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<document id="BEE886399BBC97D02A1664A7FAFC0FD9" ID-DOI="10.11646/zootaxa.5514.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849826" ID-ZooBank="D745608E-E51C-401E-A1A1-3573A24C51BD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1727441634877" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany, Merchan-Vargas, Geraldine Lucia &amp; Soares, Matheus M. M." docDate="2024" docId="03A71175FFCB3C6DCB966A8E81E2FAC2" docLanguage="en" docName="zootaxa.5514.3.3.pdf" docOrigin="Zootaxa 5514 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus Loew 1857" docType="treatment" docVersion="4" lastPageNumber="258" masterDocId="FF9E690DFFCA3C6CCB01695C832BFF88" masterDocTitle="First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species" masterLastPageNumber="272" masterPageNumber="257" pageNumber="258" updateTime="1727718066609" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="DAD073FFEF69781D5B151F9836A62858">First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species</mods:title>
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<mods:affiliation id="E8A7A48E4650A5EBFB675E263CF202F5">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<treatment id="03A71175FFCB3C6DCB966A8E81E2FAC2" ID-DOI="http://doi.org/10.5281/zenodo.13849922" ID-Zenodo-Dep="13849922" LSID="urn:lsid:plazi:treatment:03A71175FFCB3C6DCB966A8E81E2FAC2" httpUri="http://treatment.plazi.org/id/03A71175FFCB3C6DCB966A8E81E2FAC2" lastPageNumber="258" pageId="1" pageNumber="258">
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<paragraph id="8BB1A063FFCB3C6DCB966A8E828FFC65" blockId="1.[151,420,978,1005]" box="[151,420,978,1005]" pageId="1" pageNumber="258">
<heading id="D0F9170FFFCB3C6DCB966A8E828FFC65" bold="true" box="[151,420,978,1005]" fontSize="11" level="1" pageId="1" pageNumber="258" reason="1">
<emphasis id="B97A7C71FFCB3C6DCB966A8E828FFC65" bold="true" box="[151,420,978,1005]" pageId="1" pageNumber="258">
Genus
<taxonomicName id="4C0EDBE0FFCB3C6DCBEF6A8F828FFC65" ID-CoL="7RNG" authority="Loew" authorityName="Loew" authorityYear="1857" box="[238,420,979,1005]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B97A7C71FFCB3C6DCBEF6A8F8273FC65" bold="true" box="[238,344,979,1005]" italics="true" pageId="1" pageNumber="258">Systenus</emphasis>
Loew
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<paragraph id="8BB1A063FFCB3C6DCB966D4B82B7FBA7" blockId="1.[151,1437,1047,1102]" box="[151,412,1047,1071]" pageId="1" pageNumber="258">
<treatmentCitationGroup id="AB1E874DFFCB3C6DCB966D4B82B7FBA7" box="[151,412,1047,1071]" pageId="1" pageNumber="258">
<treatmentCitation id="0AAF8672FFCB3C6DCB966D4B82B3FBA7" author="Loew, H." box="[151,408,1047,1071]" page="34" pageId="1" pageNumber="258" year="1857">
<taxonomicName id="4C0EDBE0FFCB3C6DCB966D4B82B3FBA7" ID-CoL="7RNG" authority="Loew, 1857: 34" authorityName="Loew" authorityPageNumber="34" authorityYear="1857" box="[151,408,1047,1071]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B97A7C71FFCB3C6DCB966D4B83C5FBA6" box="[151,238,1047,1070]" italics="true" pageId="1" pageNumber="258">Systenus</emphasis>
<bibRefCitation id="EF9FDD92FFCB3C6DCBF46D4482B3FBA7" author="Loew, H." box="[245,408,1047,1071]" pageId="1" pageNumber="258" pagination="1 - 56" refId="ref11207" refString="Loew, H. (1857) Neue Beitrage zur Kenntniss der Dipteren. Funfter Beitrage. Programme der Koniglichen Realschule zu Meseritz, 1857, 1 - 56." type="journal article" year="1857">Loew, 1857: 34</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C314F3E8FFCB3C6DCAA36D448246FBC6" pageId="1" pageNumber="258" type="type_taxon">
<paragraph id="8BB1A063FFCB3C6DCAA36D448246FBC6" blockId="1.[151,1437,1047,1102]" pageId="1" pageNumber="258">
<typeStatus id="54B51EC1FFCB3C6DCAA36D4482FFFBB8" box="[418,468,1048,1072]" pageId="1" pageNumber="258">Type</typeStatus>
species,
<taxonomicName id="4C0EDBE0FFCB3C6DC9336D4B8092FBA7" ID-CoL="BB2MC" authority="Loew, 1857" authorityName="Loew" authorityYear="1857" box="[562,953,1047,1071]" class="Insecta" family="Dolichopodidae" genus="Rhaphium" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="adpropinquans">
<emphasis id="B97A7C71FFCB3C6DC9336D4B8012FBA7" box="[562,825,1047,1071]" italics="true" pageId="1" pageNumber="258">Rhaphium adpropinquans</emphasis>
<bibRefCitation id="EF9FDD92FFCB3C6DC8416D448092FBA7" author="Loew, H." box="[832,953,1047,1071]" pageId="1" pageNumber="258" pagination="1 - 56" refId="ref11207" refString="Loew, H. (1857) Neue Beitrage zur Kenntniss der Dipteren. Funfter Beitrage. Programme der Koniglichen Realschule zu Meseritz, 1857, 1 - 56." type="journal article" year="1857">Loew, 1857</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C0EDBE0FFCB3C6DC8DC6D4B860AFBA7" ID-CoL="8DML5" authority="Roser, 1840" authorityName="Roser" authorityYear="1840" box="[989,1313,1047,1071]" class="Insecta" family="Dolichopodidae" genus="Rhaphium" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B97A7C71FFCB3C6DC8DC6D4B87B7FBA7" box="[989,1180,1047,1071]" italics="true" pageId="1" pageNumber="258">Rhaphium pallipes</emphasis>
<bibRefCitation id="EF9FDD92FFCB3C6DCFA26D44860AFBA7" author="Roser, K. L. F. von" box="[1187,1313,1047,1071]" pageId="1" pageNumber="258" pagination="49 - 64" refId="ref11286" refString="Roser, K. L. F. von (1840) Erster Nachtrag zu dem im Jahre 1834 bekannt gemachten Verzeichnisse in Wurttemberg vorkommender zweiflugliger Insekten. Correspondenzblatt de Koniglich Wurttembergischen Landwirtschaftlichen Vereins, Stuttgart, New Series 17, 37 (1), 49 - 64." type="journal article" year="1840">Roser, 1840</bibRefCitation>
</taxonomicName>
), des. Foote
<emphasis id="B97A7C71FFCB3C6DCBC66D6B83DFFBC6" box="[199,244,1078,1102]" italics="true" pageId="1" pageNumber="258">et al</emphasis>
. 1965: 517.
</paragraph>
</subSubSection>
<subSubSection id="C314F3E8FFCB3C6DCB966D2081E2FAC2" pageId="1" pageNumber="258" type="diagnosis">
<paragraph id="8BB1A063FFCB3C6DCB966D2081E2FAC2" blockId="1.[151,1437,1148,1355]" pageId="1" pageNumber="258">
<emphasis id="B97A7C71FFCB3C6DCB966D208239FB1E" bold="true" box="[151,274,1148,1174]" pageId="1" pageNumber="258">Diagnosis.</emphasis>
The genus is easily recognized by the males with postpedicel triangular or subrectangular at base, elongate about 2–4 times longer than basal width, slightly to strongly tapered at apical 1/2 to 2/3 (
<figureCitation id="1335BCE6FFCB3C6DCFDF6DFC8612FB32" box="[1246,1337,1184,1210]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCE476DFC8642FB32" box="[1350,1385,1184,1210]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCE756DFC86BCFB32" box="[1396,1431,1184,1210]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">3B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCB966D988390FB56" box="[151,187,1220,1246]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); proepisternum with 1 pale seta (
<figureCitation id="1335BCE6FFCB3C6DC9316D9881A0FB56" box="[560,651,1220,1246]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">Figs 3B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9976D988191FB56" box="[662,698,1220,1246]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); R
<subScript id="178AA226FFCB3C6DC9E16D8A81D1FB6C" attach="left" box="[736,762,1238,1252]" fontSize="6" pageId="1" pageNumber="258">4+5</subScript>
and M
<subScript id="178AA226FFCB3C6DC8466D8A8064FB6C" attach="left" box="[839,847,1238,1252]" fontSize="6" pageId="1" pageNumber="258">1</subScript>
subapically bowed, M
<subScript id="178AA226FFCB3C6DCF4A6D8A8778FB6C" attach="left" box="[1099,1107,1238,1252]" fontSize="6" pageId="1" pageNumber="258">1</subScript>
beyond dm-m crossvein with flexion (
<figureCitation id="1335BCE6FFCB3C6DCBF66DB4827BFA8A" box="[247,336,1256,1282]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1D</figureCitation>
); six strong dorsocentral setae (
<figureCitation id="1335BCE6FFCB3C6DC9B96DB4803BFA8A" box="[696,784,1256,1282]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1C</figureCitation>
); abdominal segment 7 with tergite and sternite distinctly elongate, forming a peduncle (
<figureCitation id="1335BCE6FFCB3C6DCAE96C508168FAAE" box="[488,579,1292,1318]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Figs 2C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC94F6C50815BFAAE" box="[590,624,1292,1318]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">3C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC97A6C50818BFAAE" box="[635,672,1292,1318]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4C</figureCitation>
); only one single lobe of the surstylus is present (
<figureCitation id="1335BCE6FFCB3C6DCFBF6C508638FAAE" box="[1214,1299,1292,1318]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCE1F6C508610FAAE" box="[1310,1339,1292,1318]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCE476C50864FFAAE" box="[1350,1380,1292,1318]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">3F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCE6F6C5086BBFAAE" box="[1390,1424,1292,1318]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4F</figureCitation>
). See
<bibRefCitation id="EF9FDD92FFCB3C6DCBC66C6C8270FAC2" author="Bickel, D. J." box="[199,347,1328,1354]" pageId="1" pageNumber="258" pagination="263 - 270" refId="ref10671" refString="Bickel, D. J. (1986) Australian species of Systenus (Diptera: Dolichopodidae). Records of the Australian Museum, 38, 263 - 270. https: // doi. org / 10.3853 / j. 0067 - 1975.38.1986.350" type="journal article" year="1986">Bickel (1986</bibRefCitation>
,
<bibRefCitation id="EF9FDD92FFCB3C6DCA676C6C828BFAC3" author="Bickel, D. J." box="[358,416,1328,1355]" pageId="1" pageNumber="258" pagination="169 - 182" refId="ref10716" refString="Bickel, D. J. (2015) The Costa Rican Systenus Low (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus. Zootaxa, 4020 (1), 169 - 182. https: // doi. org / 10.11646 / zootaxa. 4020.1.7" type="journal article" year="2015">2015</bibRefCitation>
) for a complete diagnosis.
</paragraph>
</subSubSection>
</treatment>
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343
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<mods:title id="D6BBE7D0CDA41A2AF69AE60384D0A236">First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species</mods:title>
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Key to species of
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<emphasis id="B97A7C71FFCB3C6DCA6F6CC082F3FA3E" bold="true" box="[366,472,1436,1462]" italics="true" pageId="1" pageNumber="258">Systenus</emphasis>
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from
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(males)
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<paragraph id="8BB1A063FFCB3C6DCB966CBD86B7FA7E" blockId="1.[151,1437,1505,2026]" box="[151,1436,1505,1527]" pageId="1" pageNumber="258">
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1 Metepimeron dark (
<emphasis id="B97A7C71FFCB3C6DCAA36CBE82EEFA7F" box="[418,453,1506,1527]" italics="true" pageId="1" pageNumber="258">e.g.</emphasis>
<figureCitation id="1335BCE6FFCB3C6DCACB6CBD8132FA7E" box="[458,537,1505,1527]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9226CBD816FFA7E" box="[547,580,1505,1526]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2A</figureCitation>
); abdominal tergites 1–3 mostly metallic green (
<emphasis id="B97A7C71FFCB3C6DCF146CBE8713FA7F" box="[1045,1080,1506,1527]" italics="true" pageId="1" pageNumber="258">e.g.</emphasis>
<figureCitation id="1335BCE6FFCB3C6DCF3C6CBD87A2FA7E" box="[1085,1161,1505,1527]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1E</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF926CBD8784FA7E" box="[1171,1199,1505,1526]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2E</figureCitation>
).................. 3
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- Metepimeron yellow (
<figureCitation id="1335BCE6FFCB3C6DCAB66CA2812DF99C" box="[439,518,1534,1556]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Figs 5C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9116CA2811BF99C" box="[528,560,1534,1556]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">6A</figureCitation>
); abdominal tergites 1–3 partly yellow (
<figureCitation id="1335BCE6FFCB3C6DC8AE6CA280D5F99C" box="[943,1022,1534,1556]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Figs 5C</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF096CA2870CF99C" box="[1032,1063,1534,1556]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">6A</figureCitation>
)............................. 2
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<keyLead id="30FF0356FFCB3C6DCB966F4086B6F9C7" pageId="1" pageNumber="258">
2 Femur III entirely yellow (
<figureCitation id="1335BCE6FFCB3C6DCAE56F40811BF9B9" box="[484,560,1563,1586]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
); abdominal tergite 3 dorsally metallic green, tergites 4–5 wholly metallic green (
<figureCitation id="1335BCE6FFCB3C6DCE456F4086BBF9B9" box="[1348,1424,1563,1586]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5C</figureCitation>
); surstylus slightly curved ventrally at apex (
<figureCitation id="1335BCE6FFCB3C6DC9816F6581E0F9C7" box="[640,715,1593,1615]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5D</figureCitation>
)............................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFEC6F6586B6F9C7" ID-CoL="544RL" authority="Naglis" authorityName="Naglis" authorityYear="2000" box="[1261,1437,1593,1615]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="flaviatus">
<emphasis id="B97A7C71FFCB3C6DCFEC6F658673F9C6" box="[1261,1368,1593,1614]" italics="true" pageId="1" pageNumber="258">S. flaviatus</emphasis>
Naglis
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966F0B86B6F902" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966F0B86B6F902" pageId="1" pageNumber="258">
- Femur III yellow with dark brown apex (
<figureCitation id="1335BCE6FFCB3C6DC9676F0A8198F9E4" box="[614,691,1622,1644]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6A</figureCitation>
); abdominal tergite 3 mostly yellow (
<figureCitation id="1335BCE6FFCB3C6DCF166F0A8748F9E4" box="[1047,1123,1622,1644]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6A</figureCitation>
), with T-shaped macula dorsally, tergites 4–5 dark brown; surstylus abruptly curved ventrally at apex (
<figureCitation id="1335BCE6FFCB3C6DC8796F2880E9F902" box="[888,962,1652,1674]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6B</figureCitation>
).....................
<taxonomicName id="4C0EDBE0FFCB3C6DCFD16F2886B6F902" ID-CoL="9HSKT" authority="Bickel" authorityName="Bickel" authorityYear="2015" box="[1232,1437,1652,1674]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="naranjensis">
<emphasis id="B97A7C71FFCB3C6DCFD16F2887F7F901" box="[1232,1244,1652,1673]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCFE96F288673F901" box="[1256,1368,1652,1673]" italics="true" pageId="1" pageNumber="258">naranjensis</emphasis>
Bickel
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966FCD86B7F94B" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966FCD86B7F92E" blockId="1.[151,1437,1505,2026]" box="[151,1436,1681,1703]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966FCD86B7F92E" box="[151,1436,1681,1703]" pageId="1" pageNumber="258">
3 Antenna wholly dark brown (
<figureCitation id="1335BCE6FFCB3C6DCAFC6FCD814EF92E" box="[509,613,1681,1703]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">Fig. 3A, B</figureCitation>
); femora mostly dark brown (
<figureCitation id="1335BCE6FFCB3C6DC8856FCD80E4F92E" box="[900,975,1681,1703]" captionStart="FIGURE 3" captionStartId="7.[151,250,1787,1811]" captionTargetBox="[162,1426,185,1760]" captionTargetId="figure-19@7.[158,1430,181,1764]" captionTargetPageId="7" captionText="FIGURE 3. Systenus fuscus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849836" httpUri="https://zenodo.org/record/13849836/files/figure.png" pageId="1" pageNumber="258">Fig. 3A</figureCitation>
).......................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF76FCD8661F92E" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1270,1354,1681,1702]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B97A7C71FFCB3C6DCFF76FCD8661F92E" box="[1270,1354,1681,1702]" italics="true" pageId="1" pageNumber="258">S. fuscus</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFCB3C6DCE516FCD86B7F92E" bold="true" box="[1360,1436,1681,1702]" pageId="1" pageNumber="258">
<taxonomicNameLabel id="A249C10AFFCB3C6DCE516FCD86B7F92E" box="[1360,1436,1681,1702]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966FF386B7F94B" blockId="1.[151,1437,1505,2026]" box="[151,1436,1710,1732]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966FF386B7F94B" box="[151,1436,1710,1732]" pageId="1" pageNumber="258">
- Antenna partly orangish yellow (
<figureCitation id="1335BCE6FFCB3C6DC91F6FF28147F94B" box="[542,620,1710,1732]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9776FF281BEF94B" box="[630,661,1710,1731]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC99F6FF28196F94B" box="[670,701,1710,1731]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); femora yellow (
<figureCitation id="1335BCE6FFCB3C6DC8666FF2809CF94B" box="[871,951,1710,1732]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Figs 2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC8C06FF280F4F94B" box="[961,991,1710,1731]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
)................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966F9086B7F894" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966F9086B6F977" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966F9086B6F977" pageId="1" pageNumber="258">
4 It
<subScript id="178AA226FFCB3C6DCBF06F8783D3F96F" attach="left" box="[241,248,1755,1767]" fontSize="5" pageId="1" pageNumber="258">3</subScript>
white (
<figureCitation id="1335BCE6FFCB3C6DCA436F9082A7F969" box="[322,396,1739,1762]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5E</figureCitation>
); wing apex brown: posterior wing membrane reduced near apex, with vein M
<subScript id="178AA226FFCB3C6DCF896F8787A4F96F" attach="left" box="[1160,1167,1755,1767]" fontSize="5" pageId="1" pageNumber="258">1</subScript>
ending on posterior margin (
<figureCitation id="1335BCE6FFCB3C6DCBE86FB5821AF977" box="[233,305,1769,1791]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="1" pageNumber="258">Fig. 5E</figureCitation>
)...........................................................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFC16FB586B6F977" ID-CoL="9HWXR" authority="Bickel" authorityName="Bickel" authorityYear="2015" box="[1216,1437,1769,1791]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B97A7C71FFCB3C6DCFC16FB587E7F976" box="[1216,1228,1769,1790]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCFD96FB58673F976" box="[1240,1368,1769,1790]" italics="true" pageId="1" pageNumber="258">maculipennis</emphasis>
Bickel
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966E5B86B7F894" blockId="1.[151,1437,1505,2026]" box="[151,1436,1798,1826]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E5B86B7F894" box="[151,1436,1798,1826]" pageId="1" pageNumber="258">
- It 3 yellow (
<figureCitation id="1335BCE6FFCB3C6DCA4D6E5A82B7F893" box="[332,412,1798,1820]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Figs 2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCAA76E5A82ECF893" box="[422,455,1798,1819]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
); wing apex unmodified, membrane hyaline or brownish (
<figureCitation id="1335BCE6FFCB3C6DC8F26E5A8768F893" box="[1011,1091,1798,1820]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1D</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF4C6E5A8747F893" box="[1101,1132,1798,1819]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2D</figureCitation>
)....................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30FAB8C6FFCB3C6DCB966E7F86B7F81A" pageId="1" pageNumber="258">
<paragraph id="8BB1A063FFCB3C6DCB966E7F86B6F8DF" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E7F86B6F8DF" pageId="1" pageNumber="258">
5 Postpedicel wholly orangish yellow at basal 1/2 (
<figureCitation id="1335BCE6FFCB3C6DC9CE6E788035F8B2" box="[719,798,1828,1850]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6C</figureCitation>
); abdominal segment 7 forming peduncle shorter than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCA366E1D82A8F8DF" box="[311,387,1857,1879]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6C</figureCitation>
); cercus rectangular, shorter than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DC87A6E1D80EDF8DF" box="[891,966,1857,1879]" captionStart="FIGURE 6" captionStartId="13.[151,250,1342,1366]" captionTargetBox="[155,1431,185,1314]" captionTargetId="figure-19@13.[151,1435,181,1318]" captionTargetPageId="13" captionText="FIGURE 6. A–B. Systenus naranjensis Bickel, 2015, identified male. A. Habitus, lateral view; B. Hypopygium, left lateral view. C–D. Systenus parkeri Bickel, 2015, identified male. C. Habitus, lateral view; D. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849843" httpUri="https://zenodo.org/record/13849843/files/figure.png" pageId="1" pageNumber="258">Fig. 6D</figureCitation>
).........................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF86E1D86B6F8DF" ID-CoL="9HYXF" authority="Bickel" authorityName="Bickel. A" authorityYear="2015" box="[1273,1437,1857,1879]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="parkeri">
<emphasis id="B97A7C71FFCB3C6DCFF86E1D862EF8DE" box="[1273,1285,1857,1878]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCE106E1D8673F8DE" box="[1297,1368,1857,1878]" italics="true" pageId="1" pageNumber="258">parkeri</emphasis>
Bickel
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</paragraph>
<paragraph id="8BB1A063FFCB3C6DCB966E0386B7F81A" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6DCB966E0386B7F81A" pageId="1" pageNumber="258">
- Postpedicel partly brownish at basal 1/2 (
<figureCitation id="1335BCE6FFCB3C6DC97F6E0281E4F8FB" box="[638,719,1886,1908]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC9DD6E0281D1F8FB" box="[732,762,1886,1907]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2B</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DC8066E02800DF8FB" box="[775,806,1886,1907]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4B</figureCitation>
); abdominal segment 7 forming peduncle longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCA366E2082ADF819" box="[311,390,1916,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCA916E20829BF819" box="[400,432,1916,1937]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2A</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCABB6E2082F0F819" box="[442,475,1916,1937]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4A</figureCitation>
); cercus digitiform, as long or longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCF296E20875AF819" box="[1064,1137,1916,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Figs 1F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF7D6E2087BEF819" box="[1148,1173,1916,1937]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">2F</figureCitation>
,
<figureCitation id="1335BCE6FFCB3C6DCF9E6E208792F819" box="[1183,1209,1916,1937]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="1" pageNumber="258">4F</figureCitation>
)................. 6
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<paragraph id="8BB1A063FFCB3C6DCB966EC586B7F843" blockId="1.[151,1437,1505,2026]" pageId="1" pageNumber="258">
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6 Palpus mostly brown (
<figureCitation id="1335BCE6FFCB3C6DCABF6EC58121F826" box="[446,522,1945,1967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1B</figureCitation>
); wing membrane light brown (
<figureCitation id="1335BCE6FFCB3C6DC8406EC580A4F826" box="[833,911,1945,1967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1D</figureCitation>
); cercus about 2X longer than epandrial capsule (
<figureCitation id="1335BCE6FFCB3C6DCE746EC583D7F843" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="1" pageNumber="258">Fig. 1F</figureCitation>
)....................................................................................
<taxonomicName id="4C0EDBE0FFCB3C6DCFF36EEB8661F843" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1266,1354,1974,1996]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
<emphasis id="B97A7C71FFCB3C6DCFF36EEB87D5F844" box="[1266,1278,1975,1996]" italics="true" pageId="1" pageNumber="258">S</emphasis>
.
<emphasis id="B97A7C71FFCB3C6DCE0B6EEA8661F843" box="[1290,1354,1974,1995]" italics="true" pageId="1" pageNumber="258">bickeli</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFCB3C6DCE516EEA86B7F843" bold="true" box="[1360,1436,1974,1995]" pageId="1" pageNumber="258">
<taxonomicNameLabel id="A249C10AFFCB3C6DCE516EEA86B7F843" box="[1360,1436,1974,1995]" pageId="1" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFCB3C6ECB966E8886B7FF25" blockId="1.[151,1437,1505,2026]" lastBlockId="2.[151,1437,151,379]" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258">
<keyLead id="30FF0356FFCB3C6ECB966E8886B7FF25" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258">
- Palpus yellow (
<figureCitation id="1335BCE6FFCB3C6DCA7D6E8882E7F861" box="[380,460,2004,2026]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Fig. 2B</figureCitation>
); wing membrane hyaline (
<figureCitation id="1335BCE6FFCB3C6DC9DF6E888005F861" box="[734,814,2004,2026]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="1" pageNumber="258">Fig. 2D</figureCitation>
); cercus as long or only slightly longer than epandrial capsule (
<figureCitation id="1335BCE6FFC83C6ECBE869CB821EFF25" box="[233,309,151,173]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Figs 2F</figureCitation>
,
<figureCitation id="1335BCE6FFC83C6ECA3E69C48272FF25" box="[319,345,152,173]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">4F</figureCitation>
)......................................................................................... 7
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<paragraph id="8BB1A063FFC83C6ECB9669E986B6FF60" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
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7 Surstylus longer than cercus and epandrial capsule, ventral edge nearly straight, only slightly narrowed and curved ventrally at apex (
<figureCitation id="1335BCE6FFC83C6ECA1D698E824DFF60" box="[284,358,210,232]" captionStart="FIGURE 5" captionStartId="11.[151,250,1904,1928]" captionTargetBox="[181,1406,312,1876]" captionTargetId="figure-81@11.[177,1410,308,1880]" captionTargetPageId="11" captionText="FIGURE 5. A–B. Systenus baetae Naglis, 2000, identified male. A. Habitus, lateral view, arrow indicating metepimeron; B. Hypopygium, left lateral view. C–D. Systenus flaviatus Naglis, 2000, identified Male. C. Habitus, lateral view; D. Hypopygium, left lateral view. Systenus maculipennis Bickel, 2015, identified male. E. Habitus, lateral view, arrow indicating the tarsus white; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849840" httpUri="https://zenodo.org/record/13849840/files/figure.png" pageId="2" pageNumber="259">Fig. 5B</figureCitation>
)............................................................................
<taxonomicName id="4C0EDBE0FFC83C6ECE01698E86B6FF60" ID-CoL="544RF" authority="Naglis" authorityName="Naglis" authorityYear="2000" box="[1280,1437,210,232]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="beatae">
<emphasis id="B97A7C71FFC83C6ECE01698E8627FF6F" box="[1280,1292,210,231]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECE19698E8673FF6F" box="[1304,1368,210,231]" italics="true" pageId="2" pageNumber="259">beatae</emphasis>
Naglis
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFC83C6ECB9669AC86B7FE8D" blockId="2.[151,1437,151,379]" box="[151,1436,239,261]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB9669AC86B7FE8D" box="[151,1436,239,261]" pageId="2" pageNumber="259">
- Surstylus shorter or as long as cercus and epandrial capsule and abruptly narrowed towards apex (
<figureCitation id="1335BCE6FFC83C6ECF8C69B387FCFE8D" box="[1165,1239,239,261]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Figs 2F</figureCitation>
,
<figureCitation id="1335BCE6FFC83C6ECFE069AC87D0FE8D" box="[1249,1275,240,261]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">4F</figureCitation>
)........... 8
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<keyStep id="30FAB8C6FFC83C6ECB96685186B7FEF2" pageId="2" pageNumber="259">
<paragraph id="8BB1A063FFC83C6ECB96685186B7FEB7" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB96685186B7FEB7" pageId="2" pageNumber="259">
8 Coxa III mostly yellow (
<figureCitation id="1335BCE6FFC83C6ECACD68518132FEAA" box="[460,537,269,291]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">Fig. 4A</figureCitation>
); surstylus as long as cercus, wide, abruptly narrowed at apex, with 1 basodorsal acute process (
<figureCitation id="1335BCE6FFC83C6ECBE868768260FEB7" box="[233,331,298,320]" captionStart="FIGURE 4" captionStartId="9.[151,250,1696,1720]" captionTargetBox="[162,1426,185,1668]" captionTargetId="figure-19@9.[158,1430,181,1672]" captionTargetPageId="9" captionText="FIGURE 4. Systenus grossus sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Left wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view; G. Hypopygium, ventral view, arrows indicating the digitiform processes on inner surface of surstylus." figureDoi="http://doi.org/10.5281/zenodo.13849838" httpUri="https://zenodo.org/record/13849838/files/figure.png" pageId="2" pageNumber="259">Fig. 4F, G</figureCitation>
)............................................................................
<taxonomicName id="4C0EDBE0FFC83C6ECFE868768662FEC8" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1257,1353,298,320]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="grossus" status="sp. nov.">
<emphasis id="B97A7C71FFC83C6ECFE8687687DEFEB7" box="[1257,1269,298,319]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECE0068778662FEC8" box="[1281,1353,299,320]" italics="true" pageId="2" pageNumber="259">grossus</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFC83C6ECE51687686B7FEB7" bold="true" box="[1360,1436,298,319]" pageId="2" pageNumber="259">
<taxonomicNameLabel id="A249C10AFFC83C6ECE51687686B7FEB7" box="[1360,1436,298,319]" pageId="2" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB1A063FFC83C6ECB96681486B7FEF2" blockId="2.[151,1437,151,379]" pageId="2" pageNumber="259">
<keyLead id="30FF0356FFC83C6ECB96681486B7FEF2" pageId="2" pageNumber="259">
- Coxa III mostly brown (
<figureCitation id="1335BCE6FFC83C6ECAD2681B8108FED5" box="[467,547,327,349]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Fig. 2A</figureCitation>
); surstylus shorter than cercus, narrow, slightly narrowed at apex, with 1 basodorsal acute process and 1 apicoventral acute process (
<figureCitation id="1335BCE6FFC83C6EC97468398197FEF2" box="[629,700,357,379]" captionStart="FIGURE 2" captionStartId="5.[151,250,1718,1742]" captionTargetBox="[155,1431,185,1690]" captionTargetId="figure-19@5.[151,1435,181,1694]" captionTargetPageId="5" captionText="FIGURE 2. Systenus colombiensis sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.13849832" httpUri="https://zenodo.org/record/13849832/files/figure.png" pageId="2" pageNumber="259">Fig. 2F</figureCitation>
).........................................
<taxonomicName id="4C0EDBE0FFC83C6ECFB568398662FEF2" authorityName="Ramos-Pastrana &amp; Merchan-Vargas &amp; Soares" authorityYear="2024" box="[1204,1353,357,378]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="colombiensis" status="sp. nov.">
<emphasis id="B97A7C71FFC83C6ECFB5683987EBFEF2" box="[1204,1216,357,378]" italics="true" pageId="2" pageNumber="259">S</emphasis>
.
<emphasis id="B97A7C71FFC83C6ECFCD68398662FEF2" box="[1228,1353,357,378]" italics="true" pageId="2" pageNumber="259">colombiensis</emphasis>
</taxonomicName>
<emphasis id="B97A7C71FFC83C6ECE51683986B7FEF2" bold="true" box="[1360,1436,357,378]" pageId="2" pageNumber="259">
<taxonomicNameLabel id="A249C10AFFC83C6ECE51683986B7FEF2" box="[1360,1436,357,378]" pageId="2" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

199
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<mods:title id="FE9CC77BEF97342B6169C724227ACCC9">Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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Subgenus
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<emphasis id="B966EAAE0B416370FE8822B6FE0DFF25" bold="true" box="[279,439,151,178]" italics="true" pageId="2" pageNumber="234">Cladoconnus</emphasis>
Reitter
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<paragraph id="8BAD36BC0B416370FF0822FDFE6EFE85" blockId="2.[151,1436,219,274]" pageId="2" pageNumber="234">
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<treatmentCitation id="0AB310AD0B416370FF0822FDFE62FF64" author="Reitter, E." box="[151,472,219,244]" page="226" pageId="2" pageNumber="234" year="1909">
<taxonomicName id="4C124D3F0B416370FF0822FDFE62FF64" authority="Reitter, 1909: 226" authorityName="Reitter" authorityYear="1909" box="[151,472,219,244]" pageId="2" pageNumber="254" rank="subGenus" subGenus="Cladoconnus">
<emphasis id="B966EAAE0B416370FF0822FDFEA7FF63" box="[151,285,220,244]" italics="true" pageId="2" pageNumber="234">Cladoconnus</emphasis>
<bibRefCitation id="EF834B4D0B416370FEBD22FDFE62FF64" author="Reitter, E." box="[290,472,219,244]" pageId="2" pageNumber="234" pagination="41 - 80" refId="ref15899" refString="Reitter, E. (1909) Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. II. Band. K. G. Lutz, Stuttgart, 392 pp., pls. 41 - 80." type="book chapter" year="1909">Reitter, 1909: 226</bibRefCitation>
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(as subgenus of
<taxonomicName id="4C124D3F0B416370FDE022FDFD59FF64" ID-CoL="8VRNT" box="[639,739,220,243]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B416370FDE022FDFD59FF64" box="[639,739,220,243]" italics="true" pageId="2" pageNumber="234">Euconnus</emphasis>
</taxonomicName>
).
<typeStatus id="54A9881E0B416370FD6822FDFC93FF63" box="[759,809,220,244]" pageId="2" pageNumber="234">Type</typeStatus>
species:
<taxonomicName id="4C124D3F0B416370FC1B22FDFAB2FF63" ID-CoL="4W44V" authority="Sturm, 1838" authorityName="Sturm" authorityYear="1838" box="[900,1288,220,244]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="234" phylum="Arthropoda" rank="species" species="motschulskii">
<emphasis id="B966EAAE0B416370FC1B22FDFB3CFF63" box="[900,1158,220,244]" italics="true" pageId="2" pageNumber="234">Scydmaenus motschulskii</emphasis>
<bibRefCitation id="EF834B4D0B416370FB1422FDFAB2FF63" author="Sturm, J." box="[1163,1288,220,244]" pageId="2" pageNumber="234" pagination="259 - 271" refId="ref15965" refString="Sturm, J. (1838) Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. V. Abtheilung. Die Insecten. Dreizehntes Bandchen. Kafer. J. Sturm, Nurnberg, 128 pp., pls. 259 - 271." type="book chapter" year="1838">Sturm, 1838</bibRefCitation>
</taxonomicName>
(des. by Franz in
<bibRefCitation id="EF834B4D0B416370FF7E22DAFE73FE85" author="Newton, A. F. &amp; Franz, H." box="[225,457,250,274]" pageId="2" pageNumber="234" pagination="137 - 165" refId="ref15865" refString="Newton, A. F. &amp; Franz, H. (1998) World catalog of the genera of Scydmaenidae (Coleoptera). Koleopterologische Rundschau, 68, 137 - 165." type="journal article" year="1998">Newton &amp; Franz 1998</bibRefCitation>
).
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</paragraph>
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<subSubSection id="C30865370B416370FF082360FDB1FE7C" pageId="2" pageNumber="234" type="discussion">
<paragraph id="8BAD36BC0B416370FF082360FDB1FE7C" blockId="2.[151,1436,320,743]" pageId="2" pageNumber="234">
<emphasis id="B966EAAE0B416370FF082360FEB1FECC" bold="true" box="[151,267,321,347]" pageId="2" pageNumber="234">Remarks.</emphasis>
Morphological characters were illustrated and the diagnosis emended in
<bibRefCitation id="EF834B4D0B416370FBA12361FA98FECD" author="Jaloszynski, P." box="[1086,1314,320,346]" pageId="2" pageNumber="234" pagination="369 - 380" refId="ref14873" refString="Jaloszynski, P. (2018 b) Taxonomy of &quot; Euconnus complex &quot;. Part XVII. Status of subgenera defined by male antennal characters: Androconnus Franz and Cladoconnus Reitter (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4414 (2), 369 - 380. https: // doi. org / 10.11646 / zootaxa. 4415.2.7" type="journal article" year="2018">Jałoszyński (2018b)</bibRefCitation>
; species of the Balkans,
<collectingCountry id="F305762C0B416370FEB52345FEC0FEE9" box="[298,378,356,382]" name="Turkey" pageId="2" pageNumber="234">Turkey</collectingCountry>
and Caucasus were revised in
<bibRefCitation id="EF834B4D0B416370FD452345FB9BFEE8" author="Hlavac, P. &amp; Stevanovic, M." box="[730,1057,356,383]" pageId="2" pageNumber="234" pagination="401 - 25" refId="ref13557" refString="Hlavac, P. &amp; Stevanovic, M. (2013) A review of the subgenus Cladoconnus Reitter of the genus Euconnus Thomson (Coleoptera: Staphylinidae: Scydmaeninae) from the Balkan Peninsula, Turkey and Caucasus. Zootaxa, 3646 (4), 401 - 25. https: // doi. org / 10.11646 / zootaxa. 3646.4.5" type="journal article" year="2013">Hlaváč &amp; Stevanović (2013)</bibRefCitation>
, Eastern Palaearctic species were treated by
<bibRefCitation id="EF834B4D0B416370FE8D23A9FE7CFE34" author="Hoshina, H." box="[274,454,392,419]" pageId="2" pageNumber="234" pagination="119 - 123" refId="ref13620" refString="Hoshina, H. (2004) First new species of the subgenus Cladoconnus of the genus Euconnus from Japan (Coleoptera: Scydmaenidae). Entomological Problems, 34 (1 - 2), 119 - 123." type="journal article" year="2004">Hoshina (2004)</bibRefCitation>
,
<bibRefCitation id="EF834B4D0B416370FE4B23A9FD59FE34" author="Hoshina, H. &amp; Park, S. - J." box="[468,739,392,419]" pageId="2" pageNumber="234" pagination="93 - 94" refId="ref13713" refString="Hoshina, H. &amp; Park, S. - J. (2020) A new species, Euconnus (Cladoconnus) odaesanensis sp. nov. (Coleoptera: Staphylinidae: Scydmaeninae), from Korea. Japanese Journal of Systematic Entomology, 26 (1), 93 - 94." type="journal article" year="2020">Hoshina &amp; Park (2020)</bibRefCitation>
,
<bibRefCitation id="EF834B4D0B416370FD6E23A9FC6DFE35" author="Jaloszynski, P." box="[753,983,392,418]" pageId="2" pageNumber="234" pagination="481 - 488" refId="ref14997" refString="Jaloszynski, P. (2019 b) The first Korean species of Euconnus (Cladoconnus), with a synopsis of the E. ussuriensis species group (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4615 (3), 481 - 488. https: // doi. org / 10.11646 / zootaxa. 4615.3.4" type="journal article" year="2019">Jałoszyński (2019b)</bibRefCitation>
, and
<bibRefCitation id="EF834B4D0B416370FB8923A9FB6CFE34" author="Kurbatov, S." box="[1046,1238,392,419]" pageId="2" pageNumber="234" pagination="1742 - 1745" refId="ref15652" refString="Kurbatov, S. (1988) On the study of beetles from the family Scydmaenidae (Coleoptera) in Far East of the USSR. Zoologicheskii Zhournal, 67 (11), 1742 - 1745." type="journal article" year="1988">Kurbatov (1988)</bibRefCitation>
, and recently the North American species were studied by
<bibRefCitation id="EF834B4D0B416370FDFD238DFCADFE50" author="Caterino, M. S." box="[610,791,428,455]" pageId="2" pageNumber="234" pagination="133 - 175" refId="ref13421" refString="Caterino, M. S. (2022) First report of the Euconnus Thomson subgenus Cladoconnus Reitter in the New World, represented by thirteen new Appalachian species (Coleoptera, Staphylinidae, Scydmaeninae). ZooKeys, 1137, 133 - 175. https: // doi. org / 10.3897 / zookeys. 1137.97068" type="journal article" year="2022">Caterino (2022)</bibRefCitation>
. One extinct species is known from Eocene Rovno amber (
<bibRefCitation id="EF834B4D0B416370FF0023F1FE45FE7C" author="Jaloszynski, P. &amp; Perkovsky, E. E." box="[159,511,464,491]" pageId="2" pageNumber="234" pagination="395 - 400" refId="ref15593" refString="Jaloszynski, P. &amp; Perkovsky, E. E. (2021) A new bizarre species of Euconnus (Cladoconnus) in Upper Eocene Rovno amber (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 5004 (2), 395 - 400. https: // doi. org / 10.11646 / zootaxa. 5004.2.8" type="journal article" year="2021">Jałoszyński &amp; Perkovsky 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30865370B416370FF5823D5FCE5FD71" pageId="2" pageNumber="234" type="distribution">
<paragraph id="8BAD36BC0B416370FF5823D5FDFAFD54" blockId="2.[151,1436,320,743]" pageId="2" pageNumber="234">
<emphasis id="B966EAAE0B416370FF5823D5FDC3FD99" bold="true" box="[199,633,500,527]" pageId="2" pageNumber="234">Distribution of extant species. Asia:</emphasis>
<collectingCountry id="F305762C0B416370FD1D23D4FD78FD98" box="[642,706,501,527]" name="Japan" pageId="2" pageNumber="234">Japan</collectingCountry>
(Honshu, Shikoku), Russian Far East (Primorie),
<collectingCountry id="F305762C0B416370FA9923D5FA22FD98" box="[1286,1432,500,527]" name="South Korea" pageId="2" pageNumber="234">South Korea</collectingCountry>
,
<collectingCountry id="F305762C0B416370FF082039FF51FDA5" box="[151,235,536,562]" name="Turkey" pageId="2" pageNumber="234">Turkey</collectingCountry>
;
<emphasis id="B966EAAE0B416370FF652038FEE1FDA4" bold="true" box="[250,347,537,563]" pageId="2" pageNumber="234">Europe:</emphasis>
<collectingCountry id="F305762C0B416370FEF92039FE78FDA5" box="[358,450,536,562]" name="Albania" pageId="2" pageNumber="234">Albania</collectingCountry>
,
<collectingCountry id="F305762C0B416370FE502039FD8EFDA5" box="[463,564,536,562]" name="Armenia" pageId="2" pageNumber="234">Armenia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FDDE2039FD2CFDA5" box="[577,662,536,562]" name="Austria" pageId="2" pageNumber="234">Austria</collectingCountry>
,
<collectingCountry id="F305762C0B416370FD392039FCB3FDA5" box="[678,777,536,562]" name="Belgium" pageId="2" pageNumber="234">Belgium</collectingCountry>
,
<collectingCountry id="F305762C0B416370FC872039FB99FDA5" box="[792,1059,536,563]" name="Bosnia and Herzegovina" pageId="2" pageNumber="234">Bosnia &amp; Herzegovina</collectingCountry>
,
<collectingCountry id="F305762C0B416370FBAD2039FB3DFDA5" box="[1074,1159,536,562]" name="Croatia" pageId="2" pageNumber="234">Croatia</collectingCountry>
, Czech Rep.,
<collectingCountry id="F305762C0B416370FAB22039FA22FDA5" box="[1325,1432,536,562]" name="Denmark" pageId="2" pageNumber="234">Denmark</collectingCountry>
,
<collectingCountry id="F305762C0B416370FF08201DFF54FDC1" box="[151,238,572,598]" name="Estonia" pageId="2" pageNumber="234">Estonia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FF67201CFEFEFDC0" box="[248,324,573,599]" name="France" pageId="2" pageNumber="234">France</collectingCountry>
(incl.
<collectingCountry id="F305762C0B416370FE14201DFE5FFDC1" box="[395,485,572,598]" name="France" pageId="2" pageNumber="234">Corsica</collectingCountry>
),
<collectingCountry id="F305762C0B416370FE68201CFDDAFDC0" box="[503,608,573,599]" name="Germany" pageId="2" pageNumber="234">Germany</collectingCountry>
,
<collectingCountry id="F305762C0B416370FDF5201CFD45FDC1" box="[618,767,572,599]" name="United Kingdom" pageId="2" pageNumber="234">Great Britain</collectingCountry>
, Georgia,
<collectingCountry id="F305762C0B416370FCEF201CFC05FDC0" box="[880,959,573,599]" name="Greece" pageId="2" pageNumber="234">Greece</collectingCountry>
(incl. Rhodos),
<collectingCountry id="F305762C0B416370FBED201CFB6FFDC0" box="[1138,1237,573,599]" name="Hungary" pageId="2" pageNumber="234">Hungary</collectingCountry>
,
<collectingCountry id="F305762C0B416370FB40201DFAA8FDC1" box="[1247,1298,572,598]" name="Italy" pageId="2" pageNumber="234">Italy</collectingCountry>
(incl. Elba),
<collectingCountry id="F305762C0B416370FF082041FF5BFDED" box="[151,225,608,634]" name="Latvia" pageId="2" pageNumber="234">Latvia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FF6E2041FEE5FDED" box="[241,351,608,634]" name="Lithuania" pageId="2" pageNumber="234">Lithuania</collectingCountry>
,
<collectingCountry id="F305762C0B416370FEF02041FE54FDED" box="[367,494,608,634]" name="Macedonia" pageId="2" pageNumber="234">Macedonia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FE612040FD30FDEC" box="[510,650,609,635]" name="Serbia" pageId="2" pageNumber="234">Montenegro</collectingCountry>
,
<collectingCountry id="F305762C0B416370FD062041FC99FDED" box="[665,803,608,634]" name="Netherlands" pageId="2" pageNumber="234">Netherlands</collectingCountry>
,
<collectingCountry id="F305762C0B416370FCAC2041FC3EFDED" box="[819,900,608,634]" name="Poland" pageId="2" pageNumber="234">Poland</collectingCountry>
,
<collectingCountry id="F305762C0B416370FC0C2041FC4FFDED" box="[915,1013,608,634]" name="Portugal" pageId="2" pageNumber="234">Portugal</collectingCountry>
,
<collectingCountry id="F305762C0B416370FB9B2041FBD6FDED" box="[1028,1132,608,634]" name="Romania" pageId="2" pageNumber="234">Romania</collectingCountry>
,
<collectingCountry id="F305762C0B416370FBE32041FB70FDED" box="[1148,1226,608,634]" name="Russia" pageId="2" pageNumber="234">Russia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FB462041FA99FDED" box="[1241,1315,608,634]" name="Serbia" pageId="2" pageNumber="234">Serbia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FAAC2041FA22FDED" box="[1331,1432,608,634]" name="Slovakia" pageId="2" pageNumber="234">Slovakia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FF0820A5FF46FD09" box="[151,252,644,670]" name="Slovenia" pageId="2" pageNumber="234">Slovenia</collectingCountry>
,
<collectingCountry id="F305762C0B416370FE9520A5FEF7FD09" box="[266,333,644,670]" name="Spain" pageId="2" pageNumber="234">Spain</collectingCountry>
,
<collectingCountry id="F305762C0B416370FEC420A5FE0CFD09" box="[347,438,644,670]" name="Sweden" pageId="2" pageNumber="234">Sweden</collectingCountry>
,
<collectingCountry id="F305762C0B416370FE5B20A5FDF7FD09" box="[452,589,644,670]" name="Switzerland" pageId="2" pageNumber="234">Switzerland</collectingCountry>
,
<collectingCountry id="F305762C0B416370FDC420A5FD02FD09" box="[603,696,644,670]" name="Ukraine" pageId="2" pageNumber="234">Ukraine</collectingCountry>
, ‘Yugoslavia’;
<emphasis id="B966EAAE0B416370FCF820A4FB9DFD09" bold="true" box="[871,1063,644,671]" pageId="2" pageNumber="234">North America:</emphasis>
<collectingCountry id="F305762C0B416370FBAE20A4FBDDFD08" box="[1073,1127,645,671]" name="United States of America" pageId="2" pageNumber="234">USA</collectingCountry>
(
<collectingRegion id="49D6F85E0B416370FBE620A5FB59FD09" box="[1145,1251,644,670]" country="United States of America" name="Alabama" pageId="2" pageNumber="234">Alabama</collectingRegion>
,
<collectingRegion id="49D6F85E0B416370FB6F20A5FAF7FD09" box="[1264,1357,644,670]" country="United States of America" name="Georgia" pageId="2" pageNumber="234">Georgia</collectingRegion>
,
<collectingRegion id="49D6F85E0B416370FAC420A5FF40FD55" country="United States of America" name="North Carolina" pageId="2" pageNumber="234">North Carolina</collectingRegion>
,
<collectingRegion id="49D6F85E0B416370FE992089FE0BFD55" box="[262,433,680,706]" country="United States of America" name="South Carolina" pageId="2" pageNumber="234">South Carolina</collectingRegion>
,
<collectingRegion id="49D6F85E0B416370FE232088FD8EFD54" box="[444,564,681,707]" country="United States of America" name="Tennessee" pageId="2" pageNumber="234">Tennessee</collectingRegion>
).
</paragraph>
<paragraph id="8BAD36BC0B416370FF5820EDFCE5FD71" blockId="2.[151,1436,320,743]" box="[199,863,716,743]" pageId="2" pageNumber="234">
<emphasis id="B966EAAE0B416370FF5820EDFEDDFD71" bold="true" box="[199,359,716,742]" pageId="2" pageNumber="234">Composition.</emphasis>
51 species (incl. 1 extinct) and 3 subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>

116
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<mods:title id="4B60E15F306417927C4E1433F100C70C">Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<subSubSection id="C30865370B426373FF0827C5FE5CFA69" box="[151,486,1508,1534]" pageId="1" pageNumber="233" type="nomenclature">
<paragraph id="8BAD36BC0B426373FF0827C5FE5CFA69" blockId="1.[151,486,1508,1534]" box="[151,486,1508,1534]" pageId="1" pageNumber="233">
<heading id="D0E581D00B426373FF0827C5FE5CFA69" bold="true" box="[151,486,1508,1534]" fontSize="11" level="1" pageId="1" pageNumber="233" reason="1">
<emphasis id="B966EAAE0B426373FF0827C5FE5CFA69" bold="true" box="[151,486,1508,1534]" pageId="1" pageNumber="233">
Genus
<taxonomicName id="4C124D3F0B426373FF7127C5FE5CFA69" ID-CoL="8VRNT" authority="Thomson" authorityName="Thomson" box="[238,486,1508,1534]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FF7127C5FED0FA69" bold="true" box="[238,362,1508,1534]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
Thomson
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30865370B426373FF082408FE74F9D7" box="[151,462,1576,1600]" pageId="1" pageNumber="233" type="reference_group">
<paragraph id="8BAD36BC0B426373FF082408FE74F9D7" blockId="1.[151,1074,1576,1601]" box="[151,462,1576,1600]" pageId="1" pageNumber="233">
<treatmentCitationGroup id="AB0211920B426373FF082408FE74F9D7" box="[151,462,1576,1600]" pageId="1" pageNumber="233">
<treatmentCitation id="0AB310AD0B426373FF082408FE70F9D7" author="Thomson, C. G." box="[151,458,1576,1600]" page="61" pageId="1" pageNumber="233" year="1859">
<taxonomicName id="4C124D3F0B426373FF082408FE70F9D7" ID-CoL="8VRNT" authority="Thomson, 1859: 61" authorityName="Thomson" authorityPageNumber="61" authorityYear="1859" box="[151,458,1576,1600]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FF082408FF41F9D7" box="[151,251,1577,1600]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
<bibRefCitation id="EF834B4D0B426373FE9E2409FE70F9D7" author="Thomson, C. G." box="[257,458,1576,1600]" pageId="1" pageNumber="233" refId="ref16010" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. 1. Berlingska Boktryckeriet, Lund, 290 pp. https: // doi. org / 10.5962 / bhl. title. 138677" type="book" year="1859">Thomson, 1859: 61</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C30865370B426373FE4C2408FB88F9D7" box="[467,1074,1576,1601]" pageId="1" pageNumber="233" type="type_taxon">
<paragraph id="8BAD36BC0B426373FE4C2408FB88F9D7" blockId="1.[151,1074,1576,1601]" box="[467,1074,1576,1601]" pageId="1" pageNumber="233">
<typeStatus id="54A9881E0B426373FE4C2408FDBFF9D6" box="[467,517,1577,1601]" pageId="1" pageNumber="233">Type</typeStatus>
species:
<taxonomicName id="4C124D3F0B426373FDFC2409FC0CF9D6" authority="Illiger, 1798" authorityName="Illiger" authorityYear="1798" box="[611,950,1576,1601]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B966EAAE0B426373FDFC2409FC8EF9D7" box="[611,820,1576,1600]" italics="true" pageId="1" pageNumber="233">Pselaphus hirticollis</emphasis>
<bibRefCitation id="EF834B4D0B426373FCA52409FC0CF9D6" author="Illiger, J. K. W." box="[826,950,1576,1601]" pageId="1" pageNumber="233" refId="ref13767" refString="Illiger, J. K. W. (1798) Verzeichniss der Kafer Preussens. Entworfen von Johann Gottlieb Kugelann, Apotheker in Osterode. Ausgearbeitet von Johann Karl Wilhelm Illiger. Mit einer Vorrede des Professors und Pagenhofmeisters Hellwig in Braunschweig und dem angehangten Versuche einer naturlichen Ordnungs- und Gattungs- Folge der Insekten. J. J. Gebauer, Halle, xlii + 510 + [1] pp." type="journal volume" year="1798">Illiger, 1798</bibRefCitation>
</taxonomicName>
(des. orig.).
</paragraph>
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<subSubSection id="C30865370B426373FF08244EFAA0F866" pageId="1" pageNumber="233" type="discussion">
<paragraph id="8BAD36BC0B426373FF08244EFCF4F962" blockId="1.[151,1437,1647,2033]" pageId="1" pageNumber="233">
<emphasis id="B966EAAE0B426373FF08244EFEB1F91E" bold="true" box="[151,267,1647,1673]" pageId="1" pageNumber="233">Remarks.</emphasis>
The subgeneric system of
<taxonomicName id="4C124D3F0B426373FDDD244EFD0AF91F" box="[578,688,1647,1672]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FDDD244EFD0AF91F" box="[578,688,1647,1672]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
</taxonomicName>
was a subject of a long series of publications (
<bibRefCitation id="EF834B4D0B426373FB4E244EFA22F91E" author="Jaloszynski, P." box="[1233,1432,1647,1673]" pageId="1" pageNumber="233" pagination="55 - 82" refId="ref13863" refString="Jaloszynski, P. (2012) Taxonomy of &quot; Euconnus complex &quot;. Part I. Morphology of Euconnus s. str. and revision of Euconnomorphus Franz and Venezolanoconnus Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3555 (1), 55 - 82. https: // doi. org / 10.11646 / zootaxa. 3555.1.3" type="journal article" year="2012">Jałoszyński 2012</bibRefCitation>
,
<bibRefCitation id="EF834B4D0B426373FF0824B2FF5AF93A" author="Jaloszynski, P." box="[151,224,1683,1709]" pageId="1" pageNumber="233" pagination="523 - 543" refId="ref13972" refString="Jaloszynski, P. (2013 b) Taxonomy of &quot; Euconnus complex &quot;. Part II. Revision of Archiconnus Franz, Parapseudoconnus Franz and Mexiconnus gen. nov. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3666 (4), 523 - 543. https: // doi. org / 10.11646 / zootaxa. 3666.4.7" type="journal article" year="2013">2013b</bibRefCitation>
,
<bibRefCitation id="EF834B4D0B426373FF7324B3FE89F93A" author="Jaloszynski, P." box="[236,307,1682,1709]" pageId="1" pageNumber="233" pagination="1 - 24" refId="ref14034" refString="Jaloszynski, P. (2015 a) Taxonomy of &quot; Euconnus complex &quot;. Part III. Morphology of Euconnus subgenus Napochus and revision of the Australian species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3925 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 3925.1.1" type="journal article" year="2015">2015a</bibRefCitation>
, b, c, 2016a, b, c, d, 2017a, b, c, d, e, f, 2018a, b, 2019a, 2020a, b, 2021a, b, 2022b,
<bibRefCitation id="EF834B4D0B426373FB6224B2FE8CF946" author="Jaloszynski, P. &amp; Newton, A. F." pageId="1" pageNumber="233" pagination="401 - 443" refId="ref15534" refString="Jaloszynski, P. &amp; Newton, A. F. (2017) Taxonomy of &quot; Euconnus complex &quot;. Part XV. Disentangling the Euconnus-Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4344 (3), 401 - 443. https: // doi. org / 10.11646 / zootaxa. 4344.3.1" type="journal article" year="2017">Jałoszyński &amp; Newton 2017</bibRefCitation>
). Many taxonomic issues have been solved, but the current subgeneric division of
<taxonomicName id="4C124D3F0B426373FB792496FAEEF947" box="[1254,1364,1719,1744]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FB792496FAEEF947" box="[1254,1364,1719,1744]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
</taxonomicName>
is not phylogeny-based and certainly will require profound changes.
</paragraph>
<paragraph id="8BAD36BC0B426373FF5824DEFAA0F866" blockId="1.[151,1437,1647,2033]" pageId="1" pageNumber="233">
<taxonomicName id="4C124D3F0B426373FF5824DEFE8FF88F" box="[199,309,1791,1816]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FF5824DEFE8FF88F" box="[199,309,1791,1816]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
</taxonomicName>
is characterized by the following set of features: eyes (if present) situated closer to mandibular bases and antennal fossae than to occipital constriction; lateral submental sutures lacking; mesoventral intercoxal process present, carinate; notosternal sutures complete; hypomeral ridges at least partly developed or at least inner regions of hypomera delimited from outer regions by abrupt change in microsculpture; each elytron with two asetose basal pits (reduced in some species); metaventral intercoxal process widely or narrowly (but clearly) separating metacoxae; and aedeagus with free rod-like parameres.
<taxonomicName id="4C124D3F0B426373FD0E2592FD45F85B" box="[657,767,1971,1996]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="233" phylum="Arthropoda" rank="genus">
<emphasis id="B966EAAE0B426373FD0E2592FD45F85B" box="[657,767,1971,1996]" italics="true" pageId="1" pageNumber="233">Euconnus</emphasis>
</taxonomicName>
can be identified using a preliminary key (and comments related to the yet unclear diagnosis) to world Stenichnini genera published recently (
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).
</paragraph>
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<mods:title id="89BB2AA21370268237AADC90CC917C59">Subgenera Cladoconnus Reitter and Tetramelus Motschulsky of Euconnus Thomson discovered in China (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<heading id="D0E581D00B466377FF08253BFDFFF8A3" bold="true" box="[151,581,1818,1845]" fontSize="11" level="1" pageId="5" pageNumber="237" reason="1">
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Subgenus
<taxonomicName id="4C124D3F0B466377FE88253BFDFFF8A3" authority="Motschulsky" authorityName="Motschulsky" box="[279,581,1818,1845]" pageId="5" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B466377FE88253BFE27F8A2" bold="true" box="[279,413,1818,1845]" italics="true" pageId="5" pageNumber="237">Tetramelus</emphasis>
Motschulsky
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</paragraph>
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<paragraph id="8BAD36BC0B466377FF082578FEF7F807" blockId="5.[151,1436,1881,1936]" pageId="5" pageNumber="237">
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<treatmentCitationGroup id="AB0211920B466377FF082578FDA9F8E6" box="[151,531,1881,1905]" pageId="5" pageNumber="237">
<treatmentCitation id="0AB310AD0B466377FF082578FDB5F8E6" author="Motschulsky, V." box="[151,527,1881,1905]" page="257" pageId="5" pageNumber="237" year="1869">
<taxonomicName id="4C124D3F0B466377FF082578FDB5F8E6" authority="Motschulsky, 1869: 257" authorityName="Motschulsky" authorityYear="1869" box="[151,527,1881,1905]" pageId="5" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B466377FF082578FEBDF8E6" box="[151,263,1881,1905]" italics="true" pageId="5" pageNumber="237">Tetramelus</emphasis>
<bibRefCitation id="EF834B4D0B466377FE8E2578FDB5F8E6" author="Motschulsky, V." box="[273,527,1881,1905]" pageId="5" pageNumber="237" pagination="252 - 272" refId="ref15819" refString="Motschulsky, V. (1869) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. 7 - ieme Article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42 (1), 252 - 272, pl. 8." type="journal article" year="1869">Motschulsky, 1869: 257</bibRefCitation>
</taxonomicName>
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.
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<typeStatus id="54A9881E0B466377FD83257BFDF4F8E5" box="[540,590,1882,1906]" pageId="5" pageNumber="237">Type</typeStatus>
species:
<taxonomicName id="4C124D3F0B466377FD2D2578FB98F8E5" authority="Sturm, 1838" authorityName="Sturm" authorityYear="1838" box="[690,1058,1881,1906]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="237" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B966EAAE0B466377FD2D2578FC2DF8E6" box="[690,919,1881,1905]" italics="true" pageId="5" pageNumber="237">Scydmaenus oblongus</emphasis>
<bibRefCitation id="EF834B4D0B466377FC3E257BFB98F8E5" author="Sturm, J." box="[929,1058,1882,1906]" pageId="5" pageNumber="237" pagination="259 - 271" refId="ref15965" refString="Sturm, J. (1838) Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. V. Abtheilung. Die Insecten. Dreizehntes Bandchen. Kafer. J. Sturm, Nurnberg, 128 pp., pls. 259 - 271." type="book chapter" year="1838">Sturm, 1838</bibRefCitation>
</taxonomicName>
(des. by Franz in
<treatmentCitation id="0AB310AD0B466377FB74257BFEFBF807" author="Newton, A. F. &amp; Franz, H." page="147" pageId="5" pageNumber="237" year="1998">
<bibRefCitation id="EF834B4D0B466377FB74257BFEB5F807" author="Newton, A. F. &amp; Franz, H." pageId="5" pageNumber="237" pagination="137 - 165" refId="ref15865" refString="Newton, A. F. &amp; Franz, H. (1998) World catalog of the genera of Scydmaenidae (Coleoptera). Koleopterologische Rundschau, 68, 137 - 165." type="journal article" year="1998">Newton &amp; Franz (1998)</bibRefCitation>
: 147
</treatmentCitation>
).
</subSubSection>
</paragraph>
<subSubSection id="C30865370B466374FF082598FC1EFED6" lastPageId="6" lastPageNumber="238" pageId="5" pageNumber="237" type="discussion">
<paragraph id="8BAD36BC0B466377FF082598FB9FF860" blockId="5.[151,1436,1977,2040]" pageId="5" pageNumber="237">
<emphasis id="B966EAAE0B466377FF082598FEB1F844" bold="true" box="[151,267,1977,2003]" pageId="5" pageNumber="237">Remarks.</emphasis>
Morphological structures were illustrated and an emended diagnosis was given in
<bibRefCitation id="EF834B4D0B466377FB2B2598FA2DF843" author="Jaloszynski, P." box="[1204,1431,1977,2004]" pageId="5" pageNumber="237" pagination="280 - 300" refId="ref14148" refString="Jaloszynski, P. (2015 c) Taxonomy of &quot; Euconnus complex &quot;. Part V. Review of type species of Euconnus subgenera Tetramelus, Paratetramelus and Heteroconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4040 (3), 280 - 300. https: // doi. org / 10.11646 / zootaxa. 4040.3.2" type="journal article" year="2015">Jałoszyński (2015c)</bibRefCitation>
; further notes on character variability can be found in
<bibRefCitation id="EF834B4D0B466377FD7325FCFC7EF860" author="Jaloszynski, P." box="[748,964,2013,2039]" pageId="5" pageNumber="237" pagination="421 - 428" refId="ref14441" refString="Jaloszynski, P. (2017 a) Taxonomy of &quot; Euconnus complex &quot;. Part X. Drastophus placed as a junior synonym of Tetramelus, and Smicrophus as a junior synonym of Napochus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4232 (3), 421 - 428. https: // doi. org / 10.11646 / zootaxa. 4232.3.10" type="journal article" year="2017">Jałoszyński (2017a</bibRefCitation>
,
<bibRefCitation id="EF834B4D0B466377FC5025FCFBA2F860" author="Jaloszynski, P." box="[975,1048,2013,2039]" pageId="5" pageNumber="237" pagination="145 - 160" refId="ref15170" refString="Jaloszynski, P. (2021 a) Taxonomy of &quot; Euconnus complex &quot;. Part XXII. Status of subgenera Allonapochus Franz, Dimorphoconnus Franz, Eupentarius Reitter, and Myrmecodytes Croissandeau (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4966 (2), 145 - 160. https: // doi. org / 10.11646 / zootaxa. 4966.2.3" type="journal article" year="2021">2021a</bibRefCitation>
).
</paragraph>
<paragraph id="8BAD36BC0B456374FF5822B6FC1EFED6" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="238">
<taxonomicName id="4C124D3F0B456374FF5822B6FEF8FF26" box="[199,322,151,177]" pageId="6" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B456374FF5822B6FEF8FF26" box="[199,322,151,177]" italics="true" pageId="6" pageNumber="238">Tetramelus</emphasis>
</taxonomicName>
, as currently defined, can be identified using the following set of co-occurring characters: hypostomal ridges running posteromesally; head ventrally lacking deep narrow longitudinal groove connecting gular plate and submentum; pronotum lacking longitudinal hypomeral grooves; pronotum broadest in front of middle; antenna in male unmodified, gradually thickened or with variously distinct tetramerous club; and tarsomere
<quantity id="4CEA9B590B456374FB592322FB4AFE8A" box="[1222,1264,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="238" unit="in" value="4.0">4 in</quantity>
all legs lacking long distoventral lobe (for key to subgenera see
<bibRefCitation id="EF834B4D0B456374FD2C2306FC1AFED6" author="Jaloszynski, P." box="[691,928,295,321]" pageId="6" pageNumber="238" pagination="343 - 391" refId="ref15360" refString="Jaloszynski, P. (2022 b) Taxonomy of &quot; Euconnus complex &quot;. Part XXIV. Intermediate forms between Psomophus, Eupentarius and Euconnus s. str. in the East Palaearctic fauna unify problematic subgenera (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5194 (3), 343 - 391. https: // doi. org / 10.11646 / zootaxa. 5194.3.2" type="journal article" year="2022">Jałoszyński (2022b))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30865370B456374FF58236BFB7DFC16" pageId="6" pageNumber="238" type="distribution">
<paragraph id="8BAD36BC0B456374FF58236BFDFDFDAA" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="238">
<emphasis id="B966EAAE0B456374FF58236BFE01FEF3" bold="true" box="[199,443,330,356]" pageId="6" pageNumber="238">Distribution. Africa:</emphasis>
Algeria, Cameroon, Comoros, Kenya, Lesotho, Madagascar, RSA, Swaziland, Tanzania;
<emphasis id="B966EAAE0B456374FF08234FFF6FFE1F" bold="true" box="[151,213,366,392]" pageId="6" pageNumber="238">Asia:</emphasis>
Asian Turkey, Nagorno-Karabakh, Nepal, “Syria”;
<emphasis id="B966EAAE0B456374FCB9234FFC1AFE1F" bold="true" box="[806,928,366,392]" pageId="6" pageNumber="238">
<collectingCountry id="F305762C0B456374FCB9234FFC20FE1F" box="[806,922,366,392]" name="Australia" pageId="6" pageNumber="238">Australia</collectingCountry>
:
</emphasis>
Australia (incl. Franklin Is., Lord Howe Is., Reevesby Is.,
<collectingRegion id="49D6F85E0B456374FEA923B2FE1CFE3A" box="[310,422,403,429]" country="Australia" name="Tasmania" pageId="6" pageNumber="238">Tasmania</collectingRegion>
);
<emphasis id="B966EAAE0B456374FE2523B3FDF9FE3B" bold="true" box="[442,579,402,428]" pageId="6" pageNumber="238">Caribbean:</emphasis>
Jamaica;
<emphasis id="B966EAAE0B456374FD2A23B2FCADFE3A" bold="true" box="[693,791,403,429]" pageId="6" pageNumber="238">Europe:</emphasis>
Albania, Andorra, Armenia, Austria, Belgium, Bosnia &amp; Herzegovina, Bulgaria, Croatia, Czech Rep., France, Georgia, Germany, Greece, Hungary, Italy, Kosovo, Lithuania, Macedonia, Montenegro, Netherlands, Poland, Portugal, Romania, European Russia, Slovakia, Slovenia, Spain, Switzerland, European Turkey, Ukraine;
<emphasis id="B966EAAE0B456374FDF723DEFC9DFD8F" bold="true" box="[616,807,510,537]" pageId="6" pageNumber="238">North America:</emphasis>
Canada, USA;
<emphasis id="B966EAAE0B456374FC4423DFFB8CFD8F" bold="true" box="[987,1078,510,536]" pageId="6" pageNumber="238">Pacific:</emphasis>
New Caledonia, New Zealand;
<emphasis id="B966EAAE0B456374FF082003FEE9FDAB" bold="true" box="[151,339,546,572]" pageId="6" pageNumber="238">South America:</emphasis>
Bolivia, Brazil, Peru.
</paragraph>
<paragraph id="8BAD36BC0B456374FF582067FC94FD3E" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="238">
<emphasis id="B966EAAE0B456374FF582067FEDDFDF7" bold="true" box="[199,359,582,608]" pageId="6" pageNumber="238">Composition.</emphasis>
At the time of submission of the present study there were 233 species and 2 subspecies. However, in a parallel study on Malagasy species some Stenichnini are transferred into
<taxonomicName id="4C124D3F0B456374FB8B204AFB35FD12" box="[1044,1167,619,645]" pageId="6" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B456374FB8B204AFB35FD12" box="[1044,1167,619,645]" italics="true" pageId="6" pageNumber="238">Tetramelus</emphasis>
</taxonomicName>
(Jałoszyński, in press), which results in 17 more species included in this subgenus.
</paragraph>
<paragraph id="8BAD36BC0B456374FF582092FB7DFC16" blockId="6.[151,1437,151,897]" pageId="6" pageNumber="238">
All new Chinese species described below are microphthalmous, wingless and have reduced wing-related exoskeletal structures, including lack of humeral calli and the elytra strongly narrowing anteriorly; males have a large and deep median impression on the metaventrite, and all species except one have modified trochanters. All species are externally very similar, they differ in body size, pigmentation, modifications of male trochanters, and genital structures. The Chinese
<taxonomicName id="4C124D3F0B456374FDAD2162FD17FCCA" box="[562,685,835,861]" pageId="6" pageNumber="237" rank="subGenus" subGenus="Tetramelus">
<emphasis id="B966EAAE0B456374FDAD2162FD17FCCA" box="[562,685,835,861]" italics="true" pageId="6" pageNumber="238">Tetramelus</emphasis>
</taxonomicName>
species are morphologically so close to their Western Palaearctic congeners that they can be included in the same
<taxonomicName id="4C124D3F0B456374FD2A2146FC28FC16" baseAuthorityName="Sturm" baseAuthorityYear="1838" box="[693,914,871,897]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B966EAAE0B456374FD2A2146FC28FC16" box="[693,914,871,897]" italics="true" pageId="6" pageNumber="238">Euconnus oblongus</emphasis>
</taxonomicName>
group (
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).
</paragraph>
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</treatment>
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