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<document ID-DOI="10.11646/palaeoentomology.6.1.7" ID-ISSN="2624-2834" ID-Zenodo-Dep="7754940" ID-ZooBank="039F76C9-00B9-47A2-8589-1FE9E4705899" approvalRequired="6" approvalRequired_for_taxonomicNames="1" approvalRequired_for_treatments="5" checkinTime="1679381240438" checkinUser="plazi" docAuthor="KIM, JUNGGON, ROCA-CUSACHS, MARCOS, PHAM, THAI HONG PHAM &amp; JUNG, SUNGHOON" docDate="2023" docId="03B9878AFFA00C1CFF60FB00DBCEF956" docLanguage="en" docName="palaeoentomology.6.1.7.pdf" docOrigin="Palaeoentomology 6 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.1.7" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Cretanabis kerzhneri KIM &amp; ROCA-CUSACHS &amp; PHAM &amp; JUNG 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="FF80FFF2FFA30C1FFFE8FF94DF10FFFC" masterDocTitle="Cretanabis kerzhneri gen. et sp. nov., the oldest nabine genus and species (Hemiptera: Heteroptera: Nabidae) from the mid-Cretaceous Burmese amber" masterLastPageNumber="48" masterPageNumber="41" pageNumber="44" updateTime="1679381663387" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Cretanabis kerzhneri gen. et sp. nov., the oldest nabine genus and species (Hemiptera: Heteroptera: Nabidae) from the mid-Cretaceous Burmese amber</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>KIM, JUNGGON</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ROCA-CUSACHS, MARCOS</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PHAM, THAI HONG PHAM</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>JUNG, SUNGHOON</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>48</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.6.1.7</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.6.1.7</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">7754940</mods:identifier>
<mods:identifier type="ZooBank">039F76C9-00B9-47A2-8589-1FE9E4705899</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7754936" ID-Zenodo-Dep="7754936" LSID="urn:lsid:plazi:treatment:03B9878AFFA00C1CFF60FB00DBCEF956" httpUri="http://treatment.plazi.org/id/03B9878AFFA00C1CFF60FB00DBCEF956" lastPageNumber="44" pageId="3" pageNumber="44">
<subSubSection box="[136,632,1172,1199]" pageId="3" pageNumber="44" type="nomenclature">
<paragraph blockId="3.[136,632,1172,1234]" box="[136,632,1172,1199]" pageId="3" pageNumber="44">
<heading bold="true" box="[136,632,1172,1199]" fontSize="10" level="2" pageId="3" pageNumber="44" reason="3">
<emphasis bold="true" box="[136,632,1172,1199]" pageId="3" pageNumber="44">
<taxonomicName authority="Kim &amp; Jung" authorityName="KIM &amp; ROCA-CUSACHS &amp; PHAM &amp; JUNG" authorityYear="2023" box="[136,536,1172,1199]" class="Insecta" family="Nabidae" genus="Cretanabis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
<emphasis bold="true" box="[136,379,1172,1199]" italics="true" pageId="3" pageNumber="44">Cretanabis kerzhneri</emphasis>
Kim &amp; Jung
</taxonomicName>
<taxonomicNameLabel box="[543,632,1172,1198]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[136,252,1208,1234]" pageId="3" pageNumber="44" type="description">
<paragraph blockId="3.[136,632,1172,1234]" box="[136,252,1208,1234]" pageId="3" pageNumber="44">
(
<figureCitation box="[144,217,1208,1234]" captionStart="FIGURE 2" captionStartId="2.[136,235,1819,1843]" captionTargetBox="[159,1426,204,1789]" captionTargetId="figure-91@2.[151,1436,198,1795]" captionTargetPageId="2" captionText="FIGURE 2. Cretanabis kerzhneri sp. nov., holotype (CNUHHNF001). A, Dorsal habitus. B, Head. C, Antennae. D, Mid-leg. E, Hind-leg. F, Ventral habitus. Scale bars = 1 mm (A, F), 0.5 mm (BE)." figureDoi="http://doi.org/10.5281/zenodo.7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="3" pageNumber="44">Figs 2</figureCitation>
,
<figureCitation box="[229,243,1208,1234]" captionStart="FIGURE 3" captionStartId="3.[136,235,769,793]" captionTargetBox="[306,1277,210,728]" captionTargetId="figure-500@3.[295,1291,189,746]" captionTargetPageId="3" captionText="FIGURE 3. Habitus drawings of Cretanabis kerzhneri sp. nov. A, Dorsal habitus. B, Ventral habitus. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7754946" httpUri="https://zenodo.org/record/7754946/files/figure.png" pageId="3" pageNumber="44">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="44" type="materials_examined">
<paragraph blockId="3.[136,778,1279,2093]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[136,314,1279,1306]" pageId="3" pageNumber="44">Type material.</emphasis>
[
<materialsCitation collectionCode="CRBA" pageId="3" pageNumber="44" specimenCount="2" typeStatus="holotype">
<collectionCode box="[331,413,1280,1306]" pageId="3" pageNumber="44">CRBA</collectionCode>
]
<typeStatus box="[430,539,1280,1306]" pageId="3" pageNumber="44">Holotype</typeStatus>
:
<typeStatus box="[553,657,1280,1306]" pageId="3" pageNumber="44">Holotype</typeStatus>
male in a 25×10×
<quantity box="[225,293,1315,1342]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="44" unit="mm" value="3.0">3 mm</quantity>
cabochon-shaped piece of amber. Dorsal and ventral surfaces flat
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="44" type="etymology">
<paragraph blockId="3.[136,778,1279,2093]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[189,324,1387,1413]" pageId="3" pageNumber="44">Etymology.</emphasis>
Named in memory of Izya M. Kerzhner, a taxonomist of
<taxonomicName box="[330,427,1423,1449]" class="Insecta" family="Nabidae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="family">Nabidae</taxonomicName>
; treated as a noun in genitive case.
</paragraph>
</subSubSection>
<subSubSection box="[189,469,1494,1521]" pageId="3" pageNumber="44" type="diagnosis">
<paragraph blockId="3.[136,778,1279,2093]" box="[189,469,1494,1521]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[189,312,1494,1520]" pageId="3" pageNumber="44">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="44" type="materials_examined">
<paragraph blockId="3.[136,778,1279,2093]" pageId="3" pageNumber="44">
<materialsCitation country="Myanmar" location="Uncertain" pageId="3" pageNumber="44" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis bold="true" box="[189,500,1530,1556]" pageId="3" pageNumber="44">
<typeStatus box="[189,248,1530,1556]" pageId="3" pageNumber="44">Type</typeStatus>
locality and horizon.
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03B9878AFFA00C1CFF60FB00DBCEF956:8ECF6047FFA00C1CFE13FA6EDD79F9E8" box="[507,617,1530,1556]" country="Myanmar" name="Uncertain" pageId="3" pageNumber="44" stateProvince="Kachin">Uncertain</location>
locality in the
<collectingRegion box="[136,286,1566,1592]" country="Myanmar" name="Kachin" pageId="3" pageNumber="44">Kachin State</collectingRegion>
, northern
<collectingCountry box="[404,517,1566,1592]" name="Myanmar" pageId="3" pageNumber="44">Myanmar</collectingCountry>
; mid-Cretaceous (
<bibRefCitation author="Mao, Y. Y. &amp; Liang, K. &amp; Su, Y. T. &amp; Li, J. G. &amp; Rao, X. &amp; Zhang, H. &amp; Xia, F. Y. &amp; Fu, Y. Z. &amp; Cai, C. Y. &amp; Huang, D. Y." pageId="3" pageNumber="44" pagination="91 - 103" refId="ref4165" refString="Mao, Y. Y., Liang, K., Su, Y. T., Li, J. G., Rao, X., Zhang, H., Xia, F. Y., Fu, Y. Z., Cai, C. Y. &amp; Huang, D. Y. (2018) Various amberground marine animals on Burmese amber with discussions on its age. Palaeoentomology, 1 (1), 91 - 103. https: // doi. org / 10.11646 / palaeoentomology. 1.1.11" type="journal article" year="2018">
Mao
<emphasis box="[136,185,1601,1627]" italics="true" pageId="3" pageNumber="44">et al</emphasis>
., 2018
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="44" type="description">
<paragraph blockId="3.[136,778,1279,2093]" lastBlockId="3.[810,1452,849,1706]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[189,335,1637,1663]" pageId="3" pageNumber="44">Description.</emphasis>
Instar III (supposed to belong to this instar). Body ovoid, length
<quantity box="[446,550,1673,1700]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" pageId="3" pageNumber="44" unit="mm" value="2.36">2.36 mm</quantity>
.
<emphasis box="[561,684,1673,1699]" italics="true" pageId="3" pageNumber="44">Coloration</emphasis>
. Mostly of brown coloration, nearly unicolorous.
<emphasis italics="true" pageId="3" pageNumber="44">Surface and vestiture</emphasis>
. Body glabrous, beset partly with long spines on frons, clypeus and mandibular plates of head, thorax laterally, and femora and tibia of legs; third and fourth antennomeres covered with long setae.
<emphasis box="[591,694,1852,1878]" italics="true" pageId="3" pageNumber="44">Structure</emphasis>
. Head: hypognathous, length less than width, more than pronotal midline; vertex narrow, narrower than a single compound eye width; ocelli absent; antennae cylindrical, linear, shorter than body length; first antennomere shortest, shorter than head width; second antennomere as long as head width and third antennomere, slightly thicker than third and fourth antennomeres; fourth antennomere longest; proportion of first to fourth segments 0.23: 0.48: 0.43: 0.61; labium missing. Thorax: pronotum trapezoid, anterior width shorter than head width, pronotal midline length as long as 1/2 of basal width,longer than mesothorax midline; mesothorax without distinct wing pad, posterior margin slightly sinuate, posterior width more than pronotal posterior width; legs generally slender; fore femur slightly incrassate and as long as fore tibia; apical part of tibia with dense spines, less than
<quantity box="[1139,1199,1174,1200]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="44" unit="in" value="10.0">10 in</quantity>
number; tarsus threesegmented, length less than 1/2 tibia in midleg, and less than 1/3 tibia in hindleg. Abdomen: ovoid, abdominal segment VII widest. Genitalia: not developed.
</paragraph>
<paragraph blockId="3.[810,1452,849,1706]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[863,1045,1319,1345]" pageId="3" pageNumber="44">Measurements.</emphasis>
(in mm) body length: 2.36; head length 0.51; head width including compound eyes: 0.53; vertex width: 0.11; first antennomere: 0.23; second antennomere: 0.48; third antennomere: 0.43; fourth antennomere: 0.61; anterior pronotal width: 0.28; pronotal midline length: 0.33; basal pronotal maximal width (straight): 0.44; hindleg (femur: tibia: tarsus): 0.88: 1.77: 0.43.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="44" type="discussion">
<paragraph blockId="3.[810,1452,849,1706]" pageId="3" pageNumber="44">
<emphasis bold="true" box="[863,979,1607,1633]" pageId="3" pageNumber="44">Remarks.</emphasis>
The nymphal instar III was determined by the shape of body and wing pad development (no wing pad in this fossil specimen (
<figureCitation box="[1122,1194,1679,1706]" captionStart="FIGURE 2" captionStartId="2.[136,235,1819,1843]" captionTargetBox="[159,1426,204,1789]" captionTargetId="figure-91@2.[151,1436,198,1795]" captionTargetPageId="2" captionText="FIGURE 2. Cretanabis kerzhneri sp. nov., holotype (CNUHHNF001). A, Dorsal habitus. B, Head. C, Antennae. D, Mid-leg. E, Hind-leg. F, Ventral habitus. Scale bars = 1 mm (A, F), 0.5 mm (BE)." figureDoi="http://doi.org/10.5281/zenodo.7754944" httpUri="https://zenodo.org/record/7754944/files/figure.png" pageId="3" pageNumber="44">Figs 2</figureCitation>
,
<figureCitation box="[1206,1220,1680,1706]" captionStart="FIGURE 3" captionStartId="3.[136,235,769,793]" captionTargetBox="[306,1277,210,728]" captionTargetId="figure-500@3.[295,1291,189,746]" captionTargetPageId="3" captionText="FIGURE 3. Habitus drawings of Cretanabis kerzhneri sp. nov. A, Dorsal habitus. B, Ventral habitus. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7754946" httpUri="https://zenodo.org/record/7754946/files/figure.png" pageId="3" pageNumber="44">3</figureCitation>
)).
</paragraph>
</subSubSection>
</treatment>
</document>