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<document id="159B70FD2AF71A7CB72F9A136DF28EE4" ID-DOI="10.17161/jom.v0i61.4721" ID-ISSN="2325-4467" ID-Zenodo-Dep="13228771" ID-ZooBank="urn:lsid:zoobank.org:pub:0F8F1836-A557-452E-8EBE-B48DBA597143" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893534446" checkinUser="felipe" docAuthor="Silva, Nicholai de &amp; Packer, Laurence" docDate="2016" docId="03C63032A877B515ED0C434BB44CF9B6" docLanguage="en" docName="JMelittology.2016.61.1-9.pdf" docOrigin="Journal of Melittology 2016 (61)" docSource="http://dx.doi.org/10.17161/jom.v0i61.4721" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Systropha (Systropha) sirikitae de Silva &amp; Packer 2016, new species" docType="treatment" docUuid="C4A22A9D-26B0-4245-AA8A-943986827960" docUuidSource="ZooBank" docVersion="2" lastPageNumber="5" masterDocId="FFFF484AA875B511EC29422AB015FFAD" masterDocTitle="A new species of Systropha from Thailand (Hymenoptera: Halictidae: Rophitinae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1722900272744" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="580C572C5475C112DA6D4B83B99251F6">A new species of Systropha from Thailand (Hymenoptera: Halictidae: Rophitinae)</mods:title>
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<mods:namePart id="45D7CE7042D0AA92E0FEBAEEE41D492B">Silva, Nicholai de</mods:namePart>
<mods:affiliation id="4EA2D25C7BA3F5F7D47DD64349BCAE97">Department of Biology, York University, 4700 Keele St., Toronto, Ontario, M 3 J 1 P 3, Canada (desilva. nicholai @ gmail. com; xeromelissa @ mail. com).</mods:affiliation>
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<mods:namePart id="BB031C6712EBBB88B63B51C2C43FA748">Packer, Laurence</mods:namePart>
<mods:affiliation id="B6D05083062F2C5FAB06083118E91866">Department of Biology, York University, 4700 Keele St., Toronto, Ontario, M 3 J 1 P 3, Canada (desilva. nicholai @ gmail. com; xeromelissa @ mail. com).</mods:affiliation>
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<mods:title id="82FEBD3A13EACB0AECF73ADB705B92FF">Journal of Melittology</mods:title>
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<mods:date id="362882051B2465C2CE5780307EBC183B">2016</mods:date>
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<mods:number id="9749A1872D805FCEA18A911244FF9B76">2016-04-22</mods:number>
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<mods:number id="B9208EAD666D502F91D2A6EB54E7F330">2016</mods:number>
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<mods:number id="359ABA4B5E433DC90615806DC64545AF">61</mods:number>
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<subSubSection id="C375D2AFA877B513ED0C434BB42FFE34" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BD08124A877B513ED0C434BB41EFED4" blockId="2.[293,1035,353,377]" box="[293,1035,353,377]" pageId="2" pageNumber="3">
<heading id="D0983648A877B513ED0C434BB41EFED4" box="[293,1035,353,377]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="4C6FFAA7A877B513ED0C434BB366FED4" authority="de Silva &amp; Packer" authorityName="de Silva &amp; Packer" authorityYear="2016" box="[293,883,353,377]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sirikitae" status="sp. nov." subGenus="Systropha">
<emphasis id="B91B5D36A877B513ED0C434BB289FED4" bold="true" box="[293,668,353,377]" pageId="2" pageNumber="3">
<emphasis id="B91B5D36A877B513ED0C434BB18AFED4" bold="true" box="[293,415,353,377]" italics="true" pageId="2" pageNumber="3">Systropha</emphasis>
(
<emphasis id="B91B5D36A877B513ED87434BB23DFED4" bold="true" box="[430,552,353,377]" italics="true" pageId="2" pageNumber="3">Systropha</emphasis>
)
<emphasis id="B91B5D36A877B513EE1E434BB289FED4" bold="true" box="[567,668,353,377]" italics="true" pageId="2" pageNumber="3">sirikitae</emphasis>
</emphasis>
de Silva &amp; Packer
</taxonomicName>
,
<taxonomicNameLabel id="A228E04DA877B513EF574348B41EFED4" box="[894,1035,353,377]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BD08124A877B513ECDF43A9B42FFE34" blockId="2.[246,1082,387,441]" box="[246,1082,387,409]" pageId="2" pageNumber="3">
ZooBank:
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urn:lsid:zoobank.org:act:
<uuid id="FFC9BBF1A877B513EE7C43A9B42FFE34" box="[597,1082,387,409]" pageId="2" pageNumber="3">C4A22A9D-26B0-4245-AA8A-943986827960</uuid>
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</paragraph>
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<subSubSection id="C375D2AFA877B513EE3A438BB308FE14" box="[531,797,417,441]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BD08124A877B513EE3A438BB308FE14" blockId="2.[246,1082,387,441]" box="[531,797,417,441]" pageId="2" pageNumber="3">
(
<figureCitation id="13549DA1A877B513EE32438BB296FE14" box="[539,643,417,441]" captionStart="Figures 14" captionStartId="3.[160,241,1388,1410]" captionTargetBox="[160,1168,246,1382]" captionTargetId="figure-186@3.[160,1168,224,1382]" captionTargetPageId="3" captionText="Figures 14. Antennae of species of Systropha to show variation in number of flagellomeres. 1. Systropha sirikitae, new species, holotype, with six normal-sized flagellomeres and three minute apical ones forming a small hook (scale bar = 0.5 mm). 2. Paratype of S. sirikitae with the api- cal hook formed from two flagellomeres (scale bar = 1 mm). 3. S. punjabensis Batra &amp; Michener, paratype, with 10 flagellomeres (scale bar = 0.5 mm). 4. S. inexspectata Ebmer, with the normal condition of 11 flagellomeres, right antenna (on the left) in the normal, spiral-horned position; left antenna with the spiral straightened after relaxing (scale bar = 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13228773" httpUri="https://zenodo.org/record/13228773/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
,
<figureCitation id="13549DA1A877B513EEA6438BB28BFE14" box="[655,670,417,441]" captionStart="Figure 5" captionStartId="4.[160,231,900,922]" captionTargetBox="[160,1168,224,894]" captionTargetId="figure-270@4.[160,1168,224,894]" captionTargetPageId="4" captionText="Figure 5. Holotype of Systropha sirikitae, new species, in lateral view (scale bar = 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13228775" httpUri="https://zenodo.org/record/13228775/files/figure.png" pageId="2" pageNumber="3">5</figureCitation>
,
<figureCitation id="13549DA1A877B513EE83438BB2FBFE14" box="[682,750,417,441]" captionStart="Figures 69" captionStartId="5.[160,241,989,1011]" captionTargetBox="[200,1153,224,954]" captionTargetId="figure-302@5.[160,1168,224,982]" captionTargetPageId="5" captionText="Figures 69. Apical metasomal terga and terminalia of species of Systropha (all scale bars = 1 mm). 6. Systropha sirikitae, new species, paratype, T7 to show lack of spines. 7. S. tropicalis Cockerell, oblique view to show spines (indicated by red lines). 8. S7 and S8, not separated, of paratype of S. sirikitae. 9. Genital capsule, dorsal view, of S. sirikitae." figureDoi="http://doi.org/10.5281/zenodo.13228777" httpUri="https://zenodo.org/record/13228777/files/figure.png" pageId="2" pageNumber="3">6, 8, 9</figureCitation>
,
<figureCitation id="13549DA1A877B513EED3438BB300FE14" box="[762,789,417,441]" captionStart="Figures 1013" captionStartId="6.[160,241,958,980]" captionTargetBox="[160,1168,224,951]" captionTargetId="figure-399@6.[160,1168,224,952]" captionTargetPageId="6" captionText="Figures 1013. Metasomal S2 or S2 and S3 of select species of Systropha (all scale bars = 0.5 mm). 10. Transverse process on S2 in profile (black line) of Systropha sirikitae, new species, paratype. 11. Elongate, medial process on S2 (black line) of S. inexspectata Ebmer, or a closely related, undescribed species. 12. absence of processes on S3 (green line), and weak paired convexities on S2 (red line) of S. punjabensis Batra &amp; Michener, paratype. 13. More strongly developed convexities on S3 (blue line), than S2 (red line) of S. tropicalis Cockerell." figureDoi="http://doi.org/10.5281/zenodo.13228779" httpUri="https://zenodo.org/record/13228779/files/figure.png" pageId="2" pageNumber="3">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA877B513ECF943CBB26CFCD4" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BD08124A877B513ECF943CBB26CFCD4" blockId="2.[160,1168,481,1785]" pageId="2" pageNumber="3">
DIAGNOSIS: Males can be differentiated from all other congeners globally by the combination of the nature of the antennal flagellum and that of S2. The flagellum has six well-defined flagellomeres of normal size and an additional two or three that are minute (
<figureCitation id="13549DA1A877B513ED19406BB18EFDF4" box="[304,411,577,601]" captionStart="Figures 14" captionStartId="3.[160,241,1388,1410]" captionTargetBox="[160,1168,246,1382]" captionTargetId="figure-186@3.[160,1168,224,1382]" captionTargetPageId="3" captionText="Figures 14. Antennae of species of Systropha to show variation in number of flagellomeres. 1. Systropha sirikitae, new species, holotype, with six normal-sized flagellomeres and three minute apical ones forming a small hook (scale bar = 0.5 mm). 2. Paratype of S. sirikitae with the api- cal hook formed from two flagellomeres (scale bar = 1 mm). 3. S. punjabensis Batra &amp; Michener, paratype, with 10 flagellomeres (scale bar = 0.5 mm). 4. S. inexspectata Ebmer, with the normal condition of 11 flagellomeres, right antenna (on the left) in the normal, spiral-horned position; left antenna with the spiral straightened after relaxing (scale bar = 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13228773" httpUri="https://zenodo.org/record/13228773/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
). The second metasomal sternum has a well-defined transverse ridge (
<figureCitation id="13549DA1A877B513ECC7404BB155FDD4" box="[238,320,609,633]" captionStart="Figures 1013" captionStartId="6.[160,241,958,980]" captionTargetBox="[160,1168,224,951]" captionTargetId="figure-399@6.[160,1168,224,952]" captionTargetPageId="6" captionText="Figures 1013. Metasomal S2 or S2 and S3 of select species of Systropha (all scale bars = 0.5 mm). 10. Transverse process on S2 in profile (black line) of Systropha sirikitae, new species, paratype. 11. Elongate, medial process on S2 (black line) of S. inexspectata Ebmer, or a closely related, undescribed species. 12. absence of processes on S3 (green line), and weak paired convexities on S2 (red line) of S. punjabensis Batra &amp; Michener, paratype. 13. More strongly developed convexities on S3 (blue line), than S2 (red line) of S. tropicalis Cockerell." figureDoi="http://doi.org/10.5281/zenodo.13228779" httpUri="https://zenodo.org/record/13228779/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
). Only
<taxonomicName id="4C6FFAA7A877B513EDB74048B281FDD4" authority="Friese" authorityName="Friese" authorityYear="1922" box="[414,660,609,633]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="glabriventris">
<emphasis id="B91B5D36A877B513EDB74048B1B8FDD4" box="[414,429,610,633]" italics="true" pageId="2" pageNumber="3">S</emphasis>
.
<emphasis id="B91B5D36A877B513ED92404BB250FDD4" box="[443,581,609,633]" italics="true" pageId="2" pageNumber="3">glabriventris</emphasis>
Friese
</taxonomicName>
, from
<collectingCountry id="F378C1B4A877B513EEC8404BB35DFDD4" box="[737,840,609,633]" name="Namibia" pageId="2" pageNumber="3">Namibia</collectingCountry>
and
<collectingCountry id="F378C1B4A877B513EFAD404BB410FDD4" box="[900,1029,609,633]" name="Zimbabwe" pageId="2" pageNumber="3">Zimbabwe</collectingCountry>
, has a similarly structured flagellum but it lacks protuberances on S2.
<taxonomicName id="4C6FFAA7A877B513EF7740ABB44EFD34" authorityName="Friese" authorityYear="1922" box="[862,1115,641,665]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="glabriventris">
<emphasis id="B91B5D36A877B513EF7740ABB44EFD34" box="[862,1115,641,665]" italics="true" pageId="2" pageNumber="3">Systropha glabriventris</emphasis>
</taxonomicName>
also differs in having a lateral tooth on each of T6 and T7 (
<bibRefCitation id="EFFEFCD5A877B513EF09408BB43BFD14" author="Patiny, S. &amp; D. Michez" box="[800,1070,673,697]" pageId="2" pageNumber="3" pagination="27 - 44" refId="ref4105" refString="Patiny, S., &amp; D. Michez. 2006. Phylogenetic analysis of the [sic] Systropha Illiger 1806 (Hymenoptera: Apoidea: Halictidae) and description of a new subgenus. Annales de la Societe Entomologique de France 42 (1): 27 - 44." type="journal article" year="2006">Patiny &amp; Michez, 2006</bibRefCitation>
:
<emphasis id="B91B5D36A877B513E811408BB45FFD14" box="[1080,1098,673,697]" italics="true" pageId="2" pageNumber="3">cf</emphasis>
. their figure 4 and
<figureCitation id="13549DA1A877B513ED1F40EBB16DFD77" box="[310,376,705,730]" captionStart="Figures 69" captionStartId="5.[160,241,989,1011]" captionTargetBox="[200,1153,224,954]" captionTargetId="figure-302@5.[160,1168,224,982]" captionTargetPageId="5" captionText="Figures 69. Apical metasomal terga and terminalia of species of Systropha (all scale bars = 1 mm). 6. Systropha sirikitae, new species, paratype, T7 to show lack of spines. 7. S. tropicalis Cockerell, oblique view to show spines (indicated by red lines). 8. S7 and S8, not separated, of paratype of S. sirikitae. 9. Genital capsule, dorsal view, of S. sirikitae." figureDoi="http://doi.org/10.5281/zenodo.13228777" httpUri="https://zenodo.org/record/13228777/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
);
<taxonomicName id="4C6FFAA7A877B513EDA440E8B215FD74" authorityName="de Silva &amp; Packer" authorityYear="2016" box="[397,512,705,729]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sirikitae">
<emphasis id="B91B5D36A877B513EDA440E8B189FD74" box="[397,412,706,729]" italics="true" pageId="2" pageNumber="3">S</emphasis>
.
<emphasis id="B91B5D36A877B513ED8340EBB215FD74" box="[426,512,705,729]" italics="true" pageId="2" pageNumber="3">sirikitae</emphasis>
</taxonomicName>
lacks such teeth (
<figureCitation id="13549DA1A877B513EEF840EBB30AFD74" box="[721,799,705,729]" captionStart="Figure 5" captionStartId="4.[160,231,900,922]" captionTargetBox="[160,1168,224,894]" captionTargetId="figure-270@4.[160,1168,224,894]" captionTargetPageId="4" captionText="Figure 5. Holotype of Systropha sirikitae, new species, in lateral view (scale bar = 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13228775" httpUri="https://zenodo.org/record/13228775/files/figure.png" pageId="2" pageNumber="3">Figs. 5</figureCitation>
,
<figureCitation id="13549DA1A877B513EF0240EBB32EFD74" box="[811,827,705,729]" captionStart="Figures 69" captionStartId="5.[160,241,989,1011]" captionTargetBox="[200,1153,224,954]" captionTargetId="figure-302@5.[160,1168,224,982]" captionTargetPageId="5" captionText="Figures 69. Apical metasomal terga and terminalia of species of Systropha (all scale bars = 1 mm). 6. Systropha sirikitae, new species, paratype, T7 to show lack of spines. 7. S. tropicalis Cockerell, oblique view to show spines (indicated by red lines). 8. S7 and S8, not separated, of paratype of S. sirikitae. 9. Genital capsule, dorsal view, of S. sirikitae." figureDoi="http://doi.org/10.5281/zenodo.13228777" httpUri="https://zenodo.org/record/13228777/files/figure.png" pageId="2" pageNumber="3">6</figureCitation>
). There are only two known
<taxonomicName id="4C6FFAA7A877B513EC8940CBB119FD54" authorityName="Illiger" authorityYear="1806" box="[160,268,737,761]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91B5D36A877B513EC8940CBB119FD54" box="[160,268,737,761]" italics="true" pageId="2" pageNumber="3">Systropha</emphasis>
</taxonomicName>
from southeastern Asia:
<taxonomicName id="4C6FFAA7A877B513EE1C40C8B2EAFD54" authority="Smith" authorityName="Smith" authorityYear="1879" box="[565,767,737,761]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="difformis">
<emphasis id="B91B5D36A877B513EE1C40C8B251FD54" box="[565,580,738,761]" italics="true" pageId="2" pageNumber="3">S</emphasis>
.
<emphasis id="B91B5D36A877B513EE7840CBB2A4FD54" box="[593,689,737,761]" italics="true" pageId="2" pageNumber="3">difformis</emphasis>
Smith
</taxonomicName>
known only from the Republic of the Union of
<collectingCountry id="F378C1B4A877B513ED124128B1BAFCB7" box="[315,431,770,794]" name="Myanmar" pageId="2" pageNumber="3">Myanmar</collectingCountry>
[
<collectingCountry id="F378C1B4A877B513ED954128B205FCB4" box="[444,528,770,793]" name="Myanmar" pageId="2" pageNumber="3">Burma</collectingCountry>
] and
<taxonomicName id="4C6FFAA7A877B513EE784128B2E4FCB4" box="[593,753,770,793]" class="Insecta" genus="Systrophidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="inexspectata">
<emphasis id="B91B5D36A877B513EE784128B275FCB4" box="[593,608,770,793]" italics="true" pageId="2" pageNumber="3">S</emphasis>
.
<emphasis id="B91B5D36A877B513EE444128B2E4FCB4" box="[621,753,770,793]" italics="true" pageId="2" pageNumber="3">inexspectata</emphasis>
</taxonomicName>
from
<collectingCountry id="F378C1B4A877B513EF11412BB3B0FCB4" box="[824,933,769,793]" name="Thailand" pageId="2" pageNumber="3">Thailand</collectingCountry>
. The males of these both have the normal 11 flagellomeres (
<figureCitation id="13549DA1A877B513EE56410BB2D7FC94" box="[639,706,801,825]" captionStart="Figures 14" captionStartId="3.[160,241,1388,1410]" captionTargetBox="[160,1168,246,1382]" captionTargetId="figure-186@3.[160,1168,224,1382]" captionTargetPageId="3" captionText="Figures 14. Antennae of species of Systropha to show variation in number of flagellomeres. 1. Systropha sirikitae, new species, holotype, with six normal-sized flagellomeres and three minute apical ones forming a small hook (scale bar = 0.5 mm). 2. Paratype of S. sirikitae with the api- cal hook formed from two flagellomeres (scale bar = 1 mm). 3. S. punjabensis Batra &amp; Michener, paratype, with 10 flagellomeres (scale bar = 0.5 mm). 4. S. inexspectata Ebmer, with the normal condition of 11 flagellomeres, right antenna (on the left) in the normal, spiral-horned position; left antenna with the spiral straightened after relaxing (scale bar = 2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13228773" httpUri="https://zenodo.org/record/13228773/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
) (although the apical ones are narrow and form a spiral in both species), and a single, median process on S2 (
<emphasis id="B91B5D36A877B513EFF74168B3E8FCF4" box="[990,1021,834,857]" italics="true" pageId="2" pageNumber="3">e.g</emphasis>
.,
<figureCitation id="13549DA1A877B513E838416BB44AFCF4" box="[1041,1119,833,857]" captionStart="Figures 1013" captionStartId="6.[160,241,958,980]" captionTargetBox="[160,1168,224,951]" captionTargetId="figure-399@6.[160,1168,224,952]" captionTargetPageId="6" captionText="Figures 1013. Metasomal S2 or S2 and S3 of select species of Systropha (all scale bars = 0.5 mm). 10. Transverse process on S2 in profile (black line) of Systropha sirikitae, new species, paratype. 11. Elongate, medial process on S2 (black line) of S. inexspectata Ebmer, or a closely related, undescribed species. 12. absence of processes on S3 (green line), and weak paired convexities on S2 (red line) of S. punjabensis Batra &amp; Michener, paratype. 13. More strongly developed convexities on S3 (blue line), than S2 (red line) of S. tropicalis Cockerell." figureDoi="http://doi.org/10.5281/zenodo.13228779" httpUri="https://zenodo.org/record/13228779/files/figure.png" pageId="2" pageNumber="3">Fig. 11</figureCitation>
; see also
<bibRefCitation id="EFFEFCD5A877B513ECFF414BB17FFCD4" author="Ebmer, A. W." box="[214,362,865,889]" pageId="2" pageNumber="3" pagination="807 - 821" refId="ref3873" refString="Ebmer, A. W. 1994. Systropha difformis Smith, 1879 und Systropha inexspectata n. sp., die beiden ostlichen Vertreter der altweltlichen Gattung Systropha Illiger 1806 (Insecta: Hymenoptera: Apoidea: Halictidae: Rophitinae). Linzer Biologische Beitrage 26 (2): 807 - 821." type="journal article" year="1994">Ebmer, 1994</bibRefCitation>
: figures 10, 11 therein).
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA877B515ECF941ABB17AFB36" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BD08124A877B513ECF941ABB1E5FC14" blockId="2.[160,1168,481,1785]" pageId="2" pageNumber="3">
DESCRIPTION:
<specimenCount id="9D694AADA877B513ED5941A8B191FC31" box="[368,388,898,924]" count="1" pageId="2" pageNumber="3" type="male"></specimenCount>
: Body length
<quantity id="4C972CC1A877B513EE0241ABB26BFC34" box="[555,638,897,921]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="3" unit="mm" value="11.0">11 mm</quantity>
, forewing length
<quantity id="4C972CC1A877B513EF7941ABB3BFFC34" box="[848,938,897,921]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="3" unit="mm" value="6.8">6.8 mm</quantity>
, intertegular width
<quantity id="4C972CC1A877B513EC89418BB0ECFC14" box="[160,249,929,953]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="mm" value="1.8">1.8 mm</quantity>
, head width
<quantity id="4C972CC1A877B513EDBB418BB1F9FC14" box="[402,492,929,953]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="3" unit="mm" value="2.3">2.3 mm</quantity>
.
</paragraph>
<paragraph id="8BD08124A877B513ECF941EBB42BFB94" blockId="2.[160,1168,481,1785]" pageId="2" pageNumber="3">
<emphasis id="B91B5D36A877B513ECF941EBB14CFC74" bold="true" box="[208,345,961,985]" pageId="2" pageNumber="3">Coloration.</emphasis>
Black except as follows: F7F9 very pale to light brown; tegula brown, translucent posteriorly and laterally; wing veins translucent brown; T6 entirely red-brown, somewhat translucent apically; pretarsal claws red-brown; metatibial spurs brown, becoming darker distally; antennal cleaner almost entirely transparent.
</paragraph>
<paragraph id="8BD08124A877B513ECF9466BB16BFA34" blockId="2.[160,1168,481,1785]" pageId="2" pageNumber="3">
<emphasis id="B91B5D36A877B513ECF9466BB152FBF4" bold="true" box="[208,327,1089,1113]" pageId="2" pageNumber="3">Structure.</emphasis>
Head broader than long, width to length ratio 4:3. Labrum slightly shiny, basal area with irregular carinulae, apical area distinctly punctate. Mouthparts long; length of labial palpus approximately 1.5X length of head; glossa approximately 1.5X as long as labial palpus. Clypeus basal two fifths with dense, small punctures, i&lt;d, rest shiny with large, sparse, longitudinally effaced punctures, i&lt;d; with dull yellow hairs. Supraclypeal area densely and finely punctured (i&lt;d). Frontal area densely punctate, i&lt;d; upper paraocular area minutely and sparsely punctate, ocellocular area largely impunctate. Scape minutely and densely punctate; F1 twice as long as greatest breadth. Antenna reduced apically, F7F9 minute, superficially appearing as if with only 7 flagellomeres. Genal area shiny, obscurely and sparsely punctate (i&gt;3d); genal beard long (L&lt;2D).
</paragraph>
<paragraph id="8BD08124A877B513ECF94788B485F914" blockId="2.[160,1168,481,1785]" pageId="2" pageNumber="3">Mesoscutum doubly and finely punctate, i&lt;1.5d between median and parapsidal lines; large punctures sparse except on anterior one third (i=13d); pubescence sparse, short (L&lt;1.3D, somewhat longer anteriorly L&lt;2D). Mesoscutellum with punctures as on mesoscutum except smaller and somewhat sparser. Metanotum with transversely effaced punctures i&lt;2d. Metapostnotum imbricate, dull, margins smooth, lacking microsculpture, side of propodeum shiny, distinctly punctate, i=2d; pubescence long (L&lt;2D). Mesosoma lacking ventral protuberances. Fm1 and Fm3 thickest at basal one quarter, Fm2 thickest at midlength. Tb1 and Tb2 unmodified, Tb3 somewhat swollen, thickest subapically. Anterior metatibial spur curved apically, shorter than posterior spur.</paragraph>
<paragraph id="8BD08124A877B515ECF944E8B385FBD6" blockId="2.[160,1168,481,1785]" lastBlockId="4.[160,1169,963,1563]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">Terga without lateral spines or teeth. T1 distinctly and regularly punctate (i ≈ d), apical impressed area more finely and sparsely punctate (i&gt;d); pilosity pale, longest apicolaterally (L ≈1.5D), remaining hairs short (L&lt;D). T2T3 as for T1 except basal areas depressed and with punctures similar to those on apical impressed areas, apical impressed area longer, hairs towards side of disc short, thick, and dark brown; T4T6 lacking basal depressed areas, punctures on discs smaller. Hairs increasing in length from T2 to T6, on T6 ≈3D, proportionately more dark brown hairs on more posterior terga; hairs simple except somewhat plumose towards sides of terga. Sterna shiny, with row of hairs subapically; sparsely and irregularly punctate (i&gt;2d). S2 with a pronounced transverse ridge extending almost entirely across exposed portion of sternum. S5 weakly concave basomedially. S6 with raised area medially ending in short blunt process. Erect hairs longest on S1 and towards sides of S3S4 (L&lt;2D). T7 lacking lateral tooth but with broadly rounded convexity on either side.</paragraph>
<caption id="DF10D1ACA876B512EC894746B3BCF983" ID-DOI="http://doi.org/10.5281/zenodo.13228773" ID-Zenodo-Dep="13228773" httpUri="https://zenodo.org/record/13228773/files/figure.png" pageId="3" pageNumber="4" startId="3.[160,241,1388,1410]" targetBox="[160,1168,246,1382]" targetPageId="3" targetType="figure">
<paragraph id="8BD08124A876B512EC894746B3BCF983" blockId="3.[160,1175,1388,1583]" pageId="3" pageNumber="4">
<emphasis id="B91B5D36A876B512EC894746B134FA2C" bold="true" box="[160,289,1388,1410]" pageId="3" pageNumber="4">Figures 14.</emphasis>
Antennae of species of
<taxonomicName id="4C6FFAA7A876B512EE034746B285FA2C" authorityName="Illiger" authorityYear="1806" box="[554,656,1388,1409]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91B5D36A876B512EE034746B285FA2C" box="[554,656,1388,1409]" italics="true" pageId="3" pageNumber="4">Systropha</emphasis>
</taxonomicName>
to show variation in number of flagellomeres.
<emphasis id="B91B5D36A876B512EC8947A3B0A7FA33" bold="true" box="[160,178,1417,1438]" pageId="3" pageNumber="4">1.</emphasis>
<taxonomicName id="4C6FFAA7A876B512EC9147A3B178FA33" authorityName="de Silva &amp; Packer" authorityYear="2016" box="[184,365,1417,1438]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sirikitae" status="sp. nov.">
<emphasis id="B91B5D36A876B512EC9147A3B178FA33" box="[184,365,1417,1438]" italics="true" pageId="3" pageNumber="4">Systropha sirikitae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A228E04DA876B512ED5347A0B1E9FA32" box="[378,508,1417,1439]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
, holotype, with six normal-sized flagellomeres and three minute apical ones forming a small hook (scale bar = 0.5 mm).
<emphasis id="B91B5D36A876B512EF20478CB30EFA16" bold="true" box="[777,795,1446,1467]" pageId="3" pageNumber="4">2.</emphasis>
Paratype of
<taxonomicName id="4C6FFAA7A876B512EFB7478CB411FA16" authorityName="de Silva &amp; Packer" authorityYear="2016" box="[926,1028,1446,1467]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sirikitae">
<emphasis id="B91B5D36A876B512EFB7478CB3BEFA16" box="[926,939,1446,1467]" italics="true" pageId="3" pageNumber="4">S</emphasis>
.
<emphasis id="B91B5D36A876B512EF9E478CB411FA16" box="[951,1028,1446,1467]" italics="true" pageId="3" pageNumber="4">sirikitae</emphasis>
</taxonomicName>
with the apical hook formed from two flagellomeres (scale bar = 1 mm).
<emphasis id="B91B5D36A876B512EF0D47E9B323FA75" bold="true" box="[804,822,1475,1496]" pageId="3" pageNumber="4">3.</emphasis>
<taxonomicName id="4C6FFAA7A876B512EF1547E9B499FA74" authority="Batra &amp; Michener" authorityName="Batra &amp; Michener" authorityYear="1966" box="[828,1164,1475,1497]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="punjabensis">
<emphasis id="B91B5D36A876B512EF1547E9B35CFA75" box="[828,841,1475,1496]" italics="true" pageId="3" pageNumber="4">S</emphasis>
.
<emphasis id="B91B5D36A876B512EF7C47E9B3DDFA75" box="[853,968,1475,1496]" italics="true" pageId="3" pageNumber="4">punjabensis</emphasis>
Batra &amp; Michener
</taxonomicName>
, paratype, with 10 flagellomeres (scale bar = 0.5 mm).
<emphasis id="B91B5D36A876B512EEC847F5B2E6FA59" bold="true" box="[737,755,1503,1524]" pageId="3" pageNumber="4">4.</emphasis>
<taxonomicName id="4C6FFAA7A876B512EED047CAB3CCFA58" authority="Ebmer" authorityName="Ebmer" box="[761,985,1503,1525]" class="Insecta" genus="Systrophidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="7" phylum="Arthropoda" rank="species" species="inexspectata">
S.
<emphasis id="B91B5D36A876B512EF3A47CAB39FFA58" box="[787,906,1504,1525]" italics="true" pageId="3" pageNumber="4">inexspectata</emphasis>
Ebmer
</taxonomicName>
, with the normal condition of 11 flagellomeres, right antenna (on the left) in the normal, spiral-horned position; left antenna with the spiral straightened after relaxing (scale bar = 2 mm).
</paragraph>
</caption>
<caption id="DF10D1ACA871B515EC8941AEB454FC37" ID-DOI="http://doi.org/10.5281/zenodo.13228775" ID-Zenodo-Dep="13228775" box="[160,1089,900,923]" httpUri="https://zenodo.org/record/13228775/files/figure.png" pageId="4" pageNumber="5" startId="4.[160,231,900,922]" targetBox="[160,1168,224,894]" targetPageId="4" targetType="figure">
<paragraph id="8BD08124A871B515EC8941AEB454FC37" blockId="4.[160,1089,900,923]" box="[160,1089,900,923]" pageId="4" pageNumber="5">
<emphasis id="B91B5D36A871B515EC8941AEB0EAFC37" bold="true" box="[160,255,900,922]" pageId="4" pageNumber="5">Figure 5.</emphasis>
Holotype of
<taxonomicName id="4C6FFAA7A871B515EDB941AFB251FC37" authorityName="de Silva &amp; Packer" authorityYear="2016" box="[400,580,901,922]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sirikitae" status="sp. nov.">
<emphasis id="B91B5D36A871B515EDB941AFB251FC37" box="[400,580,901,922]" italics="true" pageId="4" pageNumber="5">Systropha sirikitae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A228E04DA871B515EE7841AFB2C7FC36" box="[593,722,901,923]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
, in lateral view (scale bar = 2 mm).
</paragraph>
</caption>
<paragraph id="8BD08124A871B515ECF946AEB17AFB36" blockId="4.[160,1169,963,1563]" box="[208,367,1155,1182]" pageId="4" pageNumber="5">
<specimenCount id="9D694AADA871B515ECF946AEB0F1FB33" box="[208,228,1156,1182]" count="1" pageId="4" pageNumber="5" type="female"></specimenCount>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA871B515ECF94689B3F1FA96" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BD08124A871B515ECF94689B30EFB56" blockId="4.[160,1169,963,1563]" pageId="4" pageNumber="5">
<materialsCitation id="3B078B79A871B515ECF94689B30EFB56" collectingDate="2006-09-19" collectingDateMax="2006-09-26" collectingDateMin="2006-09-19" collectionCode="QSBG" collectorName="Pha Hin Ngam NP &amp; Sa-nog &amp; Adnafai. &amp; B. To &amp; Queen Sirikit Botanical Gardens" country="Thailand" elevation="681" latitude="15.62805" location="Chaiyaphum" longLatPrecision="1" longitude="101.38872" pageId="4" pageNumber="5" specimenCode="T856" specimenCount="682" specimenCount-male="1" stateProvince="Chaiyaphum" typeStatus="holotype">
<typeStatus id="54D43F86A871B515ECF94689B15DFB16" box="[208,328,1187,1211]" pageId="4" pageNumber="5" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D694AADA871B515ED74468EB164FB13" box="[349,369,1188,1214]" count="1" pageId="4" pageNumber="5" type="male"></specimenCount>
,
<collectingCountry id="F378C1B4A871B515EDA34689B234FB16" box="[394,545,1187,1211]" name="Thailand" pageId="4" pageNumber="5">THAILAND</collectingCountry>
:
<collectingRegion id="49AB4FC6A871B515EE1E4689B2C6FB16" box="[567,723,1187,1211]" country="Thailand" name="Chaiyaphum" pageId="4" pageNumber="5">Chaiyaphum</collectingRegion>
,
<collectorName id="269AE4F2A871B515EEC34689B3FBFB16" box="[746,1006,1187,1212]" pageId="4" pageNumber="5">Pha Hin Ngam NP</collectorName>
.
<geoCoordinate id="EE5BE7E3A871B515E82D4689B485FB16" box="[1028,1168,1187,1211]" degrees="15" direction="north" minutes="37.683" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="15.62805">15°37.683N</geoCoordinate>
<geoCoordinate id="EE5BE7E3A871B515EC8946E9B126FB76" box="[160,307,1219,1243]" degrees="101" direction="east" minutes="23.323" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="101.38872">101°23.323E</geoCoordinate>
<specimenCount id="9D694AADA871B515ED1246E9B19DFB76" box="[315,392,1219,1243]" count="681" pageId="4" pageNumber="5" type="generic">
<quantity id="4C972CC1A871B515ED1246E9B191FB76" box="[315,388,1219,1243]" metricMagnitude="2" metricUnit="m" metricValue="6.81" pageId="4" pageNumber="5" unit="m" value="681.0">
<elevation id="00426617A871B515ED1246E9B191FB76" box="[315,388,1219,1243]" metricMagnitude="2" metricUnit="m" metricValue="6.81" pageId="4" pageNumber="5" unit="m" value="681.0">681 m</elevation>
</quantity>
.
</specimenCount>
<collectingMethod id="522EF933A871B515EDB946E9B231FB76" box="[400,548,1219,1243]" pageId="4" pageNumber="5">malaise trap</collectingMethod>
.
<date id="FFD1A7E4A871B515EE0646E9B2D8FB76" box="[559,717,1219,1243]" pageId="4" pageNumber="5" value="2006-09-19" valueMax="2006-09-26" valueMin="2006-09-19">
<collectingDate id="EF955E0CA871B515EE0646E9B2D8FB76" box="[559,717,1219,1243]" pageId="4" pageNumber="5" value="2006-09-19" valueMax="2006-09-26" valueMin="2006-09-19">1926.ix.2006</collectingDate>
</date>
. coll:
<collectorName id="269AE4F2A871B515EF3946E9B374FB76" box="[784,865,1219,1243]" pageId="4" pageNumber="5">Sa-nog</collectorName>
&amp;
<collectorName id="269AE4F2A871B515EFAA46E9B3FFFB76" box="[899,1002,1219,1243]" pageId="4" pageNumber="5">Adnafai.</collectorName>
<specimenCode id="DBC9295FA871B515EFD846E9B439FB76" box="[1009,1068,1219,1243]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T856</specimenCode>
<collectorName id="269AE4F2A871B515E80546E9B478FB76" box="[1068,1133,1219,1243]" pageId="4" pageNumber="5">B. To</collectorName>
be deposited in
<collectorName id="269AE4F2A871B515ED1046C9B2A1FB56" box="[313,692,1251,1275]" pageId="4" pageNumber="5">Queen Sirikit Botanical Gardens</collectorName>
(
<collectionCode id="ED7E19E1A871B515EEEB46C9B31AFB56" box="[706,783,1251,1275]" pageId="4" pageNumber="5">QSBG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD08124A871B515ECF94729B3F1FA96" blockId="4.[160,1169,963,1563]" pageId="4" pageNumber="5">
<materialsCitation id="3B078B79A871B515ECF94729B0F7FA96" collectorName="Phu Kradeung NP" country="Thailand" elevation="275" latitude="16.818317" location="Loei" longLatPrecision="1" longitude="101.79373" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
<typeStatus id="54D43F86A871B515ECF94729B156FAB6" box="[208,323,1283,1307]" pageId="4" pageNumber="5" type="paratype">PARATYPE</typeStatus>
:
<specimenCount id="9D694AADA871B515ED7B472EB173FAB3" box="[338,358,1284,1310]" count="1" pageId="4" pageNumber="5" type="male"></specimenCount>
,
<collectingCountry id="F378C1B4A871B515ED534729B204FAB6" box="[378,529,1283,1307]" name="Thailand" pageId="4" pageNumber="5">THAILAND</collectingCountry>
:
<collectingRegion id="49AB4FC6A871B515EE0B4729B24DFAB6" box="[546,600,1283,1307]" country="Thailand" name="Loei" pageId="4" pageNumber="5">Loei</collectingRegion>
,
<collectorName id="269AE4F2A871B515EE404729B347FAB6" box="[617,850,1283,1307]" pageId="4" pageNumber="5">Phu Kradeung NP</collectorName>
.
<geoCoordinate id="EE5BE7E3A871B515EF4A4729B3FAFAB6" box="[867,1007,1283,1307]" degrees="16" direction="north" minutes="49.099" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="16.818317">16°49.099N</geoCoordinate>
<geoCoordinate id="EE5BE7E3A871B515EFD54729B485FAB6" box="[1020,1168,1283,1307]" degrees="101" direction="east" minutes="47.624" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="101.79373">101°47.624E</geoCoordinate>
<quantity id="4C972CC1A871B515EC894709B0F7FA96" box="[160,226,1315,1339]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="4" pageNumber="5" unit="m" value="275.0">
<specimenCount id="9D694AADA871B515EC894709B0F7FA96" box="[160,226,1315,1339]" count="275" pageId="4" pageNumber="5" type="generic">
<elevation id="00426617A871B515EC894709B0F7FA96" box="[160,226,1315,1339]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="4" pageNumber="5" unit="m" value="275.0">275m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B078B79A871B515ECCA4709B3F5FA96" box="[227,992,1315,1339]" collectingDate="2006-11-14" collectionCode="MT, PCYU" collectorName="S. Gong-lasae. &amp; A. Deposited" country="Thailand" pageId="4" pageNumber="5" specimenCode="T1074" specimenCount="1" typeStatus="paratype">
.
<collectionCode id="ED7E19E1A871B515ECC4470EB10CFA96" box="[237,281,1316,1339]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="4" pageNumber="5">MT</collectionCode>
.
<date id="FFD1A7E4A871B515ED0A4709B18CFA96" box="[291,409,1315,1339]" pageId="4" pageNumber="5" value="2006-11-14">
<collectingDate id="EF955E0CA871B515ED0A4709B18CFA96" box="[291,409,1315,1339]" pageId="4" pageNumber="5" value="2006-11-14">14.xi.2006</collectingDate>
</date>
. coll:
<collectorName id="269AE4F2A871B515EDF34709B294FA96" box="[474,641,1315,1339]" pageId="4" pageNumber="5">S. Gong-lasae.</collectorName>
<specimenCode id="DBC9295FA871B515EEA14709B2C4FA96" box="[648,721,1315,1339]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="5">T1074</specimenCode>
<collectorName id="269AE4F2A871B515EEF84709B37AFA96" box="[721,879,1315,1339]" pageId="4" pageNumber="5">A. Deposited</collectorName>
in
<collectionCode id="ED7E19E1A871B515EFBA4709B3F5FA96" box="[915,992,1315,1339]" country="Canada" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="4" pageNumber="5" type="University or college">PCYU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA871B515ECF94769B0DAFA36" pageId="4" pageNumber="5" type="description">
<paragraph id="8BD08124A871B515ECF94769B0DAFA36" blockId="4.[160,1169,963,1563]" pageId="4" pageNumber="5">
VARIATION:
<typeStatus id="54D43F86A871B515ED74476EB1D3FAF1" box="[349,454,1348,1372]" pageId="4" pageNumber="5" type="paratype">Paratype</typeStatus>
as in
<typeStatus id="54D43F86A871B515EE3A4769B269FAF6" box="[531,636,1347,1371]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
except: only 8 flagellomeres, F7F8 minute, forming a hook, and bee slightly smaller, head width
<quantity id="4C972CC1A871B515EF3A4749B378FAD6" box="[787,877,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="5" unit="mm" value="2.2">2.2 mm</quantity>
. Terminalia as in figures 89.
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA871B515ECF94789B22BFA76" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BD08124A871B515ECF94789B22BFA76" blockId="4.[160,1169,963,1563]" pageId="4" pageNumber="5">
ETYMOLOGY: The species is named after the botanical gardens where the
<typeStatus id="54D43F86A871B515E80E4789B485FA16" box="[1063,1168,1443,1467]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
will be deposited, QSBG,
<collectingCountry id="F378C1B4A871B515EDE447E9B22FFA76" box="[461,570,1475,1499]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375D2AFA871B515ECF947C9B44CF9B6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BD08124A871B515ECF947C9B44CF9B6" blockId="4.[160,1169,963,1563]" pageId="4" pageNumber="5">
COMMENTS: In the locality labels, NP refers to National Park: both of the known specimens are from national parks. The two localities are less than
<quantity id="4C972CC1A871B515EF904429B418F9B6" box="[953,1037,1539,1563]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="km" value="150.0">150 km</quantity>
apart.
</paragraph>
</subSubSection>
</treatment>
</document>