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<document id="BCE02C92FD20F80D9E3CCA0D363DBBA7" ID-CLB-Dataset="302556" ID-DOI="10.11646/phytotaxa.372.4.4" ID-GBIF-Dataset="23351b50-e69f-4e42-bb45-8d347973d352" ID-ISSN="1179-3163" ID-Zenodo-Dep="13727338" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725678166433" checkinUser="felipe" docAuthor="Baranow, Przemyslaw &amp; Dudek, Magdalena" docDate="2018" docId="03F1DF0DFFFDFFD3FF6CFD4AFD59AE4B" docLanguage="en" docName="phytotaxa.372.4.4.pdf" docOrigin="Phytotaxa 372 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.372.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Sobralia fugax Baranow &amp; Dudek 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="278" masterDocId="FFC8A775FFFEFFD6FFE4FFFEFF9EA901" masterDocTitle="Sobralia fugax (Orchidaceae: Sobraliae) - a new species from Colombia described on the basis of morphological study and phylogenetic analyses" masterLastPageNumber="282" masterPageNumber="273" pageNumber="276" updateTime="1725690704332" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="396F1090AB776B7F7EBEBFA9FCCAA1EA">Sobralia fugax (Orchidaceae: Sobraliae) - a new species from Colombia described on the basis of morphological study and phylogenetic analyses</mods:title>
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<mods:namePart id="F3057B433DA455B75D93B35B4B3A8BFE">Baranow, Przemyslaw</mods:namePart>
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<subSubSection id="C3423D90FFFDFFD5FF6CFD4AFCB9ABCE" box="[136,807,692,719]" pageId="3" pageNumber="276" type="nomenclature">
<paragraph id="8BE76E1BFFFDFFD5FF6CFD4AFCB9ABCE" blockId="3.[136,807,692,719]" box="[136,807,692,719]" pageId="3" pageNumber="276">
<heading id="D0AFD977FFFDFFD5FF6CFD4AFCB9ABCE" box="[136,807,692,719]" fontSize="11" level="2" pageId="3" pageNumber="276" reason="3">
<taxonomicName id="4C581598FFFDFFD5FF6CFD4AFD91ABCE" ID-CoL="8X6CF" authority="Baranow &amp; Dudek" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[136,527,692,719]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="fugax" status="sp. nov.">
<emphasis id="B92CB209FFFDFFD5FF6CFD4AFEAAABCF" bold="true" box="[136,308,692,719]" italics="true" pageId="3" pageNumber="276">Sobralia fugax</emphasis>
Baranow &amp; Dudek
</taxonomicName>
<taxonomicNameLabel id="A21F0F72FFFDFFD5FDF1FD48FDF2ABCE" box="[533,620,694,719]" pageId="3" pageNumber="276" rank="species">
<emphasis id="B92CB209FFFDFFD5FDF1FD48FDFAABCE" box="[533,612,694,719]" italics="true" pageId="3" pageNumber="276">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="1363729EFFFDFFD5FD9EFD48FD7BABCE" box="[634,741,694,719]" captionStart="FIGURE 1" captionStartId="4.[136,229,1914,1935]" captionTargetBox="[176,1407,224,1859]" captionTargetId="figure-15@4.[151,1436,190,1889]" captionTargetPageId="4" captionText="FIGURE 1. Sobralia fugax Baranow &amp; Dudek:A. Habit. B. Apical part of stem with the inflorescence. C. Dorsal sepal. D. Lateral sepal. E. Petal. F. Lip. Natural position. G. Lip, spread (drawn from the holotype)." figureDoi="http://doi.org/10.5281/zenodo.13727340" httpUri="https://zenodo.org/record/13727340/files/figure.png" pageId="3" pageNumber="276">Figures 1</figureCitation>
&amp;
<figureCitation id="1363729EFFFDFFD5FCEDFD48FC84ABCE" box="[777,794,694,719]" captionStart="FIGURE 2" captionStartId="5.[136,229,1124,1145]" captionTargetBox="[151,1436,190,1099]" captionTargetId="figure-14@5.[151,1436,190,1099]" captionTargetPageId="5" captionText="FIGURE 2. Sobralia fugax Baranow &amp; Dudek: A. Flower, side view. B. Dorsal sepal. C. Lateral sepal. D. Petal. E. Lip. F. Interior surface of lip. G. Column (photographs taken from the holotype)" figureDoi="http://doi.org/10.5281/zenodo.13727342" httpUri="https://zenodo.org/record/13727342/files/figure.png" pageId="3" pageNumber="276">2</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3423D90FFFDFFD5FF6CFD22FAF5AA80" pageId="3" pageNumber="276" type="materials_examined">
<paragraph id="8BE76E1BFFFDFFD5FF6CFD22FEB2AA14" blockId="3.[136,1453,732,897]" pageId="3" pageNumber="276">
Type:—
<collectingCountry id="F34F2E8BFFFDFFD5FF32FD22FEC4ABF0" box="[214,346,732,753]" name="Colombia" pageId="3" pageNumber="276">COLOMBIA</collectingCountry>
.
<collectingRegion id="499CA0F9FFFDFFD5FE8EFD22FD8DABF0" box="[362,531,732,753]" country="Colombia" name="Valle del Cauca" pageId="3" pageNumber="276">Valle del Cauca</collectingRegion>
,
<date id="FFE648DBFFFDFFD5FDC1FD22FD16ABF0" box="[549,648,732,753]" pageId="3" pageNumber="276" value="2011-02">Feb 2011</date>
,
<emphasis id="B92CB209FFFDFFD5FD7EFD22FD7BABF0" box="[666,741,732,753]" italics="true" pageId="3" pageNumber="276">Niessen</emphasis>
cult. sub
<emphasis id="B92CB209FFFDFFD5FCBEFD22FC6EABF0" box="[858,1008,732,753]" italics="true" pageId="3" pageNumber="276">Szlachetko s.n.</emphasis>
(
<typeStatus id="54E3D0B9FFFDFFD5FBE0FD22FBC4ABF0" box="[1028,1114,732,753]" pageId="3" pageNumber="276" type="holotype">holotype</typeStatus>
<collectionCode id="ED49F6DEFFFDFFD5FB8CFD22FB2BABF0" box="[1128,1205,732,753]" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14889" lsid="urn:lsid:biocol.org:col:14889" name="Gdansk University" pageId="3" pageNumber="276" type="Herbarium">UGDA</collectionCode>
!,
<typeStatus id="54E3D0B9FFFDFFD5FB2DFD22FA90ABF0" box="[1225,1294,732,753]" pageId="3" pageNumber="276" type="isotype">isotype</typeStatus>
<collectionCode id="ED49F6DEFFFDFFD5FAF9FD22FAFCABF0" box="[1309,1378,732,753]" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14889" lsid="urn:lsid:biocol.org:col:14889" name="Gdansk University" pageId="3" pageNumber="276" type="Herbarium">UGDA</collectionCode>
—spirit collection!)
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF6CFCDAFAF5AA80" blockId="3.[136,1453,732,897]" pageId="3" pageNumber="276">
<emphasis id="B92CB209FFFDFFD5FF6CFCDAFAF5AA80" italics="true" pageId="3" pageNumber="276">
Different from other species of
<taxonomicName id="4C581598FFFDFFD5FE5AFCDAFD26AA38" authority="section Sobralia" box="[446,696,804,825]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="section" section="Sobralia">Sobralia section Sobralia</taxonomicName>
by the greenish, fleshy sepals and strongly bent ovary which determines the orientation of the flower making it hanged down in contrast to other species of the group which produce flowers directed forward. Additionally that sepals and petals of the new species are not reflexed making the flower appearance distinctly more slender.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3423D90FFFDFFD3FF6CFC4CFD59AE4B" lastPageId="5" lastPageNumber="278" pageId="3" pageNumber="276" type="etymology">
<paragraph id="8BE76E1BFFFDFFD5FF6CFC4CFD42AAEE" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
<emphasis id="B92CB209FFFDFFD5FF6CFC4CFEB0AACA" bold="true" box="[136,302,946,971]" pageId="3" pageNumber="276">Etymology:—</emphasis>
From the Latin
<taxonomicName id="4C581598FFFDFFD5FE3AFC4CFD82AACA" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[478,540,946,971]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFDFFD5FE3AFC4CFD82AACA" box="[478,540,946,971]" italics="true" pageId="3" pageNumber="276">fugax</emphasis>
</taxonomicName>
(ashamed). Named in an allusion to the barely open and hanged down flowers of the species what makes the plant looking “ashamed”.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF59FC04FDE8ACC2" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
Terrestrial plant
<quantity id="4CA0C3FEFFFDFFD5FE91FC04FE6FAD12" box="[373,497,1018,1043]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.25" metricValueMin="0.8" pageId="3" pageNumber="276" unit="cm" value="102.5" valueMax="125.0" valueMin="80.0">80125 cm</quantity>
tall, growing in clumps of few stems which are erect, covered by sheaths in basal two thirds and leafy above. Leaves 35 on each stem, 1016 ×
<quantity id="4CA0C3FEFFFDFFD5FCC8FBE0FC1DAD36" box="[812,899,1054,1079]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="3" pageNumber="276" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
, obovate, long acuminate, tridentate at the apex, leathery, glabrous, plicate. Inflorescence subtended by foliaceous bracts, covered with two narrowly, dark-brown, triangular, acute bracts, producing single flower at a time. Ovary
<quantity id="4CA0C3FEFFFDFFD5FC94FB98FC74AD7E" box="[880,1002,1126,1151]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="276" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">5060 mm</quantity>
long, white basally, becoming greenish towards the flower, strongly recurvate in the apical half what makes the flower directed downwards. Sepals apically retrose outwards, basally connate for about
<quantity id="4CA0C3FEFFFDFFD5FD94FB50FD5FADC6" box="[624,705,1198,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="276" unit="mm" value="10.0">10 mm</quantity>
, distinctly thicker than the petals and lip, light green, darker on the outer surface than on the inner one. Dorsal sepal 75 ×
<quantity id="4CA0C3FEFFFDFFD5FD0FFB2CFCA3ADEA" box="[747,829,1234,1259]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="276" unit="mm" value="18.0">18 mm</quantity>
, oblanceolate, acute, mucronate. Lateral sepals 75 ×
<quantity id="4CA0C3FEFFFDFFD5FA74FB2CFF28AC0E" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="276" unit="mm" value="18.0">18 mm</quantity>
, oblanceolate, acute, mucronate. Petals 68 ×
<quantity id="4CA0C3FEFFFDFFD5FD5AFB08FC8FAC0E" box="[702,785,1270,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="276" unit="mm" value="24.0">24 mm</quantity>
, widely oblanceolate, rounded, with somewhat curled and irregular margins in the apical half, white. Lip 71 ×
<quantity id="4CA0C3FEFFFDFFD5FD2EFAE4FC85AC32" box="[714,795,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="276" unit="mm" value="50.0">50 mm</quantity>
, obovate when spread, bilobed apically, basal half forming a tube in natural position, margins of apical half spread and strongly curled and irregular, apically bilobed, white outside, the inner surface bright yellow except the inconspicuous thickenings running along the central nerves and the spreading apical part of the lip. Lip margins entire or irregular. Column
<quantity id="4CA0C3FEFFFDFFD5FC5DFA78FB97AC9E" box="[953,1033,1414,1439]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="276" unit="mm" value="43.0">43 mm</quantity>
tall, club-shaped, with oblong, acute lateral stelidia not exceeding the anther cup.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF59FA30FCFCACE6" blockId="3.[136,1452,946,2051]" box="[189,866,1486,1511]" pageId="3" pageNumber="276">
<emphasis id="B92CB209FFFDFFD5FF59FA30FE46ACE6" bold="true" box="[189,472,1486,1511]" pageId="3" pageNumber="276">General distribution:—</emphasis>
Known only from the
<typeStatus id="54E3D0B9FFFDFFD5FD35FA30FC9FACE6" box="[721,769,1486,1511]" pageId="3" pageNumber="276">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF59FA0CFF4FAF76" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
<emphasis id="B92CB209FFFDFFD5FF59FA0CFEB8AF0A" bold="true" box="[189,294,1522,1547]" pageId="3" pageNumber="276">Notes:—</emphasis>
The phylogenetic trees obtained placed the newly described species among other representatives of
<taxonomicName id="4C581598FFFDFFD5FF6CF9E8FE31AF2E" authority="section Sobralia" box="[136,431,1558,1583]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="section" section="Sobralia">
<emphasis id="B92CB209FFFDFFD5FF6CF9E8FF76AF2E" box="[136,232,1558,1583]" italics="true" pageId="3" pageNumber="276">Sobralia</emphasis>
section
<emphasis id="B92CB209FFFDFFD5FEABF9E8FE31AF2E" box="[335,431,1558,1583]" italics="true" pageId="3" pageNumber="276">Sobralia</emphasis>
</taxonomicName>
. The structure of the inflorescence which is the basis of the infrageneric classification proposed by
<bibRefCitation id="EFC913EAFFFDFFD5FEC5F9C4FE54AF52" author="Brieger, F. G." box="[289,458,1594,1619]" pageId="3" pageNumber="276" pagination="780 - 800" refId="ref4310" refString="Brieger, F. G. (1983) Subtribus Sobraliinae. In: Brieger, F. G., Maatsch, R. &amp; Senghas, K. (Eds.) R. Schlechter, Die Orchideen, I, 13. Lieferung, Verlag Paul Parey, Berlin &amp; Hamburg, pp. 780 - 800." type="book chapter" year="1983">Brieger (1983)</bibRefCitation>
confirms the results. Thus, we treat the new species into the nominal section of the genus.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF59F97CFC43AE06" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
According to the phylogenetic results, the new species is related to
<taxonomicName id="4C581598FFFDFFD5FC7DF97CFA9CAF9A" authority="Archila (1999: 14)" authorityName="Archila" authorityPageNumber="14" authorityYear="1999" box="[921,1282,1666,1691]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="cobanensis">
<emphasis id="B92CB209FFFDFFD5FC7DF97CFBB1AF9A" box="[921,1071,1666,1691]" italics="true" pageId="3" pageNumber="276">S. cobanensis</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FBD7F97CFA9CAF9A" author="Archila, F." box="[1075,1282,1666,1691]" pageId="3" pageNumber="276" pagination="10 - 30" refId="ref4070" refString="Archila, F. (1999) Exploraciones botanicas en Guatemala (Parte 1). Revista Guatemalensis 2: 10 - 30." type="journal article" year="1999">Archila (1999: 14)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FAE8F97CFEC7AFBE" authority="Dodson (1998: 22)" authorityName="Dodson" authorityPageNumber="22" authorityYear="1998" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="ecuadorana">
<emphasis id="B92CB209FFFDFFD5FAE8F97CFA32AF9A" box="[1292,1452,1666,1691]" italics="true" pageId="3" pageNumber="276">S. ecuadorana</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FF6CF958FEC7AFBE" author="Dodson, C. H." box="[136,345,1702,1727]" pageId="3" pageNumber="276" pagination="3 - 67" refId="ref4500" refString="Dodson, C. H. (1998) New orchid species and combinations from Ecuador. Orquideologia 21: 3 - 67." type="journal article" year="1998">Dodson (1998: 22)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FE8FF958FCC6AFBE" authority="Poeppig &amp; Endlicher (1836: 54)" authorityName="Poeppig &amp; Endlicher" authorityPageNumber="54" authorityYear="1836" box="[363,856,1702,1727]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="fimbriata">
<emphasis id="B92CB209FFFDFFD5FE8FF958FE73AFBE" box="[363,493,1702,1727]" italics="true" pageId="3" pageNumber="276">S. fimbriata</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FE17F958FCC6AFBE" author="Poeppig E. F. &amp; Endlicher S. F. L." box="[499,856,1702,1727]" pageId="3" pageNumber="276" pagination="1 - 62" refId="ref5272" refString="Poeppig E. F. &amp; Endlicher S. F. L. (1836) Nova Genera ac Species Plantarum 1: 1 - 62." type="book chapter" year="1836">Poeppig &amp; Endlicher (1836: 54)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FC87F958FB53AFBE" authority="Schlechter (1922: 11)" authorityName="Schlechter" authorityPageNumber="11" authorityYear="1922" box="[867,1229,1702,1727]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="powellii">
<emphasis id="B92CB209FFFDFFD5FC87F958FC49AFBE" box="[867,983,1702,1727]" italics="true" pageId="3" pageNumber="276">S. powellii</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FC39F958FB53AFBE" author="Schlechter, F. R. R." box="[989,1229,1702,1727]" pageId="3" pageNumber="276" pagination="4 - 95" refId="ref5574" refString="Schlechter, F. R. R. (1922) Orchidaceae Powellianae Panamensis. Repertorium Specierum Novarum Regni Vegetabilis, Beihefte 17: 4 - 95" type="journal article" year="1922">Schlechter (1922: 11)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FB3CF958FE92AFE2" authority="Dressler (2005: 941)" authorityName="Dressler" authorityPageNumber="941" authorityYear="2005" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="tricolor">
<emphasis id="B92CB209FFFDFFD5FB3CF958FAD7AFBE" box="[1240,1353,1702,1727]" italics="true" pageId="3" pageNumber="276">S. tricolor</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FAAAF958FE92AFE2" author="Dressler, R. L." pageId="3" pageNumber="276" pagination="937 - 941" refId="ref4647" refString="Dressler, R. L. (2005) Six new species of Sobralia from Panama. Orchids, The Magazine of the American Orchid Society, West Palm Beach, Florida 74: 937 - 941." type="journal article" year="2005">Dressler (2005: 941)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FEF3F934FD7AAFE2" authority="Reichenbach (1852: 714)" authorityName="Reichenbach" authorityPageNumber="714" authorityYear="1852" box="[279,740,1738,1763]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="warscewiczii">
<emphasis id="B92CB209FFFDFFD5FEF3F934FE5DAFE2" box="[279,451,1738,1763]" italics="true" pageId="3" pageNumber="276">S. warscewiczii</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FE2CF934FD7AAFE2" author="Reichenbach, H. G." box="[456,740,1738,1763]" pageId="3" pageNumber="276" pagination="705 - 715" refId="ref5318" refString="Reichenbach, H. G. (1852) Neue Orchideen der Expedition des Herrn J. de Warszewicz. Botanische Zeitung (Berlin) 10: 705 - 715." type="journal article" year="1852">Reichenbach (1852: 714)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C581598FFFDFFD5FD0AF934FB6DAFE2" authority="Bennett &amp; Christenson (2001: 764)" authorityName="Bennett &amp; Christenson" authorityPageNumber="764" authorityYear="2001" box="[750,1267,1738,1763]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="withneri">
<emphasis id="B92CB209FFFDFFD5FD0AF934FCF8AFE2" box="[750,870,1738,1763]" italics="true" pageId="3" pageNumber="276">S. withneri</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FC88F934FB6DAFE2" author="Bennett, D. E. &amp; Christenson, E. A." box="[876,1267,1738,1763]" pageId="3" pageNumber="276" pagination="601 - 800" refId="ref4270" refString="Bennett, D. E. &amp; Christenson, E. A. (2001) Icones Orchidacearum Peruvianum. Part 4. A. Pastorelli de Bennett ed. Sarasota, USA, pl. 601 - 800." type="book chapter" year="2001">Bennett &amp; Christenson (2001: 764)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C581598FFFDFFD5FAC2F934FE46AE06" authority="Peeters &amp; Cogniaux (1899: 3)" authorityName="Peeters &amp; Cogniaux" authorityPageNumber="3" authorityYear="1899" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="virginalis">
<emphasis id="B92CB209FFFDFFD5FAC2F934FA35AFE2" box="[1318,1451,1738,1763]" italics="true" pageId="3" pageNumber="276">S. virginalis</emphasis>
<bibRefCitation id="EFC913EAFFFDFFD5FF6CF910FE46AE06" author="Peeters, F. &amp; Cogniaux, C. A." box="[136,472,1774,1799]" pageId="3" pageNumber="276" refId="ref5213" refString="Peeters, F. &amp; Cogniaux, C. A. (1899) Sobralia. Dictionnaire Iconographique des Orchidees, Bruxelles (Brussels) 9: pl. 3." type="journal volume" year="1899">Peeters &amp; Cogniaux (1899: 3)</bibRefCitation>
</taxonomicName>
but none of the taxa resemble
<taxonomicName id="4C581598FFFDFFD5FCD4F910FC48AE06" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[816,982,1774,1799]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFDFFD5FCD4F910FC48AE06" box="[816,982,1774,1799]" italics="true" pageId="3" pageNumber="276">Sobralia fugax</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF59F8ECFB9AAE72" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
<taxonomicName id="4C581598FFFDFFD5FF59F8ECFE3EAE2A" authorityName="Archila" authorityYear="1999" box="[189,416,1810,1835]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="cobanensis">
<emphasis id="B92CB209FFFDFFD5FF59F8ECFE3EAE2A" box="[189,416,1810,1835]" italics="true" pageId="3" pageNumber="276">Sobralia cobanensis</emphasis>
</taxonomicName>
differs in larger leaves (20 ×
<quantity id="4CA0C3FEFFFDFFD5FD00F8ECFC87AE2A" box="[740,793,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="276" unit="cm" value="9.0">9 cm</quantity>
versus 1016 ×
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), wider (
<quantity id="4CA0C3FEFFFDFFD5FB60F8ECFB4AAE2A" box="[1156,1236,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="276" unit="mm" value="30.0">30 mm</quantity>
versus
<quantity id="4CA0C3FEFFFDFFD5FAC2F8ECFAEBAE2A" box="[1318,1397,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="276" unit="mm" value="24.0">24 mm</quantity>
) and distinctly stronger crenate petals which are always recurved backwards and distinctly shorter column (
<quantity id="4CA0C3FEFFFDFFD5FAE8F8C8FAC3AE4E" box="[1292,1373,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="276" unit="mm" value="30.0">30 mm</quantity>
versus
<quantity id="4CA0C3FEFFFDFFD5FF6CF8A4FF45AE72" box="[136,219,1882,1907]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="276" unit="mm" value="43.0">43 mm</quantity>
). Apart of that, the flowers of
<taxonomicName id="4C581598FFFDFFD5FDC8F8A4FC91AE72" authorityName="Archila" authorityYear="1999" box="[556,783,1882,1907]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="cobanensis">
<emphasis id="B92CB209FFFDFFD5FDC8F8A4FC91AE72" box="[556,783,1882,1907]" italics="true" pageId="3" pageNumber="276">Sobralia cobanensis</emphasis>
</taxonomicName>
are purple and white.
</paragraph>
<paragraph id="8BE76E1BFFFDFFD5FF22F880FC6DA102" blockId="3.[136,1452,946,2051]" pageId="3" pageNumber="276">
The characteristic features of
<taxonomicName id="4C581598FFFDFFD5FDFEF880FC8BAE96" box="[538,789,1918,1943]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="ecuadoreana">
<emphasis id="B92CB209FFFDFFD5FDFEF880FC8BAE96" box="[538,789,1918,1943]" italics="true" pageId="3" pageNumber="276">Sobralia ecuadoreana</emphasis>
</taxonomicName>
are the thin, leathery leaves, the short floral bracts barely exceeding the sheath subtending the leaf-like bract, the long fleshy lamellae on the basal part of lip reaching third of its length and the distinctly shorter column than the one observed in new species (
<quantity id="4CA0C3FEFFFDFFD5FBF7F838FBFDAEDE" box="[1043,1123,1990,2015]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="276" unit="mm" value="30.0">30 mm</quantity>
versus
<quantity id="4CA0C3FEFFFDFFD5FB53F838FA97AEDE" box="[1207,1289,1990,2015]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="276" unit="mm" value="43.0">43 mm</quantity>
). While the lip color of
<taxonomicName id="4C581598FFFDFFD5FF03F814FE17A102" authorityName="Dodson" authorityYear="1998" box="[231,393,2026,2051]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="ecuadorana">
<emphasis id="B92CB209FFFDFFD5FF03F814FE17A102" box="[231,393,2026,2051]" italics="true" pageId="3" pageNumber="276">S. ecuadorana</emphasis>
</taxonomicName>
is similar with
<taxonomicName id="4C581598FFFDFFD5FDD3F814FD0FA102" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[567,657,2026,2051]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFDFFD5FDD3F814FD0FA102" box="[567,657,2026,2051]" italics="true" pageId="3" pageNumber="276">S. fugax</emphasis>
</taxonomicName>
, its sepals are white or pinkish.
</paragraph>
<caption id="DF273E93FFFAFFD2FF6CF884FCC1AEB2" ID-DOI="http://doi.org/10.5281/zenodo.13727340" ID-Zenodo-Dep="13727340" httpUri="https://zenodo.org/record/13727340/files/figure.png" pageId="4" pageNumber="277" startId="4.[136,229,1914,1935]" targetBox="[176,1407,224,1859]" targetPageId="4" targetType="figure">
<paragraph id="8BE76E1BFFFAFFD2FF6CF884FCC1AEB2" blockId="4.[136,1452,1913,1971]" pageId="4" pageNumber="277">
<emphasis id="B92CB209FFFAFFD2FF6CF884FDC7AE8E" bold="true" box="[136,601,1913,1935]" pageId="4" pageNumber="277">
FIGURE 1.
<taxonomicName id="4C581598FFFAFFD2FEE6F887FDC7AE8E" authority="Baranow &amp; Dudek" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[258,601,1913,1935]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="277" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFAFFD2FEE6F887FE0AAE8E" bold="true" box="[258,404,1913,1935]" italics="true" pageId="4" pageNumber="277">Sobralia fugax</emphasis>
Baranow &amp; Dudek
</taxonomicName>
</emphasis>
: A. Habit. B. Apical part of stem with the inflorescence. C. Dorsal sepal. D. Lateral sepal. E. Petal. F. Lip. Natural position. G. Lip, spread (drawn from the holotype).
</paragraph>
</caption>
<caption id="DF273E93FFFBFFD3FF6CFB9AFD68AD9C" ID-DOI="http://doi.org/10.5281/zenodo.13727342" ID-Zenodo-Dep="13727342" httpUri="https://zenodo.org/record/13727342/files/figure.png" pageId="5" pageNumber="278" startId="5.[136,229,1124,1145]" targetBox="[151,1436,190,1099]" targetPageId="5" targetType="figure">
<paragraph id="8BE76E1BFFFBFFD3FF6CFB9AFD68AD9C" blockId="5.[136,1452,1123,1181]" pageId="5" pageNumber="278">
<emphasis id="B92CB209FFFBFFD3FF6CFB9AFDF4AD78" bold="true" box="[136,618,1123,1145]" pageId="5" pageNumber="278">
FIGURE 2.
<taxonomicName id="4C581598FFFBFFD3FEE3FB9DFDF4AD78" authority="Baranow &amp; Dudek" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[263,618,1123,1145]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFBFFD3FEE3FB9DFE02AD78" bold="true" box="[263,412,1123,1145]" italics="true" pageId="5" pageNumber="278">Sobralia fugax</emphasis>
Baranow &amp; Dudek
</taxonomicName>
</emphasis>
: A. Flower, side view. B. Dorsal sepal. C. Lateral sepal. D. Petal. E. Lip. F. Interior surface of lip. G. Column (photographs taken from the holotype)
</paragraph>
</caption>
<paragraph id="8BE76E1BFFFBFFD3FF59FB33FAFCAC0B" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
The distinctness among the new species and
<taxonomicName id="4C581598FFFBFFD3FD4FFB33FCE7ADE7" authorityName="Poeppig &amp; Endlicher" authorityYear="1836" box="[683,889,1229,1254]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="fimbriata">
<emphasis id="B92CB209FFFBFFD3FD4FFB33FCE7ADE7" box="[683,889,1229,1254]" italics="true" pageId="5" pageNumber="278">Sobralia fimbriata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C581598FFFBFFD3FC62FB33FC69ADE7" authorityName="Dressler" authorityYear="2005" box="[902,1015,1229,1254]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="tricolor">
<emphasis id="B92CB209FFFBFFD3FC62FB33FC69ADE7" box="[902,1015,1229,1254]" italics="true" pageId="5" pageNumber="278">S. tricolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C581598FFFBFFD3FBCFFB33FB3DADE7" authorityName="Bennett &amp; Christenson" authorityYear="2001" box="[1067,1187,1229,1254]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="withneri">
<emphasis id="B92CB209FFFBFFD3FBCFFB33FB3DADE7" box="[1067,1187,1229,1254]" italics="true" pageId="5" pageNumber="278">S. withneri</emphasis>
</taxonomicName>
is evident from the first glance. All the three species have lips furnished with fimbriate lamellae and the margin of the lip is fimbriated.
</paragraph>
<paragraph id="8BE76E1BFFFBFFD3FF59FAEBFD64AC77" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
<taxonomicName id="4C581598FFFBFFD3FF59FAEBFE1FAC2F" authorityName="Schlechter" authorityYear="1922" box="[189,385,1301,1326]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="powellii">
<emphasis id="B92CB209FFFBFFD3FF59FAEBFE1FAC2F" box="[189,385,1301,1326]" italics="true" pageId="5" pageNumber="278">Sobralia powellii</emphasis>
</taxonomicName>
also can be distinguished from the new entity by the larger leaves (up to 22 ×
<quantity id="4CA0C3FEFFFBFFD3FAF3FAEBFACBAC2F" box="[1303,1365,1301,1326]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="278" unit="cm" value="6.0">6 cm</quantity>
), larger flowers (floral segments up to
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long) and wider petals (ca.
<quantity id="4CA0C3FEFFFBFFD3FC89FAC7FC5EAC53" box="[877,960,1337,1362]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="278" unit="mm" value="30.0">30 mm</quantity>
). The species also differs on the color of its flowers which are white with yellow lip throat and disc.
</paragraph>
<paragraph id="8BE76E1BFFFBFFD3FF59FA7FFB50ACBF" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
<taxonomicName id="4C581598FFFBFFD3FF59FA7FFE2DAC9B" box="[189,435,1409,1434]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="warszewiczii">
<emphasis id="B92CB209FFFBFFD3FF59FA7FFE2DAC9B" box="[189,435,1409,1434]" italics="true" pageId="5" pageNumber="278">Sobralia warszewiczii</emphasis>
</taxonomicName>
has purple flowers with deeply erose margins of apical half of lip. According to the original drawings of the
<typeStatus id="54E3D0B9FFFBFFD3FED9FA5BFEF3ACBF" box="[317,365,1445,1470]" pageId="5" pageNumber="278">type</typeStatus>
material, its lip has no protuberances on its surface except the basal papillae.
</paragraph>
<paragraph id="8BE76E1BFFFBFFD3FF59FA37FBA3AF07" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
<taxonomicName id="4C581598FFFBFFD3FF59FA37FE0EACE3" authorityName="Peeters &amp; Cogniaux" authorityYear="1899" box="[189,400,1481,1506]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="virginalis">
<emphasis id="B92CB209FFFBFFD3FF59FA37FE0EACE3" box="[189,400,1481,1506]" italics="true" pageId="5" pageNumber="278">Sobralia virginalis</emphasis>
</taxonomicName>
is a species with white flowers with orange base and disc of lip ornamented with white lines running along the nerves. Additionally, its lip margin is deeply dentate in apical part.
</paragraph>
<paragraph id="8BE76E1BFFFBFFD3FF59F9EFFE26AF97" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
Taking into account that not all the species are included in the molecular study, it is possible that the plant described as a new species represents one of the already known taxa. However, the comparison of its morphology with all other Colombian species of the section based on our recent study (
<bibRefCitation id="EFC913EAFFFBFFD3FC6AF9A7FBEBAF73" author="Baranow, P. &amp; Szlachetko, D. L. &amp; Dudek, M." box="[910,1141,1625,1650]" pageId="5" pageNumber="278" pagination="1 - 8" refId="ref4094" refString="Baranow, P., Szlachetko, D. L. &amp; Dudek, M. (2014) New species of Sobralia section Abbreviatae Brieger (Orchidaceae) from Colombia. A morphological and molecular evidence. Plant Systematics and Evolution 300: 1 - 8. https: // doi. org / 10.1007 / s 00606 - 014 - 0991 - 2" type="journal article" year="2014">
Baranow
<emphasis id="B92CB209FFFBFFD3FC1FF9A7FBADAF73" box="[1019,1075,1625,1650]" italics="true" pageId="5" pageNumber="278">et al.</emphasis>
2014
</bibRefCitation>
,
<bibRefCitation id="EFC913EAFFFBFFD3FB9BF9A7FABBAF73" author="Baranow, P." box="[1151,1317,1625,1650]" pageId="5" pageNumber="278" pagination="41 - 60" refId="ref4159" refString="Baranow, P. (2015) Taxonomic notes on Sobralia section Abbreviatae (Orchidaceae) in Colombia, with description of a new species. Plant Systematics and Evolution 301: 41 - 60. https: // doi. org / 10.1007 / s 00606 - 014 - 1053 - 5" type="journal article" year="2015">Baranow 2015</bibRefCitation>
) allowed us to exclude such possibility.
</paragraph>
<paragraph id="8BE76E1BFFFBFFD3FF59F95FFD59AE4B" blockId="5.[136,1452,1229,1866]" pageId="5" pageNumber="278">
<taxonomicName id="4C581598FFFBFFD3FF59F95FFEFDAFBB" authorityName="Baranow &amp; Dudek" authorityYear="2018" box="[189,355,1697,1722]" class="Liliopsida" family="Orchidaceae" genus="Sobralia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="278" phylum="Tracheophyta" rank="species" species="fugax">
<emphasis id="B92CB209FFFBFFD3FF59F95FFEFDAFBB" box="[189,355,1697,1722]" italics="true" pageId="5" pageNumber="278">Sobralia fugax</emphasis>
</taxonomicName>
is unique by the greenish, fleshy sepals and strongly bent ovary which determines the orientation of the flower. It is hanged down in contrast to other species of the group which produce flowers directed forward. It is also worthy to mention, that sepals and petals of the new species are not bent backwards what makes its flowers distinctly more slender. The mentioned observations were made on the basis of three flowering periods thus we can assume that these are constant features of the plant.
</paragraph>
</subSubSection>
</treatment>
</document>