treatments-xml/data/03/DB/87/03DB878C37097E4EFF706DEBFE2AE87C.xml

<document id="098602E7620AA85E4BFA0FAA8D4B151F" ID-DOI="10.5281/zenodo.399638" ID-GBIF-Dataset="6916e2ea-19c9-4427-85bf-f43840857485" ID-ISSN="1175­5326" ID-Zenodo-Dep="399638" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459810352782" checkinUser="plazi" docAuthor="Sz, Sándor Cs Ő, Radchenko, Alexander &amp; Schulz, Andreas" docDate="2007" docId="03DB878C37097E4EFF706DEBFE2AE87C" docLanguage="en" docName="zt01405p038.pdf" docOrigin="Zootaxa 1405" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tetramorium alternans Santschi 1929, n. sp." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFE2FFF437007E40FFE76B3BFFA5EF31" masterDocTitle="Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="10" updateTime="1698220957542" updateUser="plazi">
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<mods:title id="C93D5D23F12A7E8724FFC512CAD6F504">Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="D97CE20B09C211D7DF5896052B8CEE66">Sz, Sándor Cs Ő</mods:namePart>
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<mods:namePart id="5E0E3244D668B94E427CC5CEC7CAE0EB">Radchenko, Alexander</mods:namePart>
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<mods:namePart id="0E3CCD280A4610E576B1816916C9B133">Schulz, Andreas</mods:namePart>
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<mods:title id="895AA2CF8DDEF3838712640E5481553B">Zootaxa</mods:title>
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<mods:date id="08F8759E4867BF4319BF3882776509D7">2007</mods:date>
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<subSubSection id="C368651137097E49FF706DEBFCB9E9DB" box="[151,796,1744,1770]" pageId="9" pageNumber="10" type="key">
<paragraph id="8BCD369A37097E49FF706DEBFCB9E9DB" blockId="9.[151,796,1744,1770]" box="[151,796,1744,1770]" pageId="9" pageNumber="10">
<heading id="D08581F637097E49FF706DEBFCB9E9DB" bold="true" box="[151,796,1744,1770]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<keyStep id="30862E3F37097E49FF706DEBFCB9E9DB" box="[151,796,1744,1770]" pageId="9" pageNumber="10">
<key id="FFE361AB37097E49FF706DEBFCB9E9DB" box="[151,796,1744,1770]" pageId="9" pageNumber="10">
<emphasis id="B906EA8837097E49FF706DEBFCB9E9DB" bold="true" box="[151,796,1744,1770]" pageId="9" pageNumber="10">
<keyLead id="308395AF37097E49FF706DEBFCB9E9DB" box="[151,796,1744,1770]" pageId="9" pageNumber="10">
1. 
<taxonomicName id="4C724D1937097E49FF536DEBFD22E9DB" authority="Santschi, 1929" authorityName="Santschi" authorityYear="1929" box="[180,647,1744,1770]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA8837097E49FF536DEBFE69E9DB" bold="true" box="[180,460,1744,1770]" italics="true" pageId="9" pageNumber="10">Tetramorium alternans</emphasis>
<bibRefCitation id="EFE34B6B37097E49FE346DEBFD22E9DB" author="Santschi" box="[467,647,1744,1770]" pageId="9" pageNumber="38" refString="Santschi, F. (1929) Fourmis du Maroc, d'Algerie et de Tunisie. Bulletin et Annales de la Societe Entomologique de Belge, 69, 138 - 165." type="journal article" year="1929">Santschi, 1929</bibRefCitation>
</taxonomicName>
(figs 11–17)
</keyLead>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368651137097E4AFF706C20FC5BED8D" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BCD369A37097E49FF706C20FE1FE843" blockId="9.[151,1436,1819,2002]" pageId="9" pageNumber="10">
<taxonomicName id="4C724D1937097E49FF706C20FE31E803" ID-CoL="55PJV" box="[151,404,1819,1842]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caespitum">
<emphasis id="B906EA8837097E49FF706C20FE31E803" box="[151,404,1819,1842]" italics="true" pageId="9" pageNumber="10">Tetramorium caespitum</emphasis>
</taxonomicName>
<taxonomicName id="4C724D1937097E49FE7C6C20FCA3E803" authority="Santschi, 1929: 150" authorityName="Santschi" authorityPageNumber="150" authorityYear="1929" box="[411,774,1819,1842]" pageId="9" pageNumber="10" rank="variety" variety="alternans">
var. 
<emphasis id="B906EA8837097E49FE206C20FD89E803" box="[455,556,1819,1842]" italics="true" pageId="9" pageNumber="10">alternans</emphasis>
<bibRefCitation id="EFE34B6B37097E49FDD46C20FD76E803" author="Santschi" box="[563,723,1819,1842]" pageId="9" pageNumber="38" refString="Santschi, F. (1929) Fourmis du Maroc, d'Algerie et de Tunisie. Bulletin et Annales de la Societe Entomologique de Belge, 69, 138 - 165." type="journal article" year="1929">Santschi, 1929</bibRefCitation>
: 150
</taxonomicName>
(/); 
<typeStatus id="54C9883837097E49FCDD6C20FCD8E803" box="[826,893,1819,1842]" pageId="9" pageNumber="10">TYPE</typeStatus>
MATERIAL: 
<typeStatus id="54C9883837097E49FBFE6C20FB14E803" box="[1049,1201,1819,1842]" pageId="9" pageNumber="10" type="lectotype">LECTOTYPE</typeStatus>
/ and PARALECTO­ 
<typeStatus id="54C9883837097E49FF206C00FEAFE863" box="[199,266,1851,1874]" pageId="9" pageNumber="10">TYPE</typeStatus>
// designated below (3/ / NHMB); for the locality details see 
<typeStatus id="54C9883837097E49FC5F6C00FBBFE863" box="[952,1050,1851,1874]" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
designation; raised to species rank: 
<bibRefCitation id="EFE34B6B37097E49FF206C60FE2AE843" author="Gusten" box="[199,399,1883,1906]" pageId="9" pageNumber="37" refString="Gusten, R., Schulz, A. &amp; Sanetra, M. (2006) Redescription of Tetramorium forte Forel, 1904 (Insecta: Hymenoptera: Formicidae), a western Mediterranean ant species. Zootaxa, 1310, 1 - 35." type="journal article" year="2006">
Güsten 
<emphasis id="B906EA8837097E49FEFE6C60FEEAE843" box="[281,335,1883,1906]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2006
</bibRefCitation>
: 27;
</paragraph>
<paragraph id="8BCD369A37097E49FF706C40FB42E8E3" blockId="9.[151,1436,1819,2002]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB6211B437097E49FF706C40FB42E8E3" pageId="9" pageNumber="10">
<taxonomicName id="4C724D1937097E49FF706C40FE37E8A3" box="[151,402,1915,1938]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="biskrensis">
<emphasis id="B906EA8837097E49FF706C40FE37E8A3" box="[151,402,1915,1938]" italics="true" pageId="9" pageNumber="10">Tetramorium biskrensis</emphasis>
</taxonomicName>
<taxonomicName id="4C724D1937097E49FE706C40FD5EE8A3" authority="Menozzi, 1934: 162" authorityName="Menozzi" authorityPageNumber="162" authorityYear="1934" box="[407,763,1915,1938]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="variety" species="biskrensis" variety="kahenae">
var. 
<emphasis id="B906EA8837097E49FE236C40FDB8E8A3" box="[452,541,1915,1938]" italics="true" pageId="9" pageNumber="10">kahenae</emphasis>
<treatmentCitation id="0AD3108B37097E49FDC46C40FD5EE8A3" author="Menozzi" box="[547,763,1915,1938]" page="162" pageId="9" pageNumber="10" year="1934">
<bibRefCitation id="EFE34B6B37097E49FDC46C40FD63E8A3" author="Menozzi" box="[547,710,1915,1938]" pageId="9" pageNumber="38" refString="Menozzi, C. (1934) Reperti mirmecofaunistici raccolte dal Prof. L. di Caporiacco nelle oasi di Cufra e in altre localite del deserto Libico. Atti della Societa dei Naturalisti e Matematici di Modena, 13, 153 - 166." type="journal article" year="1934">Menozzi, 1934</bibRefCitation>
: 162
</treatmentCitation>
</taxonomicName>
; first available use of 
<taxonomicName id="4C724D1937097E49FC036C40FB45E8A3" ID-CoL="55PJV" box="[996,1248,1915,1938]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caespitum">
<emphasis id="B906EA8837097E49FC036C40FB45E8A3" box="[996,1248,1915,1938]" italics="true" pageId="9" pageNumber="10">Tetramorium caespitum</emphasis>
</taxonomicName>
<taxonomicName id="4C724D1937097E49FB016C40FE5CE883" authority="Santschi, 1918: 155" authorityName="Santschi" authorityPageNumber="155" authorityYear="1918" pageId="9" pageNumber="10" rank="variety" subSpecies="biskrensis" variety="kahenae">
st. 
<emphasis id="B906EA8837097E49FAE26C40FACAE8A3" box="[1285,1391,1915,1938]" italics="true" pageId="9" pageNumber="10">biskrensis</emphasis>
var. 
<emphasis id="B906EA8837097E49FF206CA0FE85E883" box="[199,288,1947,1970]" italics="true" pageId="9" pageNumber="10">kahenae</emphasis>
<treatmentCitation id="0AD3108B37097E49FEC16CA0FE5CE883" author="Santschi" box="[294,505,1947,1970]" page="155" pageId="9" pageNumber="10" year="1918">
<bibRefCitation id="EFE34B6B37097E49FEC16CA0FE62E883" author="Santschi" box="[294,455,1947,1970]" pageId="9" pageNumber="38" refString="Santschi, F. (1918) Noveaux Tetramorium africains. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord, 9, 153 - 156." type="journal article" year="1918">Santschi, 1918</bibRefCitation>
: 155
</treatmentCitation>
</taxonomicName>
(/); 
<typeStatus id="54C9883837097E49FDCA6CA0FDD5E883" box="[557,624,1947,1970]" pageId="9" pageNumber="10">TYPE</typeStatus>
MATERIAL: 
<typeStatus id="54C9883837097E49FCEC6CA0FC06E883" box="[779,931,1947,1970]" pageId="9" pageNumber="10" type="lectotype">LECTOTYPE</typeStatus>
/ designated below, for locality details see 
<typeStatus id="54C9883837097E49FA936CA0FEAFE8E3" pageId="9" pageNumber="10" type="lectotype">lectotype</typeStatus>
designation (1/ / MCSN); 
<taxonomicName id="4C724D1937097E49FDCA6C80FB8AE8E3" ID-CoL="5LGG9" authority="Mei 1995: 764" authorityName="Mei" authorityPageNumber="764" authorityYear="1995" box="[557,1071,1979,2002]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="biskrense" subSpecies="kahenae">
<emphasis id="B906EA8837097E49FDCA6C80FC24E8E3" box="[557,897,1979,2002]" italics="true" pageId="9" pageNumber="10">Tetramorium biskrense kahenae</emphasis>
: 
<treatmentCitation id="0AD3108B37097E49FC686C80FB8AE8E3" author="Mei" box="[911,1071,1979,2002]" page="764" pageId="9" pageNumber="10" year="1995">
<bibRefCitation id="EFE34B6B37097E49FC686C80FC5CE8E3" author="Mei" box="[911,1017,1979,2002]" lastPageId="9" lastPageNumber="38" pageId="9" pageNumber="37" refString="Mei, M. (1995) Arthropoda di Lampedusa, Linosa e Pantelleria (Canale di Sicilia, Mar Mediterraneo). Hymenoptera, Formicidae (con diagnosi di due nuove specie). Naturalista Siciliano, 19 (Suppl.), 753 - 772." type="journal article" year="1995">Mei 1995</bibRefCitation>
: 764
</treatmentCitation>
</taxonomicName>
; 
<emphasis id="B906EA8837097E49FBDD6C81FB42E8E3" bold="true" box="[1082,1255,1978,2002]" pageId="9" pageNumber="10">New synonymy</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCD369A370A7E4AFF706BACFC5BED8D" blockId="10.[151,1440,151,1867]" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFF706BACFE6CEF80" bold="true" box="[151,457,151,177]" pageId="10" pageNumber="11">Redescription of worker</emphasis>
(figs 14–17). Small to medium size, CS 733 [652, 820]. Whole body and appendages greyish black to black.Head nearly square CL/CW 1.01 [0.97, 1.05], with very feebly convex sides, straight, or slightly concave occipital margin and narrowly rounded occipital corners. Eyes small, EYE 0.168 [0.155, 0.176]. Frons moderately wide, FR/CS 0.39 [0.37, 0.41], frontal lobes as wide as frons, FL/FR 1.0 [1.0, 1.0]. Scape very short, SL/CS 0.71 [0.69, 0.74], without dorsal carina basally, surface feebly costulate. Promesonotal dorsum slightly convex, metanotal groove shallow, but distinct. Propodeal teeth moderately long, slightly curving upwards. Dorsal surface of petiole steeply rounded backward, NOH/NOL 1.03 [0.94, 1.13], petiole relatively high, PEH/NOL 1.84 [1.68, 2.04]. Whole body and appendages greyish black to black. General appearance coarsely rugose, ground surface microreticulate. Head dorsum longitudinally rugose and microreticulate, occiput and sides rugoso­reticulate and microreticulate. Alitrunk dorsum rugose or rugoso­reticulate and microreticulate. Mesopleuron usually coarsely microreticulate. Dorsum of petiolar node ruguloso­reticulate and microreticulate, medially microreticulate. Dorsum of postpetiole microreticulate. Polygonal striation continuous on 1st gastral tergite (see fig. 8.). Ventral surface of head with several short and few moderately long setae arising posteriorly to buccal cavity (see fig. 5.).
</paragraph>
</subSubSection>
<subSubSection id="C3686511370A7E4AFF2169F1FD88E902" pageId="10" pageNumber="11" type="description">
<paragraph id="8BCD369A370A7E4AFF2169F1FEF3EBC1" blockId="10.[151,1440,151,1867]" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFF2169F1FE66EDD5" bold="true" box="[198,451,714,740]" pageId="10" pageNumber="11">Description of Gyne</emphasis>
(figs 11–13). Small size, CS 999 [948, 1025]. Whole body and appendages greyish black. Head wider than long, CL/CW 0.92 [0.90, 0.95], with straight subparallel sides, straight occipital margin and narrowly rounded occipital corners. Frons moderately wide, FR/CS 0.41 [0.40, 0.42], frontal lobes as wide as frons, FL/FR 1.0 [1.0, 1.0]. Scape very short, SL/CS 0.69 [0.65, 0.71], without dorsal carina basally, moderately smooth and shiny. Head wider than scutum, MW/CS 0.94 [0.92, 0.97]. Propodeal teeth very short. Dorsal crest of petiolar node in frontal view straight. Petiolar node dorsum steeply rounded backward. Petiole and postpetiole relatively narrow, WAIST 0.93 [0.83, 1.0]. General appearance rugulose, ground surface smooth, more or less shiny. Head dorsum, occiput and sides ruguloso­reticulate, ground surface feebly microreticulate. Frons longitudinally rugulose and feebly microreticulate. Scutum and scutellum longitudinally rugulose, scutellum more or less smooth medially. Sides of alitrunk ruguloso­reticulate and feebly microreticulate, ventral part of katepisternum always smooth. Dorsum of petiolar node feebly reticulate and smooth, dorsum of postpetiole usually smooth. Polygonal striation disrupted on 1st gastral tergite, occasionally superficially striate basally. Ventral surface of head with several short and few longer setae, arising posteriorly to buccal cavity.
</paragraph>
<paragraph id="8BCD369A370A7E4AFF216FC6FD88E902" blockId="10.[151,1440,151,1867]" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFF216FC6FE76EA26" bold="true" box="[198,467,1277,1303]" pageId="10" pageNumber="11">Description of Male.</emphasis>
Whole body and appendages black. Head with feebly convex sides, slightly rounded occipital margin and widely rounded occipital corners. Scutum wider than head. Propodeal teeth very short, propodeum angulate in profile. Dorsal crest of petiolar node with sharp transversal edge, slightly emarginated in frontal view. Head, alitrunk and waist finely sculptured, ground surface microreticulate. Head feebly reticulate ground surface microreticulate, dull. Scutum feebly rugulose, laterally and anteriorly smooth and shiny. Scutellum longitudinally rugulose, usually shiny medially. Sides of alitrunk, smooth and shiny. Dorsum of petiolar node finely reticulate and microreticulate, postpetiole smooth and shiny. Polygonal striation disrupted on 1st gastral tergite.
</paragraph>
</subSubSection>
<subSubSection id="C3686511370A7E4BFF216D7BFC9FE833" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BCD369A370A7E4AFF216D7BFEA7E87A" blockId="10.[151,1440,151,1867]" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFF216D7BFE10E96B" bold="true" box="[198,437,1600,1626]" pageId="10" pageNumber="11">Material examined</emphasis>
(8 nest series including 
<specimenCount id="9D74FD13370A7E4AFD3C6D7AFCC1E96A" box="[731,868,1601,1627]" pageId="10" pageNumber="11" type="worker">34 workers</specimenCount>
, 6 gynes and 
<specimenCount id="9D74FD13370A7E4AFBED6D7AFBC9E96A" box="[1034,1132,1601,1627]" pageId="10" pageNumber="11" type="male">3 males</specimenCount>
). 
<collectingCountry id="F365760A370A7E4AFB6E6D7BFAB7E96B" box="[1161,1298,1600,1626]" name="Algeria" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFB6E6D7BFAB7E96B" bold="true" box="[1161,1298,1600,1626]" pageId="10" pageNumber="11">ALGERIA</emphasis>
</collectingCountry>
– Takersan 0 4.1893 leg. Anonymous (4/ / 
<collectionCode id="ED63AE5F370A7E4AFDE76D52FDFAE9B2" LSID="urn:lsid:biocol.org:col:34988" box="[512,607,1641,1667]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
); 
<collectingCountry id="F365760A370A7E4AFD956D53FCB7E9B3" box="[626,786,1640,1666]" name="Morocco" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFD956D53FCB7E9B3" bold="true" box="[626,786,1640,1666]" pageId="10" pageNumber="11">MOROCCO</emphasis>
</collectingCountry>
– Moyen Atlas, Rd 3211 14 Rkm N. Rd 3485, 40 km S. Ain Leuh, 1100 masl, leg. Aßmuth, Güsten, Sanetra, Schulz, Schumann (6/ / 
<collectionCode id="ED63AE5F370A7E4AFBD16DAAFB21E99A" box="[1078,1156,1681,1707]" pageId="10" pageNumber="11">PCAS</collectionCode>
); 
<collectingCountry id="F365760A370A7E4AFB7C6DABFAB0E99B" box="[1179,1301,1680,1706]" name="Tunisia" pageId="10" pageNumber="11">
<emphasis id="B906EA88370A7E4AFB7C6DABFAB0E99B" bold="true" box="[1179,1301,1680,1706]" pageId="10" pageNumber="11">TUNISIA</emphasis>
</collectingCountry>
– Hammamet, 
<date id="FFCC105A370A7E4AFF346D82FEFEE9E2" box="[211,347,1721,1747]" pageId="10" pageNumber="11" value="1982-04-26">26.04.1982</date>
. nr. 8. leg. H. Sheratin (2/, 1Ψ, 1ɗ / 
<collectionCode id="ED63AE5F370A7E4AFCFC6D82FCDCE9E2" LSID="urn:lsid:biocol.org:col:34991" box="[795,889,1721,1747]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM</collectionCode>
), nr. 9, 
<date id="FFCC105A370A7E4AFC306D82FBF8E9E2" box="[983,1117,1721,1747]" pageId="10" pageNumber="11" value="1982-04-27">27.04.1982</date>
. leg. H. Sheratin (7/, 3Ψ / 
<collectionCode id="ED63AE5F370A7E4AFF706DDAFF50E9CA" box="[151,245,1761,1787]" pageId="10" pageNumber="11">MSNM</collectionCode>
), nr. 23, 
<date id="FFCC105A370A7E4AFE866DDAFE4DE9CA" box="[353,488,1761,1787]" pageId="10" pageNumber="11" value="1982-04-30">30.04.1982</date>
. leg. H. Sheratin (2/, 1Ψ, 2ɗ / 
<collectionCode id="ED63AE5F370A7E4AFC866DDAFC65E9CA" box="[865,960,1761,1787]" pageId="10" pageNumber="11">MSNM</collectionCode>
); Hammamet, Sousse, 0 3.1974. nr. 804. leg. Novellini (4/ / 
<collectionCode id="ED63AE5F370A7E4AFE666C32FE45E812" box="[385,480,1801,1827]" pageId="10" pageNumber="11">MSNM</collectionCode>
); Kabylie, 
<date id="FFCC105A370A7E4AFD856C32FD69E812" box="[610,716,1801,1827]" pageId="10" pageNumber="11" value="65-05-31">31.05.65</date>
. leg. Cagniant (4/ / 
<collectionCode id="ED63AE5F370A7E4AFC596C32FBB8E812" box="[958,1053,1801,1827]" pageId="10" pageNumber="11">MSNM</collectionCode>
); Sfax, 1904. leg. Bíró (5/; 1Ψ / 
<collectionCode id="ED63AE5F370A7E4AFF706C0AFF53E87A" LSID="urn:lsid:biocol.org:col:34991" box="[151,246,1841,1867]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM</collectionCode>
).
</paragraph>
<caption id="DF0D6612370B7E4BFF706D66FDE7E985" httpUri="https://zenodo.org/record/399640/files/figure.png" pageId="11" pageNumber="12" targetBox="[264,1318,180,1608]" targetPageId="11">
<paragraph id="8BCD369A370B7E4BFF706D66FDE7E985" blockId="11.[151,1437,1629,1716]" pageId="11" pageNumber="12">
<emphasis id="B906EA88370B7E4BFF706D66FEFBE944" bold="true" box="[151,350,1629,1653]" pageId="11" pageNumber="12">FIGURES 11–17.</emphasis>
<taxonomicName id="4C724D19370B7E4BFE816D66FCACE945" authority="Santschi, 1929" authorityName="Santschi" authorityYear="1929" box="[358,777,1629,1652]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370B7E4BFE816D66FDFBE945" box="[358,606,1629,1652]" italics="true" pageId="11" pageNumber="12">Tetramorium alternans</emphasis>
Santschi, 1929
</taxonomicName>
. Gyne: alitrunk petiole and postpetiole, Fig. 11. dorsal view, Fig. 12. lateral view, Fig. 13. head. Worker: Fig. 14. head. Alitrunk petiole and postpetiole, Fig. 15. dorsal view, Fig. 16. lateral view, Fig. 17. scape, dorsal view.
</paragraph>
</caption>
<paragraph id="8BCD369A370B7E4BFF216DD3FC9FE833" blockId="11.[151,1437,1768,2038]" box="[198,826,1768,1794]" pageId="11" pageNumber="12">
<emphasis id="B906EA88370B7E4BFF216DD3FE2CE833" bold="true" box="[198,393,1768,1794]" pageId="11" pageNumber="12">Morphometrics</emphasis>
(
<specimenCount id="9D74FD13370B7E4BFE7E6DD3FDB8E833" box="[409,541,1768,1794]" pageId="11" pageNumber="12" type="worker">34 workers</specimenCount>
and 5 gynes measured).
</paragraph>
</subSubSection>
<subSubSection id="C3686511370B7E4CFF216C2BFAD8EBAB" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BCD369A370B7E4BFF216C2BFA25E8C3" blockId="11.[151,1437,1768,2038]" pageId="11" pageNumber="12">
<emphasis id="B906EA88370B7E4BFF216C2BFEE3E81B" bold="true" box="[198,326,1808,1834]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Workers of 
<taxonomicName id="4C724D19370B7E4BFE3A6C2BFDC9E818" box="[477,620,1808,1833]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370B7E4BFE3A6C2BFDC9E818" box="[477,620,1808,1833]" italics="true" pageId="11" pageNumber="12">T. alternans</emphasis>
</taxonomicName>
can be separated from related species by the lack of psammophore, relatively small eyes, (EYE, 
<tableCitation id="C6F00321370B7E4BFE096C03FDE0E863" box="[494,581,1848,1874]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="11" pageNumber="12" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.), very short scape (SL/CS, 
<tableCitation id="C6F00321370B7E4BFC7D6C03FC54E863" box="[922,1009,1848,1874]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="11" pageNumber="12" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.) without dorsal carina basally, less coarse body sculpture (i.e. dorsum of petiolar node is feebly ruguloso­reticulate and microreticulate), rounded petiolar node (NOH/NOL and PEH/NOL, 
<tableCitation id="C6F00321370B7E4BFD6C6CB3FD46E893" box="[651,739,1928,1954]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="11" pageNumber="12" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.). Workers of 
<taxonomicName id="4C724D19370B7E4BFC6C6CB3FBB2E890" box="[907,1047,1928,1953]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370B7E4BFC6C6CB3FBB2E890" box="[907,1047,1928,1953]" italics="true" pageId="11" pageNumber="12">T. alternans</emphasis>
</taxonomicName>
mostly resemble those of 
<taxonomicName id="4C724D19370B7E4BFAAB6CB3FF6BE8F8" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sanetrai" status="sp. nov.">
<emphasis id="B906EA88370B7E4BFAAB6CB3FF6BE8F8" italics="true" pageId="11" pageNumber="12">T. sanetrai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370B7E4BFF326C8BFEB2E8FB" box="[213,279,1968,1994]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B906EA88370B7E4BFF326C8BFEB2E8FB" bold="true" box="[213,279,1968,1994]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, but petiole and frons characters (NOH/NOL, PEH/NOL,FR/CS, 
<tableCitation id="C6F00321370B7E4BFBF16C8BFBCEE8FB" box="[1046,1131,1968,1994]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="11" pageNumber="12" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.) give appropriate separation between them. To separate these species see Discriminant D(2 
<emphasis id="B906EA88370B7E4BFC426CD3FC08E8C7" box="[933,941,2024,2038]" italics="true" pageId="11" pageNumber="12">b</emphasis>
) function given under 
<taxonomicName id="4C724D19370B7E4BFB5D6CE3FA91E8C0" box="[1210,1332,2008,2033]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sanetrai" status="sp. nov.">
<emphasis id="B906EA88370B7E4BFB5D6CE3FA91E8C0" box="[1210,1332,2008,2033]" italics="true" pageId="11" pageNumber="12">T. sanetrai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370B7E4BFAD96CE3FA25E8C3" box="[1342,1408,2008,2034]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B906EA88370B7E4BFAD96CE3FA25E8C3" bold="true" box="[1342,1408,2008,2034]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216BA3FD3AEE33" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
Gynes of 
<taxonomicName id="4C724D19370C7E4CFEDD6BA3FE62EF80" box="[314,455,152,177]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370C7E4CFEDD6BA3FE62EF80" box="[314,455,152,177]" italics="true" pageId="12" pageNumber="13">T. alternans</emphasis>
</taxonomicName>
can be recognised by the lack of psammophore, very short and smooth scape (SL/ CS, 
<tableCitation id="C6F00321370C7E4CFF2D6BFBFE83EFEB" box="[202,294,192,218]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="12" pageNumber="13" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), wide scutum (MW/CS, 
<tableCitation id="C6F00321370C7E4CFDB96BFBFD1CEFEB" box="[606,697,192,218]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="12" pageNumber="13" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), relatively narrow petiole and postpetiole (WAIST, 
<tableCitation id="C6F00321370C7E4CFAC86BFBFA2FEFEB" box="[1327,1418,192,218]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="12" pageNumber="13" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), and feebly reticulate petiole and postpetiole.
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216A2BFC73EE1B" blockId="12.[151,1439,152,1258]" box="[198,982,272,298]" pageId="12" pageNumber="13">
For further combination of morphometric characters see 
<tableCitation id="C6F00321370C7E4CFCB86A2BFC74EE1B" box="[863,977,272,298]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="5.[151,239,741,765]" captionStartId-1="6.[151,239,157,181]" captionTargetBox-0="[153,1441,955,2022]" captionTargetBox-1="[159,1420,372,1474]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." captionText-1="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri-0="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" httpUri-1="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="12" pageNumber="13" tableUuid-0="DF0D661237057E45FF7069DEFB78ECB9" tableUuid-1="DF0D661237067E46FF706BA6FDCFEFCB">Table 1­2</tableCitation>
.
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216A0CFD64EEC3" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
<emphasis id="B906EA88370C7E4CFF216A0CFC4DEE60" bold="true" box="[198,1000,311,337]" pageId="12" pageNumber="13">
<typeStatus id="54C98838370C7E4CFF216A0CFEE7EE60" box="[198,322,311,337]" pageId="12" pageNumber="13" type="lectotype">Lectotype</typeStatus>
designation of 
<taxonomicName id="4C724D19370C7E4CFDE06A0CFC46EE60" authority="Santschi, 1929" authorityName="Santschi" authorityYear="1929" box="[519,995,311,337]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370C7E4CFDE06A0CFC84EE60" bold="true" box="[519,801,311,337]" italics="true" pageId="12" pageNumber="13">Tetramorium alternans</emphasis>
<bibRefCitation id="EFE34B6B370C7E4CFCCC6A0CFC46EE60" author="Santschi" box="[811,995,311,337]" pageId="12" pageNumber="38" refString="Santschi, F. (1929) Fourmis du Maroc, d'Algerie et de Tunisie. Bulletin et Annales de la Societe Entomologique de Belge, 69, 138 - 165." type="journal article" year="1929">Santschi, 1929</bibRefCitation>
</taxonomicName>
.
</emphasis>
In order to avoid further nomenclatural problems it is necessary to designate a 
<typeStatus id="54C98838370C7E4CFD7D6A5BFCACEE4B" box="[666,777,352,378]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
. We investigated three 
<typeStatus id="54C98838370C7E4CFBFF6A5BFBD6EE4B" box="[1048,1139,352,378]" pageId="12" pageNumber="13" type="syntype">syntype</typeStatus>
workers mounted on one pin, labeled as: “ 
<typeStatus id="54C98838370C7E4CFEBC6AB3FE3EEE93" box="[347,411,392,418]" pageId="12" pageNumber="13">Type</typeStatus>
” [red label] [—] “Rabat” [
<collectingCountry id="F365760A370C7E4CFD316AB3FCE3EE93" box="[726,838,392,418]" name="Morocco" pageId="12" pageNumber="13">Morocco</collectingCountry>
] “Otin” [—] 1.quinz. 
<date id="FFCC105A370C7E4CFBAD6AB3FB0FEE93" box="[1098,1194,392,418]" pageId="12" pageNumber="13" value="1928-08">Aout 28</date>
[in original description: 04.1928] [—] “ 
<taxonomicName id="4C724D19370C7E4CFE6F6A8BFDBBEEFB" box="[392,542,432,458]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="caespitum">T. caespitum</taxonomicName>
<taxonomicName id="4C724D19370C7E4CFDC26A8BFD71EEFB" authority="Sant." authorityName="Sant." box="[549,724,432,458]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alternans">alternans Sant.</taxonomicName>
” SANTSCHI det. 19“29” [last two numbers are hardly readable] [—] Sammlung Dr. F. Santschi Kairouan;
</paragraph>
<paragraph id="8BCD369A370C7E4CFF21693BFE5BED5B" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
These data correspond to the original description (
<bibRefCitation id="EFE34B6B370C7E4CFCC5693BFBBEED2B" author="Santschi" box="[802,1051,512,538]" pageId="12" pageNumber="38" refString="Santschi, F. (1929) Fourmis du Maroc, d'Algerie et de Tunisie. Bulletin et Annales de la Societe Entomologique de Belge, 69, 138 - 165." type="journal article" year="1929">Santschi, 1929: 150.</bibRefCitation>
). The 
<typeStatus id="54C98838370C7E4CFB8E693BFB70ED2B" box="[1129,1237,512,538]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
is positioned on the distal end of the upper card (this is mentioned on the reverse side of 
<typeStatus id="54C98838370C7E4CFC0E6913FBF0ED73" box="[1001,1109,552,578]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
label). The 
<typeStatus id="54C98838370C7E4CFB046913FAEAED73" box="[1251,1359,552,578]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
specimen is in very good condition.
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216943FD7BEDD3" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
Morphometric data of 
<taxonomicName id="4C724D19370C7E4CFE336943FDE6EDA0" box="[468,579,632,657]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B906EA88370C7E4CFE336943FDE6EDA0" box="[468,579,632,657]" italics="true" pageId="12" pageNumber="13">alternans</emphasis>
</taxonomicName>
<typeStatus id="54C98838370C7E4CFDAB6943FD12EDA3" box="[588,695,632,658]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
worker: CL: 700; CW: 685; FR: 275; FL: 275; SL: 485; ML: 770; MW: 440; PEW: 230; PEH: 255; NOH: 145; NOL: 140; PEL: 150; PPW: 265; PPL: 165; PPH: 210; SPL: 90; SPSP: 140; EL: 135; EH: 100; ED: 180.
</paragraph>
<paragraph id="8BCD369A370C7E4CFF2169D4FE75ECE3" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
<emphasis id="B906EA88370C7E4CFF2169D4FBCEEC38" bold="true" box="[198,1131,751,777]" pageId="12" pageNumber="13">
<typeStatus id="54C98838370C7E4CFF2169D4FEE7EC38" box="[198,322,751,777]" pageId="12" pageNumber="13" type="lectotype">Lectotype</typeStatus>
designation of 
<taxonomicName id="4C724D19370C7E4CFDE869D4FBC2EC38" authority="Menozzi, 1934" authorityName="Menozzi" authorityYear="1934" box="[527,1127,751,777]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="biskrense" subSpecies="kahenae">
<emphasis id="B906EA88370C7E4CFDE869D4FC05EC38" bold="true" box="[527,928,751,777]" italics="true" pageId="12" pageNumber="13">Tetramorium biskrense kahenae</emphasis>
<bibRefCitation id="EFE34B6B370C7E4CFC4B69D4FBC2EC38" author="Menozzi" box="[940,1127,751,777]" pageId="12" pageNumber="38" refString="Menozzi, C. (1934) Reperti mirmecofaunistici raccolte dal Prof. L. di Caporiacco nelle oasi di Cufra e in altre localite del deserto Libico. Atti della Societa dei Naturalisti e Matematici di Modena, 13, 153 - 166." type="journal article" year="1934">Menozzi, 1934</bibRefCitation>
</taxonomicName>
.
</emphasis>
Santschi described this taxon based on several workers and gynes from different localities (Kairouan, Hammalif, Monastir, Pichon, Mines de Garn el Fyriat), and in order to prevent further nomenclatural problems it is necessary to designate a 
<typeStatus id="54C98838370C7E4CFF706853FEA3ECB3" box="[151,262,872,898]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
. Only one 
<typeStatus id="54C98838370C7E4CFE6F6853FE46ECB3" box="[392,483,872,898]" pageId="12" pageNumber="13" type="syntype">syntype</typeStatus>
worker specimen was investigated, labeled as: “ 
<taxonomicName id="4C724D19370C7E4CFBC36853FB1AECB3" box="[1060,1215,872,898]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="caespitum">T. caespitum</taxonomicName>
” [/] “st. 
<taxonomicName id="4C724D19370C7E4CFAC16853FA3CECB3" box="[1318,1433,872,898]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="biskrense">Biskrense</taxonomicName>
For” [/] ìv. 
<taxonomicName id="4C724D19370C7E4CFEF868ABFE23EC9B" box="[287,390,912,938]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="31" phylum="Arthropoda" rank="species" species="kahenae">Kahenae</taxonomicName>
Santì [/] “Garn el Fayah” [now Garn Alfaya mines] [/] “ 
<collectingCountry id="F365760A370C7E4CFBCB68ABFB3DEC9B" box="[1068,1176,912,938]" name="Tunisia" pageId="12" pageNumber="13">Tunisien</collectingCountry>
” [/] “(Santschi)” [this label written with red ink].
</paragraph>
<paragraph id="8BCD369A370C7E4CFF2168DBFA8DEB13" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
These data correspond to the original description (
<bibRefCitation id="EFE34B6B370C7E4CFCF468DBFBA2ECCB" author="Santschi" box="[787,1031,992,1018]" pageId="12" pageNumber="38" refString="Santschi, F. (1918) Noveaux Tetramorium africains. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord, 9, 153 - 156." type="journal article" year="1918">Santschi, 1918: 155.</bibRefCitation>
). In the present designation we fix this specimen as the 
<typeStatus id="54C98838370C7E4CFE6E6F33FE5CEB13" box="[393,505,1032,1058]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
. The 
<typeStatus id="54C98838370C7E4CFDDF6F33FD06EB13" box="[568,675,1032,1058]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
specimen is in good condition, partly covered by glue.
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216F0BFCA3EB7B" blockId="12.[151,1439,152,1258]" box="[198,774,1072,1098]" pageId="12" pageNumber="13">
Morphometric data of 
<taxonomicName id="4C724D19370C7E4CFE286F0BFD94EB78" box="[463,561,1072,1097]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kahenae">
<emphasis id="B906EA88370C7E4CFE286F0BFD94EB78" box="[463,561,1072,1097]" italics="true" pageId="12" pageNumber="13">kahenae</emphasis>
</taxonomicName>
<typeStatus id="54C98838370C7E4CFDDF6F0BFD06EB7B" box="[568,675,1072,1098]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
worker:
</paragraph>
<paragraph id="8BCD369A370C7E4CFF216F63FAD8EBAB" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">CL: 790; CW: 790; FR: 305; FL: 305; SL: 555; ML: 880; MW: 485; PEW: 240; PEH: 285; NOH: 170; NOL: 165; PEL: 175; PPW: 305; PPL: 185; PPH: 265; SPL: 110; SPSP: 135; EL: 165; EH: 115; ED: 200.</paragraph>
</subSubSection>
<subSubSection id="C3686511370C7E4CFF216F9CFAA2EBDB" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BCD369A370C7E4CFF216F9CFAA2EBDB" blockId="12.[151,1439,152,1258]" pageId="12" pageNumber="13">
<emphasis id="B906EA88370C7E4CFF216F9CFEC2EBF0" bold="true" box="[198,359,1191,1217]" pageId="12" pageNumber="13">Distribution.</emphasis>
This species seems to be endemic to the Mediterranean coast of NW Africa. It is reported from 
<collectingCountry id="F365760A370C7E4CFF306FEBFEE1EBDB" box="[215,324,1232,1258]" name="Morocco" pageId="12" pageNumber="13">Morocco</collectingCountry>
, 
<collectingCountry id="F365760A370C7E4CFEA86FEBFE0FEBDB" box="[335,426,1232,1258]" name="Algeria" pageId="12" pageNumber="13">Algeria</collectingCountry>
, 
<collectingCountry id="F365760A370C7E4CFE516FEBFDA8EBDB" box="[438,525,1232,1258]" name="Tunisia" pageId="12" pageNumber="13">Tunisia</collectingCountry>
and a dubious record from Lampedusa (Aria Rossa) (
<bibRefCitation id="EFE34B6B370C7E4CFB646FEBFB5FEBDB" author="Mei" box="[1155,1274,1232,1258]" lastPageId="12" lastPageNumber="38" pageId="12" pageNumber="37" refString="Mei, M. (1995) Arthropoda di Lampedusa, Linosa e Pantelleria (Canale di Sicilia, Mar Mediterraneo). Hymenoptera, Formicidae (con diagnosi di due nuove specie). Naturalista Siciliano, 19 (Suppl.), 753 - 772." type="journal article" year="1995">Mei 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3686511370C7E4CFF706E06FC42EA66" box="[151,999,1341,1367]" pageId="12" pageNumber="13" type="key">
<paragraph id="8BCD369A370C7E4CFF706E06FC42EA66" blockId="12.[151,999,1341,1367]" box="[151,999,1341,1367]" pageId="12" pageNumber="13">
<keyStep id="30862E3F370C7E4CFF706E06FC42EA66" box="[151,999,1341,1367]" pageId="12" pageNumber="13">
<key id="FFE361AB370C7E4CFF706E06FC42EA66" box="[151,999,1341,1367]" pageId="12" pageNumber="13">
<keyLead id="308395AF370C7E4CFF706E06FC42EA66" box="[151,999,1341,1367]" pageId="12" pageNumber="13">
<emphasis id="B906EA88370C7E4CFF706E06FDAEEA66" bold="true" box="[151,523,1341,1367]" pageId="12" pageNumber="13">
2. 
<taxonomicName id="4C724D19370C7E4CFF536E06FDAEEA66" authority="Cs" authorityName="Cs" box="[180,523,1341,1367]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anatolicum">
<emphasis id="B906EA88370C7E4CFF536E06FE46EA66" bold="true" box="[180,483,1341,1367]" italics="true" pageId="12" pageNumber="13">Tetramorium anatolicum</emphasis>
Cs
</taxonomicName>
</emphasis>
õ 
<emphasis id="B906EA88370C7E4CFDFE6E06FC42EA66" bold="true" box="[537,999,1341,1367]" pageId="12" pageNumber="13">sz &amp; Schulz, New species (figs 18–24)</emphasis>
</keyLead>
</key>
</keyStep>
</paragraph>
</subSubSection>
<subSubSection id="C3686511370C7E4EFF706E46FB36EDD6" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="8BCD369A370C7E4CFF706E46FE81E8C3" blockId="12.[151,1440,1405,2034]" pageId="12" pageNumber="13">
<emphasis id="B906EA88370C7E4CFF706E46FE10EAA6" bold="true" box="[151,437,1405,1431]" pageId="12" pageNumber="13">Description of Worker</emphasis>
(figs 21–24). Morphometric data of 
<typeStatus id="54C98838370C7E4CFC896E46FC70EAA6" box="[878,981,1405,1431]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
worker: CL: 720; CW: 695; FR: 245; FL: 265; SL: 550; ML: 770; MW: 430; PEW: 200; PEH: 240; NOH: 140; NOL: 115; PEL: 155; PPW: 250; PPL: 155; PPH: 220; SPL: 80; SPSP: 140; EL: 145; EH: 100; ED: 190; Small size, CS 702 [635, 743]. Whole body and appendages light brown to brown. Head slightly longer than broad, CL/CW 1.02 [0.99, 1.05], with very feebly convex or straight sides and occipital margin, and rounded occipital corners. Eyes small, EYE 0.176 [0.167, 0.182]. Frons moderately narrow, FR/CS 0.37 [0.35, 0.39], frontal lobes usually wider, FL/FR 1.02 [1.0, 1.08]. Scape moderately long, SL/CS 0.78 [0.76, 0.81], without longitudinal dorsal carina basally, smooth and shiny. Promesonotal dorsum convex, metanotal groove shallow, but visible. Propodeal teeth moderately long, thin and acute. Petiolar node trapezoidal in profile, NOH/NOL 1.15 [1.0, 1.30], petiole relatively high, PEH/NOL 2.01 [1.74, 2.30]. General appearance finely rugose, or rugulose. Head dorsum longitudinally rugulose, ground surface feebly microreticulate, shiny. Alitrunk dorsum longitudinally rugulose and finely microreticulate. Mesopleuron usually feebly rugulose and microreticulate. Dorsum of petiolar node rugulosoreticulate and microreticulate, dorsum of postpetiole feebly rugulose and microreticulate. Polygonal striation usually continuous on 1st gastral tergite, sometimes slightly disrupted posteriorly (see fig. 8.). Ventral surface of head with several short and few moderately long, straight, or few C­shape setae posteriorly to buccal cavity (see fig. 5.).
</paragraph>
<caption id="DF0D6612370D7E4DFF706D44FD5AE9ED" httpUri="https://zenodo.org/record/399641/files/figure.png" pageId="13" pageNumber="14" targetBox="[264,1321,197,1634]" targetPageId="13">
<paragraph id="8BCD369A370D7E4DFF706D44FD5AE9ED" blockId="13.[151,1437,1663,1756]" pageId="13" pageNumber="14">
<emphasis id="B906EA88370D7E4DFF706D44FEFAE9A6" bold="true" box="[151,351,1663,1687]" pageId="13" pageNumber="14">FIGURES 18–24.</emphasis>
<taxonomicName id="4C724D19370D7E4DFE8F6DBBFCC1E9A6" authority="Csosz &amp; Schulz New" authorityName="Csosz &amp; Schulz New" box="[360,868,1664,1687]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="anatolicum">
<emphasis id="B906EA88370D7E4DFE8F6DBBFDD7E9A6" box="[360,626,1664,1687]" italics="true" pageId="13" pageNumber="14">Tetramorium anatolicum</emphasis>
Csösz &amp; Schulz 
<taxonomicNameLabel id="A23557F3370D7E4DFCD66DBBFCC1E9A6" box="[817,868,1664,1687]" pageId="13" pageNumber="14" rank="species">New</taxonomicNameLabel>
</taxonomicName>
species. Gyne: alitrunk petiole and postpetiole, Fig. 18. dorsal view, Fig. 19. lateral view, Fig. 20. head. Worker: Fig. 21. head. Alitrunk petiole and postpetiole, Fig. 22. dorsal view, Fig. 23. lateral view, Fig. 24. scape, dorsal view.
</paragraph>
</caption>
<paragraph id="8BCD369A370D7E4EFF216C34FD42EEFD" blockId="13.[151,1436,1807,2034]" lastBlockId="14.[151,1438,152,1869]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B906EA88370D7E4DFF216C34FE64E818" bold="true" box="[198,449,1807,1833]" pageId="13" pageNumber="14">Description of Gyne</emphasis>
(figs 18–20). Medium to small size, CS 1012 [980, 1055]. Whole body and appendages dark brown to black. Head wider than long, CL/CW 0.93 [0.90, 0.98], with feebly convex sides, straight occipital margin and rounded occipital corners. Frons moderately narrow, FR/CS 0.37 [0.36, 0.40], frontal lobes as wide as frons, or slightly wider, FL/FR 1.01 [1.0, 1.03]. Scape moderately long, SL/CS 0.75 [0.71, 0.77], without longitudinal dorsal carina basally, smooth and shiny. Head wider than scutum, MW/CS 0.96 [0.92, 1.01]. Propodeal teeth moderately long. Dorsal crest of petiolar node in frontal view straight. Petiolar node dorsum steeply rounded backward. Petiole and postpetiole relatively narrow, WAIST 0.85 [0.83, 0.87]. General appearance rugulose, ground surface microreticulate, dull. Head dorsum, occiput and sides rugosoreticulate, ground surface microreticulate. Frons longitudinally rugulose and microreticulate. Scutum longitudinally rugose, anteriorly smooth, scutellum medially more or less smooth; laterally finely rugulose. Sides of alitrunk ruguloso­reticulate and microreticulate, katepisternum smooth and shiny ventrally. Dorsum of petiolar node and postpetiole reticulate, petiolar node smooth medially. Polygonal striation disrupted on 1st gastral tergite, sometimes continuous basally. Ventral surface of head with several short and few longer, straight or “C”­shape setae, arising posterior to buccal cavity.
</paragraph>
<paragraph id="8BCD369A370E7E4EFF216AE1FB36EDD6" blockId="14.[151,1438,152,1869]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF216AE1FE63EEC5" bold="true" box="[198,454,474,500]" pageId="14" pageNumber="15">Description of Male.</emphasis>
Whole body and appendages brownish black. ead with convex sides, round occipital margin and widely rounded occipital corners. Scutum wider than head. Propodeal teeth very short, propodeum nearly rounded in profile. Dorsal crest of petiolar node with sharp transversal edge, slightly emarginated in frontal view. Head, alitrunk and waist rugulose, ground surface reticulate to microreticulate, dull. Scutum finely rugulose, antero­laterally smooth and shiny. Scutellum transversally rugulose and microreticulate. Sides of alitrunk finely rugose and microreticulate. Dorsum of petiolar node finely reticulate and microreticulate. Postpetiole shiny, feebly striate. Polygonal striation disrupted on 1st gastral tergite.
</paragraph>
</subSubSection>
<subSubSection id="C3686511370E7E4EFF2169CEFCE6EB49" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BCD369A370E7E4EFF2169CEFC42EB7E" blockId="14.[151,1438,152,1869]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF2169CEFE11EC3E" bold="true" box="[198,436,757,783]" pageId="14" pageNumber="15">Material examined</emphasis>
: (6 nest series including 
<specimenCount id="9D74FD13370E7E4EFD3E69CEFCFAEC3E" box="[729,863,757,783]" pageId="14" pageNumber="15" type="worker">52 workers</specimenCount>
, 3 gynes and 
<specimenCount id="9D74FD13370E7E4EFC1869CEFBF5EC3E" box="[1023,1104,757,783]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
) 
<typeStatus id="54C98838370E7E4EFBB869CEFB5FEC3E" box="[1119,1274,757,783]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
/: 
<collectingCountry id="F365760A370E7E4EFAF969CEFA3EEC3E" box="[1310,1435,757,783]" name="Turkey" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFAF969CEFA3EEC3E" bold="true" box="[1310,1435,757,783]" pageId="14" pageNumber="15">TURKEY</emphasis>
</collectingCountry>
– Erzurum, 
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SW Aydogdu 
<quantity id="4C8A9B7F370E7E4EFDFF6826FDFFEC06" box="[536,602,797,823]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="km" value="20.0">20km</quantity>
SW Göle, 1400 mH nr.1148 leg. Schulz 
<date id="FFCC105A370E7E4EFBBB6826FB45EC06" box="[1116,1248,797,823]" pageId="14" pageNumber="15" value="1993-06-26">26.06.1993</date>
(1/ / 
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); 
<typeStatus id="54C98838370E7E4EFF70687EFE9BEC6E" box="[151,318,837,863]" pageId="14" pageNumber="15" type="paratype">PARATYPES</typeStatus>
: 
<collectingCountry id="F365760A370E7E4EFEAC687EFE6DEC6E" box="[331,456,837,863]" name="Turkey" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFEAC687EFE6DEC6E" bold="true" box="[331,456,837,863]" pageId="14" pageNumber="15">TURKEY</emphasis>
</collectingCountry>
– Digor 1650 mH 
<date id="FFCC105A370E7E4EFD54687EFC92EC6E" box="[691,823,837,863]" pageId="14" pageNumber="15" value="1991-06-13">13.06.1991</date>
leg. Lˆbl (4/ / 
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, 4/, 1Ψ 
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); Erzurum, 
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SW Aydogdu 
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SW Göle, 1400 mH nr.1148 leg. Schulz 
<date id="FFCC105A370E7E4EFC2B6856FBEAECB6" box="[972,1103,877,903]" pageId="14" pageNumber="15" value="1993-06-26">26.06.1993</date>
(5/, 1ɗ / 
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, 2/, 1Ψ, 
<collectionCode id="ED63AE5F370E7E4EFF7068AEFF40EC9E" box="[151,229,917,943]" pageId="14" pageNumber="15">PCAS</collectionCode>
); Van, 
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SE Dedeli 
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SE Patnos 1700mH, 
<emphasis id="B906EA88370E7E4EFCA868ADFC7CEC9E" box="[847,985,918,943]" italics="true" pageId="14" pageNumber="15">Hochsteppe</emphasis>
<date id="FFCC105A370E7E4EFC0268AEFBC9EC9E" box="[997,1132,917,943]" pageId="14" pageNumber="15" value="1993-06-20">20.06.1993</date>
. nr.1104. leg. Schulz (3/ 
<collectionCode id="ED63AE5F370E7E4EFF706886FF40ECE6" box="[151,229,957,983]" pageId="14" pageNumber="15">PCAS</collectionCode>
); Van­ 
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Van, 
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SE Dedeli 
<quantity id="4C8A9B7F370E7E4EFD646886FD63ECE6" box="[643,710,957,983]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="km" value="30.0">30km</quantity>
SE Patnos 1700mH, Hochsteppe 
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. nr.1102. leg. Schulz (9/ 
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, 6/ 
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); Van, 
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SE Dedeli, 
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SE Patnos, 1700 mH 
<emphasis id="B906EA88370E7E4EFB9768DDFB5FECCE" box="[1136,1274,998,1023]" italics="true" pageId="14" pageNumber="15">Hochsteppe</emphasis>
nr. 1104 leg. Schulz 
<date id="FFCC105A370E7E4EFF0A6F36FED4EB16" box="[237,369,1037,1063]" pageId="14" pageNumber="15" value="1993-06-20">20.06.1993</date>
(6/ / 
<collectionCode id="ED63AE5F370E7E4EFE546F36FDB5EB16" LSID="urn:lsid:biocol.org:col:34991" box="[435,528,1037,1063]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="15">HNHM</collectionCode>
, 3/ / 
<collectionCode id="ED63AE5F370E7E4EFDAA6F36FD39EB16" box="[589,668,1037,1063]" pageId="14" pageNumber="15">PCAS</collectionCode>
); Van, 
<quantity id="4C8A9B7F370E7E4EFD0C6F36FCBAEB16" box="[747,799,1037,1063]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="km" value="5.0">5km</quantity>
SE Dedeli, 
<quantity id="4C8A9B7F370E7E4EFC4D6F36FC4BEB16" box="[938,1006,1037,1063]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="km" value="30.0">30km</quantity>
SE Patnos, 1700 mH 
<emphasis id="B906EA88370E7E4EFB0A6F35FAD2EB16" box="[1261,1399,1038,1063]" italics="true" pageId="14" pageNumber="15">Hochsteppe</emphasis>
nr. 1106 leg. Schulz 
<date id="FFCC105A370E7E4EFE866F0EFE40EB7E" box="[353,485,1077,1103]" pageId="14" pageNumber="15" value="1993-06-20">20.06.1993</date>
(3/, 1Ψ / 
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, 6/ / 
<collectionCode id="ED63AE5F370E7E4EFD126F0EFCE4EB7E" box="[757,833,1077,1103]" pageId="14" pageNumber="15">PCAS</collectionCode>
, 3/ / 
<collectionCode id="ED63AE5F370E7E4EFC666F0EFC7EEB7E" box="[897,987,1077,1103]" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="14" pageNumber="15">SMNK</collectionCode>
).
</paragraph>
<paragraph id="8BCD369A370E7E4EFF216F66FCE6EB49" blockId="14.[151,1438,152,1869]" box="[198,835,1117,1144]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF216F66FE36EB46" bold="true" box="[198,403,1117,1143]" pageId="14" pageNumber="15">Morphometrics:</emphasis>
(
<specimenCount id="9D74FD13370E7E4EFE446F65FD82EB49" box="[419,551,1118,1144]" pageId="14" pageNumber="15" type="worker">38 workers</specimenCount>
and 3 gynes measured).
</paragraph>
</subSubSection>
<subSubSection id="C3686511370E7E4EFF216FBEFC7FE9E4" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BCD369A370E7E4EFF216FBEFD80E904" blockId="14.[151,1438,152,1869]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF216FBEFEE3EBAE" bold="true" box="[198,326,1157,1183]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
Workers of 
<taxonomicName id="4C724D19370E7E4EFE306FBDFDDCEBAE" box="[471,633,1158,1183]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFE306FBDFDDCEBAE" box="[471,633,1158,1183]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFD676FBEFD67EBAE" box="[640,706,1157,1183]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFD676FBEFD67EBAE" bold="true" box="[640,706,1157,1183]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
can be separated from related species by the lack of psammophore, relatively small eyes, (EYE, 
<tableCitation id="C6F00321370E7E4EFDDC6F95FD34EBF9" box="[571,657,1198,1224]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="14" pageNumber="15" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.), moderately long and smooth scape (SL/CS, 
<tableCitation id="C6F00321370E7E4EFB5E6F95FAAAEBF9" box="[1209,1295,1198,1224]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="14" pageNumber="15" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.), without a dorsal carina basally, relatively fine and parallel body sculpture and trapezoidal petiolar node (NOH/NOL and PEH/NOL, 
<tableCitation id="C6F00321370E7E4EFEC66FC5FEDDEA29" box="[289,376,1278,1304]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="14" pageNumber="15" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.). Workers of 
<taxonomicName id="4C724D19370E7E4EFDF96FC5FD1BEA26" box="[542,702,1278,1303]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFDF96FC5FD1BEA26" box="[542,702,1278,1303]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<emphasis id="B906EA88370E7E4EFD226FC6FCA2EA26" bold="true" box="[709,775,1277,1303]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A23557F3370E7E4EFD226FC6FCA2EA26" box="[709,775,1277,1303]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are mostly similar to 
<taxonomicName id="4C724D19370E7E4EFBE36FC5FBF0EA26" box="[1028,1109,1278,1303]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="exile" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFBE36FC5FBF0EA26" box="[1028,1109,1278,1303]" italics="true" pageId="14" pageNumber="15">T. exile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFBB96FC6FB05EA26" box="[1118,1184,1277,1303]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFBB96FC6FB05EA26" bold="true" box="[1118,1184,1277,1303]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
, but differs by its relatively shorter scape and somewhat wider frons (SL/CS and FR/CS, 
<tableCitation id="C6F00321370E7E4EFC336E1DFB8BEA71" box="[980,1070,1318,1344]" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="14" pageNumber="15" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.). Discriminant D(2a) function proves the separation between 
<taxonomicName id="4C724D19370E7E4EFDE56E6AFD06EA5B" box="[514,675,1361,1386]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFDE56E6AFD06EA5B" box="[514,675,1361,1386]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFD4C6E6BFD4BEA5B" box="[683,750,1360,1386]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFD4C6E6BFD4BEA5B" bold="true" box="[683,750,1360,1386]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C724D19370E7E4EFCC06E6AFCD9EA5B" box="[807,892,1361,1386]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="exile" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFCC06E6AFCD9EA5B" box="[807,892,1361,1386]" italics="true" pageId="14" pageNumber="15">T. exile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFC636E6BFC63EA5B" box="[900,966,1360,1386]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFC636E6BFC63EA5B" bold="true" box="[900,966,1360,1386]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
(see differential diagnosis of 
<taxonomicName id="4C724D19370E7E4EFAC26E6AFAD9EA5B" box="[1317,1404,1361,1386]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="exile" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFAC26E6AFAD9EA5B" box="[1317,1404,1361,1386]" italics="true" pageId="14" pageNumber="15">T.exile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFA636E6BFF1FEAA3" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFA636E6BFF1FEAA3" bold="true" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
). Workers of 
<taxonomicName id="4C724D19370E7E4EFEBC6E42FE5EEAA3" box="[347,507,1401,1426]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFEBC6E42FE5EEAA3" box="[347,507,1401,1426]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<emphasis id="B906EA88370E7E4EFDE56E43FDE1EAA3" bold="true" box="[514,580,1400,1426]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A23557F3370E7E4EFDE56E43FDE1EAA3" box="[514,580,1400,1426]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
may look similar to those of 
<taxonomicName id="4C724D19370E7E4EFC7D6E42FBB4EAA3" box="[922,1041,1401,1426]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chefketi">
<emphasis id="B906EA88370E7E4EFC7D6E42FBB4EAA3" box="[922,1041,1401,1426]" italics="true" pageId="14" pageNumber="15">T. chefketi</emphasis>
</taxonomicName>
, but can be distinguished by their lighter colour and shape of petiolar node: in 
<taxonomicName id="4C724D19370E7E4EFD456E9AFCE6EA8B" box="[674,835,1441,1466]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFD456E9AFCE6EA8B" box="[674,835,1441,1466]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<emphasis id="B906EA88370E7E4EFCAD6E9BFC28EA8B" bold="true" box="[842,909,1440,1466]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A23557F3370E7E4EFCAD6E9BFC28EA8B" box="[842,909,1440,1466]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
it is relatively high and short, trapezoidal, in profile, while in 
<taxonomicName id="4C724D19370E7E4EFEBB6EF2FE71EAD3" box="[348,468,1481,1506]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chefketi">
<emphasis id="B906EA88370E7E4EFEBB6EF2FE71EAD3" box="[348,468,1481,1506]" italics="true" pageId="14" pageNumber="15">T. chefketi</emphasis>
</taxonomicName>
it is relatively low and longer, cubic in profile, (PEH/NOL and NOH/NOL, 
<tableCitation id="C6F00321370E7E4EFABC6EF2FF03E93A" captionStart="TABLE 1" captionStartId="5.[151,239,741,765]" captionTargetBox="[153,1441,955,2022]" captionTargetPageId="5" captionText="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." httpUri="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" pageId="14" pageNumber="15" tableUuid="DF0D661237057E45FF7069DEFB78ECB9">Table 1</tableCitation>
.). Discriminant D(3a) function proves the separation between 
<taxonomicName id="4C724D19370E7E4EFC9C6ECAFBB9E93B" box="[891,1052,1521,1546]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFC9C6ECAFBB9E93B" box="[891,1052,1521,1546]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFBC46ECBFBC0E93B" box="[1059,1125,1520,1546]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFBC46ECBFBC0E93B" bold="true" box="[1059,1125,1520,1546]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C724D19370E7E4EFB7A6ECAFAB1E93B" box="[1181,1300,1521,1546]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chefketi">
<emphasis id="B906EA88370E7E4EFB7A6ECAFAB1E93B" box="[1181,1300,1521,1546]" italics="true" pageId="14" pageNumber="15">T. chefketi</emphasis>
</taxonomicName>
(see differential diagnosis of 
<taxonomicName id="4C724D19370E7E4EFE926D27FE6EE904" box="[373,459,1564,1589]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="exile" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFE926D27FE6EE904" box="[373,459,1564,1589]" italics="true" pageId="14" pageNumber="15">T.exile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFE346D20FDB0E904" box="[467,533,1563,1589]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFE346D20FDB0E904" bold="true" box="[467,533,1563,1589]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BCD369A370E7E4EFF216D78FBBDE99C" blockId="14.[151,1438,152,1869]" pageId="14" pageNumber="15">
Gynes of 
<taxonomicName id="4C724D19370E7E4EFEDF6D7FFE7DE96C" box="[312,472,1604,1629]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum" status="sp. nov.">
<emphasis id="B906EA88370E7E4EFEDF6D7FFE7DE96C" box="[312,472,1604,1629]" italics="true" pageId="14" pageNumber="15">T. anatolicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F3370E7E4EFE386D78FD84E96C" box="[479,545,1603,1629]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B906EA88370E7E4EFE386D78FD84E96C" bold="true" box="[479,545,1603,1629]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished by the lack of psammophore, moderately long, smooth scape (SL/CS, 
<tableCitation id="C6F00321370E7E4EFEAE6D50FE00E9B4" box="[329,421,1643,1669]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="14" pageNumber="15" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), without a dorsal carina basally, wide scutum (MW/CS, 
<tableCitation id="C6F00321370E7E4EFBBE6D50FB11E9B4" box="[1113,1204,1643,1669]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="14" pageNumber="15" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), relatively narrow petiole and postpetiole (WAIST, 
<tableCitation id="C6F00321370E7E4EFDFF6DA8FDD7E99C" box="[536,626,1683,1709]" captionStart="TABLE 2" captionStartId="6.[151,239,157,181]" captionTargetBox="[159,1420,372,1474]" captionTargetPageId="6" captionText="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="14" pageNumber="15" tableUuid="DF0D661237067E46FF706BA6FDCFEFCB">Table 2</tableCitation>
.), and partly smooth katepisternum.
</paragraph>
<paragraph id="8BCD369A370E7E4EFF216D80FC7FE9E4" blockId="14.[151,1438,152,1869]" box="[198,986,1723,1749]" pageId="14" pageNumber="15">
For further combination of morphometric characters see 
<tableCitation id="C6F00321370E7E4EFCB86D80FC73E9E4" box="[863,982,1723,1749]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="5.[151,239,741,765]" captionStartId-1="6.[151,239,157,181]" captionTargetBox-0="[153,1441,955,2022]" captionTargetBox-1="[159,1420,372,1474]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="TABLE 1. Morphometric comparison (mm) of the Tetramorium workers. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens, EYE *: following number of individuals were measured for the EYE character: T. chefketi (n = 18), T. exile (n = 30), T. moravicum (n = 18), T. anatolicum (n = 14), T. laurae (n = 15), T. alternans (n = 20), T. syriacum (n = 7), T. rhodium (n = 17), T. sulcinode (n = 13), T. annectens (n = 8), T. forte (n = 14)." captionText-1="TABLE 2. Morphometric comparison (mm) of the Tetramorium gynes. Abbreviations of morphometric characters in Material and Methods. Upper line: arithmetic mean ± standard deviation, lower line, in []: minimum and maximum values, n = number of measured specimens." httpUri-0="http://table.plazi.org/id/DF0D661237057E45FF7069DEFB78ECB9" httpUri-1="http://table.plazi.org/id/DF0D661237067E46FF706BA6FDCFEFCB" pageId="14" pageNumber="15" tableUuid-0="DF0D661237057E45FF7069DEFB78ECB9" tableUuid-1="DF0D661237067E46FF706BA6FDCFEFCB">Table 1–2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3686511370E7E4EFF216DD8FD6AE9CC" box="[198,719,1763,1789]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BCD369A370E7E4EFF216DD8FD6AE9CC" blockId="14.[151,1438,152,1869]" box="[198,719,1763,1789]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF216DD8FEC2E9CC" bold="true" box="[198,359,1763,1789]" pageId="14" pageNumber="15">Distribution.</emphasis>
It is known only from 
<collectingCountry id="F365760A370E7E4EFD926DD8FD6EE9CC" box="[629,715,1763,1789]" name="Turkey" pageId="14" pageNumber="15">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3686511370E7E4EFF216C30FE2AE87C" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BCD369A370E7E4EFF216C30FE2AE87C" blockId="14.[151,1438,152,1869]" pageId="14" pageNumber="15">
<emphasis id="B906EA88370E7E4EFF216C30FEF6E814" bold="true" box="[198,339,1803,1829]" pageId="14" pageNumber="15">Etymology.</emphasis>
This adjective [
<taxonomicName id="4C724D19370E7E4EFDE86C37FD36E814" box="[527,659,1804,1829]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="anatolicum">
<emphasis id="B906EA88370E7E4EFDE86C37FD36E814" box="[527,659,1804,1829]" italics="true" pageId="14" pageNumber="15">anatolicum</emphasis>
</taxonomicName>
(neutrum)] refers to the known distribution of this species in Anatolian part of 
<collectingCountry id="F365760A370E7E4EFED26C08FE2EE87C" box="[309,395,1843,1869]" name="Turkey" pageId="14" pageNumber="15">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>