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<document id="89210B5FBFD12D8ABA0B8682231C9550" ID-DOI="10.1371/journal.pone.0190385" ID-ISSN="1932-6203" ID-PMC="PMC5771577" ID-PubMed="29342184" ID-Zenodo-Dep="12630726" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1719974617054" checkinUser="felipe" docAuthor="Alvarado-Sizzo, Hernán, Casas, Alejandro, Parra, Fabiola, Arreola-Nava, Hilda Julieta, Terrazas, Teresa &amp; Sánchez, Cristian" docDate="2018" docId="0398344DFFBB4772FD829E9A4576FE6B" docLanguage="en" docName="PLoSONE.13.1.e0190385.pdf" docOrigin="PLoS ONE (e 0190385) 13 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0190385" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Stenocereus griseus Buxb., Botanische Studien" docType="treatment" docVersion="1" lastPageNumber="13" masterDocId="FFA14C35FFB3477EFFB798724144FFD3" masterDocTitle="Species delimitation in the Stenocereus griseus (Cactaceae) species complex reveals a new species, S. huastecorum" masterLastPageNumber="25" masterPageNumber="1" pageNumber="9" updateTime="1722020735757" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:title id="910FF6F4400D1CE249AE6BBD4908B8FB">Species delimitation in the Stenocereus griseus (Cactaceae) species complex reveals a new species, S. huastecorum</mods:title>
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<mods:namePart id="5D7443A899EA423DF546C1D3223D756C">Casas, Alejandro</mods:namePart>
<mods:affiliation id="561E9A403C304B49D0B4BD2FED398F80">Instituto de Investigaciones en Ecosistemas y Sustentabilidad (IIES), Universidad Nacional Autónoma de México, Morelia, Michoacán, México,</mods:affiliation>
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<mods:namePart id="5BC278F76E0C46BDD884BD7848D70956">Parra, Fabiola</mods:namePart>
<mods:affiliation id="9E5A6127D77EA5B04737420C77C9B952">Centro De Investigación De Zonas Áridas (CIZA), Universidad Nacional Agraria La Molina, La Molina, Lima, Perú,</mods:affiliation>
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<mods:namePart id="00E22494079A92A3A02C284C88D00442">Arreola-Nava, Hilda Julieta</mods:namePart>
<mods:affiliation id="941ABAB09F62C13E020EC8E556ABA040">Centro Universitario de Ciencias Biológicas y Agropecuarias (CUCBA), Universidad de Guadalajara, Zapopan, Jalisco, México,</mods:affiliation>
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<mods:namePart id="7D68BBFC7A80065E9DD5FF2A1ACAC149">Terrazas, Teresa</mods:namePart>
<mods:affiliation id="CB62888497EA3AA187CEBAF8736E6248">Instituto de Biología (IB), Universidad Nacional Autónoma de México, Coyoacán, Ciudad de México, México,</mods:affiliation>
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<treatmentCitation id="0A90A34AFFBB4776FD829E9A4449F8D2" author="Buxb." box="[565,1293,1768,1793]" journal="Botanische Studien" page="100" pageId="8" pageNumber="9" score="1.1836644748058363" volume="12" year="1961">
<taxonomicName id="4C31FED8FFBB4776FD829E9A453AF8D2" ID-CoL="4ZWLQ" authority="(Haw.) Buxb., Botanische Studien" authorityName="Buxb., Botanische Studien" baseAuthorityName="Haw." box="[565,1150,1768,1793]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFBB4776FD829E9A4240F8D3" bold="true" box="[565,772,1768,1792]" italics="true" pageId="8" pageNumber="9">Stenocereus griseus</emphasis>
(Haw.) Buxb., Botanische Studien
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12:100. 1961
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</paragraph>
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<subSubSection id="C32BD6D0FFBB4772FAA49E9B4576FE6B" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
<paragraph id="8B8E855BFFBB4776FAA49E9B457FF8BA" blockId="8.[533,1524,1768,1897]" pageId="8" pageNumber="9">
[
<bibRefCitation id="EFA0F8AAFFBB4776FAAA9E9B4473F8D2" author="Leinfellner W. &amp; Die Entwicklungslinien der Tribus Pachycereae F." box="[1309,1335,1769,1793]" pageId="8" pageNumber="9" refId="ref13129" refString="61. Leinfellner W. Die Entwicklungslinien der Tribus Pachycereae F. Buxbaum (Cactaceae - Cereoideae). Botanische Studien. 1961." type="book" year="1961">61</bibRefCitation>
]. [urn:lsid:ipni.
<emphasis id="B9455949FFBB4776FDA29F79421DF8F0" bold="true" box="[533,857,1803,1827]" pageId="8" pageNumber="9">org:names:244634–2:1.3.2.1].</emphasis>
Type: América meridional, (unknown).
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(here designated)
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, Departamento de
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, Municipio de Uribia,
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de Manaure.
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<collectingDate id="EFCB5A73FFBB4776FDA29F2343F3F8BA" box="[533,695,1873,1897]" pageId="8" pageNumber="9" value="1963-01-31">31-Enero-1963</collectingDate>
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,
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.
<collectingRegion id="49F54BB9FFBB4776FD699F23426FF8BA" box="[734,811,1873,1897]" country="Germany" name="Saarland" pageId="8" pageNumber="9">Saravia</collectingRegion>
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.
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˚. 2183 (
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)
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.
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<caption id="DF4ED5D3FFBA4777FEAB9F5443F0F8B1" ID-DOI="http://doi.org/10.5281/zenodo.12630736" ID-Zenodo-Dep="12630736" httpUri="https://zenodo.org/record/12630736/files/figure.png" pageId="9" pageNumber="10" startId="9.[284,312,1830,1849]" targetBox="[284,1536,208,1808]" targetPageId="9" targetType="figure">
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Fig 5. Niche suitability pairwise comparisons based on
<emphasis id="B9455949FFBA4777FCB69F54424CF8EA" bold="true" box="[769,776,1830,1849]" italics="true" pageId="9" pageNumber="10">t</emphasis>
-tests.
</emphasis>
Different letters represent levels of significant differences. https://doi.org/10.1371/journal.pone.0190385.g005
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<paragraph id="8B8E855BFFB94774FFD798A242BAFF30" blockId="10.[96,1022,208,227]" box="[96,1022,208,227]" pageId="10" pageNumber="11">Table 2. Pairwise comparisons of Ecological niche principal components and two-ways background tests.</paragraph>
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<table id="F93177FBFFB9B881FFD7988544BCFDBB" box="[96,1528,247,616]" gridcols="11" gridrows="11" pageId="10" pageNumber="11">
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<emphasis id="B9455949FFB94774FF2198854015FED9" bold="true" box="[150,337,247,266]" pageId="10" pageNumber="11">Pairwise comparison</emphasis>
</th>
<th id="76D0EE65FFB9B881FC3298854575FED9" box="[901,1073,247,266]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC2F988542B3FED9" bold="true" box="[920,1015,247,266]" pageId="10" pageNumber="11">Niche axes</emphasis>
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<emphasis id="B9455949FFB94774FE59996B4350FEFF" bold="true" box="[494,532,281,300]" pageId="10" pageNumber="11">PC1</emphasis>
</td>
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<emphasis id="B9455949FFB94774FD66996B43B3FEFF" bold="true" box="[721,759,281,300]" pageId="10" pageNumber="11">PC2</emphasis>
</td>
<td id="76D0EE65FFB9B881FC32996B42A2FEFF" box="[901,998,281,300]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC02996B429FFEFF" bold="true" box="[949,987,281,300]" pageId="10" pageNumber="11">PC3</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF996B458DFEFF" box="[1128,1225,281,300]" gridcol="7" gridrow="1" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB2F996B45FAFEFF" bold="true" box="[1176,1214,281,300]" pageId="10" pageNumber="11">PC4</emphasis>
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<td id="76D0EE65FFB9B881FAFB996B44E9FEFF" box="[1356,1453,281,300]" gridcol="9" gridrow="1" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FACB996B44E6FEFF" bold="true" box="[1404,1442,281,300]" pageId="10" pageNumber="11">PC5</emphasis>
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<taxonomicName id="4C31FED8FFB94774FFD7994841A4FE9E" box="[96,224,314,333]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="huastecorum">
<emphasis id="B9455949FFB94774FFD79948412EFE9E" box="[96,106,314,333]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FFC3994841A4FE9E" box="[116,224,314,333]" italics="true" pageId="10" pageNumber="11">huastecorum</emphasis>
</taxonomicName>
vs
<taxonomicName id="4C31FED8FFB94774FF4C99484024FE9F" box="[251,352,314,333]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="pruinosus">
<emphasis id="B9455949FFB94774FF4C99484041FE9E" box="[251,261,314,333]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FEB999484024FE9F" box="[270,352,314,332]" italics="true" pageId="10" pageNumber="11">pruinosus</emphasis>
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<emphasis id="B9455949FFB94774FE549948435AFE9E" bold="true" box="[483,542,314,333]" pageId="10" pageNumber="11">0.3369</emphasis>
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<emphasis id="B9455949FFB94774FD7099484246FE9E" bold="true" box="[711,770,314,333]" pageId="10" pageNumber="11">1.3788</emphasis>
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<td id="76D0EE65FFB9B881FC32994842A2FE9E" box="[901,998,314,333]" gridcol="5" gridrow="2" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC1C994842A2FE9E" bold="true" box="[939,998,314,333]" pageId="10" pageNumber="11">0.6422</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF9948458DFE9E" box="[1128,1225,314,333]" gridcol="7" gridrow="2" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB399948458DFE9E" bold="true" box="[1166,1225,314,333]" pageId="10" pageNumber="11">0.5282</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB994844E9FE9E" box="[1356,1453,314,333]" gridcol="9" gridrow="2" pageId="10" pageNumber="11">0.1474</td>
</tr>
<tr id="35018719FFB9B881FFD7992944BCFEBD" box="[96,1528,347,366]" gridrow="3" pageId="10" pageNumber="11">
<th id="76D0EE65FFB9B881FFD799294024FEBD" box="[96,352,347,366]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">Background tests</th>
<td id="76D0EE65FFB9B881FE099929435AFEBD" box="[446,542,347,366]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">0.0419</td>
<td id="76D0EE65FFB9B881FD989929432EFEBD" box="[559,618,347,366]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD989929432EFEBD" bold="true" box="[559,618,347,366]" pageId="10" pageNumber="11">1.6743</emphasis>
</td>
<td id="76D0EE65FFB9B881FD1699294246FEBD" box="[673,770,347,366]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD1699294398FEBD" bold="true" box="[673,732,347,366]" pageId="10" pageNumber="11">0.7520</emphasis>
</td>
<td id="76D0EE65FFB9B881FCA49929420AFEBD" box="[787,846,347,366]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FCA49929420AFEBD" bold="true" box="[787,846,347,366]" pageId="10" pageNumber="11">2.0177</emphasis>
</td>
<td id="76D0EE65FFB9B881FC32992942A2FEBD" box="[901,998,347,366]" gridcol="5" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC3299294284FEBD" bold="true" box="[901,960,347,366]" pageId="10" pageNumber="11">1.0944</emphasis>
</td>
<td id="76D0EE65FFB9B881FC4199294575FEBD" box="[1014,1073,347,366]" gridcol="6" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC4199294575FEBD" bold="true" box="[1014,1073,347,366]" pageId="10" pageNumber="11">0.3950</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF9929458DFEBD" box="[1128,1225,347,366]" gridcol="7" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FBDF992945E7FEBD" bold="true" box="[1128,1187,347,366]" pageId="10" pageNumber="11">1.2178</emphasis>
</td>
<td id="76D0EE65FFB9B881FB6D99294451FEBD" box="[1242,1301,347,366]" gridcol="8" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB6D99294451FEBD" bold="true" box="[1242,1301,347,366]" pageId="10" pageNumber="11">0.8736</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB992944E9FEBD" box="[1356,1453,347,366]" gridcol="9" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAFB992944C3FEBD" bold="true" box="[1356,1415,347,366]" pageId="10" pageNumber="11">0.2416</emphasis>
</td>
<td id="76D0EE65FFB9B881FA0A992944BCFEBD" box="[1469,1528,347,366]" gridcol="10" gridrow="3" pageId="10" pageNumber="11">0.0649</td>
</tr>
<tr id="35018719FFB9B881FFD7990E44BCFE43" box="[96,1528,380,400]" gridrow="4" pageId="10" pageNumber="11" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76D0EE65FFB9B881FFD7990E4024FE43" box="[96,352,380,400]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">
<taxonomicName id="4C31FED8FFB94774FFD7990F4182FE5C" box="[96,198,381,400]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="pruinosus">
<emphasis id="B9455949FFB94774FFD7990F412EFE43" box="[96,106,381,400]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FFC3990F4182FE5C" box="[116,198,381,399]" italics="true" pageId="10" pageNumber="11">pruinosus</emphasis>
</taxonomicName>
vs
<taxonomicName id="4C31FED8FFB94774FF56990F4003FE5C" box="[225,327,380,400]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="laevigatus">
<emphasis id="B9455949FFB94774FF56990F41AFFE43" box="[225,235,381,400]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FF43990E4003FE5C" box="[244,327,380,399]" italics="true" pageId="10" pageNumber="11">laevigatus</emphasis>
</taxonomicName>
</th>
<td id="76D0EE65FFB9B881FE09990E435AFE43" box="[446,542,380,400]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FE54990F435AFE43" bold="true" box="[483,542,381,400]" pageId="10" pageNumber="11">2.0910</emphasis>
</td>
<td id="76D0EE65FFB9B881FD16990E4246FE43" box="[673,770,380,400]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD70990F4246FE43" bold="true" box="[711,770,381,400]" pageId="10" pageNumber="11">0.3313</emphasis>
</td>
<td id="76D0EE65FFB9B881FC32990E42A2FE43" box="[901,998,380,400]" gridcol="5" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC1C990F42A2FE43" bold="true" box="[939,998,381,400]" pageId="10" pageNumber="11">0.3860</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF990E458DFE43" box="[1128,1225,380,400]" gridcol="7" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB39990F458DFE43" bold="true" box="[1166,1225,381,400]" pageId="10" pageNumber="11">0.5077</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB990E44E9FE43" box="[1356,1453,380,400]" gridcol="9" gridrow="4" pageId="10" pageNumber="11">0.0160</td>
</tr>
<tr id="35018719FFB9B881FFD799EC44BCFE62" box="[96,1528,414,433]" gridrow="5" pageId="10" pageNumber="11">
<th id="76D0EE65FFB9B881FFD799EC4024FE62" box="[96,352,414,433]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">Background tests</th>
<td id="76D0EE65FFB9B881FE0999EC435AFE62" box="[446,542,414,433]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FE0999EC40BDFE62" bold="true" box="[446,505,414,433]" pageId="10" pageNumber="11">2.7793</emphasis>
</td>
<td id="76D0EE65FFB9B881FD9899EC432EFE62" box="[559,618,414,433]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD9899EC432EFE62" bold="true" box="[559,618,414,433]" pageId="10" pageNumber="11">1.7122</emphasis>
</td>
<td id="76D0EE65FFB9B881FD1699EC4246FE62" box="[673,770,414,433]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD1699EC4398FE62" bold="true" box="[673,732,414,433]" pageId="10" pageNumber="11">1.0676</emphasis>
</td>
<td id="76D0EE65FFB9B881FCA499EC420AFE62" box="[787,846,414,433]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FCA499EC420AFE62" bold="true" box="[787,846,414,433]" pageId="10" pageNumber="11">0.2955</emphasis>
</td>
<td id="76D0EE65FFB9B881FC3299EC42A2FE62" box="[901,998,414,433]" gridcol="5" gridrow="5" pageId="10" pageNumber="11">0.0978</td>
<td id="76D0EE65FFB9B881FC4199EC4575FE62" box="[1014,1073,414,433]" gridcol="6" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC4199EC4575FE62" bold="true" box="[1014,1073,414,433]" pageId="10" pageNumber="11">0.8382</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF99EC458DFE62" box="[1128,1225,414,433]" gridcol="7" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FBDF99EC45E7FE62" bold="true" box="[1128,1187,414,433]" pageId="10" pageNumber="11">0.3178</emphasis>
</td>
<td id="76D0EE65FFB9B881FB6D99EC4451FE62" box="[1242,1301,414,433]" gridcol="8" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB6D99EC4451FE62" bold="true" box="[1242,1301,414,433]" pageId="10" pageNumber="11">1.1973</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB99EC44E9FE62" box="[1356,1453,414,433]" gridcol="9" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAFB99EC44C3FE62" bold="true" box="[1356,1415,414,433]" pageId="10" pageNumber="11">0.5972</emphasis>
</td>
<td id="76D0EE65FFB9B881FA0A99EC44BCFE62" box="[1469,1528,414,433]" gridcol="10" gridrow="5" pageId="10" pageNumber="11">0.1102</td>
</tr>
<tr id="35018719FFB9B881FFD799CD44BCFE01" box="[96,1528,447,466]" gridrow="6" pageId="10" pageNumber="11" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76D0EE65FFB9B881FFD799CD4024FE01" box="[96,352,447,466]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">
<taxonomicName id="4C31FED8FFB94774FFD799CD418CFE01" box="[96,200,447,466]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="laevigatus">
<emphasis id="B9455949FFB94774FFD799CD412EFE01" box="[96,106,447,466]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FFC399CD418CFE01" box="[116,200,447,466]" italics="true" pageId="10" pageNumber="11">laevigatus</emphasis>
</taxonomicName>
vs
<taxonomicName id="4C31FED8FFB94774FF5599CD406AFE01" box="[226,302,447,466]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFB94774FF5599CD41A8FE01" box="[226,236,447,466]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FF4299B2406AFE01" box="[245,302,448,466]" italics="true" pageId="10" pageNumber="11">griseus</emphasis>
</taxonomicName>
</th>
<td id="76D0EE65FFB9B881FE0999CD435AFE01" box="[446,542,447,466]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FE5499CD435AFE01" bold="true" box="[483,542,447,466]" pageId="10" pageNumber="11">0.6726</emphasis>
</td>
<td id="76D0EE65FFB9B881FD1699CD4246FE01" box="[673,770,447,466]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD7099CD4246FE01" bold="true" box="[711,770,447,466]" pageId="10" pageNumber="11">1.1603</emphasis>
</td>
<td id="76D0EE65FFB9B881FC3299CD42A2FE01" box="[901,998,447,466]" gridcol="5" gridrow="6" pageId="10" pageNumber="11">0.1296</td>
<td id="76D0EE65FFB9B881FBDF99CD458DFE01" box="[1128,1225,447,466]" gridcol="7" gridrow="6" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB3999CD458DFE01" bold="true" box="[1166,1225,447,466]" pageId="10" pageNumber="11">0.6940</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB99CD44E9FE01" box="[1356,1453,447,466]" gridcol="9" gridrow="6" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAC599CD44E9FE01" bold="true" box="[1394,1453,447,466]" pageId="10" pageNumber="11">0.3032</emphasis>
</td>
</tr>
<tr id="35018719FFB9B881FFD7999344BCFE27" box="[96,1528,481,500]" gridrow="7" pageId="10" pageNumber="11">
<th id="76D0EE65FFB9B881FFD799934024FE27" box="[96,352,481,500]" gridcol="0" gridrow="7" pageId="10" pageNumber="11">Background tests</th>
<td id="76D0EE65FFB9B881FE099993435AFE27" box="[446,542,481,500]" gridcol="1" gridrow="7" pageId="10" pageNumber="11">0.0157</td>
<td id="76D0EE65FFB9B881FD989993432EFE27" box="[559,618,481,500]" gridcol="2" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD989993432EFE27" bold="true" box="[559,618,481,500]" pageId="10" pageNumber="11">1.7122</emphasis>
</td>
<td id="76D0EE65FFB9B881FD1699934246FE27" box="[673,770,481,500]" gridcol="3" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD1699934398FE27" bold="true" box="[673,732,481,500]" pageId="10" pageNumber="11">0.3068</emphasis>
</td>
<td id="76D0EE65FFB9B881FCA49993420AFE27" box="[787,846,481,500]" gridcol="4" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FCA49993420AFE27" bold="true" box="[787,846,481,500]" pageId="10" pageNumber="11">1.8966</emphasis>
</td>
<td id="76D0EE65FFB9B881FC32999342A2FE27" box="[901,998,481,500]" gridcol="5" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC3299934284FE27" bold="true" box="[901,960,481,500]" pageId="10" pageNumber="11">0.4104</emphasis>
</td>
<td id="76D0EE65FFB9B881FC4199934575FE27" box="[1014,1073,481,500]" gridcol="6" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC4199934575FE27" bold="true" box="[1014,1073,481,500]" pageId="10" pageNumber="11">0.6133</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF9993458DFE27" box="[1128,1225,481,500]" gridcol="7" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FBDF999345E7FE27" bold="true" box="[1128,1187,481,500]" pageId="10" pageNumber="11">1.2786</emphasis>
</td>
<td id="76D0EE65FFB9B881FB6D99934451FE27" box="[1242,1301,481,500]" gridcol="8" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB6D99934451FE27" bold="true" box="[1242,1301,481,500]" pageId="10" pageNumber="11">0.5033</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB999344E9FE27" box="[1356,1453,481,500]" gridcol="9" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAFB999344C3FE27" bold="true" box="[1356,1415,481,500]" pageId="10" pageNumber="11">0.2753</emphasis>
</td>
<td id="76D0EE65FFB9B881FA0A999344BCFE27" box="[1469,1528,481,500]" gridcol="10" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FA0A999344BCFE27" bold="true" box="[1469,1528,481,500]" pageId="10" pageNumber="11">0.8874</emphasis>
</td>
</tr>
<tr id="35018719FFB9B881FFD79A7044BCFDC6" box="[96,1528,514,533]" gridrow="8" pageId="10" pageNumber="11" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76D0EE65FFB9B881FFD79A704024FDC6" box="[96,352,514,533]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">
<taxonomicName id="4C31FED8FFB94774FFD79A7041A4FDC6" box="[96,224,514,533]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="huastecorum">
<emphasis id="B9455949FFB94774FFD79A70412EFDC6" box="[96,106,514,533]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FFC39A7041A4FDC6" box="[116,224,514,533]" italics="true" pageId="10" pageNumber="11">huastecorum</emphasis>
</taxonomicName>
vs
<taxonomicName id="4C31FED8FFB94774FF4C9A704003FDC7" box="[251,327,514,533]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFB94774FF4C9A704041FDC6" box="[251,261,514,533]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FEB99A704003FDC7" box="[270,327,514,532]" italics="true" pageId="10" pageNumber="11">griseus</emphasis>
</taxonomicName>
</th>
<td id="76D0EE65FFB9B881FE099A70435AFDC6" box="[446,542,514,533]" gridcol="1" gridrow="8" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FE549A70435AFDC6" bold="true" box="[483,542,514,533]" pageId="10" pageNumber="11">2.4267</emphasis>
</td>
<td id="76D0EE65FFB9B881FD169A704246FDC6" box="[673,770,514,533]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD709A704246FDC6" bold="true" box="[711,770,514,533]" pageId="10" pageNumber="11">2.2078</emphasis>
</td>
<td id="76D0EE65FFB9B881FC329A7042A2FDC6" box="[901,998,514,533]" gridcol="5" gridrow="8" pageId="10" pageNumber="11">0.1266</td>
<td id="76D0EE65FFB9B881FBDF9A70458DFDC6" box="[1128,1225,514,533]" gridcol="7" gridrow="8" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB399A70458DFDC6" bold="true" box="[1166,1225,514,533]" pageId="10" pageNumber="11">0.7144</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB9A7044E9FDC6" box="[1356,1453,514,533]" gridcol="9" gridrow="8" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAC59A7044E9FDC6" bold="true" box="[1394,1453,514,533]" pageId="10" pageNumber="11">0.4376</emphasis>
</td>
</tr>
<tr id="35018719FFB9B881FFD79A5144BCFDE5" box="[96,1528,547,566]" gridrow="9" pageId="10" pageNumber="11">
<th id="76D0EE65FFB9B881FFD79A514024FDE5" box="[96,352,547,566]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">Background tests</th>
<td id="76D0EE65FFB9B881FE099A51435AFDE5" box="[446,542,547,566]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FE099A5140BDFDE5" bold="true" box="[446,505,547,566]" pageId="10" pageNumber="11">3.3591</emphasis>
</td>
<td id="76D0EE65FFB9B881FD989A51432EFDE5" box="[559,618,547,566]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD989A51432EFDE5" bold="true" box="[559,618,547,566]" pageId="10" pageNumber="11">1.0893</emphasis>
</td>
<td id="76D0EE65FFB9B881FD169A514246FDE5" box="[673,770,547,566]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FD169A514398FDE5" bold="true" box="[673,732,547,566]" pageId="10" pageNumber="11">2.8847</emphasis>
</td>
<td id="76D0EE65FFB9B881FCA49A51420AFDE5" box="[787,846,547,566]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FCA49A51420AFDE5" bold="true" box="[787,846,547,566]" pageId="10" pageNumber="11">0.7407</emphasis>
</td>
<td id="76D0EE65FFB9B881FC329A5142A2FDE5" box="[901,998,547,566]" gridcol="5" gridrow="9" pageId="10" pageNumber="11">0.1542</td>
<td id="76D0EE65FFB9B881FC419A514575FDE5" box="[1014,1073,547,566]" gridcol="6" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FC419A514575FDE5" bold="true" box="[1014,1073,547,566]" pageId="10" pageNumber="11">0.9105</emphasis>
</td>
<td id="76D0EE65FFB9B881FBDF9A51458DFDE5" box="[1128,1225,547,566]" gridcol="7" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FBDF9A5145E7FDE5" bold="true" box="[1128,1187,547,566]" pageId="10" pageNumber="11">1.2990</emphasis>
</td>
<td id="76D0EE65FFB9B881FB6D9A514451FDE5" box="[1242,1301,547,566]" gridcol="8" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FB6D9A514451FDE5" bold="true" box="[1242,1301,547,566]" pageId="10" pageNumber="11">1.0599</emphasis>
</td>
<td id="76D0EE65FFB9B881FAFB9A5144E9FDE5" box="[1356,1453,547,566]" gridcol="9" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FAFB9A5144C3FDE5" bold="true" box="[1356,1415,547,566]" pageId="10" pageNumber="11">0.4067</emphasis>
</td>
<td id="76D0EE65FFB9B881FA0A9A5144BCFDE5" box="[1469,1528,547,566]" gridcol="10" gridrow="9" pageId="10" pageNumber="11">
<emphasis id="B9455949FFB94774FA0A9A5144BCFDE5" bold="true" box="[1469,1528,547,566]" pageId="10" pageNumber="11">0.2252</emphasis>
</td>
</tr>
<tr id="35018719FFB9B881FFD79A2744BCFDBB" box="[96,1528,597,616]" gridrow="10" pageId="10" pageNumber="11">
<th id="76D0EE65FFB9B881FFD79A2744BCFDBB" box="[96,1528,597,616]" colspan="11" colspanRight="10" gridcol="0" gridrow="10" pageId="10" pageNumber="11">Boldfaces represent statistical significance.</th>
</tr>
</table>
</paragraph>
<paragraph id="8B8E855BFFB94774FFD79AF740B7FD4B" blockId="10.[96,499,645,664]" box="[96,499,645,664]" pageId="10" pageNumber="11">
<tableNote id="76D784D5FFB94774FFD79AF740B7FD4B" box="[96,499,645,664]" pageId="10" pageNumber="11" targetBox="[96,1528,247,616]" targetPageId="10">https://doi.org/10.1371/journal.pone.0190385.t002</tableNote>
</paragraph>
<paragraph id="8B8E855BFFB94774FD829AAB45D5FD22" blockId="10.[533,1536,729,1862]" box="[565,1169,729,753]" pageId="10" pageNumber="11">
Additional observed specimens: Table
<collectionCode id="ED201D9EFFB94774FC789AAB42A5FD22" box="[975,993,729,753]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="10" pageNumber="11" type="Herbarium">A</collectionCode>
in
<collectionCode id="ED201D9EFFB94774FBB49AAB4555FD22" box="[1027,1041,729,753]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="10" pageNumber="11" type="Herbarium">S</collectionCode>
3 Appendix.
</paragraph>
<paragraph id="8B8E855BFFB94774FD829A8945DBF987" blockId="10.[533,1536,729,1862]" pageId="10" pageNumber="11">
Candelabraform tree, up to
<quantity id="4CC928BEFFB94774FCEB9A8942C1FCC0" box="[860,901,763,787]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="m" value="9.0">9 m</quantity>
tall; trunk many times absent, &lt;
<quantity id="4CC928BEFFB94774FB689A894464FCC0" box="[1247,1312,763,787]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="cm" value="30.0">30 cm</quantity>
tall and &lt;
<quantity id="4CC928BEFFB94774FA389A89448BFCC0" box="[1423,1487,763,787]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="cm" value="20.0">20 cm</quantity>
width; many branches, ascending, about
<quantity id="4CC928BEFFB94774FC719B6C4542FCE5" box="[966,1030,798,822]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="cm" value="15.0">15 cm</quantity>
width, unconstrained, grayish green to glaucous; first order branching; mucilage cavities not evident in transverse branch sections; ribs 7 to 9, acute in transverse section, straight in longitudinal section,
<quantity id="4CC928BEFFB94774FB099B114474FCA8" box="[1214,1328,867,891]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.5" metricValueMin="1.2" pageId="10" pageNumber="11" unit="mm" value="23.5" valueMax="35.0" valueMin="12.0">12–35 mm</quantity>
tall by
<quantity id="4CC928BEFFB94774FACB9B1144AAFCA8" box="[1404,1518,867,891]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.5" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="23.0" valueMax="35.0" valueMin="11.0">11–35 mm</quantity>
wide at the base, without horizontal constriction between areolas within the same rib; areolas
<quantity id="4CC928BEFFB94774FDA29BDB433FFC12" box="[533,635,937,961]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.8" metricValueMin="0.8" pageId="10" pageNumber="11" unit="mm" value="18.0" valueMax="28.0" valueMin="8.0">8–28 mm</quantity>
apart each, round to obovate (scutelliform),
<quantity id="4CC928BEFFB94774FBE39BDB4588FC12" box="[1108,1228,937,961]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="11.5" metricValueMin="6.0" pageId="10" pageNumber="11" unit="mm" value="8.75" valueMax="11.5" valueMin="6.0">6–11.5 mm</quantity>
long and
<quantity id="4CC928BEFFB94774FA829BDB44E9FC12" box="[1333,1453,937,961]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="10.6" metricValueMin="6.0" pageId="10" pageNumber="11" unit="mm" value="8.3" valueMax="10.6" valueMin="6.0">6–10.6 mm</quantity>
wide, with light-colored trichomes; radial spines 5–7, subulated, divergent,
<quantity id="4CC928BEFFB94774FB459BB94413FC30" box="[1266,1367,971,995]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="3.0" metricValueMin="0.8" pageId="10" pageNumber="11" unit="mm" value="19.0" valueMax="30.0" valueMin="8.0">8–30 mm</quantity>
long, white when young, grayish when mature; 1 central spine, rarely 3, subulated, robust,
<quantity id="4CC928BEFFB94774FAE39B9C4489FBD5" box="[1364,1485,1006,1030]" metricMagnitude="-2" metricUnit="m" metricValue="3.285" metricValueMax="5.6" metricValueMin="0.97" pageId="10" pageNumber="11" unit="mm" value="32.85" valueMax="56.0" valueMin="9.7">9.7–56 mm</quantity>
long, white with reddish base when young, grayish at maturity; subapical flowers, night anthesis remaining opened until the next morning, infundibuliform,
<quantity id="4CC928BEFFB94774FB029C414452FB98" box="[1205,1302,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" metricValueMax="6.0" metricValueMin="5.6" pageId="10" pageNumber="11" unit="cm" value="5.8" valueMax="6.0" valueMin="5.6">5.6–6 cm</quantity>
long and
<quantity id="4CC928BEFFB94774FAC89C4144A4FB98" box="[1407,1504,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="5.0" metricValueMin="4.4" pageId="10" pageNumber="11" unit="cm" value="4.7" valueMax="5.0" valueMin="4.4">4.4–5 cm</quantity>
wide in anthesis; pericarpel cylindrical, green,
<quantity id="4CC928BEFFB94774FC489C2445D3FBBD" box="[1023,1175,1110,1134]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="10" pageNumber="11" unit="mm" value="13.0" valueMax="14.5" valueMin="11.5">11.5–14.5 mm</quantity>
wide, covered with slightly prominent podaria, separated, with wide oblong scales about
<quantity id="4CC928BEFFB94774FBD49C0B45A6FB42" box="[1123,1250,1145,1169]" metricMagnitude="-3" metricUnit="m" metricValue="1.8499999999999999" metricValueMax="2.3" metricValueMin="1.4" pageId="10" pageNumber="11" unit="mm" value="1.8499999999999999" valueMax="2.3" valueMin="1.4">1.4–2.3 mm</quantity>
long by
<quantity id="4CC928BEFFB94774FA8B9C0B44FEFB42" box="[1340,1466,1145,1169]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="3.2" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="2.35" valueMax="3.2" valueMin="1.5">1.5–3.2 mm</quantity>
wide at the base, reddish, areolas presenting light-colored trichomes, spines
<quantity id="4CC928BEFFB94774FAB79CE94422FB60" box="[1280,1382,1179,1203]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="10" pageNumber="11" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">7–13 mm</quantity>
long; receptacular tube
<quantity id="4CC928BEFFB94774FD329CCC43BCFB05" box="[645,760,1214,1238]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="5.4" metricValueMin="3.0" pageId="10" pageNumber="11" unit="mm" value="42.0" valueMax="54.0" valueMin="30.0">30–54 mm</quantity>
long, podaria with decurrent scales, narrow oblong to spatulate, apex obtuse to mucronated, about
<quantity id="4CC928BEFFB94774FCFA9C9342E2FB2A" box="[845,934,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="10" pageNumber="11" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
wide with few trichomes and spines; outer perianth segments narrow oblong, apex rounded to truncate,
<quantity id="4CC928BEFFB94774FBA99D7145D5FAC8" box="[1054,1169,1283,1307]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="10" pageNumber="11" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">16–20 mm</quantity>
long and
<quantity id="4CC928BEFFB94774FB4D9D714436FAC8" box="[1274,1394,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="10.0" metricValueMin="6.5" pageId="10" pageNumber="11" unit="mm" value="8.25" valueMax="10.0" valueMin="6.5">6.5–10 mm</quantity>
wide, green to reddish from the bottom to the top; inner segments oblong to spatulate, entire margin, up to
<quantity id="4CC928BEFFB94774FD869D3B4333FAB2" box="[561,631,1353,1377]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="cm" value="2.5">2.5 cm</quantity>
long and
<quantity id="4CC928BEFFB94774FD579D3B4250FAB2" box="[736,788,1353,1377]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="cm" value="1.0">1 cm</quantity>
wide, white to pinkish-white; stamina included, numerous, arranged in verticillated series; basifix anthers, yellowish, style
<quantity id="4CC928BEFFB94774FBF29D1E45F3FA57" box="[1093,1207,1388,1412]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.2" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="37.5" valueMax="52.0" valueMin="23.0">23–52 mm</quantity>
long and
<quantity id="4CC928BEFFB94774FA979D1E44C2FA57" box="[1312,1414,1388,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.9" metricValueMin="0.9" pageId="10" pageNumber="11" unit="mm" value="14.0" valueMax="19.0" valueMin="9.0">9–19 mm</quantity>
wide, white to pinkish white; stigma lobules 8–10,
<quantity id="4CC928BEFFB94774FC5D9DFC4507FA75" box="[1002,1091,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="10" pageNumber="11" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long, yellowish white; nectar chamber semi-closed by the lower filaments curvature,
<quantity id="4CC928BEFFB94774FC4C9DC3452AFA1A" box="[1019,1134,1457,1481]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="15.0" valueMax="19.0" valueMin="11.0">11–19 mm</quantity>
long and
<quantity id="4CC928BEFFB94774FB619DC34411FA1A" box="[1238,1365,1457,1481]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.7" metricValueMin="5.9" pageId="10" pageNumber="11" unit="mm" value="6.3" valueMax="6.7" valueMin="5.9">5.9–6.7 mm</quantity>
wide, striated; ovary
<quantity id="4CC928BEFFB94774FDE19DA6438BFA3F" box="[598,719,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.33" metricValueMin="0.8" pageId="10" pageNumber="11" unit="mm" value="10.65" valueMax="13.3" valueMin="8.0">8–13.3 mm</quantity>
long and
<quantity id="4CC928BEFFB94774FC8F9DA64287FA3F" box="[824,963,1492,1516]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="10.8" metricValueMin="5.6" pageId="10" pageNumber="11" unit="mm" value="8.2" valueMax="10.8" valueMin="5.6">5.6–10.8 mm</quantity>
wide; fruit ovoid, dehiscent when ripe, about
<quantity id="4CC928BEFFB94774FA1F9DA644B7FA3F" box="[1448,1523,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="11" unit="mm" value="48.0">48 mm</quantity>
in polar diameter by
<quantity id="4CC928BEFFB94774FD449D84427AF9DD" box="[755,830,1526,1550]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="mm" value="40.0">40 mm</quantity>
in equatorial diameter, dark red, covered by areolas with numerous setose spines, about
<quantity id="4CC928BEFFB94774FCA19E6B42CDF9E2" box="[790,905,1561,1585]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.8" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="14.5" valueMax="18.0" valueMin="11.0">11–18 mm</quantity>
long, yellowish white, deciduous at maturity, sweet flesh, red; ovoid, black seeds
<quantity id="4CC928BEFFB94774FCBE9E4E42C3F987" box="[777,903,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.2" metricValueMin="1.3" pageId="10" pageNumber="11" unit="mm" value="1.75" valueMax="2.2" valueMin="1.3">1.3–2.2 mm</quantity>
long by
<quantity id="4CC928BEFFB94774FC569E4E451BF987" box="[993,1119,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="10" pageNumber="11" unit="mm" value="1.05" valueMax="1.2" valueMin="0.9">0.9–1.2 mm</quantity>
wide.
</paragraph>
<paragraph id="8B8E855BFFB94774FD829E2C429EF94A" blockId="10.[533,1536,729,1862]" pageId="10" pageNumber="11">
Common name: In
<collectingCountry id="F326C5CBFFB94774FCB09E2C4231F9A5" box="[775,885,1630,1654]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
“cacto dato”, in
<collectingCountry id="F326C5CBFFB94774FB939E2C45D4F9A5" box="[1060,1168,1630,1654]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
: “cardón”, “cardón guajiro”, &quot;yosú&quot;, &quot;panameña&quot;, and “iguaraya” (fruit).
</paragraph>
<paragraph id="8B8E855BFFB94774FD829ED14265F90D" blockId="10.[533,1536,729,1862]" pageId="10" pageNumber="11">Phenology: flowers and fruits are produced over the year, but the peak of production lasts from November to April.</paragraph>
<paragraph id="8B8E855BFFB94774FD829E9B42CAF895" blockId="10.[533,1536,729,1862]" pageId="10" pageNumber="11">
Habitat:
<taxonomicName id="4C31FED8FFB94774FD269E9B43B4F8D3" box="[657,752,1769,1793]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFB94774FD269E9B43D9F8D3" box="[657,669,1769,1792]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B9455949FFB94774FD1E9E9B43B4F8D3" box="[681,752,1769,1792]" italics="true" pageId="10" pageNumber="11">griseus</emphasis>
</taxonomicName>
grows in tropical deciduous forest and xerophytic scrub alongside with
<taxonomicName id="4C31FED8FFB94774FDA29F7E43CAF8F0" box="[533,654,1803,1827]" class="Magnoliopsida" family="Cactaceae" genus="Cereus" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9455949FFB94774FDA29F7E431FF8F0" box="[533,603,1804,1827]" italics="true" pageId="10" pageNumber="11">Cereus</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="4C31FED8FFB94774FD2F9F7E4203F8F0" box="[664,839,1803,1827]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="juliflora">
<emphasis id="B9455949FFB94774FD2F9F7E4203F8F0" box="[664,839,1803,1827]" italics="true" pageId="10" pageNumber="11">Prosopis juliflora</emphasis>
</taxonomicName>
,
<taxonomicName id="4C31FED8FFB94774FCE49F794545F8F0" box="[851,1025,1803,1827]" class="Magnoliopsida" family="Zygophyllaceae" genus="Bulnesia" kingdom="Plantae" order="Zygophyllales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="arborea">
<emphasis id="B9455949FFB94774FCE49F794545F8F0" box="[851,1025,1803,1827]" italics="true" pageId="10" pageNumber="11">Bulnesia arborea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C31FED8FFB94774FBB89F79452AF8F0" box="[1039,1134,1803,1827]" class="Magnoliopsida" family="Malvaceae" genus="Ceiba" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9455949FFB94774FBB89F79450FF8F0" box="[1039,1099,1803,1827]" italics="true" pageId="10" pageNumber="11">Ceiba</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C31FED8FFB94774FB159F79442CF8F0" box="[1186,1384,1803,1827]" class="Magnoliopsida" family="Fabaceae" genus="Haematoxylum" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9455949FFB94774FB159F794406F8F0" box="[1186,1346,1803,1827]" italics="true" pageId="10" pageNumber="11">Haematoxylum</emphasis>
sp.
</taxonomicName>
From sea level to 1200 meters above sea level.
</paragraph>
<caption id="DF4ED5D3FFB84775FFDD9EC74345F8D8" ID-DOI="http://doi.org/10.5281/zenodo.12630738" ID-Zenodo-Dep="12630738" httpUri="https://zenodo.org/record/12630738/files/figure.png" pageId="11" pageNumber="12" startId="11.[106,134,1717,1736]" targetBox="[106,1533,208,1695]" targetPageId="11" targetType="figure">
<paragraph id="8B8E855BFFB84775FFDD9EC74061F931" blockId="11.[106,1461,1717,1803]" pageId="11" pageNumber="12">
<emphasis id="B9455949FFB84775FFDD9EC740F7F91B" bold="true" box="[106,435,1717,1736]" pageId="11" pageNumber="12">
Fig 6. Areolar morphometrics
<emphasis id="B9455949FFB84775FECF9EC7403BF91B" bold="true" box="[376,383,1717,1736]" italics="true" pageId="11" pageNumber="12">t</emphasis>
-tests.
</emphasis>
(A-D) areolar features lengths comparisons, (E-F) spines counts comparisons. Levels not connected by the same letter are significantly different.
</paragraph>
<paragraph id="8B8E855BFFB84775FFDD9E8A4345F8D8" blockId="11.[106,1461,1717,1803]" box="[106,513,1784,1803]" pageId="11" pageNumber="12">https://doi.org/10.1371/journal.pone.0190385.g006</paragraph>
</caption>
<paragraph id="8B8E855BFFBF4772FD8298A242FBFE83" blockId="12.[533,1536,208,1861]" pageId="12" pageNumber="13">
Discussion:
<taxonomicName id="4C31FED8FFBF4772FD0298A24251FF34" box="[693,789,208,232]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFBF4772FD0298A24385FF34" box="[693,705,208,231]" italics="true" pageId="12" pageNumber="13">S</emphasis>
.
<emphasis id="B9455949FFBF4772FD7998A24251FF34" box="[718,789,208,231]" italics="true" pageId="12" pageNumber="13">griseus</emphasis>
</taxonomicName>
is the sole species of the genus
<taxonomicName id="4C31FED8FFBF4772FBEB98A24590FF34" box="[1116,1236,208,231]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9455949FFBF4772FBEB98A24590FF34" box="[1116,1236,208,231]" italics="true" pageId="12" pageNumber="13">Stenocereus</emphasis>
</taxonomicName>
to occur in South America. This name was actually a homonym comprising a previously undescribed northern
<collectingCountry id="F326C5CBFFBF4772FA3D9880449DFED9" box="[1418,1497,242,266]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
species (
<taxonomicName id="4C31FED8FFBF4772FDD99967424FFEFE" box="[622,779,277,301]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="huastecorum">
<emphasis id="B9455949FFBF4772FDD99967433EFEFF" box="[622,634,277,300]" italics="true" pageId="12" pageNumber="13">S</emphasis>
.
<emphasis id="B9455949FFBF4772FD319967424FFEFE" box="[646,779,277,301]" italics="true" pageId="12" pageNumber="13">huastecorum</emphasis>
</taxonomicName>
) and a second one from northern South America, which shall conserve the name
<taxonomicName id="4C31FED8FFBF4772FDC9994A4399FE9C" box="[638,733,312,336]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="griseus">
<emphasis id="B9455949FFBF4772FDC9994A43CEFE9C" box="[638,650,312,335]" italics="true" pageId="12" pageNumber="13">S</emphasis>
.
<emphasis id="B9455949FFBF4772FD21994A4399FE9C" box="[662,733,312,335]" italics="true" pageId="12" pageNumber="13">griseus</emphasis>
</taxonomicName>
by priority principle.
</paragraph>
<paragraph id="8B8E855BFFBF4772FD8299284576FE6B" blockId="12.[533,1536,208,1861]" pageId="12" pageNumber="13">
Distribution: Departments of
<collectingRegion id="49F54BB9FFBF4772FCC5992842FAFEA1" box="[882,958,346,370]" country="Colombia" name="Boyaca" pageId="12" pageNumber="13">Boyacá</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FC7E99284543FEA1" box="[969,1031,346,370]" country="Colombia" name="Huila" pageId="12" pageNumber="13">Huila</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FBA6992845C7FEA1" box="[1041,1155,346,370]" country="Colombia" name="La Guajira" pageId="12" pageNumber="13">La Guajira</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FB3A99284441FEA1" box="[1165,1285,346,370]" country="Colombia" name="Magdalena" pageId="12" pageNumber="13">Magdalena</collectingRegion>
, and
<collectingRegion id="49F54BB9FFBF4772FA8B992844ECFEA1" box="[1340,1448,346,370]" country="Colombia" name="Santander" pageId="12" pageNumber="13">Santander</collectingRegion>
in
<collectingCountry id="F326C5CBFFBF4772FDA2990F43C4FE46" box="[533,640,381,405]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
; States of
<collectingRegion id="49F54BB9FFBF4772FD5E990F4277FE46" box="[745,819,381,405]" country="Venezuela" name="Falcon" pageId="12" pageNumber="13">Falcón</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FC8B990F422AFE46" box="[828,878,381,405]" country="Venezuela" name="Lara" pageId="12" pageNumber="13">Lara</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FCC0990F4283FE46" box="[887,967,381,405]" country="Venezuela" name="Merida" pageId="12" pageNumber="13">Mérida</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FC66990F454AFE46" box="[977,1038,381,405]" country="Venezuela" name="Sucre" pageId="12" pageNumber="13">Sucre</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FBAF990F4528FE46" box="[1048,1132,381,405]" country="Venezuela" name="Tachira" pageId="12" pageNumber="13">Táchira</collectingRegion>
,
<collectingRegion id="49F54BB9FFBF4772FBC1990F4586FE46" box="[1142,1218,381,405]" country="Venezuela" name="Vargas" pageId="12" pageNumber="13">Vargas</collectingRegion>
, and
<collectingRegion id="49F54BB9FFBF4772FB4D990F4475FE46" box="[1274,1329,381,405]" country="Venezuela" name="Zulia" pageId="12" pageNumber="13">Zulia</collectingRegion>
in
<collectingCountry id="F326C5CBFFBF4772FAE5990F4486FE46" box="[1362,1474,381,405]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
, it is also found in
<collectingCountry id="F326C5CBFFBF4772FD1199D243AFFE6B" box="[678,747,416,440]" name="Aruba" pageId="12" pageNumber="13">Aruba</collectingCountry>
,
<collectingCountry id="F326C5CBFFBF4772FD4299D2420FFE6B" box="[757,843,416,440]" name="Bonaire, Sint Eustatius and Saba" pageId="12" pageNumber="13">Bonaire</collectingCountry>
, and
<collectingCountry id="F326C5CBFFBF4772FC3499D24298FE6B" box="[899,988,416,440]" name="Curaçao" pageId="12" pageNumber="13">Curaçao</collectingCountry>
islands.
</paragraph>
</subSubSection>
</emphasis>
</heading>
</treatment>
</document>
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