of Rebbachisauridae. Acta Palaeontologica Polonica 2017; 62: 221 - 36." year="2017">
|
||
Ibiricu
|
||
<emphasis id="B90CEAF3FFE1FFBD5B9F336EFC864EE3" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). This reduction in size progresses in the transverse processes, to the point where these structures transform into heavy blunt knobs, regarded here as the beginning of the middle caudal series, especially in UFRJ-DG 527-R, UFRJ-DG 591-R, UFRJ-DG 635-R, and UFRJ-DG 704-R (
|
||
<figureCitation id="13432A64FFE1FFBD5AF533D2FA924F6E" box="[1243,1350,1773,1797]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4D–G</figureCitation>
|
||
). The transverse processes are triangular and curved posteriorly, similarly to
|
||
<emphasis id="B90CEAF3FFE1FFBD5D173212FC034F2F" box="[825,983,1837,1860]" italics="true" pageId="4" pageNumber="5">Tastavinasaurus</emphasis>
|
||
and
|
||
<taxonomicName id="4C784D62FFE1FFBD5A253213FB734F2F" box="[1035,1191,1836,1860]" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE1FFBD5A253213FB734F2F" box="[1035,1191,1836,1860]" italics="true" pageId="4" pageNumber="5">Malarguesaurus</emphasis>
|
||
</taxonomicName>
|
||
, being supported by a massive and robust PRDL, especially on the most anterior caudal vertebrae (UFRJ-DG 636-R, UFRJ-DG 638-R, and UFRJ-DG 606-R). The neural canal is subcircular being delimited by thick intraprezygapophyseal laminae (TPRL, on anterior view) and intrapost-zygapophyseal lamina (TPOL, on posterior view). The thick TPRL and TPOL of
|
||
<taxonomicName id="4C784D62FFE0FFBC5F9B3226FDE24F5B" box="[437,566,1816,1840]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="differ">
|
||
<emphasis id="B90CEAF3FFE0FFBC5F9B3226FDE24F5B" box="[437,566,1816,1840]" italics="true" pageId="5" pageNumber="6">Tiamat differ</emphasis>
|
||
</taxonomicName>
|
||
from the thin laminae presented by
|
||
<taxonomicName id="4C784D62FFE0FFBC5F1F3208FD584F3B" authority="(Gorsack et al. 2014)" baseAuthorityName="Gorsack" baseAuthorityYear="2014" box="[305,652,1847,1872]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE0FFBC5F1F3208FE704F24" box="[305,420,1847,1871]" italics="true" pageId="5" pageNumber="6">Rukwatitan</emphasis>
|
||
(Gorsack
|
||
<emphasis id="B90CEAF3FFE0FFBC5C3A3207FDEB4F24" box="[532,575,1847,1871]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||
. 2014)
|
||
</taxonomicName>
|
||
and other titanosaurs.
|
||
</paragraph>
|
||
<caption id="DF076669FFE0FFBC5E5F3362FE574EA3" ID-DOI="http://doi.org/10.5281/zenodo.14764393" ID-Zenodo-Dep="14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1629,1653]" targetBox="[115,1456,150,1597]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BC736E1FFE0FFBC5E5F3362FE574EA3" blockId="5.[113,1459,1629,1737]" pageId="5" pageNumber="6">
|
||
<emphasis id="B90CEAF3FFE0FFBC5E5F3362FF1D4E1E" bold="true" box="[113,201,1629,1653]" pageId="5" pageNumber="6">Figure 4.</emphasis>
|
||
Caudal vertebrae attributed to
|
||
<taxonomicName id="4C784D62FFE0FFBC5FC03361FDAD4E1E" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[494,633,1629,1653]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="valdecii" status="gen. et sp. nov.">
|
||
<emphasis id="B90CEAF3FFE0FFBC5FC03361FDAD4E1E" box="[494,633,1629,1653]" italics="true" pageId="5" pageNumber="6">Tiamat valdecii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F5788FFE0FFBC5C513362FCDC4E1F" box="[639,776,1629,1653]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC736E1FFE0FFBC5EA33248FCBB4FA6" blockId="5.[112,763,1785,1997]" lastBlockId="5.[810,1459,1785,1997]" pageId="5" pageNumber="6">
|
||
From the middle caudal vertebrae preserved, UFRJ-DG 635-R shows prezygapophyses dorsomedially positioned, supported ventrally by sharp CPRL. Above the prezygapophyses and the post-zygapophyses,
|
||
<taxonomicName id="4C784D62FFE0FFBC5DD733C6FB404F7A" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1017,1172,1785,1809]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="valdecii">
|
||
<emphasis id="B90CEAF3FFE0FFBC5DD733C6FB404F7A" box="[1017,1172,1785,1809]" italics="true" pageId="5" pageNumber="6">Tiamat valdecii</emphasis>
|
||
</taxonomicName>
|
||
shows a marked accessory tuberosity. The SPRL observed in the most anterior caudal vertebra and in the middle caudal are weakly developed, like the weakly developed SPRL of
|
||
<taxonomicName id="4C784D62FFE0FFBC5A7B3268FCBD4FE5" authority="(Mannion and Calvo 2011)" baseAuthorityName="Mannion and Calvo" baseAuthorityYear="2011" class="Reptilia" family="Titanosauridae" genus="Andesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE0FFBC5A7B3268FB134F04" box="[1109,1223,1879,1903]" italics="true" pageId="5" pageNumber="6">Andesaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE0FFBC5AF23268FC8A4FE5" author="Mannion PD & Calvo JO" pageId="5" pageNumber="6" refId="ref13041" refString="Mannion PD, Calvo JO. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albianearly Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 2011; 163: no - no. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00699. x" year="2011">Mannion and Calvo 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C784D62FFE0FFBC5DB13249FAFE4FE5" authority="(Martinez et al. 2004)" baseAuthorityName="Martinez" baseAuthorityYear="2004" box="[927,1322,1910,1934]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE0FFBC5DB13249FBEE4FE5" box="[927,1082,1910,1934]" italics="true" pageId="5" pageNumber="6">Epachthosaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE0FFBC5A633249FACB4FE5" author="Martinez RD & Gimenez O & Rodriguez J" box="[1101,1311,1910,1934]" pageId="5" pageNumber="6" pagination="107 - 20" refId="ref13311" refString="Martinez RD, Gimenez O, Rodriguez J et al. An articulated specimen of the basal titanosaurian (Dinosauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal formation of Chubut Province, Argentina. Journal of Vertebrate Paleontology 2004; 24: 107 - 20. https: // doi. org / 10.1671 / 9.1" year="2004">
|
||
Martínez
|
||
<emphasis id="B90CEAF3FFE0FFBC5A9C3248FB094FE5" box="[1202,1245,1910,1934]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but differing from the thick SPRL presented by
|
||
<taxonomicName id="4C784D62FFE0FFBC5ABE32A9FCBE4FA6" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE0FFBC5ABE32A9FAD54FC6" box="[1168,1281,1942,1965]" italics="true" pageId="5" pageNumber="6">Patagotitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE0FFBC5B3F32AAFC8A4FA6" author="Carballido JL & Pol D & Otero A" pageId="5" pageNumber="6" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
|
||
Carballido
|
||
<emphasis id="B90CEAF3FFE0FFBC5BAA32A9FA794FC6" box="[1412,1453,1941,1965]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF076669FFE3FFBF5EAF35B0FEFB49EC" pageId="6" pageNumber="7" startId="6.[129,184,143,167]" targetBox="[129,1472,413,1056]" targetIsTable="true" targetPageId="6" targetType="table">
|
||
<paragraph id="8BC736E1FFE3FFBF5EAF35B0FEFB49EC" blockId="6.[129,1471,142,391]" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF35B0FF1A48CD" bold="true" box="[129,206,142,167]" pageId="6" pageNumber="7">Table 2.</emphasis>
|
||
Measurements of caudal vertebrae of
|
||
<taxonomicName id="4C784D62FFE3FFBF5C1C35B0FDA148CD" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[562,629,143,166]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5C1C35B0FDA148CD" box="[562,629,143,166]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations: CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC736E1FFE3FFBF5F7E34A2FAAC4C4B" pageId="6" pageNumber="7">
|
||
<table id="F978C441FFE300465EAF34A2FA144C4B" box="[129,1472,413,1056]" gridcols="8" gridrows="18" pageId="6" pageNumber="7">
|
||
<tr id="354834A3FFE300465EAF34A2FA1449B9" box="[129,1472,413,466]" gridrow="0" pageId="6" pageNumber="7" rowspan-0="1">
|
||
<th id="76995DDFFFE300465F7E34A2FE6149B9" box="[336,437,413,466]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5F7E34A2FE5A49B9" bold="true" box="[336,437,413,466]" pageId="6" pageNumber="7">UFRJ-DG 636-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465FD334A2FDB649B9" box="[509,610,413,466]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5FD334A2FDEE49B9" bold="true" box="[509,610,413,466]" pageId="6" pageNumber="7">UFRJ-DG 638-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465C8734A2FCDA49B9" box="[681,782,413,466]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5C8734A2FD3349B9" bold="true" box="[681,782,413,466]" pageId="6" pageNumber="7">UFRJ-DG 606-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465D7834A2FC6F49B9" box="[854,955,413,466]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5D7834A2FC4749B9" bold="true" box="[854,955,413,466]" pageId="6" pageNumber="7">UFRJ-DG 635-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465A2C34A2FBB349B9" box="[1026,1127,413,466]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5A2C34A2FBEB49B9" bold="true" box="[1026,1127,413,466]" pageId="6" pageNumber="7">UFRJ-DG 591-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465A8134A2FAC049B9" box="[1199,1300,413,466]" gridcol="6" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5A8134A2FB3849B9" bold="true" box="[1199,1300,413,466]" pageId="6" pageNumber="7">UFRJ-DG 527-R</emphasis>
|
||
</th>
|
||
<th id="76995DDFFFE300465B7534A2FA1449B9" box="[1371,1472,413,466]" gridcol="7" gridrow="0" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5B7534A2FA4D49B9" bold="true" box="[1371,1472,413,466]" pageId="6" pageNumber="7">UFRJ-DG 704-R</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF34D5FA144A69" box="[129,1472,490,514]" gridrow="1" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF34D5FEDF4A69" box="[129,267,490,514]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF34D5FF744A69" bold="true" box="[129,160,490,514]" pageId="6" pageNumber="7">CL</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E34D5FE614A69" box="[336,437,490,514]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">7,3</td>
|
||
<td id="76995DDFFFE300465FD334D5FDB64A69" box="[509,610,490,514]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">7,5</td>
|
||
<td id="76995DDFFFE300465C8734D5FCDA4A69" box="[681,782,490,514]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">8,2</td>
|
||
<td id="76995DDFFFE300465D7834D5FC6F4A69" box="[854,955,490,514]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">7</td>
|
||
<td id="76995DDFFFE300465A2C34D5FBB34A69" box="[1026,1127,490,514]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">6,4</td>
|
||
<td id="76995DDFFFE300465A8134D5FAC04A69" box="[1199,1300,490,514]" gridcol="6" gridrow="1" pageId="6" pageNumber="7">6</td>
|
||
<td id="76995DDFFFE300465B7534D5FA144A69" box="[1371,1472,490,514]" gridcol="7" gridrow="1" pageId="6" pageNumber="7">6,1</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3733FA144A4F" box="[129,1472,524,548]" gridrow="2" pageId="6" pageNumber="7" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="76995DDFFFE300465EAF3733FEDF4A4F" box="[129,267,524,548]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3733FEDF4A4F" bold="true" box="[129,267,524,548]" pageId="6" pageNumber="7">CL + condyle</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E3733FE614A4F" box="[336,437,524,548]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">9</td>
|
||
<td id="76995DDFFFE300465FD33733FDB64A4F" box="[509,610,524,548]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">10</td>
|
||
<td id="76995DDFFFE300465C873733FCDA4A4F" box="[681,782,524,548]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">10</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3711FA144A2D" box="[129,1472,558,582]" gridrow="3" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF3711FEDF4A2D" box="[129,267,558,582]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3711FF614A2D" bold="true" box="[129,181,558,582]" pageId="6" pageNumber="7">ACH</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E3711FE614A2D" box="[336,437,558,582]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465FD33711FDB64A2D" box="[509,610,558,582]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465C873711FCDA4A2D" box="[681,782,558,582]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">8</td>
|
||
<td id="76995DDFFFE300465D783711FC6F4A2D" box="[854,955,558,582]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">8,4</td>
|
||
<td id="76995DDFFFE300465A2C3711FBB34A2D" box="[1026,1127,558,582]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">8</td>
|
||
<td id="76995DDFFFE300465A813711FAC04A2D" box="[1199,1300,558,582]" gridcol="6" gridrow="3" pageId="6" pageNumber="7">8,1</td>
|
||
<td id="76995DDFFFE300465B753711FA144A2D" box="[1371,1472,558,582]" gridcol="7" gridrow="3" pageId="6" pageNumber="7">6,6</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF376FFA144A03" box="[129,1472,592,616]" gridrow="4" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF376FFEDF4A03" box="[129,267,592,616]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF376FFF6D4A03" bold="true" box="[129,185,592,616]" pageId="6" pageNumber="7">ACW</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E376FFE614A03" box="[336,437,592,616]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">12,5</td>
|
||
<td id="76995DDFFFE300465FD3376FFDB64A03" box="[509,610,592,616]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">12</td>
|
||
<td id="76995DDFFFE300465C87376FFCDA4A03" box="[681,782,592,616]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">10</td>
|
||
<td id="76995DDFFFE300465D78376FFC6F4A03" box="[854,955,592,616]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465A2C376FFBB34A03" box="[1026,1127,592,616]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465A81376FFAC04A03" box="[1199,1300,592,616]" gridcol="6" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465B75376FFA144A03" box="[1371,1472,592,616]" gridcol="7" gridrow="4" pageId="6" pageNumber="7">8,5</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF374EFA144AE1" box="[129,1472,625,650]" gridrow="5" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF374EFEDF4AE1" box="[129,267,625,650]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF374EFF604AE2" bold="true" box="[129,180,625,649]" pageId="6" pageNumber="7">PCH</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E374EFE614AE1" box="[336,437,625,650]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">10,8</td>
|
||
<td id="76995DDFFFE300465FD3374EFDB64AE1" box="[509,610,625,650]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">10</td>
|
||
<td id="76995DDFFFE300465C87374EFCDA4AE1" box="[681,782,625,650]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">9</td>
|
||
<td id="76995DDFFFE300465D78374EFC6F4AE1" box="[854,955,625,650]" gridcol="4" gridrow="5" pageId="6" pageNumber="7">9</td>
|
||
<td id="76995DDFFFE300465A2C374EFBB34AE1" box="[1026,1127,625,650]" gridcol="5" gridrow="5" pageId="6" pageNumber="7">8</td>
|
||
<td id="76995DDFFFE300465A81374EFAC04AE1" box="[1199,1300,625,650]" gridcol="6" gridrow="5" pageId="6" pageNumber="7">7,5</td>
|
||
<td id="76995DDFFFE300465B75374EFA144AE1" box="[1371,1472,625,650]" gridcol="7" gridrow="5" pageId="6" pageNumber="7">6</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF37ACFA144AC7" box="[129,1472,659,684]" gridrow="6" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF37ACFEDF4AC7" box="[129,267,659,684]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF37ACFF6C4AC0" bold="true" box="[129,184,659,683]" pageId="6" pageNumber="7">PCW</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E37ACFE614AC7" box="[336,437,659,684]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">12,3</td>
|
||
<td id="76995DDFFFE300465FD337ACFDB64AC7" box="[509,610,659,684]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">12</td>
|
||
<td id="76995DDFFFE300465C8737ACFCDA4AC7" box="[681,782,659,684]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465D7837ACFC6F4AC7" box="[854,955,659,684]" gridcol="4" gridrow="6" pageId="6" pageNumber="7">11,5</td>
|
||
<td id="76995DDFFFE300465A2C37ACFBB34AC7" box="[1026,1127,659,684]" gridcol="5" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465A8137ACFAC04AC7" box="[1199,1300,659,684]" gridcol="6" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||
<td id="76995DDFFFE300465B7537ACFA144AC7" box="[1371,1472,659,684]" gridcol="7" gridrow="6" pageId="6" pageNumber="7">7,4</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF378AFA144AA6" box="[129,1472,693,717]" gridrow="7" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF378AFEDF4AA6" box="[129,267,693,717]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF378AFF644AA6" bold="true" box="[129,176,693,717]" pageId="6" pageNumber="7">DFA</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E378AFE614AA6" box="[336,437,693,717]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">4,7</td>
|
||
<td id="76995DDFFFE300465FD3378AFDB64AA6" box="[509,610,693,717]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">5</td>
|
||
<td id="76995DDFFFE300465C87378AFCDA4AA6" box="[681,782,693,717]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465D78378AFC6F4AA6" box="[854,955,693,717]" gridcol="4" gridrow="7" pageId="6" pageNumber="7">5,3</td>
|
||
<td id="76995DDFFFE300465A2C378AFBB34AA6" box="[1026,1127,693,717]" gridcol="5" gridrow="7" pageId="6" pageNumber="7">4,8</td>
|
||
<td id="76995DDFFFE300465A81378AFAC04AA6" box="[1199,1300,693,717]" gridcol="6" gridrow="7" pageId="6" pageNumber="7">4,8</td>
|
||
<td id="76995DDFFFE300465B75378AFA144AA6" box="[1371,1472,693,717]" gridcol="7" gridrow="7" pageId="6" pageNumber="7">5,4</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF37E8FA144A84" box="[129,1472,727,751]" gridrow="8" pageId="6" pageNumber="7">
|
||
<th id="76995DDFFFE300465EAF37E8FEDF4A84" box="[129,267,727,751]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF37E8FF654A84" bold="true" box="[129,177,727,751]" pageId="6" pageNumber="7">DFP</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E37E8FE614A84" box="[336,437,727,751]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">2,1</td>
|
||
<td id="76995DDFFFE300465FD337E8FDB64A84" box="[509,610,727,751]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||
<td id="76995DDFFFE300465C8737E8FCDA4A84" box="[681,782,727,751]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||
<td id="76995DDFFFE300465D7837E8FC6F4A84" box="[854,955,727,751]" gridcol="4" gridrow="8" pageId="6" pageNumber="7">2</td>
|
||
<td id="76995DDFFFE300465A2C37E8FBB34A84" box="[1026,1127,727,751]" gridcol="5" gridrow="8" pageId="6" pageNumber="7">2,1</td>
|
||
<td id="76995DDFFFE300465A8137E8FAC04A84" box="[1199,1300,727,751]" gridcol="6" gridrow="8" pageId="6" pageNumber="7">2,5</td>
|
||
<td id="76995DDFFFE300465B7537E8FA144A84" box="[1371,1472,727,751]" gridcol="7" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF37C6FA144B7A" box="[129,1472,761,785]" gridrow="9" pageId="6" pageNumber="7" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF37C6FEDF4B7A" box="[129,267,761,785]" gridcol="0" gridrow="9" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF37C6FF624B7A" bold="true" box="[129,182,761,785]" pageId="6" pageNumber="7">NAH</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E37C6FE614B7A" box="[336,437,761,785]" gridcol="1" gridrow="9" pageId="6" pageNumber="7">2,1</td>
|
||
<td id="76995DDFFFE300465FD337C6FDB64B7A" box="[509,610,761,785]" gridcol="2" gridrow="9" pageId="6" pageNumber="7">2,2</td>
|
||
<td id="76995DDFFFE300465D7837C6FC6F4B7A" box="[854,955,761,785]" gridcol="4" gridrow="9" pageId="6" pageNumber="7">2</td>
|
||
<td id="76995DDFFFE300465A2C37C6FBB34B7A" box="[1026,1127,761,785]" gridcol="5" gridrow="9" pageId="6" pageNumber="7">2</td>
|
||
<td id="76995DDFFFE300465A8137C6FAC04B7A" box="[1199,1300,761,785]" gridcol="6" gridrow="9" pageId="6" pageNumber="7">1,8</td>
|
||
<td id="76995DDFFFE300465B7537C6FA144B7A" box="[1371,1472,761,785]" gridcol="7" gridrow="9" pageId="6" pageNumber="7">1,7</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3624FA144B58" box="[129,1472,795,819]" gridrow="10" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF3624FEDF4B58" box="[129,267,795,819]" gridcol="0" gridrow="10" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3624FF7B4B58" bold="true" box="[129,175,795,819]" pageId="6" pageNumber="7">PRL</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465D783624FC6F4B58" box="[854,955,795,819]" gridcol="4" gridrow="10" pageId="6" pageNumber="7">5,6</td>
|
||
<td id="76995DDFFFE300465A2C3624FBB34B58" box="[1026,1127,795,819]" gridcol="5" gridrow="10" pageId="6" pageNumber="7">4,6</td>
|
||
<td id="76995DDFFFE300465A813624FAC04B58" box="[1199,1300,795,819]" gridcol="6" gridrow="10" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B753624FA144B58" box="[1371,1472,795,819]" gridcol="7" gridrow="10" pageId="6" pageNumber="7">4,5</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3602FA144B3E" box="[129,1472,829,853]" gridrow="11" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF3602FEDF4B3E" box="[129,267,829,853]" gridcol="0" gridrow="11" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3602FF134B3E" bold="true" box="[129,199,829,853]" pageId="6" pageNumber="7">PRFW</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465D783602FC6F4B3E" box="[854,955,829,853]" gridcol="4" gridrow="11" pageId="6" pageNumber="7">3,9</td>
|
||
<td id="76995DDFFFE300465A2C3602FBB34B3E" box="[1026,1127,829,853]" gridcol="5" gridrow="11" pageId="6" pageNumber="7">4</td>
|
||
<td id="76995DDFFFE300465A813602FAC04B3E" box="[1199,1300,829,853]" gridcol="6" gridrow="11" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B753602FA144B3E" box="[1371,1472,829,853]" gridcol="7" gridrow="11" pageId="6" pageNumber="7">3,5</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3661FA144B1C" box="[129,1472,862,887]" gridrow="12" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF3661FEDF4B1C" box="[129,267,862,887]" gridcol="0" gridrow="12" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3660FF164B1C" bold="true" box="[129,194,863,887]" pageId="6" pageNumber="7">PRFH</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465D783661FC6F4B1C" box="[854,955,862,887]" gridcol="4" gridrow="12" pageId="6" pageNumber="7">3,6</td>
|
||
<td id="76995DDFFFE300465A2C3661FBB34B1C" box="[1026,1127,862,887]" gridcol="5" gridrow="12" pageId="6" pageNumber="7">3,5</td>
|
||
<td id="76995DDFFFE300465A813661FAC04B1C" box="[1199,1300,862,887]" gridcol="6" gridrow="12" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B753661FA144B1C" box="[1371,1472,862,887]" gridcol="7" gridrow="12" pageId="6" pageNumber="7">3</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF36BFFA144BF2" box="[129,1472,896,921]" gridrow="13" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF36BFFEDF4BF2" box="[129,267,896,921]" gridcol="0" gridrow="13" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF36BFFF174BF3" bold="true" box="[129,195,896,920]" pageId="6" pageNumber="7">PSFW</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465D7836BFFC6F4BF2" box="[854,955,896,921]" gridcol="4" gridrow="13" pageId="6" pageNumber="7">4</td>
|
||
<td id="76995DDFFFE300465A2C36BFFBB34BF2" box="[1026,1127,896,921]" gridcol="5" gridrow="13" pageId="6" pageNumber="7">3,2</td>
|
||
<td id="76995DDFFFE300465A8136BFFAC04BF2" box="[1199,1300,896,921]" gridcol="6" gridrow="13" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B7536BFFA144BF2" box="[1371,1472,896,921]" gridcol="7" gridrow="13" pageId="6" pageNumber="7">3,2</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF369DFA144BD1" box="[129,1472,930,954]" gridrow="14" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF369DFEDF4BD1" box="[129,267,930,954]" gridcol="0" gridrow="14" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF369DFF6A4BD1" bold="true" box="[129,190,930,954]" pageId="6" pageNumber="7">PSFH</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465D78369DFC6F4BD1" box="[854,955,930,954]" gridcol="4" gridrow="14" pageId="6" pageNumber="7">3,9</td>
|
||
<td id="76995DDFFFE300465A2C369DFBB34BD1" box="[1026,1127,930,954]" gridcol="5" gridrow="14" pageId="6" pageNumber="7">3,6</td>
|
||
<td id="76995DDFFFE300465A81369DFAC04BD1" box="[1199,1300,930,954]" gridcol="6" gridrow="14" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B75369DFA144BD1" box="[1371,1472,930,954]" gridcol="7" gridrow="14" pageId="6" pageNumber="7">3,4</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF36FBFA144BB7" box="[129,1472,964,988]" gridrow="15" pageId="6" pageNumber="7" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF36FBFEDF4BB7" box="[129,267,964,988]" gridcol="0" gridrow="15" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF36FBFF7A4BB7" bold="true" box="[129,174,964,988]" pageId="6" pageNumber="7">NSL</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E36FBFE614BB7" box="[336,437,964,988]" gridcol="1" gridrow="15" pageId="6" pageNumber="7">6,3</td>
|
||
<td id="76995DDFFFE300465FD336FBFDB64BB7" box="[509,610,964,988]" gridcol="2" gridrow="15" pageId="6" pageNumber="7">5,4</td>
|
||
<td id="76995DDFFFE300465D7836FBFC6F4BB7" box="[854,955,964,988]" gridcol="4" gridrow="15" pageId="6" pageNumber="7">5</td>
|
||
<td id="76995DDFFFE300465A2C36FBFBB34BB7" box="[1026,1127,964,988]" gridcol="5" gridrow="15" pageId="6" pageNumber="7">6,5</td>
|
||
<td id="76995DDFFFE300465A8136FBFAC04BB7" box="[1199,1300,964,988]" gridcol="6" gridrow="15" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B7536FBFA144BB7" box="[1371,1472,964,988]" gridcol="7" gridrow="15" pageId="6" pageNumber="7">6,1</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF36D9FA144B95" box="[129,1472,998,1022]" gridrow="16" pageId="6" pageNumber="7" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF36D9FEDF4B95" box="[129,267,998,1022]" gridcol="0" gridrow="16" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF36D9FF634B95" bold="true" box="[129,183,998,1022]" pageId="6" pageNumber="7">NSW</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E36D9FE614B95" box="[336,437,998,1022]" gridcol="1" gridrow="16" pageId="6" pageNumber="7">4</td>
|
||
<td id="76995DDFFFE300465FD336D9FDB64B95" box="[509,610,998,1022]" gridcol="2" gridrow="16" pageId="6" pageNumber="7">4,5</td>
|
||
<td id="76995DDFFFE300465D7836D9FC6F4B95" box="[854,955,998,1022]" gridcol="4" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||
<td id="76995DDFFFE300465A2C36D9FBB34B95" box="[1026,1127,998,1022]" gridcol="5" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||
<td id="76995DDFFFE300465A8136D9FAC04B95" box="[1199,1300,998,1022]" gridcol="6" gridrow="16" pageId="6" pageNumber="7">-</td>
|
||
<td id="76995DDFFFE300465B7536D9FA144B95" box="[1371,1472,998,1022]" gridcol="7" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||
</tr>
|
||
<tr id="354834A3FFE300465EAF3137FA144C4B" box="[129,1472,1032,1056]" gridrow="17" pageId="6" pageNumber="7" rowspan-3="1">
|
||
<th id="76995DDFFFE300465EAF3137FEDF4C4B" box="[129,267,1032,1056]" gridcol="0" gridrow="17" pageId="6" pageNumber="7">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3137FF654C4B" bold="true" box="[129,177,1032,1056]" pageId="6" pageNumber="7">CRL</emphasis>
|
||
</th>
|
||
<td id="76995DDFFFE300465F7E3137FE614C4B" box="[336,437,1032,1056]" gridcol="1" gridrow="17" pageId="6" pageNumber="7">5,2</td>
|
||
<td id="76995DDFFFE300465FD33137FDB64C4B" box="[509,610,1032,1056]" gridcol="2" gridrow="17" pageId="6" pageNumber="7">5,8</td>
|
||
<td id="76995DDFFFE300465D783137FC6F4C4B" box="[854,955,1032,1056]" gridcol="4" gridrow="17" pageId="6" pageNumber="7">2</td>
|
||
<td id="76995DDFFFE300465A2C3137FBB34C4B" box="[1026,1127,1032,1056]" gridcol="5" gridrow="17" pageId="6" pageNumber="7">2</td>
|
||
<td id="76995DDFFFE300465A813137FAC04C4B" box="[1199,1300,1032,1056]" gridcol="6" gridrow="17" pageId="6" pageNumber="7">1,2</td>
|
||
<td id="76995DDFFFE300465B753137FA144C4B" box="[1371,1472,1032,1056]" gridcol="7" gridrow="17" pageId="6" pageNumber="7">1,2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE3FFBF5EB2315BFE5B4D9F" blockId="6.[128,780,1124,1994]" pageId="6" pageNumber="7">
|
||
Concerning one of the diagnostic features proposed, the marked accessory tuberosity dorsoventrally developed located above the prezygapophyses; is an unusual feature. To our knowledge, no titanosaur presents this tuberosity. The post-zygapophyses are slightly displaced laterally and have concave articular facets, a condition also observed on more anterior caudal vertebrae of
|
||
<taxonomicName id="4C784D62FFE3FFBF5FEE301FFCD14D53" authority="(Gomani 2005)" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[448,773,1311,1336]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5FEE301FFD9E4D53" box="[448,586,1312,1336]" italics="true" pageId="6" pageNumber="7">Malawisaurus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5C4C301FFD2D4D5C" author="Gomani EM" box="[610,761,1311,1336]" pageId="6" pageNumber="7" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, those concave articular facets, being deeply excavated, in the prezygapophyses and the post-zygapophyses of
|
||
<taxonomicName id="4C784D62FFE3FFBF5EAF3041FF1E4DFE" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[129,202,1406,1429]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF3041FF1E4DFE" box="[129,202,1406,1429]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
are a unique feature among titanosaurs, especially with the contemporaneous (or temporally close) taxa compared here, as
|
||
<taxonomicName id="4C784D62FFE3FFBF5EFE3083FE974DBF" box="[208,323,1468,1492]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EFE3083FE974DBF" box="[208,323,1468,1492]" italics="true" pageId="6" pageNumber="7">Rukwatitan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D62FFE3FFBF5F603083FE0C4DBF" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[334,472,1468,1492]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5F603083FE0C4DBF" box="[334,472,1468,1492]" italics="true" pageId="6" pageNumber="7">Malawisaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D62FFE3FFBF5FCC3083FD414DBF" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[482,661,1468,1492]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5FCC3083FD414DBF" box="[482,661,1468,1492]" italics="true" pageId="6" pageNumber="7">Mnyamawamtuka</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D62FFE3FFBF5C8E3083FCD04DBF" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" box="[672,772,1468,1492]" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5C8E3083FCD04DBF" box="[672,772,1468,1492]" italics="true" pageId="6" pageNumber="7">Volgatitan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C784D62FFE3FFBF5EAF30E3FF264D98" box="[129,242,1500,1523]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EAF30E3FF264D98" box="[129,242,1500,1523]" italics="true" pageId="6" pageNumber="7">Patagotitan</emphasis>
|
||
</taxonomicName>
|
||
, among others.
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE3FFBF5EB230C4FDBF4F27" blockId="6.[128,780,1124,1994]" pageId="6" pageNumber="7">
|
||
The new species also presents a hyposphene–hypantrum articulation
|
||
<taxonomicName id="4C784D62FFE3FFBF5EC43325FE514E59" box="[234,389,1562,1586]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5EC43325FE514E59" box="[234,389,1562,1586]" italics="true" pageId="6" pageNumber="7">Epachthosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5FB63325FDCC4E59" author="Powell JE" box="[408,536,1562,1586]" pageId="6" pageNumber="7" pagination="123 - 8" refId="ref13985" refString="Powell JE. Epachthosaurus sciutoi g et sp. nov., un Nuevo dinosaurio titanosaurido del Cretacico de Patagonia. Actas V Congreso Argentino de Paleontologia y Bioestratigrafia 1990; 1: 123 - 8." year="1990">Powell 1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5C083325FD2D4E59" author="Martinez RD & Gimenez O & Rodriguez J" box="[550,761,1562,1586]" pageId="6" pageNumber="7" pagination="107 - 20" refId="ref13311" refString="Martinez RD, Gimenez O, Rodriguez J et al. An articulated specimen of the basal titanosaurian (Dinosauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal formation of Chubut Province, Argentina. Journal of Vertebrate Paleontology 2004; 24: 107 - 20. https: // doi. org / 10.1671 / 9.1" year="2004">
|
||
Martínez
|
||
<emphasis id="B90CEAF3FFE3FFBF5CA23324FD634E59" box="[652,695,1562,1586]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
). However, in
|
||
<taxonomicName id="4C784D62FFE3FFBF5F3F3305FE784E39" box="[273,428,1594,1618]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5F3F3305FE784E39" box="[273,428,1594,1618]" italics="true" pageId="6" pageNumber="7">Epachthosaurus</emphasis>
|
||
</taxonomicName>
|
||
, the hyposphene–hypantrum is observable in some of the anterior caudal vertebrae, but not in caudal vertebrae sequence, which differs from the new species proposed here, where in almost all vertebrae this feature was observed. Both prespinal and postspinal fossae are dorsoventrally elongated but relatively shallow in the anteriormost caudal vertebrae (
|
||
<figureCitation id="13432A64FFE3FFBF5EFF33C9FEEE4F65" box="[209,314,1782,1806]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="6" pageNumber="7">Fig. 4A–A</figureCitation>
|
||
<figureCitation id="13432A64FFE3FFBF5F1733C9FE924F65" box="[313,326,1782,1806]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
|
||
) and in the isolated neural spine associated (
|
||
<figureCitation id="13432A64FFE3FFBF5EA5322AFF034F46" box="[139,215,1813,1837]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="6" pageNumber="7">Fig. 4H</figureCitation>
|
||
). There is no sign of the prespinal or postspinal laminae as in some neosauropods (
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5FBE320BFD8F4F27" author="Mannion PD & Upchurch P & Barnes RN" box="[400,603,1844,1868]" pageId="6" pageNumber="7" pagination="98 - 206" refId="ref13110" refString="Mannion PD, Upchurch P, Barnes RN et al. Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms. Zoological Journal of the Linnean Society 2013; 168: 98 - 206. https: // doi. org / 10.1111 / zoj. 12029" year="2013">
|
||
Mannion
|
||
<emphasis id="B90CEAF3FFE3FFBF5FDA320AFDF64F27" box="[500,546,1844,1868]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE3FFBF5EB2326BFC314D9F" blockId="6.[128,780,1124,1994]" lastBlockId="6.[825,1475,1124,1618]" pageId="6" pageNumber="7">
|
||
Lastly, as in
|
||
<taxonomicName id="4C784D62FFE3FFBF5F00326BFE1F4F07" box="[302,459,1876,1900]" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5F00326BFE1F4F07" box="[302,459,1876,1900]" italics="true" pageId="6" pageNumber="7">Malarguesaurus</emphasis>
|
||
</taxonomicName>
|
||
, the neural spine of
|
||
<taxonomicName id="4C784D62FFE3FFBF5CEF326BFCDE4F00" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[705,778,1876,1899]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5CEF326BFCDE4F00" box="[705,778,1876,1899]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
is rhomboid shaped in dorsal view but is proportionally lower, more elongated, and relatively more robust than the Argentinean species. The neural spine preserved lacks the incipient bifurcation with anteriorly directed tips presented by
|
||
<taxonomicName id="4C784D62FFE3FFBF5D1731BBFB794CF0" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" box="[825,1197,1155,1179]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5D1731BBFC7E4CF0" box="[825,938,1156,1179]" italics="true" pageId="6" pageNumber="7">Patagotitan</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5D9331BCFB754CF0" author="Carballido JL & Pol D & Otero A" box="[957,1185,1155,1179]" pageId="6" pageNumber="7" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
|
||
Carballido
|
||
<emphasis id="B90CEAF3FFE3FFBF5A1D31BBFB8A4CF0" box="[1075,1118,1155,1179]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The isolated neural spine presents a weakly spinoprezygapophyseal lamina (SPRL) while the spinopost-zygapophyseal lamina (SPOL) is comparatively more robust than the SPRL, but the SPOL merges with the rough structure presented on this surface. To our knowledge, this rough structure is unique for titanosaurs, and here is interpreted as an autapomorphy of
|
||
<taxonomicName id="4C784D62FFE3FFBF5A59307FFB6A4D3C" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1143,1214,1344,1367]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5A59307FFB6A4D3C" box="[1143,1214,1344,1367]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
. Lastly, the overall SPRL and SPOL morphology looks like those presented in some derived titanosaurs, such as
|
||
<taxonomicName id="4C784D62FFE3FFBF5A6B3041FB164DFD" authorityName="Díez Díaz, Mocho, Páramo, Escaso, Marcos-Fernández, Sanz & Ortega" authorityYear="2016" box="[1093,1218,1406,1430]" class="Reptilia" genus="Lohuecotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5A6B3041FB164DFD" box="[1093,1218,1406,1430]" italics="true" pageId="6" pageNumber="7">Lohuecotitan</emphasis>
|
||
</taxonomicName>
|
||
. However, a marked difference is that in
|
||
<taxonomicName id="4C784D62FFE3FFBF5DC130A1FBE24DDE" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1007,1078,1438,1461]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5DC130A1FBE24DDE" box="[1007,1078,1438,1461]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
, these laminae are more robust, while those of
|
||
<taxonomicName id="4C784D62FFE3FFBF5DBD3083FBC64DBF" authorityName="Díez Díaz, Mocho, Páramo, Escaso, Marcos-Fernández, Sanz & Ortega" authorityYear="2016" box="[915,1042,1468,1492]" class="Reptilia" genus="Lohuecotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5DBD3083FBC64DBF" box="[915,1042,1468,1492]" italics="true" pageId="6" pageNumber="7">Lohuecotitan</emphasis>
|
||
</taxonomicName>
|
||
present a more complex lamination (
|
||
<bibRefCitation id="EFE94B10FFE3FFBF5BBC3083FC014D9F" author="Diez Diaz V & Mocho P & Paramo A" pageId="6" pageNumber="7" pagination="49 - 60" refId="ref14970" refString="Diez Diaz V, Mocho P, Paramo A, et al. 2016. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Research, 68, 49 - 60. doi: 10.1016 / j. cretres. 2016.08.001" type="journal article" year="2016">
|
||
Díez Díaz
|
||
<emphasis id="B90CEAF3FFE3FFBF5D4030E3FC484D9F" box="[878,924,1500,1524]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736E1FFE3FFBF5D7B30C4FA9D4E39" blockId="6.[825,1475,1124,1618]" pageId="6" pageNumber="7">
|
||
The anterior vertebrae of
|
||
<taxonomicName id="4C784D62FFE3FFBF5A7030C3FB734E78" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1118,1191,1532,1555]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90CEAF3FFE3FFBF5A7030C3FB734E78" box="[1118,1191,1532,1555]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||
</taxonomicName>
|
||
do not have well-preserved zygapophyses; however, the middle caudal vertebrae have characteristics that warrant biomechanical observations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |