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<document id="1C5AEB1678C9A780AA32AEEC03C7A576" ID-DOI="10.5252/zoosystema2024v46a25" ID-ISSN="1638-9387" ID-Zenodo-Dep="13907152" ID-ZooBank="D4AC1F89-AC34-43C4-9761-3F2015A02265" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728463297646" checkinUser="plazi" docAuthor="Segniagbeto, Gabriel Hoinsoudé, Mark-Oliver, Annemarie Ohler, Luca, Mark-Oliver Rödel, Alain, Luca Luiselli &amp; Dubois, Alain" docDate="2024" docId="03F187A4FFA9FFFFA9E9FE6BFC31F967" docLanguage="en" docName="zoosystema.2024.46.25.pdf" docOrigin="Zoosystema 46 (25)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a25.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Ptychadena oxyrhynchus Tchamba" docType="treatment" docVersion="2" lastPageNumber="658" masterDocId="FFC8FFDCFFB4FFE2AA73FFBDFFEFFFCF" masterDocTitle="Amphibians of Togo: taxonomy, distribution and conservation status" masterLastPageNumber="670" masterPageNumber="631" pageNumber="658" updateTime="1728463900544" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="67BFAC38A58DD7556696468398382C6A">Amphibians of Togo: taxonomy, distribution and conservation status</mods:title>
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<mods:namePart id="7E5958C128F0551ACCE606B669E9A3D8">Segniagbeto, Gabriel Hoinsoudé</mods:namePart>
<mods:affiliation id="05F4CE934CBB0A975A6A5977D3DC2D60">Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="9D14E23886FB718C23BE9492CB876AEC">Mark-Oliver, Annemarie Ohler</mods:namePart>
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<mods:namePart id="85C1909030B94B7BE239322D08465F7A">Luca, Mark-Oliver Rödel</mods:namePart>
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<mods:namePart id="A8B0493EE3B3A59862FA67E7299B9F9A">Alain, Luca Luiselli</mods:namePart>
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<mods:namePart id="4DE6816C0CF4A017A277A11133683046">Dubois, Alain</mods:namePart>
<mods:affiliation id="56D62D51BBE8BE49E5472C4DB23E2124">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
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<mods:title id="7283F7CAA71F7F82F09198F3C95AE74A">Zoosystema</mods:title>
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<mods:date id="81AC00774F9F7F33B2CA190C1D2C8A32">2024</mods:date>
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<taxonomicName id="4C584D31FFA9FFFFA9E9FE6BFAADFE3F" ID-CoL="4Q9R8" authority="(Smith, 1849)" authorityName="Tchamba" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[922,1346,470,497]" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" kingdom="Animalia" order="Anura" pageId="29" pageNumber="658" phylum="Chordata" rank="species" species="oxyrhynchus">
<emphasis id="B92CEAA0FFA9FFFFA9E9FE6BFB4DFE3E" bold="true" box="[922,1186,470,497]" italics="true" pageId="29" pageNumber="658">Ptychadena oxyrhynchus</emphasis>
(Smith, 1849)
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(
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)
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. —
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<collectingCountry id="F34F7622FFA9FFFFAE4CFD8CFB9AFD86" box="[1087,1141,561,585]" name="Togo" pageId="29" pageNumber="658">Togo</collectingCountry>
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,
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; Agou; MNHN- RA-1979.7145, MNHN-RA-1979.7146, MNHN-RA-1979.7149 •
<specimenCount id="9D5EFD3BFFA9FFFFA958FDDBFCBFFD4E" box="[811,848,614,641]" count="2" pageId="29" pageNumber="658" type="male">2 ♂</specimenCount>
,
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; Agoté; MNHN-RA-2006.2156-2158 •
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; Kanté; MNHN- RA-1979.7148 •
<specimenCount id="9D5EFD3BFFA9FFFFA9BFFD3DFC1EFD54" box="[972,1009,640,667]" count="3" pageId="29" pageNumber="658" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D5EFD3BFFA9FFFFA989FD3DFBF3FD54" box="[1018,1052,640,667]" count="2" pageId="29" pageNumber="658" type="female">2 ♀</specimenCount>
; Lomé; MNHN-RA-1979.7147, MNHN- RA-1979.7150, MNHN-RA-1979.7151, MNHN-RA-2006.2208 •
<specimenCount id="9D5EFD3BFFA9FFFFA94EFD08FC8BFD1F" box="[829,868,693,720]" count="1" pageId="29" pageNumber="658" type="male">1 ♂</specimenCount>
; Tové; MNHN-RA-1979.7144.
</paragraph>
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. — Large frog (SVL
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,
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), with robust body. Snout pointed, protruding beyond mouth slit. Head longer than broad (HW 30-35% SVL; HL 37-41% SVL). Tympanum distinct (TYD 6-8% SVL). Hind legs very long (TL 62-75% SVL). Webbing moderate; webbing formula: I 0 1 ½ II 0 2 III ½ 3 IV 2 ½ V. Toe tips pointed. Back marked by three pairs of glandular folds: the inner pair starting at the rear of the interorbital space, without contact with the eyelids and extending to the posterior end of the body; the middle pair starting at the posterior edge of the eyelids and ending at the groin; and the outer pair constituting the supra-tympanic folds, extending to the inguinal region; slightly shagreen skin on the back and flanks.
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<paragraph id="8BE736B2FFA9FFFFA95EFB99FC25FB99" blockId="29.[811,1458,742,1110]" pageId="29" pageNumber="658">The most conspicuous characters of this species are its extremely long hind limbs</paragraph>
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. — The dorsal colouration is grayish or olive-greenish, brownish in some specimens, with irregularly set white spots. The vertebral line is distinct. The dorsal sides of the thighs and legs have brownish transverse bars. The belly is white. In West Africa it is the only
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<emphasis id="B92CEAA0FFA9FFFFA9EFFB5CFBE8FB36" box="[924,1031,1249,1273]" italics="true" pageId="29" pageNumber="658">Ptychadena</emphasis>
</taxonomicName>
which combines a snout with colour paler than the rest of the head and reddish colouration on the posterior part of the eyelids.
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. — Males have two pairs of infer vocal sac openings, laterally on either side of the throat, and nuptial pads on first to third fingers.
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. — This species is characteristic of savannah associated with forest areas. Very frequent in ecological zones II, III and IV, it is observed in the regions of Kpélé, Adélé (MPoti and Koui), Fazao (Kalaré), Tchamba, Alibi, etc. Females lay eggs in the rainy season in small pools or water reservoirs. The species was reported by
<bibRefCitation id="EFC94B43FFA9FFFFAE51F9FCFB50F997" author="BOURGAT R." box="[1058,1215,1600,1625]" pageId="29" pageNumber="658" pagination="597 - 624" refId="ref33022" refString="BOURGAT R. 1979. - Trematodes d'amphibiens du Togo. Bulletin du Museum national d'Histoire naturelle (4 e serie) 3: 597 - 624." type="journal article" year="1979">Bourgat (1979)</bibRefCitation>
from Kovié, Lomé and Kanté, and by
<bibRefCitation id="EFC94B43FFA9FFFFA9B3F9E7FB5CF9BC" author="SEGNIAGBETO G. H. &amp; DEKAWOLE J. K. &amp; KETOH G. K. &amp; DENDI D. &amp; LUISELLI L." box="[960,1203,1626,1651]" pageId="29" pageNumber="658" refId="ref38700" refString="SEGNIAGBETO G. H., DEKAWOLE J. K., KETOH G. K., DENDI D. &amp; LUISELLI L. 2022. - Herpetofaunal diversity in a Dahomey Gap savannah of Togo (West Africa): effects of seasons on the populations of amphibians and reptiles. Diversity 14, 964." type="journal volume" year="2022">
Segniagbeto
<emphasis id="B92CEAA0FFA9FFFFAE49F9E6FB85F9BD" box="[1082,1130,1626,1650]" italics="true" pageId="29" pageNumber="658">et al.</emphasis>
(2022)
</bibRefCitation>
from Tetetou to Nagbeto dam.
<bibRefCitation id="EFC94B43FFA9FFFFA917F9C8FC09F942" author="RODEL M. - O." box="[868,998,1653,1677]" pageId="29" pageNumber="658" refId="ref37146" refString="RODEL M. - O. 2000. - Herpetofauna of West Africa. Vol 1. Amphibians of theWest African savanna. ̿ rankfurt am Main, Chimaira, 335 p." type="book" year="2000">Rödel (2000)</bibRefCitation>
reported its presence in ecological zones II and IV of the country.
</paragraph>
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