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<document id="387B7722978B89628F578A3ED83F31FC" ID-DOI="10.11646/phytotaxa.383.3.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13725779" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725674548172" checkinUser="felipe" docAuthor="Rodda, Michele &amp; Rahayu, Sri" docDate="2018" docId="03F187F4FFD7FFEAFF0438D4FF14FC1B" docLanguage="en" docName="phytotaxa.383.3.2.pdf" docOrigin="Phytotaxa 383 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.383.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Hoya uncinata Teijsm. &amp; Binn., Tijdschr. Nederl. Ind." docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="FFC8FF8CFFD6FFEEFF8C3A79FFDEFFCA" masterDocTitle="A revision of the Hoya uncinata complex (Apocynaceae, Asclepiadoideae), with description of a new species" masterLastPageNumber="258" masterPageNumber="252" pageNumber="253" updateTime="1725686874424" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1139B147DAEEE754F43ED9B59AA70148">A revision of the Hoya uncinata complex (Apocynaceae, Asclepiadoideae), with description of a new species</mods:title>
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<mods:namePart id="63820488996049BCDE3C99952C74E076">Rodda, Michele</mods:namePart>
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<mods:namePart id="7C2503A58EC7BAD6F42117E96B1E3B0C">Rahayu, Sri</mods:namePart>
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<heading id="D0AF818EFFD7FFEFFF0438D4FC2AFD0D" box="[136,1012,685,711]" fontSize="11" level="2" pageId="1" pageNumber="253" reason="3">
<treatmentCitation id="0AF910F3FFD7FFEFFF0438D4FC47FD0D" author="Teijsm. &amp; Binn." box="[136,921,685,711]" journal="Tijdschr. Nederl. Ind." page="408" pageId="1" pageNumber="253" score="0.1312722213955852" volume="25" year="1863">
<taxonomicName id="4C584D61FFD7FFEFFF0438D4FD32FD0D" ID-CoL="3MS5B" authority="Teijsm. &amp; Binn., Tijdschr. Nederl. Ind." box="[136,748,685,711]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD7FFEFFF0438D4FEEAFD0D" bold="true" box="[136,308,685,711]" italics="true" pageId="1" pageNumber="253">Hoya uncinata</emphasis>
Teijsm. &amp; Binn., Tijdschr. Nederl. Ind.
</taxonomicName>
25: 408 (1863)
</treatmentCitation>
(
<figureCitation id="13632A67FFD7FFEFFC2438D7FC36FD0D" box="[936,1000,686,711]" captionStart="FIGURE 1" captionStartId="1.[136,229,1428,1449]" captionTargetBox="[151,1436,949,1400]" captionTargetId="figure-218@1.[151,1437,949,1400]" captionTargetPageId="1" captionText="FIGURE 1. Comparison between the species in the Hoya uncinata complex. A, Hoya corneri. Based on M. Rodda MR647 (SING); B, Hoya uncinata, based on Rodda M MR609 (SING); C. Hoya ruthiae, based on M. Rodda MR606 (SING). Photos by M. Rodda." figureDoi="http://doi.org/10.5281/zenodo.13725781" httpUri="https://zenodo.org/record/13725781/files/figure.png" pageId="1" pageNumber="253">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426569FFD7FFEFFF0438ACFC09FCC4" pageId="1" pageNumber="253" type="materials_examined">
<paragraph id="8BE736E2FFD7FFEFFF0438ACFC09FCC4" blockId="1.[136,1451,725,782]" pageId="1" pageNumber="253">
<emphasis id="B92CEAF0FFD7FFEFFF0438ACFF64FD20" bold="true" box="[136,186,725,746]" pageId="1" pageNumber="253">Type</emphasis>
:—
<materialsCitation id="3B303CBFFFD7FFEFFF5B38ACFB8CFD20" box="[215,1106,725,746]" country="Indonesia" location="Java" pageId="1" pageNumber="253" specimenCount="1" stateProvince="Jawa Timur">
<collectingCountry id="F34F7672FFD7FFEFFF5B38ACFEE7FD20" box="[215,313,725,746]" name="Indonesia" pageId="1" pageNumber="253">Indonesia</collectingCountry>
,
<collectingRegion id="499CF800FFD7FFEFFECF38ACFEB0FD20" box="[323,366,725,746]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="253">Java</collectingRegion>
and
<collectingRegion id="499CF800FFD7FFEFFE1138ACFE33FD20" box="[413,493,725,746]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="253">Sumatra</collectingRegion>
“In silvis Javae et Sumatrae ins. prov. Palembang (not traced)
</materialsCitation>
;
<materialsCitation id="3B303CBFFFD7FFEFFBD038ACFC0AFCC4" collectingDate="1907-01-20" collectionCode="R, B" country="Indonesia" elevation="900" location="Java" pageId="1" pageNumber="253" specimenCode="B100277224" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">
<emphasis id="B92CEAF0FFD7FFEFFBD038ACFB57FD20" box="[1116,1161,725,746]" italics="true" pageId="1" pageNumber="253">ibid.</emphasis>
, auf Bäumen bei Padang,
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<elevation id="0075D1D1FFD7FFEFFA0B38ACFF0CFCC4" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="253" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<date id="FFE61022FFD7FFEFFF503880FE86FCC4" box="[220,344,761,782]" pageId="1" pageNumber="253" value="1907-01-20">
<collectingDate id="EFA2E9CAFFD7FFEFFF503880FE86FCC4" box="[220,344,761,782]" pageId="1" pageNumber="253" value="1907-01-20">20 Jan. 1907</collectingDate>
</date>
,
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<collectionCode id="ED49AE27FFD7FFEFFEEE3880FEADFCC4" box="[354,371,761,782]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="253">R</collectionCode>
. Schlechter 15916
</emphasis>
(
<typeStatus id="54E38840FFD7FFEFFDBD3880FD5FFCC4" box="[561,641,761,782]" pageId="1" pageNumber="253" type="neotype">neotype</typeStatus>
,
<collectionCode id="ED49AE27FFD7FFEFFD063880FD44FCC4" box="[650,666,761,782]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="253" type="Herbarium">B</collectionCode>
[
<specimenCode id="DBFE9E99FFD7FFEFFD293880FCF8FCC4" box="[677,806,761,782]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="253" type="Herbarium">B100277224</specimenCode>
] here designated)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426569FFD7FFEFFF043947FC19FC9D" box="[136,967,830,855]" pageId="1" pageNumber="253" type="reference_group">
<paragraph id="8BE736E2FFD7FFEFFF043947FC19FC9D" blockId="1.[136,1451,830,926]" box="[136,967,830,855]" pageId="1" pageNumber="253">
<treatmentCitation id="0AF910F3FFD7FFEFFF043947FC80FC9D" author="" box="[136,862,830,855]" issue="2" journal="Bot. Centralbl." page="15" pageId="1" pageNumber="253" score="0.001166684644447956" volume="34" year="1916">
<taxonomicName id="4C584D61FFD7FFEFFF043947FD40FC9D" ID-CoL="3MRXP" authority="Schltr., Beih. Bot. Centralbl." box="[136,670,830,855]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="padangensis">
<emphasis id="B92CEAF0FFD7FFEFFF043947FE86FC9D" box="[136,344,830,855]" italics="true" pageId="1" pageNumber="253">Hoya padangensis</emphasis>
Schltr., Beih. Bot. Centralbl.
</taxonomicName>
34(2): 15 (1916)
</treatmentCitation>
<emphasis id="B92CEAF0FFD7FFEFFCE93947FC1EFC9D" box="[869,960,830,855]" italics="true" pageId="1" pageNumber="253">syn. nov</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426569FFD7FFEAFF04391CFE5EFD8F" lastPageId="4" lastPageNumber="256" pageId="1" pageNumber="253" type="materials_examined">
<paragraph id="8BE736E2FFD7FFEFFF04391CFEEDFC54" blockId="1.[136,1451,830,926]" pageId="1" pageNumber="253">
<emphasis id="B92CEAF0FFD7FFEFFF04391CFF64FCB0" bold="true" box="[136,186,869,890]" pageId="1" pageNumber="253">Type</emphasis>
:—
<collectingCountry id="F34F7672FFD7FFEFFF5B391CFEE7FCB0" box="[215,313,869,890]" name="Indonesia" pageId="1" pageNumber="253">Indonesia</collectingCountry>
,
<collectingRegion id="499CF800FFD7FFEFFECB391CFE47FCB0" box="[327,409,869,890]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="253">Sumatra</collectingRegion>
, auf Bäumen bei Padang,
<quantity id="4CA09B07FFD7FFEFFD25391CFD32FCB0" box="[681,748,869,890]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="253" unit="m" value="900.0">900 m</quantity>
,
<date id="FFE61022FFD7FFEFFD76391CFCA0FCB0" box="[762,894,869,890]" pageId="1" pageNumber="253" value="1907-01-20">20 Jan. 1907</date>
,
<emphasis id="B92CEAF0FFD7FFEFFC01391CFB89FCB0" box="[909,1111,869,890]" italics="true" pageId="1" pageNumber="253">
<collectionCode id="ED49AE27FFD7FFEFFC01391CFC40FCB0" box="[909,926,869,890]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="253">R</collectionCode>
. Schlechter 15916
</emphasis>
(
<typeStatus id="54E38840FFD7FFEFFBE4391CFB1AFCB0" box="[1128,1220,869,890]" pageId="1" pageNumber="253" type="lectotype">lectotype</typeStatus>
,
<collectionCode id="ED49AE27FFD7FFEFFB5E391CFB3CFCB0" box="[1234,1250,869,890]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="253" type="Herbarium">B</collectionCode>
[
<collectionCode id="ED49AE27FFD7FFEFFB7D391CFADEFCB0" box="[1265,1280,869,890]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="253" type="Herbarium">B</collectionCode>
100277224] here designated).
</paragraph>
<caption id="DF27666AFFD7FFEFFF043FEDFA8EFA07" ID-DOI="http://doi.org/10.5281/zenodo.13725781" ID-Zenodo-Dep="13725781" httpUri="https://zenodo.org/record/13725781/files/figure.png" pageId="1" pageNumber="253" startId="1.[136,229,1428,1449]" targetBox="[151,1436,949,1400]" targetPageId="1" targetType="figure">
<paragraph id="8BE736E2FFD7FFEFFF043FEDFA8EFA07" blockId="1.[136,1451,1428,1485]" pageId="1" pageNumber="253">
<emphasis id="B92CEAF0FFD7FFEFFF043FEDFF21FA63" bold="true" box="[136,255,1428,1449]" pageId="1" pageNumber="253">FIGURE 1.</emphasis>
Comparison between the species in the
<taxonomicName id="4C584D61FFD7FFEFFD043FEDFCC9FA63" box="[648,791,1428,1449]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD7FFEFFD043FEDFCC9FA63" box="[648,791,1428,1449]" italics="true" pageId="1" pageNumber="253">Hoya uncinata</emphasis>
</taxonomicName>
complex. A,
<taxonomicName id="4C584D61FFD7FFEFFC173FEDFBC3FA63" box="[923,1053,1428,1449]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="corneri">
<emphasis id="B92CEAF0FFD7FFEFFC173FEDFBC3FA63" box="[923,1053,1428,1449]" italics="true" pageId="1" pageNumber="253">Hoya corneri</emphasis>
</taxonomicName>
. Based on
<emphasis id="B92CEAF0FFD7FFEFFB073FEDFAE6FA63" box="[1163,1336,1428,1449]" italics="true" pageId="1" pageNumber="253">M. Rodda MR647</emphasis>
(SING); B,
<taxonomicName id="4C584D61FFD7FFEFFF043FC1FEC8FA07" box="[136,278,1464,1485]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD7FFEFFF043FC1FEC8FA07" box="[136,278,1464,1485]" italics="true" pageId="1" pageNumber="253">Hoya uncinata</emphasis>
</taxonomicName>
, based on
<emphasis id="B92CEAF0FFD7FFEFFEF13FC1FDFCFA07" box="[381,546,1464,1485]" italics="true" pageId="1" pageNumber="253">Rodda M MR609</emphasis>
(SING); C.
<taxonomicName id="4C584D61FFD7FFEFFD153FC1FCC6FA07" authorityName="Rodda" authorityYear="2015" box="[665,792,1464,1485]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="253" phylum="Tracheophyta" rank="species" species="ruthiae">
<emphasis id="B92CEAF0FFD7FFEFFD153FC1FCC6FA07" box="[665,792,1464,1485]" italics="true" pageId="1" pageNumber="253">Hoya ruthiae</emphasis>
</taxonomicName>
, based on
<emphasis id="B92CEAF0FFD7FFEFFCF33FC1FBF4FA07" box="[895,1066,1464,1485]" italics="true" pageId="1" pageNumber="253">M. Rodda MR606</emphasis>
(SING). Photos by M. Rodda.
</paragraph>
</caption>
<paragraph id="8BE736E2FFD7FFECFF043F84FAEEFEA7" blockId="1.[136,1452,1533,2062]" lastBlockId="2.[136,1452,160,365]" lastPageId="2" lastPageNumber="254" pageId="1" pageNumber="253">
Delicate epiphytic climber with clear latex in all vegetative parts; all vegetative parts glabrous. Leafy
<emphasis id="B92CEAF0FFD7FFEFFB653F84FAF9F9DC" box="[1257,1319,1533,1558]" italics="true" pageId="1" pageNumber="253">stems</emphasis>
cylindrical, slender, up to
<quantity id="4CA09B07FFD7FFEFFEAD3C58FEBEF9F0" box="[289,352,1569,1594]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="253" unit="mm" value="4.0">4 mm</quantity>
in diam., dark brown or grey, with membranaceous peeling bark;
<emphasis id="B92CEAF0FFD7FFEFFBBE3C58FB76F9F0" box="[1074,1192,1569,1594]" italics="true" pageId="1" pageNumber="253">internodes</emphasis>
<quantity id="4CA09B07FFD7FFEFFB223C58FAD3F9F0" box="[1198,1293,1569,1594]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="253" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">515 cm</quantity>
long.
<emphasis id="B92CEAF0FFD7FFEFFADD3C58FA75F9F0" box="[1361,1451,1569,1594]" italics="true" pageId="1" pageNumber="253">Petioles</emphasis>
terete, fleshy, 510 ×
<quantity id="4CA09B07FFD7FFEFFE0F3C3CFE3CF994" box="[387,482,1605,1630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="253" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
in diam.;
<emphasis id="B92CEAF0FFD7FFEFFDD53C3CFD79F994" box="[601,679,1605,1630]" italics="true" pageId="1" pageNumber="253">lamina</emphasis>
elliptic-lanceolate, fleshy, 710 ×
<quantity id="4CA09B07FFD7FFEFFBB93C3CFB7CF994" box="[1077,1186,1605,1630]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="253" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 cm</quantity>
, apex acute-acuminate, base rounded or cuneate, dark green above with numerous grey spots, lighter green underneath, venation pinnate, very faintly visible above, not visible beneath.
<emphasis id="B92CEAF0FFD7FFEFFDE33CF4FCCEF96C" box="[623,784,1677,1702]" italics="true" pageId="1" pageNumber="253">Inflorescences</emphasis>
sciadioidal, ca.
<quantity id="4CA09B07FFD7FFEFFC433CF4FBD7F96C" box="[975,1033,1677,1702]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="253" unit="cm" value="4.0">4 cm</quantity>
in diam., 822 flowered;
<emphasis id="B92CEAF0FFD7FFEFFAB73CF4FA75F96C" box="[1339,1451,1677,1702]" italics="true" pageId="1" pageNumber="253">peduncles</emphasis>
extra-axillary, negatively geotropic or horizontal, terete,
<quantity id="4CA09B07FFD7FFEFFC983CC8FCA1F900" box="[788,895,1713,1738]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="1" pageNumber="253" unit="cm" value="3.25" valueMax="5.0" valueMin="1.5">1.55 cm</quantity>
×
<quantity id="4CA09B07FFD7FFEFFC2E3CC8FBDCF900" box="[930,1026,1713,1738]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="253" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
in diam., glabrous;
<emphasis id="B92CEAF0FFD7FFEFFB623CC8FA94F900" box="[1262,1354,1713,1738]" italics="true" pageId="1" pageNumber="253">pedicels</emphasis>
upright,
<quantity id="4CA09B07FFD7FFEFFF043CACFEDFF924" box="[136,257,1749,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="253" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
×
<quantity id="4CA09B07FFD7FFEFFE923CACFE7BF924" box="[286,421,1749,1774]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="253" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
in diam., purplish-green, glabrous.
<taxonomicName id="4C584D61FFD7FFEFFCB93CACFCA8F924" box="[821,886,1749,1774]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="1" pageNumber="253" phylum="Porifera" rank="genus">
<emphasis id="B92CEAF0FFD7FFEFFCB93CACFCA8F924" box="[821,886,1749,1774]" italics="true" pageId="1" pageNumber="253">Calyx</emphasis>
</taxonomicName>
lobes ovate, slightly spatulate, red or pink 1.82.5 ×
<quantity id="4CA09B07FFD7FFEFFF113C80FEFBF8D8" box="[157,293,1785,1810]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="1" pageNumber="253" unit="mm" value="1.0" valueMax="1.3" valueMin="0.7">0.71.3 mm</quantity>
, apex obtuse or round, glabrous; basal
<emphasis id="B92CEAF0FFD7FFEFFD5C3C80FCF8F8D8" box="[720,806,1785,1810]" italics="true" pageId="1" pageNumber="253">colleter</emphasis>
<quantity id="4CA09B07FFD7FFEFFCA03C80FC8BF8D8" box="[812,853,1785,1810]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="253" unit="in" value="1.0">1 in</quantity>
each sepal sinus, triangular,
<quantity id="4CA09B07FFD7FFEFFB1D3C80FAC9F8D8" box="[1169,1303,1785,1810]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="253" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.20.3 mm</quantity>
long.
<emphasis id="B92CEAF0FFD7FFEFFAD63C80FF63F8FC" italics="true" pageId="1" pageNumber="253">Flower buds</emphasis>
globular, truncate at top, with 5 distinct lateral ridges, c.
<quantity id="4CA09B07FFD7FFEFFCC73D64FC53F8FC" box="[843,909,1821,1846]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="253" unit="mm" value="6.0">6 mm</quantity>
in diam. before anthesis.
<emphasis id="B92CEAF0FFD7FFEFFB3F3D64FAD7F8FC" box="[1203,1289,1821,1846]" italics="true" pageId="1" pageNumber="253">Corolla</emphasis>
rotate, deeply lobed,
<quantity id="4CA09B07FFD7FFEFFF583D38FE8FF890" box="[212,337,1857,1882]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="1" pageNumber="253" unit="cm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 cm</quantity>
in diam. when flattened, white tinged pink, thinly and minutely pubescent inside, outside glabrous, tube
<quantity id="4CA09B07FFD7FFEFFF4C3D1CFEC0F8B4" box="[192,286,1893,1918]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="253" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long,
<quantity id="4CA09B07FFD7FFEFFEE93D1CFE1DF8B4" box="[357,451,1893,1918]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="253" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
in diam.,
<emphasis id="B92CEAF0FFD7FFEFFDB93D1CFDAEF8B4" box="[565,624,1893,1918]" italics="true" pageId="1" pageNumber="253">lobes</emphasis>
laterally recurved, ovate with a triangular acute apex, slightly ascending, 67(9) ×
<quantity id="4CA09B07FFD7FFEFFE8F3DF0FE51F868" box="[259,399,1929,1954]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.5" metricValueMin="4.4" pageId="1" pageNumber="253" unit="mm" value="4.95" valueMax="5.5" valueMin="4.4">4.45.5 mm</quantity>
.
<emphasis id="B92CEAF0FFD7FFEFFE173DF0FDF1F868" box="[411,559,1929,1954]" italics="true" pageId="1" pageNumber="253">Gynostegium</emphasis>
stalked, stalk
<quantity id="4CA09B07FFD7FFEFFD593DF0FC80F868" box="[725,862,1929,1954]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="253" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
tall,
<quantity id="4CA09B07FFD7FFEFFC163DF0FBFDF868" box="[922,1059,1929,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="1" pageNumber="253" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
in diam. (base);
<emphasis id="B92CEAF0FFD7FFEFFB643DF0FAE3F868" box="[1256,1341,1929,1954]" italics="true" pageId="1" pageNumber="253">Corona</emphasis>
staminal,
<quantity id="4CA09B07FFD7FFEFFF043DD4FF39F80C" box="[136,231,1965,1990]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="253" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
tall,
<quantity id="4CA09B07FFD7FFEFFEA83DD4FE46F80C" box="[292,408,1965,1990]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.0" metricValueMin="5.5" pageId="1" pageNumber="253" unit="mm" value="6.25" valueMax="7.0" valueMin="5.5">5.57 mm</quantity>
in diam., fleshy;
<emphasis id="B92CEAF0FFD7FFEFFDE83DD4FD28F80C" box="[612,758,1965,1990]" italics="true" pageId="1" pageNumber="253">corona lobes</emphasis>
ovate, laterally compressed and slightly convex above, with revolute margins beneath, 34 ×
<quantity id="4CA09B07FFD7FFEFFE703DA8FDAFF820" box="[508,625,2001,2026]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="253" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
,
<quantity id="4CA09B07FFD7FFEFFDF13DA8FD05F820" box="[637,731,2001,2026]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="253" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
tall, inner process erecto-patent with an acute tip c.
<quantity id="4CA09B07FFD7FFEFFAA63DA8FAB2F820" box="[1322,1388,2001,2026]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="253" unit="mm" value="1.0">1 mm</quantity>
long, as long or slightly longer than the anthers, light pink, outer process ending in an incurved membranaceous appendage c.
<quantity id="4CA09B07FFD4FFECFF2E3AD9FF27FF73" box="[162,249,160,185]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="254" unit="mm" value="0.5">0.5 mm</quantity>
, sometimes much reduced, white or light yellow.
<emphasis id="B92CEAF0FFD4FFECFCAD3AD9FCA6FF73" box="[801,888,160,185]" italics="true" pageId="2" pageNumber="254">Anthers</emphasis>
ovate, 0.80.85 ×
<quantity id="4CA09B07FFD4FFECFBCB3AD9FB00FF73" box="[1095,1246,160,185]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="254" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.450.5 mm</quantity>
, with apical round membranaceous appendage as long as the style-head apex.
<taxonomicName id="4C584D61FFD4FFECFCBF3ABDFC53FF17" box="[819,909,196,221]" class="Magnoliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="2" pageNumber="254" phylum="Tracheophyta" rank="genus">
<emphasis id="B92CEAF0FFD4FFECFCBF3ABDFC53FF17" box="[819,909,196,221]" italics="true" pageId="2" pageNumber="254">Pollinia</emphasis>
</taxonomicName>
oblong, with round apex and base and evident pellucid margin, 600650 × 150180 μm;
<emphasis id="B92CEAF0FFD4FFECFDE33A91FD07FECB" box="[623,729,232,257]" italics="true" pageId="2" pageNumber="254">caudicles</emphasis>
attached at the base of the corpusculum, winged,
<emphasis id="B92CEAF0FFD4FFECFA973A91FA72FECB" box="[1307,1452,232,257]" italics="true" pageId="2" pageNumber="254">corpusculum</emphasis>
600650 × 250300 μm; style-head 5 angled in cross section, with 5 spreading lobes alternating with the stamens,
<emphasis id="B92CEAF0FFD4FFECFF043B49FF25FE83" box="[136,251,304,329]" italics="true" pageId="2" pageNumber="254">style-head</emphasis>
convex, stipitate, as tall as the anther appendages, c. 0.8 ×
<quantity id="4CA09B07FFD4FFECFC083B49FC02FE83" box="[900,988,304,329]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="254" unit="mm" value="0.6">0.6 mm</quantity>
.
<emphasis id="B92CEAF0FFD4FFECFC693B49FBF5FE83" box="[997,1067,304,329]" italics="true" pageId="2" pageNumber="254">Ovary</emphasis>
linear,
<quantity id="4CA09B07FFD4FFECFBF13B49FB33FE83" box="[1149,1261,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="254" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
long, ca.
<quantity id="4CA09B07FFD4FFECFADB3B49FF6AFEA7" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="254" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50.7 mm</quantity>
wide at the base.
<emphasis id="B92CEAF0FFD4FFECFEF03B2DFDF4FEA7" box="[380,554,340,365]" italics="true" pageId="2" pageNumber="254">Immature fruits</emphasis>
linear, c.
<quantity id="4CA09B07FFD4FFECFD143B2DFD03FEA7" box="[664,733,340,365]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="254" unit="cm" value="12.0">12 cm</quantity>
long, c.
<quantity id="4CA09B07FFD4FFECFCB33B2DFC5EFEA7" box="[831,896,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="254" unit="mm" value="5.0">5 mm</quantity>
diam., brownish-violet;
<emphasis id="B92CEAF0FFD4FFECFB1F3B2DFB1BFEA7" box="[1171,1221,340,365]" italics="true" pageId="2" pageNumber="254">seed</emphasis>
not seen.
</paragraph>
<caption id="DF27666AFFD4FFECFF043DF5FC04F823" ID-DOI="http://doi.org/10.5281/zenodo.13725783" ID-Zenodo-Dep="13725783" httpUri="https://zenodo.org/record/13725783/files/figure.png" pageId="2" pageNumber="254" startId="2.[136,229,1932,1953]" targetBox="[215,1372,389,1907]" targetPageId="2" targetType="figure">
<paragraph id="8BE736E2FFD4FFECFF043DF5FC04F823" blockId="2.[136,1452,1932,2025]" pageId="2" pageNumber="254">
<emphasis id="B92CEAF0FFD4FFECFF043DF5FF20F86B" bold="true" box="[136,254,1932,1953]" pageId="2" pageNumber="254">FIGURE 2.</emphasis>
<taxonomicName id="4C584D61FFD4FFECFE883DF5FE58F86B" box="[260,390,1932,1953]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="254" phylum="Tracheophyta" rank="species" species="corneri">
<emphasis id="B92CEAF0FFD4FFECFE883DF5FE58F86B" box="[260,390,1932,1953]" italics="true" pageId="2" pageNumber="254">Hoya corneri</emphasis>
</taxonomicName>
. A, inflorescence with buds and just opened flowers; B, Inflorescence with fully open flowers; C, Flower, top view; D, flower, side view; E, flower bud just before anthesis. F, leaf, adaxial side; G, leaf, abaxial side. A, CG by M. Rodda, based on
<emphasis id="B92CEAF0FFD4FFECFF043DADFEEDF823" box="[136,307,2004,2025]" italics="true" pageId="2" pageNumber="254">M. Rodda MR647</emphasis>
(SING). B by S. Somadee, based on
<emphasis id="B92CEAF0FFD4FFECFD153DADFC5BF823" box="[665,901,2004,2025]" italics="true" pageId="2" pageNumber="254">S. Somadee SR2012-017</emphasis>
(SING).
</paragraph>
</caption>
<caption id="DF27666AFFD5FFEDFF043C1DFA87F957" ID-DOI="http://doi.org/10.5281/zenodo.13725787" ID-Zenodo-Dep="13725787" httpUri="https://zenodo.org/record/13725787/files/figure.png" pageId="3" pageNumber="255" startId="3.[136,229,1636,1657]" targetBox="[204,1436,247,1611]" targetPageId="3" targetType="figure">
<paragraph id="8BE736E2FFD5FFEDFF043C1DFA87F957" blockId="3.[136,1451,1636,1693]" pageId="3" pageNumber="255">
<emphasis id="B92CEAF0FFD5FFEDFF043C1DFEDEF9B3" bold="true" box="[136,256,1636,1657]" pageId="3" pageNumber="255">FIGURE 3.</emphasis>
<taxonomicName id="4C584D61FFD5FFEDFE853C1DFE53F9B3" box="[265,397,1636,1657]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="corneri">
<emphasis id="B92CEAF0FFD5FFEDFE853C1DFE53F9B3" box="[265,397,1636,1657]" italics="true" pageId="3" pageNumber="255">Hoya corneri</emphasis>
</taxonomicName>
. A, corona, view from above; B, corona, view from below; C, calyx and ovary; D, corona, side view; E, pollinarium, F, transverse section of gynostegium. Based on
<emphasis id="B92CEAF0FFD5FFEDFD413CF1FC67F957" box="[717,953,1672,1693]" italics="true" pageId="3" pageNumber="255">S. Somadee SR2012-017</emphasis>
(SING). Photos by Songkran Upasontanon
</paragraph>
</caption>
<paragraph id="8BE736E2FFD5FFEAFF313CB7FB0DFEEF" blockId="3.[136,1452,1742,2091]" lastBlockId="4.[136,1452,160,977]" lastPageId="4" lastPageNumber="256" pageId="3" pageNumber="255">
<emphasis id="B92CEAF0FFD5FFEDFF313CB7FED7F92D" bold="true" box="[189,265,1742,1767]" pageId="3" pageNumber="255">Notes:</emphasis>
<taxonomicName id="4C584D61FFD5FFEDFEAA3CB7FE10F92D" box="[294,462,1742,1767]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD5FFEDFEAA3CB7FE10F92D" box="[294,462,1742,1767]" italics="true" pageId="3" pageNumber="255">Hoya uncinata</emphasis>
</taxonomicName>
was described based on materials collected from
<collectingRegion id="499CF800FFD5FFEDFB983CB7FB98F92D" box="[1044,1094,1742,1767]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="255">Java</collectingRegion>
and
<collectingRegion id="499CF800FFD5FFEDFB0E3CB7FB3DF92D" box="[1154,1251,1742,1767]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="255">Sumatra</collectingRegion>
. Its flowers were originally described as being negatively geotropic (i.e.,
<emphasis id="B92CEAF0FFD5FFEDFC8F3C8BFC56F8C1" box="[771,904,1778,1803]" italics="true" pageId="3" pageNumber="255">flores erecti</emphasis>
), a rare occurrence in
<taxonomicName id="4C584D61FFD5FFEDFB023C8BFB15F8C1" authorityName="Brown" authorityYear="1810" box="[1166,1227,1778,1803]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="genus">
<emphasis id="B92CEAF0FFD5FFEDFB023C8BFB15F8C1" box="[1166,1227,1778,1803]" italics="true" pageId="3" pageNumber="255">Hoya</emphasis>
</taxonomicName>
. Peduncles in
<taxonomicName id="4C584D61FFD5FFEDFAE33C8BFA72F8C1" authorityName="Brown" authorityYear="1810" box="[1391,1452,1778,1803]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="genus">
<emphasis id="B92CEAF0FFD5FFEDFAE33C8BFA72F8C1" box="[1391,1452,1778,1803]" italics="true" pageId="3" pageNumber="255">Hoya</emphasis>
</taxonomicName>
are commonly positively geotropic, less frequently neither positively or negatively geotropic “laterally held” and very rarely negatively geotropic (
<bibRefCitation id="EFC94B13FFD5FFEDFE4D3D43FD6FF899" author="Lamb, A. L. &amp; Rodda, M." box="[449,689,1850,1875]" pageId="3" pageNumber="255" refId="ref4520" refString="Lamb, A. L. &amp; Rodda, M. (2016) A guide to Hoyas of Borneo. Natural History Publications Borneo, Kota Kinabalu, 204 pp." type="book" year="2016">Lamb &amp; Rodda 2016</bibRefCitation>
). Within each inflorescence the orientation of flowers depends on the inflorescence
<typeStatus id="54E38840FFD5FFEDFEA93D27FE87F8BD" box="[293,345,1886,1911]" pageId="3" pageNumber="255">type</typeStatus>
. In globose inflorescences individual flowers range from positively to negatively geotropic, while in flat or concave/convex inflorescences all flowers are either positively or negatively geotropic. Species with both peduncles and individual flowers negatively geotropic are for example
<taxonomicName id="4C584D61FFD5FFEDFC3D3DDFFB0FF875" authority="Kerr (1940" authorityName="Kerr" authorityYear="1940" box="[945,1233,1958,1983]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="mitrata">
<emphasis id="B92CEAF0FFD5FFEDFC3D3DDFFB96F875" box="[945,1096,1958,1983]" italics="true" pageId="3" pageNumber="255">Hoya mitrata</emphasis>
<bibRefCitation id="EFC94B13FFD5FFEDFBDC3DDFFB0FF875" author="Kerr, A. F. G." box="[1104,1233,1958,1983]" pageId="3" pageNumber="255" refId="ref4475" refString="Kerr, A. F. G. (1940) Hoya mitrata. Hooker's Icones Plantarum 35: t. 3406" type="journal volume" year="1940">Kerr (1940</bibRefCitation>
</taxonomicName>
: t. 3406) and
<taxonomicName id="4C584D61FFD5FFEDFAE33DDFFDC7F829" authority="Schlechter (1913: 126)" authorityName="Schlechter" authorityPageNumber="126" authorityYear="1913" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="oreostemma">
<emphasis id="B92CEAF0FFD5FFEDFAE33DDFFECEF829" italics="true" pageId="3" pageNumber="255">Hoya oreostemma</emphasis>
<bibRefCitation id="EFC94B13FFD5FFEDFE9A3DB3FDC7F829" author="Schlechter, R." box="[278,537,1994,2019]" pageId="3" pageNumber="255" pagination="81 - 164" refId="ref4646" refString="Schlechter, R. (1913) Die Asclepiadaceen von Deutsch-Neu-Guinea. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50: 81 - 164." type="journal article" year="1913">Schlechter (1913: 126)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C584D61FFD5FFEDFDAE3DB3FD19F829" box="[546,711,1994,2019]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD5FFEDFDAE3DB3FD19F829" box="[546,711,1994,2019]" italics="true" pageId="3" pageNumber="255">Hoya uncinata</emphasis>
</taxonomicName>
has negatively geotropic peduncles (sometimes laterally held) and individual flowers that range from negatively geotropic to horizontal (rarely some flowers are positively geotropic in very rich inflorescences).
<taxonomicName id="4C584D61FFD5FFEDFE21326BFD8AF7E1" box="[429,596,2066,2091]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD5FFEDFE21326BFD8AF7E1" box="[429,596,2066,2091]" italics="true" pageId="3" pageNumber="255">Hoya uncinata</emphasis>
</taxonomicName>
was observed alive by Teijsmann and Binnendijk as they noted the tropism of the flowers, impossible to study on herbarium materials, as well as leaves and flowers colours. Apart from the erect flowers the species is characterised by laterally compressed corona lobes, convex above and bearing an apiculate, hooked and inflexed outer process that however is sometimes much reduced.
<taxonomicName id="4C584D61FFD2FFEAFC6E3A91FADDFECB" authority="Schltr." authorityName="Schlechter" authorityYear="1916" box="[994,1283,232,257]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="padangensis">
<emphasis id="B92CEAF0FFD2FFEAFC6E3A91FB6FFECB" box="[994,1201,232,257]" italics="true" pageId="4" pageNumber="256">Hoya padangensis</emphasis>
Schltr.
</taxonomicName>
, also described from
<collectingRegion id="499CF800FFD2FFEAFF493B75FEFBFEEF" box="[197,293,268,293]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="256">Sumatra</collectingRegion>
, bears this set of characters and is here treated as a new synonym of
<taxonomicName id="4C584D61FFD2FFEAFBAA3B75FB12FEEF" box="[1062,1228,268,293]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFBAA3B75FB12FEEF" box="[1062,1228,268,293]" italics="true" pageId="4" pageNumber="256">Hoya uncinata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE736E2FFD2FFEAFF313B49FE5EFD8F" blockId="4.[136,1452,160,977]" pageId="4" pageNumber="256">
At
<collectionCode id="ED49AE27FFD2FFEAFF6F3B49FED4FE83" box="[227,266,304,329]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="4" pageNumber="256" type="Herbarium">BO</collectionCode>
only one original specimen of
<taxonomicName id="4C584D61FFD2FFEAFDFA3B49FBD1FE83" authority="Scheffer (1876: 37)" authorityName="Scheffer" authorityPageNumber="37" authorityYear="1876" box="[630,1039,304,329]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="apiculata">
<emphasis id="B92CEAF0FFD2FFEAFDFA3B49FCF8FE83" box="[630,806,304,329]" italics="true" pageId="4" pageNumber="256">Hoya apiculata</emphasis>
<bibRefCitation id="EFC94B13FFD2FFEAFCA33B49FBD1FE83" author="Scheffer, L. C." box="[815,1039,304,329]" pageId="4" pageNumber="256" pagination="1 - 60" refId="ref4699" refString="Scheffer, L. C. (1876) Enumeration des plantes de la Nouvelle-Guinee, avec description des especes nouvelles. Annales du Jardin Botanique de Buitenzorg 1: 1 - 60." type="journal article" year="1876">Scheffer (1876: 37)</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C584D61FFD2FFEAFBBE3B49FA7DFE83" authority="Traill [1830: 28]" authorityName="Traill" authorityPageNumber="28" authorityYear="1830" box="[1074,1443,304,329]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B92CEAF0FFD2FFEAFBBE3B49FB04FE83" box="[1074,1242,304,329]" italics="true" pageId="4" pageNumber="256">Hoya australis</emphasis>
<bibRefCitation id="EFC94B13FFD2FFEAFB6F3B49FA7DFE83" author="Traill, J." box="[1251,1443,304,329]" pageId="4" pageNumber="256" pagination="16 - 30" refId="ref4765" refString="Traill, J. (1830) Accounts and descriptions of the several plants belonging to the genus Hoya, which are cultivated in the garden of the Horticultural Society at Chiswick. Transactions of the Horticultural Society of London 7: 16 - 30." type="journal article" year="1830">Traill [1830: 28]</bibRefCitation>
</taxonomicName>
) was found, which is a Teijsmann collection (
<bibRefCitation id="EFC94B13FFD2FFEAFD0C3B2DFC55FEA7" author="Forster, P. I. &amp; Liddle, D. J." box="[640,907,340,365]" pageId="4" pageNumber="256" pagination="507 - 519" refId="ref4382" refString="Forster, P. I. &amp; Liddle, D. J. (1994) Type Collections of Asclepiadaceae at Herbarium Bogoriense (BO). Australian Systematic Botany 7: 507 - 519. https: // doi. org / 10.1071 / SB 9940507" type="journal article" year="1994">Forster &amp; Liddle, 1994</bibRefCitation>
). Numerous original specimens of
<taxonomicName id="4C584D61FFD2FFEAFA9A3B2DFA75FEA7" box="[1302,1451,340,365]" pageId="4" pageNumber="256">
<emphasis id="B92CEAF0FFD2FFEAFA9A3B2DFA8DFEA7" box="[1302,1363,340,365]" italics="true" pageId="4" pageNumber="256">Hoya</emphasis>
species
</taxonomicName>
described by Teijsmann and Binnendijk have been located at
<collectionCode id="ED49AE27FFD2FFEAFCDE3B01FCBCFE5B" box="[850,866,376,401]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="4" pageNumber="256" type="Herbarium">L</collectionCode>
and
<collectionCode id="ED49AE27FFD2FFEAFC123B01FC6BFE5B" box="[926,949,376,401]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="4" pageNumber="256" type="Herbarium">U</collectionCode>
. After an accurate examination of all
<taxonomicName id="4C584D61FFD2FFEAFAE33B01FA72FE5B" authorityName="Brown" authorityYear="1810" box="[1391,1452,376,401]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="genus">
<emphasis id="B92CEAF0FFD2FFEAFAE33B01FA72FE5B" box="[1391,1452,376,401]" italics="true" pageId="4" pageNumber="256">Hoya</emphasis>
</taxonomicName>
materials at
<collectionCode id="ED49AE27FFD2FFEAFE943BE5FE9FFE7F" box="[280,321,412,437]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="4" pageNumber="256" type="Herbarium">BO</collectionCode>
,
<collectionCode id="ED49AE27FFD2FFEAFEDC3BE5FEBEFE7F" box="[336,352,412,437]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="4" pageNumber="256" type="Herbarium">L</collectionCode>
and
<collectionCode id="ED49AE27FFD2FFEAFE123BE5FE6CFE7F" box="[414,434,412,437]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="4" pageNumber="256" type="Herbarium">U</collectionCode>
we were unable to trace any original material of
<taxonomicName id="4C584D61FFD2FFEAFC773BE5FB7AFE7F" box="[1019,1188,412,437]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFC773BE5FB7AFE7F" box="[1019,1188,412,437]" italics="true" pageId="4" pageNumber="256">Hoya uncinata</emphasis>
</taxonomicName>
. It is possible that the original material of
<taxonomicName id="4C584D61FFD2FFEAFEE33BB9FE28FE13" box="[367,502,448,473]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFEE33BB9FE28FE13" box="[367,502,448,473]" italics="true" pageId="4" pageNumber="256">H. uncinata</emphasis>
</taxonomicName>
was lost or that the taxon description was based on live plants only. We therefore designate
<emphasis id="B92CEAF0FFD2FFEAFF753B9DFE02FE37" box="[249,476,484,509]" italics="true" pageId="4" pageNumber="256">
<collectionCode id="ED49AE27FFD2FFEAFF753B9DFED3FE37" box="[249,269,484,509]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="256">R</collectionCode>
. Schlechter 15916
</emphasis>
(
<collectionCode id="ED49AE27FFD2FFEAFE603B9DFDDEFE37" box="[492,512,484,509]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="256" type="Herbarium">B</collectionCode>
) as
<typeStatus id="54E38840FFD2FFEAFDA33B9DFD56FE37" box="[559,648,484,509]" pageId="4" pageNumber="256" type="neotype">neotype</typeStatus>
for
<taxonomicName id="4C584D61FFD2FFEAFD3B3B9DFCE5FE37" box="[695,827,484,509]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFD3B3B9DFCE5FE37" box="[695,827,484,509]" italics="true" pageId="4" pageNumber="256">H. uncinata</emphasis>
</taxonomicName>
. The same specimen is here designated as
<typeStatus id="54E38840FFD2FFEAFA913B9DFA5AFE37" box="[1309,1412,484,509]" pageId="4" pageNumber="256" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C584D61FFD2FFEAFF043871FE87FDEB" authorityName="Schlechter" authorityYear="1916" box="[136,345,520,545]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="padangensis">
<emphasis id="B92CEAF0FFD2FFEAFF043871FE87FDEB" box="[136,345,520,545]" italics="true" pageId="4" pageNumber="256">Hoya padangensis</emphasis>
</taxonomicName>
, in accordance with Art 9.1 &amp; 9.3 of the
<collectionCode id="ED49AE27FFD2FFEAFCA23871FC80FDEB" box="[814,862,520,545]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="256">ICN</collectionCode>
(
<bibRefCitation id="EFC94B13FFD2FFEAFCE23871FB93FDEB" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[878,1101,520,545]" pageId="4" pageNumber="256" refId="ref4811" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Glashutten: Koeltz Botanical Books, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B92CEAF0FFD2FFEAFC433871FBDCFDEB" box="[975,1026,520,545]" italics="true" pageId="4" pageNumber="256">et al</emphasis>
. 2018
</bibRefCitation>
). Therefore, the two names are homotypic synonyms.
</paragraph>
</subSubSection>
<subSubSection id="C3426569FFD2FFEAFF313829FF14FC1B" pageId="4" pageNumber="256" type="distribution">
<paragraph id="8BE736E2FFD2FFEAFF313829FE2AFD33" blockId="4.[136,1452,160,977]" pageId="4" pageNumber="256">
<emphasis id="B92CEAF0FFD2FFEAFF313829FE30FDA3" bold="true" box="[189,494,592,617]" pageId="4" pageNumber="256">Distribution and ecology:</emphasis>
<collectingCountry id="F34F7672FFD2FFEAFD873829FDA4FDA3" box="[523,634,592,617]" name="Indonesia" pageId="4" pageNumber="256">Indonesia</collectingCountry>
,
<collectingRegion id="499CF800FFD2FFEAFD0A3829FD66FDA3" box="[646,696,592,617]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="256">Java</collectingRegion>
and
<collectingRegion id="499CF800FFD2FFEAFD633829FC8EFDA3" box="[751,848,592,617]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="256">Sumatra</collectingRegion>
. Records of
<taxonomicName id="4C584D61FFD2FFEAFC523829FB5AFDA3" box="[990,1156,592,617]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFC523829FB5AFDA3" box="[990,1156,592,617]" italics="true" pageId="4" pageNumber="256">Hoya uncinata</emphasis>
</taxonomicName>
are few and based mostly on old and / or unlocalised materials. It was described from materials collected in
<collectingRegion id="499CF800FFD2FFEAFBAF380DFB8BFD47" box="[1059,1109,628,653]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="256">Java</collectingRegion>
and
<collectingRegion id="499CF800FFD2FFEAFB00380DFB32FD47" box="[1164,1260,628,653]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="256">Sumatra</collectingRegion>
, without specific location and elevation. According to the herbarium records
<taxonomicName id="4C584D61FFD2FFEAFCA638E1FC0EFD7B" box="[810,976,664,689]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFCA638E1FC0EFD7B" box="[810,976,664,689]" italics="true" pageId="4" pageNumber="256">Hoya uncinata</emphasis>
</taxonomicName>
collected at Gunung Pantjar, Bogor,
<collectingRegion id="499CF800FFD2FFEAFAF838E1FF62FD1F" country="Indonesia" name="Jawa Barat" pageId="4" pageNumber="256">West Java</collectingRegion>
, was growing close to warm spring water, at
<quantity id="4CA09B07FFD2FFEAFD4E38C5FCD4FD1F" box="[706,778,700,725]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="256" unit="m" value="500.0">500 m</quantity>
altitude and a specimen collected at Padang Panjang,
<collectingRegion id="499CF800FFD2FFEAFAF838C5FF36FD33" country="Indonesia" name="Sumatera Barat" pageId="4" pageNumber="256">West Sumatra</collectingRegion>
, was growing at
<specimenCount id="9D5EFD6BFFD2FFEAFE293899FE2AFD33" box="[421,500,736,761]" count="900" pageId="4" pageNumber="256" type="generic">
<quantity id="4CA09B07FFD2FFEAFE293899FE31FD33" box="[421,495,736,761]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="256" unit="m" value="900.0">900 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BE736E2FFD2FFEAFF31397DFF14FC1B" blockId="4.[136,1452,160,977]" pageId="4" pageNumber="256">
<emphasis id="B92CEAF0FFD2FFEAFF31397DFE6AFCD7" bold="true" box="[189,436,772,797]" pageId="4" pageNumber="256">Conservation status:</emphasis>
<materialsCitation id="3B303CBFFFD2FFEAFE43397DFC9DFC8B" collectionCode="A, SING" location="Java and Sumatra" pageId="4" pageNumber="256" specimenCount="1" stateProvince="Java and Sumatra">
<collectionCode id="ED49AE27FFD2FFEAFE43397DFE3CFCD7" box="[463,482,772,797]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="256" type="Herbarium">A</collectionCode>
rarely collected species from
<collectingRegion id="499CF800FFD2FFEAFCBE397DFC25FCD7" box="[818,1019,772,797]" isEnumeration="true" pageId="4" pageNumber="256">Java and Sumatra</collectingRegion>
. The most recently collected specimen was collected in
<quantity id="4CA09B07FFD2FFEAFECF3951FE46FC8B" box="[323,408,808,833]" metricMagnitude="1" metricUnit="m" metricValue="4.98602" pageId="4" pageNumber="256" unit="in" value="1963.0">1963 in</quantity>
<collectingRegion id="499CF800FFD2FFEAFE133951FE0FFC8B" box="[415,465,808,833]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="256">Java</collectingRegion>
is
<emphasis id="B92CEAF0FFD2FFEAFE7D3951FD3BFC8B" box="[497,741,808,833]" italics="true" pageId="4" pageNumber="256">Nengah Wirawan 128</emphasis>
(
<collectionCode id="ED49AE27FFD2FFEAFD783951FCE5FC8B" box="[756,827,808,833]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="4" pageNumber="256" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B303CBFFFD2FFEAFCC23951FF14FC1B" collectionCode="BO, B, DD" county="The" location="In" municipality="Backer" pageId="4" pageNumber="256" specimenCode="IUCN 2017" specimenCount="1" stateProvince="Sumatera Utara">
however, because of the heavily damaged flowers it is impossible to confirm its identity.
<collectingCounty id="62864E6EFFD2FFEAFD8A3935FDECFCAF" box="[518,562,844,869]" pageId="4" pageNumber="256">The</collectingCounty>
earlier record for the island is
<emphasis id="B92CEAF0FFD2FFEAFC0B3935FBCBFCAF" box="[903,1045,844,869]" italics="true" pageId="4" pageNumber="256">
<collectingMunicipality id="6B83AC98FFD2FFEAFC0B3935FC08FCAF" box="[903,982,844,869]" pageId="4" pageNumber="256">Backer</collectingMunicipality>
9040
</emphasis>
(
<collectionCode id="ED49AE27FFD2FFEAFBA83935FB91FCAF" box="[1060,1103,844,869]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="4" pageNumber="256" type="Herbarium">BO</collectionCode>
), collected in 1913.
<location id="8E876039FFD2FFEAFABC3935FA99FCAF" LSID="urn:lsid:plazi:treatment:03F187F4FFD7FFEAFF0438D4FF14FC1B:8E876039FFD2FFEAFABC3935FA99FCAF" box="[1328,1351,844,869]" county="The" municipality="Backer" name="In" pageId="4" pageNumber="256" stateProvince="Sumatera Utara">In</location>
<collectingRegion id="499CF800FFD2FFEAFAC23935FA75FCAF" box="[1358,1451,844,869]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="256">Sumatra</collectingRegion>
no collections have been made since
<location id="8E876039FFD2FFEAFDBC3909FD64FC43" LSID="urn:lsid:plazi:treatment:03F187F4FFD7FFEAFF0438D4FF14FC1B:8E876039FFD2FFEAFDBC3909FD64FC43" box="[560,698,880,905]" county="The" municipality="Backer" name="Schlechter's" pageId="4" pageNumber="256" stateProvince="Sumatera Utara">Schlechters</location>
<typeStatus id="54E38840FFD2FFEAFD4E3909FD2CFC43" box="[706,754,880,905]" pageId="4" pageNumber="256">type</typeStatus>
specimen
<emphasis id="B92CEAF0FFD2FFEAFCE13909FBECFC43" box="[877,1074,880,905]" italics="true" pageId="4" pageNumber="256">Schlechter 15916</emphasis>
(
<collectionCode id="ED49AE27FFD2FFEAFBC83909FB87FC43" box="[1092,1113,880,905]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="256" type="Herbarium">B</collectionCode>
). Without any information on population size and decline, the preliminary conservation assessment for
<taxonomicName id="4C584D61FFD2FFEAFC3439EDFB83FC67" box="[952,1117,916,941]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="uncinata">
<emphasis id="B92CEAF0FFD2FFEAFC3439EDFB83FC67" box="[952,1117,916,941]" italics="true" pageId="4" pageNumber="256">Hoya uncinata</emphasis>
</taxonomicName>
is Data Deficient (
<collectionCode id="ED49AE27FFD2FFEAFABC39EDFA82FC67" box="[1328,1372,916,941]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="4" pageNumber="256" type="Herbarium">DD</collectionCode>
,
<bibRefCitation id="EFC94B13FFD2FFEAFAEB39EDFF1CFC1B" author="IUCN Standards &amp; Petitions Subcommittee" pageId="4" pageNumber="256" refId="ref4429" refString="IUCN Standards and Petitions Subcommittee (2017) Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Available from: http: // cmsdocs. s 3. amazonaws. com / RedListGuidelines. pdf (accessed 29 September 2017)" type="book" year="2017">
<specimenCode id="DBFE9E99FFD2FFEAFAEB39EDFF1CFC1B" collectionCode="DD" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="4" pageNumber="256" type="Herbarium">IUCN 2017</specimenCode>
</bibRefCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>