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<mods:title id="D6BBE7D0CDA41A2AF69AE60384D0A236">First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species</mods:title>
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Key to species of
<taxonomicName id="4C0EDBE0FFCB3C6DCA6F6CC082F3FA3E" ID-CoL="7RNG" authorityName="Loew" authorityYear="1857" box="[366,472,1436,1462]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B97A7C71FFCB3C6DCA6F6CC082F3FA3E" bold="true" box="[366,472,1436,1462]" italics="true" pageId="1" pageNumber="258">Systenus</emphasis>
</taxonomicName>
from
<collectingCountry id="F319E0F3FFCB3C6DC9256CC081B5FA3E" box="[548,670,1436,1462]" name="Colombia" pageId="1" pageNumber="258">Colombia</collectingCountry>
(males)
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<keyStep id="30FAB8C6FFCB3C6DCB966CBD86B7F99B" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966CBD86B7FA7E" blockId="1.[151,1437,1505,2026]" box="[151,1436,1505,1527]" pageId="1" pageNumber="258">
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1 Metepimeron dark (
<emphasis id="B97A7C71FFCB3C6DCAA36CBE82EEFA7F" box="[418,453,1506,1527]" italics="true" pageId="1" pageNumber="258">e.g.</emphasis>
<figureCitation id="1335BCE6FFCB3C6DCACB6CBD8132FA7E" box="[458,537,1505,1527]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9226CBD816FFA7E" box="[547,580,1505,1526]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2A</figureCitation>
); abdominal tergites 13 mostly metallic green (
<emphasis id="B97A7C71FFCB3C6DCF146CBE8713FA7F" box="[1045,1080,1506,1527]" italics="true" pageId="1" pageNumber="258">e.g.</emphasis>
<figureCitation id="1335BCE6FFCB3C6DCF3C6CBD87A2FA7E" box="[1085,1161,1505,1527]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1E</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF926CBD8784FA7E" box="[1171,1199,1505,1526]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2E</figureCitation>
).................. 3
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<paragraph id="8BB1A063FFCB3C6DCB966CA386B7F99B" blockId="1.[151,1437,1505,2026]" box="[151,1436,1534,1556]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966CA386B7F99B" box="[151,1436,1534,1556]" pageId="1" pageNumber="258">
- Metepimeron yellow (
<figureCitation id="1335BCE6FFCB3C6DCAB66CA2812DF99C" box="[439,518,1534,1556]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Figs 5C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9116CA2811BF99C" box="[528,560,1534,1556]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">6A</figureCitation>
); abdominal tergites 13 partly yellow (
<figureCitation id="1335BCE6FFCB3C6DC8AE6CA280D5F99C" box="[943,1022,1534,1556]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Figs 5C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF096CA2870CF99C" box="[1032,1063,1534,1556]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">6A</figureCitation>
)............................. 2
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<keyStep id="30FAB8C6FFCB3C6DCB966F4086B6F902" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966F4086B6F9C7" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966F4086B6F9C7" pageId="1" pageNumber="258">
2 Femur III entirely yellow (
<figureCitation id="1335BCE6FFCB3C6DCAE56F40811BF9B9" box="[484,560,1563,1586]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
); abdominal tergite 3 dorsally metallic green, tergites 45 wholly metallic green (
<figureCitation id="1335BCE6FFCB3C6DCE456F4086BBF9B9" box="[1348,1424,1563,1586]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
); surstylus slightly curved ventrally at apex (
<figureCitation id="1335BCE6FFCB3C6DC9816F6581E0F9C7" box="[640,715,1593,1615]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5D</figureCitation>
)............................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFEC6F6586B6F9C7" ID-CoL="544RL" authority="Naglis" authorityName="Naglis" authorityYear="2000" box="[1261,1437,1593,1615]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="flaviatus">
<emphasis id="B97A7C71FFCB3C6DCFEC6F658673F9C6" box="[1261,1368,1593,1614]" italics="true" pageId="1" pageNumber="258">S. flaviatus</emphasis>
Naglis
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</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966F0B86B6F902" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966F0B86B6F902" pageId="1" pageNumber="258">
- Femur III yellow with dark brown apex (
<figureCitation id="1335BCE6FFCB3C6DC9676F0A8198F9E4" box="[614,691,1622,1644]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6A</figureCitation>
); abdominal tergite 3 mostly yellow (
<figureCitation id="1335BCE6FFCB3C6DCF166F0A8748F9E4" box="[1047,1123,1622,1644]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6A</figureCitation>
), with T-shaped macula dorsally, tergites 45 dark brown; surstylus abruptly curved ventrally at apex (
<figureCitation id="1335BCE6FFCB3C6DC8796F2880E9F902" box="[888,962,1652,1674]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6B</figureCitation>
).....................
<taxonomicName id="4C0EDBE0FFCB3C6DCFD16F2886B6F902" ID-CoL="9HSKT" authority="Bickel" authorityName="Bickel" authorityYear="2015" box="[1232,1437,1652,1674]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="naranjensis">
<emphasis id="B97A7C71FFCB3C6DCFD16F2887F7F901" box="[1232,1244,1652,1673]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCFE96F288673F901" box="[1256,1368,1652,1673]" italics="true" pageId="1" pageNumber="258">naranjensis</emphasis>
Bickel
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966FCD86B7F94B" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966FCD86B7F92E" blockId="1.[151,1437,1505,2026]" box="[151,1436,1681,1703]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966FCD86B7F92E" box="[151,1436,1681,1703]" pageId="1" pageNumber="258">
3 Antenna wholly dark brown (
<figureCitation id="1335BCE6FFCB3C6DCAFC6FCD814EF92E" box="[509,613,1681,1703]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">Fig. 3A, B</figureCitation>
); femora mostly dark brown (
<figureCitation id="1335BCE6FFCB3C6DC8856FCD80E4F92E" box="[900,975,1681,1703]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">Fig. 3A</figureCitation>
).......................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF76FCD8661F92E" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1270,1354,1681,1702]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B97A7C71FFCB3C6DCFF76FCD8661F92E" box="[1270,1354,1681,1702]" italics="true" pageId="1" pageNumber="258">S. fuscus</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFCB3C6DCE516FCD86B7F92E" bold="true" box="[1360,1436,1681,1702]" pageId="1" pageNumber="258">
<taxonomicNameLabel id="A249C10AFFCB3C6DCE516FCD86B7F92E" box="[1360,1436,1681,1702]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966FF386B7F94B" blockId="1.[151,1437,1505,2026]" box="[151,1436,1710,1732]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966FF386B7F94B" box="[151,1436,1710,1732]" pageId="1" pageNumber="258">
- Antenna partly orangish yellow (
<figureCitation id="1335BCE6FFCB3C6DC91F6FF28147F94B" box="[542,620,1710,1732]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9776FF281BEF94B" box="[630,661,1710,1731]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC99F6FF28196F94B" box="[670,701,1710,1731]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); femora yellow (
<figureCitation id="1335BCE6FFCB3C6DC8666FF2809CF94B" box="[871,951,1710,1732]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Figs 2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC8C06FF280F4F94B" box="[961,991,1710,1731]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
)................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966F9086B7F894" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966F9086B6F977" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966F9086B6F977" pageId="1" pageNumber="258">
4 It
<subScript id="178AA226FFCB3C6DCBF06F8783D3F96F" attach="left" box="[241,248,1755,1767]" fontSize="5" pageId="1" pageNumber="258">3</subScript>
white (
<figureCitation id="1335BCE6FFCB3C6DCA436F9082A7F969" box="[322,396,1739,1762]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5E</figureCitation>
); wing apex brown: posterior wing membrane reduced near apex, with vein M
<subScript id="178AA226FFCB3C6DCF896F8787A4F96F" attach="left" box="[1160,1167,1755,1767]" fontSize="5" pageId="1" pageNumber="258">1</subScript>
ending on posterior margin (
<figureCitation id="1335BCE6FFCB3C6DCBE86FB5821AF977" box="[233,305,1769,1791]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5E</figureCitation>
)...........................................................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFC16FB586B6F977" ID-CoL="9HWXR" authority="Bickel" authorityName="Bickel" authorityYear="2015" box="[1216,1437,1769,1791]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B97A7C71FFCB3C6DCFC16FB587E7F976" box="[1216,1228,1769,1790]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCFD96FB58673F976" box="[1240,1368,1769,1790]" italics="true" pageId="1" pageNumber="258">maculipennis</emphasis>
Bickel
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966E5B86B7F894" blockId="1.[151,1437,1505,2026]" box="[151,1436,1798,1826]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E5B86B7F894" box="[151,1436,1798,1826]" pageId="1" pageNumber="258">
- It 3 yellow (
<figureCitation id="1335BCE6FFCB3C6DCA4D6E5A82B7F893" box="[332,412,1798,1820]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Figs 2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCAA76E5A82ECF893" box="[422,455,1798,1819]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
); wing apex unmodified, membrane hyaline or brownish (
<figureCitation id="1335BCE6FFCB3C6DC8F26E5A8768F893" box="[1011,1091,1798,1820]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1D</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF4C6E5A8747F893" box="[1101,1132,1798,1819]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2D</figureCitation>
)....................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966E7F86B7F81A" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966E7F86B6F8DF" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E7F86B6F8DF" pageId="1" pageNumber="258">
5 Postpedicel wholly orangish yellow at basal 1/2 (
<figureCitation id="1335BCE6FFCB3C6DC9CE6E788035F8B2" box="[719,798,1828,1850]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6C</figureCitation>
); abdominal segment 7 forming peduncle shorter than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCA366E1D82A8F8DF" box="[311,387,1857,1879]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6C</figureCitation>
); cercus rectangular, shorter than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DC87A6E1D80EDF8DF" box="[891,966,1857,1879]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. AB. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. CD. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6D</figureCitation>
).........................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF86E1D86B6F8DF" ID-CoL="9HYXF" authority="Bickel" authorityName="Bickel. A" authorityYear="2015" box="[1273,1437,1857,1879]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="parkeri">
<emphasis id="B97A7C71FFCB3C6DCFF86E1D862EF8DE" box="[1273,1285,1857,1878]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCE106E1D8673F8DE" box="[1297,1368,1857,1878]" italics="true" pageId="1" pageNumber="258">parkeri</emphasis>
Bickel
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966E0386B7F81A" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E0386B7F81A" pageId="1" pageNumber="258">
- Postpedicel partly brownish at basal 1/2 (
<figureCitation id="1335BCE6FFCB3C6DC97F6E0281E4F8FB" box="[638,719,1886,1908]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9DD6E0281D1F8FB" box="[732,762,1886,1907]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC8066E02800DF8FB" box="[775,806,1886,1907]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); abdominal segment 7 forming peduncle longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCA366E2082ADF819" box="[311,390,1916,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCA916E20829BF819" box="[400,432,1916,1937]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCABB6E2082F0F819" box="[442,475,1916,1937]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
); cercus digitiform, as long or longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCF296E20875AF819" box="[1064,1137,1916,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF7D6E2087BEF819" box="[1148,1173,1916,1937]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF9E6E208792F819" box="[1183,1209,1916,1937]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4F</figureCitation>
)................. 6
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<paragraph id="8BB1A063FFCB3C6DCB966EC586B7F843" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966EC586B7F843" pageId="1" pageNumber="258">
6 Palpus mostly brown (
<figureCitation id="1335BCE6FFCB3C6DCABF6EC58121F826" box="[446,522,1945,1967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1B</figureCitation>
); wing membrane light brown (
<figureCitation id="1335BCE6FFCB3C6DC8406EC580A4F826" box="[833,911,1945,1967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1D</figureCitation>
); cercus about 2X longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCE746EC583D7F843" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1F</figureCitation>
)....................................................................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF36EEB8661F843" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1266,1354,1974,1996]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
<emphasis id="B97A7C71FFCB3C6DCFF36EEB87D5F844" box="[1266,1278,1975,1996]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCE0B6EEA8661F843" box="[1290,1354,1974,1995]" italics="true" pageId="1" pageNumber="258">bickeli</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFCB3C6DCE516EEA86B7F843" bold="true" box="[1360,1436,1974,1995]" pageId="1" pageNumber="258">
<taxonomicNameLabel id="A249C10AFFCB3C6DCE516EEA86B7F843" box="[1360,1436,1974,1995]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6ECB966E8886B7FF25" blockId="1.[151,1437,1505,2026]" lastBlockId="2.[151,1437,151,379]" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6ECB966E8886B7FF25" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258">
- Palpus yellow (
<figureCitation id="1335BCE6FFCB3C6DCA7D6E8882E7F861" box="[380,460,2004,2026]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Fig. 2B</figureCitation>
); wing membrane hyaline (
<figureCitation id="1335BCE6FFCB3C6DC9DF6E888005F861" box="[734,814,2004,2026]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Fig. 2D</figureCitation>
); cercus as long or only slightly longer than epandrial capsule (
<figureCitation id="1335BCE6FFC83C6ECBE869CB821EFF25" box="[233,309,151,173]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Figs 2F</figureCitation>
,
<figureCitation id="1335BCE6FFC83C6ECA3E69C48272FF25" box="[319,345,152,173]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">4F</figureCitation>
)......................................................................................... 7
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<keyStep id="30FAB8C6FFC83C6ECB9669E986B7FE8D" pageId="2" pageNumber="259">
<paragraph id="8BB1A063FFC83C6ECB9669E986B6FF60" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB9669E986B6FF60" pageId="2" pageNumber="259">
7 Surstylus longer than cercus and epandrial capsule, ventral edge nearly straight, only slightly narrowed and curved ventrally at apex (
<figureCitation id="1335BCE6FFC83C6ECA1D698E824DFF60" box="[284,358,210,232]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. AB. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. CD. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="2" pageNumber="259">Fig. 5B</figureCitation>
)............................................................................
<taxonomicName id="4C0EDBE0FFC83C6ECE01698E86B6FF60" ID-CoL="544RF" authority="Naglis" authorityName="Naglis" authorityYear="2000" box="[1280,1437,210,232]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="beatae">
<emphasis id="B97A7C71FFC83C6ECE01698E8627FF6F" box="[1280,1292,210,231]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECE19698E8673FF6F" box="[1304,1368,210,231]" italics="true" pageId="2" pageNumber="259">beatae</emphasis>
Naglis
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFC83C6ECB9669AC86B7FE8D" blockId="2.[151,1437,151,379]" box="[151,1436,239,261]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB9669AC86B7FE8D" box="[151,1436,239,261]" pageId="2" pageNumber="259">
- Surstylus shorter or as long as cercus and epandrial capsule and abruptly narrowed towards apex (
<figureCitation id="1335BCE6FFC83C6ECF8C69B387FCFE8D" box="[1165,1239,239,261]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Figs 2F</figureCitation>
,
<figureCitation id="1335BCE6FFC83C6ECFE069AC87D0FE8D" box="[1249,1275,240,261]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">4F</figureCitation>
)........... 8
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<keyStep id="30FAB8C6FFC83C6ECB96685186B7FEF2" pageId="2" pageNumber="259">
<paragraph id="8BB1A063FFC83C6ECB96685186B7FEB7" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB96685186B7FEB7" pageId="2" pageNumber="259">
8 Coxa III mostly yellow (
<figureCitation id="1335BCE6FFC83C6ECACD68518132FEAA" box="[460,537,269,291]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">Fig. 4A</figureCitation>
); surstylus as long as cercus, wide, abruptly narrowed at apex, with 1 basodorsal acute process (
<figureCitation id="1335BCE6FFC83C6ECBE868768260FEB7" box="[233,331,298,320]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">Fig. 4F, G</figureCitation>
)............................................................................
<taxonomicName id="4C0EDBE0FFC83C6ECFE868768662FEC8" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1257,1353,298,320]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="grossus" status="sp. nov.">
<emphasis id="B97A7C71FFC83C6ECFE8687687DEFEB7" box="[1257,1269,298,319]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECE0068778662FEC8" box="[1281,1353,299,320]" italics="true" pageId="2" pageNumber="259">grossus</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFC83C6ECE51687686B7FEB7" bold="true" box="[1360,1436,298,319]" pageId="2" pageNumber="259">
<taxonomicNameLabel id="A249C10AFFC83C6ECE51687686B7FEB7" box="[1360,1436,298,319]" pageId="2" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BB1A063FFC83C6ECB96681486B7FEF2" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB96681486B7FEF2" pageId="2" pageNumber="259">
- Coxa III mostly brown (
<figureCitation id="1335BCE6FFC83C6ECAD2681B8108FED5" box="[467,547,327,349]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Fig. 2A</figureCitation>
); surstylus shorter than cercus, narrow, slightly narrowed at apex, with 1 basodorsal acute process and 1 apicoventral acute process (
<figureCitation id="1335BCE6FFC83C6EC97468398197FEF2" box="[629,700,357,379]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Fig. 2F</figureCitation>
).........................................
<taxonomicName id="4C0EDBE0FFC83C6ECFB568398662FEF2" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1204,1353,357,378]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="B97A7C71FFC83C6ECFB5683987EBFEF2" box="[1204,1216,357,378]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECFCD68398662FEF2" box="[1228,1353,357,378]" italics="true" pageId="2" pageNumber="259">colombiensis</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFC83C6ECE51683986B7FEF2" bold="true" box="[1360,1436,357,378]" pageId="2" pageNumber="259">
<taxonomicNameLabel id="A249C10AFFC83C6ECE51683986B7FEF2" box="[1360,1436,357,378]" pageId="2" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
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</treatment>
</document>