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@ -56,7 +57,7 @@
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<paragraph id="8BDC315FFFABFFF13626FC1A5B96FEC5" blockId="2.[822,1427,257,442]" box="[822,935,289,313]" pageId="2" pageNumber="101">
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(
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<figureCitation id="13582DDAFFABFFF1362EFC1A5BAFFEC5" box="[830,926,289,313]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[159,192,1704,1725]" captionStartId-1="4.[157,190,1302,1323]" captionTargetBox-0="[265,1286,263,1647]" captionTargetBox-1="[160,766,262,1270]" captionTargetId-0="figure-104@3.[232,1367,260,1726]" captionTargetId-1="figure-579@4.[151,811,238,1292]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 2. A. carnosus - A. Korean specimen from Geoman (NIBR IV261234 - DNA w37). B. Japanese neotype of A. carnosus (Tokyo An- 435 from Blakemore, 2012a). These two specimens are almost identical morphologically allowing genetic comparison(Fig. 1)." captionText-1="Fig. 3. A. carnosus - sketch of Hikone specimen (Tokyo An-460 - DNA JET-112) showing ventral view of anterior and dorsal views of prostomium, clitellum and flared male pores posteriorly (other specimen An-461 and JET samples 113-115 agreed 100%). This specimen resembles P. pingi (cf Fig. 11) but its DNA agrees with A. carnosus (as in Fig. 1)." pageId="2" pageNumber="101">Figs. 2-3</figureCitation>
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<figureCitation id="13582DDAFFABFFF1362EFC1A5BAFFEC5" box="[830,926,289,313]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[159,192,1704,1725]" captionStartId-1="4.[157,190,1302,1323]" captionTargetBox-0="[265,1286,263,1647]" captionTargetBox-1="[160,766,262,1270]" captionTargetId-0="figure-104@3.[232,1367,260,1726]" captionTargetId-1="figure-579@4.[151,811,238,1292]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 2. A. carnosus - A. Korean specimen from Geoman (NIBR IV261234 - DNA w37). B. Japanese neotype of A. carnosus (Tokyo An- 435 from Blakemore, 2012a). These two specimens are almost identical morphologically allowing genetic comparison(Fig. 1)." captionText-1="Fig. 3. A. carnosus - sketch of Hikone specimen (Tokyo An-460 - DNA JET-112) showing ventral view of anterior and dorsal views of prostomium, clitellum and flared male pores posteriorly (other specimen An-461 and JET samples 113-115 agreed 100%). This specimen resembles P. pingi (cf Fig. 11) but its DNA agrees with A. carnosus (as in Fig. 1)." figureDoi-0="http://doi.org/10.5281/zenodo.13144619" figureDoi-1="http://doi.org/10.5281/zenodo.13144623" httpUri-0="https://zenodo.org/record/13144619/files/figure.png" httpUri-1="https://zenodo.org/record/13144623/files/figure.png" pageId="2" pageNumber="101">Figs. 2-3</figureCitation>
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)
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</paragraph>
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@ -75,7 +76,7 @@ and
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<emphasis id="B917ED4DFFABFFF131D2FC595D2CFE85" box="[1218,1309,354,377]" italics="true" pageId="2" pageNumber="101">A. pingi</emphasis>
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</taxonomicName>
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described below). DNA data (
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<figureCitation id="13582DDAFFABFFF1310EFCB95C6AFE66" box="[1054,1115,386,410]" captionStart="Fig" captionStartId="2.[157,190,1924,1945]" captionTargetBox="[199,1380,1077,1894]" captionTargetId="figure-304@2.[190,1400,1065,1897]" captionTargetPageId="2" captionText="Fig. 1. Phylotree from MEGA5 program Maximum Likelihood defaults - sample codes explained in the Appendix; ◆- samples have figured specimens; ◇- ex GenBank. Appears to show significant genetic difference between specimens superficially compliant with broad A. corticis definition as discussed below that taxon herein. Synonym priority for either Gp 1 or Gp 2 is currently unresolved." pageId="2" pageNumber="101">Fig. 1</figureCitation>
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<figureCitation id="13582DDAFFABFFF1310EFCB95C6AFE66" box="[1054,1115,386,410]" captionStart="Fig" captionStartId="2.[157,190,1924,1945]" captionTargetBox="[199,1380,1077,1894]" captionTargetId="figure-304@2.[190,1400,1065,1897]" captionTargetPageId="2" captionText="Fig. 1. Phylotree from MEGA5 program Maximum Likelihood defaults - sample codes explained in the Appendix; ◆- samples have figured specimens; ◇- ex GenBank. Appears to show significant genetic difference between specimens superficially compliant with broad A. corticis definition as discussed below that taxon herein. Synonym priority for either Gp 1 or Gp 2 is currently unresolved." figureDoi="http://doi.org/10.5281/zenodo.13144617" httpUri="https://zenodo.org/record/13144617/files/figure.png" pageId="2" pageNumber="101">Fig. 1</figureCitation>
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) indicates a new sub-species as next described.
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</paragraph>
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<figureCitation id="13582DDAFFB9FFE336B2FF785B8CFDA7" box="[930,957,579,603]" captionStart="Fig" captionStartId="17.[159,192,1237,1258]" captionTargetBox="[186,731,257,1204]" captionTargetId="figure-704@17.[186,731,257,1204]" captionTargetPageId="17" captionText="Fig. 13. Prior Amynthas hongkongensis after Michaelsen (1910: fig. 22), for comparison." pageId="16" pageNumber="115">13</figureCitation>
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@ -145,7 +146,7 @@ Shen
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]:-
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<taxonomicName id="4C634ADCFFB9FFE335FDFAA9599BF85A" baseAuthorityName="Gates" baseAuthorityYear="1925" box="[237,426,1938,1958]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="16" pageNumber="115" phylum="Annelida" rank="species" species="fornicatus">
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@ -154,7 +155,7 @@ Shen
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showing Gates’ (1935: figs. 5-6) most uninformative and irrelevant details of spermathecae invoking adage “missing the forest for the trees”.
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<subSubSection id="65C66517D700A921FF50FA4FFD53E020" box="[157,698,1528,1553]" pageId="2" pageNumber="129" type="nomenclature">
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|
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@ -63,7 +64,7 @@ Blakemore
|
|||
<paragraph id="2D63369CD700A921FF7AF9AFFEE6E001" blockId="2.[183,271,1560,1584]" box="[183,271,1560,1584]" pageId="2" pageNumber="129">
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<emphasis id="1FA8EA8ED700A921FF7AF9AFFEE6E001" bold="true" box="[183,271,1560,1584]" pageId="2" pageNumber="129">
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(
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<figureCitation id="B5E72A19D700A921FF72F9AFFEEBE001" box="[191,258,1560,1584]" captionStart="Fig" captionStartId="3.[159,192,1272,1293]" captionTargetBox="[249,1303,261,1228]" captionTargetId="figure-350@3.[231,1426,241,1254]" captionTargetPageId="3" captionText="Fig. 2. A. Drawida songae yeongdo subsp. n. holotype (H, DNA H1) showing prostomium, gizzards, pygidium, habitus with reproductive organs in situ and all four genital marking glands in 8-11lhs plus a nephridium from 15rhs. B. D. songae songae after Hong (2002: fig. 1) with GMs in 7, 10-11(but not 12?) for fair use comparison." pageId="2" pageNumber="129">Fig. 2</figureCitation>
|
||||
<figureCitation id="B5E72A19D700A921FF72F9AFFEEBE001" box="[191,258,1560,1584]" captionStart="Fig" captionStartId="3.[159,192,1272,1293]" captionTargetBox="[249,1303,261,1228]" captionTargetId="figure-350@3.[231,1426,241,1254]" captionTargetPageId="3" captionText="Fig. 2. A. Drawida songae yeongdo subsp. n. holotype (H, DNA H1) showing prostomium, gizzards, pygidium, habitus with reproductive organs in situ and all four genital marking glands in 8-11lhs plus a nephridium from 15rhs. B. D. songae songae after Hong (2002: fig. 1) with GMs in 7, 10-11(but not 12?) for fair use comparison." figureDoi="http://doi.org/10.5281/zenodo.13144592" httpUri="https://zenodo.org/record/13144592/files/figure.png" pageId="2" pageNumber="129">Fig. 2</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -139,7 +140,7 @@ with fewer segments (viz. 172). Both taxa have simple, superficial spermathecal
|
|||
<emphasis id="1FA8EA8ED700A921FAEEFBB2FA7AE22D" box="[1315,1427,1029,1052]" italics="true" pageId="2" pageNumber="129">D. songae</emphasis>
|
||||
</taxonomicName>
|
||||
was described with a pink colour (but possibly due to preservation?) rather than green as in the current specimen. The genital markings were twice described as from 7-11,12 (although it is ambiguous since only some were figured - see
|
||||
<figureCitation id="B5E72A19D700A921FC73FB12FBE6E28C" box="[958,1039,1189,1213]" captionStart="Fig" captionStartId="3.[159,192,1272,1293]" captionTargetBox="[249,1303,261,1228]" captionTargetId="figure-350@3.[231,1426,241,1254]" captionTargetPageId="3" captionText="Fig. 2. A. Drawida songae yeongdo subsp. n. holotype (H, DNA H1) showing prostomium, gizzards, pygidium, habitus with reproductive organs in situ and all four genital marking glands in 8-11lhs plus a nephridium from 15rhs. B. D. songae songae after Hong (2002: fig. 1) with GMs in 7, 10-11(but not 12?) for fair use comparison." pageId="2" pageNumber="129">Fig. 2B</figureCitation>
|
||||
<figureCitation id="B5E72A19D700A921FC73FB12FBE6E28C" box="[958,1039,1189,1213]" captionStart="Fig" captionStartId="3.[159,192,1272,1293]" captionTargetBox="[249,1303,261,1228]" captionTargetId="figure-350@3.[231,1426,241,1254]" captionTargetPageId="3" captionText="Fig. 2. A. Drawida songae yeongdo subsp. n. holotype (H, DNA H1) showing prostomium, gizzards, pygidium, habitus with reproductive organs in situ and all four genital marking glands in 8-11lhs plus a nephridium from 15rhs. B. D. songae songae after Hong (2002: fig. 1) with GMs in 7, 10-11(but not 12?) for fair use comparison." figureDoi="http://doi.org/10.5281/zenodo.13144592" httpUri="https://zenodo.org/record/13144592/files/figure.png" pageId="2" pageNumber="129">Fig. 2B</figureCitation>
|
||||
), whereas the current specimen lacks markings in both 7 and 12 but has them clearly in 8-11. The markings in
|
||||
<taxonomicName id="EADC4D1FD700A921FC23FB51FBB2E2CC" authorityName="Hong" authorityYear="2002" box="[1006,1115,1254,1277]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="2" pageNumber="129" phylum="Annelida" rank="species" species="songae">
|
||||
<emphasis id="1FA8EA8ED700A921FC23FB51FBB2E2CC" box="[1006,1115,1254,1277]" italics="true" pageId="2" pageNumber="129">D. songae</emphasis>
|
||||
|
|
|
|||
|
|
@ -1,48 +1,49 @@
|
|||
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@ -99,7 +100,7 @@ to
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<emphasis id="1FA8EA8ED700A921FE20FB2CFD8EE283" box="[493,615,1178,1202]" italics="true" pageId="2" pageNumber="129">D. koreana</emphasis>
|
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|
||||
is a variable species. Several other specimens in current studies are superficially similar but appear to differ molecularly (
|
||||
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|
||||
<figureCitation id="B5E72A19D700A921FD02FB6DFF40E323" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="2" pageNumber="129">Fig. 1</figureCitation>
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).
|
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<typeStatus id="F267883ED700A921FF0DFB4CFEE9E322" box="[192,256,1275,1299]" pageId="2" pageNumber="129">Types</typeStatus>
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of
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@ -1,50 +1,51 @@
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@ -63,7 +64,7 @@ Blakemore
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<paragraph id="2D63369CD700A921FC9CF940FC40E13E" blockId="2.[849,937,1783,1807]" box="[849,937,1783,1807]" pageId="2" pageNumber="129">
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<figureCitation id="B5E72A19D700A921FC95F940FC72E13E" box="[856,923,1783,1807]" captionStart="Fig" captionStartId="4.[157,190,1574,1595]" captionTargetBox="[325,1258,259,1545]" captionTargetId="figure-137@4.[310,1294,249,1559]" captionTargetPageId="4" captionText="Fig. 3. A. Amynthas carnosus roki subsp. n. holotype (H, DNA w56) showing habitus, prostomium, spermathecae, caecum and prostate of 18lhs in situ with X2 enlargements of post-intersegmental spermathecal pores of 8rhs and simple male pore of 18rhs. B. Sketch of these from Busan paratype (P, DNA H4 agreeing 100%). 3C and 3D are comparison sketches with similar X2 enlargements for sympatric A. carnosus carnosus (Goto and Hatai, 1899) specimens from Incheon (200 and 280mm long) with pre-intersegmental spermathecal pores that provided DNA samples w55 and w57, respectively, agreeing>99% with each other (see Fig. 1 and Appendix)." figureDoi="http://doi.org/10.5281/zenodo.13144594" httpUri="https://zenodo.org/record/13144594/files/figure.png" pageId="2" pageNumber="129">Fig. 3</figureCitation>
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).
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</emphasis>
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|
|
@ -98,7 +99,7 @@ April, 2013 from
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.
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<emphasis id="1FA8EA8ED701A920FF52FB4FFF35E33C" bold="true" box="[159,220,1272,1293]" pageId="3" pageNumber="130">Fig. 2.</emphasis>
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A.
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|
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@ -181,7 +182,7 @@ and
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, respectively.
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<emphasis id="1FA8EA8ED706A927FF50F991FF33E00A" bold="true" box="[157,218,1574,1595]" pageId="4" pageNumber="131">Fig. 3.</emphasis>
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A.
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|
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@ -256,7 +257,7 @@ that clearly separate out on their genetics from the current taxon.
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<paragraph id="2D63369CD707A926FF78FD74FEB2E5AD" blockId="5.[159,764,257,1984]" pageId="5" pageNumber="132">
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From the results in the DNA phylogram (
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<figureCitation id="B5E72A19D707A926FDA4FD74FD4FE4EA" box="[617,678,707,731]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." pageId="5" pageNumber="132">Fig. 1</figureCitation>
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<figureCitation id="B5E72A19D707A926FDA4FD74FD4FE4EA" box="[617,678,707,731]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="5" pageNumber="132">Fig. 1</figureCitation>
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) it appears that the specimen from Ulleungdo (providing DNA sample w54) labeled as “?
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<taxonomicName id="EADC4D1FD707A926FE0FFCB4FF34E50A" authority="(Stephenson, 1925)" baseAuthorityName="Stephenson" baseAuthorityYear="1925" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="132" phylum="Annelida" rank="species" species="pingi">
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<emphasis id="1FA8EA8ED707A926FE0FFCB4FD82E52A" box="[450,619,771,795]" italics="true" pageId="5" pageNumber="132">Amynthas pingi</emphasis>
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@ -1,48 +1,49 @@
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@ -62,7 +63,7 @@ Blakemore, 2012
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<paragraph id="2D63369CD704A925FC9CFA7CFC53E3D2" blockId="6.[849,954,1483,1507]" box="[849,954,1483,1507]" pageId="6" pageNumber="133">
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(
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<figureCitation id="B5E72A19D704A925FC95FA7CFC44E3D2" box="[856,941,1483,1507]" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." pageId="6" pageNumber="133">Fig. 4B</figureCitation>
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<figureCitation id="B5E72A19D704A925FC95FA7CFC44E3D2" box="[856,941,1483,1507]" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." figureDoi="http://doi.org/10.5281/zenodo.13144596" httpUri="https://zenodo.org/record/13144596/files/figure.png" pageId="6" pageNumber="133">Fig. 4B</figureCitation>
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).
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</emphasis>
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</paragraph>
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@ -71,7 +72,7 @@ Blakemore, 2012
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<paragraph id="2D63369CD704A925FCFBF9BCFC0EE055" blockId="6.[822,1428,1547,1990]" pageId="6" pageNumber="133">
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<emphasis id="1FA8EA8ED704A925FCFBF9BCFC72E012" bold="true" box="[822,923,1547,1571]" pageId="6" pageNumber="133">Material.</emphasis>
|
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IV0000261294 (DNA HY2, this data not includ- ed in phylogram
|
||||
<figureCitation id="B5E72A19D704A925FC35F99BFBD1E075" box="[1016,1080,1580,1604]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." pageId="6" pageNumber="133">Fig. 1</figureCitation>
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<figureCitation id="B5E72A19D704A925FC35F99BFBD1E075" box="[1016,1080,1580,1604]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="6" pageNumber="133">Fig. 1</figureCitation>
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) collected from Jungang Park, 26
|
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<superScript id="DAA99BD4D704A925FC9CF9FDFCB7E06B" attach="left" box="[849,862,1610,1626]" fontSize="7" pageId="6" pageNumber="133">th</superScript>
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April, 2013.
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@ -1,50 +1,51 @@
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@ -77,7 +78,7 @@ Body has ‘classic’ transversely coloured stripes alternating with pale inter
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Part of the species-complex (
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<figureCitation id="B5E72A19D705A924FDF0FAF6FD93E368" box="[573,634,1345,1369]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." pageId="7" pageNumber="134">Fig. 1</figureCitation>
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<figureCitation id="B5E72A19D705A924FDF0FAF6FD93E368" box="[573,634,1345,1369]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="7" pageNumber="134">Fig. 1</figureCitation>
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||||
and Appendix), agreeing 100% with GenBank sample (FJ214216) from “
|
||||
<collectingCountry id="55CB760CD705A924FF2BFA36FEB9E3A8" box="[230,336,1409,1433]" name="Denmark" pageId="7" pageNumber="134">Denmark</collectingCountry>
|
||||
: Svendborg”. Obviously this specimen differs from some Korean samples variously named “
|
||||
|
|
|
|||
|
|
@ -1,48 +1,49 @@
|
|||
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<treatment id="A575878AD705A92BFF52F9D2FE0AE0F7" LSID="urn:lsid:plazi:treatment:A575878AD705A92BFF52F9D2FE0AE0F7" httpUri="http://treatment.plazi.org/id/A575878AD705A92BFF52F9D2FE0AE0F7" lastPageId="8" lastPageNumber="135" pageId="7" pageNumber="134">
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@ -63,7 +64,7 @@ sub-sp. n.
|
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<paragraph id="2D63369CD705A924FF0AF933FEF6E0AD" blockId="7.[199,287,1668,1692]" box="[199,287,1668,1692]" pageId="7" pageNumber="134">
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(
|
||||
<figureCitation id="B5E72A19D705A924FF03F933FEF8E0AD" box="[206,273,1668,1692]" captionStart="Fig" captionStartId="8.[157,190,1441,1462]" captionTargetBox="[251,1322,268,1411]" captionTargetId="figure-183@8.[232,1518,246,1414]" captionTargetPageId="8" captionText="Fig. 5. A. Eisenia japonica vaga sub-sp. n. Busan holotype (H, DNA H5). B. Eisenia j. japonica (Michaelsen, 1892) Berlin syntype Nr. 2117 (as anticipated, DNA was not extractable from this older material), author’s sketch after Blakemore and Grygier (2011, fig. 2) that compares to Hamburg syntypes V119-V121 (see Blakemore and Grygier, 2011, fig. 3)." pageId="7" pageNumber="134">Fig. 5</figureCitation>
|
||||
<figureCitation id="B5E72A19D705A924FF03F933FEF8E0AD" box="[206,273,1668,1692]" captionStart="Fig" captionStartId="8.[157,190,1441,1462]" captionTargetBox="[251,1322,268,1411]" captionTargetId="figure-183@8.[232,1518,246,1414]" captionTargetPageId="8" captionText="Fig. 5. A. Eisenia japonica vaga sub-sp. n. Busan holotype (H, DNA H5). B. Eisenia j. japonica (Michaelsen, 1892) Berlin syntype Nr. 2117 (as anticipated, DNA was not extractable from this older material), author’s sketch after Blakemore and Grygier (2011, fig. 2) that compares to Hamburg syntypes V119-V121 (see Blakemore and Grygier, 2011, fig. 3)." figureDoi="http://doi.org/10.5281/zenodo.13144598" httpUri="https://zenodo.org/record/13144598/files/figure.png" pageId="7" pageNumber="134">Fig. 5</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -129,7 +130,7 @@ species-complex. It is not clear which part is closest, however it is remarkably
|
|||
specimen described by
|
||||
<bibRefCitation id="494D4B6DD705A924FB57FCF4FA8DE56D" author="Blakemore, R. J." box="[1178,1380,835,860]" pageId="7" pageNumber="134" pagination="105 - 132" refId="ref6132" refString="Blakemore, R. J. 2012 c. On Schmarda's lost worm and some newly found New Zealand species. Journal of Species Research 1 (2): 105 - 132 [Available from: http: // www. nibr. go. kr / english / event / journal _ spe _ 3. jsp]." type="journal article" year="2012">Blakemore (2012c</bibRefCitation>
|
||||
: fig. 3, 2013a: 40, 45). The DNA data is unequivocal that the Busan specimen objectively represents an as yet unknown taxon, and ditto sample WO20 having 99% agreement (
|
||||
<figureCitation id="B5E72A19D705A924FC8DFC73FC68E5ED" box="[832,897,964,988]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." pageId="7" pageNumber="134">Fig. 1</figureCitation>
|
||||
<figureCitation id="B5E72A19D705A924FC8DFC73FC68E5ED" box="[832,897,964,988]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="7" pageNumber="134">Fig. 1</figureCitation>
|
||||
and Appendix). The current specimen sequence is only 92-93% similar to GenBank specimens unidentifi- ed as “
|
||||
<taxonomicName id="EADC4D1FD705A924FCB0FBB3FBEEE22D" box="[893,1031,1028,1052]" class="Clitellata" family="Lumbricidae" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="134" phylum="Annelida" rank="family">Lumbricidae</taxonomicName>
|
||||
sp.” (AB607078) or as
|
||||
|
|
@ -168,7 +169,7 @@ are perhaps the tumid setae ab only on 25 and 26 (cf. on 22 and
|
|||
a
|
||||
<typeStatus id="F267883ED705A924FCF5FA31FC62E3AF" box="[824,907,1414,1438]" pageId="7" pageNumber="134" type="syntype">syntype</typeStatus>
|
||||
-
|
||||
<figureCitation id="B5E72A19D705A924FC6BFA31FC00E3AF" box="[934,1001,1414,1438]" captionStart="Fig" captionStartId="8.[157,190,1441,1462]" captionTargetBox="[251,1322,268,1411]" captionTargetId="figure-183@8.[232,1518,246,1414]" captionTargetPageId="8" captionText="Fig. 5. A. Eisenia japonica vaga sub-sp. n. Busan holotype (H, DNA H5). B. Eisenia j. japonica (Michaelsen, 1892) Berlin syntype Nr. 2117 (as anticipated, DNA was not extractable from this older material), author’s sketch after Blakemore and Grygier (2011, fig. 2) that compares to Hamburg syntypes V119-V121 (see Blakemore and Grygier, 2011, fig. 3)." pageId="7" pageNumber="134">Fig. 5</figureCitation>
|
||||
<figureCitation id="B5E72A19D705A924FC6BFA31FC00E3AF" box="[934,1001,1414,1438]" captionStart="Fig" captionStartId="8.[157,190,1441,1462]" captionTargetBox="[251,1322,268,1411]" captionTargetId="figure-183@8.[232,1518,246,1414]" captionTargetPageId="8" captionText="Fig. 5. A. Eisenia japonica vaga sub-sp. n. Busan holotype (H, DNA H5). B. Eisenia j. japonica (Michaelsen, 1892) Berlin syntype Nr. 2117 (as anticipated, DNA was not extractable from this older material), author’s sketch after Blakemore and Grygier (2011, fig. 2) that compares to Hamburg syntypes V119-V121 (see Blakemore and Grygier, 2011, fig. 3)." figureDoi="http://doi.org/10.5281/zenodo.13144598" httpUri="https://zenodo.org/record/13144598/files/figure.png" pageId="7" pageNumber="134">Fig. 5</figureCitation>
|
||||
), and the TP on 27 and 29 maybe less triangular. Differences from
|
||||
<taxonomicName id="EADC4D1FD705A924FBBEFA10FC9EE3EF" authority="(Oishi, 1934)" baseAuthorityName="Oishi" baseAuthorityYear="1934" class="Clitellata" family="Lumbricidae" genus="Eisenia" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="134" phylum="Annelida" rank="subSpecies" species="japonica" subSpecies="minuta">
|
||||
<emphasis id="1FA8EA8ED705A924FBBEFA10FAACE38F" box="[1139,1349,1447,1470]" italics="true" pageId="7" pageNumber="134">E. japonica minuta</emphasis>
|
||||
|
|
@ -196,7 +197,7 @@ had specimens from “Keiki-dô” identified as
|
|||
<bibRefCitation id="494D4B6DD705A924FBA3F910FB02E08E" author="Oishi, M." box="[1134,1259,1703,1727]" pageId="7" pageNumber="134" pagination="133 - 134" refId="ref7259" refString="Oishi, M. 1934. Earthworms. Zoological Magazine, Tokyo 46: 133 - 134 [in Japanese]." type="journal article" year="1934">Oishi, 1934</bibRefCitation>
|
||||
</taxonomicName>
|
||||
- they also had setae ab of 25 and 26 on oval papillae - although it is now certain neither that this complies with the Japanese originals nor that his identification is correct in light of the new information presented here. Moreover, that the specimen providing DNA sample WO19 (
|
||||
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|
||||
<figureCitation id="B5E72A19D705A924FB7BF8FFFB1AE151" box="[1206,1267,1864,1888]" captionStart="Fig" captionStartId="1.[159,192,1905,1926]" captionTargetBox="[267,1321,605,1871]" captionTargetId="figure-126@1.[257,1331,585,1877]" captionTargetPageId="1" captionText="Fig. 1. MEGA5 Maximum Likelihood phylogram of DNA code results for Busan specimens compared to similar taxa. Solid squares (■) are Busan samples; open circles (○) are unconfirmed GenBank names (from China, Korea or Japan); closed circles(●) are from the author’s Japanese studies.(Sample codes and provisional or new species names are explained in the Appendix legend)." figureDoi="http://doi.org/10.5281/zenodo.13144589" httpUri="https://zenodo.org/record/13144589/files/figure.png" pageId="7" pageNumber="134">Fig. 1</figureCitation>
|
||||
and Appendix) represents
|
||||
<taxonomicName id="EADC4D1FD705A924FC67F8DEFACDE1B1" authority="(Oishi, 1934)" baseAuthorityName="Oishi" baseAuthorityYear="1934" box="[938,1316,1896,1920]" class="Clitellata" family="Lumbricidae" genus="Eisenia" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="134" phylum="Annelida" rank="subSpecies" species="japonica" subSpecies="gigantica">
|
||||
<emphasis id="1FA8EA8ED705A924FC67F8DEFB78E1B1" box="[938,1169,1897,1920]" italics="true" pageId="7" pageNumber="134">E. japonica gigantica</emphasis>
|
||||
|
|
@ -216,7 +217,7 @@ nor
|
|||
<emphasis id="1FA8EA8ED70AA92BFD97F999FD56E074" box="[602,703,1582,1605]" italics="true" pageId="8" pageNumber="135">gigantica</emphasis>
|
||||
have since been found and reported sufficiently for comparison by subsequent Japanese workers.
|
||||
</paragraph>
|
||||
<caption id="79A36614D70AA92BFF50FA16FC5BE3DA" pageId="8" pageNumber="135" startId="8.[157,190,1441,1462]" targetBox="[251,1322,268,1411]" targetPageId="8" targetType="figure">
|
||||
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|
||||
<paragraph id="2D63369CD70AA92BFF50FA16FC5BE3DA" blockId="8.[157,1428,1441,1515]" pageId="8" pageNumber="135">
|
||||
<emphasis id="1FA8EA8ED70AA92BFF50FA16FF32E387" bold="true" box="[157,219,1441,1462]" pageId="8" pageNumber="135">Fig. 5.</emphasis>
|
||||
A.
|
||||
|
|
|
|||
|
|
@ -1,48 +1,49 @@
|
|||
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|
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@ -62,7 +63,7 @@
|
|||
<paragraph id="2D63369CD707A926FC9EF950FC57E0CE" blockId="5.[851,958,1767,1791]" box="[851,958,1767,1791]" pageId="5" pageNumber="132">
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(
|
||||
<figureCitation id="B5E72A19D707A926FC97F950FC58E0CE" box="[858,945,1767,1791]" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." pageId="5" pageNumber="132">Fig. 4A</figureCitation>
|
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<figureCitation id="B5E72A19D707A926FC97F950FC58E0CE" box="[858,945,1767,1791]" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." figureDoi="http://doi.org/10.5281/zenodo.13144596" httpUri="https://zenodo.org/record/13144596/files/figure.png" pageId="5" pageNumber="132">Fig. 4A</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
|
|
@ -71,7 +72,7 @@
|
|||
<paragraph id="2D63369CD707A926FCF5F890FC26E191" blockId="5.[824,1429,1831,1984]" pageId="5" pageNumber="132">
|
||||
<emphasis id="1FA8EA8ED707A926FCF5F890FC4BE10E" bold="true" box="[824,930,1831,1855]" pageId="5" pageNumber="132">Material.</emphasis>
|
||||
NIBR- IV0000261289 (DNA sample H3;
|
||||
<figureCitation id="B5E72A19D707A926FAA7F89FFCB0E151" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." pageId="5" pageNumber="132">Fig. 4A</figureCitation>
|
||||
<figureCitation id="B5E72A19D707A926FAA7F89FFCB0E151" captionStart="Fig" captionStartId="6.[157,190,1150,1171]" captionTargetBox="[266,1329,261,1115]" captionTargetId="figure-365@6.[247,1367,251,1279]" captionTargetPageId="6" captionText="Fig. 4. A. Amynthas masatakae (Beddard, 1892) Busan specimen showing spermathecal and male regions. B. Metaphire ryunome Blakemore, 2012 Busan specimen [boxed is X2 enlargement of its18rhs male pore that appears elevated, non-superficial, i.e.,=Metaphire]." figureDoi="http://doi.org/10.5281/zenodo.13144596" httpUri="https://zenodo.org/record/13144596/files/figure.png" pageId="5" pageNumber="132">Fig. 4A</figureCitation>
|
||||
) from Jungang Park, 26
|
||||
<superScript id="DAA99BD4D707A926FB9EF8F1FB89E167" attach="left" box="[1107,1120,1862,1878]" fontSize="7" pageId="5" pageNumber="132">th</superScript>
|
||||
April. Natural History Museum, London
|
||||
|
|
@ -89,7 +90,7 @@ Length
|
|||
. Clitellum 14-16. Spermathecal pores 7/8/9. Male pores on 18, superficial. GMs typically present, paired near spermathecal and male pores. Spermathecae with elongated, paprika-shaped diverticular bulbs on long stalks. Prostates (always?) reduced to muscular duct. Glands associated with GMs, where present. Caeca ventrally incised.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
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<paragraph id="2D63369CD704A925FF50FBC9FAF4E29F" blockId="6.[157,1427,1150,1198]" pageId="6" pageNumber="133">
|
||||
<emphasis id="1FA8EA8ED704A925FF50FBC9FF31E2A2" bold="true" box="[157,216,1150,1171]" pageId="6" pageNumber="133">Fig. 4.</emphasis>
|
||||
A.
|
||||
|
|
|
|||
Loading…
Reference in a new issue