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<document id="F4B111273AE3410A7E9E338C62B4406C" ID-DOI="10.12651/JSR.2013.2.2.099" ID-ISSN="2713-8615" ID-Zenodo-Dep="13144615" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722438108010" checkinUser="felipe" docAuthor="Blakemore, Robert J." docDate="2013" docId="03CA8049FFB9FFE23626FF195C50F85C" docLanguage="en" docName="JSpeciesRes.2.2.99-126.pdf" docOrigin="Journal of Species Research 2 (2)" docSource="http://dx.doi.org/10.12651/jsr.2013.2.2.099" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Pheretima fornicata Gates 1935" docType="treatment" docVersion="2" lastPageNumber="116" masterDocId="FFF3F831FFA9FFF33510FD3B5831FFFC" masterDocTitle="Megascolex (Perichaeta) diffringens Baird, 1869 and Pheretima pingi Stephenson, 1925 types compared to the Amynthas corticis (Kinberg, 1867) and A. carnosus (Goto &amp; Hatai, 1899) species-groups (Oligochaeta: Megadrilacea: Megascolecidae)" masterLastPageNumber="126" masterPageNumber="99" pageNumber="115" updateTime="1722438559216" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0B1B248276250915EB1B0004E5B7ABA6">Megascolex (Perichaeta) diffringens Baird, 1869 and Pheretima pingi Stephenson, 1925 types compared to the Amynthas corticis (Kinberg, 1867) and A. carnosus (Goto &amp; Hatai, 1899) species-groups (Oligochaeta: Megadrilacea: Megascolecidae)</mods:title>
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<mods:namePart id="DA548060EB4FFD05255FEF4237818295">Blakemore, Robert J.</mods:namePart>
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<taxonomicName id="4C634ADCFFB9FFE33626FF195CAFFDC6" authority="Gates, 1935" authorityName="Gates" authorityYear="1935" box="[822,1182,546,570]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="115" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB9FFE33626FF195CAFFDC6" bold="true" box="[822,1182,546,570]" pageId="16" pageNumber="115">
<emphasis id="B917ED4DFFB9FFE33626FF195C25FDC6" bold="true" box="[822,1044,546,570]" italics="true" pageId="16" pageNumber="115">Pheretima fornicata</emphasis>
<bibRefCitation id="EFF24CAEFFB9FFE3310AFF195CAFFDC6" author="Gates, G. E." box="[1050,1182,546,570]" pageId="16" pageNumber="115" pagination="1 - 19" refId="ref15126" refString="Gates, G. E. 1935. New earthworms from China, with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Miscellaneous Collections 93: 1 - 19." type="journal article" year="1935">Gates, 1935</bibRefCitation>
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(
<figureCitation id="13582DDAFFB9FFE3362EFF795BA6FDA7" box="[830,919,578,603]" captionStart="Fig" captionStartId="16.[157,190,1938,1959]" captionTargetBox="[320,1268,1211,1906]" captionTargetId="figure-487@16.[315,1268,1211,1906]" captionTargetPageId="16" captionText="Fig. 12. Amynthas fornicatus showing Gates (1935: figs. 5-6) most uninformative and irrelevant details of spermathecae invoking adage “missing the forest for the trees”." figureDoi="http://doi.org/10.5281/zenodo.13144641" httpUri="https://zenodo.org/record/13144641/files/figure.png" pageId="16" pageNumber="115">Figs. 12</figureCitation>
,
<figureCitation id="13582DDAFFB9FFE336B2FF785B8CFDA7" box="[930,957,579,603]" captionStart="Fig" captionStartId="17.[159,192,1237,1258]" captionTargetBox="[186,731,257,1204]" captionTargetId="figure-704@17.[186,731,257,1204]" captionTargetPageId="17" captionText="Fig. 13. Prior Amynthas hongkongensis after Michaelsen (1910: fig. 22), for comparison." figureDoi="http://doi.org/10.5281/zenodo.13144643" httpUri="https://zenodo.org/record/13144643/files/figure.png" pageId="16" pageNumber="115">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37962D4FFB9FFE33626FF585D12FC60" pageId="16" pageNumber="115" type="reference_group">
<paragraph id="8BDC315FFFB9FFE33626FF585BE5FCA0" blockId="16.[822,1427,546,924]" pageId="16" pageNumber="115">
<treatmentCitationGroup id="AB731671FFB9FFE33626FF585BE5FCA0" pageId="16" pageNumber="115">
<treatmentCitation id="0AC2174EFFB9FFE33626FF585C96FD87" author="Gates, G. E." box="[822,1191,610,635]" page="9" pageId="16" pageNumber="115" year="1935">
<taxonomicName id="4C634ADCFFB9FFE33626FF585C96FD87" authority="Gates, 1935: 9" authorityName="Gates" authorityPageNumber="9" authorityYear="1935" box="[822,1191,610,635]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="115" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB9FFE33626FF585C38FD87" box="[822,1033,611,635]" italics="true" pageId="16" pageNumber="115">Pheretima fornicata</emphasis>
<bibRefCitation id="EFF24CAEFFB9FFE33100FF585C96FD87" author="Gates, G. E." box="[1040,1191,610,635]" pageId="16" pageNumber="115" pagination="1 - 19" refId="ref15126" refString="Gates, G. E. 1935. New earthworms from China, with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Miscellaneous Collections 93: 1 - 19." type="journal article" year="1935">Gates, 1935: 9</bibRefCitation>
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figs. 5-6;
<treatmentCitation id="0AC2174EFFB9FFE33003FF585B44FD67" author="Chen, Y." page="296" pageId="16" pageNumber="115" year="1936">
<bibRefCitation id="EFF24CAEFFB9FFE33003FF585B44FD67" author="Chen, Y." pageId="16" pageNumber="115" pagination="269 - 306" refId="ref14831" refString="Chen, Y. 1936. On the terrestrial Oligochaeta from Szechuan II with the notes on Gates' types. Contribibutions of Biological Laboratory of Science Society of China (Zoology) 11: 269 - 306." type="journal article" year="1936">Chen, 1936: 296</bibRefCitation>
</treatmentCitation>
(re-examination of
<typeStatus id="54D88FFDFFB9FFE33158FFB85C47FD67" box="[1096,1142,643,667]" pageId="16" pageNumber="115">type</typeStatus>
in US National Museum);
<treatmentCitation id="0AC2174EFFB9FFE3365DFF985C33FD47" author="Gates, G. E." box="[845,1026,675,699]" page="434" pageId="16" pageNumber="115" year="1939">
<bibRefCitation id="EFF24CAEFFB9FFE3365DFF985C33FD47" author="Gates, G. E." box="[845,1026,675,699]" pageId="16" pageNumber="115" pagination="405 - 507" refId="ref15162" refString="Gates, G. E. 1939. On some species of Chinese earthworms with special reference to specimens collected in Szechuan by Dr. D. C. Graham. Proceedings of the United States National Museum 85: 405 - 507 [Available from: si-dppr. si. edu / dspace / bitstream / 10088 / 16290 / USNMP- 85 _ 3040 _ 1939. pdf]." type="journal article" year="1939">Gates, 1939: 434</bibRefCitation>
</treatmentCitation>
(changing
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locality to “Tatsienlu, Szechwan” “
<emphasis id="B917ED4DFFB9FFE336C8FFFF5D64FD07" italics="true" pageId="16" pageNumber="115">
From Dr Graham: 3 clitellate specimens labeled Tatsienlu,
<quantity id="4C9B9CBAFFB9FFE33136FFD85C95FD07" box="[1062,1188,739,763]" metricMagnitude="3" metricUnit="m" metricValue="3.6576" pageId="16" pageNumber="115" unit="ft" value="12000.0">12,000 feet</quantity>
,
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</emphasis>
” and adding 1 clitellate specimen in poor condition labeled “
<emphasis id="B917ED4DFFB9FFE33649FE1F5BF0FCA0" italics="true" pageId="16" pageNumber="115">
Between Gin Keo Ho and Dawei,
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feet
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, August 1-2
</emphasis>
”.
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</paragraph>
<paragraph id="8BDC315FFFB9FFE33626FE5F5D12FC60" blockId="16.[822,1427,546,924]" pageId="16" pageNumber="115">
<treatmentCitationGroup id="AB731671FFB9FFE33626FE5F5D12FC60" pageId="16" pageNumber="115">
<taxonomicName id="4C634ADCFFB9FFE33626FE5F5C3CFC80" baseAuthorityName="Gates" baseAuthorityYear="1925" box="[822,1037,868,892]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="115" phylum="Annelida" rank="species" species="fornicatus">
<emphasis id="B917ED4DFFB9FFE33626FE5F5C3CFC80" box="[822,1037,868,892]" italics="true" pageId="16" pageNumber="115">Amynthas fornicatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AC2174EFFB9FFE3310BFE5F5D33FC80" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" box="[1051,1282,868,892]" page="482" pageId="16" pageNumber="115" year="2003">
<bibRefCitation id="EFF24CAEFFB9FFE3310BFE5F5D33FC80" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" box="[1051,1282,868,892]" pageId="16" pageNumber="115" pagination="479 - 490" refId="ref15882" refString="Shen, H. - P., C. - F. Tsai and S. - C. Tsai. 2003. Six new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Central Taiwan. Zoolological Studies 42 (4): 479 - 490 [Available from: // zoolstud. sinica. edu. tw / Journals / 42.4 / 479. pdf]." type="journal article" year="2003">
Shen
<emphasis id="B917ED4DFFB9FFE33146FE5F5CA1FC80" box="[1110,1168,868,892]" italics="true" pageId="16" pageNumber="115">et al.,</emphasis>
2003: 482
</bibRefCitation>
</treatmentCitation>
, tab. 1;
<bibRefCitation id="EFF24CAEFFB9FFE33044FE5F5BE7FC60" author="Wang, H. - J. &amp; J. - P. Qiu" pageId="16" pageNumber="115" refId="ref16424" refString="Wang, H. - J. and J. - P. Qiu. 2005. A Preliminary Report on" type="book" year="2005">Wang &amp; Qiu, 2005</bibRefCitation>
: tab. 1;
<bibRefCitation id="EFF24CAEFFB9FFE3313AFEBF5CE7FC60" author="Sun, J. &amp; Q. Zhao &amp; J. - P. Qiu" box="[1066,1238,900,924]" pageId="16" pageNumber="115" pagination="149 - 158" refId="ref16335" refString="Sun, J., Q. Zhao and J. - P. Qiu. 2012. Four new species of the Amynthas corticis-group (Oligochaeta: Megascolecidae) from Hainan Island, China. Zootaxa 2680: 149 - 158." type="journal article" year="2012">
Sun
<emphasis id="B917ED4DFFB9FFE3314AFEBE5CA6FC60" box="[1114,1175,900,924]" italics="true" pageId="16" pageNumber="115">et al.,</emphasis>
2012
</bibRefCitation>
: tab. 1.
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</paragraph>
</subSubSection>
<subSubSection id="C37962D4FFB9FFE23626FEFF5CC9FE06" lastPageId="17" lastPageNumber="116" pageId="16" pageNumber="115" type="description">
<paragraph id="8BDC315FFFB9FFE33626FEFF5D33FB81" blockId="16.[822,1427,964,1149]" pageId="16" pageNumber="115">
<emphasis id="B917ED4DFFB9FFE33626FEFF5BF3FC20" bold="true" box="[822,962,964,988]" pageId="16" pageNumber="115">Description.</emphasis>
The original description in full in quotes with Gates (1939) amendments added in square braces that yet ignore
<bibRefCitation id="EFF24CAEFFB9FFE336C3F93F5C56FBE0" author="Chen, Y." box="[979,1127,1028,1052]" pageId="16" pageNumber="115" pagination="269 - 306" refId="ref14831" refString="Chen, Y. 1936. On the terrestrial Oligochaeta from Szechuan II with the notes on Gates' types. Contribibutions of Biological Laboratory of Science Society of China (Zoology) 11: 269 - 306." type="journal article" year="1936">Chens (1936)</bibRefCitation>
additional and contradictory data from reinspection of Gates
<typeStatus id="54D88FFDFFB9FFE33189F91E5CE0FBC1" box="[1177,1233,1061,1085]" pageId="16" pageNumber="115">types</typeStatus>
<emphasis id="B917ED4DFFB9FFE331C8F91E5D33FB81" bold="true" pageId="16" pageNumber="115">
[data from
<bibRefCitation id="EFF24CAEFFB9FFE3304AF91F5B8CFBA0" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" pageId="16" pageNumber="115" pagination="479 - 490" refId="ref15882" refString="Shen, H. - P., C. - F. Tsai and S. - C. Tsai. 2003. Six new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Central Taiwan. Zoolological Studies 42 (4): 479 - 490 [Available from: // zoolstud. sinica. edu. tw / Journals / 42.4 / 479. pdf]." type="journal article" year="2003">
Shen
<emphasis id="B917ED4DFFB9FFE33626F97E5B40FBA0" bold="true" box="[822,881,1092,1116]" italics="true" pageId="16" pageNumber="115">et al.</emphasis>
(2003
</bibRefCitation>
: tab. 1) bolded and also put in square braces but presumably not from
<typeStatus id="54D88FFDFFB9FFE331BCF95E5CDCFB81" box="[1196,1261,1125,1149]" pageId="16" pageNumber="115">types</typeStatus>
]:-
</emphasis>
</paragraph>
<caption id="DF1C61D7FFB9FFE3358DFAA959FAF83E" ID-DOI="http://doi.org/10.5281/zenodo.13144641" ID-Zenodo-Dep="13144641" httpUri="https://zenodo.org/record/13144641/files/figure.png" pageId="16" pageNumber="115" startId="16.[157,190,1938,1959]" targetBox="[320,1268,1211,1906]" targetPageId="16" targetType="figure">
<paragraph id="8BDC315FFFB9FFE3358DFAA959FAF83E" blockId="16.[157,1427,1938,1986]" pageId="16" pageNumber="115">
<emphasis id="B917ED4DFFB9FFE3358DFAA958D4F85B" bold="true" box="[157,229,1938,1959]" pageId="16" pageNumber="115">Fig. 12.</emphasis>
<taxonomicName id="4C634ADCFFB9FFE335FDFAA9599BF85A" baseAuthorityName="Gates" baseAuthorityYear="1925" box="[237,426,1938,1958]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="115" phylum="Annelida" rank="species" species="fornicatus">
<emphasis id="B917ED4DFFB9FFE335FDFAA9599BF85A" box="[237,426,1938,1958]" italics="true" pageId="16" pageNumber="115">Amynthas fornicatus</emphasis>
</taxonomicName>
showing Gates (1935: figs. 5-6) most uninformative and irrelevant details of spermathecae invoking adage “missing the forest for the trees”.
</paragraph>
</caption>
<caption id="DF1C61D7FFB8FFE2358FF9EE594FFAF9" ID-DOI="http://doi.org/10.5281/zenodo.13144643" ID-Zenodo-Dep="13144643" httpUri="https://zenodo.org/record/13144643/files/figure.png" pageId="17" pageNumber="116" startId="17.[159,192,1237,1258]" targetBox="[186,731,257,1204]" targetPageId="17" targetType="figure">
<paragraph id="8BDC315FFFB8FFE2358FF9EE594FFAF9" blockId="17.[159,763,1237,1285]" pageId="17" pageNumber="116">
<emphasis id="B917ED4DFFB8FFE2358FF9EE58D8FB16" bold="true" box="[159,233,1237,1258]" pageId="17" pageNumber="116">Fig. 13.</emphasis>
Prior
<taxonomicName id="4C634ADCFFB8FFE23439F9EE5A25FB15" authorityName="Michaelsen" authorityYear="1910" box="[297,532,1237,1257]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="hongkongensis">
<emphasis id="B917ED4DFFB8FFE23439F9EE5A25FB15" box="[297,532,1237,1257]" italics="true" pageId="17" pageNumber="116">Amynthas hongkongensis</emphasis>
</taxonomicName>
after Michaelsen (1910: fig. 22), for comparison.
</paragraph>
</caption>
<paragraph id="8BDC315FFFB8FFE235A5F87C5BD9FEC5" blockId="17.[159,764,1350,1985]" lastBlockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
<taxonomicName id="4C634ADCFFB8FFE235AFF87C59CCFAA3" authorityName="Blakemore" authorityYear="2013" box="[191,509,1351,1375]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="115" phylum="Annelida" rank="species" species="fornicata" status="n. sp.">PHERETIMA FORNICATA</taxonomicName>
,
<taxonomicNameLabel id="A2245036FFB8FFE2371AF87C5A74FAA2" box="[522,581,1351,1374]" pageId="17" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
Length
<quantity id="4C9B9CBAFFB8FFE237B1F87C58F9FA82" metricMagnitude="-2" metricUnit="m" metricValue="8.4" metricValueMax="9.0" metricValueMin="7.8" pageId="17" pageNumber="116" unit="mm" value="84.0" valueMax="90.0" valueMin="78.0">78 to 90 mm</quantity>
[
<quantity id="4C9B9CBAFFB8FFE235C5F85C5904FA82" box="[213,309,1383,1407]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="116" unit="mm" value="100.0">100 mm</quantity>
], diameter
<quantity id="4C9B9CBAFFB8FFE234BAF85C5A2AFA82" box="[426,539,1382,1407]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="116" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4 to 6 mm</quantity>
.
<emphasis id="B917ED4DFFB8FFE23735F85D5AC4FA82" bold="true" box="[549,757,1382,1406]" pageId="17" pageNumber="116">[Segments 90-105]</emphasis>
. Setae: vi/17-24 [16-26]
<emphasis id="B917ED4DFFB8FFE234B6F8BC59C7FA63" bold="true" box="[422,502,1415,1439]" pageId="17" pageNumber="116">[43-50]</emphasis>
, vii/19-21 [19-23], viii/ 18-23 [18-27]
<emphasis id="B917ED4DFFB8FFE23452F89C59A2FA43" bold="true" box="[322,403,1447,1471]" pageId="17" pageNumber="116">[42-52]</emphasis>
,
<emphasis id="B917ED4DFFB8FFE234B4F89C5A28FA43" bold="true" box="[420,537,1447,1471]" pageId="17" pageNumber="116">[xii/42-52]</emphasis>
, xvii/13-14 [13-19], xviii/9-14, xix/12-15, [x, 56,] xx/56,
<emphasis id="B917ED4DFFB8FFE23738F8FC5A94FA23" bold="true" box="[552,677,1479,1503]" pageId="17" pageNumber="116">[xxv 45-55]</emphasis>
; a wide dorsal gap in the setal circle of ii. First dorsal pore in 12/ 13 [with pore like marking in 11/12]. [Clitellum annular, 13/14-16/17]. Spermathecal pores minute and superficial, four pairs, in 5/6-8/9 [on tiny, grayish, transversely oval markings]. Male pores superficial, on circular to transversely oval, disk-shaped porophores [that are about
<quantity id="4C9B9CBAFFB8FFE237CAFBB358F9F943" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="116" unit="mm" value="0.5">0.5 mm</quantity>
wide]. No genital markings. Septum 8/9 present but membranous [9/10 laking]. [Intestine from 15]. Intestinal caeca simple. [Hearts in 9-13 to ventral blood vessel, those in 10 and 11 within testis sac]. Testis sacs of x and xi unpaired and horseshoe-shaped. [Seminal vesicles are in 11 &amp; 12, pseudovesicles in 13 and sometimes 14]. [Spermathecal duct as long as the ampulla, diverticulum longer than duct and ampulla]. Spermathecal diverticulum with a long, slender stalk and a spheroidal or asymmetrical seminal chamber.
<emphasis id="B917ED4DFFB8FFE23118FC395C0CFEE5" box="[1032,1085,258,281]" italics="true" pageId="17" pageNumber="116">
<typeStatus id="54D88FFDFFB8FFE23118FC395C0CFEE5" box="[1032,1085,258,281]" pageId="17" pageNumber="116">Type</typeStatus>
</emphasis>
. -
<collectingCountry id="F37471CFFFB8FFE23171FC3A5CBEFEE5" box="[1121,1167,257,281]" name="United States of America" pageId="17" pageNumber="116">U.S.</collectingCountry>
N.M no. 200099, from Tatsienlu, Tibet.
</paragraph>
<paragraph id="8BDC315FFFB8FFE2365FFC7A5CC9FE06" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
Distinguished from
<taxonomicName id="4C634ADCFFB8FFE23132FC795CECFEA5" authorityName="Michaelsen" authorityYear="1910" box="[1058,1245,321,345]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="hongkongensis">
<emphasis id="B917ED4DFFB8FFE23132FC795CECFEA5" box="[1058,1245,321,345]" italics="true" pageId="17" pageNumber="116">P. hongkongensis</emphasis>
</taxonomicName>
by the gap in the setal circle of ii, the exclusion of the seminal vesicles of xi from the posterior testis sac, and the absence of genital markings.” [None of these latter three characters seems sufficient justification for their separation in context with the current revisions of
<taxonomicName id="4C634ADCFFB8FFE2317BFCD95CD8FE06" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[1131,1257,482,506]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE2317BFCD95CD8FE06" box="[1131,1257,482,506]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C37962D4FFB8FFE23628FF395C50F85C" pageId="17" pageNumber="116" type="discussion">
<paragraph id="8BDC315FFFB8FFE23628FF395BE6FD27" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
<emphasis id="B917ED4DFFB8FFE23628FF395B96FDE6" bold="true" box="[824,935,514,538]" pageId="17" pageNumber="116">Remarks.</emphasis>
<bibRefCitation id="EFF24CAEFFB8FFE236BEFF395C59FDE6" author="Chen, Y." box="[942,1128,514,538]" pageId="17" pageNumber="116" pagination="269 - 306" refId="ref14831" refString="Chen, Y. 1936. On the terrestrial Oligochaeta from Szechuan II with the notes on Gates' types. Contribibutions of Biological Laboratory of Science Society of China (Zoology) 11: 269 - 306." type="journal article" year="1936">Chen (1936: 298)</bibRefCitation>
said
<taxonomicName id="4C634ADCFFB8FFE231B2FF385D12FDE6" authorityName="Blakemore" authorityYear="2013" box="[1186,1315,514,538]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE231B2FF385D12FDE6" box="[1186,1315,514,538]" italics="true" pageId="17" pageNumber="116">P. fornicata</emphasis>
</taxonomicName>
“is probably identical with
<taxonomicName id="4C634ADCFFB8FFE236EAFF185CADFDC6" authority="Steph." authorityName="Steph." box="[1018,1180,546,570]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B917ED4DFFB8FFE236EAFF185C7EFDC6" box="[1018,1103,547,570]" italics="true" pageId="17" pageNumber="116">P. pingi</emphasis>
Steph.
</taxonomicName>
” but
<bibRefCitation id="EFF24CAEFFB8FFE231C6FF195DA5FDC6" author="Gates, G. E." box="[1238,1428,546,570]" pageId="17" pageNumber="116" pagination="405 - 507" refId="ref15162" refString="Gates, G. E. 1939. On some species of Chinese earthworms with special reference to specimens collected in Szechuan by Dr. D. C. Graham. Proceedings of the United States National Museum 85: 405 - 507 [Available from: si-dppr. si. edu / dspace / bitstream / 10088 / 16290 / USNMP- 85 _ 3040 _ 1939. pdf]." type="journal article" year="1939">Gates (1939: 436)</bibRefCitation>
responded tersely with “
<taxonomicName id="4C634ADCFFB8FFE23150FF785CF0FDA7" authorityName="Blakemore" authorityYear="2013" box="[1088,1217,579,603]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE23150FF785CF0FDA7" box="[1088,1217,579,603]" italics="true" pageId="17" pageNumber="116">P. fornicata</emphasis>
</taxonomicName>
is, on the contrary, clearly distinguished from
<taxonomicName id="4C634ADCFFB8FFE23179FF585CF2FD86" box="[1129,1219,611,634]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B917ED4DFFB8FFE23179FF585CF2FD86" box="[1129,1219,611,634]" italics="true" pageId="17" pageNumber="116">P. pingi</emphasis>
</taxonomicName>
by the horseshoe-shaped testis sacs of x and xi.” However, I think this is an unreliable character for separation and am obliged to combine them.
</paragraph>
<paragraph id="8BDC315FFFB8FFE2365FFFD85CABFC00" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
<bibRefCitation id="EFF24CAEFFB8FFE2365FFFD85BE4FD07" author="Gates, G. E." box="[847,981,739,763]" pageId="17" pageNumber="116" pagination="1 - 19" refId="ref15126" refString="Gates, G. E. 1935. New earthworms from China, with notes on the synonymy of some Chinese species of Drawida and Pheretima. Smithsonian Miscellaneous Collections 93: 1 - 19." type="journal article" year="1935">Gates (1935)</bibRefCitation>
had implicated his
<taxonomicName id="4C634ADCFFB8FFE231B6FFD85D37FD07" authorityName="Blakemore" authorityYear="2013" box="[1190,1286,739,763]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE231B6FFD85D37FD07" box="[1190,1286,739,763]" italics="true" pageId="17" pageNumber="116">fornicata</emphasis>
</taxonomicName>
as being closest to
<taxonomicName id="4C634ADCFFB8FFE23699FE3F5D7FFCE7" authority="Michaelsen, 1910: 107" authorityName="Michaelsen" authorityPageNumber="107" authorityYear="1910" box="[905,1358,771,795]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="hongkongensis">
<emphasis id="B917ED4DFFB8FFE23699FE3F5C78FCE7" box="[905,1097,771,795]" italics="true" pageId="17" pageNumber="116">P. hongkongensis</emphasis>
Michaelsen, 1910: 107
</taxonomicName>
based on a probable immature and abnormal type that has only a pair of small marking immediately median to male pores on 18 which is all that separates it from
<taxonomicName id="4C634ADCFFB8FFE231E6FE5F5D44FC87" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[1270,1397,868,891]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE231E6FE5F5D44FC87" box="[1270,1397,868,891]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
as redescribed by
<bibRefCitation id="EFF24CAEFFB8FFE236CDFEBF5C9BFC60" author="Blakemore, R. J." box="[989,1194,900,924]" pageId="17" pageNumber="116" pagination="35 - 43" refId="ref14125" refString="Blakemore, R. J. 2012 a. Amynthas carnosus (Goto &amp; Hatai, 1899) redescribed on its neotype (Oligochaeta: Megadrilacea: Megascolecidae). Journal of Species Research 1 (1): 35 - 43 [Available from: www. nibr. go. kr / download? attach _ path = journal &amp; attach = 2012 vol 1 no 1 _ 4. pdf]." type="journal article" year="2012">Blakemore (2012a)</bibRefCitation>
based on its
<typeStatus id="54D88FFDFFB8FFE2302AFEBF5DA0FC60" box="[1338,1425,900,924]" pageId="17" pageNumber="116" type="neotype">neotype</typeStatus>
, and on
<bibRefCitation id="EFF24CAEFFB8FFE2369DFE9F5C50FC40" author="Kobayashi, S." box="[909,1121,932,956]" pageId="17" pageNumber="116" pagination="115 - 138" refId="ref15652" refString="Kobayashi, S. 1936. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashis (1936)</bibRefCitation>
excellent redescription in a publication that
<bibRefCitation id="EFF24CAEFFB8FFE236F9FEFF5C9BFC20" author="Gates, G. E." box="[1001,1194,964,988]" pageId="17" pageNumber="116" pagination="405 - 507" refId="ref15162" refString="Gates, G. E. 1939. On some species of Chinese earthworms with special reference to specimens collected in Szechuan by Dr. D. C. Graham. Proceedings of the United States National Museum 85: 405 - 507 [Available from: si-dppr. si. edu / dspace / bitstream / 10088 / 16290 / USNMP- 85 _ 3040 _ 1939. pdf]." type="journal article" year="1939">Gates (1939: 469)</bibRefCitation>
was clearly aware of but rejected for unsound reasons.
</paragraph>
<paragraph id="8BDC315FFFB8FFE2365FF93F5BF5FA82" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
Thus differences of
<taxonomicName id="4C634ADCFFB8FFE23122F93E5C87FBE0" authorityName="Blakemore" authorityYear="2013" box="[1074,1206,1028,1052]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE23122F93E5C87FBE0" box="[1074,1206,1028,1052]" italics="true" pageId="17" pageNumber="116">P. fornicata</emphasis>
</taxonomicName>
from
<taxonomicName id="4C634ADCFFB8FFE231EFF93E5D68FBE0" box="[1279,1369,1029,1052]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B917ED4DFFB8FFE231EFF93E5D68FBE0" box="[1279,1369,1029,1052]" italics="true" pageId="17" pageNumber="116">A. pingi</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C634ADCFFB8FFE2306EF93E5BA9FBC0" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE2306EF93E5BA9FBC0" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
) as reinspected here from its type, are mainly a lack of genital markings which is irrelevant for
<emphasis id="B917ED4DFFB8FFE23021F97E5B53FB80" italics="true" pageId="17" pageNumber="116">
<taxonomicName id="4C634ADCFFB8FFE23021F97E5B6FFB80" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">A. carnosus</taxonomicName>
,
</emphasis>
and the uncertainty about spermathecal papillae. However, the name
<taxonomicName id="4C634ADCFFB8FFE2311DF9BE5C5FFB61" authorityName="Blakemore" authorityYear="2013" box="[1037,1134,1157,1181]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE2311DF9BE5C5FFB61" box="[1037,1134,1157,1181]" italics="true" pageId="17" pageNumber="116">fornicata</emphasis>
</taxonomicName>
is complicit with the redescription of
<taxonomicName id="4C634ADCFFB8FFE236A4F99E5C04FB41" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[948,1077,1189,1213]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE236A4F99E5C04FB41" box="[948,1077,1189,1213]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
based on its
<typeStatus id="54D88FFDFFB8FFE231DCF99E5D11FB41" box="[1228,1312,1189,1213]" pageId="17" pageNumber="116" type="neotype">neotype</typeStatus>
by
<bibRefCitation id="EFF24CAEFFB8FFE2305EF99E5BF9FB21" author="Blakemore, R. J." pageId="17" pageNumber="116" refId="ref14323" refString="Blakemore, R. J. 2012 d. Cosmopolitan earthworms. 5 th Edition. VermEcology, Yokohama, Japan. pp. 950 [CD publication]." type="journal volume" year="2012">Blakemore (2012d)</bibRefCitation>
and on subsequent records: in particular it agrees with
<bibRefCitation id="EFF24CAEFFB8FFE236ACF9DE5CBFFB01" author="Kobayashi, S." box="[956,1166,1253,1277]" pageId="17" pageNumber="116" pagination="115 - 138" refId="ref15652" refString="Kobayashi, S. 1936. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashis (1936)</bibRefCitation>
absent GMs form types I &amp; I. Thus
<taxonomicName id="4C634ADCFFB8FFE236D5F83E5C1BFAE1" authorityName="Blakemore" authorityYear="2013" box="[965,1066,1285,1309]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicata">
<emphasis id="B917ED4DFFB8FFE236D5F83E5C1BFAE1" box="[965,1066,1285,1309]" italics="true" pageId="17" pageNumber="116">fornicata</emphasis>
</taxonomicName>
should probably be included in
<taxonomicName id="4C634ADCFFB8FFE23628F81D5B5EFAC1" box="[824,879,1318,1341]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B917ED4DFFB8FFE23628F81D5B5EFAC1" box="[824,879,1318,1341]" italics="true" pageId="17" pageNumber="116">pingi</emphasis>
</taxonomicName>
as Chen suggested and both now seemingly merit inclusion in synonymy of the prior
<taxonomicName id="4C634ADCFFB8FFE231A6F87D5BF0FA82" authority="(Goto &amp; Hatai, 1899)" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE231A6F87D5D04FAA1" box="[1206,1333,1350,1373]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
(
<bibRefCitation id="EFF24CAEFFB8FFE23051F87D5B89FA82" author="Goto, S. &amp; S. Hatai" pageId="17" pageNumber="116" pagination="13 - 24" refId="ref15374" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BDC315FFFB8FFE2365FF8BD5C89F963" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
Another recently described taxon,
<taxonomicName id="4C634ADCFFB8FFE231A7F8BD5C2AFA42" authority="Shen et al. (2013)" authorityName="Shen" authorityYear="2013" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="taiwumontis">
<emphasis id="B917ED4DFFB8FFE231A7F8BD5B60FA42" italics="true" pageId="17" pageNumber="116">Amynthas taiwumontis</emphasis>
<bibRefCitation id="EFF24CAEFFB8FFE23649F89D5C2AFA42" author="Shen, H. - P. &amp; C. - H. Chang &amp; C. - L. Li &amp; W. - J. Chih &amp; J. - H. Chen" box="[857,1051,1446,1470]" pageId="17" pageNumber="116" pagination="471 - 482" refId="ref16042" refString="Shen, H. - P., C. - H. Chang, C. - L. Li, W. - J. Chih and J. - H. Chen. 2013. Four new earthworms species of the genus Amynthas (Oligochaeta: Megascolecidae) from Kinmen, Taiwan. Zootaxa 3599: 471 - 482." type="journal article" year="2013">
Shen
<emphasis id="B917ED4DFFB8FFE23687F89C5BF6FA42" box="[919,967,1446,1470]" italics="true" pageId="17" pageNumber="116">et al</emphasis>
. (2013)
</bibRefCitation>
</taxonomicName>
was compared by those authors to
<taxonomicName id="4C634ADCFFB8FFE23628F8FD5B8EFA22" baseAuthorityName="Gates" baseAuthorityYear="1925" box="[824,959,1478,1502]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="fornicatus">
<emphasis id="B917ED4DFFB8FFE23628F8FD5B8EFA22" box="[824,959,1478,1502]" italics="true" pageId="17" pageNumber="116">A. fornicatus</emphasis>
</taxonomicName>
. However its description appears to comply with
<taxonomicName id="4C634ADCFFB8FFE2367DF8DC5BDBFA02" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[877,1002,1511,1534]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE2367DF8DC5BDBFA02" box="[877,1002,1511,1534]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
having Kobayashis spermathecal pores
<typeStatus id="54D88FFDFFB8FFE23628FB3C5B55F9E3" box="[824,868,1543,1567]" pageId="17" pageNumber="116">type</typeStatus>
I or II and male pores
<typeStatus id="54D88FFDFFB8FFE2315FFB3C5C4AF9E3" box="[1103,1147,1543,1567]" pageId="17" pageNumber="116">type</typeStatus>
I, as redescribed by
<bibRefCitation id="EFF24CAEFFB8FFE23040FB3D5BF5F9C3" author="Blakemore, R. J." pageId="17" pageNumber="116" pagination="35 - 43" refId="ref14125" refString="Blakemore, R. J. 2012 a. Amynthas carnosus (Goto &amp; Hatai, 1899) redescribed on its neotype (Oligochaeta: Megadrilacea: Megascolecidae). Journal of Species Research 1 (1): 35 - 43 [Available from: www. nibr. go. kr / download? attach _ path = journal &amp; attach = 2012 vol 1 no 1 _ 4. pdf]." type="journal article" year="2012">Blakemore (2012a)</bibRefCitation>
(this overlooked despite my sending a copy to Dr Huei-Ping Shen) and thus this name too is another probable inclusion in the long synonym list of
<taxonomicName id="4C634ADCFFB8FFE2303FFB5C5B6BF963" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE2303FFB5C5B6BF963" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
presented in the Abstract above.
</paragraph>
<paragraph id="8BDC315FFFB8FFE2365FFB935C50F85C" blockId="17.[824,1429,257,1952]" pageId="17" pageNumber="116">
However, any of these specimens with spermathecal pores in 5/6/7/8/9
<emphasis id="B917ED4DFFB8FFE23111FBFC5C17F923" bold="true" box="[1025,1062,1735,1759]" pageId="17" pageNumber="116">not</emphasis>
on papillae/porophores described for
<taxonomicName id="4C634ADCFFB8FFE23670FBD35BD1F903" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[864,992,1768,1791]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B917ED4DFFB8FFE23670FBD35BD1F903" box="[864,992,1768,1791]" italics="true" pageId="17" pageNumber="116">A. carnosus</emphasis>
</taxonomicName>
probably equate more to
<taxonomicName id="4C634ADCFFB8FFE231EEFBD35D5EF903" baseAuthorityName="Kinberg" baseAuthorityYear="1876" box="[1278,1391,1768,1791]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="corticis">
<emphasis id="B917ED4DFFB8FFE231EEFBD35D5EF903" box="[1278,1391,1768,1791]" italics="true" pageId="17" pageNumber="116">A. corticis</emphasis>
</taxonomicName>
, as indeed does
<taxonomicName id="4C634ADCFFB8FFE236ADFA335D3DF8DC" authority="Shen et al., 2003" authorityName="Shen" authorityYear="2003" box="[957,1292,1799,1824]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="17" pageNumber="116" phylum="Annelida" rank="species" species="penpuensis">
<emphasis id="B917ED4DFFB8FFE236ADFA335C60F8E3" box="[957,1105,1800,1823]" italics="true" pageId="17" pageNumber="116">A. penpuensis</emphasis>
<bibRefCitation id="EFF24CAEFFB8FFE23148FA3C5D3DF8DC" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" box="[1112,1292,1799,1824]" pageId="17" pageNumber="116" pagination="479 - 490" refId="ref15882" refString="Shen, H. - P., C. - F. Tsai and S. - C. Tsai. 2003. Six new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Central Taiwan. Zoolological Studies 42 (4): 479 - 490 [Available from: // zoolstud. sinica. edu. tw / Journals / 42.4 / 479. pdf]." type="journal article" year="2003">
Shen
<emphasis id="B917ED4DFFB8FFE23185FA335CFBF8E3" box="[1173,1226,1799,1823]" italics="true" pageId="17" pageNumber="116">et al.</emphasis>
, 2003
</bibRefCitation>
</taxonomicName>
as
<emphasis id="B917ED4DFFB8FFE23021FA335DA4F8DC" bold="true" box="[1329,1429,1800,1824]" pageId="17" pageNumber="116">syn. nov.</emphasis>
since all that currently separates it is its dorsal pores slightly more in advance (from 5/6 or 6/7 rather than from 11/12 abouts), this single character is doubtfully adequate for species level separation.
</paragraph>
</subSubSection>
</treatment>
</document>