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<taxonomicName id="4C2E13FDFFFD4C31D25EFA7202252582" ID-CoL="3R53D" authorityName="Tornabene & Deis & Erdmann" authorityYear="2018" box="[273,527,1493,1516]" family="Gobiidae" genus="Kelloggella" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="145" phylum="Chordata" rank="species" species="avaiki" status="sp. nov.">
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.
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(
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<figureCitation id="131574FBFFFD4C31D26CFA5901A2267A" box="[291,392,1533,1555]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStartId-0="3.[163,245,1168,1190]" captionStartId-1="3.[163,243,1819,1841]" captionStartId-2="4.[145,227,1295,1317]" captionStartId-3="4.[145,225,1819,1841]" captionStartId-4="4.[809,888,1847,1869]" captionStartId-5="5.[163,245,1098,1120]" captionTargetBox-0="[182,760,195,1128]" captionTargetBox-1="[217,759,1295,1739]" captionTargetBox-2="[149,757,905,1255]" captionTargetBox-3="[149,757,1414,1779]" captionTargetBox-4="[813,1421,1121,1789]" captionTargetBox-5="[231,711,195,1058]" captionTargetId-0="figure-513@3.[182,760,195,1128]" captionTargetId-1="figure-485@3.[183,759,1272,1779]" captionTargetId-2="figure-451@4.[149,757,905,1255]" captionTargetId-3="figure-480@4.[149,757,1414,1779]" captionTargetId-4="figure-510@4.[813,1421,1121,1808]" captionTargetId-5="figure-279@5.[231,711,195,1058]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="Figure 1. Kelloggella avaiki sp. nov., live, photo by Mark V.Erdmann." captionText-1="Figure 2. Kelloggella avaiki sp. nov., dorsal profile of head showing layer of gelatinous tissue. USNM 440504, male (one of the six specimens), 19.8 mm SL." captionText-2="Figure 3. Kelloggella avaiki sp. nov., preserved. A, USNM 440501, holotype, 20 mm SL, female. B, USNM 440504, male (one of the six specimens), 19.8 mm SL." captionText-3="Figure 4. Kelloggella avaiki sp. nov., arrangement of head pores. USNM 440504, male (one of the six specimens), 19.8 mm SL. AN, anterior nare; PN, posterior nare." captionText-4="Figure 5. Kelloggella avaiki sp. nov., urogenital papillae. A, female. B, male." captionText-5="Figure 6. Kelloggella avaiki sp. nov., cranial bones, USNM 440502. EO, exoccipital; EPO, epiotic; F, frontal; PT, posttemporal; PTO, pterotic; SOC, supraoccipital; SPH, sphenotic." pageId="2" pageNumber="145">
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1–6
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</figureCitation>
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,
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<figureCitation id="131574FBFFFD4C31D2DBFA5A0188267B" box="[404,418,1533,1554]" captionStart="Figure 9" captionStartId="6.[809,888,691,713]" captionTargetBox="[809,1417,194,640]" captionTargetId="figure-500@6.[809,1417,194,640]" captionTargetPageId="6" captionText="Figure 9. Suspensorium of Kelloggella avaiki sp. nov., USNM 440502. ART, articular; CH, ceratohyal; D, dentary; HYO, hyomandibular; LE, lateral ethmoid; ME, median ethmoid; MPT, metapterygoid; MX, maxilla; PAL, palatine; PMX, premaxilla; POP, preopercle; PT, ectopterygoid; QU, quadrate; SYM, symplectic." pageId="2" pageNumber="145">9</figureCitation>
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,
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<figureCitation id="131574FBFFFD4C31D2E0FA5A01E0267B" box="[431,458,1533,1554]" captionStart="Figure10" captionStartId="7.[164,242,615,637]" captionTargetBox="[291,1315,198,576]" captionTargetId="figure-268@7.[291,1315,198,576]" captionTargetPageId="7" captionText="Figure10. Pectoral girdle of Kelloggella avaiki sp. nov., USNM 440502. CL, cleithrum; COR, coracoid; EN, epineural; PR, pleural rib; PTM, posttemporal; RAD, radial; SCA, scapula; SCL, supracleithrum; VPCL, ventral postcleithrum. Dotted lines are added to highlight the position of the SCL. Note that the VPCL is visible behind the pectoral fin radials in the left frame." pageId="2" pageNumber="145">10</figureCitation>
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,
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<figureCitation id="131574FBFFFD4C31D298FA5A022E267B" box="[471,516,1533,1554]" captionStart="Figure 12" captionStartId="8.[144,222,981,1003]" captionTargetBox="[145,1425,195,941]" captionTargetId="figure-433@8.[145,1425,195,941]" captionTargetPageId="8" captionText="Figure 12. Posttemporals, in situ. A,Kelloggella avaiki sp. nov., USNM 440502. B, Gobiodon citrinus, USNM 306266. C, Kraemeria bryani, USNM 143153. D, Bryaninops sp., USNM 435307. E, Oplopomus oplopomus, USNM 287159. F, Gobiopsis macrostomus, USNM 216191. Morphotypes D–F are the typical gobiid form." pageId="2" pageNumber="145">12A</figureCitation>
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,
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<figureCitation id="131574FBFFFD4C31D15EFA5A0206267B" box="[529,556,1533,1554]" captionStart="Figure 14" captionStartId="9.[821,901,681,703]" captionTargetBox="[825,1433,195,594]" captionTargetId="figure-404@9.[825,1433,195,641]" captionTargetPageId="9" captionText="Figure 14. Vertebral column of Kelloggella avaiki sp. nov., USNM 440502. HA, haemal arch; PR, pleural rib; PTG, pterygiophore; V, vertebra." pageId="2" pageNumber="145">14</figureCitation>
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,
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<figureCitation id="131574FBFFFD4C31D176FA5A024D267A" box="[569,615,1533,1555]" captionStart="Figure 15" captionStartId="10.[145,225,1134,1156]" captionTargetBox="[262,644,181,1093]" captionTargetId="figure-640@10.[261,645,180,1094]" captionTargetPageId="10" captionText="Figure 15. Caudal skeletons. A, Kelloggella avaiki sp. nov., USNM440502.B,Kraemeria samoensis, USNM 439365. C, typical gobiid form, Oplopomus oplopomus, USNM 287159. EPU, epural; HS, haemal spine; HYP, hypural; PC, procurrent cartilage; PHYP, parahypural; NS, neural spine; US, urostyle." pageId="2" pageNumber="145">15A</figureCitation>
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)
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</paragraph>
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@ -1,52 +1,54 @@
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<mods:affiliation id="5AD511DE21B1FD2C0D5844336372400A">School of the Environment and Life Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 1 3 QL, UK & Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:affiliation id="989429D095B8E1A3DEB8B3F6B2181CB5">School of the Environment and Life Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 1 3 QL, UK & Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:affiliation id="61294AA9C23DC8A93A2C497BB1D68B70">Natuurhistorisch Museum Maastricht, De Bosquetplein 6 - 7, 6211 KJ Maastricht, the Netherlands</mods:affiliation>
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<mods:namePart id="8F62F41302B5A1A444C6E0FA5388EAFC">Jagt, John W. M.</mods:namePart>
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<mods:affiliation id="E8E9FA8DABF92EA383F96520591FE85D">Natuurhistorisch Museum Maastricht, De Bosquetplein 6 - 7, 6211 KJ Maastricht, the Netherlands</mods:affiliation>
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<mods:title id="48FB764D12D6A20A4E313441EB51CBF4">Zootaxa</mods:title>
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<mods:title id="518AA47CF3D4E7C8B48CA8D2BD9591F3">Zootaxa</mods:title>
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<mods:part id="2525898717F7C32A5442BC0E7E9CA764">
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<mods:date id="2495FCA57C5133B853050FE64AB3A9E9">2025</mods:date>
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<mods:number id="B3C81BC3D7286011AA475F564E18388F">2025-02-03</mods:number>
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<mods:number id="2740B57081FC80C638798BEACCECA93A">5583</mods:number>
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<mods:location id="BA1BE3CCC788B3830E9382047E0FEEBC">
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<mods:url id="4479894B07F4A8C09F7DEB047D79F399">https://doi.org/10.11646/zootaxa.5583.2.9</mods:url>
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|
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<mods:classification id="E7067A7105DEEEE280C6AB5365E2B07A">journal article</mods:classification>
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<mods:classification id="9A265C7390336610BB6B0809F18CAAC5">journal article</mods:classification>
|
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|
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<mods:identifier id="FBC53D1ED8A46BACAF2B990778E91861" type="DOI">10.11646/zootaxa.5583.2.9</mods:identifier>
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<mods:identifier id="2AF64CF33EC748A990C2B17D7F1B19ED" type="ZooBank">EB8D8B5E-87D9-4299-B90A-2CED350EB581</mods:identifier>
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<treatment id="03A6BE4C1B4BFF9BFF3FF9B4FC720CB2" ID-DOI="http://doi.org/10.5281/zenodo.14797293" ID-Zenodo-Dep="14797293" LSID="urn:lsid:plazi:treatment:03A6BE4C1B4BFF9BFF3FF9B4FC720CB2" httpUri="http://treatment.plazi.org/id/03A6BE4C1B4BFF9BFF3FF9B4FC720CB2" lastPageId="6" lastPageNumber="389" pageId="3" pageNumber="386">
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<subSubSection id="C3155CD11B4BFF9EFF3FF9B4FC710BAE" pageId="3" pageNumber="386" type="nomenclature">
|
||||
|
|
@ -69,7 +71,7 @@ urn:lsid:zoobank.org:act:
|
|||
</subSubSection>
|
||||
<subSubSection id="C3155CD11B4BFF9EFF3FF97DFEED0B82" box="[151,333,1694,1721]" pageId="3" pageNumber="386" type="description">
|
||||
<paragraph id="8BB00F5A1B4BFF9EFF3FF97DFEED0B82" blockId="3.[151,977,1622,1721]" box="[151,333,1694,1721]" pageId="3" pageNumber="386">
|
||||
<figureCitation id="133413DF1B4BFF9EFF3FF97DFEED0B82" box="[151,333,1694,1721]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="3" pageNumber="386">Fig. 2A, B, F–V</figureCitation>
|
||||
<figureCitation id="133413DF1B4BFF9EFF3FF97DFEED0B82" box="[151,333,1694,1721]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="3" pageNumber="386">Fig. 2A, B, F–V</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3155CD11B4BFF9EFF3FF93CFB610A25" pageId="3" pageNumber="386" type="diagnosis">
|
||||
|
|
@ -87,17 +89,15 @@ which bears a striate to complex cancellate sculpture on the fixed scutum, and i
|
|||
<emphasis id="B97BD3481B4BFF9EFF6FF8CBFED50A7A" bold="true" box="[199,373,1831,1858]" pageId="3" pageNumber="386">Type material.</emphasis>
|
||||
<materialsCitation id="3B6705071B4BFF9EFED5F8CBFC0B0A5D" collectionCode="NHMM" pageId="3" pageNumber="386" specimenCode="JJ 6503, JJ 5886, JJ 9619" specimenCount="2" typeStatus="holotype">
|
||||
The large fixed scutum (
|
||||
<figureCitation id="133413DF1B4BFF9EFD38F8CBFCAA0A79" box="[656,778,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="3" pageNumber="386">Fig. 2A, B</figureCitation>
|
||||
<figureCitation id="133413DF1B4BFF9EFD38F8CBFCAA0A79" box="[656,778,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="3" pageNumber="386">Fig. 2A, B</figureCitation>
|
||||
;
|
||||
<collectionCode id="ED1E979F1B4BFF9EFCBFF8CBFCD10A79" box="[791,881,1832,1858]" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="3" pageNumber="386">NHMM</collectionCode>
|
||||
<specimenCode id="DBA9A7211B4BFF9EFCD0F8CBFC6F0A79" box="[888,975,1831,1858]" collectionCode="NHMM" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="3" pageNumber="386">JJ 6503</specimenCode>
|
||||
) is the
|
||||
<typeStatus id="54B4B1F81B4BFF9EFB8AF8CBFB270A79" box="[1058,1159,1832,1858]" pageId="3" pageNumber="386" type="holotype">holotype</typeStatus>
|
||||
, the other figured valves (
|
||||
<figureCitation id="133413DF1B4BFF9EFF37F8AFFEB60A5D" box="[159,278,1868,1894]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="3" pageNumber="386">Fig. 2F–V</figureCitation>
|
||||
) are
|
||||
<typeStatus id="54B4B1F81B4BFF9EFEE5F8AFFE180A5D" box="[333,440,1868,1894]" pageId="3" pageNumber="386" type="paratype">paratypes</typeStatus>
|
||||
(
|
||||
<figureCitation id="133413DF1B4BFF9EFF37F8AFFEB60A5D" box="[159,278,1868,1894]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="3" pageNumber="386">Fig. 2F–V</figureCitation>
|
||||
) are paratypes (
|
||||
<collectionCode id="ED1E979F1B4BFF9EFE6FF8AFFD830A5D" box="[455,547,1868,1894]" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="3" pageNumber="386">NHMM</collectionCode>
|
||||
<specimenCode id="DBA9A7211B4BFF9EFD82F8AFFD220A5D" box="[554,642,1867,1894]" collectionCode="NHMM" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="3" pageNumber="386">JJ 5886</specimenCode>
|
||||
,
|
||||
|
|
@ -130,7 +130,7 @@ locality and horizon.
|
|||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF705FD21B4DFF98FF3FFF3FFE7E0E0F" pageId="5" pageNumber="388" startId="5.[151,250,220,244]" targetType="figure">
|
||||
<caption id="DF705FD21B4DFF98FF3FFF3FFE7E0E0F" ID-DOI="http://doi.org/10.5281/zenodo.14813042" ID-Zenodo-Dep="14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="5" pageNumber="388" startId="5.[151,250,220,244]" targetBox="[151,1435,181,1890]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8BB00F5A1B4DFF98FF3FFF3FFE7E0E0F" blockId="5.[151,1437,220,820]" pageId="5" pageNumber="388">
|
||||
<emphasis id="B97BD3481B4DFF98FF3FFF3FFEAD0DCF" bold="true" box="[151,269,220,244]" pageId="5" pageNumber="388">FIGURE 3</emphasis>
|
||||
. A–E, I, J, O–R,
|
||||
|
|
@ -210,45 +210,45 @@ A total of 16 valves, including 7 fixed scuta, 4 carinae, 1 moveable scutum, 1 m
|
|||
<paragraph id="8BB00F5A1B4DFF98FF6FFC2DFC7C096F" blockId="5.[151,1437,902,2009]" pageId="5" pageNumber="388">
|
||||
<emphasis id="B97BD3481B4DFF98FF6FFC2DFEF90ED3" bold="true" box="[199,345,974,1000]" pageId="5" pageNumber="388">Description.</emphasis>
|
||||
The fixed scuta (
|
||||
<figureCitation id="133413DF1B4DFF98FDBFFC2DFC9E0ED3" box="[535,830,974,1001]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2A–C, E, F, I–K, O–S</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FDBFFC2DFC9E0ED3" box="[535,830,974,1001]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2A–C, E, F, I–K, O–S</figureCitation>
|
||||
) are triangular to subrectangular in outline and display considerable variation in sculpture and shape with ontogeny. The Smallest specimens, approximately
|
||||
<quantity id="4CF7A2BF1B4DFF98FA8DFC11FADD0936" box="[1317,1405,1010,1037]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="388" unit="mm" value="2.5">2.5 mm</quantity>
|
||||
in length (
|
||||
<figureCitation id="133413DF1B4DFF98FF45FBF5FE22090B" box="[237,386,1046,1073]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2C, E, F</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FF45FBF5FE22090B" box="[237,386,1046,1073]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2C, E, F</figureCitation>
|
||||
), are nearly equilaterally triangular and is inclined dorsally. The external sculpture comprises interrupted apicobasal ridges, and the internal scutal adductor scar is large.
|
||||
</paragraph>
|
||||
<paragraph id="8BB00F5A1B4DFF98FF6FFBBDFAD50887" blockId="5.[151,1437,902,2009]" pageId="5" pageNumber="388">
|
||||
The moveable scutum (
|
||||
<figureCitation id="133413DF1B4DFF98FE65FBBDFDE70943" box="[461,583,1118,1144]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2L, M</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FE65FBBDFDE70943" box="[461,583,1118,1144]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2L, M</figureCitation>
|
||||
) is triangular in outline, slightly inclined dorsally, and the height (tip broken off) is greater than the breadth. The external surface carries 6 coarse apicobasal ribs, interrupted by regular growth increments. On the interior, an oval scutal adductor is present adjacent to the occludent margin. The moveable tergum (
|
||||
<figureCitation id="133413DF1B4DFF98FF51FB29FEF509DF" box="[249,341,1226,1252]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2N</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FF51FB29FEF509DF" box="[249,341,1226,1252]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2N</figureCitation>
|
||||
) is asymmetrically kite-shaped and bears a low, sinuous apicobasal ridge which separates the carinal and scutal surfaces. The carinal and occludent margins are long and straight, and angled at 120
|
||||
<superScript id="7C7AA2121B4DFF98FA9EFB0DFA9E09C7" attach="left" box="[1334,1342,1262,1276]" fontSize="6" pageId="5" pageNumber="388">o</superScript>
|
||||
and the scutal margin is weakly convex. The tergal margin (contacting the fixed tergum) is short and convex. The apical part of the occludent/carinal surface carries weak apicobasal ridges, and the scutal surface bears tubercles aligned with the scutal margin. The carina (
|
||||
<figureCitation id="133413DF1B4DFF98FE59FAB9FD73084F" box="[497,723,1370,1397]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2D, G, H, T, U</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FE59FAB9FD73084F" box="[497,723,1370,1397]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2D, G, H, T, U</figureCitation>
|
||||
) is robust, hemiconical and inclined ventrally, and the external surfaces carries weak, interrupted apicobasal ridges. The margin contacting the fixed tergum is broad and serrate, bearing transverse ridges and intervening grooves (
|
||||
<figureCitation id="133413DF1B4DFF98FD7CFA41FCD30887" box="[724,883,1442,1469]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2D, H, U</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FD7CFA41FCD30887" box="[724,883,1442,1469]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2D, H, U</figureCitation>
|
||||
). The rostrum and fixed tergum are unknown.
|
||||
</paragraph>
|
||||
<paragraph id="8BB00F5A1B4DFF98FF64FA24FDD60A3B" blockId="5.[151,1437,902,2009]" pageId="5" pageNumber="388">
|
||||
As the fixed scuta grew during ontogeny, the adductor scar enlarged to occupy most of the internal surface (
|
||||
<figureCitation id="133413DF1B4DFF98FAC7FA25FF540B3F" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2J, Q, S</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FAC7FA25FF540B3F" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2J, Q, S</figureCitation>
|
||||
) and the ventral margin of the valve elongated. The occludent surface became inturned (
|
||||
<figureCitation id="133413DF1B4DFF98FB42FA09FAC20B3F" box="[1258,1378,1514,1540]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2K, S</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FB42FA09FAC20B3F" box="[1258,1378,1514,1540]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2K, S</figureCitation>
|
||||
) and the basal part of this region articulates with the rostrum (see
|
||||
<figureCitation id="133413DF1B4DFF98FCE9F9EDFC1A0B13" box="[833,954,1550,1576]" captionStart="FIGURE 1" captionStartId="1.[151,250,684,708]" captionTargetBox="[158,1430,188,655]" captionTargetId="figure-17@1.[151,1436,181,661]" captionTargetPageId="1" captionText="FIGURE 1. Structure of the capitulum in Proverruca. A, Proverruca vinculum Withers, 1914. B, Proverruca dentifer Gale, 2015 (reconstruction)." figureDoi="http://doi.org/10.5281/zenodo.14797287" httpUri="https://zenodo.org/record/14797287/files/figure.png" pageId="5" pageNumber="388">Fig. 1A, B</figureCitation>
|
||||
). In very large specimens of a length of
|
||||
<quantity id="4CF7A2BF1B4DFF98FA28F9EDFF630B76" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="388" unit="mm" value="10.0">10 mm</quantity>
|
||||
(
|
||||
<figureCitation id="133413DF1B4DFF98FF7AF9D1FEE90B77" box="[210,329,1586,1612]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2A, B</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FF7AF9D1FEE90B77" box="[210,329,1586,1612]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2A, B</figureCitation>
|
||||
) the occludent surface is sharply inflexed, and the rostral margin is angled to the basal margin. The external sculpture changes with growth, and the apicobasal ridges became more numerous and finer (
|
||||
<figureCitation id="133413DF1B4DFF98FAACF9B5FF090BAF" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2J, K, Q, S</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FAACF9B5FF090BAF" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2J, K, Q, S</figureCitation>
|
||||
). These are weaker and interrupted by growth increments in some specimens (
|
||||
<figureCitation id="133413DF1B4DFF98FB8EF999FB220BAF" box="[1062,1154,1658,1684]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2O</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FB8EF999FB220BAF" box="[1062,1154,1658,1684]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2O</figureCitation>
|
||||
). In the largest specimen (
|
||||
<figureCitation id="133413DF1B4DFF98FF37F97DFEB90B83" box="[159,281,1694,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2A, B</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FF37F97DFEB90B83" box="[159,281,1694,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2A, B</figureCitation>
|
||||
), the apicobasal ridges on the lateral (tergal) surface are replaced by transverse rows of closely spaced, irregularly shaped nodes, and a cancellate sculpture comprising weak apicobasal ridges and transverse ridges is present on the occludent surface (
|
||||
<figureCitation id="133413DF1B4DFF98FDA7F905FDCA0A3B" box="[527,618,1766,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="5" pageNumber="388">Fig. 2A</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FDA7F905FDCA0A3B" box="[527,618,1766,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="5" pageNumber="388">Fig. 2A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -267,7 +267,7 @@ differs from
|
|||
<bibRefCitation id="EF9E72AB1B4DFF98FC0AF8E9FB800A1F" author="Gale, A. S." box="[930,1056,1802,1828]" pageId="5" pageNumber="388" pagination="243 - 282" refId="ref3740" refString="Gale, A. S. (2020) New Cretaceous cirripedes from Europe and north Africa. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen, 295 (3), 243-282. https://doi.org/10.1127/njgpa/2020/0886" type="journal article" year="2020">Gale, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133413DF1B4DFF98FB9CF8E9FB160A1F" box="[1076,1206,1802,1828]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="5" pageNumber="388">Fig. 3K–M</figureCitation>
|
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<figureCitation id="133413DF1B4DFF98FB9CF8E9FB160A1F" box="[1076,1206,1802,1828]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="5" pageNumber="388">Fig. 3K–M</figureCitation>
|
||||
) in its larger scutal adductor scar on the fixed tergum and its finer apicobasal ribbing. It is differentiated from
|
||||
<taxonomicName id="4C0F74D91B4DFF98FB33F8CCFA8D0A73" authorityName="Withers" authorityYear="1935" box="[1179,1325,1838,1864]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="cancellata">
|
||||
<emphasis id="B97BD3481B4DFF98FB33F8CCFA8D0A73" box="[1179,1325,1838,1864]" italics="true" pageId="5" pageNumber="388">P. cancellata</emphasis>
|
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|
|
@ -275,14 +275,14 @@ differs from
|
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(
|
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<bibRefCitation id="EF9E72AB1B4DFF98FA94F8CDFF720A56" author="Withers, T. H." pageId="5" pageNumber="388" refId="ref4458" refString="Withers, T. H. (1935) British Museum (Natural History). Catalogue of fossil Cirripedia in the Department of Geology. Vol. 2. Cretaceous. Trustees of the British Museum (Natural History), London, xiii + 534 pp., 50 pls." type="book" year="1935">Withers, 1935</bibRefCitation>
|
||||
;
|
||||
<figureCitation id="133413DF1B4DFF98FF77F8B1FE6D0A57" box="[223,461,1874,1901]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="5" pageNumber="388">Fig. 3A–E, I, J, O–R</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FF77F8B1FE6D0A57" box="[223,461,1874,1901]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="5" pageNumber="388">Fig. 3A–E, I, J, O–R</figureCitation>
|
||||
) in its possession of apicobasal ribbing, and in the presence of tubercles on the scutal surface of the moveable tergum. In
|
||||
<taxonomicName id="4C0F74D91B4DFF98FD97F894FCE00AAA" authority="Gale, 2015" authorityName="Gale" authorityYear="2015" box="[575,832,1910,1937]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="dentifer">
|
||||
<emphasis id="B97BD3481B4DFF98FD97F894FD160AAB" box="[575,694,1910,1936]" italics="true" pageId="5" pageNumber="388">P. dentifer</emphasis>
|
||||
<bibRefCitation id="EF9E72AB1B4DFF98FD69F895FCE00AAA" author="Gale, A. S. & Sorensen, A. M." box="[705,832,1910,1937]" pageId="5" pageNumber="388" pagination="212 - 242" refId="ref3817" refString="Gale, A. S. & Sorensen, A. M. (2015) Taxonomy and palaeoecology of thoracican cirripedes (Crustacea) from a Campanian rocky shoreline at Ivo Klack, southern Sweden. Cretaceous Research, 54, 212-242. https://doi.org/10.1016/j.cretres.2014.09.004" type="journal article" year="2015">Gale, 2015</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133413DF1B4DFF98FCFCF895FC770AAB" box="[852,983,1910,1936]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="5" pageNumber="388">Fig. 3X, Y</figureCitation>
|
||||
<figureCitation id="133413DF1B4DFF98FCFCF895FC770AAB" box="[852,983,1910,1936]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="5" pageNumber="388">Fig. 3X, Y</figureCitation>
|
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), the moveable scutum carries regular, apicobasal rows of thorns on the scutal surface; these are absent in
|
||||
<taxonomicName id="4C0F74D91B4DFF98FC29F878FC480A8F" authorityName="Gale & Jagt" authorityYear="2025" box="[897,1000,1947,1972]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
|
||||
<emphasis id="B97BD3481B4DFF98FC29F878FC480A8F" box="[897,1000,1947,1972]" italics="true" pageId="5" pageNumber="388">P. ornata</emphasis>
|
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|
|
@ -295,7 +295,7 @@ in which the few tubercles are arranged in margin-parallel rows. Additionally, t
|
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<emphasis id="B97BD3481B4DFF98FC7AF85CFBE30AE3" box="[978,1091,1982,2008]" italics="true" pageId="5" pageNumber="388">P. dentifer</emphasis>
|
||||
</taxonomicName>
|
||||
is smaller and positioned close to the basal margin (
|
||||
<figureCitation id="133413DF1B4EFF9BFED2FF74FE6E0D8A" box="[378,462,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="6" pageNumber="389">Fig. 3S</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFED2FF74FE6E0D8A" box="[378,462,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="6" pageNumber="389">Fig. 3S</figureCitation>
|
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).
|
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<taxonomicName id="4C0F74D91B4EFF9BFE4AFF74FD140D8B" authorityName="Gale & Jagt" authorityYear="2025" box="[482,692,151,176]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
|
||||
<emphasis id="B97BD3481B4EFF9BFE4AFF74FD140D8B" box="[482,692,151,176]" italics="true" pageId="6" pageNumber="389">Proverruca ornata</emphasis>
|
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|
|
@ -309,14 +309,14 @@ is distinguished from
|
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<bibRefCitation id="EF9E72AB1B4EFF9BFB3BFF74FA900D8A" author="Withers, T. H." box="[1171,1328,151,177]" pageId="6" pageNumber="389" pagination="945 - 953" refId="ref4366" refString="Withers, T. H. (1914) A remarkable new Cirripede from the Chalk of Surrey and Hertfordshire. Proceedings of the Zoological Society, London, 1914, 945-953, pl. 1. https://doi.org/10.1111/j.1469-7998.1914.tb07721.x" type="book chapter" year="1914">Withers, 1914</bibRefCitation>
|
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</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133413DF1B4EFF9BFA97FF74FF0E0DEE" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="6" pageNumber="389">Fig. 3G, H</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFA97FF74FF0E0DEE" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="6" pageNumber="389">Fig. 3G, H</figureCitation>
|
||||
),
|
||||
<taxonomicName id="4C0F74D91B4EFF9BFF6CFF58FE760DEE" authority="Withers, 1935" authorityName="Withers" authorityYear="1935" box="[196,470,186,213]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="nodosa">
|
||||
<emphasis id="B97BD3481B4EFF9BFF6CFF58FE910DEE" box="[196,305,187,213]" italics="true" pageId="6" pageNumber="389">P. nodosa</emphasis>
|
||||
<bibRefCitation id="EF9E72AB1B4EFF9BFE90FF58FE760DEE" author="Withers, T. H." box="[312,470,186,213]" pageId="6" pageNumber="389" refId="ref4458" refString="Withers, T. H. (1935) British Museum (Natural History). Catalogue of fossil Cirripedia in the Department of Geology. Vol. 2. Cretaceous. Trustees of the British Museum (Natural History), London, xiii + 534 pp., 50 pls." type="book" year="1935">Withers, 1935</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133413DF1B4EFF9BFE4EFF58FDC00DEE" box="[486,608,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="6" pageNumber="389">Fig. 3Z, A</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFE4EFF58FDC00DEE" box="[486,608,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="6" pageNumber="389">Fig. 3Z, A</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFDC8FF58FDCA0DEE" box="[608,618,187,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,684,708]" captionTargetBox="[158,1430,188,655]" captionTargetId="figure-17@1.[151,1436,181,661]" captionTargetPageId="1" captionText="FIGURE 1. Structure of the capitulum in Proverruca. A, Proverruca vinculum Withers, 1914. B, Proverruca dentifer Gale, 2015 (reconstruction)." figureDoi="http://doi.org/10.5281/zenodo.14797287" httpUri="https://zenodo.org/record/14797287/files/figure.png" pageId="6" pageNumber="389">1</figureCitation>
|
||||
) and
|
||||
<taxonomicName id="4C0F74D91B4EFF9BFD03FF58FC160DEE" authority="Withers, 1923" authorityName="Withers" authorityYear="1923" box="[683,950,187,213]" class="Maxillopoda" family="Proverrucidae" genus="Proverruca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="389" phylum="Arthropoda" rank="species" species="laurae">
|
||||
|
|
@ -324,8 +324,8 @@ is distinguished from
|
|||
<bibRefCitation id="EF9E72AB1B4EFF9BFCB0FF58FC160DEE" author="Withers, T. H." box="[792,950,187,213]" pageId="6" pageNumber="389" pagination="1 - 54" refId="ref4424" refString="Withers, T. H. (1923) Die Cirripedien der Kreide Rugens. Abhandlungen aus dem Geologisch-Palaontologischen Institut der Universitat zu Greifswald, 3, 1-54, 3 pls." type="book chapter" year="1923">Withers, 1923</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="133413DF1B4EFF9BFC6EFF58FBBE0DEE" box="[966,1054,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." pageId="6" pageNumber="389">Fig. 3A</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFBB6FF58FB8C0DEE" box="[1054,1068,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." pageId="6" pageNumber="389">2</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFC6EFF58FBBE0DEE" box="[966,1054,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,220,244]" captionTargetId="figure-25@4.[151,1435,181,1890]" captionTargetPageId="4" captionText="FIGURE 3. A–E, I, J, O–R, Proverruca cancellata Withers, 1935. A, fixed scutum, in apical view (NHMUK PI In 65335). B, C, fixed terga, in apical view (NHMUK PI In 65336 and 65337). D, E, moveable terga, external views (NHMUK PI In 65337 and 65338). I, J, fixed scuta, external views (NHMUK PI In 65343 and 65344). O–R, moveable scuta, in external (Q, R) and internal (O, P) views (NHMUK PI In 65339, 65340, 65345 and 65349, respectively), lower Campanian, West Sussex (UK). F– H, Proverruca vinculum Withers, 1914, holotype capitulum (BGS 3204) in oblique apical (F), lateral (G) and apical (H) views; the original of Withers (1914, text-fig. 1C–F; Withers 1935, figs 33–36; Gale 2014, fig. 14a–c), probably from Coniacian chalk, Watford Tunnel (Hertfordshire, UK).K–N, Proverruca anglica Gale, 2020; fixed scuta, in external (K, M) and internal (L,N) views. M, N, holotype, the original of Gale (2020, fig. 7P, Q) (NHMUK IC 1478). K, L, paratype, the original of Gale (2020, fig. 7N, O) (NHMUK IC 1477), upper Campanian, Norwich Chalk, Cringleford, Norwich (Norfolk, UK). S–Y, Proverruca dentiferGale, in Gale & Sørensen, 2015. S, T, fixed scutum, in internal (S) and external (T) views, the original of Gale, in Gale & Sørensen (2015, fig. 14J, K) (NHMUK IC 1046). U–W, carina, in apical (U), internal (V) and external (W) views, the original of Gale, in Gale & Sørensen (2015, fig. 14A–C) (NHMUK IC 1040). X, Y, external views of moveable terga, the originals of Gale, in Gale & Sørensen (2015, fig. 14D, E) (NHMUK IC 1041 [holotype] and IC 1042), upper lower Campanian, Ivö Klack, southern Sweden. Z, A1, Proverruca nodosa Withers, 1935, moveable tergum, external view, “Chalk detritus, Charing, Kent” (UK), probably Cenomanian (NHMUK I.14720). A2, Proverruca laurae Withers, 1923, moveable tergum, external view, lower Maastrichtian, Rügen, northeastern Germany (SNSB SNHMB- G 5677). Scale bars equal 0.2 mm (A–E, I, J, O–R), 1 mm (F–H, O, P, S–A2) and 0.5 mm (K–N)." figureDoi="http://doi.org/10.5281/zenodo.14813042" httpUri="https://zenodo.org/record/14813042/files/figure.png" pageId="6" pageNumber="389">Fig. 3A</figureCitation>
|
||||
<figureCitation id="133413DF1B4EFF9BFBB6FF58FB8C0DEE" box="[1054,1068,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,220,244]" captionTargetId="figure-25@2.[151,1435,181,1826]" captionTargetPageId="2" captionText="FIGURE 2. Proverruca ornata sp. nov. (A–U):A, B, large fixed scutum, external (A) and internal (B) views (holotype, NHMM JJ 6503). C, small fixed scutum, external view (paratype, NHMM JJ 9619). D, fragmentary carina, internal view (paratype, NHMM JJ 9619). E, F, fragmentary fixed scutum, external (E) and internal (F) views (paratype, NHMM JJ 9619a). G, H, large carina, external (G) and internal (H) views (paratype, NHMM JJ 9619b). I–K, fixed scutum, in internal (I), external (J) and apical (K) views (paratype, NHMM JJ 9619c). L, M, moveable scutum, in internal (L) and external (M) views (paratype, NHMM JJ 9619d). N, moveable tergum, external view (paratype, NHMM JJ 9619e). O, P, fixed scutum, external (O) and internal (P) views (paratype, NHMM JJ 13426). Q, fixed scutum, external view (paratype, NHMM JJ 9619). R, S, fixed scutum, in internal (R) and apical (S) views (paratype, NHMM JJ 9619f). T, U, carina, in external (T) and internal (U) views (paratype, NHMM JJ 5886) Provenance: A, B, G–T, Zeven Wegen Member (Gulpen Formation), lower upper Campanian, former CPL quarry, Haccourt (northwestern Belgium). Scale bars equal 5 mm (A, B), 1 mm (H, I, O, P), 0.4 mm (D, I–K, O, P, R, S) and 0.3 mm (C, E, F, N, T, U)." figureDoi="http://doi.org/10.5281/zenodo.14813040" httpUri="https://zenodo.org/record/14813040/files/figure.png" pageId="6" pageNumber="389">2</figureCitation>
|
||||
) in both the shape and sculpture of the valves.
|
||||
</paragraph>
|
||||
<paragraph id="8BB00F5A1B4EFF9BFF6FFEE0FC720CB2" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="389">
|
||||
|
|
|
|||
122
data/03/C3/87/03C387B9A008FFCFC059FAC4FA931C47.xml
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122
data/03/C3/87/03C387B9A008FFCFC059FAC4FA931C47.xml
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<mods:title id="2A175F6C290F25D7FB215A7FCBAA5861">Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification</mods:title>
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<taxonomicName id="4C6A4D2CA008FFCFC020FAC4FDFA184D" ID-CoL="7L7P" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[360,542,1334,1358]" class="Gastropoda" family="Aglajidae" genus="Spinophallus" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus" status="gen. nov.">
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.
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.
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</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13512A2AA008FFCFC065FAAFFE741876" box="[301,400,1373,1398]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">
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<smallCapsWord id="8D33A073A008FFCFC065FAAFFE871875" baselines="1392,1392" box="[301,355,1373,1398]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="39" pageNumber="40">FIGS</smallCapsWord>
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3M
|
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</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA008FFCFC0D5FAAFFDC41875" box="[413,544,1373,1398]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">
|
||||
5B;
|
||||
<smallCapsWord id="8D33A073A008FFCFC081FAAFFDC41875" baselines="1392,1392" box="[457,544,1373,1398]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Tables" pageId="39" pageNumber="40">TABLES</smallCapsWord>
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||||
</figureCitation>
|
||||
1
|
||||
<smallCapsWord id="8D33A073A008FFCFC374FA93FD8F1875" baselines="1392" box="[572,619,1377,1397]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="39" pageNumber="40">AND</smallCapsWord>
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2)
|
||||
</paragraph>
|
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<paragraph id="8BD536AFA008FFCFC1EBFA74FE5318BA" blockId="39.[163,778,1414,1466]" pageId="39" pageNumber="40">
|
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<emphasis id="B91EEABDA008FFCFC1EBFA74FED2189B" box="[163,310,1414,1435]" italics="true" pageId="39" pageNumber="40">
|
||||
<typeStatus id="54D1880DA008FFCFC1EBFA74FF3D189B" box="[163,217,1414,1435]" pageId="39" pageNumber="40">Type</typeStatus>
|
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species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC076FA74FD04189B" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[318,736,1414,1436]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="coronata">
|
||||
<emphasis id="B91EEABDA008FFCFC076FA74FDCA189B" box="[318,558,1414,1435]" italics="true" pageId="39" pageNumber="40">Philinopsis coronata</emphasis>
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC37EFA74FD04189B" author="Gosliner TM" box="[566,736,1414,1436]" pageId="39" pageNumber="40" pagination="1 - 24" refId="ref27539" refString="Gosliner TM. 2011. Six new species of aglajids opisthobranch molluscs from the tropical Indo-Pacific. Zootaxa 2751: 1-24." type="journal article" year="2011">Gosliner, 2011</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. By subsequent designation.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC1EBFA26FEBE1A58" blockId="39.[163,779,1491,1881]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B91EEABDA008FFCFC1EBFA26FEF918E9" box="[163,285,1492,1513]" italics="true" pageId="39" pageNumber="40">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA008FFCFC34CFA21FDB718E8" box="[516,595,1491,1513]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="mm" value="15.0">15 mm</quantity>
|
||||
in length. Body elongated, wide; cephalic shield blunt, quadrangular; reduced sensory bristles, not visible to the naked eye; posterior shield rounded, terminates in a medial, elongate, conical or bulbous posterior projection (autapomorphy); symmetrical caudal lobes, short, blunt; parapodia short (
|
||||
<figureCitation id="13512A2AA008FFCFC0F2F979FDF71BA0" box="[442,531,1675,1697]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">Fig. 3M</figureCitation>
|
||||
). Shell calcified, convex-dilated with open whorl (
|
||||
<figureCitation id="13512A2AA008FFCFC344F958FD871BBF" box="[524,611,1705,1728]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">Fig. 5B</figureCitation>
|
||||
). Buccal bulb large, muscular, bulb-shaped. Penial sac with internal spines; penial papilla covered by scattered series of large spines (autapomorphy), or with a ring of rounded tubercles (autapomorphy) (
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC303F8D6FCE01A39" author="Rudman WB" box="[587,772,1828,1850]" pageId="39" pageNumber="40" pagination="381 - 399" refId="ref29205" refString="Rudman WB. 1972 a. A comparative study of the genus philinopsis Pease 1860 (Aglajidae, Opisthobranchia). Pacific Science 26: 381-399." type="journal article" year="1972">Rudman, 1972a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC1EBF8B1FEAE1A58" author="Gosliner TM" box="[163,330,1859,1881]" pageId="39" pageNumber="40" pagination="1 - 24" refId="ref27539" refString="Gosliner TM. 2011. Six new species of aglajids opisthobranch molluscs from the tropical Indo-Pacific. Zootaxa 2751: 1-24." type="journal article" year="2011">Gosliner, 2011</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC273FF34FAED1DDB" blockId="39.[827,1289,197,219]" box="[827,1289,197,219]" pageId="39" pageNumber="40">
|
||||
<materialsCitation id="3B023CF2A008FFCFC273FF34FAE21DDB" box="[827,1286,197,219]" country="Philippines" location="The" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
|
||||
<emphasis id="B91EEABDA008FFCFC273FF34FC2E1DDA" box="[827,970,197,219]" italics="true" pageId="39" pageNumber="40">
|
||||
<typeStatus id="54D1880DA008FFCFC273FF34FC941DDB" box="[827,880,198,219]" pageId="39" pageNumber="40">Type</typeStatus>
|
||||
locality
|
||||
</emphasis>
|
||||
:
|
||||
<collectingRegion id="49AEF84DA008FFCFC29FFF34FBA01DDB" box="[983,1092,198,219]" country="Philippines" name="Batangas" pageId="39" pageNumber="40">Batangas</collectingRegion>
|
||||
,
|
||||
<location id="8EB56074A008FFCFC506FF37FB9E1DDB" LSID="urn:lsid:plazi:treatment:03C387B9A008FFCFC059FAC4FA931C47:8EB56074A008FFCFC506FF37FB9E1DDB" box="[1102,1146,197,219]" country="Philippines" name="The" pageId="39" pageNumber="40" stateProvince="Batangas">The</location>
|
||||
<collectingCountry id="F37D763FA008FFCFC5C9FF37FAE21DDB" box="[1153,1286,197,219]" name="Philippines" pageId="39" pageNumber="40">Philippines</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC273FF06FA931C47" blockId="39.[827,1442,244,327]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B91EEABDA008FFCFC273FF06FC251C09" box="[827,961,244,265]" italics="true" pageId="39" pageNumber="40">Etymology:</emphasis>
|
||||
The genus name (Lat.
|
||||
<emphasis id="B91EEABDA008FFCFC5AFFF06FAD31C09" box="[1255,1335,244,265]" italics="true" pageId="39" pageNumber="40">schedo</emphasis>
|
||||
= penis;
|
||||
<emphasis id="B91EEABDA008FFCFC273FEE1FC931C28" box="[827,887,275,296]" italics="true" pageId="39" pageNumber="40">spino</emphasis>
|
||||
= spines) refers to the spines present on the penial papilla or in the penial sac of species in this genus.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
143
data/03/C3/87/03C387B9A008FFCFC07AFF36FF1019E2.xml
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143
data/03/C3/87/03C387B9A008FFCFC07AFF36FF1019E2.xml
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<mods:title id="D171747FBFDF3C2DAC0E50D662A72252">Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification</mods:title>
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<mods:namePart id="EA033D8DA417276B28D2A60DC70047DF">Malaquias, Manuel António E.</mods:namePart>
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<heading id="D09D81C3A008FFCFC07AFF36FD981DDC" box="[306,636,196,221]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="40" reason="2">
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<taxonomicName id="4C6A4D2CA008FFCFC0C1FF36FE1A1DDA" ID-CoL="7P9VJ" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[393,510,196,219]" class="Gastropoda" family="Aglajidae" genus="Niparaya" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus" status="gen. nov.">
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.
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.
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<paragraph id="8BD536AFA008FFCFC064FF18FD651C03" blockId="39.[300,641,234,259]" box="[300,641,234,259]" pageId="39" pageNumber="40">
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(
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<figureCitation id="13512A2AA008FFCFC07DFF1FFE6F1C01" box="[309,395,236,258]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">
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3J
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</figureCitation>
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,
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||||
<figureCitation id="13512A2AA008FFCFC0D0FF1EFDFF1C01" box="[408,539,234,259]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">
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5D;
|
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</figureCitation>
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1
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2)
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</paragraph>
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<typeStatus id="54D1880DA008FFCFC1EBFEE6FF3C1C29" box="[163,216,276,297]" pageId="39" pageNumber="40">Type</typeStatus>
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species:
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</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC07EFEE1FD481C28" authority="Bertsch, 1972" authorityName="Bertsch" authorityYear="1972" box="[310,684,275,297]" class="Gastropoda" family="Aglajidae" genus="Aglaja" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="regiscorona">
|
||||
<emphasis id="B91EEABDA008FFCFC07EFEE1FDED1C29" box="[310,521,275,297]" italics="true" pageId="39" pageNumber="40">Aglaja regiscorona</emphasis>
|
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<bibRefCitation id="EFFB4B5EA008FFCFC347FEE1FD481C28" author="Bertsch H" box="[527,684,275,297]" pageId="39" pageNumber="40" pagination="103 - 108" refId="ref26720" refString="Bertsch H. 1972. Two additions to the Opisthobranch fauna of the Southern Gulf of California. The Veliger 34: 103-108." type="journal article" year="1972">Bertsch, 1972</bibRefCitation>
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</taxonomicName>
|
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. By subsequent designation.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC1EBFE93FD671E60" blockId="39.[163,779,352,865]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B91EEABDA008FFCFC1EBFE93FECD1C76" box="[163,297,353,374]" italics="true" pageId="39" pageNumber="40">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA008FFCFC371FE92FD651C76" box="[569,641,352,374]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="40" unit="mm" value="5.0">5 mm</quantity>
|
||||
in length. Cephalic shield acuminated, with translucent velum along the front side (autapomorphy) and a posterior, small, three-pointed elevated crown-like projection (autapomorphy); reduced sensory bristles, not visible to the naked eye; posterior shield with bulbous pustules (autapomorphy); symmetrical caudal lobes, rounded, flat; parapodia reduced (
|
||||
<figureCitation id="13512A2AA008FFCFC1E4FDA4FF1B1F6B" box="[172,255,598,620]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">Fig. 3J</figureCitation>
|
||||
). Shell calcified, concave, constricted, with closed nuclear whorl, with ridges separated by a slight indentation (
|
||||
<figureCitation id="13512A2AA008FFCFC0C1FD61FE3A1FA9" box="[393,478,659,681]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">Fig. 5D</figureCitation>
|
||||
). Small ampulla, slightly coiled; mucous gland faintly curved; receptaculum seminis entering the hermaphroditic duct at the middle of its length. Penial papilla with a rounded distal bulb (autapomorphy); prostatic region with two sections varying in shape and texture (autapomorphy) (
|
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<bibRefCitation id="EFFB4B5EA008FFCFC05FFCB9FE5F1E60" author="Bertsch H" box="[279,443,843,865]" pageId="39" pageNumber="40" pagination="103 - 108" refId="ref26720" refString="Bertsch H. 1972. Two additions to the Opisthobranch fauna of the Southern Gulf of California. The Veliger 34: 103-108." type="journal article" year="1972">Bertsch, 1972</bibRefCitation>
|
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;
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC081FCB9FD971E60" author="Gosliner TM" box="[457,627,843,865]" pageId="39" pageNumber="40" pagination="325 - 360" refId="ref27454" refString="Gosliner TM. 1980. Systematics and phylogeny of the aglajidae (Opisthobranchia: Mollusca). Zoological Journal of the Linnaean Society 68: 325-360." type="journal article" year="1980">Gosliner, 1980</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
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<paragraph id="8BD536AFA008FFCFC1EBFC78FF1E1EBE" blockId="39.[163,778,906,958]" pageId="39" pageNumber="40">
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<materialsCitation id="3B023CF2A008FFCFC1EBFC78FF121EBE" country="Mexico" location="Bahia" municipality="Cruces" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Baja California" typeStatus="holotype">
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<emphasis id="B91EEABDA008FFCFC1EBFC78FEA61E9F" box="[163,322,906,927]" italics="true" pageId="39" pageNumber="40">
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<typeStatus id="54D1880DA008FFCFC1EBFC78FF3E1E9F" box="[163,218,906,927]" pageId="39" pageNumber="40">Type</typeStatus>
|
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locality:
|
||||
</emphasis>
|
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<location id="8EB56074A008FFCFC003FC78FE761EA0" LSID="urn:lsid:plazi:treatment:03C387B9A008FFCFC07AFF36FF1019E2:8EB56074A008FFCFC003FC78FE761EA0" box="[331,402,906,928]" country="Mexico" municipality="Cruces" name="Bahia" pageId="39" pageNumber="40" stateProvince="Baja California">Bahía</location>
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||||
de las
|
||||
<collectingMunicipality id="6BB1ACD5A008FFCFC0A3FC78FDDA1E9F" box="[491,574,906,927]" pageId="39" pageNumber="40">Cruces</collectingMunicipality>
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,
|
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<collectingRegion id="49AEF84DA008FFCFC304FC78FCE21EA0" box="[588,774,906,928]" country="Mexico" name="Baja California" pageId="39" pageNumber="40">Baja California</collectingRegion>
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||||
,
|
||||
<collectingCountry id="F37D763FA008FFCFC1EBFC5AFF121EBE" box="[163,246,936,958]" name="Mexico" pageId="39" pageNumber="40">México</collectingCountry>
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.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC1EBFC25FF1019E2" blockId="39.[163,778,983,1250]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B91EEABDA008FFCFC1EBFC25FECE1EEC" box="[163,298,983,1004]" italics="true" pageId="39" pageNumber="40">Etymology:</emphasis>
|
||||
The word ‘niparaya’ means ‘creator of everything’ in Pericú, the language of the aboriginal inhabitants of the southern part of the
|
||||
<collectingRegion id="49AEF84DA008FFCFC39AFBE6FEFB1949" country="Mexico" name="Baja California" pageId="39" pageNumber="40">Baja California</collectingRegion>
|
||||
Peninsula (
|
||||
<collectingCountry id="F37D763FA008FFCFC0FBFBC1FDE91949" box="[435,525,1075,1097]" name="Mexico" pageId="39" pageNumber="40">México</collectingCountry>
|
||||
), the
|
||||
<typeStatus id="54D1880DA008FFCFC31BFBC1FD631948" box="[595,647,1075,1096]" pageId="39" pageNumber="40">type</typeStatus>
|
||||
locality of
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC1EBFBA0FEAE1967" authorityName="Bertsch" authorityYear="1972" box="[163,330,1106,1127]" class="Gastropoda" family="Aglajidae" genus="Aglaja" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="regiscorona">
|
||||
<emphasis id="B91EEABDA008FFCFC1EBFBA0FEAE1967" box="[163,330,1106,1127]" italics="true" pageId="39" pageNumber="40">N. regiscorona</emphasis>
|
||||
</taxonomicName>
|
||||
and is also the name of the main deity and king of the Pericú mythology. Thus, the genus name
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC1A3FB62FEB819A5" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[235,348,1168,1189]" class="Gastropoda" family="Aglajidae" genus="Niparaya" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B91EEABDA008FFCFC1A3FB62FEB819A5" box="[235,348,1168,1189]" italics="true" pageId="39" pageNumber="40">Niparaya</emphasis>
|
||||
</taxonomicName>
|
||||
is a metaphor for the ‘crowned slug’
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC1EBFB5CFEAB19C3" authorityName="Bertsch" authorityYear="1972" box="[163,335,1198,1219]" class="Gastropoda" family="Aglajidae" genus="Aglaja" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="regiscorona">
|
||||
<emphasis id="B91EEABDA008FFCFC1EBFB5CFEAB19C3" box="[163,335,1198,1219]" italics="true" pageId="39" pageNumber="40">N. regiscorona</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName id="4C6A4D2CA008FFCFC028FB5CFE1419C3" authorityName="Bertsch" authorityYear="1972" box="[352,496,1198,1219]" class="Gastropoda" family="Aglajidae" genus="Aglaja" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="regiscorona">
|
||||
‘
|
||||
<emphasis id="B91EEABDA008FFCFC02EFB5CFE1419C3" box="[358,496,1198,1219]" italics="true" pageId="39" pageNumber="40">regiscorona</emphasis>
|
||||
</taxonomicName>
|
||||
’ means king’s crown in Latin).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
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127
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127
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.
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<paragraph id="8BD536AFA008FFCFC235FE5DFA841CC8" blockId="39.[893,1376,431,456]" box="[893,1376,431,456]" pageId="39" pageNumber="40">
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(
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<figureCitation id="13512A2AA008FFCFC2CDFE5DFC071CC8" box="[901,995,431,456]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">
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<smallCapsWord id="8D33A073A008FFCFC2CDFE5DFC5F1CC7" baselines="450,450" box="[901,955,431,456]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="39" pageNumber="40">FIGS</smallCapsWord>
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3O
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</figureCitation>
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,
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||||
<figureCitation id="13512A2AA008FFCFC2B8FE5DFBD51CC8" box="[1008,1073,431,456]" captionStart="Figure 4" captionStartId="40.[145,226,1590,1612]" captionText="Figure 4. Live images of species of Tubulophilinopsis gen. nov., with details of the morphology of the anterior part of the cephalic shield. A, T. lineolata (Thailand, photography by Nick Chaloklum). B, T. reticulata (Thailand, photography by Nick Chaloklum), C, T. pilsbryi (Thailand, photography by Nick Chaloklum). D, E, detail of the cephalic hump of the type species T. pilsbryi (D, Kwajalein Atoll, photography by S. Johnson and J. Johnson; E, East Timor, photography by Brian Francisco). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">4A–D</figureCitation>
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,
|
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<figureCitation id="13512A2AA008FFCFC576FE5DFB1C1CC7" box="[1086,1272,431,456]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">
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5B, E, I;
|
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</figureCitation>
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1
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<smallCapsWord id="8D33A073A008FFCFC45CFE41FAA71CC7" baselines="450" box="[1300,1347,435,455]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="39" pageNumber="40">AND</smallCapsWord>
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2)
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</paragraph>
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<paragraph id="8BD536AFA008FFCFC273FE2BFBEB1F0C" blockId="39.[827,1443,472,954]" pageId="39" pageNumber="40">
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<emphasis id="B91EEABDA008FFCFC273FE2BFC2D1CED" box="[827,969,472,494]" italics="true" pageId="39" pageNumber="40">
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<typeStatus id="54D1880DA008FFCFC273FE2BFC941CEE" box="[827,880,473,494]" pageId="39" pageNumber="40">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C6A4D2CA008FFCFC286FE2AFAC81CED" authority="Eliot, 1900" authorityName="Eliot" authorityYear="1900" box="[974,1324,472,494]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pilsbryi">
|
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<emphasis id="B91EEABDA008FFCFC286FE2AFB4F1CED" box="[974,1195,472,493]" italics="true" pageId="39" pageNumber="40">Philinopsis pilsbryi</emphasis>
|
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Eliot, 1900
|
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</taxonomicName>
|
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. By subsequent designation.
|
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</paragraph>
|
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<emphasis id="B91EEABDA008FFCFC273FDE4FC561F2B" box="[827,946,534,555]" italics="true" pageId="39" pageNumber="40">Diagnosis:</emphasis>
|
||||
Living animals up to
|
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<quantity id="4C929B4AA008FFCFC5EEFDE7FB171F2B" box="[1190,1267,533,555]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="40" unit="mm" value="30.0">30 mm</quantity>
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in length. Body elongated, oval, reduced sensory bristles, not visible to the naked eye; cephalic shield well differentiated from the body, with frontal elevation, crest-like (autapomorphy); eyes frontal, conspicuous, inserted in small circular unpigmented periocular areas (autapomorphy); posterior shield with symmetrical, rounded, short caudal lobes (
|
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<figureCitation id="13512A2AA008FFCFC2FCFD1EFBF41E01" box="[948,1040,748,770]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">Figs 3O</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA008FFCFC556FD1EFBC81E01" box="[1054,1068,748,769]" captionStart="Figure 4" captionStartId="40.[145,226,1590,1612]" captionText="Figure 4. Live images of species of Tubulophilinopsis gen. nov., with details of the morphology of the anterior part of the cephalic shield. A, T. lineolata (Thailand, photography by Nick Chaloklum). B, T. reticulata (Thailand, photography by Nick Chaloklum), C, T. pilsbryi (Thailand, photography by Nick Chaloklum). D, E, detail of the cephalic hump of the type species T. pilsbryi (D, Kwajalein Atoll, photography by S. Johnson and J. Johnson; E, East Timor, photography by Brian Francisco). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="39" pageNumber="40">4</figureCitation>
|
||||
). Shell variable, convex-dilated with open whorl (
|
||||
<figureCitation id="13512A2AA008FFCFC540FCF8FBBF1E20" box="[1032,1115,778,800]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">Fig. 5B</figureCitation>
|
||||
), convex-concealed (
|
||||
<figureCitation id="13512A2AA008FFCFC400FCF8FA7E1E20" box="[1352,1434,778,800]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">Fig. 5E</figureCitation>
|
||||
) or spoon-shaped with prolonged and semi-wide open whorl (
|
||||
<figureCitation id="13512A2AA008FFCFC2DCFCBAFC001E5E" box="[916,996,840,862]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="39" pageNumber="40">Fig. 5I</figureCitation>
|
||||
). Buccal bulb tubular, long, muscular (autapomorphy). Penial papilla conical; prostate formed by a long narrow duct, not coiled (autapomorphy) (
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC234FC56FBCB1EB9" author="Rudman WB" box="[892,1071,932,954]" pageId="39" pageNumber="40" pagination="381 - 399" refId="ref29205" refString="Rudman WB. 1972 a. A comparative study of the genus philinopsis Pease 1860 (Aglajidae, Opisthobranchia). Pacific Science 26: 381-399." type="journal article" year="1972">Rudman, 1972a</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFFB4B5EA008FFCFC574FC56FB3B1EB9" author="Gosliner TM" box="[1084,1247,932,954]" pageId="39" pageNumber="40" pagination="325 - 360" refId="ref27454" refString="Gosliner TM. 1980. Systematics and phylogeny of the aglajidae (Opisthobranchia: Mollusca). Zoological Journal of the Linnaean Society 68: 325-360." type="journal article" year="1980">Gosliner, 1980</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
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<materialsCitation id="3B023CF2A008FFCFC273FC21FB5D1EE8" box="[827,1209,978,1000]" country="Fiji" location="Island" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Rotuma" typeStatus="holotype">
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<typeStatus id="54D1880DA008FFCFC273FC21FC941EE8" box="[827,880,979,1000]" pageId="39" pageNumber="40">Type</typeStatus>
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locality:
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</emphasis>
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<collectingRegion id="49AEF84DA008FFCFC29FFC21FBD51EE8" box="[983,1073,979,1000]" country="Fiji" name="Rotuma" pageId="39" pageNumber="40">Rotuma</collectingRegion>
|
||||
<location id="8EB56074A008FFCFC571FC20FB601EE8" LSID="urn:lsid:plazi:treatment:03C387B9A008FFCFC2CFFE7BFB961954:8EB56074A008FFCFC571FC20FB601EE8" box="[1081,1156,978,1000]" country="Fiji" name="Island" pageId="39" pageNumber="40" stateProvince="Rotuma">Island</location>
|
||||
,
|
||||
<collectingCountry id="F37D763FA008FFCFC5C6FC20FB5D1EE8" box="[1166,1209,978,1000]" name="Fiji" pageId="39" pageNumber="40">Fiji</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA008FFCFC273FBF3FB961954" blockId="39.[827,1442,1025,1108]" pageId="39" pageNumber="40">
|
||||
<emphasis id="B91EEABDA008FFCFC273FBF3FC201916" box="[827,964,1025,1046]" italics="true" pageId="39" pageNumber="40">Etymology:</emphasis>
|
||||
The genus name (Lat.
|
||||
<emphasis id="B91EEABDA008FFCFC5B8FBF3FADA1916" box="[1264,1342,1025,1046]" italics="true" pageId="39" pageNumber="40">tubule</emphasis>
|
||||
= tube) refers to the tubular buccal bulb present in all species included in this new genus.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
120
data/03/C3/87/03C387B9A00AFFCDC00EFDD9FD0B1806.xml
Normal file
120
data/03/C3/87/03C387B9A00AFFCDC00EFDD9FD0B1806.xml
Normal file
|
|
@ -0,0 +1,120 @@
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|||
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<mods:title id="DD04C726EE443AE655EAB5FC27E9CBBA">Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification</mods:title>
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<mods:namePart id="555039A06D319D293781D8F6930183B4">Zamora-Silva, Andrea</mods:namePart>
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<mods:roleTerm id="E15A7AE84DF37E039AD169D459C8534F">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="FD617EB0DB060E0D2767A4A80879DFF3">Malaquias, Manuel António E.</mods:namePart>
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<treatment id="03C387B9A00AFFCDC00EFDD9FD0B1806" ID-DOI="http://doi.org/10.5281/zenodo.14813120" ID-Zenodo-Dep="14813120" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC00EFDD9FD0B1806" httpUri="http://treatment.plazi.org/id/03C387B9A00AFFCDC00EFDD9FD0B1806" lastPageNumber="38" pageId="37" pageNumber="38">
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<paragraph id="8BD536AFA00AFFCDC00EFDD9FD831F43" blockId="37.[326,615,555,580]" box="[326,615,555,580]" pageId="37" pageNumber="38">
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<heading id="D09D81C3A00AFFCDC00EFDD9FD831F43" box="[326,615,555,580]" centered="true" fontSize="9" level="2" pageId="37" pageNumber="38" reason="2">
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|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC0D6FDDEFE0D1F42" ID-CoL="7NTCC" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[414,489,556,579]" class="Gastropoda" family="Aglajidae" genus="Biuve" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||||
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|
||||
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|
||||
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|
||||
.
|
||||
<smallCapsWord id="8D33A073A00AFFCDC367FDDDFD841F42" baselines="573" box="[559,608,559,578]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="37" pageNumber="38">NOV</smallCapsWord>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3706524A00AFFCDC078FDA6FD0B1806" pageId="37" pageNumber="38" type="description">
|
||||
<paragraph id="8BD536AFA00AFFCDC078FDA6FD991F69" blockId="37.[304,637,596,618]" box="[304,637,596,618]" pageId="37" pageNumber="38">
|
||||
(
|
||||
<figureCitation id="13512A2AA00AFFCDC070FDA6FE751F69" box="[312,401,596,617]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">
|
||||
<smallCapsWord id="8D33A073A00AFFCDC070FDA6FE881F69" baselines="612,613" box="[312,364,596,617]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Figs" pageId="37" pageNumber="38">FIGS</smallCapsWord>
|
||||
3B
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC0D6FDA6FDF31F69" box="[414,535,596,618]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">
|
||||
5I;
|
||||
<smallCapsWord id="8D33A073A00AFFCDC08BFDA6FDF31F69" baselines="612,613" box="[451,535,596,617]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Tables" pageId="37" pageNumber="38">TABLES</smallCapsWord>
|
||||
</figureCitation>
|
||||
1
|
||||
<smallCapsWord id="8D33A073A00AFFCDC37BFDAAFD851F69" baselines="613" box="[563,609,600,617]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="and" pageId="37" pageNumber="38">AND</smallCapsWord>
|
||||
2)
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC1EBFD89FE391FAF" blockId="37.[163,778,634,687]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC1EBFD89FED31F90" box="[163,311,635,656]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC1EBFD89FF3D1F90" box="[163,217,635,656]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC077FD88FCE21F90" authority="Baba, 1938" authorityName="Baba" authorityYear="1938" box="[319,774,634,656]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="fulvipunctata">
|
||||
<emphasis id="B91EEABDA00AFFCDC077FD88FD9D1F8F" box="[319,633,634,655]" italics="true" pageId="37" pageNumber="38">Chelidonura fulvipunctata</emphasis>
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC3C9FD88FCE21F90" author="Baba K" box="[641,774,634,656]" pageId="37" pageNumber="38" pagination="1 - 50" refId="ref26605" refString="Baba K. 1938. Opisthobranchia of the Ryukyu (Okinawa) Islands. Journal of the Faculty of Agriculture 5: 1-50." type="journal article" year="1938">Baba, 1938</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. By subsequent designation.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC1EBFD3AFE43196C" blockId="37.[163,778,712,1132]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC1EBFD3AFEC31FDD" box="[163,295,712,733]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA00AFFCDC367FD3AFD601FDD" box="[559,644,712,733]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="37" pageNumber="38" unit="mm" value="30.0">30 mm</quantity>
|
||||
in length. Body narrow, elongate, flat; cephalic shield larger than the posterior shield; anterior edge of cephalic shield trilobed, lobes rounded; posterior shield with two asymmetrical caudal lobes, right one short, nearly inconspicuous, and rounded, left prolonged, thin, flagellum-like. Whitish W-shaped mark on cephalic region (autapomorphy) (
|
||||
<figureCitation id="13512A2AA00AFFCDC319FC6CFD491EB4" box="[593,685,926,948]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">Fig. 3B</figureCitation>
|
||||
). Shell spoon-shaped with prolonged, semi-wide open whorl, solid, yellowish-white, oldest region brown (
|
||||
<figureCitation id="13512A2AA00AFFCDC1E4FC08FF1E1910" box="[172,250,1018,1040]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">Fig. 5I</figureCitation>
|
||||
). Buccal bulb reduced; penial papilla cylindrical, longer than prostate; prostate with single lobe, short and rounded (
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC0BBFBCAFD60194D" author="Baba K" box="[499,644,1080,1102]" pageId="37" pageNumber="38" pagination="1 - 50" refId="ref26605" refString="Baba K. 1938. Opisthobranchia of the Ryukyu (Okinawa) Islands. Journal of the Faculty of Agriculture 5: 1-50." type="journal article" year="1938">Baba, 1938</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC3D0FBCAFF38196C" author="Rudman WB" pageId="37" pageNumber="38" pagination="185 - 212" refId="ref29366" refString="Rudman WB. 1974. A comparison of Chelidonura, Navanax and Aglaja with other genera of the Aglajidae (Opisthobranchia: Gastropoda). Zoological Journal of the Linnaean Society 54: 185-212." type="journal article" year="1974">Rudman, 1974</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC1A3FBA4FE72196C" author="Gosliner TM" box="[235,406,1110,1132]" pageId="37" pageNumber="38" refId="ref27515" refString="Gosliner TM. 1987. Nudibranchs of Southern Africa: a guide to opisthobranch molluscs of Southern Africa. Monterey: Sea Challengers." type="book" year="1987">Gosliner, 1987</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC1EBFB74FE58199B" blockId="37.[163,444,1157,1179]" box="[163,444,1157,1179]" pageId="37" pageNumber="38">
|
||||
<materialsCitation id="3B023CF2A00AFFCDC1EBFB74FE5C199B" box="[163,440,1157,1179]" country="Japan" location="Kii" pageId="37" pageNumber="38" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B91EEABDA00AFFCDC1EBFB74FEDC199A" box="[163,312,1157,1179]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC1EBFB74FF3C199B" box="[163,216,1158,1179]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
locality:
|
||||
</emphasis>
|
||||
<location id="8EB56074A00AFFCDC077FB77FE80199B" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC00EFDD9FD0B1806:8EB56074A00AFFCDC077FB77FE80199B" box="[319,356,1157,1179]" country="Japan" name="Kii" pageId="37" pageNumber="38">Kii</location>
|
||||
,
|
||||
<collectingCountry id="F37D763FA00AFFCDC027FB74FE5C199B" box="[367,440,1158,1179]" name="Japan" pageId="37" pageNumber="38">Japan</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC1EBFB46FD0B1806" blockId="37.[163,778,1204,1287]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC1EBFB46FEC219C9" box="[163,294,1204,1225]" italics="true" pageId="37" pageNumber="38">Etymology:</emphasis>
|
||||
The name stems from the presence of a W-shape mark on the head region of animals (Lat.
|
||||
<emphasis id="B91EEABDA00AFFCDC1EBFB03FF5D1806" box="[163,185,1265,1286]" italics="true" pageId="37" pageNumber="38">bi</emphasis>
|
||||
= two or double; ‘uve’ Spanish name for letter ‘v’).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
122
data/03/C3/87/03C387B9A00AFFCDC04EFAA8FC761CA3.xml
Normal file
122
data/03/C3/87/03C387B9A00AFFCDC04EFAA8FC761CA3.xml
Normal file
|
|
@ -0,0 +1,122 @@
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|||
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<mods:title id="075788F93220806EDECB8EC745FB42D0">Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification</mods:title>
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.
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(
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<figureCitation id="13512A2AA00AFFCDC07EFA8DFE751894" box="[310,401,1407,1428]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">
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3C
|
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</figureCitation>
|
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,
|
||||
<figureCitation id="13512A2AA00AFFCDC0D5FA8CFDFD1893" box="[413,537,1406,1428]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">
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5J;
|
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</figureCitation>
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1
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2)
|
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</paragraph>
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<paragraph id="8BD536AFA00AFFCDC1EBFA54FE3918DA" blockId="37.[163,779,1445,1498]" pageId="37" pageNumber="38">
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<typeStatus id="54D1880DA00AFFCDC1EBFA54FF3D18BB" box="[163,217,1446,1467]" pageId="37" pageNumber="38">Type</typeStatus>
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species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC076FA57FCE318BB" authority="Pruvot-Fol, 1953" authorityName="Pruvot-Fol" authorityYear="1953" box="[318,775,1445,1467]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="africana">
|
||||
<emphasis id="B91EEABDA00AFFCDC076FA57FDDE18BA" box="[318,570,1445,1466]" italics="true" pageId="37" pageNumber="38">Chelidonura africana</emphasis>
|
||||
Pruvot-Fol, 1953
|
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</taxonomicName>
|
||||
. By subsequent designation.
|
||||
</paragraph>
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<paragraph id="8BD536AFA00AFFCDC1EBFA01FE501A59" blockId="37.[163,778,1522,1881]" pageId="37" pageNumber="38">
|
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<emphasis id="B91EEABDA00AFFCDC1EBFA01FEC31B08" box="[163,295,1523,1544]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA00AFFCDC367FA01FD601B08" box="[559,644,1523,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="mm" value="10.0">10 mm</quantity>
|
||||
in length. Body elongate, blunt, flat; cephalic and posterior shields nearly equal in length; anterior edge of cephalic shield bilobed, lobes rounded; posterior shield with two asymmetrical caudal lobes, right one minute, rounded, left triangular (
|
||||
<figureCitation id="13512A2AA00AFFCDC32AF97EFD531BA1" box="[610,695,1676,1698]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">Fig. 3C</figureCitation>
|
||||
). Shell calcified, white or translucent, apex with external fringe, projections at protoconch level (probably an autapomorphy) (
|
||||
<figureCitation id="13512A2AA00AFFCDC0D1F91AFE091BFE" box="[409,493,1768,1790]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">Fig. 5J</figureCitation>
|
||||
). Buccal bulb reduced; penial papilla conical, with two frontal lobes; prostate cylindrical, not bifurcated, longer than penis (
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC1E3F8B6FE401A59" author="Martinez E & Malaquias MAE & Cervera JL" box="[171,420,1860,1881]" pageId="37" pageNumber="38" pagination="349 - 353" refId="ref28277" refString="Martinez E, Malaquias MAE, Cervera JL. 2002. Chelidonura africana Pruvot-Fol, 1953 (Mollusca, Gastropoda): proposed designation of a neotype. Journal of Conchology 37: 349-353." type="journal article" year="2002">
|
||||
Martínez
|
||||
<emphasis id="B91EEABDA00AFFCDC068F8B6FE871A59" box="[288,355,1860,1881]" italics="true" pageId="37" pageNumber="38">et al.,</emphasis>
|
||||
2002
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FF34FB801DFA" blockId="37.[827,1442,197,250]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FF34FC301DDA" box="[827,980,197,219]" italics="true" pageId="37" pageNumber="38">Type locality:</emphasis>
|
||||
<materialsCitation id="3B023CF2A00AFFCDC293FF37FB841DFA" country="Spain" location="Tenerife Island" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Canarias" typeStatus="neotype">
|
||||
<location id="8EB56074A00AFFCDC293FF37FB691DDB" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC04EFAA8FC761CA3:8EB56074A00AFFCDC293FF37FB691DDB" box="[987,1165,197,219]" country="Spain" name="Tenerife Island" pageId="37" pageNumber="38" stateProvince="Canarias">Tenerife Island</location>
|
||||
,
|
||||
<collectingRegion id="49AEF84DA00AFFCDC5D2FF37FAA91DDB" box="[1178,1357,197,219]" country="Spain" name="Canarias" pageId="37" pageNumber="38">Canary Islands</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F37D763FA00AFFCDC412FF37FA7B1DDB" box="[1370,1439,197,219]" name="Spain" pageId="37" pageNumber="38">Spain</collectingCountry>
|
||||
, after
|
||||
<typeStatus id="54D1880DA00AFFCDC230FF16FC351DF9" box="[888,977,228,249]" pageId="37" pageNumber="38" type="neotype">neotype</typeStatus>
|
||||
designation
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FEE0FC761CA3" blockId="37.[827,1443,274,419]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FEE0FC2C1C27" box="[827,968,274,295]" italics="true" pageId="37" pageNumber="38">Etymology:</emphasis>
|
||||
This genus is named after Yolanda Camacho-García for her studies on the phylogeny of
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC273FEA2FC4D1C66" authorityName="PILSBRY" authorityYear="1895" box="[827,937,336,358]" class="Gastropoda" family="Aglajidae" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="family">Aglajidae</taxonomicName>
|
||||
gastropods and for being the first to reveal the existence of this clade of exclusively Atlantic species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
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130
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<mods:roleTerm id="B024E00A37070DF0B51397E4516E9652">Author</mods:roleTerm>
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||||
<mods:namePart id="17F73FCDAC21F9AB5D1E34FD11623335">Malaquias, Manuel António E.</mods:namePart>
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<mods:title id="B08D7A2D92ADB21DE455866128CE62D3">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="875C9898CB50791A203E6A4DFD1DE719">
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<mods:date id="E33C47FE70A5DF1A165FD845850943DE">2018</mods:date>
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<mods:number id="CF95C2CDA257F0FAAB798E221C400244">183</mods:number>
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<treatment id="03C387B9A00AFFCDC28DFE2DFB47199C" ID-DOI="http://doi.org/10.5281/zenodo.14813128" ID-Zenodo-Dep="14813128" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC28DFE2DFB47199C" httpUri="http://treatment.plazi.org/id/03C387B9A00AFFCDC28DFE2DFB47199C" lastPageNumber="38" pageId="37" pageNumber="38">
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<taxonomicName id="4C6A4D2CA00AFFCDC554FE12FB7E1CF6" ID-CoL="7P8DJ" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[1052,1178,480,503]" class="Gastropoda" family="Aglajidae" genus="Mannesia" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus" status="gen. nov.">
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.
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.
|
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</emphasis>
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</heading>
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</paragraph>
|
||||
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<subSubSection id="C3706524A00AFFCDC2E3FDF4FB47199C" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BD536AFA00AFFCDC2E3FDF4FAD61F1F" blockId="37.[939,1330,518,543]" box="[939,1330,518,543]" pageId="37" pageNumber="38">
|
||||
(
|
||||
<figureCitation id="13512A2AA00AFFCDC2FBFDFAFBEA1F1D" box="[947,1038,520,541]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">
|
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<smallCapsWord id="8D33A073A00AFFCDC2FBFDFAFC0C1F1D" baselines="536,537" box="[947,1000,520,541]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Figs" pageId="37" pageNumber="38">FIGS</smallCapsWord>
|
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3E
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC553FDFAFBD81F1D" box="[1051,1084,520,542]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">5G</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC501FDFAFB281F1D" box="[1097,1228,518,543]" captionStart="Figure 6" captionStartId="44.[145,228,1691,1713]" captionTargetBox="[308,1263,199,1649]" captionTargetId="figure-27@44.[305,1265,195,1651]" captionTargetPageId="44" captionText="Figure 6. Scanning electron micrographs and macrophotographs of the shell. A, Navanax inermis (CNMO 3461). B, Nakamigawaia felis (ZMBN 84913). C, Nakamigawaia spiralis [NSMT-39805, ventral (upper) and dorsal (lower) views]. D, Odontoglaja guamensis [ZMBN 95961; dorsal (left) and ventral (right) views]. E, Spinoaglaja orientalis (ZMBN 95999). F, Mariaglaja sandrana (ZMBN 94028). G, Mannesia sabadiega (photography by Manuel Caballer). Scale bars: 200 μm." pageId="37" pageNumber="38">
|
||||
6G;
|
||||
<smallCapsWord id="8D33A073A00AFFCDC53FFDFAFB281F1D" baselines="536,537" box="[1143,1228,520,541]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Tables" pageId="37" pageNumber="38">TABLES</smallCapsWord>
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||||
</figureCitation>
|
||||
1
|
||||
<smallCapsWord id="8D33A073A00AFFCDC5A0FDFEFAF21F1D" baselines="537" box="[1256,1302,524,541]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="and" pageId="37" pageNumber="38">AND</smallCapsWord>
|
||||
2)
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FDDDFAC91F63" blockId="37.[827,1442,558,611]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FDDDFC351F44" box="[827,977,559,580]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC273FDDDFC961F44" box="[827,882,559,580]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC292FDDCFC0C1F63" authority="Ortea, Moro & Espinosa, 1996" authorityName="Ortea, Moro & Espinosa" authorityYear="1996" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="sabadiega">
|
||||
<emphasis id="B91EEABDA00AFFCDC292FDDCFB091F43" box="[986,1261,558,579]" italics="true" pageId="37" pageNumber="38">Chelidonura sabadiega</emphasis>
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC5BEFDDDFC0C1F63" author="Ortea J & Moro L & Espinosa J" pageId="37" pageNumber="38" pagination="215 - 229" refId="ref28604" refString="Ortea J, Moro L, Espinosa J. 1996. Descripcion de dos nuevas especies del genero Chelidonura A. Adams, 1850 (Opisthobranchia, Cephalaspidea, Aglajidae) colectadas en la isla del Hierro. Estudio comparado con C. africana Pruvot-Fol, 1953. Revista de la Academia Canaria de Ciencias 7: 215-229." type="journal article" year="1996">Ortea, Moro & Espinosa, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. By subsequent designation.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FD8EFC221901" blockId="37.[827,1444,636,1026]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FD8EFC571F91" box="[827,947,636,657]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA00AFFCDC5D3FD8EFB0F1F91" box="[1179,1259,636,658]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="mm" value="15.0">15 mm</quantity>
|
||||
in length. Body elongated, narrow; cephalic shield bilobed, lobes quadrangular with straight sides; posterior shield about half length of cephalic shield, with two asymmetrical caudal lobes, the left larger, flat, tail-like, the right minute, triangular, acuminate (
|
||||
<figureCitation id="13512A2AA00AFFCDC5ACFCE7FAD21E2A" box="[1252,1334,789,811]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">Fig. 3E</figureCitation>
|
||||
). Shell of amber- or chestnut colour, spoon-shaped with pointed shoulder protruding slightly above the apex (
|
||||
<figureCitation id="13512A2AA00AFFCDC40CFCA0FA781E68" box="[1348,1436,850,872]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">Figs 5G</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC273FC83FCBF1E87" box="[827,859,881,903]" captionStart="Figure 6" captionStartId="44.[145,228,1691,1713]" captionTargetBox="[308,1263,199,1649]" captionTargetId="figure-27@44.[305,1265,195,1651]" captionTargetPageId="44" captionText="Figure 6. Scanning electron micrographs and macrophotographs of the shell. A, Navanax inermis (CNMO 3461). B, Nakamigawaia felis (ZMBN 84913). C, Nakamigawaia spiralis [NSMT-39805, ventral (upper) and dorsal (lower) views]. D, Odontoglaja guamensis [ZMBN 95961; dorsal (left) and ventral (right) views]. E, Spinoaglaja orientalis (ZMBN 95999). F, Mariaglaja sandrana (ZMBN 94028). G, Mannesia sabadiega (photography by Manuel Caballer). Scale bars: 200 μm." pageId="37" pageNumber="38">6G</figureCitation>
|
||||
). Radula present, with few rows of teeth (8 × 1.0.1) (autapomorphy); one single lateral tooth on each side, rachidian tooth absent; lateral teeth hook-shaped. Prostate cylindrical, unilobed (
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC5EFFC3FFAB51EE2" author="Rudman WB" box="[1191,1361,973,995]" pageId="37" pageNumber="38" pagination="89 - 107" refId="ref29403" refString="Rudman WB. 1978. A new species and genus of the Aglajidae and the evolution of the Philinacean opisthobranch molluscs. Zoological Journal of the Linnaean Society 62: 89-107." type="journal article" year="1978">Rudman, 1978</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC417FC3FFC531902" author="Ortea J & Moro L & Espinosa J" pageId="37" pageNumber="38" pagination="215 - 229" refId="ref28604" refString="Ortea J, Moro L, Espinosa J. 1996. Descripcion de dos nuevas especies del genero Chelidonura A. Adams, 1850 (Opisthobranchia, Cephalaspidea, Aglajidae) colectadas en la isla del Hierro. Estudio comparado con C. africana Pruvot-Fol, 1953. Revista de la Academia Canaria de Ciencias 7: 215-229." type="journal article" year="1996">
|
||||
Ortea
|
||||
<emphasis id="B91EEABDA00AFFCDC273FC1EFC9D1901" box="[827,889,1004,1025]" italics="true" pageId="37" pageNumber="38">et al.,</emphasis>
|
||||
1996
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FBE9FA471930" blockId="37.[827,1443,1050,1072]" box="[827,1443,1050,1072]" pageId="37" pageNumber="38">
|
||||
<materialsCitation id="3B023CF2A00AFFCDC273FBE9FA441930" box="[827,1440,1050,1072]" country="Spain" location="El Hierro Island" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Canarias" typeStatus="holotype">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FBE9FC34192F" box="[827,976,1050,1072]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC273FBE9FC941930" box="[827,880,1051,1072]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
locality:
|
||||
</emphasis>
|
||||
<location id="8EB56074A00AFFCDC29FFBE8FB701930" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC28DFE2DFB47199C:8EB56074A00AFFCDC29FFBE8FB701930" box="[983,1172,1050,1072]" country="Spain" name="El Hierro Island" pageId="37" pageNumber="38" stateProvince="Canarias">El Hierro Island</location>
|
||||
,
|
||||
<collectingRegion id="49AEF84DA00AFFCDC5D7FBE9FAAB1930" box="[1183,1359,1050,1072]" country="Spain" name="Canarias" pageId="37" pageNumber="38">Canary Islands</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="F37D763FA00AFFCDC412FBE8FA441930" box="[1370,1440,1050,1072]" name="Spain" pageId="37" pageNumber="38">Spain</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FBBBFB47199C" blockId="37.[827,1443,1097,1180]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FBBBFC58195E" box="[827,956,1097,1118]" italics="true" pageId="37" pageNumber="38">Etymology:</emphasis>
|
||||
This genus is dedicated to John Kenneth
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC273FB9AFC7C197D" box="[827,920,1128,1149]" family="Mannes" pageId="37" pageNumber="38" rank="family">Mannes</taxonomicName>
|
||||
, husband of Andrea Zamora-Silva, and to the members of the family
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC509FB75FB7B199C" box="[1089,1183,1159,1180]" family="Mannes" pageId="37" pageNumber="38" rank="family">Mannes</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
121
data/03/C3/87/03C387B9A00AFFCDC2F0FB2BFA971A58.xml
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121
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|
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@ -0,0 +1,121 @@
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<mods:namePart id="DC7E3A05E61A69957619916802DCA478">Zamora-Silva, Andrea</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name id="AB3DA5BDBF0CF648575FFBAD7C575EB0" type="personal">
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<mods:role id="46DB63681BC9AE628D3A8B02BC7E7554">
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<mods:roleTerm id="1BF7D16D1E42AA2C5E0EEAB0AA101433">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="9DD5C258FE1B0FDF8CE9C05B43E0DBAE">Malaquias, Manuel António E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="6D9747140C8DB68041643375F7F95A0E">text</mods:typeOfResource>
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<mods:titleInfo id="B67251ECBACFAFFCF188F3A156A420EC">
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<mods:title id="1338B0E9743D8B27888C280857C32686">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
|
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<mods:part id="1D34923E7C6FA567F5E8487408B5637B">
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<mods:date id="10945F235D1E09C6EED21D61D8DD13A5">2018</mods:date>
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<mods:detail id="7AA49EEEF80DE9FE989C8BA9AD8A42C3" type="volume">
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<mods:number id="F3A0516BBBF28BB3C936A7C59CD87F60">183</mods:number>
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<mods:classification id="A58086B57D1D63947D27A4DF3102BA00">journal article</mods:classification>
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<mods:identifier id="1F9ADE60AE7A691C11EF4E5111C9CC6B" type="ISSN">0024-4082</mods:identifier>
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</mods:mods>
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<treatment id="03C387B9A00AFFCDC2F0FB2BFA971A58" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC2F0FB2BFA971A58" httpUri="http://treatment.plazi.org/id/03C387B9A00AFFCDC2F0FB2BFA971A58" lastPageNumber="38" pageId="37" pageNumber="38">
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<subSubSection id="C3706524A00AFFCDC2F0FB2BFAC219F1" box="[952,1318,1241,1266]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BD536AFA00AFFCDC2F0FB2BFAC219F1" blockId="37.[952,1318,1241,1266]" box="[952,1318,1241,1266]" pageId="37" pageNumber="38">
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<heading id="D09D81C3A00AFFCDC2F0FB2BFAC219F1" box="[952,1318,1241,1266]" centered="true" fontSize="9" level="2" pageId="37" pageNumber="38" reason="2">
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<taxonomicName id="4C6A4D2CA00AFFCDC547FB2BFB4C19EF" ID-CoL="7P8GB" authorityName="Zamora-Silva & Malaquias" authorityYear="2018" box="[1039,1192,1241,1264]" class="Gastropoda" family="Aglajidae" genus="Mariaglaja" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus" status="gen. nov.">
|
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<smallCapsWord id="8D33A073A00AFFCDC547FB2BFB4C19EF" baselines="1258,1258" box="[1039,1192,1241,1264]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Mariaglaja" pageId="37" pageNumber="38">MARIAGLAJA</smallCapsWord>
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</taxonomicName>
|
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<emphasis id="B91EEABDA00AFFCDC5E7FB2EFAC219F1" bold="true" box="[1199,1318,1241,1265]" pageId="37" pageNumber="38">
|
||||
<taxonomicNameLabel id="A22D57C6A00AFFCDC5E7FB2EFAC219F1" box="[1199,1318,1241,1265]" pageId="37" pageNumber="38" rank="genus">
|
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<smallCapsWord id="8D33A073A00AFFCDC5E7FB2EFB0419EF" baselines="1258" box="[1199,1248,1244,1263]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="gen" pageId="37" pageNumber="38">GEN</smallCapsWord>
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.
|
||||
<smallCapsWord id="8D33A073A00AFFCDC5A6FB2EFAFB19EF" baselines="1258" box="[1262,1311,1244,1263]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="37" pageNumber="38">NOV</smallCapsWord>
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||||
.
|
||||
</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3706524A00AFFCDC2FAFB0DFA971A58" pageId="37" pageNumber="38" type="description">
|
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<paragraph id="8BD536AFA00AFFCDC2FAFB0DFACF1818" blockId="37.[946,1323,1279,1304]" box="[946,1323,1279,1304]" pageId="37" pageNumber="38">
|
||||
(
|
||||
<figureCitation id="13512A2AA00AFFCDC2F2FAF0FBF71816" box="[954,1043,1281,1303]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">
|
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<smallCapsWord id="8D33A073A00AFFCDC2F2FAF0FC0B1816" baselines="1298,1298" box="[954,1007,1282,1303]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Figs" pageId="37" pageNumber="38">FIGS</smallCapsWord>
|
||||
3F
|
||||
</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC569FAF3FBDC1817" box="[1057,1080,1281,1303]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">5I</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC50DFAF3FB211816" box="[1093,1221,1279,1304]" captionStart="Figure 6" captionStartId="44.[145,228,1691,1713]" captionTargetBox="[308,1263,199,1649]" captionTargetId="figure-27@44.[305,1265,195,1651]" captionTargetPageId="44" captionText="Figure 6. Scanning electron micrographs and macrophotographs of the shell. A, Navanax inermis (CNMO 3461). B, Nakamigawaia felis (ZMBN 84913). C, Nakamigawaia spiralis [NSMT-39805, ventral (upper) and dorsal (lower) views]. D, Odontoglaja guamensis [ZMBN 95961; dorsal (left) and ventral (right) views]. E, Spinoaglaja orientalis (ZMBN 95999). F, Mariaglaja sandrana (ZMBN 94028). G, Mannesia sabadiega (photography by Manuel Caballer). Scale bars: 200 μm." pageId="37" pageNumber="38">
|
||||
6F;
|
||||
<smallCapsWord id="8D33A073A00AFFCDC539FAF0FB211816" baselines="1298,1298" box="[1137,1221,1282,1303]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Tables" pageId="37" pageNumber="38">TABLES</smallCapsWord>
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</figureCitation>
|
||||
1
|
||||
<smallCapsWord id="8D33A073A00AFFCDC5A9FAF7FAEB1816" baselines="1298" box="[1249,1295,1285,1302]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="and" pageId="37" pageNumber="38">AND</smallCapsWord>
|
||||
2)
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FADBFBAB185C" blockId="37.[827,1442,1320,1372]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FADBFC34183D" box="[827,976,1320,1342]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC273FADBFC96183E" box="[827,882,1321,1342]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C6A4D2CA00AFFCDC291FADAFA92183E" authority="Gosliner, 2015" authorityName="Gosliner" authorityYear="2015" box="[985,1398,1320,1342]" class="Gastropoda" family="Aglajidae" genus="Chelidonura" kingdom="Animalia" order="Cephalaspidea" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="alexisi">
|
||||
<emphasis id="B91EEABDA00AFFCDC291FADAFB26183D" box="[985,1218,1320,1341]" italics="true" pageId="37" pageNumber="38">Chelidonura alexisi</emphasis>
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC582FADAFA92183E" author="Gosliner TM" box="[1226,1398,1320,1342]" pageId="37" pageNumber="38" pagination="191 - 205" refId="ref27563" refString="Gosliner TM. 2015. Three new species of aglajid Cephalaspidean mollusks from the tropical Indo-pacific of the Verde Island passage. Proceedings of the California Academy of Sciences 6: 191-205." type="journal article" year="2015">Gosliner, 2015</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. By subsequent designation.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273FA84FB601BDC" blockId="37.[827,1443,1397,1756]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273FA84FC57188B" box="[827,947,1398,1419]" italics="true" pageId="37" pageNumber="38">Diagnosis:</emphasis>
|
||||
Live animals up to
|
||||
<quantity id="4C929B4AA00AFFCDC5D3FA87FB0F188B" box="[1179,1259,1397,1419]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="mm" value="10.0">10 mm</quantity>
|
||||
in length. Body elongated-narrow; cephalic shield trilobed, middle lobe sometimes with small indent, lateral lobes rounded to triangular, expanded outwards, elongated, comprising two thirds of body length; posterior shield with asymmetrical caudal lobes, left one longer and fleshier, right lobe reduced, nearly invisible (
|
||||
<figureCitation id="13512A2AA00AFFCDC5DDF9DFFB001B42" box="[1173,1252,1581,1603]" captionStart="Figure 3" captionStartId="38.[144,222,1561,1583]" captionTargetBox="[208,1360,197,1520]" captionTargetId="figure-27@38.[207,1362,195,1521]" captionTargetPageId="38" captionText="Figure 3. Live images of the type species of all valid genera of Aglajidae. A, Aglaja tricolorata (Southern Portugal, photography by M.A. E. Malaquias). B, Biuve fulvipunctata (Kwajalein Atoll, photography by Jeanette Johnson). C, Camachoaglaja africana (São Miguel I., Azores, photography by M. A. E. Malaquias). D, Chelidonura hirundinina (Réunion Island, photography by Philibert Bidgrain). E, Mannesia sabadiega (Tenerife, Canary Is, photography by Manuel Caballer). F, Mariaglaja alexisi (the Philippines, photography by Sven Kahlbrock). G, Melanochlamys cylindrica (New Zealand, photography by Simon Franicevic). H, Nakamigawaia spiralis (the Philippines, photography by Philibert Bidgrain). I, Navanax inermis (California, USA, photography by Douglas Mason). J, Niparaya regiscorona (Maui, Hawaii, photography by Cory Pittman). K, Odontoglaja guamensis (Guam, photography by Jun Imamoto). L, Philinopsis speciosa (Réunion Island, photography by Philibert Bidgrain). M, Spinophallus coronata (Indo-West Pacific, photography by Nick Hobgood). N, Spinoaglaja petra (Bermuda, photography by Linda Ianiello). O, Tubulophilinopsis pilsbryi (Marshall Islands, photography by S. & J. Johnson). Copyright images are shared under the Creative Commons Attribution 4.0 International Public License." pageId="37" pageNumber="38">Fig. 3F</figureCitation>
|
||||
). Brown-reddish shell, calcified, spoon-shaped with acuminated shoul- der or with extended whorl (
|
||||
<figureCitation id="13512A2AA00AFFCDC5CFF998FB3C1B80" box="[1159,1240,1642,1664]" captionStart="Figure 5" captionStartId="40.[809,894,1555,1577]" captionTargetBox="[155,1384,193,1549]" captionTargetPageId="40" captionText="Figure 5. Shell morphology in Aglajidae. A, bulloid (adapted from Burn & Thompson, 1998). B, convex-dilated with open whorl. C, spiral evolute. D, concave-constricted with moderate close whorl. E, convex-concealed. F, spoonshaped with inward spiral whorl. G, spoon-shaped with acuminated or rounded whorl. H, spoon-shaped with conspicuous-broad and extended whorl. I, spoon-shaped with prolonged and semi-wide open whorl. J, modified version of the typical spoon-shaped with outward-pointed fringes of the protoconch and well-defined acuminated whorl. K, spoon-shaped with extended whorl and conspicuous upward-pointed spines on the protochonch." pageId="37" pageNumber="38">Figs 5I</figureCitation>
|
||||
,
|
||||
<figureCitation id="13512A2AA00AFFCDC5AEF998FAE01B80" box="[1254,1284,1642,1664]" captionStart="Figure 6" captionStartId="44.[145,228,1691,1713]" captionTargetBox="[308,1263,199,1649]" captionTargetId="figure-27@44.[305,1265,195,1651]" captionTargetPageId="44" captionText="Figure 6. Scanning electron micrographs and macrophotographs of the shell. A, Navanax inermis (CNMO 3461). B, Nakamigawaia felis (ZMBN 84913). C, Nakamigawaia spiralis [NSMT-39805, ventral (upper) and dorsal (lower) views]. D, Odontoglaja guamensis [ZMBN 95961; dorsal (left) and ventral (right) views]. E, Spinoaglaja orientalis (ZMBN 95999). F, Mariaglaja sandrana (ZMBN 94028). G, Mannesia sabadiega (photography by Manuel Caballer). Scale bars: 200 μm." pageId="37" pageNumber="38">6F</figureCitation>
|
||||
). Buccal bulb reduced, with microscopic labial spines on the lips. Penial papilla conical, with small chitinous hooks or spines (
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC2D9F934FBD71BDC" author="Gosliner TM" box="[913,1075,1734,1756]" pageId="37" pageNumber="38" refId="ref27515" refString="Gosliner TM. 1987. Nudibranchs of Southern Africa: a guide to opisthobranch molluscs of Southern Africa. Monterey: Sea Challengers." type="book" year="1987">Gosliner 1987</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFFB4B5EA00AFFCDC575F934FB9C1BDC" author="Gosliner TM" box="[1085,1144,1734,1756]" pageId="37" pageNumber="38" pagination="191 - 205" refId="ref27563" refString="Gosliner TM. 2015. Three new species of aglajid Cephalaspidean mollusks from the tropical Indo-pacific of the Verde Island passage. Proceedings of the California Academy of Sciences 6: 191-205." type="journal article" year="2015">2015</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273F904FAEE1A0B" blockId="37.[827,1290,1781,1803]" box="[827,1290,1781,1803]" pageId="37" pageNumber="38">
|
||||
<materialsCitation id="3B023CF2A00AFFCDC273F904FAE31A0B" box="[827,1287,1781,1803]" country="Philippines" location="The" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
|
||||
<emphasis id="B91EEABDA00AFFCDC273F904FC341A0A" box="[827,976,1781,1803]" italics="true" pageId="37" pageNumber="38">
|
||||
<typeStatus id="54D1880DA00AFFCDC273F904FC941A0B" box="[827,880,1782,1803]" pageId="37" pageNumber="38">Type</typeStatus>
|
||||
locality:
|
||||
</emphasis>
|
||||
<collectingRegion id="49AEF84DA00AFFCDC29FF904FBA01A0B" box="[983,1092,1782,1803]" country="Philippines" name="Batangas" pageId="37" pageNumber="38">Batangas</collectingRegion>
|
||||
,
|
||||
<location id="8EB56074A00AFFCDC507F907FB9F1A0B" LSID="urn:lsid:plazi:treatment:03C387B9A00AFFCDC2F0FB2BFA971A58:8EB56074A00AFFCDC507F907FB9F1A0B" box="[1103,1147,1781,1803]" country="Philippines" name="The" pageId="37" pageNumber="38" stateProvince="Batangas">The</location>
|
||||
<collectingCountry id="F37D763FA00AFFCDC5CAF907FAE31A0B" box="[1154,1287,1781,1803]" name="Philippines" pageId="37" pageNumber="38">Philippines</collectingCountry>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD536AFA00AFFCDC273F8D6FA971A58" blockId="37.[827,1442,1828,1880]" pageId="37" pageNumber="38">
|
||||
<emphasis id="B91EEABDA00AFFCDC273F8D6FC5B1A39" box="[827,959,1828,1849]" italics="true" pageId="37" pageNumber="38">Etymology:</emphasis>
|
||||
This genus is named after María Luisa Silva Villanueva, mother of Andrea Zamora-Silva.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
107
data/03/D1/87/03D187F7FFE1FFBD5F4135AFFDD049DA.xml
Normal file
107
data/03/D1/87/03D187F7FFE1FFBD5F4135AFFDD049DA.xml
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|
|
@ -0,0 +1,107 @@
|
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<mods:title id="84EDD3E88CC028EF178D9DEDB8EB6EC2">A new sauropod species from north-western Brazil: biomechanics and the radiation of Titanosauria (Sauropoda: Somphospondyli)</mods:title>
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<mods:namePart id="661D940AC73F17B26887E42548003DE3">Pereira, Paulo V L G C</mods:namePart>
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<mods:name id="70B4530F1F36AA602C94811A3894AD13" type="personal">
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<emphasis id="B90CEAF3FFE1FFBD5EAF35FCFF2448B0" box="[129,240,195,219]" italics="true" pageId="4" pageNumber="5">Etymology:</emphasis>
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‘
|
||||
<taxonomicName id="4C784D62FFE1FFBD5ED535FBFE9048B0" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[251,324,196,219]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
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’ is a goddess from Sumerian and Babylonian mythologies, often described as a dragon or a great serpent.
|
||||
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<emphasis id="B90CEAF3FFE1FFBD5EAF343DFF1E4972" box="[129,202,258,281]" italics="true" pageId="4" pageNumber="5">Tiamat</emphasis>
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</taxonomicName>
|
||||
is also identified as the mother of dragons (and other mythical creatures) and of all gods, alluding to the basal positioning of this new species into the group to which belong the largest animals that have ever walked on earth.
|
||||
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<emphasis id="B90CEAF3FFE1FFBD5EAF34A5FF3349DA" box="[129,231,410,433]" italics="true" pageId="4" pageNumber="5">Diagnosis:</emphasis>
|
||||
The same as for the species.
|
||||
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|
||||
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722
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ZooBank LSID for the species:
|
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urn: lsid:zoobank.org:act:
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|
||||
<emphasis id="B90CEAF3FFE1FFBD5EAF376FFF244A03" box="[129,240,592,616]" italics="true" pageId="4" pageNumber="5">Etymology:</emphasis>
|
||||
The specific epithet is in honour of Prof. Dr Valdeci dos Santos Júnior of Universidade Estadual do
|
||||
<collectingRegion id="49BCF803FFE1FFBD5C403750FF144ACD" country="Brazil" name="Rio Grande do Norte" pageId="4" pageNumber="5">Rio Grande do Norte</collectingRegion>
|
||||
, for the discovery of the fossil site and for his fundamental support during the fieldworks.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656AFFE1FFBD5EAF37D8FD514B36" pageId="4" pageNumber="5" type="materials_examined">
|
||||
<paragraph id="8BC736E1FFE1FFBD5EAF37D8FD514B36" blockId="4.[128,777,743,861]" pageId="4" pageNumber="5">
|
||||
<materialsCitation id="3B103CBCFFE1FFBD5EAF37D8FDFB4B36" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5EAF37D8FF354A94" box="[129,225,743,767]" italics="true" pageId="4" pageNumber="5">
|
||||
<typeStatus id="54C38843FFE1FFBD5EAF37D8FF084A94" box="[129,220,743,767]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
UFRJ-DG 636-R, UFRJ-DG 638-R, UFRJ-DG 606-R (anterior caudal vertebrae), UFRJ-DG 635-R, UFRJ-DG 591-R, UFRJ-DG 527-R, UFRJ-DG 704-R (middle caudal vertebrae), and UFRJ-DG 574-R (partial neural arch)
|
||||
</materialsCitation>
|
||||
(
|
||||
<figureCitation id="13432A64FFE1FFBD5C13367AFDA14B36" box="[573,629,837,861]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656AFFE1FFBD5EAF3640FE2E4D07" pageId="4" pageNumber="5" type="diagnosis">
|
||||
<paragraph id="8BC736E1FFE1FFBD5EAF3640FE2E4D07" blockId="4.[128,779,894,1388]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5EAF3640FF334BFD" box="[129,231,895,918]" italics="true" pageId="4" pageNumber="5">Diagnosis:</emphasis>
|
||||
A titanosaur sauropod diagnosed by four diagnostic features, all in the caudal vertebrae: marked accessory tuberosity dorsoventrally developed, located above the prezygapophyses in the posteriormost and in middle caudal vertebrae; deeply medioventral excavated articulation facets of the prezygapophysis and post-zygapophysis (see fig.07); presence of hypanthrum–hyposphene articulation; and short middle caudal centra with well-marked articular facets for the haemal arch. Additionally, some of the following combinations of characters assist the placement of
|
||||
<taxonomicName id="4C784D62FFE1FFBD5F4431A6FDD54CDB" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[362,513,1176,1200]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="valdecii">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5F4431A6FDD54CDB" box="[362,513,1176,1200]" italics="true" pageId="4" pageNumber="5">Tiamat valdecii</emphasis>
|
||||
</taxonomicName>
|
||||
as a titanosaurian, such as strong procoelous anterior caudal vertebrae with the apex of the posterior condyle positioned on the dorsal half of the centrum; wider than high centra which, progressively widen to the posterior vertebrae; pedicles of the neural arch attached to the anterior half of the centrum and presence of a ventral longitudinal hollow on the caudal vertebral series.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C362656AFFE1FFBF5FA130B2FA9D4E39" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
|
||||
<paragraph id="8BC736E1FFE1FFBD5FA130B2FE2F4DCD" blockId="4.[127,780,1421,1984]" box="[399,507,1421,1446]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5FA130B2FE2F4DCD" box="[399,507,1421,1446]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBD5EAF308DFB3748AC" blockId="4.[127,780,1421,1984]" lastBlockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||||
There are three anterior caudal vertebrae, four middle caudal vertebrae, and a neural spine (possibly an anterior one). The anterior caudal vertebrae (UFRJ-DG 636-R, UFRJ-DG 638-R, UFRJ-DG 606-R) are strongly procoelous, with subcircular outline, while one of the middle caudal vertebrae (UFRJ-DG 635-R) is slightly procoelous, and the other middle caudal (UFRJ-DG 591-R, UFRJ-DG 527-R, UFRJ-DG 704-R) are amphicoelous. This condition (transition of procoelous to slightly procoelous) is observed in
|
||||
<taxonomicName id="4C784D62FFE1FFBD5F353392FE614EAE" box="[283,437,1709,1733]" class="Reptilia" family="Titanosauridae" genus="Mendozasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5F353392FE614EAE" box="[283,437,1709,1733]" italics="true" pageId="4" pageNumber="5">Mendozasaurus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5FE63392FD4F4EAE" author="Gonzalez-Riga BJ" box="[456,667,1709,1733]" pageId="4" pageNumber="5" pagination="155 - 72" refId="ref12321" refString="Gonzalez-Riga BJ. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza Province, Argentina. Ameghiniana 2003; 40: 155 - 72." year="2003">González-Riga 2003</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5C873392FEF04E8F" author="Gonzalez Riga BJ & Mannion PD & Poropat SF" pageId="4" pageNumber="5" pagination="136 - 81" refId="ref12412" refString="Gonzalez Riga BJ, Mannion PD, Poropat SF et al. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 2018; 184: 136 - 81. https: // doi. org / 10.1093 / zoolinnean / zlx 103" year="2018">
|
||||
Gonzàlez Riga
|
||||
<emphasis id="B90CEAF3FFE1FFBD5E9933F2FF3D4E8F" box="[183,233,1740,1764]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
) and
|
||||
<taxonomicName id="4C784D62FFE1FFBD5F4833F2FD0C4E8F" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" box="[358,728,1740,1764]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5F4833F2FE034E8F" box="[358,471,1741,1764]" italics="true" pageId="4" pageNumber="5">Patagotitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5FC733F3FD184E8F" author="Carballido JL & Pol D & Otero A" box="[489,716,1740,1764]" pageId="4" pageNumber="5" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
|
||||
Carballido
|
||||
<emphasis id="B90CEAF3FFE1FFBD5C7033F2FD5E4E8F" box="[606,650,1740,1764]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||||
. 2017
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
; but differs from other taxa such as
|
||||
<taxonomicName id="4C784D62FFE1FFBD5F9433D3FF144F48" authority="(Mannion and Calvo 2011)" baseAuthorityName="Mannion and Calvo" baseAuthorityYear="2011" class="Reptilia" family="Titanosauridae" genus="Andesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5F9433D3FDF84F6F" box="[442,556,1772,1796]" italics="true" pageId="4" pageNumber="5">Andesaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5C1333D3FF604F48" author="Mannion PD & Calvo JO" pageId="4" pageNumber="5" refId="ref13041" refString="Mannion PD, Calvo JO. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albianearly Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 2011; 163: no - no. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00699. x" year="2011">Mannion and Calvo 2011</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D62FFE1FFBD5EE33234FE834F48" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[205,343,1803,1827]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5EE33234FE834F48" box="[205,343,1803,1827]" italics="true" pageId="4" pageNumber="5">Malawisaurus</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C784D62FFE1FFBD5FBC3234FDD14F48" box="[402,517,1803,1827]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5FBC3234FDD14F48" box="[402,517,1803,1827]" italics="true" pageId="4" pageNumber="5">Rukwatitan</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5C393234FD734F48" author="Gomani EM" box="[535,679,1803,1827]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5C9A3234FF384F29" author="Gorscak E & O'Connor PM & Stevens NJ" pageId="4" pageNumber="5" pagination="1133 - 54" refId="ref12567" refString="Gorscak E, O'Connor PM, Stevens NJ et al. The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the Middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 2014; 34: 1133 - 54. https: // doi. org / 10.1080 / 02724634.2014.845568" year="2014">
|
||||
Gorscak
|
||||
<emphasis id="B90CEAF3FFE1FFBD5EAF3214FF7F4F29" box="[129,171,1834,1858]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||||
. 2014
|
||||
</bibRefCitation>
|
||||
), since in these taxa, the anteriormost caudal vertebrae exhibits strongly procoelous condition, while the other anterior caudal vertebrae and, consequently the middle ones, turn to an amphiplatyan condition. The most anterior caudal vertebra recovered for the new Brazilian taxa also differs from
|
||||
<taxonomicName id="4C784D62FFE1FFBD5D1735AFFC3848C3" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[825,1004,144,168]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D1735AFFC3848C3" box="[825,1004,144,168]" italics="true" pageId="4" pageNumber="5">Mnyamawamtuka</emphasis>
|
||||
</taxonomicName>
|
||||
, since the African taxon presents a ‘mildprocoely’ (
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5D873590FB0648AC" author="Gorscak E & O'Connor PM" box="[937,1234,175,199]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBD5D7B35F0FA16498A" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||||
In lateral view, all vertebrae centra are anteroposteriorly short, bearing a concave lateral surface, which, in turn, is developed in the anteroposterior axis. The preserved neural arches are small and placed slightly anterior to the midpoint of the vertebral body. The vertebrae vary in height and width when observed on anterior and posterior views (
|
||||
<tableCitation id="C6FA035AFFE1FFBD5A203454FB8D49E8" box="[1038,1113,363,387]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||||
). The centrum of the more anterior caudal vertebra (UFRJ-DG 636-R) is taller than wide (
|
||||
<tableCitation id="C6FA035AFFE1FFBD5B4934B4FA6649C8" box="[1383,1458,395,419]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||||
), while it progressively dorsoventrally compresses in the middle caudal preserved (e.g.UFRJ-DG 704-R) (
|
||||
<figureCitation id="13432A64FFE1FFBD5AFF34F6FACF498A" box="[1233,1307,457,481]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4A</figureCitation>
|
||||
<figureCitation id="13432A64FFE1FFBD5B3534F6FA954989" box="[1307,1345,457,482]" captionStart="Figure 1" captionStartId="1.[114,179,1926,1950]" captionTargetBox="[117,1459,1429,1896]" captionTargetId="figure-639@1.[114,1461,1424,1898]" captionTargetPageId="1" captionText="Figure 1. Representation of the recovery process of the middle caudal vertebrae UFRJ-DG 591-R. A, absence of the left prezygapophysis in the vertebra; B, using the right prezygapophysis present in the UFRJ-DG 591-R, it was possible to infer the left prezygapophysis." figureDoi="http://doi.org/10.5281/zenodo.14764385" httpUri="https://zenodo.org/record/14764385/files/figure.png" pageId="4" pageNumber="5">1–A</figureCitation>
|
||||
<figureCitation id="13432A64FFE1FFBD5B6F34F6FA9A498A" box="[1345,1358,457,481]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="13432A64FFE1FFBD5B7634F6FAAC498A" box="[1368,1400,457,481]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">4G</figureCitation>
|
||||
<figureCitation id="13432A64FFE1FFBD5B5634F6FA71498A" box="[1400,1445,457,481]" captionStart="Figure 1" captionStartId="1.[114,179,1926,1950]" captionTargetBox="[117,1459,1429,1896]" captionTargetId="figure-639@1.[114,1461,1424,1898]" captionTargetPageId="1" captionText="Figure 1. Representation of the recovery process of the middle caudal vertebrae UFRJ-DG 591-R. A, absence of the left prezygapophysis in the vertebra; B, using the right prezygapophysis present in the UFRJ-DG 591-R, it was possible to infer the left prezygapophysis." figureDoi="http://doi.org/10.5281/zenodo.14764385" httpUri="https://zenodo.org/record/14764385/files/figure.png" pageId="4" pageNumber="5">1–G</figureCitation>
|
||||
<figureCitation id="13432A64FFE1FFBD5B8B34F6FA66498A" box="[1445,1458,457,481]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBD5D7B34D6FC654AB7" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||||
As in
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D8E34D6FBEA4A6B" box="[928,1086,489,512]" italics="true" pageId="4" pageNumber="5">Tastavinasaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5A7934D6FAE54A6A" author="Canudo JI & Royo-Torres R & Cuenca-Bescos G" box="[1111,1329,489,513]" pageId="4" pageNumber="5" pagination="712 - 31" refId="ref11762" refString="Canudo JI, Royo-Torres R, Cuenca-Bescos G. A new sauropod: Tastavinsaurus sanzi gen. et sp. nov. from the Early Cretaceous (Aptian) of Spain. Journal of Vertebrate Paleontology 2008; 28: 712 - 31. https: // doi. org / 10.1671 / 0272 - 4634 (2008) 28 [712: anstsg] 2.0. co; 2" type="journal article" year="2008">
|
||||
Canudo
|
||||
<emphasis id="B90CEAF3FFE1FFBD5A9634D6FB244A6A" box="[1208,1264,489,513]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2008
|
||||
</bibRefCitation>
|
||||
) and in the anteriormost caudal preserved of
|
||||
<taxonomicName id="4C784D62FFE1FFBD5A833737FC2B4A54" authority="(Gonzalez Riga et al. 2009)" baseAuthorityName="Gonzalez Riga" baseAuthorityYear="2009" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5A833737FA9E4A4B" box="[1197,1354,520,544]" italics="true" pageId="4" pageNumber="5">Malarguesaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B4C3737FC204A54" author="Gonzalez Riga BJ & Previtera E & Pirrone CA" pageId="4" pageNumber="5" pagination="135 - 48" refId="ref12351" refString="Gonzalez Riga BJ, Previtera E, Pirrone CA. Malarguesaurus florenciae gen. et sp. nov., a new titanosauriform (Dinosauria, Sauropoda) from the Upper Cretaceous of Mendoza, Argentina. Cretaceous Research 2009; 30: 135 - 48. https: // doi. org / 10.1016 / j. cretres. 2008.06.006" year="2009">
|
||||
González Riga
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D563717FC654A54" box="[888,945,551,575]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2009
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, the ventral surface of the centrum is anteroposteriorly concave, lacking an articular surface for the haemal arch at the anteriormost vertebrae. However, the other caudal vertebrae have a visible articular surface for a haemal arch with very pronounced end, in a similar way to many other titanosaurs.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBD5D7B37DBFC324CD9" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||||
No signal of the ventrolateral ridges is preserved for
|
||||
<taxonomicName id="4C784D62FFE1FFBD5B5737DBFC564B70" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="valdecii">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5B5737DBFC564B70" italics="true" pageId="4" pageNumber="5">Tiamat valdecii</emphasis>
|
||||
</taxonomicName>
|
||||
. The lack of this structure is also observed in several titanosaurs, e.g. in the African titanosaurs
|
||||
<taxonomicName id="4C784D62FFE1FFBD5AF8361DFC414B32" authority="(Gorscak et al. 2014)" baseAuthorityName="Gorscak" baseAuthorityYear="2014" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5AF8361DFA9D4B51" box="[1238,1353,802,826]" italics="true" pageId="4" pageNumber="5">Rukwatitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B76361DFC5E4B32" author="Gorscak E & O'Connor PM & Stevens NJ" pageId="4" pageNumber="5" pagination="1133 - 54" refId="ref12567" refString="Gorscak E, O'Connor PM, Stevens NJ et al. The basal titanosaurian Rukwatitan bisepultus (Dinosauria, Sauropoda) from the Middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 2014; 34: 1133 - 54. https: // doi. org / 10.1080 / 02724634.2014.845568" year="2014">
|
||||
Gorscak
|
||||
<emphasis id="B90CEAF3FFE1FFBD5B9F361CFC864B32" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C784D62FFE1FFBD5DEB367EFA684B32" authority="(Gorscak and O'Connor 2019)" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[965,1468,833,857]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5DEB367EFBAC4B32" box="[965,1144,833,857]" italics="true" pageId="4" pageNumber="5">Mnyamawamtuka</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5AA6367EFA654B32" author="Gorscak E & O'Connor PM" box="[1160,1457,833,857]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, as well as in
|
||||
<taxonomicName id="4C784D62FFE1FFBD5D91365EFAB74B12" authority="(Averianov and Efimov 2018)" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" box="[959,1379,865,889]" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D91365EFBF74B12" box="[959,1059,865,889]" italics="true" pageId="4" pageNumber="5">Volgatitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5A1B365EFA834B12" author="Averianov A & Efimov V" box="[1077,1367,865,889]" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. But this differs from the well-developed ventrolateral ridges observed in the basal lithostrotians
|
||||
<taxonomicName id="4C784D62FFE1FFBD5A0C369FFA9F4BDC" authority="(Wang et al. 2021)" baseAuthorityName="Wang" baseAuthorityYear="2021" box="[1058,1355,927,952]" genus="Hamititan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5A0C369FFB5D4BDC" box="[1058,1161,928,951]" italics="true" pageId="4" pageNumber="5">Hamititan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5AB7369FFAEB4BDC" author="Wang X & Bandeira KLN & Qiu R" box="[1177,1343,927,952]" pageId="4" pageNumber="5" refId="ref14686" refString="Wang X, Bandeira KLN, Qiu R et al. The first dinosaurs from the Early Cretaceous Hami Pterosaur Fauna, China. Scientific Reports 2021; 11: 14962." type="journal volume" year="2021">
|
||||
Wang
|
||||
<emphasis id="B90CEAF3FFE1FFBD5AF6369FFAD44BDC" box="[1240,1280,927,951]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Also, it differs from the less developed ventrolateral ridges in
|
||||
<taxonomicName id="4C784D62FFE1FFBD5B193680FC364B9D" authority="(Gomani 2005)" baseAuthorityName="Gomani" baseAuthorityYear="2005" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5B193680FA154BBC" box="[1335,1473,959,983]" italics="true" pageId="4" pageNumber="5">Malawisaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5D6D36E1FC034B9D" author="Gomani EM" box="[835,983,990,1014]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and in the derived lithostrotian
|
||||
<taxonomicName id="4C784D62FFE1FFBD5B6236E1FB1D4C7E" authority="(Santucci and Arruda-Campos 2011)" baseAuthorityName="Santucci and Arruda-Campos" baseAuthorityYear="2011" class="Reptilia" genus="Arrudatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5B6236E1FA164B9D" box="[1356,1474,990,1014]" italics="true" pageId="4" pageNumber="5">Arrudatitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5D6A36C2FB694C7E" author="Santucci RM & Arruda-Campos AC de" box="[836,1213,1021,1045]" pageId="4" pageNumber="5" pagination="1 - 33" refId="ref14368" refString="Santucci RM, Arruda-Campos AC de. A new sauropod (Macronaria, Titanosauria) from the Adamantina Formation, Bauru Group, Upper Cretaceous of Brazil and the phylogenetic relationships of Aeolosaurini. Zootaxa 2011; 3085: 1 - 33." year="2011">Santucci and Arruda-Campos 2011</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Also, there is no ventral longitudinal groove as presented in some titanosauriforms from the Lower Cretaceous, e.g. in
|
||||
<taxonomicName id="4C784D62FFE1FFBD5A9A3103FC1D4C18" authority="(Averianov and Efimov 2018)" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5A9A3103FACC4C3F" box="[1204,1304,1084,1108]" italics="true" pageId="4" pageNumber="5">Volgatitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B023103FC694C18" author="Averianov A & Efimov V" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Also ventrally, some few small vascular foramina (‘pneumatopore’) are abundant over the ventral surface of all caudal vertebrae.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBD5D7B3186FA6B4D24" blockId="4.[824,1477,144,1985]" pageId="4" pageNumber="5">
|
||||
One of the features of the new species, the short middle caudal centra (
|
||||
<tableCitation id="C6FA035AFFE1FFBD5DA631E6FC074C9A" box="[904,979,1241,1265]" captionStart="Table 1" captionStartId="3.[113,168,143,167]" captionTargetPageId="3" captionText="Table 1. Breakages and wear in the vertebrae of Tiamat resulted from taphonomic processes." pageId="4" pageNumber="5">Table 1</tableCitation>
|
||||
) with well-marked articular facets for the haemal arch, is also distinctive from other titanosaurian taxa. This is not observed in other titanosauriforms from the Lower Cretaceous, for example, and even in other taxa from the Upper Cretaceous.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE1FFBC5D7B3069FF3C4F04" blockId="4.[824,1477,144,1985]" lastBlockId="5.[112,763,1785,1997]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||||
The transverse processes are relatively short (
|
||||
<tableCitation id="C6FA035AFFE1FFBD5B163069FA534D05" box="[1336,1415,1366,1390]" captionStart="Table 2" captionStartId="6.[129,184,143,167]" captionTargetPageId="6" captionText="Table 2. Measurements of caudal vertebrae of Tiamat. Abbreviations:CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||||
) and dorsolaterally projected from the dorsal margin of the vertebral centra, as in many titanosaurs (e.g.
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5A8A30AAFAE64DC7" author="Gomani EM" box="[1188,1330,1428,1453]" pageId="4" pageNumber="5" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B1030AAFBCA4DA7" author="Santucci RM & Arruda-Campos AC de" pageId="4" pageNumber="5" pagination="1 - 33" refId="ref14368" refString="Santucci RM, Arruda-Campos AC de. A new sauropod (Macronaria, Titanosauria) from the Adamantina Formation, Bauru Group, Upper Cretaceous of Brazil and the phylogenetic relationships of Aeolosaurini. Zootaxa 2011; 3085: 1 - 33." year="2011">Santucci and Arruda-Campos 2011</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5A02308BFA874DA7" author="Averianov A & Efimov V" box="[1068,1363,1460,1484]" journalOrPublisher="Biological Communications" pageId="4" pageNumber="5" pagination="145 - 62" part="63" refId="ref11328" refString="Averianov A, Efimov V. The oldest titanosaurian sauropod of the Northern Hemisphere. Biological Communications 2018; 63: 145 - 62." title="The oldest titanosaurian sauropod of the Northern Hemisphere" type="journal article" year="2018">Averianov and Efimov 2018</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B4C308BFC334D80" author="Gonzalez Riga BJ & Mannion PD & Poropat SF" pageId="4" pageNumber="5" pagination="136 - 81" refId="ref12412" refString="Gonzalez Riga BJ, Mannion PD, Poropat SF et al. Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 2018; 184: 136 - 81. https: // doi. org / 10.1093 / zoolinnean / zlx 103" year="2018">
|
||||
Gonzàlez Riga
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D5D30EBFC7D4D80" box="[883,937,1491,1515]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2018
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5DD730ECFAE24D80" author="Gorscak E & O'Connor PM" box="[1017,1334,1491,1515]" pageId="4" pageNumber="5" refId="ref12514" refString="Gorscak E, O'Connor PM. A new African titanosaurian sauropod dinosaur from the middle cretaceous galula formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLoS One 2019; 14: e 0211412. https: // doi. org / 10.1371 / journal. pone. 0211412" year="2019">Gorscak and O’Connor 2019</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B6930EBFCB94E60" author="Wang X & Bandeira KLN & Qiu R" pageId="4" pageNumber="5" refId="ref14686" refString="Wang X, Bandeira KLN, Qiu R et al. The first dinosaurs from the Early Cretaceous Hami Pterosaur Fauna, China. Scientific Reports 2021; 11: 14962." type="journal volume" year="2021">
|
||||
Wang
|
||||
<emphasis id="B90CEAF3FFE1FFBD5BA330EBFA684D80" box="[1421,1468,1491,1515]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||||
. 2021
|
||||
</bibRefCitation>
|
||||
). This is indicative of the positioning of the caudal vertebrae, due to the development of the transverse processes, especially in the middle caudal vertebrae, where they progressively diminish in size until becoming vestigial (e.g.
|
||||
<bibRefCitation id="EFE94B10FFE1FFBD5B4D336EFC5B4EE3" author="Ibiricu LM & Lamanna MC & Martinez RDF" pageId="4" pageNumber="5" pagination="221 - 36" refId="ref12761" refString="Ibiricu LM, Lamanna MC, Martinez RDF et al. A novel form of postcranial skeletal pneumaticity in a sauropod dinosaur: Implications for the paleobiology of Rebbachisauridae. Acta Palaeontologica Polonica 2017; 62: 221 - 36." year="2017">
|
||||
Ibiricu
|
||||
<emphasis id="B90CEAF3FFE1FFBD5B9F336EFC864EE3" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
). This reduction in size progresses in the transverse processes, to the point where these structures transform into heavy blunt knobs, regarded here as the beginning of the middle caudal series, especially in UFRJ-DG 527-R, UFRJ-DG 591-R, UFRJ-DG 635-R, and UFRJ-DG 704-R (
|
||||
<figureCitation id="13432A64FFE1FFBD5AF533D2FA924F6E" box="[1243,1350,1773,1797]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="4" pageNumber="5">Fig. 4D–G</figureCitation>
|
||||
). The transverse processes are triangular and curved posteriorly, similarly to
|
||||
<emphasis id="B90CEAF3FFE1FFBD5D173212FC034F2F" box="[825,983,1837,1860]" italics="true" pageId="4" pageNumber="5">Tastavinasaurus</emphasis>
|
||||
and
|
||||
<taxonomicName id="4C784D62FFE1FFBD5A253213FB734F2F" box="[1035,1191,1836,1860]" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE1FFBD5A253213FB734F2F" box="[1035,1191,1836,1860]" italics="true" pageId="4" pageNumber="5">Malarguesaurus</emphasis>
|
||||
</taxonomicName>
|
||||
, being supported by a massive and robust PRDL, especially on the most anterior caudal vertebrae (UFRJ-DG 636-R, UFRJ-DG 638-R, and UFRJ-DG 606-R). The neural canal is subcircular being delimited by thick intraprezygapophyseal laminae (TPRL, on anterior view) and intrapost-zygapophyseal lamina (TPOL, on posterior view). The thick TPRL and TPOL of
|
||||
<taxonomicName id="4C784D62FFE0FFBC5F9B3226FDE24F5B" box="[437,566,1816,1840]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="differ">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5F9B3226FDE24F5B" box="[437,566,1816,1840]" italics="true" pageId="5" pageNumber="6">Tiamat differ</emphasis>
|
||||
</taxonomicName>
|
||||
from the thin laminae presented by
|
||||
<taxonomicName id="4C784D62FFE0FFBC5F1F3208FD584F3B" authority="(Gorsack et al. 2014)" baseAuthorityName="Gorsack" baseAuthorityYear="2014" box="[305,652,1847,1872]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5F1F3208FE704F24" box="[305,420,1847,1871]" italics="true" pageId="5" pageNumber="6">Rukwatitan</emphasis>
|
||||
(Gorsack
|
||||
<emphasis id="B90CEAF3FFE0FFBC5C3A3207FDEB4F24" box="[532,575,1847,1871]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||||
. 2014)
|
||||
</taxonomicName>
|
||||
and other titanosaurs.
|
||||
</paragraph>
|
||||
<caption id="DF076669FFE0FFBC5E5F3362FE574EA3" ID-DOI="http://doi.org/10.5281/zenodo.14764393" ID-Zenodo-Dep="14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1629,1653]" targetBox="[115,1456,150,1597]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8BC736E1FFE0FFBC5E5F3362FE574EA3" blockId="5.[113,1459,1629,1737]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5E5F3362FF1D4E1E" bold="true" box="[113,201,1629,1653]" pageId="5" pageNumber="6">Figure 4.</emphasis>
|
||||
Caudal vertebrae attributed to
|
||||
<taxonomicName id="4C784D62FFE0FFBC5FC03361FDAD4E1E" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[494,633,1629,1653]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="valdecii" status="gen. et sp. nov.">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5FC03361FDAD4E1E" box="[494,633,1629,1653]" italics="true" pageId="5" pageNumber="6">Tiamat valdecii</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A23F5788FFE0FFBC5C513362FCDC4E1F" box="[639,776,1629,1653]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BC736E1FFE0FFBC5EA33248FCBB4FA6" blockId="5.[112,763,1785,1997]" lastBlockId="5.[810,1459,1785,1997]" pageId="5" pageNumber="6">
|
||||
From the middle caudal vertebrae preserved, UFRJ-DG 635-R shows prezygapophyses dorsomedially positioned, supported ventrally by sharp CPRL. Above the prezygapophyses and the post-zygapophyses,
|
||||
<taxonomicName id="4C784D62FFE0FFBC5DD733C6FB404F7A" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1017,1172,1785,1809]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="valdecii">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5DD733C6FB404F7A" box="[1017,1172,1785,1809]" italics="true" pageId="5" pageNumber="6">Tiamat valdecii</emphasis>
|
||||
</taxonomicName>
|
||||
shows a marked accessory tuberosity. The SPRL observed in the most anterior caudal vertebra and in the middle caudal are weakly developed, like the weakly developed SPRL of
|
||||
<taxonomicName id="4C784D62FFE0FFBC5A7B3268FCBD4FE5" authority="(Mannion and Calvo 2011)" baseAuthorityName="Mannion and Calvo" baseAuthorityYear="2011" class="Reptilia" family="Titanosauridae" genus="Andesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5A7B3268FB134F04" box="[1109,1223,1879,1903]" italics="true" pageId="5" pageNumber="6">Andesaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE0FFBC5AF23268FC8A4FE5" author="Mannion PD & Calvo JO" pageId="5" pageNumber="6" refId="ref13041" refString="Mannion PD, Calvo JO. Anatomy of the basal titanosaur (Dinosauria, Sauropoda) Andesaurus delgadoi from the mid-Cretaceous (Albianearly Cenomanian) Rio Limay Formation, Neuquen Province, Argentina: implications for titanosaur systematics. Zoological Journal of the Linnean Society 2011; 163: no - no. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00699. x" year="2011">Mannion and Calvo 2011</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C784D62FFE0FFBC5DB13249FAFE4FE5" authority="(Martinez et al. 2004)" baseAuthorityName="Martinez" baseAuthorityYear="2004" box="[927,1322,1910,1934]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5DB13249FBEE4FE5" box="[927,1082,1910,1934]" italics="true" pageId="5" pageNumber="6">Epachthosaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE0FFBC5A633249FACB4FE5" author="Martinez RD & Gimenez O & Rodriguez J" box="[1101,1311,1910,1934]" pageId="5" pageNumber="6" pagination="107 - 20" refId="ref13311" refString="Martinez RD, Gimenez O, Rodriguez J et al. An articulated specimen of the basal titanosaurian (Dinosauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal formation of Chubut Province, Argentina. Journal of Vertebrate Paleontology 2004; 24: 107 - 20. https: // doi. org / 10.1671 / 9.1" year="2004">
|
||||
Martínez
|
||||
<emphasis id="B90CEAF3FFE0FFBC5A9C3248FB094FE5" box="[1202,1245,1910,1934]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||||
. 2004
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
but differing from the thick SPRL presented by
|
||||
<taxonomicName id="4C784D62FFE0FFBC5ABE32A9FCBE4FA6" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE0FFBC5ABE32A9FAD54FC6" box="[1168,1281,1942,1965]" italics="true" pageId="5" pageNumber="6">Patagotitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE0FFBC5B3F32AAFC8A4FA6" author="Carballido JL & Pol D & Otero A" pageId="5" pageNumber="6" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
|
||||
Carballido
|
||||
<emphasis id="B90CEAF3FFE0FFBC5BAA32A9FA794FC6" box="[1412,1453,1941,1965]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||||
. 2017
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF076669FFE3FFBF5EAF35B0FEFB49EC" pageId="6" pageNumber="7" startId="6.[129,184,143,167]" targetBox="[129,1472,413,1056]" targetIsTable="true" targetPageId="6" targetType="table">
|
||||
<paragraph id="8BC736E1FFE3FFBF5EAF35B0FEFB49EC" blockId="6.[129,1471,142,391]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF35B0FF1A48CD" bold="true" box="[129,206,142,167]" pageId="6" pageNumber="7">Table 2.</emphasis>
|
||||
Measurements of caudal vertebrae of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5C1C35B0FDA148CD" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[562,629,143,166]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5C1C35B0FDA148CD" box="[562,629,143,166]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
. Abbreviations: CL, centrum length; CL + condyle, centrum length with length condyle; ACH, anterior centrum height (excluding chevron facets); ACW, anterior centrum width; PCH, posterior centrum height (excluding chevron facets); PCW, posterior centrum width; DFA, distance from anterior end of centrum to anterior end of neural arch; DFP, distance from posterior end of centrum to posterior end of neural arch; NAH, neural arch height (measured from the dorsal surface of the centrum up to the post-zygapophyses); PRL, prezygapophysis length; PRFW, length of the articular surface of the prezygapophyses; PRFH height of the articular surface of the post-zygapophysis; PSFW, width of the articular facet of the post-zygapophysis; PSFH, height of the articular facet of the post-zygapophysis; NSL, neural spine anteroposterior length (measured along base of spine); NSW, neural spine mediolateral width (measured along base of spine at the posterior margin); CRL, caudal rib length (transverse width between distal ends of ribs). Measurements are in centimetres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BC736E1FFE3FFBF5F7E34A2FAAC4C4B" pageId="6" pageNumber="7">
|
||||
<table id="F978C441FFE300465EAF34A2FA144C4B" box="[129,1472,413,1056]" gridcols="8" gridrows="18" pageId="6" pageNumber="7">
|
||||
<tr id="354834A3FFE300465EAF34A2FA1449B9" box="[129,1472,413,466]" gridrow="0" pageId="6" pageNumber="7" rowspan-0="1">
|
||||
<th id="76995DDFFFE300465F7E34A2FE6149B9" box="[336,437,413,466]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5F7E34A2FE5A49B9" bold="true" box="[336,437,413,466]" pageId="6" pageNumber="7">UFRJ-DG 636-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465FD334A2FDB649B9" box="[509,610,413,466]" gridcol="2" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5FD334A2FDEE49B9" bold="true" box="[509,610,413,466]" pageId="6" pageNumber="7">UFRJ-DG 638-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465C8734A2FCDA49B9" box="[681,782,413,466]" gridcol="3" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5C8734A2FD3349B9" bold="true" box="[681,782,413,466]" pageId="6" pageNumber="7">UFRJ-DG 606-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465D7834A2FC6F49B9" box="[854,955,413,466]" gridcol="4" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5D7834A2FC4749B9" bold="true" box="[854,955,413,466]" pageId="6" pageNumber="7">UFRJ-DG 635-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465A2C34A2FBB349B9" box="[1026,1127,413,466]" gridcol="5" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A2C34A2FBEB49B9" bold="true" box="[1026,1127,413,466]" pageId="6" pageNumber="7">UFRJ-DG 591-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465A8134A2FAC049B9" box="[1199,1300,413,466]" gridcol="6" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A8134A2FB3849B9" bold="true" box="[1199,1300,413,466]" pageId="6" pageNumber="7">UFRJ-DG 527-R</emphasis>
|
||||
</th>
|
||||
<th id="76995DDFFFE300465B7534A2FA1449B9" box="[1371,1472,413,466]" gridcol="7" gridrow="0" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5B7534A2FA4D49B9" bold="true" box="[1371,1472,413,466]" pageId="6" pageNumber="7">UFRJ-DG 704-R</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF34D5FA144A69" box="[129,1472,490,514]" gridrow="1" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF34D5FEDF4A69" box="[129,267,490,514]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF34D5FF744A69" bold="true" box="[129,160,490,514]" pageId="6" pageNumber="7">CL</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E34D5FE614A69" box="[336,437,490,514]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">7,3</td>
|
||||
<td id="76995DDFFFE300465FD334D5FDB64A69" box="[509,610,490,514]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">7,5</td>
|
||||
<td id="76995DDFFFE300465C8734D5FCDA4A69" box="[681,782,490,514]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">8,2</td>
|
||||
<td id="76995DDFFFE300465D7834D5FC6F4A69" box="[854,955,490,514]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">7</td>
|
||||
<td id="76995DDFFFE300465A2C34D5FBB34A69" box="[1026,1127,490,514]" gridcol="5" gridrow="1" pageId="6" pageNumber="7">6,4</td>
|
||||
<td id="76995DDFFFE300465A8134D5FAC04A69" box="[1199,1300,490,514]" gridcol="6" gridrow="1" pageId="6" pageNumber="7">6</td>
|
||||
<td id="76995DDFFFE300465B7534D5FA144A69" box="[1371,1472,490,514]" gridcol="7" gridrow="1" pageId="6" pageNumber="7">6,1</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3733FA144A4F" box="[129,1472,524,548]" gridrow="2" pageId="6" pageNumber="7" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||||
<th id="76995DDFFFE300465EAF3733FEDF4A4F" box="[129,267,524,548]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3733FEDF4A4F" bold="true" box="[129,267,524,548]" pageId="6" pageNumber="7">CL + condyle</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E3733FE614A4F" box="[336,437,524,548]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">9</td>
|
||||
<td id="76995DDFFFE300465FD33733FDB64A4F" box="[509,610,524,548]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">10</td>
|
||||
<td id="76995DDFFFE300465C873733FCDA4A4F" box="[681,782,524,548]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">10</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3711FA144A2D" box="[129,1472,558,582]" gridrow="3" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF3711FEDF4A2D" box="[129,267,558,582]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3711FF614A2D" bold="true" box="[129,181,558,582]" pageId="6" pageNumber="7">ACH</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E3711FE614A2D" box="[336,437,558,582]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465FD33711FDB64A2D" box="[509,610,558,582]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465C873711FCDA4A2D" box="[681,782,558,582]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">8</td>
|
||||
<td id="76995DDFFFE300465D783711FC6F4A2D" box="[854,955,558,582]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">8,4</td>
|
||||
<td id="76995DDFFFE300465A2C3711FBB34A2D" box="[1026,1127,558,582]" gridcol="5" gridrow="3" pageId="6" pageNumber="7">8</td>
|
||||
<td id="76995DDFFFE300465A813711FAC04A2D" box="[1199,1300,558,582]" gridcol="6" gridrow="3" pageId="6" pageNumber="7">8,1</td>
|
||||
<td id="76995DDFFFE300465B753711FA144A2D" box="[1371,1472,558,582]" gridcol="7" gridrow="3" pageId="6" pageNumber="7">6,6</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF376FFA144A03" box="[129,1472,592,616]" gridrow="4" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF376FFEDF4A03" box="[129,267,592,616]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF376FFF6D4A03" bold="true" box="[129,185,592,616]" pageId="6" pageNumber="7">ACW</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E376FFE614A03" box="[336,437,592,616]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">12,5</td>
|
||||
<td id="76995DDFFFE300465FD3376FFDB64A03" box="[509,610,592,616]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">12</td>
|
||||
<td id="76995DDFFFE300465C87376FFCDA4A03" box="[681,782,592,616]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">10</td>
|
||||
<td id="76995DDFFFE300465D78376FFC6F4A03" box="[854,955,592,616]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465A2C376FFBB34A03" box="[1026,1127,592,616]" gridcol="5" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465A81376FFAC04A03" box="[1199,1300,592,616]" gridcol="6" gridrow="4" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465B75376FFA144A03" box="[1371,1472,592,616]" gridcol="7" gridrow="4" pageId="6" pageNumber="7">8,5</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF374EFA144AE1" box="[129,1472,625,650]" gridrow="5" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF374EFEDF4AE1" box="[129,267,625,650]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF374EFF604AE2" bold="true" box="[129,180,625,649]" pageId="6" pageNumber="7">PCH</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E374EFE614AE1" box="[336,437,625,650]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">10,8</td>
|
||||
<td id="76995DDFFFE300465FD3374EFDB64AE1" box="[509,610,625,650]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">10</td>
|
||||
<td id="76995DDFFFE300465C87374EFCDA4AE1" box="[681,782,625,650]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">9</td>
|
||||
<td id="76995DDFFFE300465D78374EFC6F4AE1" box="[854,955,625,650]" gridcol="4" gridrow="5" pageId="6" pageNumber="7">9</td>
|
||||
<td id="76995DDFFFE300465A2C374EFBB34AE1" box="[1026,1127,625,650]" gridcol="5" gridrow="5" pageId="6" pageNumber="7">8</td>
|
||||
<td id="76995DDFFFE300465A81374EFAC04AE1" box="[1199,1300,625,650]" gridcol="6" gridrow="5" pageId="6" pageNumber="7">7,5</td>
|
||||
<td id="76995DDFFFE300465B75374EFA144AE1" box="[1371,1472,625,650]" gridcol="7" gridrow="5" pageId="6" pageNumber="7">6</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF37ACFA144AC7" box="[129,1472,659,684]" gridrow="6" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF37ACFEDF4AC7" box="[129,267,659,684]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF37ACFF6C4AC0" bold="true" box="[129,184,659,683]" pageId="6" pageNumber="7">PCW</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E37ACFE614AC7" box="[336,437,659,684]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">12,3</td>
|
||||
<td id="76995DDFFFE300465FD337ACFDB64AC7" box="[509,610,659,684]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">12</td>
|
||||
<td id="76995DDFFFE300465C8737ACFCDA4AC7" box="[681,782,659,684]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465D7837ACFC6F4AC7" box="[854,955,659,684]" gridcol="4" gridrow="6" pageId="6" pageNumber="7">11,5</td>
|
||||
<td id="76995DDFFFE300465A2C37ACFBB34AC7" box="[1026,1127,659,684]" gridcol="5" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465A8137ACFAC04AC7" box="[1199,1300,659,684]" gridcol="6" gridrow="6" pageId="6" pageNumber="7">11</td>
|
||||
<td id="76995DDFFFE300465B7537ACFA144AC7" box="[1371,1472,659,684]" gridcol="7" gridrow="6" pageId="6" pageNumber="7">7,4</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF378AFA144AA6" box="[129,1472,693,717]" gridrow="7" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF378AFEDF4AA6" box="[129,267,693,717]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF378AFF644AA6" bold="true" box="[129,176,693,717]" pageId="6" pageNumber="7">DFA</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E378AFE614AA6" box="[336,437,693,717]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">4,7</td>
|
||||
<td id="76995DDFFFE300465FD3378AFDB64AA6" box="[509,610,693,717]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">5</td>
|
||||
<td id="76995DDFFFE300465C87378AFCDA4AA6" box="[681,782,693,717]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465D78378AFC6F4AA6" box="[854,955,693,717]" gridcol="4" gridrow="7" pageId="6" pageNumber="7">5,3</td>
|
||||
<td id="76995DDFFFE300465A2C378AFBB34AA6" box="[1026,1127,693,717]" gridcol="5" gridrow="7" pageId="6" pageNumber="7">4,8</td>
|
||||
<td id="76995DDFFFE300465A81378AFAC04AA6" box="[1199,1300,693,717]" gridcol="6" gridrow="7" pageId="6" pageNumber="7">4,8</td>
|
||||
<td id="76995DDFFFE300465B75378AFA144AA6" box="[1371,1472,693,717]" gridcol="7" gridrow="7" pageId="6" pageNumber="7">5,4</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF37E8FA144A84" box="[129,1472,727,751]" gridrow="8" pageId="6" pageNumber="7">
|
||||
<th id="76995DDFFFE300465EAF37E8FEDF4A84" box="[129,267,727,751]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF37E8FF654A84" bold="true" box="[129,177,727,751]" pageId="6" pageNumber="7">DFP</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E37E8FE614A84" box="[336,437,727,751]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">2,1</td>
|
||||
<td id="76995DDFFFE300465FD337E8FDB64A84" box="[509,610,727,751]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||||
<td id="76995DDFFFE300465C8737E8FCDA4A84" box="[681,782,727,751]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||||
<td id="76995DDFFFE300465D7837E8FC6F4A84" box="[854,955,727,751]" gridcol="4" gridrow="8" pageId="6" pageNumber="7">2</td>
|
||||
<td id="76995DDFFFE300465A2C37E8FBB34A84" box="[1026,1127,727,751]" gridcol="5" gridrow="8" pageId="6" pageNumber="7">2,1</td>
|
||||
<td id="76995DDFFFE300465A8137E8FAC04A84" box="[1199,1300,727,751]" gridcol="6" gridrow="8" pageId="6" pageNumber="7">2,5</td>
|
||||
<td id="76995DDFFFE300465B7537E8FA144A84" box="[1371,1472,727,751]" gridcol="7" gridrow="8" pageId="6" pageNumber="7">2,3</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF37C6FA144B7A" box="[129,1472,761,785]" gridrow="9" pageId="6" pageNumber="7" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF37C6FEDF4B7A" box="[129,267,761,785]" gridcol="0" gridrow="9" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF37C6FF624B7A" bold="true" box="[129,182,761,785]" pageId="6" pageNumber="7">NAH</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E37C6FE614B7A" box="[336,437,761,785]" gridcol="1" gridrow="9" pageId="6" pageNumber="7">2,1</td>
|
||||
<td id="76995DDFFFE300465FD337C6FDB64B7A" box="[509,610,761,785]" gridcol="2" gridrow="9" pageId="6" pageNumber="7">2,2</td>
|
||||
<td id="76995DDFFFE300465D7837C6FC6F4B7A" box="[854,955,761,785]" gridcol="4" gridrow="9" pageId="6" pageNumber="7">2</td>
|
||||
<td id="76995DDFFFE300465A2C37C6FBB34B7A" box="[1026,1127,761,785]" gridcol="5" gridrow="9" pageId="6" pageNumber="7">2</td>
|
||||
<td id="76995DDFFFE300465A8137C6FAC04B7A" box="[1199,1300,761,785]" gridcol="6" gridrow="9" pageId="6" pageNumber="7">1,8</td>
|
||||
<td id="76995DDFFFE300465B7537C6FA144B7A" box="[1371,1472,761,785]" gridcol="7" gridrow="9" pageId="6" pageNumber="7">1,7</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3624FA144B58" box="[129,1472,795,819]" gridrow="10" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF3624FEDF4B58" box="[129,267,795,819]" gridcol="0" gridrow="10" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3624FF7B4B58" bold="true" box="[129,175,795,819]" pageId="6" pageNumber="7">PRL</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465D783624FC6F4B58" box="[854,955,795,819]" gridcol="4" gridrow="10" pageId="6" pageNumber="7">5,6</td>
|
||||
<td id="76995DDFFFE300465A2C3624FBB34B58" box="[1026,1127,795,819]" gridcol="5" gridrow="10" pageId="6" pageNumber="7">4,6</td>
|
||||
<td id="76995DDFFFE300465A813624FAC04B58" box="[1199,1300,795,819]" gridcol="6" gridrow="10" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B753624FA144B58" box="[1371,1472,795,819]" gridcol="7" gridrow="10" pageId="6" pageNumber="7">4,5</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3602FA144B3E" box="[129,1472,829,853]" gridrow="11" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF3602FEDF4B3E" box="[129,267,829,853]" gridcol="0" gridrow="11" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3602FF134B3E" bold="true" box="[129,199,829,853]" pageId="6" pageNumber="7">PRFW</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465D783602FC6F4B3E" box="[854,955,829,853]" gridcol="4" gridrow="11" pageId="6" pageNumber="7">3,9</td>
|
||||
<td id="76995DDFFFE300465A2C3602FBB34B3E" box="[1026,1127,829,853]" gridcol="5" gridrow="11" pageId="6" pageNumber="7">4</td>
|
||||
<td id="76995DDFFFE300465A813602FAC04B3E" box="[1199,1300,829,853]" gridcol="6" gridrow="11" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B753602FA144B3E" box="[1371,1472,829,853]" gridcol="7" gridrow="11" pageId="6" pageNumber="7">3,5</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3661FA144B1C" box="[129,1472,862,887]" gridrow="12" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF3661FEDF4B1C" box="[129,267,862,887]" gridcol="0" gridrow="12" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3660FF164B1C" bold="true" box="[129,194,863,887]" pageId="6" pageNumber="7">PRFH</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465D783661FC6F4B1C" box="[854,955,862,887]" gridcol="4" gridrow="12" pageId="6" pageNumber="7">3,6</td>
|
||||
<td id="76995DDFFFE300465A2C3661FBB34B1C" box="[1026,1127,862,887]" gridcol="5" gridrow="12" pageId="6" pageNumber="7">3,5</td>
|
||||
<td id="76995DDFFFE300465A813661FAC04B1C" box="[1199,1300,862,887]" gridcol="6" gridrow="12" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B753661FA144B1C" box="[1371,1472,862,887]" gridcol="7" gridrow="12" pageId="6" pageNumber="7">3</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF36BFFA144BF2" box="[129,1472,896,921]" gridrow="13" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF36BFFEDF4BF2" box="[129,267,896,921]" gridcol="0" gridrow="13" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF36BFFF174BF3" bold="true" box="[129,195,896,920]" pageId="6" pageNumber="7">PSFW</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465D7836BFFC6F4BF2" box="[854,955,896,921]" gridcol="4" gridrow="13" pageId="6" pageNumber="7">4</td>
|
||||
<td id="76995DDFFFE300465A2C36BFFBB34BF2" box="[1026,1127,896,921]" gridcol="5" gridrow="13" pageId="6" pageNumber="7">3,2</td>
|
||||
<td id="76995DDFFFE300465A8136BFFAC04BF2" box="[1199,1300,896,921]" gridcol="6" gridrow="13" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B7536BFFA144BF2" box="[1371,1472,896,921]" gridcol="7" gridrow="13" pageId="6" pageNumber="7">3,2</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF369DFA144BD1" box="[129,1472,930,954]" gridrow="14" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF369DFEDF4BD1" box="[129,267,930,954]" gridcol="0" gridrow="14" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF369DFF6A4BD1" bold="true" box="[129,190,930,954]" pageId="6" pageNumber="7">PSFH</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465D78369DFC6F4BD1" box="[854,955,930,954]" gridcol="4" gridrow="14" pageId="6" pageNumber="7">3,9</td>
|
||||
<td id="76995DDFFFE300465A2C369DFBB34BD1" box="[1026,1127,930,954]" gridcol="5" gridrow="14" pageId="6" pageNumber="7">3,6</td>
|
||||
<td id="76995DDFFFE300465A81369DFAC04BD1" box="[1199,1300,930,954]" gridcol="6" gridrow="14" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B75369DFA144BD1" box="[1371,1472,930,954]" gridcol="7" gridrow="14" pageId="6" pageNumber="7">3,4</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF36FBFA144BB7" box="[129,1472,964,988]" gridrow="15" pageId="6" pageNumber="7" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF36FBFEDF4BB7" box="[129,267,964,988]" gridcol="0" gridrow="15" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF36FBFF7A4BB7" bold="true" box="[129,174,964,988]" pageId="6" pageNumber="7">NSL</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E36FBFE614BB7" box="[336,437,964,988]" gridcol="1" gridrow="15" pageId="6" pageNumber="7">6,3</td>
|
||||
<td id="76995DDFFFE300465FD336FBFDB64BB7" box="[509,610,964,988]" gridcol="2" gridrow="15" pageId="6" pageNumber="7">5,4</td>
|
||||
<td id="76995DDFFFE300465D7836FBFC6F4BB7" box="[854,955,964,988]" gridcol="4" gridrow="15" pageId="6" pageNumber="7">5</td>
|
||||
<td id="76995DDFFFE300465A2C36FBFBB34BB7" box="[1026,1127,964,988]" gridcol="5" gridrow="15" pageId="6" pageNumber="7">6,5</td>
|
||||
<td id="76995DDFFFE300465A8136FBFAC04BB7" box="[1199,1300,964,988]" gridcol="6" gridrow="15" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B7536FBFA144BB7" box="[1371,1472,964,988]" gridcol="7" gridrow="15" pageId="6" pageNumber="7">6,1</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF36D9FA144B95" box="[129,1472,998,1022]" gridrow="16" pageId="6" pageNumber="7" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF36D9FEDF4B95" box="[129,267,998,1022]" gridcol="0" gridrow="16" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF36D9FF634B95" bold="true" box="[129,183,998,1022]" pageId="6" pageNumber="7">NSW</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E36D9FE614B95" box="[336,437,998,1022]" gridcol="1" gridrow="16" pageId="6" pageNumber="7">4</td>
|
||||
<td id="76995DDFFFE300465FD336D9FDB64B95" box="[509,610,998,1022]" gridcol="2" gridrow="16" pageId="6" pageNumber="7">4,5</td>
|
||||
<td id="76995DDFFFE300465D7836D9FC6F4B95" box="[854,955,998,1022]" gridcol="4" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||||
<td id="76995DDFFFE300465A2C36D9FBB34B95" box="[1026,1127,998,1022]" gridcol="5" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||||
<td id="76995DDFFFE300465A8136D9FAC04B95" box="[1199,1300,998,1022]" gridcol="6" gridrow="16" pageId="6" pageNumber="7">-</td>
|
||||
<td id="76995DDFFFE300465B7536D9FA144B95" box="[1371,1472,998,1022]" gridcol="7" gridrow="16" pageId="6" pageNumber="7">3</td>
|
||||
</tr>
|
||||
<tr id="354834A3FFE300465EAF3137FA144C4B" box="[129,1472,1032,1056]" gridrow="17" pageId="6" pageNumber="7" rowspan-3="1">
|
||||
<th id="76995DDFFFE300465EAF3137FEDF4C4B" box="[129,267,1032,1056]" gridcol="0" gridrow="17" pageId="6" pageNumber="7">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3137FF654C4B" bold="true" box="[129,177,1032,1056]" pageId="6" pageNumber="7">CRL</emphasis>
|
||||
</th>
|
||||
<td id="76995DDFFFE300465F7E3137FE614C4B" box="[336,437,1032,1056]" gridcol="1" gridrow="17" pageId="6" pageNumber="7">5,2</td>
|
||||
<td id="76995DDFFFE300465FD33137FDB64C4B" box="[509,610,1032,1056]" gridcol="2" gridrow="17" pageId="6" pageNumber="7">5,8</td>
|
||||
<td id="76995DDFFFE300465D783137FC6F4C4B" box="[854,955,1032,1056]" gridcol="4" gridrow="17" pageId="6" pageNumber="7">2</td>
|
||||
<td id="76995DDFFFE300465A2C3137FBB34C4B" box="[1026,1127,1032,1056]" gridcol="5" gridrow="17" pageId="6" pageNumber="7">2</td>
|
||||
<td id="76995DDFFFE300465A813137FAC04C4B" box="[1199,1300,1032,1056]" gridcol="6" gridrow="17" pageId="6" pageNumber="7">1,2</td>
|
||||
<td id="76995DDFFFE300465B753137FA144C4B" box="[1371,1472,1032,1056]" gridcol="7" gridrow="17" pageId="6" pageNumber="7">1,2</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE3FFBF5EB2315BFE5B4D9F" blockId="6.[128,780,1124,1994]" pageId="6" pageNumber="7">
|
||||
Concerning one of the diagnostic features proposed, the marked accessory tuberosity dorsoventrally developed located above the prezygapophyses; is an unusual feature. To our knowledge, no titanosaur presents this tuberosity. The post-zygapophyses are slightly displaced laterally and have concave articular facets, a condition also observed on more anterior caudal vertebrae of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5FEE301FFCD14D53" authority="(Gomani 2005)" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[448,773,1311,1336]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5FEE301FFD9E4D53" box="[448,586,1312,1336]" italics="true" pageId="6" pageNumber="7">Malawisaurus</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5C4C301FFD2D4D5C" author="Gomani EM" box="[610,761,1311,1336]" pageId="6" pageNumber="7" pagination="1 - 37" refId="ref12297" refString="Gomani EM. Sauropod dinosaurs from the Early Cretaceous of Malawi, Africa. Palaeontologia Electronica 2005; 8: 1 - 37." year="2005">Gomani 2005</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. However, those concave articular facets, being deeply excavated, in the prezygapophyses and the post-zygapophyses of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5EAF3041FF1E4DFE" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[129,202,1406,1429]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF3041FF1E4DFE" box="[129,202,1406,1429]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
are a unique feature among titanosaurs, especially with the contemporaneous (or temporally close) taxa compared here, as
|
||||
<taxonomicName id="4C784D62FFE3FFBF5EFE3083FE974DBF" box="[208,323,1468,1492]" genus="Rukwatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EFE3083FE974DBF" box="[208,323,1468,1492]" italics="true" pageId="6" pageNumber="7">Rukwatitan</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D62FFE3FFBF5F603083FE0C4DBF" baseAuthorityName="Gomani" baseAuthorityYear="2005" box="[334,472,1468,1492]" class="Reptilia" family="Titanosauridae" genus="Malawisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5F603083FE0C4DBF" box="[334,472,1468,1492]" italics="true" pageId="6" pageNumber="7">Malawisaurus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D62FFE3FFBF5FCC3083FD414DBF" baseAuthorityName="Gorscak and O'Connor" baseAuthorityYear="2019" box="[482,661,1468,1492]" genus="Mnyamawamtuka" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5FCC3083FD414DBF" box="[482,661,1468,1492]" italics="true" pageId="6" pageNumber="7">Mnyamawamtuka</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D62FFE3FFBF5C8E3083FCD04DBF" baseAuthorityName="Averianov and Efimov" baseAuthorityYear="2018" box="[672,772,1468,1492]" genus="Volgatitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5C8E3083FCD04DBF" box="[672,772,1468,1492]" italics="true" pageId="6" pageNumber="7">Volgatitan</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C784D62FFE3FFBF5EAF30E3FF264D98" box="[129,242,1500,1523]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EAF30E3FF264D98" box="[129,242,1500,1523]" italics="true" pageId="6" pageNumber="7">Patagotitan</emphasis>
|
||||
</taxonomicName>
|
||||
, among others.
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE3FFBF5EB230C4FDBF4F27" blockId="6.[128,780,1124,1994]" pageId="6" pageNumber="7">
|
||||
The new species also presents a hyposphene–hypantrum articulation
|
||||
<taxonomicName id="4C784D62FFE3FFBF5EC43325FE514E59" box="[234,389,1562,1586]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5EC43325FE514E59" box="[234,389,1562,1586]" italics="true" pageId="6" pageNumber="7">Epachthosaurus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5FB63325FDCC4E59" author="Powell JE" box="[408,536,1562,1586]" pageId="6" pageNumber="7" pagination="123 - 8" refId="ref13985" refString="Powell JE. Epachthosaurus sciutoi g et sp. nov., un Nuevo dinosaurio titanosaurido del Cretacico de Patagonia. Actas V Congreso Argentino de Paleontologia y Bioestratigrafia 1990; 1: 123 - 8." year="1990">Powell 1990</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5C083325FD2D4E59" author="Martinez RD & Gimenez O & Rodriguez J" box="[550,761,1562,1586]" pageId="6" pageNumber="7" pagination="107 - 20" refId="ref13311" refString="Martinez RD, Gimenez O, Rodriguez J et al. An articulated specimen of the basal titanosaurian (Dinosauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal formation of Chubut Province, Argentina. Journal of Vertebrate Paleontology 2004; 24: 107 - 20. https: // doi. org / 10.1671 / 9.1" year="2004">
|
||||
Martínez
|
||||
<emphasis id="B90CEAF3FFE3FFBF5CA23324FD634E59" box="[652,695,1562,1586]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||||
. 2004
|
||||
</bibRefCitation>
|
||||
). However, in
|
||||
<taxonomicName id="4C784D62FFE3FFBF5F3F3305FE784E39" box="[273,428,1594,1618]" class="Reptilia" family="Titanosauridae" genus="Epachthosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5F3F3305FE784E39" box="[273,428,1594,1618]" italics="true" pageId="6" pageNumber="7">Epachthosaurus</emphasis>
|
||||
</taxonomicName>
|
||||
, the hyposphene–hypantrum is observable in some of the anterior caudal vertebrae, but not in caudal vertebrae sequence, which differs from the new species proposed here, where in almost all vertebrae this feature was observed. Both prespinal and postspinal fossae are dorsoventrally elongated but relatively shallow in the anteriormost caudal vertebrae (
|
||||
<figureCitation id="13432A64FFE3FFBF5EFF33C9FEEE4F65" box="[209,314,1782,1806]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="6" pageNumber="7">Fig. 4A–A</figureCitation>
|
||||
<figureCitation id="13432A64FFE3FFBF5F1733C9FE924F65" box="[313,326,1782,1806]" captionStart="Figure 3" captionStartId="3.[113,178,1495,1519]" captionTargetBox="[117,1454,634,1464]" captionTargetId="figure-321@3.[114,1458,632,1467]" captionTargetPageId="3" captionText="Figure 3. Digital reconstruction of caudal vertebrae of the Tiamat. The vertebrae used in the range of motion analysis are from C17? to C27?." figureDoi="http://doi.org/10.5281/zenodo.14764391" httpUri="https://zenodo.org/record/14764391/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
|
||||
) and in the isolated neural spine associated (
|
||||
<figureCitation id="13432A64FFE3FFBF5EA5322AFF034F46" box="[139,215,1813,1837]" captionStart="Figure 4" captionStartId="5.[113,178,1629,1653]" captionTargetBox="[115,1456,150,1597]" captionTargetId="figure-125@5.[113,1460,144,1601]" captionTargetPageId="5" captionText="Figure 4. Caudal vertebrae attributed to Tiamat valdecii gen. et sp.nov.UFRJ-DG 636-R (A1–A3), UFRJ-DG 638-R (B1–B3), UFRJ-DG 606-R (C1–C3), UFRJ-DG 635-R (D1–D3), UFRJ-DG 591-R (E1–E3), UFRJ-DG 527-R (F1–F3), UFRJ-DG 704-R (G1–G3), UFRJ-DG 574-R (H). A1 to G3 in views lateral, anterior, and posterior, respectively. Abbreviations: ppr, prz, prezygapophysis; poz, post-zygapophysis; ns, neural spine. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.14764393" httpUri="https://zenodo.org/record/14764393/files/figure.png" pageId="6" pageNumber="7">Fig. 4H</figureCitation>
|
||||
). There is no sign of the prespinal or postspinal laminae as in some neosauropods (
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5FBE320BFD8F4F27" author="Mannion PD & Upchurch P & Barnes RN" box="[400,603,1844,1868]" pageId="6" pageNumber="7" pagination="98 - 206" refId="ref13110" refString="Mannion PD, Upchurch P, Barnes RN et al. Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms. Zoological Journal of the Linnean Society 2013; 168: 98 - 206. https: // doi. org / 10.1111 / zoj. 12029" year="2013">
|
||||
Mannion
|
||||
<emphasis id="B90CEAF3FFE3FFBF5FDA320AFDF64F27" box="[500,546,1844,1868]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||||
2013
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE3FFBF5EB2326BFC314D9F" blockId="6.[128,780,1124,1994]" lastBlockId="6.[825,1475,1124,1618]" pageId="6" pageNumber="7">
|
||||
Lastly, as in
|
||||
<taxonomicName id="4C784D62FFE3FFBF5F00326BFE1F4F07" box="[302,459,1876,1900]" class="Reptilia" genus="Malarguesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5F00326BFE1F4F07" box="[302,459,1876,1900]" italics="true" pageId="6" pageNumber="7">Malarguesaurus</emphasis>
|
||||
</taxonomicName>
|
||||
, the neural spine of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5CEF326BFCDE4F00" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[705,778,1876,1899]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5CEF326BFCDE4F00" box="[705,778,1876,1899]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
is rhomboid shaped in dorsal view but is proportionally lower, more elongated, and relatively more robust than the Argentinean species. The neural spine preserved lacks the incipient bifurcation with anteriorly directed tips presented by
|
||||
<taxonomicName id="4C784D62FFE3FFBF5D1731BBFB794CF0" authority="(Carballido et al. 2017)" baseAuthorityName="Carballido" baseAuthorityYear="2017" box="[825,1197,1155,1179]" class="Reptilia" genus="Patagotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5D1731BBFC7E4CF0" box="[825,938,1156,1179]" italics="true" pageId="6" pageNumber="7">Patagotitan</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5D9331BCFB754CF0" author="Carballido JL & Pol D & Otero A" box="[957,1185,1155,1179]" pageId="6" pageNumber="7" refId="ref11873" refString="Carballido JL, Pol D, Otero A et al. A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs. Proceedings Biological Sciences 2017; 284: 20171219. https: // doi. org / 10.1098 / rspb. 2017.1219" type="journal volume" year="2017">
|
||||
Carballido
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A1D31BBFB8A4CF0" box="[1075,1118,1155,1179]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||||
. 2017
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. The isolated neural spine presents a weakly spinoprezygapophyseal lamina (SPRL) while the spinopost-zygapophyseal lamina (SPOL) is comparatively more robust than the SPRL, but the SPOL merges with the rough structure presented on this surface. To our knowledge, this rough structure is unique for titanosaurs, and here is interpreted as an autapomorphy of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5A59307FFB6A4D3C" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1143,1214,1344,1367]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A59307FFB6A4D3C" box="[1143,1214,1344,1367]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
. Lastly, the overall SPRL and SPOL morphology looks like those presented in some derived titanosaurs, such as
|
||||
<taxonomicName id="4C784D62FFE3FFBF5A6B3041FB164DFD" authorityName="Díez Díaz, Mocho, Páramo, Escaso, Marcos-Fernández, Sanz & Ortega" authorityYear="2016" box="[1093,1218,1406,1430]" class="Reptilia" genus="Lohuecotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A6B3041FB164DFD" box="[1093,1218,1406,1430]" italics="true" pageId="6" pageNumber="7">Lohuecotitan</emphasis>
|
||||
</taxonomicName>
|
||||
. However, a marked difference is that in
|
||||
<taxonomicName id="4C784D62FFE3FFBF5DC130A1FBE24DDE" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1007,1078,1438,1461]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5DC130A1FBE24DDE" box="[1007,1078,1438,1461]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
, these laminae are more robust, while those of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5DBD3083FBC64DBF" authorityName="Díez Díaz, Mocho, Páramo, Escaso, Marcos-Fernández, Sanz & Ortega" authorityYear="2016" box="[915,1042,1468,1492]" class="Reptilia" genus="Lohuecotitan" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5DBD3083FBC64DBF" box="[915,1042,1468,1492]" italics="true" pageId="6" pageNumber="7">Lohuecotitan</emphasis>
|
||||
</taxonomicName>
|
||||
present a more complex lamination (
|
||||
<bibRefCitation id="EFE94B10FFE3FFBF5BBC3083FC014D9F" author="Diez Diaz V & Mocho P & Paramo A" pageId="6" pageNumber="7" pagination="49 - 60" refId="ref14970" refString="Diez Diaz V, Mocho P, Paramo A, et al. 2016. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Research, 68, 49 - 60. doi: 10.1016 / j. cretres. 2016.08.001" type="journal article" year="2016">
|
||||
Díez Díaz
|
||||
<emphasis id="B90CEAF3FFE3FFBF5D4030E3FC484D9F" box="[878,924,1500,1524]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC736E1FFE3FFBF5D7B30C4FA9D4E39" blockId="6.[825,1475,1124,1618]" pageId="6" pageNumber="7">
|
||||
The anterior vertebrae of
|
||||
<taxonomicName id="4C784D62FFE3FFBF5A7030C3FB734E78" authorityName="Pereira & Bandeira & Vidal & Ribeiro & R. & Candeiro & Bergqvist" authorityYear="2024" box="[1118,1191,1532,1555]" class="Reptilia" genus="Tiamat" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B90CEAF3FFE3FFBF5A7030C3FB734E78" box="[1118,1191,1532,1555]" italics="true" pageId="6" pageNumber="7">Tiamat</emphasis>
|
||||
</taxonomicName>
|
||||
do not have well-preserved zygapophyses; however, the middle caudal vertebrae have characteristics that warrant biomechanical observations.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
93
data/03/D1/87/03D187F7FFE6FFBA5DC7326BFA824FD9.xml
Normal file
93
data/03/D1/87/03D187F7FFE6FFBA5DC7326BFA824FD9.xml
Normal file
|
|
@ -0,0 +1,93 @@
|
|||
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<mods:namePart id="A7A1060568D1F78DE04A4742CD863973">Pereira, Paulo V L G C</mods:namePart>
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<mods:namePart id="05286050C79500D793993CCE53D854A1">Bandeira, Kamila L N</mods:namePart>
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<mods:namePart id="0969055EE484F0BCCAF0161D65C24E58">Vidal, Luciano S</mods:namePart>
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<mods:namePart id="7ED0DBABCA46635566472C1F4A2C6E91">Ribeiro, Theo B</mods:namePart>
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<mods:namePart id="58A002D9269C3E5798A5ADF1643FECF2">Candeiro, Carlos R dos A</mods:namePart>
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<mods:namePart id="885CEDCEF6D9BD096A26FB18ED8089E2">Bergqvist, Lilian P</mods:namePart>
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<taxonomicName id="43944D572223E364FF0F2352B09FFE06" authority="Memon, Ahmad & Perveen, 2002: 51" authorityName="Memon, Ahmad & Perveen" authorityPageNumber="51" authorityYear="2002" box="[151,681,381,403]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="fabricii">
|
||||
<emphasis id="B6E0EAC62223E364FF0F2352B314FE06" box="[151,290,381,403]" italics="true" pageId="10" pageNumber="361">Halys fabricii</emphasis>
|
||||
<treatmentCitation id="053510C52223E364FEB12351B09FFE06" author="Memon, N. & Ahmad, I. & Perveen, R." box="[297,681,381,403]" page="51" pageId="10" pageNumber="361" year="2002">
|
||||
<bibRefCitation id="E0054B252223E364FEB12351B0B2FE06" author="Memon, N. & Ahmad, I. & Perveen, R." box="[297,644,381,403]" pageId="10" pageNumber="361" pagination="50 - 54" refId="ref13618" refString="Memon, N., Ahmad, I. & Perveen, R. (2002) Redescription of the type species of Halys Fabricius nomen novum for Cimex dentatus Fabricius (Hemiptera: Pentatomidae: Pentatominae: Halyini), a preoccupied name of genus Halys Fabricius with a note on its independent status from Halys serrigera (Westwood). Journal of Natural History and Wildlife, 1, 50 - 54." type="journal article" year="2002">Memon, Ahmad & Perveen, 2002</bibRefCitation>
|
||||
|
|
@ -147,7 +147,7 @@
|
|||
</paragraph>
|
||||
<paragraph id="842B36D42223E364FF0F23B3B3C1FE27" blockId="10.[151,1354,258,464]" box="[151,503,412,434]" pageId="10" pageNumber="361">
|
||||
<treatmentCitationGroup id="A48411FA2223E364FF0F23B3B3C1FE27" box="[151,503,412,434]" pageId="10" pageNumber="361">
|
||||
<taxonomicName id="43944D572223E364FF0F23B3B3C5FE27" authority=": Ghauri, 1988: 80" authorityName="Ghauri" authorityPageNumber="80" authorityYear="1988" box="[151,499,412,434]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="361" phylum="Chordata" rank="species" species="serrigera">
|
||||
<taxonomicName id="43944D572223E364FF0F23B3B3C5FE27" authority=": Ghauri, 1988: 80" authorityName="Ghauri" authorityPageNumber="80" authorityYear="1988" box="[151,499,412,434]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC62223E364FF0F23B3B300FE27" box="[151,310,412,434]" italics="true" pageId="10" pageNumber="361">Halys serrigera</emphasis>
|
||||
:
|
||||
<treatmentCitation id="053510C52223E364FEDC23B3B3C5FE27" author="Ghauri, M. S. K." box="[324,499,412,434]" page="80" pageId="10" pageNumber="361" year="1988">
|
||||
|
|
@ -160,7 +160,7 @@
|
|||
</paragraph>
|
||||
<paragraph id="842B36D42223E364FF0F2395B3DEFE45" blockId="10.[151,1354,258,464]" box="[151,488,442,464]" pageId="10" pageNumber="361">
|
||||
<treatmentCitationGroup id="A48411FA2223E364FF0F2395B3DEFE45" box="[151,488,442,464]" pageId="10" pageNumber="361">
|
||||
<taxonomicName id="43944D572223E364FF0F2395B3D2FE45" authority=": Rider, 2006: 308" authorityName=": Rider" authorityPageNumber="308" authorityYear="2006" box="[151,484,442,464]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="361" phylum="Chordata" rank="species" species="serriger">
|
||||
<taxonomicName id="43944D572223E364FF0F2395B3D2FE45" authority=": Rider, 2006: 308" authorityName=": Rider" authorityPageNumber="308" authorityYear="2006" box="[151,484,442,464]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="serriger">
|
||||
<emphasis id="B6E0EAC62223E364FF0F2395B31FFE45" box="[151,297,442,464]" italics="true" pageId="10" pageNumber="361">Halys serriger</emphasis>
|
||||
:
|
||||
<treatmentCitation id="053510C52223E364FEAF2395B3D2FE45" author="Rider, D. A." box="[311,484,442,464]" page="308" pageId="10" pageNumber="361" year="2006">
|
||||
|
|
@ -256,16 +256,6 @@ by the close examination of terminalia (even in natural condition without macera
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D0EB665C2222E365FF0F26DEB103FAD0" httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="11" pageNumber="362" startId="11.[151,264,1265,1288]" targetBox="[185,1415,375,1192]" targetPageId="11">
|
||||
<paragraph id="842B36D42222E365FF0F26DEB103FAD0" blockId="11.[151,1436,1265,1349]" pageId="11" pageNumber="362">
|
||||
<emphasis id="B6E0EAC62222E365FF0F26DEB365FA92" bold="true" box="[151,339,1265,1288]" pageId="11" pageNumber="362">FIGURES 42–48.</emphasis>
|
||||
<taxonomicName id="43944D572222E365FEC126DCB0B5FA9D" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[345,643,1266,1288]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="362" phylum="Chordata" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC62222E365FEC126DCB3E3FA9D" box="[345,469,1267,1288]" italics="true" pageId="11" pageNumber="362">H. serrigera</emphasis>
|
||||
Westwood, 1837
|
||||
</taxonomicName>
|
||||
—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CC8E655F2222E365FF5F2743B714F95C" pageId="11" pageNumber="362" type="description">
|
||||
<paragraph id="842B36D42222E365FF5F2743B714F95C" blockId="11.[151,1437,1388,2025]" pageId="11" pageNumber="362">
|
||||
<emphasis id="B6E0EAC62222E365FF5F2743B3DBFA10" bold="true" box="[199,493,1388,1413]" pageId="11" pageNumber="362">Measurements (in mm).</emphasis>
|
||||
|
|
@ -887,16 +877,6 @@ and
|
|||
, Veenakumari, K.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D0EB665C2225E362FF0F2483B752F88A" httpUri="https://zenodo.org/record/2644570/files/figure.png" pageId="12" pageNumber="363" startId="12.[151,264,1708,1731]" targetBox="[172,1426,811,1630]" targetPageId="12">
|
||||
<paragraph id="842B36D42225E362FF0F2483B752F88A" blockId="12.[151,1436,1708,1823]" pageId="12" pageNumber="363">
|
||||
<emphasis id="B6E0EAC62225E362FF0F2483B365F957" bold="true" box="[151,339,1708,1731]" pageId="12" pageNumber="363">FIGURES 49–56.</emphasis>
|
||||
<taxonomicName id="43944D572225E362FEC12481B0B5F956" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[345,643,1709,1731]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="363" phylum="Chordata" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC62225E362FEC12481B3E3F956" box="[345,469,1710,1731]" italics="true" pageId="12" pageNumber="363">H. serrigera</emphasis>
|
||||
Westwood, 1837
|
||||
</taxonomicName>
|
||||
—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CC8E655F2225E363FF5F2563B7BDFE45" lastPageId="13" lastPageNumber="364" pageId="12" pageNumber="363" type="discussion">
|
||||
<paragraph id="842B36D42225E363FF5F2563B0AEFE1D" blockId="12.[151,1436,1868,2037]" lastBlockId="13.[151,1436,151,464]" lastPageId="13" lastPageNumber="364" pageId="12" pageNumber="363">
|
||||
<emphasis id="B6E0EAC62225E362FF5F2563B30DF8F0" bold="true" box="[199,315,1868,1893]" pageId="12" pageNumber="363">Remarks.</emphasis>
|
||||
|
|
@ -962,11 +942,11 @@ and
|
|||
<paragraph id="842B36D42224E363FF5F23BBB7BDFE45" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="364">
|
||||
<bibRefCitation id="E0054B252224E363FF5F23BBB361FE38" author="Rider, D. A." box="[199,343,404,429]" pageId="13" pageNumber="364" pagination="233 - 402" refId="ref14017" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Fol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402." type="book chapter" year="2006">Rider (2006)</bibRefCitation>
|
||||
listed this species as
|
||||
<taxonomicName id="43944D572224E363FDD023BBB0DFFE39" authorityName=": Rider" authorityYear="2006" box="[584,745,404,429]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="species" species="serriger">
|
||||
<taxonomicName id="43944D572224E363FDD023BBB0DFFE39" authorityName=": Rider" authorityYear="2006" box="[584,745,404,429]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="serriger">
|
||||
<emphasis id="B6E0EAC62224E363FDD023BBB0DFFE39" box="[584,745,404,429]" italics="true" pageId="13" pageNumber="364">Halys serriger</emphasis>
|
||||
</taxonomicName>
|
||||
. The gender of the generic name
|
||||
<taxonomicName id="43944D572224E363FBFA23BBB695FE38" authorityName="Fabricius" authorityYear="1803" box="[1122,1187,404,429]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="genus">
|
||||
<taxonomicName id="43944D572224E363FBFA23BBB695FE38" authorityName="Fabricius" authorityYear="1803" box="[1122,1187,404,429]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B6E0EAC62224E363FBFA23BBB695FE38" box="[1122,1187,404,429]" italics="true" pageId="13" pageNumber="364">Halys</emphasis>
|
||||
</taxonomicName>
|
||||
, however, is feminine (
|
||||
|
|
|
|||
|
|
@ -1,49 +1,49 @@
|
|||
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|
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<treatment id="0C3D87C22224E37EFF0F276AB760FE1B" ID-GBIF-Taxon="156788548" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B" httpUri="http://treatment.plazi.org/id/0C3D87C22224E37EFF0F276AB760FE1B" lastPageId="16" lastPageNumber="367" pageId="13" pageNumber="364">
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<treatment id="0C3D87C22224E37EFF0F276AB760FE1B" ID-GBIF-Taxon="156788548" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B" httpUri="http://treatment.plazi.org/id/0C3D87C22224E37EFF0F276AB760FE1B" lastPageId="16" lastPageNumber="367" pageId="13" pageNumber="364" scope_family="Pentatomidae" scope_order="Hemiptera">
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<paragraph id="842B36D42224E363FF0F276AB3D3FACA" blockId="13.[151,485,1349,1409]" box="[151,485,1349,1375]" pageId="13" pageNumber="364">
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<heading id="DF6381B82224E363FF0F276AB3D3FACA" bold="true" box="[151,485,1349,1375]" fontSize="11" level="1" pageId="13" pageNumber="364" reason="1">
|
||||
<taxonomicName id="43944D572224E363FF0F276AB3D3FACA" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[151,485,1349,1375]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="species" species="shaista">
|
||||
<taxonomicName id="43944D572224E363FF0F276AB3D3FACA" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[151,485,1349,1375]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC62224E363FF0F276AB3D3FACA" bold="true" box="[151,485,1349,1375]" pageId="13" pageNumber="364">
|
||||
<emphasis id="B6E0EAC62224E363FF0F276AB30EFACA" bold="true" box="[151,312,1349,1375]" italics="true" pageId="13" pageNumber="364">Halys shaista</emphasis>
|
||||
<bibRefCitation id="E0054B252224E363FED8276AB3D3FACA" author="Ghauri, M. S. K." box="[320,485,1349,1375]" pageId="13" pageNumber="364" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
|
||||
|
|
@ -62,7 +62,7 @@
|
|||
<subSubSection id="CC8E655F2224E363FF0F279FB3CEF971" pageId="13" pageNumber="364" type="reference_group">
|
||||
<paragraph id="842B36D42224E363FF0F279FB010FA53" blockId="13.[151,550,1456,1478]" box="[151,550,1456,1478]" pageId="13" pageNumber="364">
|
||||
<treatmentCitationGroup id="A48411FA2224E363FF0F279FB010FA53" box="[151,550,1456,1478]" pageId="13" pageNumber="364">
|
||||
<taxonomicName id="43944D572224E363FF0F279FB3E2FA53" authority="Ghauri, 1988: 78" authorityName="Ghauri" authorityPageNumber="78" authorityYear="1988" box="[151,468,1456,1478]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="species" species="shaista">
|
||||
<taxonomicName id="43944D572224E363FF0F279FB3E2FA53" authority="Ghauri, 1988: 78" authorityName="Ghauri" authorityPageNumber="78" authorityYear="1988" box="[151,468,1456,1478]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC62224E363FF0F279FB316FA53" box="[151,288,1456,1478]" italics="true" pageId="13" pageNumber="364">Halys shaista</emphasis>
|
||||
<treatmentCitation id="053510C52224E363FEBE279FB3E2FA53" author="Ghauri, M. S. K." box="[294,468,1456,1478]" page="78" pageId="13" pageNumber="364" year="1988">
|
||||
<bibRefCitation id="E0054B252224E363FEBE279FB399FA53" author="Ghauri, M. S. K." box="[294,431,1456,1478]" pageId="13" pageNumber="364" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
|
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|
|
@ -117,16 +117,6 @@ Articulatory apparatus with basal plates and support bridge complex, dorsal conn
|
|||
<figureCitation id="1CAF2A512227E360FD4720A0B10EFD3D" box="[735,824,655,680]" captionStart="FIGURES 69–76" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Fig. 72</figureCitation>
|
||||
); aedeagus obliquely bent upwards and shorter than processes of aedeagus.
|
||||
</paragraph>
|
||||
<caption id="D0EB665C2227E360FF0F244CB097F922" httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365" startId="14.[151,264,1635,1658]" targetBox="[205,1394,783,1545]" targetPageId="14">
|
||||
<paragraph id="842B36D42227E360FF0F244CB097F922" blockId="14.[151,1436,1635,1719]" pageId="14" pageNumber="365">
|
||||
<emphasis id="B6E0EAC62227E360FF0F244CB361F9ED" bold="true" box="[151,343,1635,1658]" pageId="14" pageNumber="365">FIGURES 61–68.</emphasis>
|
||||
<taxonomicName id="43944D572227E360FEF8244BB06AF9EC" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[352,604,1635,1657]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="14" pageNumber="365" phylum="Chordata" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC62227E360FEF8244BB3FDF9EC" box="[352,459,1635,1657]" italics="true" pageId="14" pageNumber="365">H. shaista</emphasis>
|
||||
Ghauri, 1988
|
||||
</taxonomicName>
|
||||
—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="842B36D42227E360FF5F24F1B2D9F85A" blockId="14.[151,1437,1758,1999]" pageId="14" pageNumber="365">
|
||||
<emphasis id="B6E0EAC62227E360FF5F24F1B094F962" bold="true" box="[199,674,1758,1783]" pageId="14" pageNumber="365">
|
||||
External female genitalia (
|
||||
|
|
@ -143,22 +133,11 @@ Valvifers VIII roughly triangular, with mesal margin straight, inner posterior a
|
|||
) apical receptacle orbicular with two finger-like processes, directed towards flanges.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D0EB665C2226E361FF0F264EB2FCFB20" httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="15" pageNumber="366" startId="15.[151,264,1121,1144]" targetBox="[181,1411,227,1050]" targetPageId="15">
|
||||
<paragraph id="842B36D42226E361FF0F264EB2FCFB20" blockId="15.[151,1436,1121,1205]" pageId="15" pageNumber="366">
|
||||
<emphasis id="B6E0EAC62226E361FF0F264EB361FBE2" bold="true" box="[151,343,1121,1144]" pageId="15" pageNumber="366">FIGURES 69–76.</emphasis>
|
||||
<taxonomicName id="43944D572226E361FEF8264CB06AFBED" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[352,604,1122,1144]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="366" phylum="Chordata" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC62226E361FEF8264CB34CFBED" box="[352,378,1123,1144]" italics="true" pageId="15" pageNumber="366">H.</emphasis>
|
||||
<emphasis id="B6E0EAC62226E361FE1B264DB3FDFBED" box="[387,459,1122,1144]" italics="true" pageId="15" pageNumber="366">shaista</emphasis>
|
||||
Ghauri, 1988
|
||||
</taxonomicName>
|
||||
—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CC8E655F2226E361FF5F26CDB608FA1F" pageId="15" pageNumber="366" type="diagnosis">
|
||||
<paragraph id="842B36D42226E361FF5F26CDB608FA1F" blockId="15.[151,1437,1250,2031]" pageId="15" pageNumber="366">
|
||||
<emphasis id="B6E0EAC62226E361FF5F26CDB3F8FB6E" bold="true" box="[199,462,1250,1275]" pageId="15" pageNumber="366">Differential diagnosis.</emphasis>
|
||||
This species can be differentiated from other species of
|
||||
<taxonomicName id="43944D572226E361FBDF26CDB6BEFB6E" authorityName="Fabricius" authorityYear="1803" box="[1095,1160,1250,1275]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="366" phylum="Chordata" rank="genus">
|
||||
<taxonomicName id="43944D572226E361FBDF26CDB6BEFB6E" authorityName="Fabricius" authorityYear="1803" box="[1095,1160,1250,1275]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B6E0EAC62226E361FBDF26CDB6BEFB6E" box="[1095,1160,1250,1275]" italics="true" pageId="15" pageNumber="366">Halys</emphasis>
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||||
</taxonomicName>
|
||||
by the peculiar shape of paramere and absence of inverted U-shaped process on ventral rim of genital capsule but with a median tumescence. The parameral crown is apically angulate, possessing a single tooth around the middle of its dorsolateral magin, triangular valvifers VIII with posterior margin nearly straight or slightly convex (
|
||||
|
|
|
|||
|
|
@ -1,197 +0,0 @@
|
|||
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|
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: 180
|
||||
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||||
.
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species by subsequent designation (
|
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</treatmentCitation>
|
||||
):
|
||||
<taxonomicName id="43944D57222BE36CFBD027C2B748F996" authority="Fabricius, 1803" authorityName=": Fabricius" authorityYear="1803" box="[1096,1406,1517,1539]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="353" phylum="Chordata" rank="species" species="dentata">
|
||||
<emphasis id="B6E0EAC6222BE36CFBD027C2B6E1F996" box="[1096,1239,1517,1539]" italics="true" pageId="2" pageNumber="353">Halys dentata</emphasis>
|
||||
<bibRefCitation id="E0054B25222BE36CFB4727C2B748F996" author="Fabricius, J. C." box="[1247,1406,1517,1539]" pageId="2" pageNumber="353" refId="ref13060" refString="Fabricius, J. C. (1803) Systema Rhyngotorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae, x + 335 pp." type="book" year="1803">Fabricius, 1803</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(=
|
||||
<taxonomicName id="43944D57222BE36CFF5F2423B02CF9B7" authority="Westwood, 1837" authorityName="Westwood" authorityYear="1837" box="[199,538,1548,1570]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="353" phylum="Chordata" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC6222BE36CFF5F2423B350F9B7" box="[199,358,1548,1570]" italics="true" pageId="2" pageNumber="353">Halys serrigera</emphasis>
|
||||
<bibRefCitation id="E0054B25222BE36CFEF52423B02CF9B7" author="Westwood, J. O." box="[365,538,1548,1570]" pageId="2" pageNumber="353" refId="ref14641" refString="Westwood, J. O. (1837) A Catalogue of Hemiptera in the collection of the Rev. F. W. Hope, M. A. with short Latin diagnoses of the new species. Pt. 1. Printed by J. C. Bridgewater, London, 46 pp. [Hope, F. W. (Eds.)]" type="book" year="1837">Westwood, 1837</bibRefCitation>
|
||||
</taxonomicName>
|
||||
).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CC8E655F222BE36CFF0F2461B0DBF8F6" pageId="2" pageNumber="353" type="diagnosis">
|
||||
<paragraph id="842B36D4222BE36CFF0F2461B0DBF8F6" blockId="2.[151,1436,1614,2035]" pageId="2" pageNumber="353">
|
||||
<emphasis id="B6E0EAC6222BE36CFF0F2461B397F9F2" bold="true" box="[151,417,1614,1639]" pageId="2" pageNumber="353">Differential diagnosis.</emphasis>
|
||||
Members of this genus are usually dull coloured, characteristic for their elongate head, with at least one pair of teeth on lateral margins of head, round apex of bucculae (lacking tooth), elongate labium resting in the median longitudinal groove on the ventral side of the abdomen, well exposed connexivum and spoutshaped (crescent-shaped) peritreme (
|
||||
<figureCitation id="1CAF2A51222BE36CFDAC2495B0B0F946" box="[564,646,1722,1747]" captionStart="FIGURES 91–93" captionStartId="21.[151,264,1372,1395]" captionTargetBox="[170,1394,494,1295]" captionTargetId="figure@21.[151,1435,443,1351]" captionTargetPageId="21" captionText="FIGURES 91–93. FIGURE 91. External scent efferent system showing peritreme and associated structures. FIGURES 92– 93. Genital capsule (caudal). 92, H. mudigerensis sp. nov.; 93, H. serrigera Westwood, 1837. Lettering: E—evaporatorium; Mf—median furrow; Mp—Metepimeral pseudosuture; Ms—metathoracic spiracle with filter process; O—Ostiole; Op— Ostiolar plate; P—Peritreme; t—tooth; ts—tooth like serrations." httpUri="https://zenodo.org/record/2644582/files/figure.png" pageId="2" pageNumber="353">Fig. 91</figureCitation>
|
||||
), which is usually not extending beyond middle of metapleuron.
|
||||
<collectingRegion id="4650F836222BE36CFAFA2495B7AAF946" box="[1378,1436,1722,1747]" country="Maldives" name="Maale" pageId="2" pageNumber="353">Male</collectingRegion>
|
||||
genitalia with genital capsule quadrangular, paramere roughly C-shaped with crown expanded, with or without ridges, provided with one or two teeth in various positions; inflated phallus with a pair of process of aedeagus, either free or fused; usually with two pairs of membranous conjunctival processes. Laterotergite IX of female terminalia is distinctly shorter than apex of abdomen.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CC8E655F222BE36DFF5F2541B188F891" lastPageId="3" lastPageNumber="354" pageId="2" pageNumber="353" type="description">
|
||||
<paragraph id="842B36D4222BE36DFF5F2541B32EFEF1" blockId="2.[151,1436,1614,2035]" lastBlockId="3.[151,1437,151,1941]" lastPageId="3" lastPageNumber="354" pageId="2" pageNumber="353">
|
||||
<emphasis id="B6E0EAC6222BE36CFF5F2541B342F812" bold="true" box="[199,372,1902,1927]" pageId="2" pageNumber="353">Redescription.</emphasis>
|
||||
<emphasis id="B6E0EAC6222BE36CFEE32541B381F812" box="[379,439,1902,1927]" italics="true" pageId="2" pageNumber="353">Head</emphasis>
|
||||
(
|
||||
<figureCitation id="1CAF2A51222BE36CFE5E2541B073F812" box="[454,581,1902,1927]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="2" pageNumber="353">Figs 10–16</figureCitation>
|
||||
) flattened dorsally, coplanar with pronotum in lateral view; lateral margin of mandibular plates with one or two pairs of teeth, first pair just in front of compound eyes, second pair (if present) at anterior third; anterior third of mandibular plates abruptly narrowing, extreme apex rounded, not meeting in front of clypeus; clypeus nearly of equal width throughout and open anteriorly, usually slightly longer than mandibular plates, sometimes of equal length. Antenniferous tubercles (
|
||||
<figureCitation id="1CAF2A51222AE36DFCA722B8B1A6FF25" box="[831,912,151,176]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="3" pageNumber="354">Fig. 11</figureCitation>
|
||||
) small, visible from above along with base of antennae. Antennae five segmented, slender. Antennal segments of variable length, segment I cylindrical, shortest and stoutest, not reaching apex of head, remaining segments slender, nearly of uniform thickness. Bucculae (
|
||||
<figureCitation id="1CAF2A51222AE36DFAF722F0B282FE88" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="3" pageNumber="354">Fig. 16</figureCitation>
|
||||
) rounded anteriorly. Labrum (
|
||||
<figureCitation id="1CAF2A51222AE36DFD8C232BB05FFE89" box="[532,617,260,285]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="3" pageNumber="354">Fig. 12</figureCitation>
|
||||
) narrow. Labial segment I stoutest, not surpassing posterior margin of bucculae, apex of labium surpassing anterior margin of ventrite V to reaching or surpassing posterior margin of ventrite VI.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F235FB0BAFDF5" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F235FB30AFE12" bold="true" box="[199,316,368,391]" italics="true" pageId="3" pageNumber="354">Pronotum</emphasis>
|
||||
(
|
||||
<figureCitation id="1CAF2A51222AE36DFEC82340B39FFE1D" box="[336,425,367,392]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="3" pageNumber="354">Fig. 17</figureCitation>
|
||||
). Pronotum more or less flattened; anterior half of pronotal disc slightly sloping and sometimes with prominent undulations or pit-like depressions; posterior half slightly convex or elevated. Anterior pronotal margin deeply incised to accommodate base of head, lacking collar; anterolateral angles rectangular, unarmed; lateral margin nearly straight or sometimes slightly concave medially, serrated,serrations prominent on anterolateral margin, compared to posterolateral margin. Humeri angulate, sometimes broadly angulate, prominent and slightly elevated; posterolateral margin of pronotum obliquely straight or medially slightly convex; posterior margin straight or slightly concave medially.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F2043B61FFD58" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2043B30AFD10" bold="true" box="[199,316,620,645]" italics="true" pageId="3" pageNumber="354">Scutellum</emphasis>
|
||||
(
|
||||
<figureCitation id="1CAF2A51222AE36DFED22043B3ADFD10" box="[330,411,620,645]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="3" pageNumber="354">Fig. 15</figureCitation>
|
||||
) elongate triangular, longer than width at base, sides slightly convex in frenal portion; distal third abruptly narrowed; much shorter than anterodistal angles of corium; apex narrowly rounded; disc more or less uniformly and slightly convex to flat, sometimes slightly elevated at basal region.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F20F8B0EAFC80" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F20F8B377FD65" bold="true" box="[199,321,727,752]" italics="true" pageId="3" pageNumber="354">Hemelytra</emphasis>
|
||||
Clavus narrowly triangular, corium laterally slightly convex, gradually widening posteriad. Membrane with reticulate venation, length variable.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F2130B6F9FC5D" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2130B006FCA2" bold="true" box="[199,560,799,824]" italics="true" pageId="3" pageNumber="354">Thoracic pleuron and sternum</emphasis>
|
||||
Prosternum and metasternum lacking carina, with indistinct median groove. Mesosternum lacking prominent longitudinal groove or keel but with obsolete longitudinal carina. External scent efferent system with peritreme spout-shaped (crescent-shaped) (
|
||||
<figureCitation id="1CAF2A51222AE36DFCE82148B1F5FC15" box="[880,963,871,896]" captionStart="FIGURES 91–93" captionStartId="21.[151,264,1372,1395]" captionTargetBox="[170,1394,494,1295]" captionTargetId="figure@21.[151,1435,443,1351]" captionTargetPageId="21" captionText="FIGURES 91–93. FIGURE 91. External scent efferent system showing peritreme and associated structures. FIGURES 92– 93. Genital capsule (caudal). 92, H. mudigerensis sp. nov.; 93, H. serrigera Westwood, 1837. Lettering: E—evaporatorium; Mf—median furrow; Mp—Metepimeral pseudosuture; Ms—metathoracic spiracle with filter process; O—Ostiole; Op— Ostiolar plate; P—Peritreme; t—tooth; ts—tooth like serrations." httpUri="https://zenodo.org/record/2644582/files/figure.png" pageId="3" pageNumber="354">Fig. 91</figureCitation>
|
||||
) usually not extending beyond middle of metapleuron; evaporatorium developed as roughly triangular and slightly elevated patch on metapleuron, encompassing ostiole and peritreme. Metathoracic spiracle long and well visible in ventral view.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F21FBB06AFBA0" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F21FBB2C8FC7E" bold="true" box="[199,254,980,1003]" italics="true" pageId="3" pageNumber="354">Legs</emphasis>
|
||||
elongate, slender; femora cylindrical, unarmed. All tibiae unarmed, more slender than adjoining femora, outer surface with median longitudinal groove. All tarsi with segment II shortest, III as long as I; all tarsal segments dorsally regularly rounded, not grooved.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F2610B333FB7D" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2610B380FBCD" bold="true" box="[199,438,1087,1112]" italics="true" pageId="3" pageNumber="354">Pregenital abdomen</emphasis>
|
||||
. Connexivum wide, fully exposed dorsally, each ventrite with posterolateral angles angulate, not toothed. Abdominal venter convex, with prominent median, longitudinal groove. Ventrite II lacking spine or tubercle but with median longitudinal groove. Spiracle on ventrite II nearly exposed, only anterior margin covered by metapleuron, sometimes unexposed. A pair of trichobothria posteriad of spiracular line on each side of abdomen.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F26DBB126F9C5" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F26DBB343FA98" bold="true" box="[199,373,1268,1293]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F26DBB35BFA98" bold="true" box="[199,365,1268,1293]" italics="true" pageId="3" pageNumber="354">
|
||||
<collectingRegion id="4650F836222AE36DFF5F26DBB334FA98" box="[199,258,1268,1293]" country="Maldives" name="Maale" pageId="3" pageNumber="354">Male</collectingRegion>
|
||||
genitalia
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<emphasis id="B6E0EAC6222AE36DFEE426DBB01BFA98" box="[380,557,1268,1293]" italics="true" pageId="3" pageNumber="354">Genital capsule</emphasis>
|
||||
roughly quadrangular; dorsal rim widely and concavely excavated; infoldings sublaterally with black, sclerotized teeth or minute tooth-like serrations; ventral rim concavely excavated; infoldings with or without a short, stout and narrow, inverted U-shaped (roughly tongue-like) median process (
|
||||
<figureCitation id="1CAF2A51222AE36DFAF22713B280FAED" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1839,1862]" captionTargetBox="[181,1418,993,1795]" captionTargetId="figure@7.[151,1436,973,1818]" captionTargetPageId="7" captionText="FIGURES 21–28. Cimex dentatus Fabricius, 1775 (Lectotype)—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown." httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="3" pageNumber="354">Figs 22</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFF58274FB2E9FAED" box="[192,223,1376,1400]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="3" pageNumber="354">30</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFF72274FB33EFAED" box="[234,264,1376,1400]" captionStart="FIGURES 42–48" captionStartId="11.[151,264,1265,1288]" captionTargetBox="[185,1415,375,1192]" captionTargetId="figure@11.[151,1435,337,1245]" captionTargetPageId="11" captionText="FIGURES 42–48. H. serrigera Westwood, 1837—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="3" pageNumber="354">43</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFE8B274FB305FAED" box="[275,307,1376,1400]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="3" pageNumber="354">78</figureCitation>
|
||||
).
|
||||
<emphasis id="B6E0EAC6222AE36DFEDD274EB382FAED" box="[325,436,1377,1400]" italics="true" pageId="3" pageNumber="354">Paramere</emphasis>
|
||||
roughly C-shaped, with short stem and expanded plate-like apodeme (
|
||||
<figureCitation id="1CAF2A51222AE36DFB5D2770B72DFAED" box="[1221,1307,1375,1400]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1839,1862]" captionTargetBox="[181,1418,993,1795]" captionTargetId="figure@7.[151,1436,973,1818]" captionTargetPageId="7" captionText="FIGURES 21–28. Cimex dentatus Fabricius, 1775 (Lectotype)—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown." httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="3" pageNumber="354">Figs 26</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFABE274FB773FAED" box="[1318,1349,1376,1400]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="3" pageNumber="354">32</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFAC82770B758FAED" box="[1360,1390,1375,1400]" captionStart="FIGURES 42–48" captionStartId="11.[151,264,1265,1288]" captionTargetBox="[185,1415,375,1192]" captionTargetId="figure@11.[151,1435,337,1245]" captionTargetPageId="11" captionText="FIGURES 42–48. H. serrigera Westwood, 1837—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="3" pageNumber="354">45</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFAE12770B7AEFAED" box="[1401,1432,1375,1400]" captionStart="FIGURES 61–68" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 61–68. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="3" pageNumber="354">64</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFF0F27ABB281FA09" box="[151,183,1412,1436]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="3" pageNumber="354">80</figureCitation>
|
||||
). Crown with small, blunt and black tooth, sometimes continued as sclerotized, black ridge, position of tooth on crown varies.
|
||||
<emphasis id="B6E0EAC6222AE36DFEAC2788B3BFFA55" box="[308,393,1447,1472]" italics="true" pageId="3" pageNumber="354">Phallus</emphasis>
|
||||
: articulatory apparatus with 1+1 roughly oval or irregular or quadrangular capitate processes (
|
||||
<figureCitation id="1CAF2A51222AE36DFF0727E3B314FA70" box="[159,290,1484,1509]" captionStart="FIGURES 94–99" captionStartId="22.[151,264,1273,1296]" captionTargetBox="[170,1415,467,1204]" captionTargetId="figure@22.[151,1435,409,1253]" captionTargetPageId="22" captionText="FIGURES 94–99. FIGURES 94–95. Genital capsule (caudal). 94, H. shaista Ghauri, 1988; 95, H. sulcata (Thunberg, 1783). Lettering: t—tooth; ts—tooth like serrations. FIGURES 96–99. Capitate processes. 96, H. mudigerensis sp. nov.; 97, H. serrigera Westwood, 1837; 98, H. shaista Ghauri, 1988; 99, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644584/files/figure.png" pageId="3" pageNumber="354">Figs 96–99</figureCitation>
|
||||
). Phallosoma sclerotized, constricted towards both ends, broadest medially and concave dorsally; two pairs of membranous conjunctival processes, dorsal pair with or without bifid apex, ventral pair shorter than or subequal to dorsal pair and apically rounded. Processes of aedeagus sclerotized, free or fused along midline at proximal half encompassing short, sclerotized aedeagus.
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4222AE36DFF5F2473B188F891" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2473B02CF9E0" box="[199,538,1628,1653]" italics="true" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2473B3A3F9E0" bold="true" box="[199,405,1628,1653]" italics="true" pageId="3" pageNumber="354">Female genitalia.</emphasis>
|
||||
Terminalia
|
||||
</emphasis>
|
||||
: valvifers VIII usually quadrangular, sometimes roughly triangular. Valvulae VIII with apical (= caudal) margin either uniformly rounded (
|
||||
<figureCitation id="1CAF2A51222AE36DFCCF2450B186F90D" box="[855,944,1663,1688]" captionStart="FIGURES 49–56" captionStartId="12.[151,264,1708,1731]" captionTargetBox="[172,1426,811,1630]" captionTargetId="figure@12.[151,1435,780,1688]" captionTargetPageId="12" captionText="FIGURES 49–56. H. serrigera Westwood, 1837—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644570/files/figure.png" pageId="3" pageNumber="354">Figs 54</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFC2524AFB1ECF90D" box="[957,986,1664,1688]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="3" pageNumber="354">90</figureCitation>
|
||||
) or slightly concave medially (
|
||||
<figureCitation id="1CAF2A51222AE36DFAA72450B7A1F90D" box="[1343,1431,1663,1688]" captionStart="FIGURES 36–41" captionStartId="9.[151,264,1900,1923]" captionTargetBox="[209,1419,730,1794]" captionTargetId="figure@9.[151,1435,701,1851]" captionTargetPageId="9" captionText="FIGURES 36–41. H. mudigerensis sp. nov.—male and female genitalia. 36, phallus (dorsal); 37, phallus (ventral); 38, phallus (lateral); 39, spermatheca; 40, female terminalia; 41,valvulae VIII. Lettering: dcp—dorsal conjunctival processes; lt VIIIlaterotergite VIII; lt IX—laterotergite IX; pa—processes of aedeagus; sp—spermathecal pump; vcp—ventral conjunctival process; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644566/files/figure.png" pageId="3" pageNumber="354">Figs 41</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51222AE36DFF0F248BB281F928" box="[151,183,1700,1725]" captionStart="FIGURES 69–76" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="3" pageNumber="354">76</figureCitation>
|
||||
). Valvifers IX fused to a single median transverse sclerite. Laterotergite IX distinctly shorter than apex of abdomen, caudal lateral margin of laterotergite VIII smooth and convex. Spermathecal duct expanded into single bulb-like dilation; apical receptacle orbicular with finger-like processes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CC8E655F222AE36DFF5F2520B1B6F800" pageId="3" pageNumber="354" type="discussion">
|
||||
<paragraph id="842B36D4222AE36DFF5F2520B1B6F800" blockId="3.[151,1437,151,1941]" pageId="3" pageNumber="354">
|
||||
<emphasis id="B6E0EAC6222AE36DFF5F2520B30DF8BD" bold="true" box="[199,315,1807,1832]" pageId="3" pageNumber="354">Remarks.</emphasis>
|
||||
Most of the structural characters such as length of mandibular plates with respect to clypeus, teeth in front of eyes, lengths of antennal segments, length of labium, serrations on lateral margins of pronotum, number and length of finger-like processes on spermathecal pump etc., show continuous variation within a given species. External colouration is also highly variable within a given species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,50 +1,50 @@
|
|||
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<treatment id="0C3D87C2222EE367FF0F22B8B705FDF5" ID-GBIF-Taxon="156788546" LSID="urn:lsid:zoobank.org:act:30D2DF53-8D03-4415-B3AD-7D28147193FC" httpUri="http://treatment.plazi.org/id/0C3D87C2222EE367FF0F22B8B705FDF5" lastPageId="9" lastPageNumber="360" pageId="7" pageNumber="358">
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<taxonomicName id="43944D57222EE369FF0F22B8B3B3FF24" authority="Salini, 2019" authorityName="Salini" authorityYear="2019" box="[151,389,151,177]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="358" phylum="Chordata" rank="species" species="mudigerensis" status="sp. nov.">
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<taxonomicName id="43944D57222EE369FF0F22B8B3B3FF24" authority="Salini, 2019" authorityName="Salini" authorityYear="2019" box="[151,389,151,177]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="mudigerensis" status="sp. nov.">
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<emphasis id="B6E0EAC6222EE369FF0F22B8B3B3FF24" bold="true" box="[151,389,151,177]" italics="true" pageId="7" pageNumber="358">Halys mudigerensis</emphasis>
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</taxonomicName>
|
||||
<taxonomicNameLabel id="ADD357BD222EE369FE1422B7B3DCFF27" box="[396,490,152,178]" pageId="7" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
|
|
@ -86,16 +86,6 @@ Dorsum including head covered with dense, irregular dark brown to black puncture
|
|||
<emphasis id="B6E0EAC6222EE369FF5F2131B30FFCA2" bold="true" box="[199,313,798,823]" italics="true" pageId="7" pageNumber="358">Structure.</emphasis>
|
||||
Labium elongate, reaching or surpassing posterior margin of ventrite VI. Lateral margin of pronotum slightly concave and posterior margins of pronotum slightly concave medially. Humeri broadly triangular and elevated. Disc of pronotum with strong undulations or pit-like depressions on anterior half and posterior half of pronotal disc, elevated. Disc of scutellum slightly elevated basally.
|
||||
</paragraph>
|
||||
<caption id="D0EB665C222EE369FF0F2500B6C7F816" httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="7" pageNumber="358" startId="7.[151,264,1839,1862]" targetBox="[181,1418,993,1795]" targetPageId="7">
|
||||
<paragraph id="842B36D4222EE369FF0F2500B6C7F816" blockId="7.[151,1436,1839,1923]" pageId="7" pageNumber="358">
|
||||
<emphasis id="B6E0EAC6222EE369FF0F2500B36FF8D0" bold="true" box="[151,345,1839,1862]" pageId="7" pageNumber="358">FIGURES 21–28.</emphasis>
|
||||
<taxonomicName id="43944D57222EE369FEFC251EB100F8D3" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[356,822,1840,1862]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="dentatus">
|
||||
<emphasis id="B6E0EAC6222EE369FEFC251EB030F8D3" box="[356,518,1840,1862]" italics="true" pageId="7" pageNumber="358">Cimex dentatus</emphasis>
|
||||
Fabricius, 1775 (Lectotype)
|
||||
</taxonomicName>
|
||||
—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="842B36D4222EE369FF5F259FB7AAF878" blockId="7.[151,1436,1968,2029]" pageId="7" pageNumber="358">
|
||||
<emphasis id="B6E0EAC6222EE369FF5F259FB0A9F85C" bold="true" box="[199,671,1968,1993]" pageId="7" pageNumber="358">
|
||||
External male genitalia (
|
||||
|
|
@ -128,18 +118,6 @@ Articulatory apparatus with basal plates and support bridge complex, dorsal conn
|
|||
<figureCitation id="1CAF2A512221E366FEDC20A0B3A1FD3D" box="[324,407,655,680]" captionStart="FIGURES 36–41" captionStartId="9.[151,264,1900,1923]" captionTargetBox="[209,1419,730,1794]" captionTargetId="figure@9.[151,1435,701,1851]" captionTargetPageId="9" captionText="FIGURES 36–41. H. mudigerensis sp. nov.—male and female genitalia. 36, phallus (dorsal); 37, phallus (ventral); 38, phallus (lateral); 39, spermatheca; 40, female terminalia; 41,valvulae VIII. Lettering: dcp—dorsal conjunctival processes; lt VIIIlaterotergite VIII; lt IX—laterotergite IX; pa—processes of aedeagus; sp—spermathecal pump; vcp—ventral conjunctival process; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644566/files/figure.png" pageId="8" pageNumber="359">Fig. 38</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="D0EB665C2221E366FF0F24B4B619F97A" httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="8" pageNumber="359" startId="8.[151,264,1691,1714]" targetBox="[188,1424,813,1586]" targetPageId="8">
|
||||
<paragraph id="842B36D42221E366FF0F24B4B619F97A" blockId="8.[151,1435,1691,1775]" pageId="8" pageNumber="359">
|
||||
<emphasis id="B6E0EAC62221E366FF0F24B4B361F924" bold="true" box="[151,343,1691,1714]" pageId="8" pageNumber="359">FIGURES 29–35.</emphasis>
|
||||
<taxonomicName id="43944D572221E366FEC724B2B031F927" authority="Salini, 2019" authorityName="Salini" authorityYear="2019" box="[351,519,1692,1714]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="359" phylum="Chordata" rank="species" species="mudigerensis" status="sp. nov.">
|
||||
<emphasis id="B6E0EAC62221E366FEC724B2B031F927" box="[351,519,1692,1714]" italics="true" pageId="8" pageNumber="359">H. mudigerensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ADD357BD2221E366FD8824B3B055F924" box="[528,611,1692,1713]" pageId="8" pageNumber="359" rank="species">
|
||||
<emphasis id="B6E0EAC62221E366FD8824B3B055F924" bold="true" box="[528,611,1692,1713]" pageId="8" pageNumber="359">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="842B36D42221E367FF5F2539B386FF6D" blockId="8.[151,1436,1814,2019]" lastBlockId="9.[151,1437,151,608]" lastPageId="9" lastPageNumber="360" pageId="8" pageNumber="359">
|
||||
<emphasis id="B6E0EAC62221E366FF5F2539B0ADF8BA" bold="true" box="[199,667,1814,1839]" pageId="8" pageNumber="359">
|
||||
External female genitalia (
|
||||
|
|
|
|||
|
|
@ -1,49 +1,49 @@
|
|||
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<taxonomicName id="43944D572239E37EFF0F23CEB02DFE6E" authority="(Thunberg, 1783)" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[151,539,481,507]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcata">
|
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<taxonomicName id="43944D572239E37EFF0F23CEB02DFE6E" authority="(Thunberg, 1783)" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[151,539,481,507]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sulcata">
|
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<emphasis id="B6E0EAC62239E37EFF0F23CEB02DFE6E" bold="true" box="[151,539,481,507]" pageId="16" pageNumber="367">
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<emphasis id="B6E0EAC62239E37EFF0F23CEB30CFE6E" bold="true" box="[151,314,481,507]" italics="true" pageId="16" pageNumber="367">Halys sulcata</emphasis>
|
||||
(
|
||||
|
|
@ -72,7 +72,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="43944D572239E37EFF0F2044B3CBFD14" authority=": Kirkaldy, 1909: 196" authorityName="Kirkaldy" authorityPageNumber="196" authorityYear="1909" box="[151,509,619,641]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcata">
|
||||
<taxonomicName id="43944D572239E37EFF0F2044B3CBFD14" authority=": Kirkaldy, 1909: 196" authorityName="Kirkaldy" authorityPageNumber="196" authorityYear="1909" box="[151,509,619,641]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sulcata">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F2044B317FD14" box="[151,289,619,641]" italics="true" pageId="16" pageNumber="367">Halys sulcata</emphasis>
|
||||
:
|
||||
<treatmentCitation id="053510C52239E37EFEB72044B3CBFD14" author="Kirkaldy, G. W." box="[303,509,619,641]" page="196" pageId="16" pageNumber="367" year="1909">
|
||||
|
|
@ -81,7 +81,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="43944D572239E37EFF0F20A6B3DAFD0A" authority="Chopra, 1974: 477" authorityName="Chopra" authorityPageNumber="477" authorityYear="1974" box="[151,492,649,671]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="magnus">
|
||||
<taxonomicName id="43944D572239E37EFF0F20A6B3DAFD0A" authority="Chopra, 1974: 477" authorityName="Chopra" authorityPageNumber="477" authorityYear="1974" box="[151,492,649,671]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="magnus">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F20A6B311FD0A" box="[151,295,649,671]" italics="true" pageId="16" pageNumber="367">Halys magnus</emphasis>
|
||||
<treatmentCitation id="053510C52239E37EFEB520A6B3DAFD0A" author="Chopra, N. P." box="[301,492,649,671]" page="477" pageId="16" pageNumber="367" year="1974">
|
||||
<bibRefCitation id="E0054B252239E37EFEB520A6B38CFD0A" author="Chopra, N. P." box="[301,442,649,671]" pageId="16" pageNumber="367" pagination="473 - 479" refId="ref12757" refString="Chopra, N. P. (1974) Studies on the genus Halys (Hemiptera: Pentatomidae), Oriental Insects, 8 (4), 473 - 479. https: // doi. org / 10.1080 / 00305316.1974.10434879" type="journal article" year="1974">Chopra, 1974</bibRefCitation>
|
||||
|
|
@ -94,7 +94,7 @@
|
|||
: 81
|
||||
</treatmentCitation>
|
||||
).
|
||||
<taxonomicName id="43944D572239E37EFF0F2087B002FD2B" authority="Abbasi & Ahmad, 1976: 26" authorityName="Abbasi & Ahmad" authorityPageNumber="26" authorityYear="1976" box="[151,564,680,702]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="qadrii">
|
||||
<taxonomicName id="43944D572239E37EFF0F2087B002FD2B" authority="Abbasi & Ahmad, 1976: 26" authorityName="Abbasi & Ahmad" authorityPageNumber="26" authorityYear="1976" box="[151,564,680,702]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="qadrii">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F2087B320FD2B" box="[151,278,680,702]" italics="true" pageId="16" pageNumber="367">Halys qadrii</emphasis>
|
||||
<treatmentCitation id="053510C52239E37EFE842087B002FD2B" author="Abbasi, Q. A. & Ahmad, I." box="[284,564,680,702]" page="26" pageId="16" pageNumber="367" year="1976">
|
||||
<bibRefCitation id="E0054B252239E37EFE842087B027FD2B" author="Abbasi, Q. A. & Ahmad, I." box="[284,529,680,702]" pageId="16" pageNumber="367" pagination="26 - 31" refId="ref12358" refString="Abbasi, Q. A. & Ahmad, I. (1976) A new species of the genus Halys Fabr. (Pentatomidea: Halyini) from East Bengal with notes on its genitalia and their bearing on classification. Karachi University Journal of Science, 4 (1 - 2), 26 - 31." type="journal article" year="1976">Abbasi & Ahmad, 1976</bibRefCitation>
|
||||
|
|
@ -107,7 +107,7 @@
|
|||
: 81
|
||||
</treatmentCitation>
|
||||
).
|
||||
<taxonomicName id="43944D572239E37EFF0F20E9B3DEFD49" authority=": Ghauri, 1988: 81" authorityName="Ghauri" authorityPageNumber="81" authorityYear="1988" box="[151,488,710,732]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcatus">
|
||||
<taxonomicName id="43944D572239E37EFF0F20E9B3DEFD49" authority=": Ghauri, 1988: 81" authorityName="Ghauri" authorityPageNumber="81" authorityYear="1988" box="[151,488,710,732]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sulcatus">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F20E9B31DFD49" box="[151,299,710,732]" italics="true" pageId="16" pageNumber="367">Halys sulcatus</emphasis>
|
||||
:
|
||||
<treatmentCitation id="053510C52239E37EFEA120E9B3DEFD49" author="Ghauri, M. S. K." box="[313,488,710,732]" page="81" pageId="16" pageNumber="367" year="1988">
|
||||
|
|
@ -116,7 +116,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="43944D572239E37EFF0F20CAB3D3FD6E" authority=": Rider, 2006: 308" authorityName="Rider" authorityPageNumber="308" authorityYear="2006" box="[151,485,741,763]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sulcatus">
|
||||
<taxonomicName id="43944D572239E37EFF0F20CAB3D3FD6E" authority=": Rider, 2006: 308" authorityName="Rider" authorityPageNumber="308" authorityYear="2006" box="[151,485,741,763]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sulcatus">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F20CAB31DFD6E" box="[151,299,741,763]" italics="true" pageId="16" pageNumber="367">Halys sulcatus</emphasis>
|
||||
:
|
||||
<treatmentCitation id="053510C52239E37EFEA120CAB3D3FD6E" author="Rider, D. A." box="[313,485,741,763]" page="308" pageId="16" pageNumber="367" year="2006">
|
||||
|
|
@ -125,7 +125,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="43944D572239E37EFF0F212BB05CFC8F" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[151,618,772,794]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="sindilla">
|
||||
<taxonomicName id="43944D572239E37EFF0F212BB05CFC8F" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[151,618,772,794]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="sindilla">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F212BB312FC8F" box="[151,292,772,794]" italics="true" pageId="16" pageNumber="367">Halys sindilla</emphasis>
|
||||
<bibRefCitation id="E0054B252239E37EFEB3212AB05CFC8F" author="Memon, N. & Meier, R. & Manan, A. & Su, K. F. - Y." box="[299,618,772,794]" pageId="16" pageNumber="367" pagination="703 - 710" refId="ref13758" refString="Memon, N., Meier, R., Manan, A. & Su, K. F. - Y. (2006) On the use of DNA sequences for determining the species limits of a polymorphic new species in the stink bug genus Halys (Heteroptera: Pentatomidae) from Pakistan. Systematic Entomology, 31 (4), 703 - 710. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00350. x" type="journal article" year="2006">Memon, Meier & Manan, 2006</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -133,7 +133,7 @@
|
|||
<taxonomicNameLabel id="ADD357BD2239E37EFDED212CB0E0FC8C" box="[629,726,771,794]" pageId="16" pageNumber="367">
|
||||
<emphasis id="B6E0EAC62239E37EFDED212CB0E0FC8C" bold="true" box="[629,726,771,794]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
<taxonomicName id="43944D572239E37EFF0F210CB057FCAC" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" box="[151,609,803,825]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="spinosa">
|
||||
<taxonomicName id="43944D572239E37EFF0F210CB057FCAC" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" box="[151,609,803,825]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="spinosa">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F210CB313FCAC" box="[151,293,803,825]" italics="true" pageId="16" pageNumber="367">Halys spinosa</emphasis>
|
||||
<bibRefCitation id="E0054B252239E37EFEB4210CB057FCAC" author="Shaikh, A. M. & Memon, N. & Shah, M. A." box="[300,609,803,825]" pageId="16" pageNumber="367" pagination="157 - 160" refId="ref14285" refString="Shaikh, A. M., Memon, N. & Shah, M. A. (2011) A new halyine species of Halys Fabricius (Hemiptera, Pentatomidae, Pentatominae) from Miani forest, Sindh, Pakistan. Sindh University Research Journal, Science Series, 43 (2), 157 - 160." type="journal article" year="2011">Shaikh, Memon & Shah, 2011</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -141,7 +141,7 @@
|
|||
<taxonomicNameLabel id="ADD357BD2239E37EFDF4210DB0FBFCAD" box="[620,717,802,825]" pageId="16" pageNumber="367">
|
||||
<emphasis id="B6E0EAC62239E37EFDF4210DB0FBFCAD" bold="true" box="[620,717,802,825]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
<taxonomicName id="43944D572239E37EFF0F216EB120FCC2" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[151,790,833,855]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="mulberriensis">
|
||||
<taxonomicName id="43944D572239E37EFF0F216EB120FCC2" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[151,790,833,855]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="mulberriensis">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F216EB355FCC2" box="[151,355,833,855]" italics="true" pageId="16" pageNumber="367">Halys mulberriensis</emphasis>
|
||||
<bibRefCitation id="E0054B252239E37EFEF2216DB120FCC2" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[362,790,833,855]" pageId="16" pageNumber="367" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -149,7 +149,7 @@
|
|||
<taxonomicNameLabel id="ADD357BD2239E37EFCB8216EB1B4FCC2" box="[800,898,833,856]" pageId="16" pageNumber="367">
|
||||
<emphasis id="B6E0EAC62239E37EFCB8216EB1B4FCC2" bold="true" box="[800,898,833,856]" pageId="16" pageNumber="367">Syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
<taxonomicName id="43944D572239E37EFF0F214FB17BFCE3" authority="Memon, Parveen, Ahmad & Shaikh, 2016: 977" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityPageNumber="977" authorityYear="2016" box="[151,845,864,886]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="mulberriences">
|
||||
<taxonomicName id="43944D572239E37EFF0F214FB17BFCE3" authority="Memon, Parveen, Ahmad & Shaikh, 2016: 977" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityPageNumber="977" authorityYear="2016" box="[151,845,864,886]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="mulberriences">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F214FB35FFCE3" box="[151,361,864,886]" italics="true" pageId="16" pageNumber="367">Halys mulberriences</emphasis>
|
||||
<treatmentCitation id="053510C52239E37EFEF7214EB17BFCE3" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[367,845,864,886]" page="977" pageId="16" pageNumber="367" year="2016">
|
||||
<bibRefCitation id="E0054B252239E37EFEF7214EB128FCE3" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[367,798,864,886]" pageId="16" pageNumber="367" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||||
|
|
@ -157,7 +157,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
–979 (lapsus calami)
|
||||
<taxonomicName id="43944D572239E37EFF0F2151B0CDFC01" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" box="[151,763,894,916]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="noakoatensis">
|
||||
<taxonomicName id="43944D572239E37EFF0F2151B0CDFC01" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" box="[151,763,894,916]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="noakoatensis">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F2151B36AFC01" box="[151,348,894,916]" italics="true" pageId="16" pageNumber="367">Halys noakoatensis</emphasis>
|
||||
<bibRefCitation id="E0054B252239E37EFEFB2150B0CDFC01" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[355,763,894,916]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -169,7 +169,7 @@
|
|||
</paragraph>
|
||||
<paragraph id="842B36D42239E37EFF0F218CB625FC40" blockId="16.[151,1055,588,981]" pageId="16" pageNumber="367">
|
||||
<treatmentCitationGroup id="A48411FA2239E37EFF0F218CB625FC40" pageId="16" pageNumber="367">
|
||||
<taxonomicName id="43944D572239E37EFF0F218CB11AFC2C" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,812,931,953]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="naokotensis">
|
||||
<taxonomicName id="43944D572239E37EFF0F218CB11AFC2C" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,812,931,953]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="naokotensis">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F218CB379FC2C" box="[151,335,931,953]" italics="true" pageId="16" pageNumber="367">Halys naokotensis</emphasis>
|
||||
<treatmentCitation id="053510C52239E37EFECE218BB11AFC2C" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[342,812,931,953]" page="1304" pageId="16" pageNumber="367" year="2017">
|
||||
<bibRefCitation id="E0054B252239E37EFECE218BB0C6FC2C" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[342,752,931,953]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||||
|
|
@ -177,7 +177,7 @@
|
|||
</treatmentCitation>
|
||||
</taxonomicName>
|
||||
(lapsus calami)
|
||||
<taxonomicName id="43944D572239E37EFF0F2190B145FC40" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,883,959,981]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="367" phylum="Chordata" rank="species" species="naokotiensis">
|
||||
<taxonomicName id="43944D572239E37EFF0F2190B145FC40" authority="Memon, Parveen, Ahmad & Shah, 2017: 1304" authorityName="Memon, Parveen, Ahmad & Shah" authorityPageNumber="1304" authorityYear="2017" box="[151,883,959,981]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="naokotiensis">
|
||||
<emphasis id="B6E0EAC62239E37EFF0F2190B360FC40" box="[151,342,959,981]" italics="true" pageId="16" pageNumber="367">Halys naokotiensis</emphasis>
|
||||
<treatmentCitation id="053510C52239E37EFEC521EFB103FC40" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[349,821,959,981]" page="1304" pageId="16" pageNumber="367" year="2017">
|
||||
<bibRefCitation id="E0054B252239E37EFEC521EFB0C1FC40" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[349,759,959,981]" pageId="16" pageNumber="367" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||||
|
|
@ -231,16 +231,6 @@ Articulatory apparatus with basal plates and support bridge complex, dorsal conn
|
|||
<figureCitation id="1CAF2A512238E37FFEE122B8B3FAFF25" box="[377,460,151,176]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="17" pageNumber="368">Fig. 86</figureCitation>
|
||||
); aedeagus moderately long and broadly curved upwards.
|
||||
</paragraph>
|
||||
<caption id="D0EB665C2238E37FFF0F26AAB103FB4C" httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="17" pageNumber="368" startId="17.[151,264,1157,1180]" targetBox="[168,1425,263,1128]" targetPageId="17">
|
||||
<paragraph id="842B36D42238E37FFF0F26AAB103FB4C" blockId="17.[151,1437,1157,1241]" pageId="17" pageNumber="368">
|
||||
<emphasis id="B6E0EAC62238E37FFF0F26AAB362FB0E" bold="true" box="[151,340,1157,1180]" pageId="17" pageNumber="368">FIGURES 77–83.</emphasis>
|
||||
<taxonomicName id="43944D572238E37FFEC226A8B04FFB09" authority="(Thunberg, 1783)" authorityName="Thunberg" authorityYear="1783" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[346,633,1158,1180]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="17" pageNumber="368" phylum="Chordata" rank="species" species="sulcata">
|
||||
<emphasis id="B6E0EAC62238E37FFEC226A8B3F5FB09" box="[346,451,1158,1180]" italics="true" pageId="17" pageNumber="368">H. sulcata</emphasis>
|
||||
(Thunberg, 1783)
|
||||
</taxonomicName>
|
||||
—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="842B36D42238E37FFF5F272FB1C2FA58" blockId="17.[151,1437,1280,2025]" pageId="17" pageNumber="368">
|
||||
<emphasis id="B6E0EAC62238E37FFF5F272FB0A3FA8C" bold="true" box="[199,661,1280,1305]" pageId="17" pageNumber="368">
|
||||
External female genitalia (
|
||||
|
|
@ -340,16 +330,6 @@ from Coimbatore (
|
|||
</taxonomicName>
|
||||
), but the determination is questionable and needs verification.
|
||||
</paragraph>
|
||||
<caption id="D0EB665C223BE37CFF0F27AFB2FCFA41" httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="18" pageNumber="369" startId="18.[151,264,1408,1431]" targetBox="[175,1422,499,1319]" targetPageId="18">
|
||||
<paragraph id="842B36D4223BE37CFF0F27AFB2FCFA41" blockId="18.[151,1436,1408,1492]" pageId="18" pageNumber="369">
|
||||
<emphasis id="B6E0EAC6223BE37CFF0F27AFB360FA03" bold="true" box="[151,342,1408,1431]" pageId="18" pageNumber="369">FIGURES 84–90.</emphasis>
|
||||
<taxonomicName id="43944D57223BE37CFEC727ADB0B3FA02" authority="(Thunberg, 1783)" authorityName="Thunberg" authorityYear="1783" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[351,645,1409,1431]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="18" pageNumber="369" phylum="Chordata" rank="species" species="sulcata">
|
||||
<emphasis id="B6E0EAC6223BE37CFEC727ADB3FCFA02" box="[351,458,1409,1431]" italics="true" pageId="18" pageNumber="369">H. sulcata</emphasis>
|
||||
(Thunberg, 1783)
|
||||
</taxonomicName>
|
||||
—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="842B36D4223BE37DFF57242EB151FEF0" blockId="18.[151,1437,1537,2030]" lastBlockId="19.[151,1436,151,357]" lastPageId="19" lastPageNumber="370" pageId="18" pageNumber="369">
|
||||
There are quite a few species (see the details below) described from
|
||||
<collectingCountry id="FC837644223BE37CFC42242EB60FF98F" box="[986,1081,1537,1562]" name="Pakistan" pageId="18" pageNumber="369">Pakistan</collectingCountry>
|
||||
|
|
@ -424,7 +404,7 @@ Memon
|
|||
)
|
||||
</taxonomicName>
|
||||
=
|
||||
<taxonomicName id="43944D57223AE37DFEE6232BB1BAFE88" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[382,908,260,285]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="sindilla">
|
||||
<taxonomicName id="43944D57223AE37DFEE6232BB1BAFE88" authority="Memon, Meier & Manan, 2006" authorityName="Memon, Meier & Manan" authorityYear="2006" box="[382,908,260,285]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="370" phylum="Arthropoda" rank="species" species="sindilla">
|
||||
<emphasis id="B6E0EAC6223AE37DFEE6232BB02DFE88" box="[382,539,260,285]" italics="true" pageId="19" pageNumber="370">Halys sindilla</emphasis>
|
||||
<bibRefCitation id="E0054B25223AE37DFDBC232AB1BAFE88" author="Memon, N. & Meier, R. & Manan, A. & Su, K. F. - Y." box="[548,908,260,285]" pageId="19" pageNumber="370" pagination="703 - 710" refId="ref13758" refString="Memon, N., Meier, R., Manan, A. & Su, K. F. - Y. (2006) On the use of DNA sequences for determining the species limits of a polymorphic new species in the stink bug genus Halys (Heteroptera: Pentatomidae) from Pakistan. Systematic Entomology, 31 (4), 703 - 710. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00350. x" type="journal article" year="2006">Memon, Meier & Manan, 2006</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -433,7 +413,7 @@ Memon
|
|||
<emphasis id="B6E0EAC6223AE37DFC00232BB637FE88" bold="true" box="[920,1025,260,285]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
=
|
||||
<taxonomicName id="43944D57223AE37DFBBA232BB32AFED5" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="spinosa">
|
||||
<taxonomicName id="43944D57223AE37DFBBA232BB32AFED5" authority="Shaikh, Memon & Shah, 2011" authorityName="Shaikh, Memon & Shah" authorityYear="2011" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="370" phylum="Arthropoda" rank="species" species="spinosa">
|
||||
<emphasis id="B6E0EAC6223AE37DFBBA232BB6F7FE89" box="[1058,1217,260,285]" italics="true" pageId="19" pageNumber="370">Halys spinosa</emphasis>
|
||||
<bibRefCitation id="E0054B25223AE37DFB51232BB32AFED5" author="Shaikh, A. M. & Memon, N. & Shah, M. A." pageId="19" pageNumber="370" pagination="157 - 160" refId="ref14285" refString="Shaikh, A. M., Memon, N. & Shah, M. A. (2011) A new halyine species of Halys Fabricius (Hemiptera, Pentatomidae, Pentatominae) from Miani forest, Sindh, Pakistan. Sindh University Research Journal, Science Series, 43 (2), 157 - 160." type="journal article" year="2011">Shaikh, Memon & Shah, 2011</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -442,7 +422,7 @@ Memon
|
|||
<emphasis id="B6E0EAC6223AE37DFEB62308B3ADFED5" bold="true" box="[302,411,295,320]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
=
|
||||
<taxonomicName id="43944D57223AE37DFE5D2308B684FED5" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[453,1202,295,320]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="mulberriensis">
|
||||
<taxonomicName id="43944D57223AE37DFE5D2308B684FED5" authority="Memon, Parveen, Ahmad & Shaikh, 2016" authorityName="Memon, Parveen, Ahmad & Shaikh" authorityYear="2016" box="[453,1202,295,320]" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="370" phylum="Arthropoda" rank="species" species="mulberriensis">
|
||||
<emphasis id="B6E0EAC6223AE37DFE5D2308B09BFED5" box="[453,685,295,320]" italics="true" pageId="19" pageNumber="370">Halys mulberriensis</emphasis>
|
||||
<bibRefCitation id="E0054B25223AE37DFD222307B684FED5" author="Memon, N. & Parveen, R. & Ahmad, I. & Shaikh, A. M." box="[698,1202,295,320]" pageId="19" pageNumber="370" pagination="977 - 980" refId="ref13843" refString="Memon, N., Parveen, R., Ahmad, I. & Shaikh, A. M. (2016) Description of a new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Lahore, Punjab, Pakistan. Pakistan Journal of Zoology, 48 (4), 977 - 980." type="journal article" year="2016">Memon, Parveen, Ahmad & Shaikh, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
@ -451,7 +431,7 @@ Memon
|
|||
<emphasis id="B6E0EAC6223AE37DFB5B2308B707FED5" bold="true" box="[1219,1329,295,320]" pageId="19" pageNumber="370">syn. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
=
|
||||
<taxonomicName id="43944D57223AE37DFAC32308B0C2FEF1" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="noakoatensis">
|
||||
<taxonomicName id="43944D57223AE37DFAC32308B0C2FEF1" authority="Memon, Parveen, Ahmad & Shah, 2017" authorityName="Memon, Parveen, Ahmad & Shah" authorityYear="2017" class="Insecta" family="Pentatomidae" genus="Halys" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="370" phylum="Arthropoda" rank="species" species="noakoatensis">
|
||||
<emphasis id="B6E0EAC6223AE37DFAC32308B31FFEF0" italics="true" pageId="19" pageNumber="370">Halys noakoatensis</emphasis>
|
||||
<bibRefCitation id="E0054B25223AE37DFEA82362B0C2FEF1" author="Memon, N. & Parveen, R. & Ahmad, I. & Shah, N. A." box="[304,756,332,357]" pageId="19" pageNumber="370" pagination="1301 - 1306" refId="ref13904" refString="Memon, N., Parveen, R., Ahmad, I. & Shah, N. A. (2017) Description of two new species of Oriental genus Halys (Pentatomoidae, Pentatominae, Halyini) from Sindh, Pakistan with a key of its nine Asian species. Pakistan Journal of Zoology, 49 (4), 1301 - 1306. https: // doi. org / 10.17582 / journal. pjz / 2017.49.4.1301.1306" type="journal article" year="2017">Memon, Parveen, Ahmad & Shah, 2017</bibRefCitation>
|
||||
</taxonomicName>
|
||||
|
|
|
|||
|
|
@ -1,140 +0,0 @@
|
|||
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|
||||
Key to species of females of
|
||||
<taxonomicName id="43944D57223AE37DFE6B2149B00FFC15" authorityName="Fabricius" authorityYear="1803" box="[499,569,870,896]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="genus">
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<emphasis id="B6E0EAC6223AE37DFE6B2149B00FFC15" bold="true" box="[499,569,870,896]" italics="true" pageId="19" pageNumber="370">Halys</emphasis>
|
||||
</taxonomicName>
|
||||
from
|
||||
<collectingCountry id="FC837644223AE37DFD1D2149B0FEFC15" box="[645,712,870,896]" name="India" pageId="19" pageNumber="370">India</collectingCountry>
|
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</emphasis>
|
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|
||||
</paragraph>
|
||||
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|
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|
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|
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<paragraph id="842B36D4223AE37DFF0F2182B7ADFC4A" blockId="19.[151,1436,941,1197]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F2182B7ADFC4A" pageId="19" pageNumber="370">
|
||||
1. Valvifers IX with posterior margin incised medially giving a medially divided appearance (
|
||||
<figureCitation id="1CAF2A51223AE37DFBFA2182B698FC57" box="[1122,1198,941,962]" captionStart="FIGURES 49–56" captionStartId="12.[151,264,1708,1731]" captionTargetBox="[172,1426,811,1630]" captionTargetId="figure@12.[151,1435,780,1688]" captionTargetPageId="12" captionText="FIGURES 49–56. H. serrigera Westwood, 1837—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644570/files/figure.png" pageId="19" pageNumber="370">Figs 56</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFB222182B6E3FC57" box="[1210,1237,941,962]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="19" pageNumber="370">88</figureCitation>
|
||||
); valvulae VIII with apical (= caudal) margin uniformly rounded (
|
||||
<figureCitation id="1CAF2A51223AE37DFD0021E5B131FC4A" box="[664,775,970,991]" captionStart="FIGURES 49–56" captionStartId="12.[151,264,1708,1731]" captionTargetBox="[172,1426,811,1630]" captionTargetId="figure@12.[151,1435,780,1688]" captionTargetPageId="12" captionText="FIGURES 49–56. H. serrigera Westwood, 1837—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644570/files/figure.png" pageId="19" pageNumber="370">Figs 54, 55</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFC8921E5B11CFC4A" box="[785,810,970,991]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="19" pageNumber="370">90</figureCitation>
|
||||
).................................................. 2
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F21C8B7AAFB8F" blockId="19.[151,1436,941,1197]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F21C8B7AAFB8F" pageId="19" pageNumber="370">
|
||||
1'. Valvifers IX with posterior margin not incised medially, either uniformly rounded or concave (
|
||||
<figureCitation id="1CAF2A51223AE37DFBE021C8B6F5FC69" box="[1144,1219,999,1020]" captionStart="FIGURES 36–41" captionStartId="9.[151,264,1900,1923]" captionTargetBox="[209,1419,730,1794]" captionTargetId="figure@9.[151,1435,701,1851]" captionTargetPageId="9" captionText="FIGURES 36–41. H. mudigerensis sp. nov.—male and female genitalia. 36, phallus (dorsal); 37, phallus (ventral); 38, phallus (lateral); 39, spermatheca; 40, female terminalia; 41,valvulae VIII. Lettering: dcp—dorsal conjunctival processes; lt VIIIlaterotergite VIII; lt IX—laterotergite IX; pa—processes of aedeagus; sp—spermathecal pump; vcp—ventral conjunctival process; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644566/files/figure.png" pageId="19" pageNumber="370">Figs 40</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFB5621C8B73DFC69" box="[1230,1291,999,1020]" captionStart="FIGURES 69–76" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="19" pageNumber="370">74, 75</figureCitation>
|
||||
);valvulae VIII with apical (= caudal) margin not uniformly rounded rather slightly concave medially (
|
||||
<figureCitation id="1CAF2A51223AE37DFBBA262AB65AFB8F" box="[1058,1132,1029,1050]" captionStart="FIGURES 36–41" captionStartId="9.[151,264,1900,1923]" captionTargetBox="[209,1419,730,1794]" captionTargetId="figure@9.[151,1435,701,1851]" captionTargetPageId="9" captionText="FIGURES 36–41. H. mudigerensis sp. nov.—male and female genitalia. 36, phallus (dorsal); 37, phallus (ventral); 38, phallus (lateral); 39, spermatheca; 40, female terminalia; 41,valvulae VIII. Lettering: dcp—dorsal conjunctival processes; lt VIIIlaterotergite VIII; lt IX—laterotergite IX; pa—processes of aedeagus; sp—spermathecal pump; vcp—ventral conjunctival process; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644566/files/figure.png" pageId="19" pageNumber="370">Figs 41</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFBEE262AB6B9FB8F" box="[1142,1167,1029,1050]" captionStart="FIGURES 69–76" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="19" pageNumber="370">76</figureCitation>
|
||||
).................... 3
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="3F602E71223AE37DFF0F260DB7ADFBE7" pageId="19" pageNumber="370">
|
||||
<paragraph id="842B36D4223AE37DFF0F260DB7ACFBC1" blockId="19.[151,1436,941,1197]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F260DB7ACFBC1" pageId="19" pageNumber="370">
|
||||
2. Posterior margin of valvifers VIII undulate and with distinct concavity adjacent to inner posterior angles and mesal margin straight (
|
||||
<figureCitation id="1CAF2A51223AE37DFEAD2610B395FBC1" box="[309,419,1087,1108]" captionStart="FIGURES 49–56" captionStartId="12.[151,264,1708,1731]" captionTargetBox="[172,1426,811,1630]" captionTargetId="figure@12.[151,1435,780,1688]" captionTargetPageId="12" captionText="FIGURES 49–56. H. serrigera Westwood, 1837—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX." httpUri="https://zenodo.org/record/2644570/files/figure.png" pageId="19" pageNumber="370">Figs 55, 56</figureCitation>
|
||||
).................................................................
|
||||
<taxonomicName id="43944D57223AE37DFB23266FB7ACFBC1" authority="Westwood" authorityName="Westwood" authorityYear="1837" box="[1211,1434,1087,1109]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC6223AE37DFB23266FB707FBC0" box="[1211,1329,1088,1109]" italics="true" pageId="19" pageNumber="370">H. serrigera</emphasis>
|
||||
Westwood
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F2672B7ADFBE7" blockId="19.[151,1436,941,1197]" box="[151,1435,1116,1138]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F2672B7ADFBE7" box="[151,1435,1116,1138]" pageId="19" pageNumber="370">
|
||||
2' Posterior margin of valvifers VIII convex to straight and mesal margin slightly concave (
|
||||
<figureCitation id="1CAF2A51223AE37DFBAC2672B694FBE7" box="[1076,1186,1117,1138]" captionStart="FIGURES 84–90" captionStartId="18.[151,264,1408,1431]" captionTargetBox="[175,1422,499,1319]" captionTargetId="figure@18.[151,1435,460,1367]" captionTargetPageId="18" captionText="FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII." httpUri="https://zenodo.org/record/2644580/files/figure.png" pageId="19" pageNumber="370">Figs 88, 89</figureCitation>
|
||||
)..
|
||||
<taxonomicName id="43944D57223AE37DFB5E2672B7ADFBE7" authority="(Thunberg)" authorityName="Thunberg" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[1222,1435,1116,1138]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="sulcata">
|
||||
<emphasis id="B6E0EAC6223AE37DFB5E2672B71FFBE7" box="[1222,1321,1116,1138]" italics="true" pageId="19" pageNumber="370">H. sulcata</emphasis>
|
||||
(Thunberg)
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="3F602E71223AE37DFF0F2655B7AAFB39" pageId="19" pageNumber="370">
|
||||
<paragraph id="842B36D4223AE37DFF0F2655B7AAFB05" blockId="19.[151,1436,941,1197]" box="[151,1436,1145,1168]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F2655B7AAFB05" box="[151,1436,1145,1168]" pageId="19" pageNumber="370">
|
||||
3. Pronotum broadly angulate, disc slightly elevated, scutellum black (
|
||||
<figureCitation id="1CAF2A51223AE37DFCF42655B1F7FB1A" box="[876,961,1146,1167]" captionStart="FIGURES 1–9" captionStartId="4.[151,264,1939,1962]" captionTargetBox="[246,1377,251,1915]" captionTargetId="figure@4.[183,1409,193,1927]" captionTargetPageId="4" captionText="FIGURES 1–9. Halys Fabricius, 1803 (dorsal and ventral habitus). 1–2, Halys mudigerensis sp. nov. (male); 3, H. mudigerensis sp. nov. (female without abdomen); 4–5, H. serrigera Westwood, 1837; 6–7, H. shaista Ghauri, 1988; 8–9, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644556/files/figure.png" pageId="19" pageNumber="370">Figs 1, 3</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFC542655B63AFB1A" box="[972,1036,1146,1167]" captionStart="FIGURES 10–17" captionStartId="5.[151,264,1871,1894]" captionTargetBox="[180,1427,296,1750]" captionTargetId="figure@5.[151,1435,253,1808]" captionTargetPageId="5" captionText="FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates." httpUri="https://zenodo.org/record/2644558/files/figure.png" pageId="19" pageNumber="370">15, 17</figureCitation>
|
||||
).............
|
||||
<taxonomicName id="43944D57223AE37DFB352655B77FFB1A" authority="Salini, 2019" authorityName="Salini" authorityYear="2019" box="[1197,1353,1145,1167]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="mudigerensis" status="sp. nov.">
|
||||
<emphasis id="B6E0EAC6223AE37DFB352655B77FFB1A" box="[1197,1353,1145,1167]" italics="true" pageId="19" pageNumber="370">H. mudigerensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ADD357BD223AE37DFAD72655B7AAFB05" box="[1359,1436,1146,1168]" pageId="19" pageNumber="370" rank="species">
|
||||
<emphasis id="B6E0EAC6223AE37DFAD72655B7AAFB05" bold="true" box="[1359,1436,1146,1168]" pageId="19" pageNumber="370">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F26B8B7AAFB39" blockId="19.[151,1436,941,1197]" box="[151,1436,1175,1197]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F26B8B7AAFB39" box="[151,1436,1175,1197]" pageId="19" pageNumber="370">
|
||||
3'. Pronotum narrowly angulate, disc more or less flat, scutellum concolourous withdorsal body (
|
||||
<figureCitation id="1CAF2A51223AE37DFBFD26B8B696FB39" box="[1125,1184,1175,1196]" captionStart="FIGURES 1–9" captionStartId="4.[151,264,1939,1962]" captionTargetBox="[246,1377,251,1915]" captionTargetId="figure@4.[183,1409,193,1927]" captionTargetPageId="4" captionText="FIGURES 1–9. Halys Fabricius, 1803 (dorsal and ventral habitus). 1–2, Halys mudigerensis sp. nov. (male); 3, H. mudigerensis sp. nov. (female without abdomen); 4–5, H. serrigera Westwood, 1837; 6–7, H. shaista Ghauri, 1988; 8–9, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644556/files/figure.png" pageId="19" pageNumber="370">Fig. 6</figureCitation>
|
||||
)......
|
||||
<taxonomicName id="43944D57223AE37DFB6A26B7B7AAFB39" authority="Ghauri" authorityName="Ghauri" authorityYear="1988" box="[1266,1436,1175,1197]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC6223AE37DFB6A26B7B765FB38" box="[1266,1363,1175,1197]" italics="true" pageId="19" pageNumber="370">H. shaista</emphasis>
|
||||
Ghauri
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,144 +0,0 @@
|
|||
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Key to species of males of
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<taxonomicName id="43944D57223AE37DFE442398B014FE44" authorityName="Fabricius" authorityYear="1803" box="[476,546,439,465]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="genus">
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<emphasis id="B6E0EAC6223AE37DFE442398B014FE44" bold="true" box="[476,546,439,465]" italics="true" pageId="19" pageNumber="370">Halys</emphasis>
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</taxonomicName>
|
||||
from
|
||||
<collectingCountry id="FC837644223AE37DFDF62397B087FE47" box="[622,689,440,466]" name="India" pageId="19" pageNumber="370">India</collectingCountry>
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</emphasis>
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</heading>
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<subSubSection id="CC8E655F223AE37DFF0F23D1B7ACFC8E" pageId="19" pageNumber="370" type="key">
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<keyLead id="3F6595E1223AE37DFF0F23D1B7ADFDA5" pageId="19" pageNumber="370">
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1. Infoldings of ventral rim of genital capsule medially with median process (
|
||||
<figureCitation id="1CAF2A51223AE37DFC2523D1B63FFD86" box="[957,1033,510,531]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1839,1862]" captionTargetBox="[181,1418,993,1795]" captionTargetId="figure@7.[151,1436,973,1818]" captionTargetPageId="7" captionText="FIGURES 21–28. Cimex dentatus Fabricius, 1775 (Lectotype)—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown." httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="19" pageNumber="370">Figs 22</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFB8C23D1B618FD86" box="[1044,1070,510,531]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="19" pageNumber="370">30</figureCitation>
|
||||
, 43,78); parameral crown with apical margin convex or rounded, not angulate (
|
||||
<figureCitation id="1CAF2A51223AE37DFDF42034B081FDA5" box="[620,695,539,560]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1839,1862]" captionTargetBox="[181,1418,993,1795]" captionTargetId="figure@7.[151,1436,973,1818]" captionTargetPageId="7" captionText="FIGURES 21–28. Cimex dentatus Fabricius, 1775 (Lectotype)—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown." httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="19" pageNumber="370">Figs 26</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFD592034B0EDFDA5" box="[705,731,539,560]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="19" pageNumber="370">32</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFD7D2034B0C9FDA5" box="[741,767,539,560]" captionStart="FIGURES 42–48" captionStartId="11.[151,264,1265,1288]" captionTargetBox="[185,1415,375,1192]" captionTargetId="figure@11.[151,1435,337,1245]" captionTargetPageId="11" captionText="FIGURES 42–48. H. serrigera Westwood, 1837—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="19" pageNumber="370">45</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFC912034B114FDA5" box="[777,802,539,560]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="19" pageNumber="370">80</figureCitation>
|
||||
)................................................... 2
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F2016B7AAFDFE" blockId="19.[151,1436,510,795]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F2016B7AAFDFE" pageId="19" pageNumber="370">
|
||||
1'. Infoldings of ventral rim of genital capsule lacking a median process (
|
||||
<figureCitation id="1CAF2A51223AE37DFC122016B1E5FDDB" box="[906,979,569,590]" captionStart="FIGURES 61–68" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 61–68. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="19" pageNumber="370">Fig. 62</figureCitation>
|
||||
); parameral crown with apical margin angulate (
|
||||
<figureCitation id="1CAF2A51223AE37DFF702079B318FDFE" box="[232,302,598,619]" captionStart="FIGURES 61–68" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 61–68. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="19" pageNumber="370">Fig. 64</figureCitation>
|
||||
)................................................................................
|
||||
<taxonomicName id="43944D57223AE37DFB752079B7AAFDFE" authority="Ghauri" authorityName="Ghauri" authorityYear="1988" box="[1261,1436,597,619]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="shaista">
|
||||
<emphasis id="B6E0EAC6223AE37DFB752079B765FDFE" box="[1261,1363,597,619]" italics="true" pageId="19" pageNumber="370">H. shaista</emphasis>
|
||||
Ghauri
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="3F602E71223AE37DFF0F205BB7ADFD33" pageId="19" pageNumber="370">
|
||||
<paragraph id="842B36D4223AE37DFF0F205BB7ADFD1C" blockId="19.[151,1436,510,795]" box="[151,1435,628,649]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F205BB7ADFD1C" box="[151,1435,628,649]" pageId="19" pageNumber="370">
|
||||
2. Crown of paramere with single tooth (
|
||||
<figureCitation id="1CAF2A51223AE37DFDD7205BB0ACFD1C" box="[591,666,628,649]" captionStart="FIGURES 21–28" captionStartId="7.[151,264,1839,1862]" captionTargetBox="[181,1418,993,1795]" captionTargetId="figure@7.[151,1436,973,1818]" captionTargetPageId="7" captionText="FIGURES 21–28. Cimex dentatus Fabricius, 1775 (Lectotype)—male genitalia. 21, genital capsule (dorsal); 22, genital capsule (ventral); 23–26, paramere (different views); 27, phallus (dorsal); 28, phallus (ventral). Courtesy of H. Enghoff and A. Illum, ©Natural History Museum of Denmark. Lettering: mp—median process; t—tooth on parameral crown." httpUri="https://zenodo.org/record/2644562/files/figure.png" pageId="19" pageNumber="370">Figs 23</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFD3B205BB08BFD1C" box="[675,701,628,649]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="19" pageNumber="370">32</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFD5F205BB0D7FD1C" box="[711,737,628,649]" captionStart="FIGURES 42–48" captionStartId="11.[151,264,1265,1288]" captionTargetBox="[185,1415,375,1192]" captionTargetId="figure@11.[151,1435,337,1245]" captionTargetPageId="11" captionText="FIGURES 42–48. H. serrigera Westwood, 1837—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="19" pageNumber="370">46</figureCitation>
|
||||
,
|
||||
<figureCitation id="1CAF2A51223AE37DFD73205BB132FD1C" box="[747,772,628,649]" captionStart="FIGURES 61–68" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 61–68. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="19" pageNumber="370">65</figureCitation>
|
||||
)..................................................... 3
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F20BEB7ADFD33" blockId="19.[151,1436,510,795]" box="[151,1435,656,678]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F20BEB7ADFD33" box="[151,1435,656,678]" pageId="19" pageNumber="370">
|
||||
2'. Crown of paramere with two teeth (
|
||||
<figureCitation id="1CAF2A51223AE37DFDAF20BEB04BFD33" box="[567,637,657,678]" captionStart="FIGURES 77–83" captionStartId="17.[151,264,1157,1180]" captionTargetBox="[168,1425,263,1128]" captionTargetId="figure@17.[151,1435,229,1137]" captionTargetPageId="17" captionText="FIGURES 77–83. H. sulcata (Thunberg, 1783)—male genitalia. 77, genital capsule (dorsal); 78, genital capsule (ventral); 79, genital capsule (caudal); 80, paramere (dorsal); 81, paramere (ventrolateral); 82, paramere (ventral); 83, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644578/files/figure.png" pageId="19" pageNumber="370">Fig. 82</figureCitation>
|
||||
)................................................
|
||||
<taxonomicName id="43944D57223AE37DFB5E20BEB7ADFD33" authority="(Thunberg)" authorityName="Thunberg" baseAuthorityName="Thunberg" baseAuthorityYear="1783" box="[1222,1435,656,678]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="sulcata">
|
||||
<emphasis id="B6E0EAC6223AE37DFB5E20BEB71FFD33" box="[1222,1321,656,678]" italics="true" pageId="19" pageNumber="370">H. sulcata</emphasis>
|
||||
(Thunberg)
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="3F602E71223AE37DFF0F2081B7ACFC8E" pageId="19" pageNumber="370">
|
||||
<paragraph id="842B36D4223AE37DFF0F2081B7AAFD74" blockId="19.[151,1436,510,795]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F2081B7AAFD74" pageId="19" pageNumber="370">
|
||||
3. Crown of paramere with tooth located medially on the dorsolateral margin not continued as ridge, dorsolateral margin of parameral crown nearly straight (
|
||||
<figureCitation id="1CAF2A51223AE37DFD8720E3B0BAFD74" box="[543,652,716,737]" captionStart="FIGURES 29–35" captionStartId="8.[151,264,1691,1714]" captionTargetBox="[188,1424,813,1586]" captionTargetId="figure@8.[151,1435,750,1658]" captionTargetPageId="8" captionText="FIGURES 29–35. H. mudigerensis sp. nov.—male genitalia. 29, genital capsule (dorsal); 30, genital capsule (ventral); 31, genital capsule (caudal); 32, paramere (ventral); 33, paramere (dorsal); 34, paramere (lateral); 35, articulatory apparatus. Lettering: bcl—bifid caudal lobes; cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644564/files/figure.png" pageId="19" pageNumber="370">Figs 32, 33</figureCitation>
|
||||
)............................................
|
||||
<taxonomicName id="43944D57223AE37DFB3420E3B77EFD74" authority="Salini, 2019" authorityName="Salini" authorityYear="2019" box="[1196,1352,715,737]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="mudigerensis" status="sp. nov.">
|
||||
<emphasis id="B6E0EAC6223AE37DFB3420E3B77EFD74" box="[1196,1352,715,737]" italics="true" pageId="19" pageNumber="370">H. mudigerensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ADD357BD223AE37DFAD720E4B7AAFD74" box="[1359,1436,715,737]" pageId="19" pageNumber="370" rank="species">
|
||||
<emphasis id="B6E0EAC6223AE37DFAD720E4B7AAFD74" bold="true" box="[1359,1436,715,737]" pageId="19" pageNumber="370">sp. nov.</emphasis>
|
||||
</taxonomicNameLabel>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="842B36D4223AE37DFF0F20C6B7ACFC8E" blockId="19.[151,1436,510,795]" pageId="19" pageNumber="370">
|
||||
<keyLead id="3F6595E1223AE37DFF0F20C6B7ACFC8E" pageId="19" pageNumber="370">
|
||||
3'. Crown of paramere with tooth located medially towards apical margin and continuedas ridge, dorsolateral margin of parameral crown excavated concave (
|
||||
<figureCitation id="1CAF2A51223AE37DFE7C2129B01CFC8E" box="[484,554,774,795]" captionStart="FIGURES 42–48" captionStartId="11.[151,264,1265,1288]" captionTargetBox="[185,1415,375,1192]" captionTargetId="figure@11.[151,1435,337,1245]" captionTargetPageId="11" captionText="FIGURES 42–48. H. serrigera Westwood, 1837—male genitalia. 42, genital capsule (dorsal); 43, genital capsule (ventral); 44, genital capsule (caudal); 45, paramere (dorsal); 46, paramere (dorsolateral); 47, paramere (ventral); 48, articulatory apparatus. Lettering: cp—capitate processes; mp—median process; t—tooth." httpUri="https://zenodo.org/record/2644568/files/figure.png" pageId="19" pageNumber="370">Fig. 45</figureCitation>
|
||||
)......................................................
|
||||
<taxonomicName id="43944D57223AE37DFB222129B7ACFC8E" authority="Westwood" authorityName="Westwood" authorityYear="1837" box="[1210,1434,774,795]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="19" pageNumber="370" phylum="Chordata" rank="species" species="serrigera">
|
||||
<emphasis id="B6E0EAC6223AE37DFB222129B706FC8E" box="[1210,1328,774,795]" italics="true" pageId="19" pageNumber="370">H. serrigera</emphasis>
|
||||
Westwood
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue